taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
038987A0FFF54B09B384A65C1056FA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFF54B09B384A65C1056FA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399708/files/figure.png	https://doi.org/10.5281/zenodo.7399708	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFF54B09B384A65C1056FA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFF54B09B384A65C1056FA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399720/files/figure.png	https://doi.org/10.5281/zenodo.7399720	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFF54B09B384A65C1056FA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399728/files/figure.png	https://doi.org/10.5281/zenodo.7399728	Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFF54B09B384A65C1056FA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399744/files/figure.png	https://doi.org/10.5281/zenodo.7399744	Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom).	Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFEA4B0EB38AA5B816FBFC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFEA4B0EB38AA5B816FBFC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399708/files/figure.png	https://doi.org/10.5281/zenodo.7399708	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFEA4B0EB38AA5B816FBFC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFEA4B0EB38AA5B816FBFC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399720/files/figure.png	https://doi.org/10.5281/zenodo.7399720	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFEA4B0EB38AA5B816FBFC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399728/files/figure.png	https://doi.org/10.5281/zenodo.7399728	Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFEA4B0EB38AA5B816FBFC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399730/files/figure.png	https://doi.org/10.5281/zenodo.7399730	Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange).	Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFEA4B0EB38AA5B816FBFC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399744/files/figure.png	https://doi.org/10.5281/zenodo.7399744	Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom).	Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFED4B02B391A418119CFEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFED4B02B391A418119CFEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399706/files/figure.png	https://doi.org/10.5281/zenodo.7399706	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFED4B02B391A418119CFEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399708/files/figure.png	https://doi.org/10.5281/zenodo.7399708	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFED4B02B391A418119CFEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFED4B02B391A418119CFEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399720/files/figure.png	https://doi.org/10.5281/zenodo.7399720	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFED4B02B391A418119CFEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399728/files/figure.png	https://doi.org/10.5281/zenodo.7399728	Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFED4B02B391A418119CFEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399730/files/figure.png	https://doi.org/10.5281/zenodo.7399730	Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange).	Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFED4B02B391A418119CFEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399744/files/figure.png	https://doi.org/10.5281/zenodo.7399744	Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom).	Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE14B07B3AFA19817ADFF6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE14B07B3AFA19817ADFF6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399706/files/figure.png	https://doi.org/10.5281/zenodo.7399706	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE14B07B3AFA19817ADFF6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399708/files/figure.png	https://doi.org/10.5281/zenodo.7399708	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE14B07B3AFA19817ADFF6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE14B07B3AFA19817ADFF6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399720/files/figure.png	https://doi.org/10.5281/zenodo.7399720	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE14B07B3AFA19817ADFF6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399728/files/figure.png	https://doi.org/10.5281/zenodo.7399728	Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE14B07B3AFA19817ADFF6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399746/files/figure.png	https://doi.org/10.5281/zenodo.7399746	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE44B07B139A11816E5FECA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE44B07B139A11816E5FECA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399722/files/figure.png	https://doi.org/10.5281/zenodo.7399722	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE44B07B139A11816E5FECA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399748/files/figure.png	https://doi.org/10.5281/zenodo.7399748	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE44B07B139A11816E5FECA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399732/files/figure.png	https://doi.org/10.5281/zenodo.7399732	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE44B07B139A11816E5FECA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399746/files/figure.png	