identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038987A0FFF54B16B124A7561163F9AE.text	038987A0FFF54B16B124A7561163F9AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius fuliginosus (Gravenhorst 1802)	<div><p>Quedius fuliginosus -group</p><p>Diagnosis. Head with 4 or 6 interocular punctures between anterior frontal punctures. Generally largest species within the subgenus with entire body dark; only rarely elytra reddish. Temporal puncture situated at posterior margin of eye. Paramere does not extend over median lobe apex, peg setae arranged in single rows; median lobe without large teeth on parameral side, at most with small teeth or subapical notch; internal sac without large c-shaped sclerite. Generally widespread species, common in the Palearctic region, found in diverse ground-based debris in both forest and open landscapes. Only Quedius afrofuliginosus is rare, possibly an artefact of both misidentifications and poor sampling in parts of its range.</p></div>	https://treatment.plazi.org/id/038987A0FFF54B16B124A7561163F9AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFF54B16B11EA5801498F8B4.text	038987A0FFF54B16B11EA5801498F8B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius Stephens 1829	<div><p>Subgenus Quedius Stephens, 1829</p><p>Type species. Staphylinus levicollis Brullé, 1832 by subsequent designation in Opinion 1851 (ICZN 1996: 215).</p><p>Diagnosis. Generally large and robust species; pronotum with deflexed hypomera; head with large eyes, fully developed infraorbital ridges and entire (not bilobed or apically notched) labrum; tarsi of forelegs rather wide in both sexes; mesoscutellum with two basal carinae; abdomen almost parallel sided, not notably tapering apicad; aedeagus with sharply pointed median lobe, internal sac well developed and often having heavily sclerotized structures, with longitudinal rows of sensory peg setae on paramere apically.</p></div>	https://treatment.plazi.org/id/038987A0FFF54B16B11EA5801498F8B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFF54B09B384A65C1056FA0A.text	038987A0FFF54B09B384A65C1056FA0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius afrofuliginosus Gusarov 1991	<div><p>Quedius afrofuliginosus Gusarov, 1991</p><p>(Figs 1, 3, 6E, 7A, 11A, 18)</p><p>Quedius afrofuliginosus Gusarov, 1991: 9 [Type locality: Algeria]</p><p>Type material. Hඈඅඈඍඒඉൾ: “ Algerie, Raffray / [black circle] / c.A.Yakovleva [in Russian] / Holotypus Quedius afrofuliginosus V.Gusarov 1990 [red label] / Zoological Institute St. Petersburg INS_COL_0002505” (J ZIN). Pൺඋൺඍඒඉൾඌ: same as holotype, except “ Allotypus Quedius afrofuliginosus V. Gusarov 1990 [red label] / Zoological Institute St. Petersburg INS_COL_0002506” (1 ♀ ZIN); same as holotype, except “ Paratypus Quedius afrofuliginosus V. Gusarov 1990 [red label] / Zoological Institute St. Petersburg INS_COL_0002507-000250710” (3 JJ 1 ♀ ZIN).</p><p>Additional material examined. FRANCE: Cඈඋඌൾ: Col de Bravella, [41.83, 9.26], 490m, 25.VII.1994, leg. Zerche (1 J SDEI). ITALY: Sൺඋൽංඇංൺ: Aritzo, Mte.Gennargentu,[39.94, 9.17], leg.C. Krüger (2JJ NMW); Ottana, [40.23, 9.04], 11.X.1974 (1 ♀ cBor); MOROCCO: Ain Leuh, [33.28, -5.33], 17.V.1925, leg. M. Escalera (1 ♀ MNCN). SPAIN: Cadiz, Laguna de la Janda, [36.24,-5.84], 30.V.1966, leg. Besuchet (1J NMPC).</p><p>Redescription. Measurements JJ (n = 6): HW = 1.78– 1.89 (1.84); HL = 1.54–1.58 (1.56); HL/HW 0.82–0.89 (0.85); PW = 2.24–2.44 (2.36); PL = 2.11–2.22 (2.16); PL/PW 0.89–0.96 (0.92); EW = 2.33–2.68 (2.59); EL = 2.40–2.51 (2.47); EL/EW 0.90–1.03 (0.95); EL/PL 1.10–1.19 (1.14); PW/HW 1.44–1.57 (1.51); forebody length 6.10–6.29 (6.19). ♀♀ (n = 3): HW = 1.84–1.89 (1.87); HL = 1.56–1.62 (1.59); HL/HW 0.84–0.86 (0.85); PW = 2.38–2.40 (2.39); PL = 2.04–2.24 (2.17); PL/PW 0.86–0.94 (0.91); EW = 2.69–2.73 (2.72); EL = 2.36–2.58 (2.48); EL/EW 0.88–0.94 (0.91); EL/PL 1.06–1.26 (1.15); PW/HW 1.47–1.53 (1.50); forebody length 6.18–6.36 (6.24).</p><p>Large species; body black (Fig. 7A).</p><p>Head black, distinctly transverse, with microsculpture of head transverse waves, four punctures between anterior frontal punctures, one additional unique puncture on either side posterior of these (Fig. 6E), eyes very large (EyL/TL = 3.17–3.91 (3.68)) and clearly protruding; antennae and palpi pale reddish, all anntennomeres elongate.</p><p>Thorax: pronotum black, slightly wider than long, clearly wider than head, with microsculpture of transverse waves, three punctures in dorsal row, two punctures in sublateral row with posteriormost puncture reaching level of middle puncture of dorsal row; scutellum smooth and glabrous; elytra black, uniformly pubescent, quadrate or slightly wider than long, slightly longer than pronotum; legs dark brown with inner face of tibia darkened and tarsi paler.</p><p>Abdomen black, tergites uniformly punctured, slightly iridescent.</p><p>Male. Aedeagus (Fig. 11A): paramere broad, rather parallel-sided, reaching apex of median lobe, with sensory peg setae forming two single rows along parameral lateral margins for 2/3 of their length; median lobe on parameral side with clear apical protrusion and two subapical small teeth, the latter positioned near end of peg setae rows of paramere; internal sac with four distal elongate medial sclerites.</p><p>Differential diagnosis. Quedius afrofuliginosus can be distinguished from all other Quedius s. str. by the unique head chaetotaxy, with a puncture on either side posterior of the anterior frontal puncture (Fig. 6E). Within the fuliginosus -group it is additionally distinguished from Q. fuliginosus by the pale antennae, from Q. levicollis by the impunctate and glabrous scutellum, and from Q. curtipennis by wider head (HW/HL = 0.74 vs 0.85) and aedeagal median lobe with apical protrusion.</p><p>Bionomics. Nothing is known about the biology of Q. afrofuliginosus . Based on collecting sites it is most likely occurring in forested areas of medium elevation (450 to 1400 m).</p><p>Distribution. Quedius afrofuliginosus was originally described from an unknown locality in Algeria. The new material studied in this paper brought to light additional specimens from Tunisia and Morocco in Northern Africa, as well as the first records of this species from the Mediterranean region in southern Spain and on the southern European islands of Corse and Sardinia (Fig. 18).</p></div>	https://treatment.plazi.org/id/038987A0FFF54B09B384A65C1056FA0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFEA4B0EB38AA5B816FBFC6A.text	038987A0FFEA4B0EB38AA5B816FBFC6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius curtipennis Bernhauer 1908	<div><p>Quedius curtipennis Bernhauer, 1908</p><p>(Figs 1, 3, 6B, 7B, 11C, 12, 18)</p><p>Quedius curtipennis Bernhauer, 1908 e: 335 [Type locality: Faroe; Romania; Buchara; Böhmen, Wran a. Moldau].</p><p>Quedius gracilis Stephens, 1832: 215 [Type locality: London], syn. rev. Quedius parallelus Hatch,1957:216 [Type locality:Washington, Seattle]</p><p>References. Gඋංൽൾඅඅං (1924): 79 (characters), SർHൾൾඋඉൾඅඍඓ (1933): 1441 (characters), Hൺඇඌൾඇ (1952): 141 (characters); Sආൾඍൺඇൺ (1958): 362 (characters and ecology); (1962): 133, (1965b): 38 (characters); (1971a): 129 (characters, synonymy of Q. parallelus); (1965a): 12, (1970): 62, (1990): 98, (1993): 50 (distribution); (1978a): 826 (biology); KඈඋGൾ (1962b): 332 (suspected synonymy of Q. parallelus); (1964): 119 (distribution); Pൺඅආ (1963): 141 (characters); LඈHඌൾ (1964): 211 (characters); Dඏඈෞගĸ (1965): 89 (distribution); Hඈඋංඈඇ (1965): 272 (distribution); Sඓඎඃൾർĸං (1968): 735 (distribution); Fඋൺඇĸ (1969): 267 (biology); Bඈඋൽඈඇං (1974b): 10, (1976b): 92 (characters); Pඈඉൾ (1977): 31 (distribution); Cඈංൿൿൺංඍ (1978): 191 (characters); Oඎඍൾඋൾඅඈ (1978): 281 (pupal characters); Bඎඋൺĸඈඐඌĸං et al. (1980): 134 (distribution); TඬඍH (1984): 118 (characters); DඋඎGආൺඇൽ (1987): 312 (distribution); Nඈඐඈඌൺൽ (1990): 145 (biology); BඈඋGൾඌ (1990): Table II (distribution); Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං (1995): 32 (distribution); Oඐൾඇ (1997): 149 (distribution); Oඎඍൾඋൾඅඈ et al. (1998): 129 (biology); AඌඌංඇG (2001): 75 (biology); UHඅංG et al. (2006): 54 (distribution); Tඋඈඇඊඎൾඍ (2006): 102 (distribution); Mൺඃĸൺ &amp; Sආൾඍൺඇൺ (2007): 429 (characters and distribution); ÖඓGൾඇ (2011):204 (distribution); Bඋඎඇĸൾ &amp; MൺඋඌHൺඅඅ (2011): 59 (characters and distribution); Wൾൻඌඍൾඋ et al. (2012): 313 (distribution); MൺGඎඋൺ et al. (2013): 719 (biology); TඬඍHආඣඋඣඌඓ et al. (2014): 688 (biology); Sൾආൾඇඈඏ et al. (2015): 127 (distribution); Sൾආංඈඇൾඇĸඈඏ et al. (2015): 330 (distribution); Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2018a): 125 (notes on types and distribution); (2019): 48 (distribution).</p><p>Type material examined. Quedius curtipennis: Lൾർඍඈඍඒඉൾ: (J, here designated): “Suderö Faroer Ins./ Dr. Cornu 1907/ v. curtipennis Brh. Typus / fuliginosus Grav. Scheerp. / Chicago NHMus M. Bernhauer Collection / D. Drugmand det. 1994 Quedius (s. str.) curtipennis Brnh. / Lectotype Quedius curtipennis Bernhauer, 1908 A.K. Hansen &amp; A. Solodovnikov des. 2021” (FMNH). Pൺඋൺඅൾർඍඈඍඒඉൾඌ: “Nördl. Faroer Ins. / Dr. Cornu 1907 / v. curtipennis Brh. Typus / fuliginosus Scheerp. [sic!] det. [illegible] / Chicago NHMus M. Bernhauer Collection / D. Drugmand det. 1994 Quedius (s. str.) curtipennis Brnh / Paralectotype Quedius curtipennis Bernhauer, 1908 A.K. Hansen &amp; A. Solodovnikov des. 2021” (1 J FMNH); “ v. curtipennis Buchara Bang Haas det. Bernh. / Chicago NHMus M. Bernhauer Collection” (1 J FMNH).</p><p>Quedius parallelus: Hඈඅඈඍඒඉൾ: “Seattle, Wash. U.W.Campus IV-12- 1949 / Del. 1954 H. Houk / Type J Quedius (s. str.) parallelus - 1951 M. Hatch / Quedius (s. str.) fuliginosus Grav.M. Hatch – 1947” (1J UWBM). Pൺඋൺඍඒඉൾඌ: “Seattle,Wash. April 12, 1934 ” (1J UWBM); “Seattle,Wash. U.W.Campus, IV-11-1947 ” (1 J UWBM, N. Vowles coll.); “Vancouver, B.C., 5 March, 1949, G.B. Rich” (1 J CNC); “Wash. Seattle, Feb. 15, 1954, B. Malkin” (1 ♀ FMNH, ex coll. B. Malkin).</p><p>Additional material examined. ARMENIA: Arzakanskaja dolina, Aghveran, [40.51, 44.56], 3.VII.1978, leg. R. Rous (1 ♀ MCZ); NE Dilizhan nr. Haghartsin Monastery, 40.8013, 44.8919, 1450 m, 21.V.2001, leg. Shaverdo &amp; Schillhammer (4 JJ NMW); 18 km E Dilizhan, 3 km Gosh, 40.7405, 45.0191, 1850 m, 27.V.2001, leg. Shaverdo (1 J NMW); 21 km N Hrazdan, Ankavan, 40.6286, 44.5533, 1000 m, 22.V.2001, leg. Shaverdo &amp; Schillhammer (1 J NMW); Teghut, Akhtala, 41.0901, 44.8120, 1009 m, 28.VI.2016, leg. J. Müller (1 J cSch); same locality, 41.0911, 44.8131, 1020m, 20.V.2016, leg.A. &amp; J. Müller (2♀♀ ZMHB). AUSTRIA: Bisamberg, [48.33, 16.36], leg. Luze (1 ♀ NMW); S Steiermark, [46.91, 15.60] (1 J NMW); Tirol, Musau, [47.53, 10.67], litter, moss, and grass, 28.C.1995, leg. M.Uhlig (1 J ZMHB); Wolfpassing, [48.08, 15.06], VI.1965, leg. Wewalka (1 ♀ NMW). BULGARIA: Rila Monastery, [42.13, 23.34], 1150 m, 14.VII.1965, leg. P. Jørum (1 J NHMD). BOSNIA AND HERZEGOVINA: Zeleznica [Zeljeznica] n. Foinica, [43.89, 17.95], 500-700m, 13.IV.1969, leg. U. Heinz (1 J ZMHB). CANADA: BඋංඍංඌH Cඈඅඎආൻංൺ: Stanley Park, Vancouver, [49.30, -123.14], 9.VI.1954, leg. D. Lazorko (1 J 1 ♀ NMW). CZECH REPUBLIC: Brandeis, [50.18, 14.65], leg. Skalitzky (2 JJ NMW). DENMARK: Vigsø Bugt, 1 km E of Vigsø, 57.0950, 8.7394, pine forest, pitfall traps, 16.-18.VI.2014, leg.A.Hansen, M.Justesen,A.Solodovnikov &amp; L.Kraemer (1 J NHMD) LඔඌØ: Laesø Klitplantage, 57.3082, 11.0081, mixed forest, sifting litter, 28.V.2013, leg.A. Solodovnikov &amp; M.Justesen (1 NHMD). FAROE ISLANDS: Suderö [Suðuroy], [61.51, -6.83], 1907, leg. Cornu (1 J FMNH). FRANCE: Ain, Torcieu n. Amberieu, [45.91, 5.39] (1 J ZMHB); Bretagne, Locronan, [48.09, -4.21], VII.1983, leg. H.Korge (1 J ZMHB); Chigny n. Oise, [49.91, 3.76], leg. Gelchert (1 ♀ ZMHB); Morlaix, Bretagne, [48.57, -3.82], 1.VIII.1950, leg. A. Nilsson (2 JJ MZLU). GEORGIA: Borzhom Distr., Bakuriani, [41.75, 43.52], 27.VII.1947, leg. Bogachev (1 J 1 ♀ ZIN); Kartliskij Chreb. [Kartalinskiy Khrebet], Sabaduris Tye [Sabaduri Pass] [41.91, 44.91], 1800m, 8.VI.1987, leg. Wrase (1 J cSch); same locality, 25.VI.1988, leg. Wrase (1 J ZMHB); Trialetski, Khrebet Bakuriani, [41.74, 43.83], 1800-2200m, 4.-7.VII.1986, leg.Wrase &amp; Schülke (8JJ 4♀♀ ZMHB); Zchneti [Tskneti], [41.69, 44.69], 1200m, 26.VI.1986, leg. Wrase (1 J cSch). GERMANY: Bayern, Garmisch, [47.48, 11.08], 27.IV.1932, leg. Ihssen (1 J ZMHB); Bonn, [50.72, 7.09], X, leg. Verhoeff (1 ♀ ZMHB); Bayern, Eschenlohe, [47.58, 11.18], 3.VIII.1937, leg.Ihssen (1J ZMHB); Brandenburg, Strausberg, [52.52, 13.85], 17.IV.1987, leg. M. Schülke (1 J ZMHB); Dresden, Kötschenbroda,[51.11, 13.61], moss, 23.I.1920, leg. Dahl (1 J ZMHB); Düsseldrof Unterbach, [51.20, 6.89], 27.VII.1974, leg. H.Bremer (1 ♀ ZMHB); Marburg, [50.80, 8.76], 16.IV.1904, leg. Strand (1 ♀ ZMHB); Sarstadt, [52.24, 9.85], 12.IX.1960, leg. Kuntze (1 J ZMHB). ITALY: Aniene, Lazio, [41.95, 12.82], 1200 m, 18.III.1964, leg. A. Vigna-Taglianti (1 J cBor); Barni, Lombardia, [45.91, 9.26], 11.II.2001, leg. Diotti (1 J 1 ♀ NHMD); Moliterno, Basilicata, [40.24, 15.86], 30.IX.2000, leg. F. Angelini (1 ♀ NHMD); Pignola, Basilicata, [40.57, 15.78], 700 m, 19.IV.1998, leg. F. Angelini (1 J NHMD); Querciola, Toscana, [43.84, 11.32], 13.IV.2005, leg. Bordoni (1 J cBor). LATVIA: Curonia Libau [Liepāja], [56.49, 21.01], 4.IV.1910 (1 J 1 ♀ ZMHB). MOROCCO: Cirque du Jaafar, [32.55, -4.91], 4.III.1964 (1 J NMW) [potentially mislabelled or introduced specimen clearly belonging to this species]. MYANMAR: Mountains, Tenasserim, Siam Border, [11.79, 99.62], II.-V.1913, leg. K.G. Gairdner (1 J BMNH) [potentially mislabelled or introduced specimen clearly belonging to this species]. PO-LAND: Dunajec River n. Sromowce Nizne, [49.39, 20.39], 18.VII.1977, leg. F. Hieke (3 JJ 4 ♀♀ ZMHB); Wyspowy, Krzozonowka River n. Poim, [49.69, 20.24] 300m, 26.VIII.1980, leg. Hieke (1 J ZMHB). PORTUGAL: Aඓඈඋൾඌ: Santa Maria, 2 km ENE Almagreira, Miradoro dos Picos, [36.97, -25.07], 440 m, 19.III.1957, leg. Brinck &amp; Dahl (1 J MZLU); São Miguel, 500 m E of Lagoa do Pau Pique, [37.83, -25.74], pond, 7.III.1957, leg. Brinck &amp; Dahl (1 J MZLU); São Miguel, Pico de Vara, 37.8119, -25.2105, 1050m, boggy slope 24.VII.2013, leg.V.Assing (4 ♀♀ ZMHB). Mൺൽൾංඋൺ: Levada da Serra do Faial, [32.68, -16.85], creek, 820m, 11.IX.2014, leg. A. Kleeberg (1 J cSch). ROMANIA: Klaussenburg [Cluj-Napoca], [46.76, 23.62], leg.Verhoeff (1 J ZMHB); Hermannstadt, Siebenbürgen [Sibiu], [45.79, 24.15], leg. Skalitzky (1 J NMW); Piatra, Virful Tiganului, Sighetu Marmației, Maramureș, [47.99, 23.53], 1300 m, pasture, in rotten logs of beech, 14.VI.2006, leg. A. Grabant, Z. György, O. Merkl &amp; A. Podlussany (1 J NHMD). RUSSIA: KൺඋൺർHൺඒ- CH ൾඋKൾඌඌංൺ: Teberda,[43.44, 41.74], VI.1912, leg.Roubal (1 ♀ SNMC). Mඈඌർඈඐ Oൻඅൺඌඍ: Bykovo village, VNIIKR institute territory, [55.61, 38.05], 16.X.2013, leg.S.A. Kurbatov (3 JJ ZIN); Odintsovsky Distr., near Zhavoronki vill., [55.64, 37.10], 6-13.VIII.2017, leg. E. M. Veselova &amp; A. B. Ryvkin (1 J ZIN). NංඓHൾGඈඋඈൽඌKൺඃൺ Oൻඅൺඌඍ: near Gorodets town, [56.64, 43.47], mixed forest, 17.VIII.1989, leg. A. Klimenko (1J ZIN). NඈඏGඈඋඈൽඌKൺඃൺ Oൻඅൺඌඍ: Torbino, Novgorod, [58.58, 32.87], 10.IV.1914, leg. A. Filiniev (1 J cBor). NඈඋඍH Oඌඌൾඍංൺ Rൾඉ.: Caucasus, Skalisty Khrebet, Fiagdon River, [42.87, 44.32], 1700-2200 m, VI.1976, leg. R. Rous (3 JJ 1 ♀ MCZ). SLOVAKIA: Belá River n. Pribylina, [49.09, 19.81], 600m, 18.VIII.1981, leg. Hieke (1 J 3 ♀♀ ZMHB). SPAIN: Bera de Bidasoa, Navarra, [43.27, -1.68], 1.-7.I.2001, leg.A.Anichtchenko (1J NHMD); Caldetas n. Barcelona, [41.57, 2.52], leg. G. Heine (1 J ZMHB); La Molina, Pyrenees, [42.33, 1.93], 11.-16. VII.1963 (1 J MZLU); O Rial, Serra do Mirador, [42.74, -7.83], 450m, 3.VI.2003, leg. J. P. Valcárcel (1 J cSch); Picos de Europa, Camping El Redondo [Fuente Dé], [43.14, 4.81], forest, 1100 m, 14.-17.VII.1996, leg. Wrase (1 J cSch); Plaiaundi, Irun, Guipuzcoa, [43.34, -1.79], V.2006, A. Anichtchenko (1 J 1 ♀ NHMD); Pyrenees, Toses, 42.3502, 1.9955, 1660m, fir-spruce forest with moss, 9.X.1997, leg. Zerche (1 J SDEI); Sierra Ancares, [42.81, -6.86], 20.X.1984, leg. J. C. Otero (6 JJ 3 ♀♀ ZMHB); Sierra Caurel, [42.59, -7.18], 20.VII.1985, leg. J. C. Otero (5 JJ 4 ♀♀ ZMHB); Sierra de Gredos, Hoyos de Collado, [40.35, -5.21], 1600m, 14.IV.1991,leg.Heinz (2♀♀ ZMHB); Sierra Guadarrama, [40.85, -3.95], 1.VI.1980, leg. C. Otero (2 JJ ZMHB); Sierra de la Demanda, Val de Zcaray [Ezcaray], [42.32, -3.02], forest, 1650m, 23.07.1996, leg. Zaballos &amp; Wrase (1 J cSch). SWEDEN: Brönnestad, [56.08, 13.70] (1 MZLU); Omberg, [58.33, 14.64] (1 MZLU); Strömholm, [59.52, 16.23] (1 MZLU); Visseltofta, [56.42, 13.85] (1 MZLU); Öඅൺඇൽ: Vickleby, [56.57, 16.46] (1 MZLU). SWITZERLAND: Wolfenschiessen, [46.91, 8.39], 5.VIII.1963, moss (1 J ZMHB). TURKEY: Abant n. Bolu, [40.61, 31.27], 30.V.1964, leg. H.Korge (1 J ZMHB); Akkus n. Ünye, [40.79, 37.01], 28.V.1964,leg. H.Korge (1J 2♀♀ ZMHB); Ilgaz Dağları, [41.11, 33.90], 17.-21.VI.1960, leg. F. Schubert (1 ♀ NMW); n. Ismetpasa &amp; Cerkes, [40.75, 32.82], 05.IV.1979, leg. Heinz (1 J NMEG); Kulakkaya n. Giresun, [40.69, 38.33], 1450m, 27.VII.1963, leg. H. Korge (2 ♀♀ ZMHB). UKRAINE: Tausban Bazar, [44.97, 34.59], 20.VI.1907, leg. B. Grigoriev (1 J ZIN). UNITED KINGDOM: EඇGඅൺඇൽ: Barnstaple, Devon, [51.07, -4.06], 12.-31.V.1954, leg. Lindroth (1J MZLU); Devon, Bidefort [51.01, -4.21], leg. H. Korge (2 ♀♀ ZMHB); Kent, Knockholt, 51.3124, 0.1191, 2.IV.2011, leg. J.J. Shaw (1 J cJen); Shrops, Edgmond Harper Adams Uni. Campus, [52.77, -2.42], 6.III.2015, leg. J.J. Shaw (1 J cJen); Sർඈඍඅൺඇൽ: Loch Hope, [58.44, -4.62], 19.VII.1987, leg. G. Gillerfors (1 J MZLU). USA: Maine: Booth Bay, [43.85, -69.62], 15.VIII.1982, leg. Wewalka (1 ♀ NMW). UZBEKISTAN: Buchara, [39.77, 64.42], leg. Bang Haas (1 J FMNH).</p><p>Redescription. Measurements JJ (n = 6): HW = 1.89– 1.96 (1.93); HL = 1.53–1.71 (1.61); HL/HW 0.81–0.88 (0.84); PW = 2.36–2.56 (2.49); PL = 2.16–2.36 (2.29); PL/PW 0.87–0.94 (0.92); EW = 2.51–2.67 (2.61); EL = 2.36–2.49 (2.44); EL/EW 0.92–0.96 (0.93); EL/PL 1.04– 1.10 (1.06); PW/HW 1.31–1.60 (1.49); forebody length 6.04–6.56 (6.34). J ♀ (n = 4): HW = 1.78–2.00 (1.89); HL = 1.51–1.64 (1.58); HL/HW 0.82–0.85 (0.84); PW = 2.22–2.56 (2.41); PL = 2.07–2.40 (2.18); PL/PW 0.86–0.94 (0.91); EW = 2.44–2.62 (2.52); EL = 2.20–2.49 (2.33); EL/EW 0.90–0.95 (0.92); EL/PL 1.03–1.15 (1.07); PW/ HW 1.47–1.55 (1.52); forebody length 5.78–6.53 (6.10).</p><p>Very large species; body black (Fig. 7B).</p><p>Head black, slightly transverse; eyes large (EyL/TL = 3.00–4.10 (3.55)) not protruding; microsculpture of transverse waves, four punctures between anterior frontal punctures (Fig. 6B); antennae and palpi pale reddish, antennae long, all antennomeres clearly elongate.</p><p>Thorax: pronotum black, slightly wider than long, clearly wider than head, microsculpture of transverse waves, three punctures in dorsal row and two in sublateral row with posteriormost puncture reaching level of middle puncture of dorsal row; scutellum smooth and glabrous; elytra black, uniformly pubescent, wider than long, of equal length as pronotum; legs dark brown with inner face of tibia darkened and tarsi paler.</p><p>Abdomen black, tergites uniformly punctured, slightly iridescent.</p><p>Male. Aedeagus (Fig. 11C): paramere broad, rather parallel-sided, reaching apex of median lobe, with sensory peg setae forming two single rows following parameral lateral margins for 2/3 of their length; median lobe with apical part dorsoventrally flattened to form a plate-like structure; internal sac with two pairs of sclerites.</p><p>Differential diagnosis. Within the fuliginosus- group Q. curtipennis is very close to Q. fuliginosus . Both species are very frequently encountered, often co-occurring, nearly non-distinguishable from each other based on external morphological characters alone. Our study of hundreds of specimens across their entire distribution ranges revealed that Q. curtipennis always have pale antennomeres 1–3, whereas Q. fuliginosus have them at least slightly darkened. A negligible fraction (ca. 2 %) of Q. fuliginosus with pale basal antennomeres are mostly teneral. Antennae are generally more elongate in Q. curtipennis, as opposed to being stouter in Q. fuliginosus (Fig. 7). Also Q. curtipennis has an almost smoothly rounded head with the eyes protruding while Q. fuliginosus has eyes clearly protruding and thus making its head seem more quadrate (Fig. 6). Only some rare specimens, maybe hybrids, faded and very old museum specimens, or actual rare intraspecific variants, do not fit these trends and their identity can be ascertained only through examination of the aedeagi. In Q. curtipennis, contrary to Q. fuliginosus, the paramere lacks clear medial attenuation, the apical part of median lobe dorso-ventrally flattened, and internal sac with 2 pairs of clear sclerites.</p><p>The examined material for both species clearly shows that Q. curtipennis is much more common in the south and west and becomes increasingly rare towards the north and east of its range, while Q. fuliginosus shows an opposite trend. Quedius curtipennis seems to be more common in warmer and drier areas with more sandy soils, where Q. fuliginosus seems to prefer wetter, colder and more humus rich soils.