https://doi.org/10.5281/zenodo.7399746	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE44B39B3DEA17816FDFA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE44B39B3DEA17816FDFA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE44B39B3DEA17816FDFA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399722/files/figure.png	https://doi.org/10.5281/zenodo.7399722	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE44B39B3DEA17816FDFA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399732/files/figure.png	https://doi.org/10.5281/zenodo.7399732	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE44B39B3DEA17816FDFA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399746/files/figure.png	https://doi.org/10.5281/zenodo.7399746	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFE44B39B3DEA17816FDFA0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDA4B3BB36CA5B114E4F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDA4B3BB36CA5B114E4F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399706/files/figure.png	https://doi.org/10.5281/zenodo.7399706	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDA4B3BB36CA5B114E4F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDA4B3BB36CA5B114E4F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399722/files/figure.png	https://doi.org/10.5281/zenodo.7399722	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDA4B3BB36CA5B114E4F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399732/files/figure.png	https://doi.org/10.5281/zenodo.7399732	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDA4B3BB36CA5B114E4F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399748/files/figure.png	https://doi.org/10.5281/zenodo.7399748	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDA4B3BB36CA5B114E4F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD84B3DB392A0B917E2F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD84B3DB392A0B917E2F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD84B3DB392A0B917E2F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399724/files/figure.png	https://doi.org/10.5281/zenodo.7399724	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD84B3DB392A0B917E2F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399722/files/figure.png	https://doi.org/10.5281/zenodo.7399722	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD84B3DB392A0B917E2F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399730/files/figure.png	https://doi.org/10.5281/zenodo.7399730	Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange).	Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD84B3DB392A0B917E2F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399732/files/figure.png	https://doi.org/10.5281/zenodo.7399732	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD84B3DB392A0B917E2F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399748/files/figure.png	https://doi.org/10.5281/zenodo.7399748	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD84B3DB392A0B917E2F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDE4B3CB11AA7781113FACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDE4B3CB11AA7781113FACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399706/files/figure.png	https://doi.org/10.5281/zenodo.7399706	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDE4B3CB11AA7781113FACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDE4B3CB11AA7781113FACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399720/files/figure.png	https://doi.org/10.5281/zenodo.7399720	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDE4B3CB11AA7781113FACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399732/files/figure.png	https://doi.org/10.5281/zenodo.7399732	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDE4B3CB11AA7781113FACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399748/files/figure.png	https://doi.org/10.5281/zenodo.7399748	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDE4B3CB11AA7781113FACA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDF4B31B3BDA57817D6FA8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDF4B31B3BDA57817D6FA8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDF4B31B3BDA57817D6FA8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399720/files/figure.png	https://doi.org/10.5281/zenodo.7399720	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDF4B31B3BDA57817D6FA8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399722/files/figure.png	https://doi.org/10.5281/zenodo.7399722	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDF4B31B3BDA57817D6FA8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399732/files/figure.png	https://doi.org/10.5281/zenodo.7399732	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDF4B31B3BDA57817D6FA8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399736/files/figure.png	https://doi.org/10.5281/zenodo.7399736	Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens.	Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDF4B31B3BDA57817D6FA8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399748/files/figure.png	https://doi.org/10.5281/zenodo.7399748	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFDF4B31B3BDA57817D6FA8A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD24B30B0EFA5381011FEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD24B30B0EFA5381011FEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399706/files/figure.png	https://doi.org/10.5281/zenodo.7399706	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD24B30B0EFA5381011FEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD24B30B0EFA5381011FEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399720/files/figure.png	https://doi.org/10.5281/zenodo.7399720	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD24B30B0EFA5381011FEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399734/files/figure.png	https://doi.org/10.5281/zenodo.7399734	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD24B30B0EFA5381011FEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399750/files/figure.png	https://doi.org/10.5281/zenodo.7399750	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD24B30B0EFA5381011FEEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD34B33B3BBA198114BFCEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD34B33B3BBA198114BFCEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD34B33B3BBA198114BFCEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399722/files/figure.png	https://doi.org/10.5281/zenodo.7399722	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD34B33B3BBA198114BFCEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399732/files/figure.png	https://doi.org/10.5281/zenodo.7399732	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD34B33B3BBA198114BFCEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399748/files/figure.png	https://doi.org/10.5281/zenodo.7399748	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD34B33B3BBA198114BFCEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD04B33B3FCA3A9118CFB4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD04B35B388A4FE1015F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD04B35B388A4FE1015F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399706/files/figure.png	https://doi.org/10.5281/zenodo.7399706	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD04B35B388A4FE1015F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399708/files/figure.png	https://doi.org/10.5281/zenodo.7399708	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD04B35B388A4FE1015F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399724/files/figure.png	https://doi.org/10.5281/zenodo.7399724	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD04B35B388A4FE1015F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399734/files/figure.png	https://doi.org/10.5281/zenodo.7399734	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD04B35B388A4FE1015F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399746/files/figure.png	https://doi.org/10.5281/zenodo.7399746	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD64B37B3DCA6981720F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD64B37B3DCA6981720F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399708/files/figure.png	https://doi.org/10.5281/zenodo.7399708	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD64B37B3DCA6981720F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399724/files/figure.png	https://doi.org/10.5281/zenodo.7399724	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD64B37B3DCA6981720F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399734/files/figure.png	https://doi.org/10.5281/zenodo.7399734	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD64B37B3DCA6981720F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399746/files/figure.png	https://doi.org/10.5281/zenodo.7399746	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD64B37B3DCA6981720F9EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD44B29B136A69814BAFBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD44B29B136A69814BAFBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399740/files/figure.png	https://doi.org/10.5281/zenodo.7399740	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD44B29B136A69814BAFBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399742/files/figure.png	https://doi.org/10.5281/zenodo.7399742	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD44B29B136A69814BAFBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD44B29B136A69814BAFBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399750/files/figure.png	https://doi.org/10.5281/zenodo.7399750	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFD44B29B136A69814BAFBCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399754/files/figure.png	https://doi.org/10.5281/zenodo.7399754	Fig. 22. Distribution of Quedius molochinoides Smetana, 1965 (top), and Q. laticollis (Gravenhorst, 1802) and Q. strenuus Casey, 1915 (bottom).	Fig. 22. Distribution of Quedius molochinoides Smetana, 1965 (top), and Q. laticollis (Gravenhorst, 1802) and Q. strenuus Casey, 1915 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCA4B28B0C5A4781772FAEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCA4B28B0C5A4781772FAEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCA4B28B0C5A4781772FAEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399726/files/figure.png	https://doi.org/10.5281/zenodo.7399726	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCA4B28B0C5A4781772FAEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399740/files/figure.png	https://doi.org/10.5281/zenodo.7399740	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCA4B28B0C5A4781772FAEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399742/files/figure.png	https://doi.org/10.5281/zenodo.7399742	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCA4B28B0C5A4781772FAEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399750/files/figure.png	https://doi.org/10.5281/zenodo.7399750	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCA4B28B0C5A4781772FAEA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCB4B2BB126A59817E3FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCB4B2BB126A59817E3FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCB4B2BB126A59817E3FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399714/files/figure.png	https://doi.org/10.5281/zenodo.7399714	Fig. 5. Network analysis of Quedius molochinoides-group: Quedius horni Hatch, 1957 (purple), Q. lanei Hatch, 1957 (green), Q. molochinoides-group 1 (dark blue), Q. molochinoides-group 2 (light blue), and Q. molochinoides-group 3 (teal).	Fig. 5. Network analysis of Quedius molochinoides-group: Quedius horni Hatch, 1957 (purple), Q. lanei Hatch, 1957 (green), Q. molochinoides-group 1 (dark blue), Q. molochinoides-group 2 (light blue), and Q. molochinoides-group 3 (teal).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCB4B2BB126A59817E3FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399726/files/figure.png	https://doi.org/10.5281/zenodo.7399726	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCB4B2BB126A59817E3FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399740/files/figure.png	https://doi.org/10.5281/zenodo.7399740	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCB4B2BB126A59817E3FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399742/files/figure.png	https://doi.org/10.5281/zenodo.7399742	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCB4B2BB126A59817E3FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399750/files/figure.png	https://doi.org/10.5281/zenodo.7399750	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCB4B2BB126A59817E3FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC84B2AB0E3A4191140FC0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC84B2AB0E3A4191140FC0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC84B2AB0E3A4191140FC0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399726/files/figure.png	https://doi.org/10.5281/zenodo.7399726	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC84B2AB0E3A4191140FC0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399740/files/figure.png	https://doi.org/10.5281/zenodo.7399740	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC84B2AB0E3A4191140FC0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399742/files/figure.png	https://doi.org/10.5281/zenodo.7399742	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC84B2AB0E3A4191140FC0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399750/files/figure.png	https://doi.org/10.5281/zenodo.7399750	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC84B2AB0E3A4191140FC0A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC94B2CB3CEA3A917E9F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC94B2CB3CEA3A917E9F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC94B2CB3CEA3A917E9F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399714/files/figure.png	https://doi.org/10.5281/zenodo.7399714	Fig. 5. Network analysis of Quedius molochinoides-group: Quedius horni Hatch, 1957 (purple), Q. lanei Hatch, 1957 (green), Q. molochinoides-group 1 (dark blue), Q. molochinoides-group 2 (light blue), and Q. molochinoides-group 3 (teal).	Fig. 5. Network analysis of Quedius molochinoides-group: Quedius horni Hatch, 1957 (purple), Q. lanei Hatch, 1957 (green), Q. molochinoides-group 1 (dark blue), Q. molochinoides-group 2 (light blue), and Q. molochinoides-group 3 (teal).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC94B2CB3CEA3A917E9F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399726/files/figure.png	https://doi.org/10.5281/zenodo.7399726	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC94B2CB3CEA3A917E9F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399740/files/figure.png	https://doi.org/10.5281/zenodo.7399740	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC94B2CB3CEA3A917E9F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399742/files/figure.png	https://doi.org/10.5281/zenodo.7399742	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC94B2CB3CEA3A917E9F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399750/files/figure.png	https://doi.org/10.5281/zenodo.7399750	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC94B2CB3CEA3A917E9F8CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCF4B2EB0E6A7781752F94A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCF4B2EB0E6A7781752F94A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCF4B2EB0E6A7781752F94A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399714/files/figure.png	https://doi.org/10.5281/zenodo.7399714	Fig. 5. Network analysis of Quedius molochinoides-group: Quedius horni Hatch, 1957 (purple), Q. lanei Hatch, 1957 (green), Q. molochinoides-group 1 (dark blue), Q. molochinoides-group 2 (light blue), and Q. molochinoides-group 3 (teal).	Fig. 5. Network analysis of Quedius molochinoides-group: Quedius horni Hatch, 1957 (purple), Q. lanei Hatch, 1957 (green), Q. molochinoides-group 1 (dark blue), Q. molochinoides-group 2 (light blue), and Q. molochinoides-group 3 (teal).