</p><p>Quedius curtipennis is also very similar to Q. afrofuliginosus, from which it is easily recognized by having no additional punctures between anterior and posterior frontal punctures. Besides, distributions of Q. curtipennis and Q. afrofuliginosus, as far as known, are allopatric.</p><p>From Q. levicollis, Q. curtipennis is easily recognized by unpunctured and glabrous scutellum.</p><p>Lectotype designation. BൾඋඇHൺඎൾඋ (1908) described Q. curtipennis as a variety of Q. fuliginosus without giving any exact data on the type material. He had stressed that it is very common at Faroe Islands and mentioned its occurrence in Central Europe as well as in ‘Romania’ and even ‘Buchara’. Here (see above), we have listed specimens from Faroe Islands and ‘Buchara’ at the FMNH which qualify as syntypes of Q. curtipennis and which we recorded long ago but were unable to re-examine in detail recently. There is no doubt about the identity of the syntypes from Faroe Islands. However, a syntype from ‘Buchara’ must be carefully checked. Given, additionally, some ambiguity about the geographic origin of that Middle Asian syntype (see Distribution), a syntype male from Faroe Islands is here designated as a lectotype for Q. curtipennis .</p><p>Synonymic notes. Quedius parallelus was described by HൺඍർH (1957) in his work on beetles of the Pacific Northwest of North America without reference to European species. Based on the description of Q. parallelus, KඈඋGൾ (1962b) suspected it to be identical to the European Q. curtipennis . This was confirmed by Sආൾඍൺඇൺ (1971a) who studied the type material of Q. parallelus and found them conspecific with 2 male syntypes of Q. curtipennis from Faroe Islands at the FMNH. It is now firmly known that Quedius curtipennis occurs in the North American West (Sආൾඍൺඇൺ 1971a) and East Coasts (Sආൾඍൺඇൺ 1990, Bඋඎඇĸൾ &amp; MൺඋඌHൺඅඅ 2011).</p><p>Quedius gracilis Stephens, 1832 was described from London, UK. Later, SඍൾඉHൾඇඌ (1837) had it as a variety of Q. tristis (now Q. levicollis). In more recent works it was cited as a synonym of Q. fuliginosus . Based on examination of its two syntypes by Roger Booth (BMNH, pers. comm.) on our request, it was established that these specimens are conspecific with Q. curtipennis . As the name is older than Q. curtipennis we here establish the prevailing use of Q. curtipennis as opposed to the rule of priority (ICZN Article 23.9). Quedius curtipennis has been used at least 26 times in the last 50 years by more than ten authors (Bඈඋൽඈඇං 1974b, 1976b; Pඈඉൾ 1977; Cඈංൿൿൺංඍ 1978; Oඎඍൾඋൾඅඈ 1978; Bඎඋൺĸඈඐඌĸං et al. 1980; TඬඍH 1984; DඋඎGආൺඇൽ 1987; Nඈඐඈඌൺൽ 1990; BඈඋGൾඌ 1990; Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං 1995; Oඐൾඇ 1997; Oඎඍൾඋൾඅඈ et al. 1998; AඌඌංඇG 2001; UHඅංG et al. 2006; Tඋඈඇඊඎൾඍ 2006; Mൺඃĸൺ &amp; Sආൾඍൺඇൺ 2007; ÖඓGൾඇ 2011; Bඋඎඇĸൾ &amp; MൺඋඌHൺඅඅ 2011; Wൾൻඌඍൾඋ et al. 2012; MൺGඎඋൺ et al. 2013; TඬඍHආඣඋඣඌඓ et al. 2014; Sൾආൾඇඈඏ et al. 2015; Sൾආංඈඇൾඇĸඈඏ et al. 2015; Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ 2018a, 2019). On the other hand, Q. gracilis has only been used once after its description as a valid species which was more than 150 years ago (Cඎඋඍංඌ 1837). This deems the name Quedius gracilis Stephens, 1832 a nomen oblitum in relation to Quedius curtipennis Bernhauer 1908 (nomen protectum). Quedius gracilis Stephens, 1832 syn. rev. is thus moved from synonymy with Quedius fuliginosus (Gravenhorst, 1802) to synonymy with Quedius curtipennis Bernhauer 1908 .</p><p>Bionomics. Based on published data as well as our observations, Q. curtipenni s is a polytopic, somewhat thermophilous species found in a large range of habitats and various ground-based microhabitats with a tendency to occur in open woodlands. For example, in Central and Eastern Europe it was found under stones in dry pastures in Poland (Dඏඈෞගĸ 1965), in xerothermous grassland in Germany (AඌඌංඇG 2001), and in closed oak forest with extensive ground vegetation and shrub cover in Hungary (TඬඍHආඣඋඣඌඓ et. al 2014). We have collected this species in a mixed forest on the sandy dry island of Laesø in Denmark. There are also records from mole ( Talpa europaea) nests built from grasses and reeds on wet meadow in Poland (Nඈඐඈඌൺൽ 1990). In south-western Europe it was recorded from under rock in pine forest at an altitude of 1550 m in Central Spain (Oඎඍൾඋൾඅඈ 1978) and from a cave in Northern Spain (Oඎඍൾඋൾඅඈ et al. 1998). On the Azores (Sආൾඍൺඇൺ 1970) it was found under rocks at the shore of a freshwater lake at an elevation of 400– 500 m. Presumably, the nest or cave-based subterranean records are accidental occurrences, as there is no evidence of the species preferring these habitats. The species is found within the wide range of elevations from sea level in the northern part of its range up to about 2.000 m the southern part.</p><p>As an introduced species in North America, it was found especially around human settlements in various debris, under rocks, and in greenhouses. Some specimens were also collected by sifting barnyard litter on a pasture in Oregon (Sආൾඍൺඇൺ 1978a) and in natural habitats away from settlements (in moss, in leaf litter, etc.).</p><p>Pupae of the species are described in Oඎඍൾඋൾඅඈ (1978). Adult females from Wytham, UK were observed to contain eggs between October and January (Fඋൺඇĸ 1969).</p><p>Distribution. Quedius curtipennis is distributed from the Atlantic coast of Europe to Asia Minor, east to the Ural Mountains and Middle Asia (Fig. 18). It is also present on the islands of Madeira, the Azores, and the Faroe Islands. One specimen, which we studied was found in the central Morrocan mountains, and one old specimen from the type series is from ‘Buchara’, Uzbekistan. These are the only specimens we know from North Africa and Middle Asia, respectively. The species’ occurrence in these areas needs confirmation to rule out possible mislabelling. Most previous records of Q. curtipennis (and Q. fuliginosus) from North Africa could be assigned to Q. afrofuliginosus, which is externally very similar. As Q. curtipennis does not seem to occur in southern Spain, Corse and Sardinia, it is very likely absent from North Africa too.</p><p>Quedius curtipennis is also known as introduced to at least four regions of North America (Fig. 12). The presence of Q. curtipennis in the Nearctic was first recognized by Sආൾඍൺඇൺ (1971a) when he reviewed specimens of Q. parallelus, a newly described species from the Nearctic. Quedius parallelus was found to be identical to the Palearctic Q. curtipennis, something already hinted by KඈඋGൾ (1962b). One specimen from the type series of Q. parallelus was collected in 1934 near the city of Seattle in Washington, which is currently the oldest known record of the species from North America. In 1971, the species was already found in a relatively large area along the Pacific coast of Oregon, Washington, and British Columbia (Sආൾ-ඍൺඇൺ 1971a), suggesting that it may have been established in North America even prior to 1934. Currently, Q. curtipennis in western North America is still known only from these states and provinces, where it is found as far East as Cranbrook, British Columbia in the Canadian Rockies (from web www.boldsystems.org) (teal in Fig. 12). One to three additional introductions of Q. curtipenni s were later reported from the East Coast, in New Hampshire, USA, as well as Ontario and Nova Scotia, Canada. Specimens from near Guelph in Ontario were collected in 1976 (Bඋඎඇĸൾ &amp; MൺඋඌHൺඅඅ 2011), from New Hampshire in 1983 (Sආൾඍൺඇൺ 1990) and from Nova Scotia in 1997 (Mൺඃĸൺ &amp; Kඅංආൺ-ඌඓൾඐඌĸං 2008).</p></div>	https://treatment.plazi.org/id/038987A0FFEA4B0EB38AA5B816FBFC6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFED4B02B391A418119CFEEA.text	038987A0FFED4B02B391A418119CFEEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius fuliginosus (Gravenhorst 1802)	<div><p>Quedius fuliginosus (Gravenhorst, 1802)</p><p>(Figs 1, 2A,E, 3, 6C, 7C, 11D, 12, 18)</p><p>Staphylinus fuliginosus Gravenhorst, 1802: 34 [Type locality:Brunsviga]. Note: See lectotype designation by Gඎඌൺඋඈඏ (1993)</p><p>Quedius granulipennis Motschulsky, 1858: 656 [Type locality: Autriche]. Note: See lectotype designation by Gඎඌൺඋඈඏ (1993)</p><p>Quedius impunctifrons Delahon, 1915: 395 . [Type locality: Mark Brandenburg]</p><p>Quedius jelineki Krása, 1904: 81 [Type locality: Vrané n. Vlt, Bohemia]</p><p>Quedius latus Hochhuth, 1851: 30 [Type locality: Kaukasien, Alagez- Gebirge], syn. nov.</p><p>Quedius molochinicolor Roubal, 1931: 1 [Type locality:RCS.:Harmanec]</p><p>Quedius picicornis Stephens, 1832: 215 [Type locality: London]</p><p>Quedius subfuliginosus Britten, 1944: 290 [Type locality: England: Cotterill Clough, Cheshire]</p><p>Quedius viduus Sawada, 1965: 17 [Type locality: Japan, Pref. Nagano, Mt. Jônen], syn. nov.</p><p>References. MൺඋඌHൺආ (1802): 504 (characters); Lൺඍඋൾංඅඅൾ (1804): 319 (characters); GඒඅඅൾඇHൺඅ (1810):301 (as tristis; characters);MൺඇඇൾඋHൾංආ (1830): 25 (distribution); (1831): 439 (distribution); Nඈඋൽආൺඇඇ (1837): 78 (distribution); EඋංർHඌඈඇ (1839a): 490 (distribution); (1840): 537 (characters); Hൾൾඋ (1839): 276 (characters); Dඎൿඈඎඋ (1843): 33 (characters); Kඈඅൾඇൺඍං (1846): 22 (distribution); GඋൺඏൾඇHඈඋඌඍ (1847): 232 (characters); HඈർHHඎඍH (1849): 151 (characters); (1862): 46 (distribution); RൾൽඍൾඇൻൺർHൾඋ (1849): 710, (1857): 203, (1874): 200 (characters); Kංൾඌൾඇඐൾඍඍൾඋ (1851): 419 (distribution); Hൺඋൽඒ (1851): 33 (biology); Kඳඌඍൾඋ (1853): 62 (characters); Fൺංඋආൺංඋൾ &amp; Lൺൻඈඎඅൻජඇൾ (1856): 539 (characters); Kඋൺൺඍඓ (1857): 503 (characters); (1858): 58 (distribution); Pൾඒඋඈඇ (1858): 428 (distribution); Fൺඎඏൾඅ (1865):293 (biology); (1874): 289 (characters); Sൾංൽඅංඍඓ (1875): 267 (characters); Mඎඅඌൺඇඍ &amp; Rൾඒ (1876): 686 (characters); SൺHඅൻൾඋG (1876): 24 (distribution); Fඈඐඅൾඋ (1888): 237 (characters); GൺඇGඅൻൺඎൾඋ (1895): 403 (characters); Pඈඉඉංඎඌ (1905): 9 (distribution); Pඈඋඍൺ (1907): 128 (characters); Rൾංඍඍൾඋ (1909): 111 (characters); JඈHൺඇඌൾඇ (1914): 363 (characters); EංർHൾඅൻൺඎආ (1914): 339, (1915): 91 (morphology); Fංඈඋං (1915): 16 (distribution); Mඎඇඌඍൾඋ (1923): 195 (distribution); Gඋංൽൾඅඅං (1924): 79 (characters); Pඈඋඍൾඏංඇ (1929): 341 (characters); Wඳs ඍHඈൿൿ (1938): fig. 21 (morphology); Pൺඎඅංൺඇ (1941): 263 (larval characters); Hංඇඍඈඇ (1945): 70 (characters); FൺGൾඅ (1948): 196 (characters); Jൾൺඇඇൾඅ &amp; JൺඋඋංGൾ (1949): 378 (biology); Hൺඇඌൾඇ (1952): 140 (characters); Sආൾඍൺඇൺ (1958): 361 (characters and biology); (1962): 133 (characters and distribution); (1964): 79 (distribution); (1976): 22 (distribution); (1978c): 85 (distribution); (1993): 50 (distribution); Pൺඅආ (1963): 139 (characters); LඈHඌൾ (1964):211 (characters); Hඈඋංඈඇ (1965): 272 (distribution); Pඈඍඈඍඌĸൺංൺ (1967): 84, 87 (larval characters); Kൺඌඎඅൾ (1968): 53 (biology); (1970): 63 (larval characters); KඈඋGൾ (1968): 52 (characters); Sඓඎඃൾർĸං (1968): 735 (distribution); Fඋൺඇĸ (1969): 266 (biology); (1982): 46 (list of parasites); Bඈඋൽඈඇං (1974): 12, (1976a): 89 (characters); (1976b): 238 (distribution); Oඌൾඅඅൺ &amp; Zൺඇൾඍඍං (1975): 120 (biology); Pඈඉൾ (1977): 31 (distribution); Cඈංൿൿൺංඍ (1978): 190 (distribution); Tඈඉඉ (1979): 19 (development); Bඎඋൺĸඈඐඌĸං et al. (1980): 134 (distribution); TඬඍH (1984): 117 (characters); Nඈඐඈඌൺൽ (1990):143 (biology);Aඅඅൾඇ (1990): 4 (biology); WHංඍൾHൾൺൽ (1991): 6 (biology); Gඎඌൺඋඈඏ (1991):9 (identity); (1993): 72 (synonymic notes, lectotype designation); WൾඅർH (1993): 229 (morphology); HඈൽGൾ &amp; Jඈඇൾඌ (1995): 42 (characters); Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං (1995):32 (distribution); Sඍൺඇංൾർ (1996):117 (pupal characters); (2010): 22 (distribution); Oඐൾඇ (1997): 149 (biology); Sඍൺඇංൾർ (1999): 52 (characters of pupa); Dൾඋඎඇĸඈඏ (2005): 279 (distribution); UHඅංG et al. (2006): 54 (distribution); Tඋඈඇඊඎൾඍ (2006): 102 (distribution); Mൺඃĸൺ &amp; Sආൾඍൺඇൺ (2007): 428 (distribution); Mൺඓඎඋ et al. (2007): 29 (distribution);Sඍൺඇ (2009):242 (distribution);Sൾආൾඇඈඏ et al.(2015): 127 (distribution); Sൾආංඈඇൾඇĸඈඏ et al. (2015): 331 (distribution); Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2018a): 125, (2019): 48 (distribution); PඎർHĸඈඏ et al. (2020): 43 (distribution).</p><p>Type material examined. Quedius fuliginosus: Lൾർඍඈඍඒඉൾ J (ZMHB), labelled: “coll. Hellwig / Paratypus fuliginosus Grav. ”.</p><p>Quedius granulipennis: Lൾർඍඈඍඒඉൾ J (ZMMU), labelled: “ Austria / Quedius granulipennis Motsch. Austria ”.</p><p>Quedius latus: Nൾඈඍඒඉൾ J (ZMHB), designated here, labelled: “AR-MENIA (AR-16-19) 35 km NW Sisian, 39°40’59”N 45°46’50”E, 2070 m, stream valley, litter beneath bushes near stream sifited 3.VII.2016, leg. M.Schülke / Quedius (Quedius) fuliginosus (Gravenhrost) det. M. Schülke 2019 / Museum für Naturkunde Berlin Sammlung M. Schülke / Neotype Quedius latus Hochhuth, 1851 A.K. Hansen &amp; A. Solodovnikov des. 2021”.</p><p>Quedius molochinicolor: Sඒඇඍඒඉൾ ♀ (SNMC), labelled: “ Slovensko Harmanec Roubal 26.IX.1930 / f. molochinicolor mi [my] type / [red label] / fuliginosus f. molochinicolor cotypus”.</p><p>Additional material examined. ARMENIA: N Yerevan, NW Hrazdan, 40.6350 44.4602, grassy W slope with scattered Salix, litter and roots of grass sifted, 2500m, 26.VI.2016, leg. M. Schülke (1 ♀ cSch); same locality, 40.6944, 44.4878, stream valley, mixed decidious forest, litter and grass roots sifted, 2110m, 28.VI.2016, leg. M. Schülke (1 ♀ cSch). AUS-TRIA: Arbesbach; [48.49, 14.95], 1.-13.VII.1951, leg. Schubert (1 ♀ NMW); Brunnlust, Moosbrunn, Schwechat, [48.01, 16.45], 12.VI.1997, leg.Schillhammer (1J NMW);Faak, Kärnten,[46.56, 13.91], 5.VIII.1968, leg. Wewalka (1 ♀ NMW); Immerkrems, [46.96, 13.72], subalpine, 1400 - 2000 m, VI.1954, leg. Lindroth (1 ♀ MZLU); Plesch-Kogel n. Rein, [47.13, 15.28], 590m, sifting forest litter, 20.VI.1995, leg. Zerche &amp; Behne (1 J SDEI); Rekawinkel, [48.17, 16.02], leg. Briet (1 J NMW). BULGARIA: Sofia, Tschernia [Cherni Vrah], [42.56, 23.27], 600m, 22.VI.1988,leg.Behne (1J SDEI); Sredna Gora,S Koprivschtiza, 42.8777, 25.0544, moss under Picea, 1000m, 29.VI.1997, leg. Zerche &amp; Behne (1 ♀ SDEI); Stara Plania, 5km S Ribaritza, 42.75, 24.38, 750m, 7.VI.1997, Beech forest, sifting, leg. Zerche &amp; Behne (1 J SDEI); SW-Pirin, Kurort Popina Laka, 16 km NO Sandanski, [41.56, 23.27], ca. 1350m, Pinus and Picea, under snow remains, sifted, 8.IV.2005 leg. Zerche &amp; Behne (1 J SDEI); W Rhodopen, S Velingrad, Tschernowroh, [41.68, 24.05], 1600m 10.VI.1987, leg. Zerche &amp; Behne (1 ♀ SDEI). CZECH REPUBLIC: Podyjí Nat. Park, forest at river Dyje valley ca. 2.5 km NW of Havraníky, 48.8300, 15.9804, h 240m, sifted flood debris, 8.VI.2016, leg.A. Solodovnikov, J. Jenkins Shaw,A. Hansen (1 ♀ NHMD). DENMARK: Amager, Kalvebod Faelled, Store Høj Søen, 55.6159, 12.5606, sifting leaf litter, debris near lake, 17.III.2019, leg.A. K. Hansen (1 J 1 ♀ NHMD); Lisjerg Forest, 8 km N of Aarhus, 56.2335, 10.1688, h 85m, edges of forest lakes, hand collected in leaf litter and moss, 17.IV.2019, leg.A. K.Hansen (4JJ 4 ♀♀ NHMD); Nors Sø, S of Nors Sø, 57.0197, 8.6197, pitfall trap, dry pine forest, 16-18.VI.2014, leg.Solodovnikov, Hansen, Kraemer, &amp; Justesen (1 J NHMD). GEORGIA: N of Sviri, [41.73, 42.99], pitfall traps, 14.IV.-11.VI.2018, leg. D. Formynikh (1 J cSch); Tusheti, n. Omalo, Abanos Pass, [42.27, 45.51], river valley, 1700m, 14.-17.VII.2016, leg. Heinz (1 J NMEG). GERMANY: Bayern, Eschenlohe, [47.58, 11.18], 3.VIII.1937,leg.Ihssen (1J 1♀ SDEI);Berlin-Blankefelde,[52.62,13.39], 8.V.1994, leg. D. Wrase (1 J SDEI); Biesenthal, [52.76, 13.61], 500m, field sandish-clayish ruderal, 1.IV.2002, leg. D. Wrase (1 ♀ SDEI); Brandenburg Kiesfläche 1km S Oderberg, [52.84, 14.04], 13.X.1990, leg. F. Hieke (1 J SDEI); Brandenburg, Oranienburg, Briesetal, [52.71, 13.31], 12.IV.1992, leg.J. Ziegler (1J SDEI); Dars, Zingst, Freesenbruch, [54.43, 12.68], 20.VII.1987, leg. D. Wrase (1♀ SDEI); Erzgebirge,Prinzgehöhle, [50.63, 12.67], 29.IX.1921, leg. Uhmann (1 ♀ SDEI); Erzgebirge, W Teil, SW Kammweg &amp; Morgenröthe, [50.43, 12.51], 4.V.1990 leg. O. Sorge (1 J SDEI); Finkenkrug n. Berlin, [52.56, 13.03], 12.IV.1898, leg. Dahl (1 J SDEI); Kehlheim Dürnbacher Forst, [48.74, 11.75], (1 ♀ SDEI); Liepnitzsee, n.Wandlitz,[52.74, 13.51], 01.X.1989, leg. M. Uhlig (1J SDEI); Marburg i Hess, [50.81, 8.77], 16.IV.1904, leg. Strand (1 J 1 ♀ SDEI); n. Märkische Buchholz, Königs Wusterhausen, [52.11, 13.76], 12.VI.1980, leg. H.Wendt (1 ♀ SDEI); Martinfeld, Krs. Heiligenstadt, [51.28, 10.18], 21.-29.VI.1990, leg. M. Uhlig (1 J SDEI); München-Grünwald, [48.04, 11.53], 3.VII.1931, leg. Ihssen (1 ♀ SDEI); Nauen, [52.61, 12.87], 19.III.1910, leg.M.Ude (1J SDEI);Sächsischen Schweiz, Rathen,[50.95, 14.08], 1.V.1977, leg.K. H.Mohr (1♀ SDEI); Schwäbisch Gmünd,Röben, [48.79, 9.81], 4.V.1953 (1 J SDEI); Seilershof, Gransee, [53.06, 13.18], 22.V.1988, leg. M. Schülke (1 ♀ SDEI); Suekow, [53.07, 11.79], XI.1974, leg. M. Moritz (3 JJ SDEI); Wandlitz, Liepnitzsee, [52.74, 13.51], 6.I.1981, leg. M. Uhlig (1 J 2 ♀♀ SDEI); Wildeschütz n. Deuben Silbersee Schilfgürtel, [51.12, 12.07], 11.X.1987, leg. M. Uhlig (1 ♀ SDEI). ITALY: Alpie Orobie,Bergamo Castione &amp;Passo delia Presolana, 45.9202, 10.1001, 1170m, 18.-21.VI.2006, leg.P. Schnitter (1 J NMEG); Basilicata, Pognola Ris. L. Pignola, [40.58, 15.74], 700 m, 23.XII.1996, leg. F. Angelini (1 cBor); Trentino, Lago di Tenno,N Riva, [45.93, 10.81], 560m, 2.VII.1994, leg. L. Zerche (1 ♀ SDEI). KAZAKHSTAN: Altai, Bukhtarma riv., Chingistai, [49.19, 85.88], 30.VIII.2010, leg. V.A. Kastcheev (1 J ZIN);Akshatau Mt., NW Ayaguz, Semipalat, [48.25, 79.64], forest leaf litter, 17.VII.1962, leg. L. V.Arnoldi (1 J ZIN); Ivanovsky Mt. Ridge, 32 km S Leninogorsk, [50.04, 83.51], 1300 m, 14.VIII.1986, leg. I. I. Kabak (1 J ZIN). RUSSIA: Aඅඍൺං Kඋൺං: Talmenskij Distr., E of vill. Litvinovka 53.6918, 83.7309, 200m, mixed forest, sifted leaf litter, 24.VI.2019, leg. A. Solodovnikov, A. K. Hansen &amp; A. Tokareva (1 NHMD); Chemalskij Distr., SE of vill. Edigan 51.0546, 86.3791, 1030m, larch-fur forest, sifted leaf litter, 27.VI.2019, leg.A. Solodovnikov &amp; A. K. Hansen (1 NHMD); Chemalskij Distr., Verkhnij Berulu river, 51.3035, 86.1221, 790m, pine-birch forest, sifted leaf litter, 28.VI.2019, leg. A. Solodovnikov, A. K. Hansen &amp; M. J. Justesen (1 NHMD). BൺඌHKඈඋඈඌඍൺඇ Rൾඉ.: Buzdyakskij Distr., nr vill. Arslanovo 54.6485, 54.3789, 229m, at creek bank, 11. VI.2019, leg.A. Solodovnikov &amp; A. Tokareva (1 NHMD); Mechetlinskij Distr., Oka river nr vill. Bolshaya Oka, 56.1085, 58.1977, 224m, debris and ground at river bank, 12.VI.2019, leg.A. Solodovnikov &amp; A. Tokareva (1 NHMD). IඋKඎඍඌK Oൻඅൺඌඍ: Baikal, Chamar-Daban, Solsan Valley, [51.36, 104.15], 17.-21.VI.1978, leg. Schilenkov (1 J cSch); Baikal, Baikalsk, Solsan River near mouth, [51.47, 104.37], 20.VII.1989, leg. Hieke (1 J 1 ♀ SDEI); Khamar-Daban Mts., valley of Snezhnaya River, [51.06, 103.99], 5.-8.VI.2007. leg.A. Shavrin (3 JJ 6 ♀♀ cSha); Podvoloshino, Valley of Nizhnyaya Tungska River, [58.25, 108.42], 4.-9. VIII.2008, leg.Shavrin &amp; Enustschenko 18 km N Ust-Kut,Valley of Lena River, [56.95, 106.14], 26.-28.VII.2008, leg. Shavrin &amp; Enustschenko (1 ♀ ZIN) Valley of Bodchakta River, [57.11, 106.61], 10.-11.VIII.2008, leg. Shavrin &amp; Enustschenko (1 J ZIN). KൺඅඎGൺ Pඋඈඏ.: 4-5 km SE from Chernaya Grayz, [54.96, 36.86], V.1988, leg. I. Ushakovl (1 J cRyv); Kozelsk Reg., Berezich glass factory, Novaya Derevnya village, NR “Ugra”,[56.27, 40.25], broad leaved forest,leaf litter near stream (stagnant channel), 13.VII.2017, leg. M. Salnitska (1 ♀ ZIN). KൾආൾඋඈඏඌKൺඃൺ Oൻඅൺඌඍ: Tashtagolsky Distr.,Shorsky Nat.Park, 5 km SSE Tajmet [Verkhnij Tajmet] village, [52.48, 88.27], cedar forest, 8-18.VI. 2012, leg. L. A. Trilikauskas (1 J 1♀ ZIN). KඋൺඌඇඈඒൺඋඌKංඃ Kඋൺං: nr.Divnogorsk town, left tributary of Enissey river, [55.95, 92.38] cedar-birch forest, in leaf litter, 1.VI.1988, leg.A. B. Ryvkin (2 JJ ZIN); Enissey distr., environs of vill. Ust-Pit, [58.98, 91.76]‚ 02-10.VII.1995, leg. A. Rybalov (2 JJ 1 ♀ ZIN); Nature Preserve ‘Stolby’ [Kranoyarskie Stolby], river Mana near Berly hut, [54.98, 94.00], stream edge, in moss and litter, 20.VI.1990, leg. A. B. Ryvkin (1 J ZIN); Niznaya Lebedyanka river, [61.96, 89.45], 22.VI.1992, leg.V.S. (2JJ cRyv). LൾඇංඇGඋൺൽ Oൻඅൺඌඍ: Kirovsky Distr., Turishkino vill., Mga River, 59.684306, 31.2202, flood plain, meadow in forest, leaf litter, 02.VI.2018, leg. M. Salnitska, K. Fadeev (1 J ZIN); Leningrad [Sankt Petersborg], in litter, 29.VII.1976, leg. Schilow (3 JJ SDEI). Mඈඌർඈඐ Oൻඅൺඌඍ: Odintsovsky Distr., Zvenigorod Biological Station of MSU, [55.70, 36.72], soil sample, Betula forest, 7.VIII.1981, leg. K. G. Mikhailov (1 J cRyv). NඈඏGඈඋඈൽ Oൻඅൺඌඍ: Staraja Russa, [57.99, 31.35], leg. Kessler (1 J NMW). Pൾඋආ Kඋൺං: Perm town, ravine meadow on the right bank of Kama river, Motovilikha, [58.04, 56.30], 1989, leg. V. O. Kozminykh (1 J ZIN). Rൾඉ. ඈൿ AൽඒGൾൺ: Caucasus, Adygeja, Guzeripl, Belaja River, [43.99, 40.13], 2000m, 23.VI.1999, leg. Putchkov (1 ♀ cSch). Rൾඉ. ඈൿ N. Oඌඌൾඍංൺ- Aඅൺඇංൺ: N Ossetia Nat. Res., mt. Kariuchoch, Kora pass. (Alansky), 42.8301, 44.2123, ~ 2200m, leaf litter Betula sp., 24.V.2017, leg. M. Salnitska (1 J ZIN). SඏൾඋൽඅඈඏඌK Oൻඅൺඌඍ: Visimskij Nature Res., 57.3733, 59.7734, 570m, spruce dominated forest, pitfalls, 18.VI.2019, leg. A. Solodovnikov, A. K. Hansen &amp; A. Tokareva (1 NHMD); W of vill. Elan’, 57.6406, 63.6220, 100m, mixed forest, sifted leaf litter, 20.VI.2019, leg.A. Solodovnikov,A. K. Hansen &amp; A. Tokareva (1 NHMD). Tඒඎආൾඇ Oൻඅൺඌඍ: Uvatsky Distr., 8-14 km S from vill. Gornoslinkino, near Tobolsk station of IEE RAS,[58.68, 68.79], 11.VI.2004, leg. A. B. Ryvkin (2 JJ 1 ♀ cRyv); Tyumenskij Distr., Tura river nr vill. Konyashina, 57.2925, 65.0281, 80m, flood forest, sifted leaf litter, 21.VI.2019, leg. A. Solodovnikov, A. K. Hansen &amp; A. Tokareva (1 NHMD); nr Zavodoukovsk, 56.5329, 66.5689, 116m, pine-birch grove, sifted leaf litter, 21.VI.2019, leg. A. Solodovnikov, A. K. Hansen &amp; A. Tokareva (1 NHMD). SLOVAKIA: Belá River n. Vavrisovo n. Lipt. Hrádek,[49.06, 19.77], 18.VII.1980, leg. Hieke (12JJ 1♀ SDEI); Gombasek n. Rožňava, [48.57, 20.45], 6.VIII.1981, leg. Hieke (1 ♀ SDEI); Harmanec, [48.79, 19.07], 26.IX.1930, leg. Roubal (1 ♀ SNMC); Pieninsky National Park, [49.38, 20.51], 18.VII.1993, leg. Zerche (1 J SDEI); Rudohorie, Hnilec-Tal n. Švedlár, [48.83, 20.51], 8.VII.1981 leg. Hieke (1 ♀ SDEI). SLOVENIA: Triglav Nat. Park, river Soča valley, ca. 3 km SW Trenta; Obcina Bovec, 46.3667, 13.7255, sifting flood debris and under stones, 03.VI.2016, leg. A. Solodovnikov, A. Hansen. &amp; M. Salnitska (1 NHMD). SWEDEN: Månsatorp, 5km SW of Vittsjö, Skåne; 56.3105, 13.5955, hand collected in/under moss, 21.I.2018, leg. A. K. Hansen (1 J 2 ♀♀ NHMD); Ödeshög V, Ekopark Omberg, Storpissan nature reserve, 58.3344, 14.6508 sifting leaf litter on stream edge; 30.V.2017, leg.D. Żyła, K. Koszela, A. Solodovnikov (1 J NHMD); Vegesjö, Skåne, 56.3031, 13.6473, banks of bog in wet sphagnum, 100 m, 22.IV.2018, leg. A. K. Hansen (1 ♀ NHMD). TURKEY: Harçbeli Pass, Ordu, [40.59, 37.64], 1900 m, 28.V.1989,leg. Schönmann &amp; Schillhammer (1♀ NMW); Sümela, Altındere, [40.68, 39.65], 10.