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCF4B2EB0E6A7781752F94A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399726/files/figure.png	https://doi.org/10.5281/zenodo.7399726	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCF4B2EB0E6A7781752F94A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399740/files/figure.png	https://doi.org/10.5281/zenodo.7399740	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCF4B2EB0E6A7781752F94A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399742/files/figure.png	https://doi.org/10.5281/zenodo.7399742	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCF4B2EB0E6A7781752F94A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399750/files/figure.png	https://doi.org/10.5281/zenodo.7399750	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	Fig. 21. Distribution of Quedius unicolor Kiesenwetter, 1847 (top),.Q. labradorensis Smetana, 1965 (middle), Quedius molochinoides-group (bottom). Barcoded specimens are denoted with +.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCF4B2EB0E6A7781752F94A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399754/files/figure.png	https://doi.org/10.5281/zenodo.7399754	Fig. 22. Distribution of Quedius molochinoides Smetana, 1965 (top), and Q. laticollis (Gravenhorst, 1802) and Q. strenuus Casey, 1915 (bottom).	Fig. 22. Distribution of Quedius molochinoides Smetana, 1965 (top), and Q. laticollis (Gravenhorst, 1802) and Q. strenuus Casey, 1915 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCF4B2EB0E6A7781752F94A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCD4B2EB157A6F817E3F7BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCD4B21B380A0B91106F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCD4B21B380A0B91106F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCD4B21B380A0B91106F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399724/files/figure.png	https://doi.org/10.5281/zenodo.7399724	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCD4B21B380A0B91106F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399734/files/figure.png	https://doi.org/10.5281/zenodo.7399734	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCD4B21B380A0B91106F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399754/files/figure.png	https://doi.org/10.5281/zenodo.7399754	Fig. 22. Distribution of Quedius molochinoides Smetana, 1965 (top), and Q. laticollis (Gravenhorst, 1802) and Q. strenuus Casey, 1915 (bottom).	Fig. 22. Distribution of Quedius molochinoides Smetana, 1965 (top), and Q. laticollis (Gravenhorst, 1802) and Q. strenuus Casey, 1915 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFCD4B21B380A0B91106F78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC74B27B137A5DF14ADF78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC74B27B137A5DF14ADF78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399710/files/figure.png	https://doi.org/10.5281/zenodo.7399710	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC74B27B137A5DF14ADF78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399724/files/figure.png	https://doi.org/10.5281/zenodo.7399724	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC74B27B137A5DF14ADF78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399734/files/figure.png	https://doi.org/10.5281/zenodo.7399734	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC74B27B137A5DF14ADF78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399754/files/figure.png	https://doi.org/10.5281/zenodo.7399754	Fig. 22. Distribution of Quedius molochinoides Smetana, 1965 (top), and Q. laticollis (Gravenhorst, 1802) and Q. strenuus Casey, 1915 (bottom).	Fig. 22. Distribution of Quedius molochinoides Smetana, 1965 (top), and Q. laticollis (Gravenhorst, 1802) and Q. strenuus Casey, 1915 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC74B27B137A5DF14ADF78A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399704/files/figure.png	https://doi.org/10.5281/zenodo.7399704	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399706/files/figure.png	https://doi.org/10.5281/zenodo.7399706	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	Fig. 2. Habitats of Quedius s. str. A – Q. fuliginosus (Gravenhorst, 1802) in mossy habitat in a forest on Zealand, Denmark; B – Oak forest habitat of Q. levicollis Brullé, 1832 found in litter in Extremadura, Spain; C – Saline lake habitat of Q. meridiocarpathicus Smetana, 1958 found in bank litter on Peloponese, Greece; D – River bank habitat of Q. hispanicus Bernhauer, 1898 found in flood debris in Extremadura, Spain; E – Wet mixed forest habitat of Q. fuliginosus found in litter in Southern Ural, Russia; F – Alpine near tree line habitat of Q. subunicolor Korge, 1961 found under stones at 1700 m in Altai Mountains, Russia; G – Alpine mossy creek habitat of Q. unicolor Kiesenwetter, 1847 found in moss in Italian Alps (Photo:A. Zanetti).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399708/files/figure.png	https://doi.org/10.5281/zenodo.7399708	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	Fig. 3. Phylogeny of Quedius s. str. based on Bayesian Inference (BI) and Maximum Likelihood (ML) analyses of COI barcodes (part 1). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399714/files/figure.png	https://doi.org/10.5281/zenodo.7399714	Fig. 5. Network analysis of Quedius molochinoides-group: Quedius horni Hatch, 1957 (purple), Q. lanei Hatch, 1957 (green), Q. molochinoides-group 1 (dark blue), Q. molochinoides-group 2 (light blue), and Q. molochinoides-group 3 (teal).	