-11.VI.1969, leg. P. Brignoli (1 ♀ cBor). UNITED KINGDOM: EඇGඅൺඇൽ: Norwich, Norfolk, UEA Campus, [52.62, 1.23], 3.III.2013, leg. C. Billet (1 J cJen). UKRAINE: Kanev [Kaniv] near Kiew [49.75,31.46], 28.I.1988, leg. Zerche (2 ♀♀ SDEI); Ivano-Frankovo, ca. 40 km WNW of Lvov, Rostochye, State Reserve, [49.94,23.65] Quercus, Pinus etc.forest, 16-20.IX.1999, leg.S. Golovatch (1 J cRyv); N Yablunytsia vill., S Gorgany Mnts., valley Prut riv., [48.34, 24.45], 800m, pitfalls, 22-24.V.2018, leg.Panin R.(1J cGon). UZBEKIS-TAN: Tashkent, [41.29, 69.25], near railway station, plant residues, 24.V.1986, leg. S. A. Kurbatov (1 J 1 ♀ ZIN).</p><p>Redescription. Measurements JJ (n = 5): HW = 1.78– 1.89 (1.81); HL = 1.40–1.51 (1.46); HL/HW 0.79–0.83 (0.80); PW = 2.22–2.42 (2.31); PL = 2.04–2.20 (2.13); PL/PW 0.90–0.98 (0.92); EW = 2.44–2.51 (2.48); EL = 2.29–2.38 (2.32); EL/EW 0.91–0.96 (0.93); EL/PL 1.05– 1.14 (1.09); PW/HW 1.54–1.60 (1.58); forebody length 5.78–6.09 (5.91). ♀♀ (n = 5): HW = 1.80–1.96 (1.84); HL = 1.42–1.51 (1.46); HL/HW 0.77–0.81 (0.80); PW = 2.22–2.42 (2.32); PL = 2.09–2.13 (2.10); PL/PW 0.88–0.94 (0.91); EW = 2.40–2.60 (2.50); EL = 2.29–2.44 (2.34); EL/EW 0.91–0.97 (0.94); EL/PL 1.08–1.15 (1.11); PW/ HW 1.54–1.66 (1.59); forebody length 5.82–6.09 (5.90).</p><p>Large species; body black (Fig. 7C).</p><p>Head black, clearly transverse, with eyes very large (EyL/TL = 3.00–3.90 (3.54)) and clearly protruding, microsculpture of transverse waves, four punctures between anterior frontal punctures on frons (Fig. 6C); antennae and palpi darkened, except the joints, which are slightly paler, antenna with all antennomeres elongate. Thorax: pronotum black, slightly wider than long, wider than head, microsculpture of transverse waves, three punctures in dorsal row and two in sublateral row with the posteriormost puncture not reaching level of second puncture of dorsal row; scutellum smooth and glabrous, elytra black, uniformly pubescent, slightly wider than long, slightly longer than pronotum; legs dark brown to black with tarsi slightly paler.</p><p>Abdomen black, tergites uniformly punctured, clearly iridescent.</p><p>Male. Aedeagus (Fig. 11D): paramere broadly lanceolate, with clear medial attenuation, sensory peg setae forming two subapical, longitudinal rows along paramere edge apically but slightly turning inwards at half their length. Median lobe with two small but distinct teeth in front of basal part of parameral peg setae. Internal sac without clear sclerites.</p><p>Differential diagnosis. Even though Quedius fuliginosus differs from all other members of the fuliginosus -group by having the basal antennomeres at least slightly darkened, this is not always obvious in teneral or old museum specimens as coloration is altered. It is easily distinguished from Q. levicollis by the glabrous scutellum. It can be safely distinguished from Q. curtipennis by the aedeagi. In Q. fuliginosus, contrary to Q. curtipennis, the paramere has clear medial attenuation, a median lobe with the apical part being robustly pyramid-shaped and not laterally flattened, and an internal sac with no clear sclerotization. Subtle external difference between Q. fuliginosus and Q. curtipennis includes slightly stouter antennae, more transverse head, larger eyes clearly protruding thus making the head shape more quadrate, as well as slightly longer elytra in Q. fuliginosus . Also, Q. fuliginosus is very similar to Q. afrofuliginosus, but is easily recognized from the latter by having no additional punctures between anterior and posterior frontal punctures. It is also unclear if the two species have a distributional overlap.</p><p>Synonymic notes and neotype designation. Quedius fuliginosus was described from environs of Braunschweig, North Germany (GඋൺඏൾඇHඈඋඌඍ 1802). Two syntypes from the collection of Johann Christian Ludwig Hellwig (1743–1831) were revised by Gඎඌൺඋඈඏ (1993), who designated a lectotype to represent our current concept of Q. fuliginosus, while the paralectotype was identified as Q. curtipennis . Gඎඌൺඋඈඏ (1993) also studied the type of Quedius granulipennis; he found it to be a junior subjective synonym of Q. fuliginosus .</p><p>Quedius latus was described from “Alagez-Gebirge” (HඈർHHඎඍH 1851), the Aragatz mountain area in Armenia and has never been recorded since the original description. Based on the rather extensive faunistic knowledge of the Caucasus region, Q. latus could either be conspecific to Q. fuliginosus or Q. curtipennis, both common in the area (AඌඌංඇG &amp; SർHඳඅĸൾ 2019) and both fitting the original description of Q. latus . Since no type or any other authentic material of Q. latus was found in Hochhuth’s collection in Kyiv, Ukraine (S. Glotov, pers. comm.), apparently it is lost. To secure current and future stability of the name Quedius latus Hochhuth, 1851, in accordance with the ICZN, Article 75, we designate its neotype. For this, a male specimen of Q. fuliginosus collected in the vicinity of the original type locality (for details see Type Material section above) has been selected. The neotype fits the original description of Q. latus and thus allows us to consider the so far dubious name Quedius latus Hochhuth, 1851 syn. nov. as a junior subjective synonym of Quedius fuliginosus Gravenhorst, 1802 .</p><p>The specimen we studied from the syntype series of Quedius molochinicolor Roubal, 1931 represents Quedius fuliginosus with slightly brownish elytra, thus confirming the currently accepted synonymy.</p><p>Bඋංඍඍൾඇ (1944) recognized that there were two closely related species of Quedius s.str. in Britain. Based on his descriptions and illustration, it is clear that Q. fuliginosus sensu Britten is Q. curtipennis while the new species he described, Q. subfuliginosus, is identical to Q. fuliginosus . This is in agreement with the currently accepted synonymy.</p><p>We were not able to check the types of Quedius impunctifrons Delahon, 1915 and Quedius jelineki Krása, 1904 from Germany and Czech Republic, respectively. Both names are currently in synonymy with Q. fuliginosus . Since the descriptions state that they lack the interocular punctures on frons characteristic of the Q. fuliginosus -group, they either represent variability in chaetotaxy – or more likely, they are dark variants of Q. molochinus, the latter possibility calling for a revised synonymy.</p><p>Sൺඐൺൽൺ (1965) described Q. viduus as being very similar to Q. fuliginosus, but different in the broader head, larger and more convex eyes, pronotum with lateral sides less convergent anteriorly, broader elytra, and rougher punctation of abdomen. This original description based on the single female specimen is the only record of Quedius s. str. from Japan. In spite of the thorough searches in the collections of National Museum of Nature and Science, Tokyo (Shuhei Nomura), Osaka Museum of Natural History, Osaka (Shigehiko Shiyake), Nakane Collection at Hokkaido University (Masahiro Ohara), and Sawada personal collection (Yasuhiko Hayashi), all performed by respective curators upon our request, the holotype of Q. viduus could not be located. Based on the original description and lack of any other species of Quedius s. str. in Japan, we suspect this is either a mislabeled specimen not from Japan, or an introduced specimen of Q. fuliginosus, which was not established or recollected in Japan. Thus, Q. viduus Sawada, 1965 syn. nov. is considered as a junior subjective synonym of Q. fuliginosus Gravenhorst, 1802 .</p><p>Bionomics. Quedius fuliginosus is widespread and eurytopic species found in a variety of forests (both deciduous and coniferous) and in the open habitats too. Usually, it occurs in moss and leaf litter or in various other kinds of ground debris (Sඈඅඈൽඈඏඇංĸඈඏ 2012). We have commonly collected this species in moss near the edge of a lake in Denmark (Fig. 2A), in various forests often with dense fern or grassy undergrowth, across larger parts of Western Siberia (Fig. 2E), in flood debris of a forested mountain stream in Triglav, Slovenia and along river forested on either side in the southern Czech Republic. It seems to prefer mossy patches and moist litter, and is thus to some degree often associated with water bodies, although it is not found at the very edge of these. It has also been reported from mole ( Talpa europaea) nests (Nඈඐඈඌൺൽ 1990), from bracket fungi ( Polyporus squamosus) (Sൾආൾඇඈඏ et al. 2015), and from ant nests (Gඈඋൾඌඅൺඏൾඍඌ 2010, 2016), all of which are presumably accidental occurrences, as there is no indication of the species preferring these habitats.</p><p>Larvae were described by Pൺඎඅංൺඇ (1941), Pඈඍඈඍඌĸൺඒൺ (1967) and Kൺඌඎඅൾ (1970). Pupae of the species are described in Sඍൺඇංൾർ (1999); Sඓඎඃൾർĸං (1966) described the egg. Adult females from Wytham, UK were observed containing eggs between May and June (Fඋൺඇĸ 1969).</p><p>Distribution. Quedius fuliginosus can be found across a large part of the Palearctic region (Fig. 18). In Europe it is common in the North-East, but becomes rarer south--westwards and is probably absent from the southern parts of the Iberian Peninsula and Italy. It is found across most of Siberia from the Urals to at least the Baikal region. In the southern part of the range the species is confined to the forested zone in mountains. Here it is found along the Pyrenees, the Alps, the North Anatolian mountains, the Caucasus, and Transcaucasia. Eastwards it extends to the western forested regions of the Tian Shan Mountain Range in central Asia. The northern distributional border of the species seems to be determined by the forested boreal zone, as it does not continue into the tundra.</p><p>Records of Q. fuliginosus from Tunisia and Algeria (Fൺඎඏൾඅ 1902) most likely refer to either Q. levicollis or Q. afrofuliginosus, as we did not come across any specimens of this species from northern Africa or nearby regions. Moreover, Q. fuliginosus is apparently absent in the south Mediterreanean coastal areas, which makes its presence in North Africa even more unlikely.</p><p>Introduced to eastern North America in an area of Nova Scotia where the first specimens were found in 1996 (Mൺඃ- ĸൺ &amp; Sආൾඍൺඇൺ 2007) (Fig. 12).</p></div>	https://treatment.plazi.org/id/038987A0FFED4B02B391A418119CFEEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFE14B07B3AFA19817ADFF6A.text	038987A0FFE14B07B3AFA19817ADFF6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius levicollis (Brulle 1832)	<div><p>Quedius levicollis (Brullé, 1832)</p><p>(Figs 1, 2B, 3, 6D, 7D, 11B, 19)</p><p>Staphylinus levicollis Brullé, 1832: 131 [Type locality: Morée: plaine de Modon]</p><p>Philonthus frontalis Nordmann, 1837: 76 [Type locality: Gallia; Symperopoli]</p><p>Quedius balthasari Mařan, 1935: 9 [Type locality: Knossos, Creta]</p><p>Staphylinus nigritarsis Ménétriés, 1832: 144 [Type locality: Caucase]</p><p>References. GඋൺඏൾඇHඈඋඌඍ (1802):34,(1847):231 (characters); SඍൾඉHൾඇඌ (1832): 215, (1839): 389 (characters); Fൺඅൽൾඋආൺඇඇ (1835): 123 (characters); Lൺർඈඋൽൺංඋൾ (1835): 376 (characters); Nඈඋൽආൺඇඇ (1837): 75 (distribution); Hඈඅආൾ (1837): 63 (distribution); Wൾඌඍඐඈඈൽ (1838): 167 (larva and pupa); EඋංർHඌඈඇ (1840): 537 (characters); Kඈඅൾඇൺඍං (1846): 22 (characters); Kංൾඌൾඇඐൾඍඍൾඋ (1848): 54 (characters); HඈർHHඎඍH (1849): 154, (1862): 46 (characters); RൾൽඍൾඇൻൺർHൾඋ (1849): 709, (1857): 201, (1874): 198 (characters); Fൺංඋආൺංඋൾ &amp; Lൺൻඈඎඅൻජඇൾ (1856): 536 (characters); RඈඌൾඇHൺඎൾඋ (1856): 71 (distribution); Kඋൺൺඍඓ (1857): 502 (characters); Fൺංඋආൺංඋൾ &amp; Cඈඊඎൾඋൾඅ (1860): 154 (distribution); Fൺඎඏൾඅ (1865): 293 (ecology); (1874): 289 (characters and distribution); (1878): 128, (1886): 58, (1902): 115 (distribution); Mඎඅඌൺඇඍ &amp; Rൾඒ (1876): 688 (characters); Fඈඐඅൾඋ (1888): 237 (characters); GൺඇGඅൻൺඎൾඋ (1895): 403 (characters); Sൺංඇඍൾ- Cඅൺංඋൾ Dൾඏංඅඅൾ (1906): 102 (distribution); Pඈඋඍൺ (1907): 129 (characters); Rൾංඍඍൾඋ (1909): 111 (characters); JඈHൺඇඌൾඇ (1914): 363 (characters); Mඎඇඌඍൾඋ (1923): 196 (distribution); Gඋංൽൾඅඅං (1924):80 (characters and distribution); Pඈඋඍൾඏංඇ (1929):341 (characters); WඳඌඍHඈൿൿ (1938): fig. 22 (characters); TඈඍඍൾඇHൺආ (1949): 376 (characters); Hൺඋൽඒ (1851): 32 (distribution); Hൺඇඌൾඇ (1952): 142 (characters); SർHൾൾඋඉൾඅඍඓ (1956): 1102, (1965a): 407, (1965b): 511 (distribution); (1961): 133 (characters); Sආൾඍൺඇൺ (1958): 362 (characters and ecology); (1962): 134 (characters and distribution); (1963): 39, (1967): 558, (1970): 62, (1993):50 (distribution);(1978b):121 (distribution and ecology); Pൺඅආ (1963): 141 (characters and distribution); LඈHඌൾ (1964): 211 (characters); Fඈർൺඋංඅൾ (1964): 61 (distribution); Hඈඋංඈඇ (1965): 273 (distribution); Cඈංൿൿൺංඍ (1970):62 (distribution); (1978):192 (characters and distribution); Bඈඋൽඈඇං (1976a):93 (characters);(1982):140 (ecology); (1984):342 (distribution); Pඈඉൾ (1977): 31 (characters and distribution); Hංඇඍඈඇ (1981): 662 (characters of egg); TඬඍH (1984): 118 (distribution); Fඋൺඇඓ (1987): 70 (distribution); Gඎඌൺඋඈඏ (1993): 73 (synonymy); WൾඅർH (1993): 229 (ovary structure); Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං (1995): 33 (distribution); Nൾඐඍඈඇ (1995): 51 (requested to ICZN to place Q. levicollis on the Official List of Specific Names in Zoology); ICZN (1996): 215 ( Q. levicollis placed on Official List of Specific Names in Zoology); Bඅൺඇൽ &amp; Sංඇർඅൺංඋ (1997): 204 (biology); MൺඋർHൺඅ &amp; CH ൺඋൽඈඇඇൾඍ (2000): 72 (distribution); Oඐൾඇ (2000): 252 (ecology); AඌඌංඇG &amp; Wඎඇൽൾඋඅൾ (2001): 37 (distribution); UHඅංG et al. (2006): 54 (distribution); Tඋඈඇඊඎൾඍ (2006): 102 (distribution); Aඇඅൺ෡ (2009): 229 (distribution); Sඍൺඇ (2009): 242 (distribution); Vඈඋඌඍ et al. (2009): 16 (distribution); Mൺඃඓඅൺඇ &amp; BඈHගඹ (2012): 182 (distribution); Bൾൻൾඋආൺඇඌ et al. (2016): 135 (distribution); ÖඓGൾඇ et al. (2016): 620 (distribution); Rඎඍൺ et al. (2016): 37 (distribution); Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2019): 49 (distribution).</p><p>Type material examined. Staphylinus nigritarsis: Lൾർඍඈඍඒඉൾ: designated by Gඎඌൺඋඈඏ (1993) (ZIN), labelled: “[golden square] / Caucas. / nigritarsis Menet. Cauc ”. Pൺඋൺඅൾർඍඈඍඒඉൾ: labelled: “[golden square] / Caucas.” (1 J ZIN).</p><p>Additional material examined. ALBANIA: 13km SE Elbasan, Mali i Shpatit, Mt. Bukanikut, 41.0431, 20.2172, brook bank in beech forest, 1500m, 22.V.2010, leg. D.W.Wrase (1J cSch). BOSNIAAND HERZE-GOVINA: Maklen-Pass,[43.85,17.65], 1902, leg.O.Leonhard (1 SDEI); Prozor, [43.82, 17.61], 1902, leg. O. Leonhard (1 SDEI). BULGARIA: Arkutino, S of Burgas, [42.41, 27.65], 28.V-9.VI.1971, leg. Wallis (1 J SDEI). CROATIA: Capela [Kapela], [44.89, 15.52], leg. Heyden (1 SDEI); Velebit Mountains, [44.52, 15.22], wine traps, V.2005 (1 J cJen). CYPRUS: Agios Nikolaios, [34.86, 32.75], 16.IV.1995, leg. H. Schmid (1 J NMW); Troodos Mountain, Stavros tis Psokas, [35.02, 32.63], 800- 950m, 12.-18.IV.2010, leg. I. Wolf (1 J cSch). DENMARK: Ovestrup Hede, 56.2412, 8.9297, pitfall trap, heathland grazed part, 7-22.VI.2017, leg. E. Kristensen, D. Byriel, &amp; M. Justesen (1 J NHMD). FRANCE: Avignon, [43.94, 4.81], 22.V.1949, leg. R. Skovgaard (1 J NHMD); Chigny n. Oise, [49.91, 3.76], 19.X.1918, leg. Salchert (1 ♀ ZMHB); Paris, [48.83, 2.41], 19.V.1949, leg. R. Skovgaard (1 J NHMD). Cඈඋඌൾ: Bocognano,[42.08,9.06],1905, leg.O.Leonhard (1 SDEI);Col de Bavella, [41.79, 9.23], 1240m, East slope, 25.VII.1994 leg. L. Zerche (1 J SDEI); Foret de Valdo-Niello, Evisa, [42.28, 8.91], 1000-1200 m, 27.VII.1967, leg. Langemark-Lomholdt (1 J NHMD). GERMANY: Berlin-Marzahn, Hellersdorfer Berg, [52.53, 13.58], 8.VIII.1997, leg. Hieke (1 ♀ ZMHB); Berlin, Prenzlauerberg Hof, [52.53, 13.46], 29.V.1985, leg. I. Uhlig (1 J ZMHB); Grunewald n. Zehlendorf, [52.48, 13.22], leg.Wagner (1 SDEI). GREECE: Attica, [38.05, 23.84], 17.IV.1922, leg. W. Liebmann (1 SDEI); Paleokastro, Halkidiki, [40.43, 23.42], 800m, 11.VII.1983, leg.A. Casale (1 ♀ cBor); 2 km W Panagia, Trikala, [39.79, 21.33], 15.VI.1982, leg. R Danielsson (1 J MZLU); Voria Pindos, Morfi, [40.21, 21.21], dry slope, 26.VI.1997, leg. Winkelmann (1 ♀ cSch); Cඈඋൿඎ: Pandokratoras, 39.7492, 19.8701,N-slope with scattered pine,decidious trees,and bushes, litter and roots between rocks sifted, 800 m, 4.VI.2017, leg. Schülke (1 J cSch);Pantokrator,[39.74,19.87], V.1964, leg.Palm (1♀ MZLU). Cඋൾඍൾ: Georgioupouli, [35.36, 24.26], 24.-25.IV.1995, leg. B. Jaeger (1 ♀ cSch); Ida mts., ca. 6 km S Analipsis, 35.1752, 24.8391, Quercus forest, 1260m, 21.X.2006, leg. M. Schülke (1 J cSch); Mt.Ida, 28km SW Iraklio, [35.23, 24.77], 1600m, 7.IX.1991, leg. N. Esser (1 ♀ NHMD). KൺඋඉൺඍHඈඌ: Karpathos, 35.5097, 27.2189, 40 m, terracod grassy fallow, 01.I.2016, leg. V. Assing (ZMHB). Lൾඌൻඈඌ: SW Agiassos, road to Olympos peak, [39.06, 26.36], 700 m, 10.VI.1996, leg. Jäch (1 J 1 ♀ NMW). Pൾඅඈඉඈඇඇൾඌ: Korinthia, Kyllini, Zoria, [37.94, 22.43], 2000 m, 3.VI.1995, leg. Giachino &amp; Vailati (1 J 3 ♀♀ cBor); Kyllini Mountain, [37.97, 22.39], 2100m, North side, below summit over valley, 9.VI.1996, leg. P. Wunderle (1 ♀ cSch); Sparta, [37.06, 22.43], 3.V.1962, leg. Skovgaard (1 J NHMD); N Taygetos, 6 km NE Thouria, 37.1401, 22.1284, litter and soil under bushes, sifted, 584 m, 20.IV.2015, leg. M. Schülke (1 ♀ cSch). RHඈൽൾඌ: Prophitis Ilias, 36.2734, 27.9287, pine forest, under stones &amp; sifted from litter &amp; moss, 685m, 17.V.2015, leg. Wrase &amp; Laser (1 J cSch). IRAN: Kohgiluyech-Va-Boyern-Ahmad province, 20 km SW Yasuk, 10 km SE Sepidor, [30.56, 51.51], 5.-6.V.2007, leg.A. Anistchschenko (1 ♀ SDEI). IRELAND: Inch Dingle Eire, [52.14, -9.97], IX.2012, leg. J.J. Shaw (1 J cJen); Pembury, Kent 51.1625, 0.3431, 24.VII.2011, leg. J.J. Shaw (2 JJ cJen). ISRAEL: Upper Galilee,Har Avivim,W Avivim, 33.0833, 35.4923, open stony grazing land limestone, 768m, 21.IV.2006, leg. D.W.Wrase (1 J cSch); Upper Galilee, Meron Mts. Har Meron, Kamin Rom, 32.9908, 35.4111, open stony grazing land, limestone, 1100m, 28.IV.2006,leg.D.W. Wrase (1J cSch). ITALY: CalabriaAspromonte,[38.21, 15.99],1905,leg. Pagametti (1 SDEI); Calabria, Roseto Capo Spulico, [39.98, 16.61], 20 m, 6.X.1998 leg.F.Angelini(1♀ NHMD); Dicomano, Toscana,[43.89,11.52], I.1971, leg Bordoni (1J cBor); Macerata, [43.29, 13.45], 1.X.1955, leg.D. Caccamicelo (1♀ cBor); Marche,Montemonaco, Isola San Biagio,[42.91, 13.31], fields, 900 m, 27.VII.-1.IX.2004, leg.M.Rismondo &amp; R. Fabbri (2 JJ 2♀♀ NHMD); same locality, black hornbeam forest, 1.IX.-7.X.2004, leg. M. Rismondo &amp; R. Fabbri (1 J NHMD); Modena, Prignano sulla Secchia, La Quercia, [44.45, 10.73], pitfall traps, 400 m, 5.-18.VII.2006, leg. R. Fabbri (1 J NHMD); Reggio di Calabria, river Sant Agata, [38.09, 15.66], 11.III.1959 (1 J cBor); Settecamini, Rome, Lazio, [41.93, 12.62], 23.Xi.1969 (1 J 1 ♀ cBor); Sovara, Anghiari, [43.53, 12.03], 19.V.1930, leg. A. Andreini (1 ♀ cBor). Sൺඋൽංඇංൺ: Aritzo Mountain, Gennargentu, [40.01, 9.29], leg. C.Kruger (2 SDEI); Ottana, [40.23, 9.04], 15.X.1974 (3 JJ 1♀ cBor);Oristano,[39.89,8.59], 3.V.1975,leg.S. Ripze (2♀♀ cBor); Sංർංඅඒ: Ficuzza, [37.85, 13.38], 1906, leg. O.Leonhard (1 SDEI);Messina, [38.18, 15.54], leg. Schiödte (4 ♀♀ NHMD). MOROCCO: Ain Leuh, [33.28, -5.33], 17.V.1925, leg. M. Escalera (1 J 4 ♀♀ MNCN); Ar Rif, Chefchaounen reg., env.Bad Bessen, 34.9913, -4.8555, 1124 m, 3.VI.2007, leg. P. Hlaváč (1 J cSch); Ar Rif, Chefchaounen region, rd. Bad Berret - Bad Bessen, 1392 m, 35.0169, -4.0035, 3.-6.VI.2007, leg. P. Hlaváč (1 ♀ cSch); Taza, [34.22, -4.00], leg. F. Schubert (1 ♀ NMW). PORTUGAL: Penedo da Meditação, Coimbra, [40.224371, -8.40], 24.IV.1961, leg.R.N. F. (1 J NMW). Mൺൽൾංඋൺ: Faja da Nogueira, [32.74, -16.90], laurisilva with big Ocoteas, 750 m, 20.IX.1980, leg. H. Enghoff &amp; O. Martin (2 JJ 3 ♀♀ NHMD); Paul da Serra, [32.73, -17.05], 27.XII.1987, leg.Gillerfors (1 J MZLU). SPAIN: Castanos de Calabazas, 1 km S Castanar de Ibor, Extemadura, 39.6235, -5.4128, 860 m, sifted mossy rock scree litter, chestnut and oak forest /w rock screes and small stream, 10.V.2019, leg.A. K. Hansen, J. J. Shaw, &amp; J. Kypke (NHMD); Cavarrubias [Covarrubias], Burgos,[42.05, -3.51], 10.XI.1995, leg.A.Anichtchenko(1J cSha); Lloret de Mar, [41.69, 2.83], 17.IX.-6.X.1961, leg. Palm (1 J MZLU); Sierra de Ancares,A Degrada vill., [42.81, -6.87], 1100-1250 m, 8.-9.VII.1996, leg. Wrase (2 JJ cSch); Sierra de Guadarrama, Navacerrada,Arroyo de Peca [Peña] Cabrita, [40.77, -3.99], 1800 m, 5.IX.2001, leg.A.Anichtchenko (1 J NHMD); Sierra Nevada,rio Lanjaron,Tello,[36.92, -3.47], 1500 m, 29.- 31.III.2010, leg.Shavrin &amp;Anichtchenko (1J NHMD);Toledo,El Pielago Navamorcruende, [40.15, -4.79], 4.VIII.2005 leg. Lencina &amp; Andujar (1 ZMHB). SYRIA: Krak des Chevaliers, [34.75, 36.29], 600 m, 7.V.1982 (1 J 1 ♀ MCZ); MashtalHelu [Mashta Al Hilu], E of Safita, [34.87, 36.26], 30.IV.2008, leg.Skoupý (1 J cSch); Zainie, 45 km NE of Latakia, [35.78, 36.19], 4.V.1982, leg. M. Dvorák (3♀♀ MCZ). SWEDEN: Fൺඋද: Farön, [57.94, 19.14] (1 MZLU). GඈඍඌKൺ Sൺඇൽදඇ: Sandön, [58.36, 19.24] (1 MZLU). Öඅൺඇൽ: Stenåsa, [56.51, 16.59] (1 MZLU). TURKEY: Amanos [Nur Mountains], [36.74, 36.33], 24.V.1987, leg. Jäch (1 ♀ NMW); Bolu, [40.72, 31.61], leg. F. Schubert (1 ♀ NMW); Çıkrıkçı, Turgutlu, [38.47, 27.81], 15.II.2006, leg. S.Anlaş (1 J NHMD); Çıkrıkçı, Turgutlu, [38.47, 27.81], 17.II.2006, leg.S.Anlaş (1J 3♀♀ NHMD);Danin-Danin, Hakkari Prov, [37.44,42.86], 2200 m, 1.VI.1987, leg.Schönemann &amp; Schillhammer (2 JJ 7♀♀ NMW); Izmir, Boz daglari, Boz dag köy env., [38.34, 27.75], 1500-1700 m, 30.V-3.VI.2003, leg. R. Lohaj (1 J 1 ♀ cSch); 50 km N Antalya, [37.33, 30.77], 18.IV.2015 leg. Snizek (1 ♀ NMEG); Sogukoluk, Belen, Hatay, [36.49,36.14], 24.V.1987, leg.Schönemann &amp;Schillhammer (1 ♀ NMW). UNITED KINGDOM: Sർඈඍඅൺඇൽ: Bridge of Orchy, [56.51, -4.76], 23.VII.1987, leg. Gillerfors (1 J 1 ♀ MZLU); Hawock [55.42, -2.78], VI.1964, leg. M. Paoletti (1 ♀ NMW).</p><p>Redescription. Measurements JJ (n = 6): HW = 1.78– 1.87 (1.81); HL = 1.47–1.56 (1.51); HL/HW 0.81–0.85 (0.83); PW = 2.27–2.36 (2.31); PL = 2.22–2.24 (2.23); PL/PW 0.94–0.98 (0.96); EW = 2.60–2.69 (2.63); EL = 2.49–2.60 (2.54); EL/EW 0.95–0.98 (0.96); EL/PL 1.12– 1.16 (1.14); PW/HW 1.49–1.58 (1.53); forebody length 6.20–6.33 (6.28). ♀♀ (n = 4): HW = 1.80–1.87 (1.84); HL = 1.49–1.56 (1.52); HL/HW 0.81–0.86 (0.83); PW = 2.38–2.51 (2.44); PL = 2.27–2.36 (2.31); PL/PW 0.94–0.95 (0.95); EW = 2.73–2.89 (2.82); EL = 2.58–2.69 (2.64); EL/EW 0.92–0.97 (0.94); EL/PL 1.09–1.17 (1.14); PW/ HW 1.53–1.69 (1.61); forebody length 6.42–6.56 (6.47).</p><p>Very large species; body black (Fig. 7D).</p><p>Head black, slightly transverse, with eyes large (EyL/TL = 3.64–4.78 (4.22)) following contour of head; microsculpture of transverse waves, 4 to 6 punctures between anterior frontal punctures on frons (Fig. 6D); antennae pale reddish, long, all antennomeres clearly elongate; palpi pale reddish.</p><p>Thorax: pronotum black, almost quadrate, as wide as long, clearly wider than head, microsculpture of transverse waves, three punctures in dorsal row and three in sublateral row with the most posterior one reaching the level of third puncture of dorsal row; scutellum punctured and setiferous, elytra black, densely punctured and pubescent, slightly wider than long, slightly longer than pronotum; legs dark brown with inner face of tibia darkened and tarsi paler.</p><p>Abdomen black, tergites uniformly punctured, slightly iridescent.</p><p>Male. Aedeagus (Fig. 11B): Paramere slender, lanceolate, reaching apex of median lobe, sensory peg setae forming two clear rows; median lobe with protrusion at apices and two small but distinct teeth near lower level of parameres peg setae; internal sac with two small distal medial sclerites.</p><p>Differential diagnosis. Quedius levicollis is easily distinguished from other members of the fuliginosus -group by the punctured and setiferous scutellum and three punctures in the sublateral row of pronotum with the posteriormost one reaching the level of the third (basalmost) puncture of the dorsal row. It can be additionally distinguished from Q. fuliginosus by the pale antennae, and from Q. curtipennis by the slender paramere and median lobe with apical protrusion. Quedius levicollis is mainly found in drier habitats and rarely co-occurs with any of the other species in the species group.</p><p>Synonymic notes. The name Quedius tristis (Gravenhorst, 1802) has long been applied to this species. GඋൺඏൾඇHඈඋඌඍ (1802) did not describe Staphylinus tristis as a new species but attributed it to Fabricius. The Fabrician species is now in Ocypus (Pseudocypus) . Nൾඐඍඈඇ (1995) made an appeal to the International Commission of Zoological Nomenclature requesting that the species takes the name Quedius levicollis, as this was the next available name associated with this species. As a result, Quedius levicollis (Brullé, 1832) was accepted on the Official List of Specific Names in Zoology (ICZN 1996: 215).</p><p>BൾඋඇHൺඎൾඋ &amp; SർHඎൻൾඋඍ (1916) placed Staphylinus nigritarsis Ménétriés, 1832 in synonymy to Philonthus splendens . Gඎඌൺඋඈඏ (1993) revised the type series of S. nigritarsis and designated a lectotype. He found S. nigritarsis to be conspecific with Q. levicollis (Brullé, 1832) based on both external characters and the shape of the aedeagus. We confirm Gusarov’s identification through examination of photos of the type from ZIN provided by M. Salnitska (see supplementary Fig. S1).</p><p>We were not able to examine the types of Philonthus frontalis Nordmann, 1837 and Quedius balthasari Mařan, 1935, both currently accepted synonyms of Q. levicollis . Based on the original descriptions, both synonymies appear to be correct. Philonthus frontalis was described from Italy and Greece and fits the description of Q. levicollis, with interorbital punctures and punctate scutellum. Quedius balthasari Mařan, 1935, described from central Crete as a species different from Q. levicollis in sparser punctation and reddish margins of elytra is likely a color aberration because Q. levicollis is also known from Crete.</p><p>Bionomics. Quedius levicollis is a clearly thermophilous eurytopic species, which prefers sandy soils and is found in both open habitats and dry forests. It occurs in various ground debris and under rocks, e.g., it has been found in forest litter at different elevations (200–1900 m) in Cyprus (AඌඌංඇG &amp; Wඎඇൽൾඋඅൾ 2001), under rocks in Corse (MൺඋർHൺඅ &amp; CH ൺඋൽඈඇඇൾඍ 2000), and from sifted wet leaf-litter under oleander trees, at 500–1100 m in Israel (Sආൾඍൺඇൺ 1978b). We have encountered this species under rocks in open oak and chestnut forest in Spain (Fig. 2B), and between the roots of Erica in a heathland in Denmark. It has been recorded from caves in Sardinia (Bඈඋൽඈඇං 1982), although this most likely represents an accidental occurrence.</p><p>WൾඅඌH (1993) found this species to contain six ovaries with a fasciculata type of structure.</p><p>Distribution. Quedius levicollis is widely distributed across the Western Palearctic from the Atlantic coast, including British Islands through the Mediterranean, including North Africa, in the west to the Russian Black Sea coast and the Middle East (Fig. 19). In the Middle East the species extends into the Levant region, but not into the Nile delta. The south-easternmost record of Q. levicollis is from South Iran. The species is found on all major Mediterranean islands, including Cyprus, Crete, Corse and Sardinia, as well on the island of Madeira in the Atlantic Ocean. It is most common in the southern part of its distribution range gradually becoming rarer towards the north-east where it occurs at xerothermic sites reaching its northernmost limit on the Swedish island of Gotland in the Baltic Sea.</p></div>	https://treatment.plazi.org/id/038987A0FFE14B07B3AFA19817ADFF6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFE44B07B139A11816E5FECA.text	038987A0FFE44B07B139A11816E5FECA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius molochinus (Gravenhorst 1806)	<div><p>Quedius molochinus -group</p><p>Diagnosis. Species of medium size. Coloration variable from completely dark to dark with brown, reddish, or yellowish pale elytra and appendages. Eyes smaller, temples larger, with temporal puncture separated from posterior margin of eye by a distance equal to the diameter of the puncture. Frons without interocular punctures (cf. Fig. 6F). Scutellum punctate and pubescent. Pronotum without puncture(s) in the posterior part of sublateral row. Palearctic; Q. molochinus introduced to North America.</p><p>Comments. The Quedius molochinus -group is composed of four species ( Q. molochinus, Q. meridiocarpathicus, Q. vicinus, and Q. balticus) that usually have red elytra, although some variants have them darkened. All of these are distributed within the Palearctic Region – although Q. molochinus has been introduced to the Nearctic (Sආൾඍൺඇൺ 1981). Among others of the subgenus, these species are characterized by usually having reddish-brown to orange elytra, frons without interocular punctures, scutellum punctate and pubescent, and sublateral rows on pronotum without punctures on the posterior half (Fig. 8). The species are externally very similar and have been confused for a long time, rendering their real distributions unknown. They have also been confused with the red elytral species of the Q. pallipes -group, from which they are distinguished by sublateral rows on pronotum without punctures in posterior half. Thorough investigation of representative material here demonstrates that Q. molochinus, Q. meridiocarpathicus and Q. vicinus are almost entirely allopatric (Fig. 20). Quedius molochinus is a more northern species, distributed in Europe from the UK and European Russia to the Alps and the Czech Republic, in Asia it is widespread from the Urals to Transbaikalia. In southern Europe it is replaced by Q. meridiocarpathicus, which is found in the Italian and Balkan peninsulas and continuing eastwards through Hungary to the Black Sea coast of Ukraine and Russia. Quedius vicinus is confined to Eastern Transcaucasia along the Caspian Sea, southeastern Anatolia, and the northern Middle East across to central Asia. The only overlap is in Anatolia where both Q. meridiocarpathicus and Q. vicinus can co-occur. In this area the species can be most easily distinguished by differences in male genitalia (Fig. 13). Another minor difference includes the antennal coloration, where the middle antennomeres (from fourth to sixth) are slightly lighter in Q. vicinus compared to Q. meridiocarpathicus (Fig. 8). Quedius meridiocarpathicus in turn has slightly lighter middle antennomeres as compared to Q. molochinus (Fig. 8). All three species are found in similar habitats, mainly in various types of moist litter or under rocks either near creeks and rivers or in forests. Quedius balticus is found across a large part of the Palearctic and thus has an overlapping distribution with the other three species in parts of their ranges (Fig. 19). Unlike the three former species, Q. balticus is almost exclusively found in bogs and on floodplains, to which it is highly adapted.Also, it is externally easy to recognize based on the darkened basal three antennomeres, which are pale in the three other species (Fig. 8). If there is any doubt in external characters, these four species in the Q. molochinus -group can most easily be identified through differences in male genitalia (see descriptions).</p></div>	https://treatment.plazi.org/id/038987A0FFE44B07B139A11816E5FECA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFE44B39B3DEA17816FDFA0A.text	038987A0FFE44B39B3DEA17816FDFA0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius balticus Korge 1960	<div><p>Quedius balticus Korge, 1960</p><p>(Figs 1, 4, 8E, 13E, 19)</p><p>Quedius balticus Korge, 1960a: 52 [Type locality: Südspitze der Insel Hiddensee bei Rügen]</p><p>References. Lൺඌඍ (1952): 148 (characters; cited as Q. molochinus, but misidentified according to KඈඋGൾ 1960a); (1963): 43 (characters); KඈඋGൾ (1960a): 67 (characters and notes); (1963): 87 (biology); Cඈ-ංൿൿൺංඍ (1961): 50 (characters); (1978): 192 (characters, distribution and biology); Sආൾඍൺඇൺ (1962): 135 (characters); Hൺඇඌൾඇ (1962): 319 (characters); Pൺඅආ (1963a): 160, (1963b): 117 (characters); LඈHඌൾ (1964): 212 (characters); Hඈඋංඈඇ (1965): 277 (distribution); Sඓඎඃൾർĸං (1966): 36 (distribution); Pඈඉൾ (1977): 31 (distribution); TඬඍH (1984): 119 (characters); LඎർHඍ (1987): 109 (distribution); Gඎඌൺඋඈඏ (1989): 12 (distribution); Jൺඇගĸ (1992): 91 (distribution); Sආൾඍൺඇൺ (1993): 50 (distribution); Hඒආൺඇ (1994): 194 (distribution); HඈൽGൾ &amp; Jඈඇൾඌ (1995): 41 (characters); Mඈඋൺඏൾർ &amp; Vඈඇංඹĸൺ (2000): 39 (distribution and biology); Jൾඅටඇൾĸ (2001): 200 (distribution); KඈආඈඌංŇඌĸං (2001): 19 (distribution); Fඳඅදඉ (2005): 144 (distribution); UHඅංG et al. (2006): 41 (distribution); NൺඌH (2009): 23 (distribution and biology); Cඎඉඉൾඇ et al. (2011): 12 (distribution); Mඈඇඌൾඏංඹංඎඌ (2013): 27 (distribution); Mൺඃඓඅൺඇ &amp; Lංඍൺඏඌĸප (2017): 19 (distribution and biology); Hඈൿൿආൺඇඇ et al. (2018): 199 (biology).</p><p>Material examined. AUSTRIA: Purbach, Burgenland, [47.90, 16.70], 6.VIII.1985, leg. V. Kovits (1 J NMW). BULGARIA: Sozopol, Black Sea coast, [42.42, 27.68], 16.VI.1972, leg. M. Uhlig (1 J ZMHB). CZECH REPUBLIC: Poděbrady, [50.13, 15.12], 3.IX.1936 (1 J NHMD). DENMARK: Bognaes Skov på Tuse Naes, 55.7511, 11.7695, sifted from sea debris on saline marsh, 30.XII.2013, leg. K.B. Nielsen (1 J NHMD); Engestofte, [54.76, 11.56], 20.VIII.1976 (2 JJ 1♀ NHMD); Sundby Storeskov, [54.78, 11.81], 29.VII.1962, leg. V. Hansen (9 JJ 2 ♀♀ NHMD); Vålse Vesterskov, 54.9592, 11.7728, sifted from fresh sea debris on saline marsh, 7.XII.2013 leg. K.B. Nielsen (1 ♀ NHMD). FIN-LAND: Kiiminki, [65.13, 25.78], 2011, leg. M. Mutanen (1 ♀ ZMUO). GERMANY: Degensmoor,Wesenberg, [53.27, 13.03], 31.VII.1978, leg. Stöcke (1 ♀ ZMHB); Döbitzer Heide, Ferbitzer Bruch, [52.50, 13.01] (1 J 1♀ ZMHB); Federsee,Württenberg, [48.08, 9.62], 28.VII.1977 (2♀♀ ZMHB); Korswandt n. Wolgast, [53.92, 14.16], 25.VIII.1989, leg. Kleeberg (1J 1♀ NMW); Lübars, Berlin, [52.62, 13.36], 14.VII.1969, leg. H. Korge (3JJ 4♀♀ ZMHB); Müritz,Waren, [53.44, 12.74], sifting, spaghnum with grass and leaf litter, 17.VI.1976, leg. M. Uhlig (2 JJ ZMHB); Teufelbruch n. Spandau, [52.57, 13.21], 22.VIII.1964 (1 J ZMHB). HUNGARY: Pest nr. Mariabesnyö, [47.61, 19.41], 23.VIII.1931, leg. J. Fodor (1 ♀ NHMD); Pest nr. Pécel, [47.49, 19.34], IX. -XI.1929 leg. J. Fodor (1J NHMD). ITALY: Zeccone n. Pavia, [45.26, 9.19], 17.X.1977, leg. P. Kanaar (1 J ZMHB). KAZAKHSTAN: Karatal river, [45.41, 77.81], 1.VII.2004, leg. V.A. Kastcheev (2 JJ ZIN); Kyrgyz Alatau, Koshirme konezanod, [43.04, 76.84], 9.VII.2010, leg. V.A. Kastcheev (2 JJ ZIN). NORTH MACEDONIA: Ohrid Lake, n. Struga, [41.17, 20.66], 7.VI.1980, leg. F. Hieke (1 J 2 ♀♀ ZMHB). SWEDEN: Arlöv, [55.64, 13.05], grass roots on mossy soil near sea, 3.X.1962, leg. Palm (1 J MZLU); Fotevik, [55.45, 12.95] (1 MZLU). UKRAINE: Peter &amp; Paul, VII.-VIII., leg. A. Spaney (1 J ZMHB).</p><p>Redescription. Measurements JJ (n = 5): HW = 1.64– 1.78 (1.68); HL = 1.38–1.47 (1.41); HL/HW 0.81–0.88 (0.84); PW = 2.13–2.38 (2.22); PL = 1.98–2.00 (1.98); PL/PW 0.84–0.93 (0.89); EW = 2.11–2.40 (2.21); EL = 1.91–2.27 (2.07); EL/EW 0.88–0.96 (0.93); EL/PL 0.97– 1.13 (1.04); PW/HW 1.48–1.65 (1.57); forebody length 5.36–5.71 (5.46). ♀♀ (n = 5): HW = 1.62–1.71 (1.66); HL = 1.33–1.44 (1.40); HL/HW 0.82–0.87 (0.84); PW = 2.00–2.22 (2.13); PL = 1.78–2.00 (1.88); PL/PW 0.86–0.90 (0.89); EW = 2.13–2.24 (2.20); EL = 2.02–2.18 (2.08); EL/EW 0.90–0.98 (0.94); EL/PL 1.01–1.15 (1.10); PW/ HW 1.29–1.56 (1.47); forebody length 5.18–5.51 (5.36).</p><p>Medium sized, robust species; body dark brown to black (Fig. 8E).</p><p>Head black, distinctly transverse, with eyes medium sized (EyL/TL = 2.00–2.27 (2.14)); microsculpture of transverse waves; no interocular punctures between anterior frontal punctures (cf. Fig. 6F); antennae pale reddish with base of antennomeres 1–3 clearly darkened, all antennomeres elongate; palpi pale reddish.</p><p>Thorax: pronotum dark brown to black, slightly wider than long, wider than head, with microsculpture of transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond the first puncture of dorsal row; scutellum punctured and pubescent; elytra most often reddish brown rarely fully darkened, uniformly pubescent, with punctures clearly separated, slightly wider than long, roughly as long as pronotum; legs reddish brown with inner face of tibia and femur darkened, tarsi paler.</p><p>Abdomen dark brown to black, tergites uniformly punctured, without clear iridescence.</p><p>Male. Aedeagus (Fig. 13E): paramere rather broad, with clear medial expansion and extending into a point at apex, reaching just beyond apex of median lobe, with sensory peg setae forming two short broad rows fusing together towards apex, in lateral view peg setae are easily seen on protruding medial part of parameral apex; median lobe broad with sudden constriction to a point at apex, on parameral side with two well-defined teeth directed almost medially, positioned at level near end of peg setae rows of paramere; internal sac without extension of the C-sclerite.</p><p>Differential diagnosis. Quedius balticus is very similar to Q. molochinus, Q. meridiocarpathicus and Q. vicinus, but can be distinguished by the antennomeres 1–3 being at least partially darkened. For confident identification genitalia should be examined, especially when specimens are teneral or faded (from old collections) and thus with unnatural coloration. The aedeagus of Q. balticus differs from Q. molochinus in the absence of an extension of the C-sclerite in the internal sac, in the broad median lobe suddenly constricted at tip and in the medial broadening of paramere.It can be distinguished most easily from Q. meridiocarpathicus by the shape of the paramere, which is medially broadened in Q. balticus and broadened towards apex in Q. meridiocarpathicus . Quedius balticus can be confused with Q. picipes from the subgenus Raphirus, which may occur in similar environments. It can be readily distinguished from the latter by the labrum being entire, the darkened base of antennomeres 1–3 and the abdomen being more parallel-sided.</p><p>Color variation. Quedius balticus var. funebris Coiffait, 1971 was described from a female specimen from ‘Sarthe, La Ferté-Bernard’ as a variant of Q. balticus with black elytra (Cඈංൿൿൺංඍ 1971). Based on the images of that specimen kindly made on our request by Vinicius Ferreira at the MNHP, it is Q. molochinus with dark elytra. According to the ICZN Article 45.6 Quedius funebris is unavailable name.</p><p>Quedius picipennis ab. nigribasis Korge, 1960, another unavailable name, was described from a female from Germany (“Sweertia Weise Bin Lübais”) and characterized by darkened antennomeres. So far, the form has been considered a color form of Q. molochinus (e.g., Hൾඋආൺඇ 2001), but we are inclined to believe that this is indeed Q. balticus . At the time of description H. Korge considered Q. balticus restricted to an area near the Baltic Sea, but it is now known that the species is much more widespread.</p><p>Bionomics. Quedius balticus was originally considered a littoral species found only in sea debris (KඈඋGൾ 1960a, Hൺඇඌൾඇ 1962) and litter washed up on beaches (Pൺඅආ 1963). Later it was discovered in reed litter in swamps, moors, wet meadows, as well as riverine areas subject to flooding (KඈඋGൾ 1963, Mඈඋൺඏൾർ &amp; Vඈඇංඹĸൺ 2000, NൺඌH 2009, Mൺඃඓඅൺඇ &amp; Lංඍൺඏඌĸප 2017). Based on the earlier published data and our observations, Q. balticus is not common.</p><p>KඈඋGൾ (1963) noted that specimens, which he kept in a terrarium were hiding in wet meadow litter and could not even be expelled from it by flooding. While other beetles, including closely related representatives of the same genus, quickly moved out when debris were flooded, Q. balticus stayed behind. In a study conducted in a valley of the river Peene in NE Germany, Hඈൿൿආൺඇඇ et. al (2018) found Q. balticus to live only near natural fens and be the strongest indicator of these natural habitats. Given these observations the species seems to be highly adapted to environments subject to flooding and it can be a good indicator of a natural hydrology of an area.</p><p>Distribution. Quedius balticus is currently known to occur across large parts of Europe except the Iberian and the Italian peninsulas (Fig. 19). The species is currently known from Southern Finland and Sweden in the north to Italy, Bulgaria and Macedonia in the south, and from the British Isles and Northern France in the west to Kazakhstan in the east. Newly studied material herein extends the range north- and southwards, from Finland to Italy and Macedonia.