Fig. 5. Network analysis of Quedius molochinoides-group: Quedius horni Hatch, 1957 (purple), Q. lanei Hatch, 1957 (green), Q. molochinoides-group 1 (dark blue), Q. molochinoides-group 2 (light blue), and Q. molochinoides-group 3 (teal).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399716/files/figure.png	https://doi.org/10.5281/zenodo.7399716	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399734/files/figure.png	https://doi.org/10.5281/zenodo.7399734	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399720/files/figure.png	https://doi.org/10.5281/zenodo.7399720	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	Fig. 7. Habitus of Quedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. curtipennis Bernhauer, 1908; C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. unicolor Kiesenwetter, 1847; F – Q. subunicolor Korge, 1961.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399736/files/figure.png	https://doi.org/10.5281/zenodo.7399736	Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens.	Fig. 15. Variation of the shape of the paramere of Quedius sundukovi Smetana, 2003. Locality and BOLD BIN number are indicated under respective specimens.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399722/files/figure.png	https://doi.org/10.5281/zenodo.7399722	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	Fig. 8. Habitus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. meridiocarpathicus Smetana, 1958; C – Q. molochinus (Gravenhorst, 1806) (red elytra variant); D – Q. molochinus (black elytra variant); E – Q. balticus Korge, 1960; F – Q. sundukovi Smetana, 2003.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399724/files/figure.png	https://doi.org/10.5281/zenodo.7399724	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399728/files/figure.png	https://doi.org/10.5281/zenodo.7399728	Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 11. Aedeagus ofQuedius s. str. A – Q. afrofuliginosus Gusarov, 1991; B – Q. levicollis Brullé, 1832; C – Q. curtipennis Bernhauer, 1908; D – Q. fuliginosus (Gravenhorst, 1802). I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399726/files/figure.png	https://doi.org/10.5281/zenodo.7399726	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	Fig. 10. Habitus of Quedius s. str. A – Q. horni Hatch, 1957; B – Q. lanei Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. molochinoides Smetana, 1965 (red elytra variant); E – Q. molochinoides (black elytra variant); F – Q. labradorensis Smetana, 1965.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399730/files/figure.png	https://doi.org/10.5281/zenodo.7399730	Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange).	Fig. 12. Distribution of Eurasian species introduced to North America: Quedius curtipennis Bernhauer, 1908 (teal), Q. fuliginosus (Gravenhorst, 1802) (green), and Q. molochinus (Gravenhorst, 1806) (orange).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399740/files/figure.png	https://doi.org/10.5281/zenodo.7399740	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	Fig. 16. Aedeagus of Quedius s. str. A – Q. labradorensis Smetana, 1965; B – Q. horni Hatch, 1957; C – Q. altanai Hansen & Brunke, sp. nov.; D – Q. lanei Hatch, 1957; E – Q. molochinoides Smetana, 1965. I – parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399732/files/figure.png	https://doi.org/10.5281/zenodo.7399732	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	Fig. 13. Aedeagus of Quedius s. str. A – Q. vicinus (Ménétriés, 1832); B – Q. sundukovi Smetana, 2003; C – Q. subunicolor Korge, 1961; D – Q. molochinus (Gravenhorst, 1806); E – Q. balticus Korge, 1960; F – Q. meridiocarpathicus Smetana, 1958. I – parameral view of whole aedeagus; II – apices of paramere in antiparameral view with peg setae; III – lateral view of whole aedeagus; IV – apices of median lobe in parameral view.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399742/files/figure.png	https://doi.org/10.5281/zenodo.7399742	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	Fig. 17. Variation in genitalic morphology of species from Quedius molochinoides-group.From left to right for each specimen, first paramere then median lobe (parameral view) and lastly median lobe (lateral view) are given. For each specimen locality is noted underneath.	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399744/files/figure.png	https://doi.org/10.5281/zenodo.7399744	Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom).	Fig. 18. Distribution of Quedius afrofuliginosus Gusarov, 1991 (top), Q. curtipennis Bernhauer, 1908 (middle) and Q. fuliginosus (Gravenhorst, 1802) (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399746/files/figure.png	https://doi.org/10.5281/zenodo.7399746	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	Fig. 19. Distribution of Quedius levicollis Brullé, 1832 (top), Q. hispanicus Bernhauer, 1898 and Q. pallipes Lucas, 1846 (middle), and Q.balticus Korge, 1960 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/7399748/files/figure.png	https://doi.org/10.5281/zenodo.7399748	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	Fig. 20. Distribution of Quedius molochinus (Gravenhorst, 1806), Q. meridiocarpathicus Smetana, 1958, and Q. vicinus (Ménétriés, 1832) (top), Q. subunicolor Korge, 1961 (middle), and Q. sundukovi Smetana, 2003 (bottom).	2022-11-26	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey		Zenodo	biologists	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey			