</p></div>	https://treatment.plazi.org/id/038987A0FFE44B39B3DEA17816FDFA0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFDA4B3BB36CA5B114E4F78A.text	038987A0FFDA4B3BB36CA5B114E4F78A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius meridiocarpathicus Smetana A. Solodovnikov 1958	<div><p>Quedius meridiocarpathicus Smetana, 1958</p><p>(Figs 1, 2C, 4, 8B, 13F, 20)</p><p>Quedius meridiocarpathicus Smetana, 1958: 365 [Type locality: Slovensko, Kamenný Most, Slanec]</p><p>Quedius dalmatinus Korge, 1960b: 68 [Type locality: Dalmatia: Castelnuovo]</p><p>References. Cඈංൿൿൺංඍ (1961): 53 (characters); KඈඋGൾ (1962): 153 (synonymy of Q. dalmatinus); Sආൾඍൺඇൺ (1962): 138 (characters); LඈHඌൾ (1964): 212 (characters); Hඈඋංඈඇ (1965): 277 (distribution); Sආൾඍൺඇൺ (1967): 558 (distribution); Bඈඋൽඈඇං (1976): 99 (characters); Cඈංൿൿൺංඍ (1978): 195 (characters); TඬඍH (1984): 129 (characters); Sආൾඍൺඇൺ (1993): 50 (distribution); Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං (1995): 32 (distribution); Lගඌඓඅඬ (1998): 137 (distribution); Aඇඅൺ෡ (2009): 230 (distribution); Sඈඅඈൽඈඏඇංĸඈඏ (2012): 33 (distribution); Pൾඍඋൾඇĸඈ (2015): 80 (distribution); Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2019): 49 (characters and distribution).</p><p>Type material examined. Quedius meridiocarpathicus: Lൾർඍඈඍඒඉൾ J (cSme), designated by Sඈඅඈൽඈඏඇංĸඈඏ (2012), labelled: “ Slovakia mer. Kamen. Most 5.5.1955 Smetana 1955 / Quedius meridiocarpathicus s. Smetana det. 1957 / Lectotype Quedius meridiocarpathicus Smetana A. Solodovnikov des. 2009 / Quedius meridiocarpathicus Smetana A. Solodovnikov det. 2009”.</p><p>Pൺඋൺඅൾർඍඈඍඒඉൾඌ: 1J, labelled:“ Slovakia mer. Kamen.most 3.5.1955 Smetana / Quedius meridiocarpathicus Smetana det. 1957 / Quedius meridiocarpathicus A. Solodovnikov det. 2009 / PARALECTOTYPE Quedius meridiocarpathicus SmetanaA.Solodovnikov des.2009 / NHMD 00523118” (NHMD); 1♀, labelled: “ Slovakia mer.Kamen.most 3.5.1955 Smetana / Quedius meridiocarpathicus Smetana det. 1957 / Quedius meridiocarpathicus A. Solodovnikov det. 2009 / PARALECTOTYPE Quedius meridiocarpathicus Smetana A.Solodovnikov des.2009 / NHMD 00523117”.</p><p>Additional material examined. ALBANIA: Kolonjë, 3 km ENE Leskovik, 40.1586, 20.6216, flooded pasture, 990 m, 28.V.2010, M. Schülke (1 J cSch). BOSNIA AND HERZEGOVINA: Jajce, [44.34, 17.27], 12.-23.VI.1967, leg.Baderle (1 J 3♀♀ NMW); Trebinje, [42.71, 18.35], 1903 O. Leonhard (1 ♀ SDEI). BULGARIA: Burgas, Black Sea coast, [42.50, 27.48], 8.-19.V.1985, leg. D.W. Wrase (1 J 5 ♀♀ ZMHB); Kiten, [42.24, 27.77], 29.V.1982, leg. H. Wendt (1 ♀ ZMHB); Melnik, [41.52, 23.38], 12.-20.V.1981 (2 ♀♀ ZMHB); Nessebar, [42.66, 27.71], 6.-7.VI.1982, leg. H.Wendt (1 J 2♀♀ ZMHB); Pomorie, Black Sea coast, [42.58, 27.62], 9.-18.V.1985, leg. D.W. Wrase (1 J cSch); Primorsko, [42.26, 27.75], VII.1980, leg. I. Pokorný (1 ♀ cSch); Rupite, General Todorov, [41.43, 23.24], 5.V.1984, leg. Hieke (1 ♀ ZMHB); nr. Sandanski, [41.55, 23.23], 28.IV.1985, leg. B. Jaeger (1 J 1 ♀ ZMHB); Slivno, [42.67, 26.32] (1 J NHMD); Veliko Tarnovo, [43.08, 25.63], 25.VII.1984, leg. U. Opitz (2 ♀♀ ZMHB). CROATIA: Castelnuovo [Kaštel Novi], [43.55, 16.31], 1895, leg. F. Müller (1 ♀ SDEI); nr. Plitvice, [44.86, 15.58], 19.VII.1910, leg. Hochetlinger (1 ♀ NHMD); nr. Zagreb, [45.78, 15.98] (1 J NMW). GREECE: Alexandroupolis, [40.85, 25.87], 20.-30.IV.1967, leg. Krätschmer; 4 km NO of Anthili, Thessalia, Lamia, 38.8655, 22.5224, 6.II.2015, leg.T.Wolsch (1J cSch); Essymi [Aesymi], [41.01, 25.95], 700 m, 25.IV.1978, leg. Heinz (1 J 1 ♀ ZMHB); Ioannina, Metsovo, Lago O Pigon, [39.81, 21.11], 18.V.2005, 1500m, leg.Angelini (2 ♀♀ NHMD); Janina [Ioannina], [39.66, 20.85], IV.1927, leg.C. Purkyne (2 JJ NHMD); Keramidi, Thessalia, Palamas, 39.5627, 22.0801, 5.II.2015, leg. T. Wolsch (1 ♀ cSch); Lavrio, 60 km S Athina, [37.71, 24.06], 10.IV.1993, leg. P. Jørum (1 ♀ NHMD); Parnass. [Parnassos], [38.53, 22.62] (3 JJ NHMD); Skarfia, 20 km WSW Lamia, 38.8405, 22.6988, salt meadows, 6.II.2015, leg. T. Mainda (1 J cSch). Cඈඋൿඎ: Val di Ropa, [38.24, 21.73], 1905, leg. O. Leonhard (1 ♀ SDEI). Cඋൾඍൾ: Crete [35.41, 24.75], leg. Paganetti (1 J NMW) [potentially mislabelled or introduced specimen clearly belonging to this species]. Kൾൿൺඅඈඇංൺ: Paranetti, [38.21, 20.58], leg. F. Müller (1 ♀ SDEI). Pൾඅඈඉඈඇඇൾඌ: 3 km E Kalogria, 20 km NW of Varda, [38.19, 21.39], 15.IV.1993, leg. P. Jørum (1 ♀ NHMD); Nomos Lakonias, Near Fokiano, 37.0741, 22.9676, 0 m, sifting, waters edge, stagnant water, salt marsh, 02.IV.2019, leg. M. J. Justesen &amp; A.K.Hansen (1 J NHMD). ZൺKඒඇඍHඈඌ: Kalamaki, [37.74, 20.89], 1909, leg. M. Hilf (1♀ ZMHB). HUNGARY: Hódmezővásárhely, [46.41, 20.34], 16.-22.VI.1995, leg Probst (2 ♀♀ NMW); Rapnitz [Rábca] river, Neusiedler See, [47.67, 16.81], V.1994 (1J ZMHB); Velencei, Dinnyes, [47.17, 18.54], salt lake with gravel banks, 15.VI.1979, leg. M. Uhlig (1 J 1♀ ZMHB). ITALY: Albareto, Emilia, [44.53, 10.40], 8.II.1994, leg.A. Fiori (2♀♀ ZMHB); NE of Bibione, left bank of Canale di Lugugnana mouth at Bibione, Veneto, 45.6259, 12.9791, sifting flood debris and litter, 29.V.2016, leg. A. Solodovnikov &amp; M. Salnitska (2 JJ 1 ♀ NHMD); Casinalbo, Emilia, [44.53, 10.40] 25.VII.1996, leg.A. Fiori (2 JJ ZMHB); Cascine Del Riccio, Riva Ema [43.73, 11.25], 14.I.1970, leg. Mascagnia (1 J cBor); Comacchio, [44.69, 12.17], 28.III.1974, leg. P. Garaghahi (2 ♀♀ cBor); Firenzuola, [44.12, 11.37], VII.1970, leg. Mascagnia (1 J cBor); Lido di Jesolo, [45.51, 12.62], 10.VIII.1962, leg. Salvato (1 ♀ cBor); Noghera Istria, [45.58, 13.81] (1 J NHMD); Padova, [45.39, 11.90], IX.1969 (1 J ZMHB); Roma, Portonaccio, [41.90, 12.54], IX.1937, leg. G. Sacca (1 J 4 ♀♀ cBor); Tencarola, [45.39, 11.80], 10.XII.1961, leg. Salvato (1 ♀ cBor). NORTH MACEDONIA: Bitola, Jugoslavia, [41.02, 21.32], 1962, leg. J.T. Skovgaard (1 ♀ NHMD); Katlanovo nr. Skopje, [41.89, 21.68], 17.IX.-12.X.1966, leg. Gruber (1J 1♀ ZMHB); Treska River, Sara, [41.97 21.30], 3.VI.1980, leg.F. Hieke (1 J ZMHB); Vardar River, n. Skopje, [41.98, 21.49], 26.V.1980, leg. F. Hieke (1 J 3 ♀♀ ZMHB). ROMANIA: Predeal, [45.49, 25.57], 20.VI.1977 (1 ♀ MZLU). RUSSIA: Kඋൺඌඇඈൽൺඋ Kඋൺං: 15 km S of vill. Taman, [45.21, 36.70], 15.V.1995, sandy sea shore, under logs (2 JJ NHMD); Temruk, mud volcano Gefest, 45.2566, 37.4326, leaf litter in meadows, 12.VI.2017, leg. M. Salnitska (1 NHMD) Karabetova Gryada 5 km SE of vill. Taman, [45.12, 36.76], in litter at the bank of the permanent pond (1 ♀ NHMD); Mt.Tkhab, valley of river Zhene, [44.58, 38.35], in forest litter, 21.VI.1992, leg. M. Savitsky (1 J NHMD). SLOVENIA: 9 km SE Piran, [45.48, 13.59], 26.VI.1981, leg. F. Hieke (1 ♀ ZMHB). TURKEY: Adapazarı, [40.79, 30.40], flooding, 15.VII.1965, leg. Korge &amp; Heinz (1 J ZMHB); SW Arnavutköy and Yassıören, [41.17, 28.65], 60 m, 19.IV.1990, leg. Heinz (1 J ZMHB); Caykasi River, Erfelek, Sinop, [41.89, 34.92], 26.V.1989, leg. Schönmann &amp; Schillhammer (1 J 3 ♀♀ NMW); 6 km E Gebze, Izmit, [40.78, 29.49], IV.1983, leg. Heinz (1 J 2 ♀♀ ZMHB); 20 km W Gebze, [40.87, 29.22], IV.1989, leg. Heinz (2 JJ 1 ♀ ZMHB); Gelibolu, [40.41, 26.67], 10 m, 3.V.1982, leg. Boffa (2 ♀♀ cBor) İnönü, Eskişehir, [39.82, 30.14], 850 m, 3.IV.1986, leg. Heinz (1 ♀ ZMHB); Keşan, [40.85, 26.62], 22.IV.1978, leg. Heinz (2 JJ ZMHB) E Pamucak, Selcuk, [37.96, 27.27], swamp, 28.III.1986, leg. Heinz (1 J ZMHB); 2 km E Sarısaltık, 39.1241, 39.2475, 1594 m, 19.V.2011, leg. Anlaş (1 ♀ NHMD); Saray, 30 km W Ankara, [40.07, 32.61], 23.II.1973, leg. Exp. Nat. Mus. Praha (1 J NHMD); Söğüt, SE of Bilecik, [40.02, 30.18], 1200 m, 4.IV.1992, leg. Heinz (1 J 2 ♀♀ ZMHB); Susuz-Dag, Dokuz-Göl, Elmalı, [36.50, 29.76], 1700-2000 m, 20.VII.1971, leg. Heinz (1 J ZMHB). UKRAINE: Environs Odessa, right bank of Kujalnitskij estuary, [46.57, 30.73], 10.VI.2005, under stones, leg.A. Gontarenko (3 JJ 1 ♀ NHMD); Nikitsky botanical garden, [44.50, 34.23], by the sea, stones, 16.IX.1966 (1 J cRyv).</p><p>Redescription. Measurements JJ (n = 7): HW = 1.73– 1.82 (1.78); HL = 1.47–1.56 (1.51); HL/HW 0.82–0.88 (0.85); PW = 2.27–2.47 (2.36); PL = 2.09–2.31 (2.19); PL/PW 0.89–0.95 (0.93); EW = 2.27–2.47 (2.36); EL = 2.09–2.24 (2.17); EL/EW 0.87–0.99 (0.92); EL/PL 0.92–1.05 (0.99); PW/HW 1.50–1.66 (1.56); forebody length 5.71–6.02 (5.86). ♀♀ (n = 3): HW = 1.64–1.76 (1.71); HL = 1.33–1.51 (1.43); HL/HW 0.81–0.86 (0.83); PW = 2.07–2.36 (2.21); PL = 1.96–2.13 (2.05); PL/PW 0.91–0.95 (0.93); EW = 2.16–2.38 (2.28); EL = 1.98–2.24 (2.12); EL/EW 0.90–0.97 (0.93); EL/PL 1.00–1.09 (1.03); PW/HW 1.52–1.56 (1.54); forebody length 5.27–5.78 (5.60).</p><p>Medium sized, robust species; body light to dark brown (Fig. 8B).</p><p>Head black, distinctly transverse, with eyes medium sized (EyL/TL = 1.50–1.67 (1.56)); microsculpture of transverse waves; no interocular punctures between anterior frontal punctures (cf. Fig. 6F); antennae pale reddish, antennomeres elongate; palpi pale reddish.</p><p>Thorax: pronotum light to dark brown, slightly wider than long, wider than head, with microsculpture of transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond the level of first puncture of dorsal row; scutellum punctured and pubescent; elytra most often reddish brown rarely fully darkened, uniformly pubescent, slightly wider than long, roughly same length as pronotum; legs reddish brown with inner face of tibia darker.</p><p>Abdomen light to dark brown, tergites uniformly punctured, without clear iridescence.</p><p>Male. Aedeagus (Fig. 13F): paramere spatulate with clear proximal to medial attenuation and extending into paddle-like structure near apex, reaching just beyond apex of median lobe, with sensory peg setae forming a broad horseshoe shaped band of relatively large peg setae; median lobe broad with gentle constriction to a point at apex, on parameral side with two small teeth pointing slightly basad, positioned at level near basal level of peg setae band of paramere; internal sac without a continuation of C-sclerite.</p><p>Differential diagnosis. Quedius meridiocarpathicus is very similar to other species in the molochinus -group with red elytra, viz Q. molochinus, Q. balticus, and Q. vicinus . It can be distinguished from Q. balticus by the pale base of the basal three antennomeres. Generally, it has paler middle antennomeres compared to Q. molochinus . Compared to Q. vicinus it is generally slightly darker, elytra brownish red (pale red to yellowish in Q. vicinus), abdomen dark brown (light brown in Q. vicinus). For confident discrimination of these species, their genitalia should be checked: the aedeagus of Q. meridiocarpathicus differs from Q. molochinus in the absence of an extension of the C-sclerite in the internal sac, in the spatula-shaped paramere with clear proximal to medial attenuation, and larger peg setae. It can be distinguished most easily from Q. balticus by the shape of the paramere, which is medially broadened in Q. balticus and broadened towards apex in Q. meridiocarpathicus . It differs from Q. vicinus in the absence of a blunt apical hook on the paramere (parameral side). Quedius meridiocarpathicus can be confused with Q. picipes of the subgenus Raphirus, which may occur in similar environments. It can be readily distinguished from the latter by more parallel-sided abdomen and entire labrum.</p><p>Synonymic notes. Independent of each other, Sආൾඍൺඇൺ (1958) and KඈඋGൾ (1960b) recognized that a species very similar to Q. molochinus was present in Southeastern Europe. KඈඋGൾ (1962) himself synonymized Q. dalmatinus Korge, 1960 with Q. meridiocarpathicus Smetana, 1958 . Based on the description and illustration of Q. dalmatinus, it clearly agrees with the concept of Q. meridiocarpathicus herein.</p><p>Bionomics. Quedius meridiocarpathicus appears to be found most frequently in wet debris and under rocks near water bodies, both fresh and saline. We (AKH and AS) have collected this species in the flood debris of a salt lagune near Bibione, Italy, in debris around the stagnant salt pond near the beach of Fokiano, Greece (Fig. 2C) and on the marsh debris near the Gefest Mud Volcano in the Taman Peninsula in southern Russia. Like many other Quedius s. str., the species is found at lower elevations in the northern part of its range and at much higher elevations (up to 2000 m) towards the southern part of its range.</p><p>Distribution. Quedius meridiocarpathicus is distributed from Italy through Southeastern Europe to the Crimea, Western Caucasus and Anatolia (Fig. 20). There seem to be rather well defined distributional boundaries between this species, and Q. molochinus and Q. vicinus . The northern range of Q. meridiocarpathicus is roughly delimited by the Alps and Carpathians, where it is replaced by Q. molochinus to the North of these mountain chains. The species continues north around the Black Sea to the western foothills of the Caucasus. To the South it extends to northern and western Anatolia, mainly along the coast, but it is also found more inland in some sites. In southern and eastern Anatolia Q. meridiocarpathicus is gradually replaced by Q. vicinus, with which it seems to co-occur to some extent in southern Anatolia.</p></div>	https://treatment.plazi.org/id/038987A0FFDA4B3BB36CA5B114E4F78A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFD84B3DB392A0B917E2F8CA.text	038987A0FFD84B3DB392A0B917E2F8CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius molochinus (Gravenhorst 1806)	<div><p>Quedius molochinus (Gravenhorst, 1806)</p><p>(Figs 1, 4, 9, 8C,D, 12, 13D, 20)</p><p>Staphylinus molochinus Gravenhorst, 1806: 56 [Type locality:Brunsviga]</p><p>Quedius denudatus Stephens, 1832:216 [Type locality: London; Suffolk]</p><p>Quedius lathburii Stephens, 1832: 218 [Type locality: London; Suffolk, Orwell]</p><p>Quedius niger Sahlberg, 1876: 229 [Type locality: Muonioniska]</p><p>Quedius maculicornis Mulsant &amp; Rey, 1876: 694 [Type locality: les montagnes du Lyonnais, à la Grande Chartreuse, dans les Pyrénées]</p><p>Raphirus picipennis Stephens, 1833: 243 [Type locality: London]</p><p>Staphylinus laevicollis Runde, 1835: 6 [Type locality: Halae]</p><p>Staphylinus lapponicus Zetterstedt, 1838: 61 [Type locality: Lapponia Umensis, Lycksele; Wilhelmina; Gaskelougt]</p><p>References. GඒඅඅൾඇHൺඅ (1810):302, (1827):477 (characters);SൺHඅൻൾඋG (1830): 314 (characters); (1876): 24 (distribution); MൺඇඇൾඋHൾංආ (1830): 25, (1831): 439 (distribution); (1843): 232 (characters); Rඎඇൽൾ (1835): 5 (characters); Lൺർඈඋൽൺංඋൾ (1835): 377 (characters); Cඎඋඍංඌ (1837): pl. 638 (characters); Nඈඋൽආൺඇඇ (1837): 76 (distribution); EඋංർHඌඈඇ (1839): 489, (1840): 535 (characters); Hൾൾඋ (1839): 276 (characters); SඍൾඉHൾඇඌ (1839):390 (characters);Lൺඉඈඋඍൾ (1840):177 (characters);Kංൾඌൾඇඐൾඍඍൾඋ (1848): 52 (characters); (1851): 419 (distribution); GඋൺඏൾඇHඈඋඌඍ (1847): 227 (characters); RൾൽඍൾඇൻൺർHൾඋ (1849): 709, (1857): 201, (1874): 198 (characters); Hൺඋൽඒ (1851): 32 (biology); Fൺංඋආൺංඋൾ &amp; Lൺൻඈඎඅൻජඇൾ (1856): 537 (characters), Kඋൺൺඍඓ (1857): 500 (characters); WൺඍൾඋHඈඎඌൾ (1858): 23 (synonymy); THඈආඌඈඇ (1859): 25 (notes on types); (1860): 173 (synonymy); GൾආආංඇGൾඋ &amp; Hൺඋඈඅൽ (1868): 570 (synonymy); Fൺඎඏൾඅ (1874): 290 (characters and distribution); Sൾංൽඅංඍඓ (1875): 267 (characters); Mඎඅඌൺඇඍ &amp; Rൾඒ (1876): 692 (characters); Fඈඐඅൾඋ (1888): 237 (characters); Rൾංඍඍൾඋ (1909): 112 (characters); JඈHൺඇඌൾඇ (1914): 364 (characters); Gඋංൽൾඅඅං (1924): 81, (1962a): 320 (characters); Pඈඋඍൾඏංඇ (1929): 341 (characters); MඣඊඎංGඇඈඇ (1937): 3 (synonymy); Hൺඇඌൾඇ (1952): 142 (characters); Sආൾඍൺඇൺ (1958): 364 (characters and biology); (1962a): 136, (1965a): 38 (characters); (1962b): 134 (larval characters); (1964): 80, (1976a): 172, (1978a): 85, (1981): 638, (1990): 98, (1993): 50 (distribution); (1971): 133 (distribution; introduction to Nearctic); (1973): 1427 (introduction to Nearctic);KඈඋGൾ (1960b): 53 (characters); Cඈංൿൿൺංඍ (1961): 52, (1963): 423, (1978):194 (characters); Pൺඅආ (1963a):141 (characters); LඈHඌൾ (1964): 212 (characters); Dඏඈřák 1965: 89 (distribution); Pඈඍඈඍඌĸൺංൺ (1967): 82 (larval characters) Kൺඌඎඅൾ (1968a): 66 (biology); (1970): 61 (larval characters); Sඓඎඃൾർĸං (1968): 736 (distribution); Oඌൾඅ-අൺ &amp; Zൺඇൾඍඍං (1975): 120 (biology); Bඈඋൽඈඇං (1976a): 95 (characters); SHංඅඈඏ (1975): 376 (distribution); Pඈඉൾ (1977): 31 (distribution); TඬඍH (1984):120 (characters);SൾGൾඋඌ(1987):182 (morphology);WൾඅർH (1993): 229 (morphology);Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං (1995):32 (distribution); Sඍൺඇංൾർ (1996):117 (pupal characters);Dඈඐඇංൾ &amp;Aඋඇൾඍඍ (1996):391 (characters); Oඐൾඇ (2000): 252 (biology); AඌඌංඇG (2001): 75 (biology); UHඅංG et al. (2006): 55 (distribution); Mൺඃĸൺ (2007): 949 (distribution); Dൺඉĸඎඌ &amp; Tൺආඎඍංඌ (2008): 33 (distribution); Cඎඉඉൾඇ et al. (2011): 15 (distribution); Sൾආංඈඇൾඇĸඈඏ et al. (2015): 335 (distribution).</p><p>Type material examined. Staphylinus molochinus: Sඒඇඍඒඉൾ J (ZMHB), labelled:“6230 / Braunsberg Dahl leg./ molochinus gr.gyll. / (Para)? typus Grav. / det. H. Korge Quedius molochinus Grav. Typus!”.</p><p>Quedius niger: Sඒඇඍඒඉൾ ♀ (MZH), labelled:“Tjudi / J. Sahlbg./ 1617. / Mus.Zool.H:fors Spec.typ. No 157 Qued.molochinus var.niger J. Sbg. / Syntype / http://id.luomus.fi/GZ.83 / Photographed 2020 Pekka Malinen”. Additional material examined. AUSTRIA: Arbesbach, [48.49, 14.95], 13.VII.1957, leg. Schubert (1 ♀ NMW); Admont, [47.56, 14.45], leg. H. Franz (1 ♀ NMW); Prater, [48.21, 16.39], 19.IX.1964 (1 J NHMD). CZECH REPUBLIC: Iser [Jizera] River, n. Benešov u Semil, [50.60, 15.36], 350 m, 24.VII.1982, leg.Hieke (1♀ ZMHB);Pastviny n.Žamberk, [50.09, 16.56], 31.VII.-3.VIII.1982, leg. Hieke (1 ♀ ZMHB); Podyji Nat. Park,forest, at river Dyje valley ca. 2.5 km NW of Havraníky,48.83,15.98, 240 m, sifted flood debris, 08.VI.2016, leg. A. Solodovnikov, J. Jenkins Shaw,&amp;A. Hansen (1J NHMD); Silesia Altvater [Praděd], [50.08, 17.23], leg. Letzner (1 SDEI); Teplice nad Metují, [50.59, 16.16], 3.-5.VIII.1992, leg.Hieke (1♀ ZMHB);Vysoká Hola, NW Karlov, Malá Morávka, [50.08, 17.23], sifting litter of Pinus mugo, 25.VII.1993, leg. Zerche (1 SDEI). DENMARK: Hjelm Hede, [56.49, 8.91], 20.VII.1951, leg. J. Petersen (2 ♀♀ NHMD). BඈඋඇHඈඅආ: Øle Å, 19.IV.1965, [55.05, 15.02], leg. F. Bangsholt (1 ♀ NHMD). Fඎඇൾඇ: Bjergsted, [55.66, 11.35], 1.VI.1981, leg. M.Hansen (1 J NHMD);Ristinge Klint, [54.82, 10.61], 14.VII.1978, leg. M. Hansen (2 JJ NHMD). Zൾൺඅൺඇൽ: Furesøstien, [55.81, 12.41], 27.IX.1954, leg. J. Petersen (3 ♀♀ NHMD); Holmegårds Mose, [55.29, 11.82], 26.VI.1975, leg. M. Hansen (1 J NHMD); Ormø, [55.21, 11.45], 13.IX.1997, leg. O. Martin (1 ♀ NHMD); Rude Skov, [55.83, 12.47], 19.VIII.1948, leg. J. Petersen (1 ♀ NHMD). FINLAND: Kilpisjärvi, [69.04,20.81], 21.-22.VII.1985, leg.Hieke (1♀ ZMHB);Kiuruvesi,[63.65, 26.62], 6.VII.1951, leg. Linnavuori (1 J 2 ♀♀ ZMUO); Mutenia, Sodankylä,[68.02, 27.41], 13.IX.2001, leg.J.Itämies, O. Nenonen (1♀ ZMUO); Rymättylä,[60.37, 21.94],leg.Linnavuori (1♀ ZMUO);Saana,Enontekiö, [69.04,20.85], 10.VII.2013, M.Pentinsaari (1J ZMUO). FRANCE: Chigny,Oise River, [49.91, 3.77],Selchert (1J ZMHB). GERMANY: Bayern, Schleissheim,[48.24, 11.59], 15.V.1931, leg. Ihssen (1J ZMHB); Berlin--Marzahn, Hellersdorfer Berg, [52.53,13.58], 8.VIII.1997, leg. Hieke (1J ZMHB); Brunn, Dosse [52.91, 12.51], 16.III.1980 leg. E. Rössner (2 JJ SDEI); Engolling,Auerbach, [48.79,13.10], 5.IX.1978, leg.K.Koufrauen (1 J 1 ♀ ZMHB); Erzgebirge, Flohtelberg, [50.42, 12.95], 2.VI.1925, leg Uhmann (1 J SDEI); Finkenkrug nr. Berlin, [52.56, 13.03], 12.IV.1898, leg. Dahl (1 ♀ ZMHB); Flessenow, Schwerin, [53.75, 11.49], 4.VII.1968, leg.Uhlig (1 ZMHB); Hainleite, SW Günseroda and Seega,[51.32, 11.02], 10.VIII.1993, leg. F. Hieke (1 J ZMHB); Marburg in Hess, [50.81, 8.77], 16.IV.1904, leg. Strand (1 J ZMHB); Neuendorf, Teupitz, [52.11, 13.58], 30.IV.1992, leg. Renner (1 J ZMHB); Sarnow n.Anklam, [53.75, 13.63], 29.IX.1964, leg. F. Hieke (1 J ZMHB); Strausberg, 23.VII.1981, [52.56, 13.88], leg. H. Wendt (1 ♀ ZMHB); Thüringer Wald nr. Scheibe, [50.48, 11.05], under stones, 8.I.1957, leg. Dieckmann (1 SDEI); Willersdorf, [51.02, 8.85], VII (1 ♀ ZMHB). ITALY: Gello, [43.96, 10.45], VI.1922, leg. A. Strand (1 J NHMD); Zeccone n. Pavia, [45.26, 9.19], 17.X.1977, leg. P. Kanaar (1 J ZMHB). NORWAY: Hemnesberget, [66.22, 13.61], 8.-14.VII.1903, leg. E. Strand (2 JJ ZMHB); Kongsvinger Eis, [60.19, 12.01], 27.X.1980, leg. F. Midtgaard (1 J NHMD); Pasvik Lake, Sydvaranger, [69.34, 29.53], VI.1994, leg. G. Nietsch (1 J NMW); Utgardsjön, Rønningen,Austmarka, [60.01, 12.37], 28.VI.-10.VII.1975, leg.P.Kanaar (1 ♀ ZMHB). POLAND: Misdroy, [53.91, 14.44] (1 J ZMHB); Pölitz [Police], Pommern,[53.55, 14.57], 13.VI.1920,leg.E. Hanau (1♀ ZMHB); Riesengebirge [Krkonoše], [50.79, 15.57], 750m, moss, 16.X.1902, leg. Dahl (1 J ZMHB); Tamsel, [52.62, 14.70], 20.V.1920, leg. Kuntzen (1 ♀ ZMHB). RUSSIA: CH ංඍൺ Oൻඅൺඌඍ: Malkhanskiy Khrebet, pass to Maleta, 20 km S Maloarkhangel’sk, 50.2352 108.916, 19.-25.VIII.1998, leg. A. Shavrin (NHMD). KHൺඇඍඒ- Mൺඇඌං Aඎඍඈඇඈආඈඎඌ Dංඌඍඋංർඍ: Sovetsky region, Malaya Sosva Nature Reserve, near Cordon Hangokurt, road to Belaya Gora, 61.9504, 64.2489, 81 m, nr a roadside puddle with Polytrichum sp. on clay, 28.VI.2017, leg. A.B. Ryvkin (1 J cRyv). Kඈආං Rൾඉ: Ukhta surrounding,[63.57, 53.71], 29.VI.1974, leg.V.Shulov (6JJ 6 ♀♀ ZMHB). KඋൺඌඇඈඒൺඋඌK Kඋൺං: Niznaya Lebedyanka river, [61.96, 89.45], 23.VI.1992, leg. V.S. (1 J cRyv); Turukhanskiy Distr., Mirnoye, [62.28, 89.03] 6.-17.VII.1990, leg. L.B. Rybalov (1 J 1 ♀ cRyv). Lൾඇංඇ-G උൺൽ Oൻඅൺඌඍ: Saint-Petersburg, Sestroretsk, Rzhavaya kanava, [60.12, 29.96], 16.VIII.2009, leg.V.N.Prasolov (1J ZIN); Stary Petergof, [59.87, 29.87], forest, 7.VI.1997, in leaf litter, leg.A. Solodovnikov (1J NHMD) Yaschera, [58.92, 29.98], 18.V.1996, leg.L.Anisutkin (1J NHMD). Mඈ-ඌർඈඐ Oൻඅൺඌඍ: Moscow,Izmajlovskij Park,[55.75,37.62], 7.VII.1983, leg. A.Isaev (1J NHMD); Odintsovsky Distr., Zvenigorod Biological Station of MSU, [55.70, 36.72], mixed forest, 22.VIII.1984, leg. K.G. Mikhailov (2 JJ cRyv). MඎඋආൺඇඌK Oൻඅൺඌඍ: Kyndomys [Kindo] Peninsula,[66.65, 33.22], forest floor, 16.VII.1992, leg. Wegner (1 J ZMHB). Pൾඋආ Kඋൺං: Perm city, [58.01, 56.22], pitfall, 29.VIII.1978, leg. S.L. Esunin (1 J NHMD). Rൾඉ. ඈൿ Bඎඋඒൺඍංൺ: Barguzinsky Distr., Barguzin valley, 20 km E Ust-Barguzin, Gusikha, [53.42, 109.42], aspen forest, pitfalls, leg. V. Shilenkov (1 J NHMD); Vitimsky Nat. Preserve, lake Oron, mouth or Labaznyj creek, Oron hut, [54.82, 112.26], 27-30.VII.2000, leg. A. Shavrin (2 JJ NHMD). Rඒൺඓൺඇ Oൻඅൺඌඍ: Ryazan Prov., Spassky Distr., Oka nature reserve, [54.71, 40.85], leaf litter, 26.VI.1981, leg. Kormylutsyn &amp; Eskov (1J 1♀ cRyv). SඏൾඋൽඅඈඏඌK Oൻඅൺඌඍ: Ural,Visim Nature Reserve, [57.37, 59.77], 25.I.1985, leg L.P. Titova (2 JJ cRyv). Tඒඎආൾඇ Oൻඅൺඌඍ: Surgutsky Distr., Yugansky Nature Reserve, Ai-Magromsy River Basin, near ‘Medvezhii Ugol’, [59.59, 74.71], 22.IX.2002, leg.A.B. Ryvkin (1 J cRyv). UඅඒൺඇඈඏඌKൺඃൺ Oൻඅൺඌඍ: Inzenskij Distr., Argash village, [53.96, 46.25]‚ birch forest, 9.VII.1995, leg.A.Isaev (2JJ NHMD). SLOVAKIA: Inovec, [48.77, 18.04], IX.1931, leg. Čepelák (1 J ZMHB). SWEDEN: Dagsås-trakten, Hall, [57.06, 12.48], 26.X.1953, leg. J.T. Skovgaard (2 JJ 3 ♀♀ NHMD); Södermanand, Ornö, [59.05, 18.41], 15.V.1946, leg. J.T. Skovgaard (1 J 1 ♀ NHMD); Stockholm, Djurgården 23.III.1945, leg. J.T. Skovgaard (1 ♀ NHMD); Vitamölle, [55.69, 14.21], 1.VIII.1926, leg.Stephensen (1J NHMD). Gඈඍඅൺඇൽ: Lau,[57.28, 18.61], 25.VI.1950 leg. J.T. Skovgaard (1 J NHMD). SWITZERLAND: Reussdelta, Seedorf, [46.89, 8.61], 435 m, VIII.1998, leg. Herger &amp; Rezb. (1 J ZMHB). UNITED KINGDOM: EඇGඅൺඇൽ: Stanford le Hope, Essex,[51.51, 0.42], 7.IX.1969,leg.Wewalka (1♀ NMW). Sർඈඍඅൺඇൽ: Bridge of Orchy,[56.51, -4.76], 23.VII.1987, leg.Gillerfors (1♀ MZLU);S Braemer,[56.99,-3.39], 15.VII.1987, leg. Gillerfors (1 ♀ MZLU).</p><p>Redescription. Measurements JJ (n = 5): HW = 1.60– 1.78 (1.68); HL = 1.36–1.53 (1.44); HL/HW 0.83–0.88 (0.85); PW = 2.09–2.24 (2.20); PL = 1.93–2.11 (2.04); PL/PW 0.91–0.94 (0.93); EW = 2.00–2.18 (2.11); EL = 1.73–2.07 (1.92); EL/EW 0.87–0.95 (0.91); EL/PL 0.90– 1.02 (0.94); PW/HW 1.32–1.57 (1.47); forebody length 5.02–5.58 (5.40). ♀♀ (n = 5): HW = 1.60–1.69 (1.64); HL = 1.38–1.51 (1.45); HL/HW 0.86–0.93 (0.89); PW = 2.07–2.22 (2.13); PL = 1.91–2.09 (2.00); PL/PW 0.90–0.98 (0.94); EW = 2.04–2.18 (2.11); EL = 1.84–1.93 (1.91); EL/EW 0.88–0.95 (0.91); EL/PL 0.91–1.01 (0.96); PW/ HW 1.40–1.53 (1.47); forebody length 5.22–5.51 (5.35).</p><p>Medium sized, robust species; body dark brown to black (Figs 8C,D).</p><p>Head black, distinctly transverse, with eyes medium sized (EyL/TL = 1.67–2.13 (1.76)); microsculpture of transverse waves; no interocular punctures between anterior frontal punctures (cf. Fig. 6F); antennae pale reddish with antennomeres 3–5 slightly darkened, all antennomeres elongate; palpi pale reddish.</p><p>Thorax: pronotum dark brown to black, slightly wider than long, wider than head, with microsculpture of transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond level of first puncture of dorsal row; scutellum punctured and pubescent; elytra most often reddish brown rarely fully darkened, uniformly pubescent, slightly wider than long, roughly same length as pronotum; legs reddish brown with inner face of tibia and inner femur darkened.</p><p>Abdomen dark brown to black, tergites uniformly punctured, with slight iridescence.</p><p>Male. Aedeagus (Fig. 13D): paramere lanceolate with slight medial attenuation and extending into a slight expansion broadest below slightly asymmetric apex, reaching just beyond apex of median lobe, with sensory peg setae forming two long rows fusing together towards apex; median lobe broad with gentle constriction to a point at apex, on parameral side with two small teeth pointing slightly basad, positioned at level near basal level of peg setae band of paramere; C-shaped sclerite of internal sac with spine-like basal extension.</p><p>Differential diagnosis. Quedius molochinus is very similar to the other species with red elytra in the molochinus -group: Q. meridiocarpathicus, Q. balticus, and Q. vicinus . It can be distinguished from Q. balticus by the pale base of the basal three antennomeres. In most cases it has darker middle antennomeres and is generally slightly darker with abdomen black to dark brown (dark to light brown in Q. meridiocarpathicus and Q. vicinus). For confident discrimination of these species genitalia should be examined: aedeagus of Q. molochinus has an extension of the C-sclerite in the internal sac. Quedius molochinus with black elytra can be confused with Q. subunicolor where their ranges overlap. Usually, the lighter colored appendages and the coarser microsculpture of Q. molochinus are sufficient to distinguish it from Q. subunicolor . Quedius molochinus can be confused with Q. picipes from the subgenus Raphirus, which may occur in similar environments. It can be readily distinguished from the latter by more parallel-sided abdomen and entire labrum. In North America the species can be confused with Q. laticollis, from which it can be distinguished by stouter antennae with slightly darker middle antennomeres. Also, in Q. molochinus elytra are more often brick red or almost black, whereas in Q. laticollis they are most often dark brown.</p><p>Synonymic notes. Quedius picipennis Stephens, 1832 from London, Norfolk, Suffolk, Devonshire, and Somerset and Raphirus picipennis Stephens, 1833 from London have recently been mixed up as the same species in both Hൾඋආൺඇ (2001) and Nൾඐඍඈඇ (2020). Based on the original descriptions, Raphirus picipennis clearly belongs in the Quedius subgenus Raphirus, whereas Quedius picipennis fits Q. molochinus . Original descriptions of Staphylinus laevicollis Runde, 1935 described from Halle in Central Germany and Staphylinus lapponicus Zetterstedt, 1838 described from Finland fit Q. molochinus, consistently with the currently accepted synonymy.</p><p>Bionomics. Quedius molochinus is found in meadows, heaths and grasslands, as well as in mainly coniferous forests. Most records are from under rocks or in moss. We have encountered it in flood debris near a river forested on either side in Czech Republic, in Sphagnum moss of a bog in Southern Sweden, and from pitfall traps set in a heathland in Denmark. Records from Russia (Sൾආංඈඇൾඇ-ĸ ඈඏ et al. 2015), Switzerland (UHඅංG et al. 2006) and the Netherlands (Cඎඉඉൾඇ et al. 2011) are from meadows. In North America it seems to be in more ruderal habitats as well, having been found in e.g., potato fields on Prince Edward Island (Mൺඃĸൺ 2007).</p><p>Distribution. Among the closest congeners, Quedius molochinus is a more northern species widely distributed from France and British Isles through Central and Northern Europe, across European Russia and western Siberia to Transbaicalia (Fig. 20). In the southern areas of its distribution, it gradually moves to higher elevations, for example in the Alps extending into the Apennines. More thorough sampling in Russia may widen its distribution significantly. Quedius molochinus is introduced in North America, where it is currently found in three separate areas in Canada: around the Avalon Peninsula of Newfoundland (Sආൾ-ඍൺඇൺ 1981), in the Canadian Maritime Provinces (Nova Scotia, New Brunswick and Prince Edward Island, Mൺඃĸൺ 2007) and around Quebec City (Sආൾඍൺඇൺ 1981) (Fig. 12).</p></div>	https://treatment.plazi.org/id/038987A0FFD84B3DB392A0B917E2F8CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFDE4B3CB11AA7781113FACA.text	038987A0FFDE4B3CB11AA7781113FACA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius subunicolor Korge 1961	<div><p>Quedius subunicolor Korge, 1961</p><p>(Figs 1, 2F, 4, 7F, 13C, 20)</p><p>Quedius subunicolor Korge, 1961: 81 [Type locality: Schweden, Häggenås, Jämtland]</p><p>Quedius altaicus Korge, 1962: 152 [Type locality: Zentral-Altai], syn. nov.</p><p>References. Pൺඅආ (1962): 187, (1963): 142 (characters); TංĸHඈආංඋඈඏൺ (1973): 189 (distribution); SHංඅඈඏ (1975): 376 (distribution); Cඈංൿൿൺංඍ (1978): 193 (characters); LඈHඌൾ &amp; LඎർHඍ (1989): 172 (biology); Sൺඅ-ඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2018a): 128–130 (redescription, suspected synonym of Q. altaicus); (2019): 47 (key and characters).</p><p>Type material examined. Quedius subunicolor: Pൺඋൺඍඒඉൾඌ (MZLU):1J, labelled: “Häggenäs s-n Jtl.T. Palm 4–8, 8 1945 / det. H. Korge Quedius subunicolor Korge / Paratypus subunicolor Korge / Quedius subunicolor Korge / Type no. 1202:2 MZLU/ 2016 189 MZLU”; 3♀♀, same data, but last two labels “Type no: 1202:3 MZLU/ 2016 190 MZLU”.</p><p>Quedius altaicus: Hඈඅඈඍඒඉൾ: ♀ (FMNH), labelled: “Zentral-Altai, lg. Leder, det.Bang-Haag / unicolor Kies. det.Bernhauer / ♀ Holotypus Quedius s. str. altaicus H. Korge / Chicago NHMus M. Bernhauer Collection / Holotype teste D.J. Clarke 2014 GDI Imaging Project / Photographed Kelsey Keaton 2014 Emu Catalog / FMNHINS 2819427 Field Museum”. Additional material examined. FINLAND: Kuusamo, [65.96, 29.17] (1 ♀ ZMUO); Savukoski, [67.29, 28.15], 15.-25.VI.2006, leg.M.Mutanen, P.Välimaki &amp; M.Pentisaari (1J ZMUO). KAZAKHSTAN: Altai, Listvyaga, SSW Gora Tesninskiy Belok, Seredchikha River, [49.51, 85.25], 1200-1500 m, 27.VII.1997, R. Dudko &amp; V. Zinchenko (1 J NHMD); Bukhtarma riv., Uryl-Chingistai, [49.19, 85.88], 13.VI.1987, leg. V.A. Kastcheev (1 J ZIN); Rahmanovskie Kluchi, [49.53, 86.50] 14.-16. VI.1980, leg.V.A. Kastcheev (1 J ZIN); Saur ridge, Uidene riv., [47.20, 85.32], 1600-1800 m, 15.VII.2011, leg.V.A. Kastcheev (10J 2♀♀ ZIN). RUSSIA: Aඅඍൺං Rൾඉ.: Source of Kuma river, 50.9989, 84.1647, 1479 m, mixed forest, under stones, 4.VII.2019, leg. A. Solodovnikov, A. K. Hansen, &amp; M. J. Justesen (2 NHMD); Source of Kuma river, 50.9806, 84.1652, 1800 m, under stones, timber line, 3.VII.2019,A. Solodovnikov, A. K. Hansen, &amp; M. J. Justesen (1 J NHMD). Rൾඉ. ඈൿ Bඎඋඒൺඍංൺ: Lake Baikal, Svjatoy Nos, [53.69, 108.86], 9.VIII.1997, leg. A. Shavrin (1 J NHMD). SWEDEN: Barsele, [65.04, 17.45], leg. Alan Dufberg (1 MZLU); Revsund, [62.89, 15.12], 27.V.1970, leg. R. Bergvall (3 MZLU).</p><p>Redescription. Measurements JJ (n = 6): HW = 1.67– 1.78 (1.74); HL = 1.42–1.56 (1.49); HL/HW 0.80–0.89 (0.86); PW = 2.24–2.40 (2.32); PL = 2.04–2.20 (2.12); PL/PW 0.90–0.96 (0.91); EW = 2.22–2.56 (2.46); EL = 2.29–2.47 (2.36); EL/EW 0.92–1.03 (0.96); EL/PL 1.06– 1.17 (1.11); PW/HW 1.50–1.64 (1.56); forebody length 5.78–6.04 (5.97). ♀♀ (n = 3): HW = 1.67–1.76 (1.73); HL = 1.44–1.49 (1.47); HL/HW 0.82–0.88 (0.85); PW = 2.13–2.33 (2.25); PL = 1.89–2.02 (1.97); PL/PW 0.83–0.94 (0.88); EW = 2.11–2.47 (2.30); EL = 2.00–2.36 (2.24); EL/EW 0.95–1.01 (0.97); EL/PL 1.00–1.25 (1.14); PW/ HW 1.45–1.62 (1.54); forebody length 5.47–5.82 (5.67).</p><p>Medium sized, robust species; body black (Fig. 7F).</p><p>Head black, distinctly transverse, with eyes medium sized (EyL/TL = 1.88–2.13 (2.02)); microsculpture of fine transverse waves, in some areas almost meshed; no interocular punctures between anterior frontal punctures (cf. Fig. 6F); antennae brown with antennomeres 1–5 clearly darkened, all antennomeres elongate; palpi brown.</p><p>Thorax: pronotum black, slightly wider than long, wi-der than head, with microsculpture of transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond level of first puncture of dorsal row; scutellum punctured and pubescent; elytra black, uniformly pubescent, slightly wider than long, roughly as long as–slightly longer than pronotum; legs dark brown to black, tarsi usually lighter pale brown.</p><p>Abdomen black, tergites uniformly punctured, with slight iridescence.</p><p>Male. Aedeagus (Fig. 13C): paramere lanceolate with no or very slight medial attenuation, reaching just beyond apex of median lobe, with rather large sensory peg setae forming two long single rows fusing together towards apex; median lobe broad with gentle constriction to a point at apex, on parameral side with two small teeth pointing slightly basad, positioned at level near basal level of peg setae band of paramere; internal sac without a continuation of C-sclerite.</p><p>Differential diagnosis. Quedius subunicolor is very similar to the species of the molochinus -group with dark elytra – Q. unicolor, Q. sundukovi – and to the dark forms of Q. molochinus . It differs from Q. unicolor by the microsculpture of head consisting of transverse waves and rarely becoming meshed (clearly meshed frons appearing dull in Q. unicolor). Also, genitalia are clearly different in both species, and they have completely non-overlapping distributions. Quedius subunicolor can be clearly distinguished from Q. sundukovi by the presence of the palisade fringe of tergite VII, which is missing in Q. sundukovi . It usually differs from the dark forms of Q. molochinus in the darker colored appendages and finer microsculpture. If in doubt the internal sacs are also clearly different in both species: in Q. subunicolor C-sclerite lacking an extension, which is present in Q. molochinus .</p><p>Synonymic notes. KඈඋGൾ (1961) described Quedius subunicolor from Sweden for fully darkened specimens that originally were assigned to Q. unicolor, a species which is actually restricted to the central and southern European mountains. Additional records showed that Q. subunicolor was rather widespread in Scandinavia (Pൺඅආ 1962, 1963; TංĸHඈආංඋඈඏൺ 1973). Nearly at the same time, KඈඋGൾ (1962) described Q. altaicus based on two females from an unspecified locality in ‘central Altai’. Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2018a) made a thorough morphological comparison between Q. altaicus and Q. subunicolor, including type material and demonstrated that the external characters claimed by KඈඋGൾ (1962) as unique for Q. altaicus compared with Q. subunicolor (microstructure of the head, proportions of the pronotum, chaetotaxy of the head and pronotum) vary continuously between both species. They suspected a synonymy and suggested that molecular comparisons of the two species should be conducted. The results of such an analysis here, that included newly collected material, suggest that specimens from Northern Europe and Altai are identical based on COI barcode (0.6% intraspecific variation among them in one clade; Table 2; Fig. 4). Since there is continuous morphological variation and no clear separation in COI, there is no evidence for separate species. We therefore propose that Quedius altaicus Korge, 1962 syn. nov. is placed in synonymy with Quedius subunicolor Korge, 1961 .</p><p>Quedius subunicolor has historically been confused with Q. unicolor . Many older specimens from Scandinavia and Northern Russia have been assigned to Q. unicolor, although these belong to Q. subunicolor . More recently, it has been erroneously reported from the Bohemian Forest in Central Europe, e.g., BඈHගඹ &amp; Mൺඍෂඃටඹൾĸ (2004).</p><p>Bionomics. In Northern Europe Q. subunicolor is almost exclusively found in bogs and fens. This is a type of habitat that historically has often been drained for agriculture and peat gathering, thus the species may be rarer now than it used to be. In the Altai Mountains all hitherto known specimens were collected at the elevations 1200–2000 m. We (AKH and AS) have spent six days in the western Altai mountains in the early summer of 2019 searching for this species. There we sifted many types of debris, but without any luck. It was not until we started turning stones just below the timber line that we found the first specimen. In spite of six days of focused collecting, we encountered only three specimens, all collected from under rocks near streams in forested areas (Fig. 2F). The low number of specimens we encountered is in agreement with the very few numbers of specimens available in museums. The species is likely very rare and patchily distributed across its range. Contrary to the morphological and molecular homogeneity of this species, we see a clear difference in habitat preference between the Scandinavian populations restricted to bogs and Altaian populations confined to the banks of mountain creeks at higher altitudes. Such variation in habitat preference within the distribution range is not unique and is also found in other members of Quedius sensu stricto, e.g., Q. labradorensis . Presumably, these different habitats provide the same or similar parameters of temperatures and moisture for a species within a wide range.</p><p>Distribution. Quedius subunicolor currently displays a disjunct distribution between Northern Europe and Altai Mountains, with few specimens known from the Ukhta District in the Komi Region of Russia (SHංඅඈඏ 1975) and from Lake Baikal listed herein (Fig. 20). It is still unknown if Q. subunicolor represents a species continuously distributed from Northern Europe to Altai, or if it is a hitherto unrecorded case of a boreo-montane disjunction between Northern Europe and Siberian mountains. Further sampling should be done in the areas of the apparently large distributional gaps.</p></div>	https://treatment.plazi.org/id/038987A0FFDE4B3CB11AA7781113FACA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFDF4B31B3BDA57817D6FA8A.text	038987A0FFDF4B31B3BDA57817D6FA8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius sundukovi Smetana 2003	<div><p>Quedius sundukovi Smetana, 2003</p><p>(Figs 1, 4, 7, 8F, 13B, 15, 20)</p><p>Quedius sundukovi Smetana 2003: 189-193 [Type locality: Badshalskyi Khrebet mountains]</p><p>References. Sආൾඍൺඇൺ &amp; SHൺඏඋංඇ (2018): 834 (distribution); Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2018a):130–131 (distribution); (2019):50 (characters and distribution).</p><p>Material examined. KAZAKHSTAN: SW Altai, East of Narymskij Mt. Ridge, upper course of Ozernaja River, 49.0478, 85.1614, 1900-2300 m, supalpine zone, 18.VII.1997, leg.R.Yu.Dudko &amp; V.K. Zinchenko (ZIN). RUSSIA: JൾඐංඌH Aඎඍඈඇඈආඈඎඌ RൾG.: Kuldur vill., 49.1854, 131.6171, pitfall trap, stream, 480m, 14.-15.VII.2017, leg.A.Bergmann (1 J cSch). Eൺඌඍ YൺKඎඍංൺ: Suntar-Khayata Mts, Valley of Tyry river valley, Kidyrki river near Khalya, [62.35, 138.46], 800-950 m, 14.-19.VIII.1991, leg. Alexeev (NHMD). KHൺൻൺඋඈඏඌK Tൾඋඋ.: Badshalskyi chr., Omot lake, [50.54, 134.26], tundra, 1850-2050 m, 7.-19.VII.1997, leg A.Plutenko (2 JJ 2♀♀ cSch); Levaja Bureya riv., estuary Don riv., [51.68, 134.51], 24.- 25.VII.2006, leg. U. Valainis (ZIN); Levaja Bureya riv., the left bank of the Imganakh river near the mouth, 51.7355, 134.572, 650-670 m, sloppy rocky hills with burrow pits, litter and mosses of Picea ajanensis, Abies nephrolepis, Larix gmelinii, Rhododendron dauricum, Betula platyphylla, Hylocomium splendens, Pleurozium schreberi, Ptilium crista-castrensis, Dircranum sp., Polytrichum sp., Sphagnum girgensohnii, Sph. sp., 26.VI.2011, leg.A.B. Ryvkin (ZIN);Verkhnebureinsky natural park, Ust--Urgal project gauging station Ust-Niman Topolevnik, 51.3995, 132.733, 1.IX.2009, leg. L.A. Trilikauskas (NHMD); Verkhnebureinsky natural park, Ust-Urgal project gauging station Ust-Niman Topolevnik, 51.3995, 132.733, 315 m, Padus, Alnus, Salix, Picea, Plagiomium, 4.VIII.2009, leg. A.B. Ryvkin (ZIN). SൺKHൺඅංඇ Iඌඅൺඇൽ: 13 km N from Korsakov, 2km N from Solovyovka, [46.74, 142.73], Acer, Betula, 5.VIII.1992, leg. V. Gusarov, (CNC); 13 km N from Korsakov, 2 km N from Solovyovka, [46.74, 142.73], Abies, Larix, Ledum, Betula, 6.VIII.1992, leg.V.Gusarov (CNC). ZൺൻൺඒKൺඅඌKඒ Kඋൺං: Stanovoy Highlands, W part of Kodar, top Chara River, 50 km WSW River Novaya Chara, [56.65, 117.54], 1700- 2000 m, 27.VII.1995, leg.A. &amp; R. Dudko, D. Lomakin (ZIN); Stanovoy Highlands, top Chara River, Lake Leprindo, 56.636, 117.537, 1000 m, 23.VII.1995, leg. A. &amp; R. Dudko, D. Lomakin (ZIN).</p><p>Redescription. Measurements JJ (n = 5): HW = 1.40– 1.69 (1.53); HL = 1.22–1.40 (1.29); HL/HW 0.79–0.89 (0.85); PW = 1.82–2.22 (2.00); PL = 1.71–2.07 (1.81); PL/PW 0.80–0.96 (0.91); EW = 1.80–2.09 (1.88); EL = 1.44–1.71 (1.60); EL/EW 0.80–0.89 (0.85); EL/PL 0.83–0.95 (0.89); PW/HW 1.46–1.67 (1.55); forebody length 4.38–5.18 (4.70). ♀♀ (n = 5): HW = 1.58–1.73 (1.67); HL = 1.33–1.42 (1.38); HL/HW 0.78–0.87 (0.83); PW = 1.93–2.24 (2.04); PL = 1.80–2.11 (1.88); PL/PW 0.90–0.94 (0.92); EW = 1.89–2.09 (1.96); EL = 1.69–1.78 (1.72); EL/EW 0.83–0.91 (0.88); EL/PL 0.82–0.96 (0.92); PW/HW 1.39–1.58 (1.48); forebody length 4.82–5.27 (4.99).</p><p>Small sized species; body dark brown to black (Fig. 8F).</p><p>Head black, distinctly transverse, with eyes medium sized (EyL/TL = 1.73–2.00 (1.83)); microsculpture of fine transverse waves; no additional punctures between anterior frontal punctures (cf. Fig. 6F); antennae dark internally becoming continuously lighter, all antennomeres slightly elongate; palpi pale with apical palpomere slightly darkened.</p><p>Thorax: pronotum black, wider than long, wider than head, with microsculpture of transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond level of first puncture of dorsal row; scutellum sparsely punctured and pubescent; elytra most often fully darkened but occasional dark reddish brown, short uniformly pubescent, clearly shortened wider than long, clearly shorter than pronotum; legs dark with tarsi lighter.</p><p>Abdomen black, tergites sparsely and uniformly punctured, without palisade fringe of tergite 7, without clear iridescence.</p><p>Male. Aedeagus (Fig. 13B): paramere lanceolate with slight medial attenuation and extending into a slight expansion broadest below apex, apex slightly asymmetric, reaching just beyond apex of median lobe, with sensory peg setae forming two long rows fusing together towards apex (Fig. 15); median lobe broad with gentle constriction to a point at apex, on parameral side with two small teeth pointing slightly basad, positioned at level near basal level of peg setae band of paramere; internal sac without a continuation of C-sclerite.</p><p>Differential diagnosis. Quedius sundukovi is most similar to Q. unicolor, Q. subunicolor, and Q. molochinus . It can easily be distinguished from all these species by its smaller size and brachypterous habitus with shortened elytra and absent palisade fringe on tergite VII. If in doubt, the aedeagus can be checked for characters listed in the diagnosis. It is also similar to Siberian species Q. (Raphirus) jenisseensis Sahlberg, 1880, from which it is easily distinguished by the entire labrum (medially incisioned in Q. jenisseensis).</p><p>Comments. Quedius sundukovi was described by Sආൾඍൺ-ඇൺ (2003) based on 13 specimens from the Badshalskyi Khrebet mountains in Khabarovsk Krai in the Far East of Russia. Sආൾඍൺඇൺ &amp; SHൺඏඋංඇ (2018) reported additional specimens from Vitimskiy Nature Reserve and the Udokan Plateau extending the species range significantly westwards in Russia to Transbaikalia. Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2018a) reported a number of records of this species from the Altai Mountains in Kazakhstan. Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈ-ൽඈඏඇංĸඈඏ (2019) reported the occurence of this species in Buryat Republic in Transbaikalia and in the Russian Far East regions of Amur Oblast and Sakhalin Island (Fig. 20). Here we add detailed information on many of these records along with COI barcodes from specimens across the range (Fig. 4). The barcodes showed high variation as we recovered five OTUs (four BINs) with as high as 7.7% divergence between the OTUs (Table 2). Morphological studies of the specimens also revealed some variation in size – but not in the shape – of the paramere (Fig. 15). We found no differences to correspond to any of the molecular clusters, which also did not cluster geographically. Lack of a clear congruence between the molecular and morphological variation, as well as no hiatus between different morphological variants, suggests that they all represent a single species. The species is known across a large area with complex montane orography and it is flightless, with the palisade fringe of abdominal segment VII absent and clearly shortened elytra.Apparently, the high degree of the intraspecific variation can be related to the flightlessness of the species leading to low dispersal abilities of members of different populations and accelerated rates of divergence in the barcode region (Mංඍඍൾඋൻඈൾർĸ &amp; Aൽൺආඈඐංർඓ 2013).</p><p>Bionomics. Little is known about the specific habitat requirement of Q. sundukovi . Based on the examined material, the species appear to occur in various leaf litter, both coniferous and deciduous, and is commonly associated with talus debris and moss, often near streams or rivers. Specimens are found within a wide range of elevations from 315 to 2300 m, at higher elevation at lower latitudes.</p><p>Distribution. Quedius sundukovi is currently known from relatively few specimens scattered across a large range comprising the Altai Mountains, Transbaikalia, Bureya and the Suntar-Khayata Range (Fig. 20). It is also found on Sakhalin Island.Although currently known from only a few localities, the species distribution is most likely continuous. The range maybe poorly known because it spans through an area with very little recent and historical entomological activity, especially for leaf-litter dwelling organisms.</p></div>	https://treatment.plazi.org/id/038987A0FFDF4B31B3BDA57817D6FA8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFD24B30B0EFA5381011FEEA.text	038987A0FFD24B30B0EFA5381011FEEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius unicolor Kiesenwetter 1847	<div><p>Quedius unicolor Kiesenwetter, 1847</p><p>(Figs 1, 2G, 4, 7E, 14A, 21)</p><p>Quedius unicolor Kiesenwetter in Kංൾඌൾඇඐൾඍඍൾඋ &amp; Mඟඋĸൾඅ (1847: 75). [Type locality: Riesengebirge]</p><p>References. Kංൾඌൾඇඐൾඍඍൾඋ (1848): 53 (characters); RൾൽඍൾඇൻൺർHൾඋ (1849): 825, (1857): 202, (1874): 199 (characters); Fൺංඋආൺංඋൾ &amp; Lൺൻඈඎඅ-ൻජඇൾ (1856):536 (characters); Fൺඎඏൾඅ (1874):291 (characters);Mඎඅඌൺඇඍ &amp; Rൾඒ (1876): 697 (characters); GൺඇGඅൻൺඎൾඋ (1895): 403 (characters); Pඈඋඍൺ (1907):132 (characters);Rൾංඍඍൾඋ (1909):112 (characters); Gඋංൽൾඅඅං (1924): 80 (characters); Pඈඋඍൾඏංඇ (1929): 341 (characters); WඳඌඍHඈൿൿ (1938): fig. 24 (characters); Sආൾඍൺඇൺ (1955): 143, (1960a): 259, (1964): 168, (1966): 331, (1993): 50 (distribution); (1958): 363 (characters and biology); (1962a): 134 (characters);Cඈංൿൿൺංඍ (1961): 49,(1978):193 (characters);LඈHඌൾ (1964):211 (characters); Hൺඏൾඅĸൺ (1964):93 (distribution); Hඈඋංඈඇ (1965): 274 (distribution); Bඈඋൽඈඇං (1976a): 94 (characters); ZൾඋർHൾ (1977): 9 (distribution), TඬඍH (1984): 119 (characters); LඎർHඍ (1987):109 (distribution);Jൺඇගĸ (1992):91 (distribution);Hൾඋආൺඇ (2001): 50 (application to ICZN), Tඋඈඇඊඎൾඍ (2006): 102 (distribution); Mൺඓඎඋ et al. (2007): 30 (distribution); MൺඍൾඅൾඌHĸඈ (2007): 184 (distribution); SർHൺඍඓ (2008): 388 (distribution); Zൺඇൾඍඍං (2015): 96 (distribution).</p><p>Material examined. AUSTRIA: Golling, [47.59, 13.16], leg. Skalitzky (1 ♀ NMW); Kerschbaumeralm, [46.75, 12.76], 24.VII.-7.VIII.1948, leg. F. Schubert (1 ♀ NMW); Koralpe, [46.88, 14.99], leg. A. Otto (3 JJ NMW); Lünersee, [47.05, 9.74], 14.VI.-3.VII.1952, leg. F. Schubert (1 ♀ NMW); Stuhleck, [47.56, 15.78], leg. Breit (1 ♀ NHMD); Stuhleck, Steiermark, [47.56, 15.78], leg.Wingelmüller (1 ♀ NMW); Tirol, [47.22, 11.51] (1 SDEI); Wechsel,[47.52, 15.91], leg. Scheerpeltz (3♀♀ NMW); Wolayersee, [46.61, 12.86], 3.-11.VIII.1949, leg. F. Schubert (2 JJ 1 ♀ NHMD); Zillertal nr. Mayrhofen, [47.16, 11.87], 4.VIII.1914, leg. H. Wagner (1 SDEI); Wolayersee, [46.61, 12.86], 3.11. VIII.1949, leg. F. Schubert (2 ♀♀ NMW). CZECH REPUBLIC: Krkonoše [50.71, 15.66], leg. Obenberger (1 J 2 ♀♀ NHMD); Spindelmühle [Špindlerův Mlýn], [50.73, 15.61], leg. Skalitzky (1 NHMD). GERMANY: Harz, Brocken, [51.79, 10.61], 9.V.1922, leg. Uhmann (1 J SDEI); Hercyn mont.[Harz Mountains], [51.74, 10.63], 14.VI.1914, leg. Beckenkamp (3 ♀♀ SDEI); Thüringer Wald Oberhof, [50.71, 10.73], 8.IX.1951, leg. W. Liebmann (1 ♀ SDEI). ITALY: Gressory [Gressoney] la Trinite, [45.87, 7.82], 15.VII.1901, leg. Künnemann (1 SDEI); Malga Preghena Alta, Trento, 46.4148, 10.8922, 2100 m, under stones along stream with moss, 26.VIII.2019, leg.A.Zanetti (NHMD);Valle Brembana, Strada di Porcile, [46.06, 9.73], 1800-2000 m, 26.VI.1961, leg. V. Rosa (1 J NMW). PO-LAND: Silesia [Bad] Flinsberg [Świeradów-Zdrój], [50.92, 15.31], leg. Kraatz (1 SDEI). SPAIN: Cantabria, Alto Campoo, W Reinosa, [43.03, -4.37], 4.VI.1991 2000m leg. Zerche (1 SDEI). SLOVAKIA: Ružomberok, [49.04, 19.23], VII.1952, leg. Přívora (1 J NHMD); High Tatras, Mengušovská dolina, Vysoke Tatry, [49.16, 20.07], 1200 m, 17.V.1991, leg. J. Frisch (1 J ZMHB); Tatra, Kráľova hoľa, [48.88, 20.13], 1300- 1600 m, 12.VIII.1981, leg. Hieke (1 J ZMHB). SWITZERLAND: Aarau, [47.38, 8.05], leg. Stierlin (1 SDEI).</p><p>Redescription. Measurements JJ (n = 5): HW = 1.47– 1.76 (1.58); HL = 1.31–1.44 (1.36); HL/HW 0.82–0.91 (0.86); PW = 1.96–2.33 (2.16); PL = 1.78–2.02 (1.91); PL/PW 0.86–0.92 (0.89); EW = 2.13–2.47 (2.32); EL = 2.18–2.47 (2.32); EL/EW 0.95–1.05 (1.00); EL/PL 1.15– 1.26 (1.22); PW/HW 1.47–1.65 (1.58); forebody length 5.36–5.93 (5.58). ♀♀ (n = 5): HW = 1.58–1.67 (1.61); HL = 1.31–1.42 (1.37); HL/HW 0.79–0.89 (0.85); PW = 2.00–2.18 (2.08); PL = 1.82–1.89 (1.87); PL/PW 0.87–0.92 (0.90); EW = 2.22–2.40 (2.31); EL = 2.20–2.40 (2.31); EL/EW 0.96–1.01 (0.99); EL/PL 1.19–1.27 (1.23); PW/ HW 1.45–1.57 (1.52); forebody length 5.36–5.69 (5.52).</p><p>Medium sized species; body black (Fig. 7E).</p><p>Head black, distinctly transverse, with eyes rather small (EyL/TL = 1.59–2.00 (1.82)); microsculpture of fine mesh, especially on frons, appears dull; no interocular punctures between anterior frontal punctures (cf. Fig. 6F); antennae dark brown to black, all antennomeres slightly elongate; palpi dark brown to black.</p><p>Thorax: pronotum black, slightly wider than long, clearly wider than head, with microsculpture of fine transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond first puncture of dorsal row; scutellum punctured and pubescent; elytra black, uniformly pubescent, as long as wide, clearly longer than prontoum; legs dark, outer face of tibia and tarsi often paler.</p><p>Abdomen black, tergites uniformly punctured, without clear iridescence.</p><p>Male. Aedeagus (Fig. 14A): paramere lanceolate with slight medial attenuation and extending into a slight expansion broadest below apex, apex extending into small blunt knob away from the median lobe, apex slightly asymmetric, reaching just beyond apex of median lobe, with sensory peg setae forming two rows fusing together towards apex; median lobe broad with gentle constriction to a point at apex, on parameral side with two small ridges continuing basad, positioned at level near basal level of peg setae band of the paramere; internal sac without C-sclerite.</p><p>Differential diagnosis. Quedius unicolor is very similar to other species with dark elytra from the molochinus- group – Q. subunicolor, Q. sundukovi and the dark form of Q. molochinus . It can be distinguished from Q. subunicolor by the dull frons due to meshed microsculpture (in Q. subunicolor the microsculpture consists of fine transverse waves or only slightly meshed). Also, the genitalia and the distributions are clearly different in the two species. It usually differs from the dark form of Q. molochinus in the darker appendages and the finer microsculpture in addition to a clearly different aedagus, which is without a C-sclerite in Q. unicolor, but with such in Q. molochinus . Quedius unicolor has non-overlapping distribution with Quedius sundukovi and can be clearly distinguished from the latter species by the presence of palisade fringe on tergite VII.</p><p>Quedius unicolor has historically been confused with Q. subunicolor . For details see comments about the latter species.</p><p>Comments. Quedius unicolor was originally described from Riesengebirge [Krkonoše] (Kංൾඌൾඇඐൾඍඍൾඋ &amp; Mඟඋĸൾඅ 1847), a mountain range at the border between Poland and Czech Republic. It was later discovered in other mountainous area, including the Alps, the Carpathians, the Pyrenees, Picos de Europa, the Harz Mountains, and possibly extending into Sierra Nevada and even central Anatolia, although the latter records are not confirmed here (Cඈංൿൿൺංඍ 1961, AൻൺർංGංඅ et. al 2009; Fig. 21). At these sites it seems to be restricted to a very specific habitat type, with most specimens being found under stones and in moss near mountain streams at higher elevations. COI barcodes of specimens from several sites, including the Austrian and Italian Alps, the Carpathians of Slovakia, and Picos de Europa in Spain displayed high genetic variability (Fig. 4), with the sites being grouped into three OTUs (the Carpathians, Picos de Europa, and the Alps) ranging from 3.5 % to 4.5 % divergence, with a 0.2% divergence between the Austrian and Italian Alps (Table 2). With such a high genetic divergence in the barcode region, we carefully checked for morphological differences. However, we found no obvious differences corresponding to the molecular clusters. Since the wings seem to be fully developed in this species, these genetic differences may be attributed to isolation by distance, the very specific habitat requirements of the species and its close association to a now fragmented habitat, which would limit gene flow among populations.</p><p>Bionomics. Quedius unicolor seems to be very habitat selective and is only found in alpine sites in sphagnum moss and under rocks near bogs, swamps and creeks (Fig. 2G). It is found at altitudes between 900 and 2300 m depending on the latitude of the locality, with higher elevations at more southern sites.</p><p>Distribution. Quedius unicolor is restricted to the montane and subalpine zone of Central and southern Europe (Fig. 21). Specifically, it is known from sites in the Carpathians, the Alps, the Pyrenees, Picos de Europa, the Sudetes, the Harz and the Ore mountain ranges. Due to its very specific habitat requirement, it may be under-collected and may actually be present in mountains further south. Cඈංൿൿൺංඍ (1961) reported it from Sierra Nevada in Southern Spain, but this record may be in fact a misidentified Q. hispanicus . Another doubtful record of Q. unicolor comes from Koçere Stream in the Kazdağları Mountain of the North-western Anatolia by AൻൺർංGංඅ et. al (2009), which is a location remote from the main range of this species.</p></div>	https://treatment.plazi.org/id/038987A0FFD24B30B0EFA5381011FEEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFD34B33B3BBA198114BFCEA.text	038987A0FFD34B33B3BBA198114BFCEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius vicinus (Menetries 1832)	<div><p>Quedius vicinus (Ménétriés, 1832)</p><p>(Figs 1, 4, 8A, 13A, 20)</p><p>Staphylinus vicinus Ménétriés, 1832: 144 [Type locality: Lenkoran]</p><p>Quedius libanicus Coiffait, 1954: 160 [Type locality: Liban: grotte de Birket Aanjar]</p><p>References. Fൺඅൽൾඋආൺඇඇ (1835): 129 (characters); HඈർHHඎඍH (1862): 44 (distribution); Cඈංൿൿൺංඍ (1955): 427 (biology); (1961): 55, (1978): 195 (characters); JൺඋඋංGൾ (1971): 497 (distribution); KඈඋGൾ (1971): 11 (characters); BඈHගඹ (1988): 554 (characters); Gඎඌൺඋඈඏ (1993): 73 (lectotype designation, synonymic notes); AඌඌංඇG &amp; Wඎඇൽൾඋඅൾ (2001): 37 (distribution); Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ (2018a): 131 (distribution); (2019): 51 (characters and distribution).</p><p>Type material examined. Quedius vicinus: Lൾർඍඈඍඒඉൾ ♀ [torso without head and thorax] (ZIN), designated by Gඎඌൺඋඈඏ (1993) [not examined], labelled: “[golden square] / Lenkoran. / vicinus Menet. Lenkor. ” (Gඎ-ඌൺඋඈඏ 1993).</p><p>Additional material examined. AFGHANISTAN: Obéh, [34.35, 63.23], 1680 m (1 J NMW). CYPRUS: Episcope, Limasol, [34.67, 32.88], 12.IV.1995, leg. Schmid (3 JJ 5 ♀♀ NMW); Larnaca, [34.89, 33.63], salt lake, 10 m, 20.IV.1995, leg.W.Suppantschitsch (1J NMW). GEORGIA: Kumisi nr. Tbilisi, [41.61, 44.78], 20.VI.1988, leg. Wrase (1 J cSch). IRAN: Darab to Estahban Rd., 20km NW Darab, 28.8525, 54.4093, 1300m, 16.IV.2006, leg. Serri &amp; Frisch (1 ♀ SDEI); Sonqor, Kermanshah, [34.78, 47.59], 1800 m, 7.VII.1969, leg. Heinz (1 J ZMHB); SE Sepidan, W Dalkhan, 30.2911, 52.0951, 2100m, 9.V.2007, leg. Frisch &amp; Serii (2 JJ SDEI). ISRAEL: Hazerim, [31.24, 34.71], 8.X.1988, leg. E. Orbach (1 J NHMD); Kfar Ha Horesh, n. Nazareth, [32.70, 35.27], 500 m, 8.IV.1997, leg.Heinz (1♀ ZMHB) Lower Galilee, ca. 4 km W Tamra, 32.8633, 35.1715, loamy field edge, 25m 25.IV.2006 leg. D.W.Wrase (1J cSch); Wadi Musrara [Ayalon River], Petah Tiqwa, [32.09, 34.80], 5.XI.1932 (3JJ NMW). JORDAN: Wádi, Hisbán, NW Ma’ dabá, 31.8286, 35.7945, 681 m, brook, slopes and field edges, under stones, 1.IV.2016, leg. Wrase &amp; Laser (1 J cSch). LEBANON: Beirut, [33.88, 35.50], leg. Kindern (1 ♀ SDEI). SYRIA: Zabadani, [33.72, 36.09], 15.IV.1978, leg. Heinz (1 ♀ ZMHB); Zainie, 45 km NE of Latakia, [35.78, 36.19], 4.V.1982, leg. M. Dvořák (1 ♀ MCZ). TURKEY: Antalya, Aksu, [36.88, 30.92], 13.IV.1977, leg. Korell (1 J 1 ♀ ZMHB), Antalya, Alara Han, [36.69, 31.72], 300m, 1997, leg. H. Winkelmann (1 ♀ cSch); Antalya, 26 km W Alanya, nr. Incecum, [36.63, 31.74], 16.IV.- 2.V.1954 leg. V. Brachat (1 J cSch); Catalan, Adana, [37.20, 35.29], 18.IV.1985, leg. Barries (1 ♀ NMW); Karaman Stream, Antalya, [36.86, 30.61], IV.1962, leg. Schweiger (1 J ZMHB); Mardin, [37.29, 40.72], 1300 m, 10.-15.V.1969, leg. Schubert (1 J NMW); Muratpaşa, 20 km E Kırıkhan, [36.47, 36.45], 20 m, 16.IV.1989, leg.Heinz (1J 1♀ ZMHB); E Pamucak, Selcuk, [37.96, 27.27], swamp, 28.III.1986, leg. Heinz (1J 1 ♀ ZMHB). TURKMENISTAN: Kopet Dag, 6 km NE Tschuli n. Firjuza, [37.98, 58.02], 18.IX.1976, leg. Hieke (5 JJ ZMHB).</p><p>Redescription. Measurements JJ (n = 5): HW = 1.64– 1.80 (1.72); HL = 1.40–1.53 (1.48); HL/HW 0.83–0.87 (0.86); PW = 2.16–2.44 (2.30); PL = 2.02–2.20 (2.11); PL/PW 0.89–0.94 (0.92); EW = 2.20–2.47 (2.33); EL = 2.07–2.24 (2.16); EL/EW 0.91–0.94 (0.93); EL/PL 0.99– 1.04 (1.02); PW/HW 1.52–1.60 (1.56); forebody length 5.49–5.96 (5.74). ♀♀ (n = 5): HW = 1.69–1.73 (1.72); HL = 1.38–1.49 (1.45); HL/HW 0.79–0.87 (0.84); PW = 2.22–2.42 (2.32); PL = 2.04–2.22 (2.12); PL/PW 0.91–0.92 (0.92); EW = 2.38–2.47 (2.44); EL = 2.18–2.36 (2.26); EL/EW 0.92–0.95 (0.93); EL/PL 1.01–1.11 (1.07); PW/ HW 1.52–1.66 (1.60); forebody length 5.73–6.07 (5.84).</p><p>Medium sized to large, robust species; body light to dark brown (Fig. 8A).</p><p>Head dark, distinctly transverse, with eyes rather small (EyL/TL = 1.45–1.82 (1.60)); microsculpture of transverse waves; no interocular punctures between anterior frontal punctures (cf. Fig. 6F); antennae and palpi pale yellowish- -brown with base of antennomeres 1–3 clearly pale, all antennomeres clearly elongate.</p><p>Thorax: pronotum slightly wider than long, wider than head, with microsculpture of transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond first puncture of dorsal row; scutellum punctured and pubescent; elytra orange to yellowish-brown, uniformly pubescent, slightly wider than long, roughly of same length as pronotum; legs pale yellowish-brown with inner face of tibia and femur darkened.</p><p>Abdomen: tergites uniformly punctured, with slight iridescence.</p><p>Male. Aedeagus (Fig. 13A): paramere rather slender, with a slight apical expansion and extending into a blunt hook oriented away from median lobe, reaching just beyond apex of median lobe, with sensory peg setae forming two short broad rows fusing together towards apex; median lobe broadly constricting to a point at apex, on parameral side folded into two teeth almost meeting at middle, positioned at level near basal end of peg setae rows of paramere; internal sac without C-sclerite.</p><p>Differential diagnosis. Quedius vicinus stands out as a rather pale member of the Q. molochinus -group and is thus clearly recognizable from Q. subunicolor, Q. unicolor, Q. sundukovi and Q. balticus, in particular by having pale antennomeres 1–3. It is very similar to Q. molochinus and Q. meridiocarpathicus, but it is often paler than these, with elytra and appendages yellowish red compared to brownish red, and abdomen brown compared to dark brown. For confident identification genitalia should be checked as some individuals of Q. molochinus and Q. meridiocarpathicus can have similar external appearance. The aedeagus of Q. vicinus is distinguished from Q. molochinus and Q. meridiocarpathicus by the absence of a C-sclerite in the internal sac, and by the paramere with blunt apical hook oriented away from the median lobe (best seen in lateral view). Quedius vicinus is recognized from some similarly looking sympatric members of the subgenera Microsaurus and Raphirus, such as Q. suramensis – with which it has long been confused (see comments) – by the entire labrum and clearly parallel-sided abdomen.</p><p>Synonymic notes. Gඎඌൺඋඈඏ (1993) designated a lectotype for Q. vicinus and clarified the identity of the species.Also he examined the female holotype of Q. libanicus Coiffait, 1954 described from ‘Lenkoran’ in Azerbaijan (see BඈHගඹ (1988) for details about this locality) and synonymized it with Q. vicinus . Our data fully corroborate the synonymy of Q. vicinus and Q. libanicus .</p><p>The identity of Q. vicinus was confused for a long time since HඈർHHඎඍH (1849) misidentified Q. vicinus as Q. (Raphirus) suramensis Eppelsheim, 1880 . This led to confusion of the two species from different subgenera of Quedius, clarified by Gඎඌൺඋඈඏ (1993). Both species still remain confused in many collections although they are easily distinguished (see Differential diagnosis).</p><p>Bionomics. Quedius vicinus is mainly found near bodies of water like streams, rivers, lakes, etc., where it is collected in litter and under rocks. In an arid region, this maybe the only suitable microhabitats. It has also been found in caves (Cඈංൿൿൺංඍ 1955, as Q. libanicus), although this is most likely accidental, as nothing suggests that it is connected to these.</p><p>Distribution. Quedius vicinus is found from central southern Anatolia across eastern Transcaucasia and most of the northern part of the Middle East into southern Central Asia (Fig. 20). It extends into the Levant region including the island of Cyprus, but does not go into the Nile delta. It is unclear to what extent it occurs in Transcaucasia, where it may only penetrate into the eastern valleys along the west Caspian Sea coast. The species continues into Iran and Afghanistan, as well as to Middle Asia where it is only known from one locality in Kazakhstan (Sൺඅඇංඍඌĸൺ &amp; Sඈඅඈൽඈඏඇංĸඈඏ 2018a).</p></div>	https://treatment.plazi.org/id/038987A0FFD34B33B3BBA198114BFCEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFD04B33B3FCA3A9118CFB4C.text	038987A0FFD04B33B3FCA3A9118CFB4C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius pallipes Lucas 1846	<div><p>Quedius pallipes -group</p><p>Diagnosis. Medium sized species with mostly brown, light brown to reddish elytra and appendages, rarely with darkened elytra. Eyes small, temples large, distance between temporal puncture and posterior margin of eye equals the diameter of the puncture; frons without additional setiferous punctures (Fig. 6F). Scutellum punctate and pubescent. Pronotum with punctures in posterior part of sublateral row. Palearctic origin.</p></div>	https://treatment.plazi.org/id/038987A0FFD04B33B3FCA3A9118CFB4C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFD04B35B388A4FE1015F9EA.text	038987A0FFD04B35B388A4FE1015F9EA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius hispanicus Bernhauer 1898	<div><p>Quedius hispanicus Bernhauer, 1898</p><p>(Figs 1, 2D, 3, 9A,B, 14C, 19)</p><p>Quedius hispanicus Bernhauer, 1898: 341 [Type locality: Cartagena und Cadix in Spanien] stat. rev.</p><p>Quedius cobosi Coiffait, 1964: 293 [Type locality: Espagne: Province de Almeria, Seron], syn. nov.</p><p>Quedius hammianus Sharp, 1911:57 [Type locality:Anglia:Deal, Strood, Lymington, and Hayling Island], syn. rev.</p><p>Quedius rufulus Blümml, 1898: 216 [Type locality: Spanien, Granada], syn. rev.</p><p>Quedius secundus Last, 1952 [Type locality:Studland, Dorset; Mudeford, Hampshire; Chichester], syn. rev.</p><p>References. Fൺඎඏൾඅ (1865): 16 (synonym of Q. molochinus); Kඋൺൺඍඓ (1867): 414 (synonym of Q. molochinus); GൺඇGඅൻൺඎൾඋ (1895): 403 (synonym of Q. molochinus); BൾඋඇHൺඎൾඋ (1900): 46 (distribution); Pඈඋඍൺ (1907): 129 (synonym of Q. molochinus); Gඋංൽൾඅඅං (1924): 81 (subspecies of Q. molochinus; characters); SർHൾൾඋඉൾඅඍඓ (1933): 1457 (synonym of Q. picipennis); MඣඊඎංGඇඈඇ (1937): 3 (as Q. molochinus pallipes); KඈඋGൾ (1960b):69 (synonym of Q. pallipes); Cඈංൿൿൺංඍ (1961): 56 (characters and distribution); (1978):196 (characters); Hൺඇඌൾඇ (1962): 319 (characters and distribution); Sආൾඍൺඇൺ (1962a): 134 (characters); (1963): 39; (1970): 61 (distribution); Lൺඌඍ (1963): 45 (synonym of Q. simplicifrons); LඈHඌൾ (1964): 211 (characters and distribution); Hඈඋං- ඈඇ (1965): 276 (distribution); Bඈඋൽඈඇං (1976a): 100 (synonym of Q. pallipes); Pඈඉൾ (1977): 31 (distribution); LඎർHඍ (1987): 109 (distribution); LඈHඌൾ &amp; LඎർHඍ (1989): 172 (distribution); Iඌඋൺൾඅඌඈඇ (1990): 3 (distribution); BඈඋGൾඌ (1990): Table II (distribution); WൾඅർH (1993): 229 (ovariole number and ovary structure); Oඎඍൾඋൾඅඈ et al. (1995): 80 (distribution; as Q. pallipes).</p><p>Type material examined. Quedius cobosi: Pൺඋൺඍඒඉൾ: J (MNHN), labelled: “ PARATYPE [red label] / Quedius cobosi Coiff. H. COIFFAIT det. 1964 / Laujar Almeria - España A. Cobos coll.”</p><p>Additional material examined. DENMARK: Lakolk, [55.14, 8.49], debris on beach, 1.XI.2013, leg. J. Pedersen &amp; A. Brunke (1 J NHMD); Skallingen, [55.50, 8.25], 17.X.1959, leg. F. Bangsholt (2 JJ NHMD). GERMANY: Archsum, [Sylt], [54.87, 8.39], IX.1901 (1 SDEI); Insel Borkum, [53.58, 6.66], 12.VIII.1937, leg. Struse (2 ♀♀ 4 JJ NMW); N Dagebüll, Nordfriesland, 54.7361, 8.7094, salt pasture among stones and clay on dam, 25.VIII.2017, leg. Wrase &amp; Laser (1 J cSch); Memmert, Juist, [53.64, 6.88], 13.VIII.1917, leg. Alfrum (1 SDEI). FRANCE: Bordeuax, St. Louis, [44.94, -0.52], 22.II.1964 leg. C. Jeanne (1 JcBor); Camargue, Etang du Fangassier, [43.42, 4.63], 20.V.1994 salt marsh, leg. Schülke &amp; Grünberg (1 J cSch); Sainte Marie la Mer, [42.72, 3.03], 28.IV.1955, leg. J.T. Skovgaard (1 ♀ NHMD). PORTUGAL: Lissabon, [38.76, -8.87], leg. Von Heyden (1 J 2 ♀♀ SDEI); Lusitania, Lissabon, [38.76, -8.87], IV.-V.1910, leg.A. Schartzmayr (1 J 1♀ SDEI). Aඓඈඋൾඌ: Faial.Baia da Areia das Fontes 1km N of Praia do Norte, [38.61, -28.74], under stones, grassy ground, near rocky shore, 2.IV.1957, leg.Brinck and Dahl (5 ♀♀ MZLU); Faial, Caldeira, [38.58, -28.71], 10.IV.1957. leg. Brinck and Dahl (3♀♀ MZLU); Faial, Horta, Porto Pim., [38.52, -28.62], sandy ground, 7.IV.1957 leg.Brinck and Dahl (1 ♀ MZLU); Faial. Nasce Agua 3 km S of Cedros, [38.60, -28.68], ravine, under stones, 5.IV.1957, leg. Brinck and Dahl (1 J MZLU); Faial, Praia do Almoxarife, [38.55, -28.61], sandy grassy ground, under stones, 31.III.1957. leg. Brinck and Dahl (1 ♀ MZLU); São Miguel,Atalhada, [37.74, -25.58], natural forest, 4.IX.1999, leg. H. Enghoff (2 ♀♀ NHMD). Mൺൽൾංඋൺ: Campanario, Fontes, [32.69, -17.01], 1200-1400m, 16.IX.2014, leg. Kleeberg (1 J cSch); Funchal, [32.67, -16.91], 0-500 m, II.1966, leg. Palm (2 JJ 3♀♀ MZLU); Pico do Castelo, [33.08, -16.33], 17.19. V.1977, leg.Palm (2 JJ MZLU); Terreiro da Luta, [32.68, -16.91], 850 m, stream, 20.IV.1957, leg. Brinck and Dahl (1 J MZLU). SPAIN: Alicante, Orihuela, [38.08, -0.94], 20.IV.1997, leg. Lauffer (2 MNCN); Altea, Rio Algar, Alicante, [38.61, -0.05], 27.II.1999, leg. Wrase (1 J cSch); Aranjuez, Madrid, [40.02, -3.59], 520 m 3.-4.V.1988, leg. Feller (3 ♀♀ cSch); Arroyo de Jurnadiel, S Brozas, Cáceres, [39.56, -6.81], 325 m, 20.VI.1991, leg. Wrase (1 J cSch); Ávila, Avila, [40.65, -4.68], leg. Sanz (3 MNCN); Badajoz: Aljucen, [39.04, -6.33], leg. Pacheco (1 MNCN); Boadilla del Monte, NW Madrid, [40.41, -3.88], 6.VII.1996, leg. D.Wrase (1J cSch); Cabo de Gata, SE Almeria, 36.7908, -2.2513, coastal area, 5.IV.2012, leg. T. Wolsch (1 ♀ cSch); Caboalles [de Abajo], [42.95, -6.37], leg. Paganetti (1 ♀ SDEI); Cádiz, [36.53, -6.22], 16.I.1894, leg. Navarro (2 MNCN); Cádiz, Algeciras, [36.16, -5.43], leg. Arias (1 MNCN); Cádiz, Vega del Barbate, Vejer de la Frontera, [36.25, -5.95], leg. De Quirós (1 MNCN); Cornudilla, [42.68, -3.42], 13.IX.1962, leg. J.T. Skovgaard (1 J 1 ♀ NHMD); Cuenca, Uclés, [39.98, -2.86] (2 MNCN); Guipúzcoa, Arechavaleta, [43.03, -2.51], leg.Antoniol (1 MNCN); Guipuzcoa, Irun, Jaitzubia, [43.34, -1.82], 10.V.2006,leg.Anichtchenko (1J 1♀ NHMD); Huelva, Cala, [37.97, -6.31], leg. C. Bolivar (1 MNCN); Jáen Mengibar, Rio Guadalquivir, [37.98, -3.79], 330m, 16.VI.1991, leg. Wrase (1 J cSch); La Coruña, Coruña, [43.36, -8.4], leg. Ricol (2 MNCN);La Coruña, Villa Rutis, [43.28, -8.38], VII.1908, leg. Bolivar (2 MNCN); Laguna de Ruiz Sánchez, N Osuna, Sevilla, [37.41, -5.09], 26.V.1991, leg.Wrase (1 J cSch); Los Guajares, Andalusien, 36.8174, -3.5546, quarry, 6.IV.2012, leg. T. Wolsch (1 J cSch); Madrid, Escorial [40.57, -4.11] (1 MNCN); Madrid, Torrelaguna. [40.82, -3.53], leg. F. Beltran (1 MNCN); Madrid, Viñuelas [40.79, -3.34] (1 MNCN); Maranchon, Guadalajara, [41.04, -2.20], 24.XI.2005, leg.Anichchenko (1 J NHMD); Murcia, Cartagena, [37.62, -0.99], leg. S. Gomez (1 MNCN); Palencia, [42.01, -4.52], leg. Paganetti (1 J SDEI); Palmones, Cádiz, [36.17, -5.43], 29.V.1991, leg. Wrase (1 J cSch); Quero, Toledo, [39.51, -3.24], 8.III.1999, saline, leg. D.W. Wrase (1 J cSch); Serrania de Ronda, Málaga, [36.74, -5.17], 11.VI.1991, leg. Wrase (1 J cSch); Sitges, [41.23, 1.84], 22.X.1922 leg. W.Liebmann (1 ♀ SDEI); Toledo, Nahalhermosa, [39.63, -4.47], 27.VII.2007 (1 J NHMD); Tres Cantos, [40.60, -3.70], 29.-30.II.2000, leg. Anichtchenko (1 ♀ NHMD). Mൺඅඅඈඋർൺ: Palma, [39.56, 2.65], IV.1957, leg. T. Palm (1 ♀ MZLU); Rabassa, [39.54, 2.69], leg. Briet (3 JJ 1♀ NMW); Valldecousa, [39.71, 2.62], 15.-20.IV.1957, leg.T. Palm (1 J 3 ♀♀ MZLU). Cൺඇൺඋඒ Iඌඅൺඇൽඌ: Gඋൺඇ Cൺඇൺඋංൺ: Las Lagunetas, [28.01, -15.57], 22.XI.1988, leg. G. Gillerfors (1 ♀ MZLU). Tൾඇൾඋංൿൾ: Anaga Mountains, nr.Cruz del Carmen, [28.53, -16.27], 14.VI.1992, leg. Hieke &amp; Wendt (1 ♀ ZMHB); Cumbre Dorsal Esperanza-Forest, [28.38, -16.44], 1250m, 13.IV.1992, leg. Zerche (1 ♀ SDEI); Erjos env., [28.32, -16.81], 1100 m, 7.XII.2004, leg. Skoupý (1 ♀ cSch); Genovés, [28.36, -16.74], 400m, 9.VI.1989, leg. Felies (1 J cSch); La Esperanza, [28.44, -16.37], 1200m, 18.VI.1992, leg. Hieke &amp; Wendt (2 JJ ZMHB); Orotava, valley Aguamansa, [28.36, -16.49], 1115m, 25.XII.1995, leg. Stüben &amp; Bahr (1 ♀ SDEI); Nr. Orotava, [28.35, -16.51], 1400m, 19.VI.1992, leg. Hieke &amp; Wendt (2 JJ 1 ♀ ZMHB); Puerto de la Cruz env., [28.36, -16.52], 300m, 20.II.-5.III.2004, leg. E. Hajdaj (1 J cSch); Teno Alto, San Jeronimo, [28.34, -16.87], 800-900 m, 20.I.1999, leg. G. Zappi (2 JJ 1 ♀ cBor). UNITED KINGDOM: EඇGඅൺඇൽ: Chichester Harbour, [50.79, -0.96], 6.V.1947, leg. H. Last (2 JJ NMW).</p><p>Redescription. Measurements JJ (n = 6): HW = 1.62– 1.76 (1.69); HL = 1.47–1.49 (1.48); HL/HW 0.86–0.92 (0.88); PW = 2.22–2.29 (2.25); PL = 2.07–2.22 (2.16); PL/PW 0.93–0.97 (0.96); EW = 2.27–2.49 (2.37); EL = 2.20–2.33 (2.26); EL/EW 0.92–1.00 (0.96); EL/PL 1.01– 1.09 (1.05); PW/HW 1.49–1.56 (1.52); forebody length 5.76–6.02 (5.90). ♀♀ (n = 4): HW = 1.58–1.73 (1.66); HL = 1.38–1.53 (1.46); HL/HW 0.87–0.89 (0.88); PW = 1.96–2.27 (2.16); PL = 1.91–2.11 (2.05); PL/PW 0.93–0.98 (0.95); EW = 2.22–2.51 (2.42); EL = 2.11–2.42 (2.29); EL/EW 0.91–0.96 (0.95); EL/PL 1.08–1.15 (1.12); PW/ HW 1.42–1.52 (1.48); forebody length 5.40–6.07 (5.80).</p><p>Medium to large sized species; body black to brown (Figs 9A,B).</p><p>Head black, distinctly transverse, eyes medium size (EyL/TL: 1.88–2.06 (1.94)), microsculpture of transverse waves, no interocular punctures between anterior frontal punctures; antennae pale.</p><p>Thorax: pronotum dark brown to brown, slightly wider than long, clearly wider than head, with microsculpture of transverse waves, three punctures in dorsal row, sublateral row of pronotum with a cluster of one to three punctures situated slightly anterior of the anterior most puncture of the dorsal row; scutellum with fine setiferous punctures and with wavy to meshed microsculpture; elytra usually light brown to brown, occasionally black with paler suture and posterior margin, rarely completely black, relatively long, uniformly pubescent, quadrate or slightly wider than long, slightly longer than pronotum; legs pale, except inner face of tibia darkened.</p><p>Abdomen black to dark brown and slightly iridescent, tergites uniformly punctured, with slight iridescence.</p><p>Male. Aedeagus (Fig. 14C): paramere elongate, narrow, lanceolate with narrowly slightly asymmetrical apical portion exceeding apex of median lobe, with numerous sensory peg setae forming two lateral longitudinal groups connected subapically; median lobe with sides of apex folded into distinct tooth situated at proximal part of fold; internal sac with a pair of very long medial sclerites, and with a pair of globular sclerites.</p><p>Differential diagnosis. Quedius hispanicus can be distinguished from all species of the Quedius s. str. except Q. pallipes, by a cluster of 1–3 additional setiferous punctures in the posterior part of the sublateral row of the pronotum. They are very rarely missing on both sides of the pronotum in Q. hispanicus . It can be distinguished from Q. pallipes only by male genitalia characters: by a slightly narrower and, in medial section, more attenuate paramere; and by a median lobe with sides of apex each folded into distinct tooth, tooth situated at base of the fold.</p><p>Synonymic notes on Q. pallipes, Q. hispanicus, Q. simplicifrons, Q. cobosi, and Q. levasseuri . Lඎർൺඌ (1849) described Quedius pallipes from the coastal town of Philippeville [now Skikda], Algeria. Fൺංඋආൺංඋൾ (1862) described Quedius simplicifrons as a very similar species from the French island of Corse. BൾඋඇHൺඎൾඋ (1898) described Quedius hispanicus, a species very similar to these, from the Spanish coastal towns of Cartagena and Cadiz. These three species have been mixed up, probably due to their very similar habitus, rendering their individual distributions unclear. Cඈංൿൿൺංඍ (1961) stated that Q. pallipes was restricted to North Africa and hypothesized that Q. simplicifrons – with Q. hispanicus as a junior synonym – could be a European subspecies of Q. pallipes without formally downgrading Q. simplicifrons to subspecific rank. Subsequently, some authors (Hൺඇඌൾඇ 1962, LඈHඌൾ 1964, Hඈඋංඈඇ 1965, Sආൾඍൺඇൺ 1970, Bඈඋ-ൽඈඇං 1976, WൾඅർH 1993) have treated Q. hispanicus as a valid species for Western European specimens, while others continued to consider it a synonym of Q. simplicifrons (Pඈඉൾ 1977, Cඈංൿൿൺංඍ 1978, Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං 1995, Sඈඅඈൽඈඏඇංĸඈඏ 2012). KඈඋGൾ (1962b) mentioned specimens from Southern Italy and North Africa as conspecific with Q. pallipes . He also reported that these were collected at higher altitudes (up to just above 1000 m) very far from the coast. Cඈංൿൿൺංඍ (1964) further described two new species based mainly on genitalia differences of the males, Quedius cobosi Coiffait, 1964 from Seron in Spain, and Quedius levasseuri Coiffait, 1964 from Corse, both similar to Q. pallipes and Q. simplicifrons / Q. hispanicus . Quedius cobosi was characterised by the presence of numerous small setae on the edge of the paramere, while Q. levasseuri was described as having paler sutural and elytral posterior margins as well as a differently shaped aedeagus, compared to Q. pallipes and Q. simplicifrons . This history implies several species- -group taxa of uncertain status, distribution and habitat preference that are similar to Q. pallipes and co-occur in the West Mediterranean. Studying a broad sample of specimens from that area, we found only two morphotypes, which differ only in the structure of the aedeagus. The first is found along the European Atlantic Coast, including Southern UK, Canaries, Madeira, Azores and inland in southern France and on the Iberian Peninsula (Figs 14C, 19). These areas include the type localities of Q. hispanicus and Q. cobosi (Fig. 19). The second is found in mainland Italy, on Corse and Sardinia, and in North Africa at both coastal and higher elevation sites (Figs 14D, 19). These areas include the type localities for Q. pallipes, Q. simplicifrons, and Q. levasseuri . There is no transition between the aedeagi of both types. The steep cliff shoreline between Nice, France and La Spezia, Italy may be unsuitable for these species and may represent a barrier for their dispersal. As the morphological difference between both types of genitalia is clear, and in agreement with the type localities, we here propose to use Q. hispanicus for the Iberian and Atlantic coast species (type 1) and Q. pallipes for the Italian and North African species (type 2) (Figs 14, 19). Quedius cobosi syn. nov. is synonymized with Q. hispanicus .</p><p>However, barcodes from specimens across the entire distribution of both types 1 and 2 show very little variation, with no clear clustering corresponding to both genitalia types (Fig. 3). A rather uniform genetic barcode among specimens morphologically corresponding to both Q. hispanicus and Q. pallipes (only 0.5% divergence, Table 2) suggests that there may be some level of mitochondrial introgression between these species. External factors such as past or present infections by endosymbionts such as Wolbachia (e.g. KඈඇൽൺඇൽൺඋൺආൺංൺH et al. 2013) cannot be ruled out as an explanation either. This should be explored more thoroughly for example with the use of nuclear genes or genomics.</p><p>The original descriptions and distribution records of Quedius hammianus Sharp, 1911 and Quedius secundus Last, 1952, both described from the UK, match the morphology and the ecological association with coastal sites of Q. hispanicus . Quedius rufulus Blümml, 1898, described from Granada in Spain as having darker base of the antennae, most likely represents a variant of Q. hispanicus . All three species were hitherto in synonymy with Q. simplicifrons . Under the revised concept of the Q. pallipes -group, we synonymize them with Q. hispanicus .</p><p>Bionomics. Quedius hispanicus mainly occurs in saline habitats along the Atlantic coast, such as salt marshes and the banks of estuaries in flood debris. Further south in the Iberian Peninsula and southern France the species seems to be present inland in non-saline habitats. Inland it occurs in the debris along the banks of creeks and rivers. For example, AKH has collected it in the debris near streams and rivers in the Extremadura region (Fig. 2D) and also in the flood debris on a salt marsh near the coast in southern Denmark. Most records are from low elevations, except a few on Madeira and the Canary Islands, where Q. hispanicus is found up to 1400 m.</p><p>Distribution. Quedius hispanicus occurs along theAtlantic coast from southern Denmark to the Iberian Peninsula where it continues to the French Mediterranean coast, with one unverified report from the northwestern Italian coast (Fig. 19). In the Iberian Peninsula it occurs inland along most rivers and creeks. It also occurs on the Azores (Faial), Madeira (Madeira and Porto Santo), the Canaries (Tenerife and Gran Canaries) and Mallorca.</p></div>	https://treatment.plazi.org/id/038987A0FFD04B35B388A4FE1015F9EA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFD64B37B3DCA6981720F9EA.text	038987A0FFD64B37B3DCA6981720F9EA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius pallipes Lucas 1849	<div><p>Quedius pallipes Lucas, 1849</p><p>(Figs 1, 3, 9C, 14D, 19)</p><p>Quedius pallipes Lucas, 1849: 113 [Type locality: Algeria: environs de Philippeville]</p><p>Quedius leonhardi Bernhauer, 1914:67 [Type locality:Gargano], syn. rev.</p><p>Quedius levasseuri Coiffait, 1964: 293 [Type locality: Corse, Porto-Vecchio], syn. nov.</p><p>Quedius sardous Gridelli, 1924: 81 [Type locality: Sardegna: Cagliari, Assemini, Decimo, Golfo Aranci, Gonnesa], syn. rev.</p><p>Quedius simplicifrons Fairmaire, 1862: 580 [Type locality: Corse], syn. nov.</p><p>References. Fൺඎඏൾඅ (1869a):487, (1869b):151, (1902):115 (distribution; synonym of Q. molochinus); Pඈඋඍൺ (1907): 130 (distribution; synonym of Q. molochinus); MඣඊඎංGඇඈඇ (1937): 3 (distribution; subspecies of Q. molochinus); Pൺඎඅංൺඇ (1941): 268 (larval characters; variety of Q. molochinus); Lൺඌඍ (1952) (characters and distribution); KඈඋGൾ (1960): 53 (characters); (1962b):335, (1973): 275 (distribution); Cඈංൿൿൺංඍ (1961): 55 (characters); (1978):196 (distribution); Pൺඅආ (1963):142 (characters); SർHൾൾඋඉൾඅඍඓ (1965): 407; (1965b): 511 (distribution); Bඈඋൽඈඇං (1974): 12; (1976): 100 (characters and distribution); 1976b: 102 (characters); Cංർൾඋඈඇං &amp; Zൺඇൾඍඍං (1995): 32 (distribution).</p><p>Type material examined. Quedius leonhardi: Hඈඅඈඍඒඉൾ J (FMNH), labelled: “Monte Gargano 6.25.5.1907 legit. M. Hill Coll. O. Leonhard / Leonhardi Brh. Typus unique / Chicago NHMus M. Bernhauer Collection / In euparal / D.Drugmand det., 1994 Quedius (S.str.) molochinus (Grav.) / FMNHINS 3982385 FIELD MUSEUM Pinned / PHOTOGRAPHED S. Ware 2020 Emu catalog”.</p><p>Additional material examined. ALGERIA: Alger [Algiers], [36.71, 3.07], 2.-10.V.1972, leg. M. Dvořák (1 J MCZ); Bona [Annaba], [36.89, 7.76], leg. Meinert (1 ♀ NHMD); Mandoura, Boumerdes, [36.83, 3.66], 17.VI.1988, leg. Sama (1 J cSch). CHAD: Tibesti, Distr. Emi, Miski, [20.18, 17.94], 29.X.1885 (1 J NMW). FRANCE: Cඈඋඌൾ: Ajaccio, [41.93, 8.73] (1 J NHMD);Aleria, [42.11, 9.49], 1905, leg.O. Leonhard (1 J 1 ♀ SDEI). ITALY: Aspromonte, Samo, [38.07, 16.05], 250m, 12.III.2000, leg. F. Angelini (1 ♀ NHMD); Barletta, [41.32, 16.27], 14.V.1962, leg. J.T. Skovgaard (4 JJ 3 ♀♀ NHMD); Calaria, Locri, [38.25, 16.28], 3.V.1975, leg. R. Mourgua (1 ♀ cSch); Canale, [44.79, 7.99], leg. Paganetti (2 ♀♀ SDEI); Gargano, [41.71, 15.72], leg. L.S. Giovanni (1 ♀ NHMD); Lago S. Floriano, Toscana, [42.42, 11.35], 30.V.2002, leg. Bordoni (1 J 1 ♀ cBor); Lido di Roma, [41.73, 12.28], 17.-18.VI.1939 Palm (1 ♀ MZLU); Monti Aurunci, Itri, [41.31, 13.49], 25.IX.1973, Leg. M. Zampetti (1 ♀ cBor); Portonaccio, Lazio, [42.02, 12.39], 9.IX.1932, leg. G. Sacca (6 ♀♀ cBor); Puglia, Riserva Naturale Le Cesine, [40.35, 18.33], 11.VI.1999 leg. F. Angelini (2 ♀♀ NHMD); Roseto Capo Spulico, [39.98, 16.61], 10.XII.2000, leg. F. Angelini (1 ♀ NHMD). Sൺඋൽංඇංൺ: Assuni, [39.87, 8.94], leg. Krausse (1 ♀ SDEI); Cagliari, [39.21, 9.11], 29.I.1967, leg. S. Riese (3 ♀♀ cBor); Lago Omodeo, Oristano, [40.13, 8.92], 2.V.1979, leg.S. Riese (1J cBor); Oristano, [39.88, 8.58], 21.IV-7.V.1978, leg.S. Riese (1J 1♀ cBor); Ottana, [40.23, 9.04], 14.V.1976 (2 ♀♀ cBor); Sant’Antioco, [39.06, 8.45], IV.1984, leg. Bordoni (1 ♀ cBor); Sorgono, [40.02, 9.11], 1912 leg. Krausse (1 J SDEI); Stagni Cobras, Oristano, [39.91, 8.49], 19.IV.1971, leg. Visua (1 ♀ cBor). Sංർංඅඒ: Catania, [37.51, 15.09], IV.1965, leg. Fello (1 JcBor); Enna, [37.56, 14.28], 1.V.1990 leg. Roland (1 J NHMD); Gibilmanna, [37.98, 14.01], 800m 23.IV.1981, leg. T. Palm (1 ♀ MZLU); Marausa, [37.95, 12.5], 8.III.1982, leg.Alique (1 JcBor); Stracusa, [37.06, 15.28], 18.VII.1969, leg. Alraub (1 J 4 ♀♀ cBor). MOROCCO: Al Hakkama, ca. 18 km SE Tanger, 35.6377, -5.6841, fields, fallow, pastures, 100m, 26.II.2004, leg. D.W. Wrase (1 ♀ cSch); Bouznika, [33.81, -7.14], 12.III.1974 (1J cBor) Douyiet, [34.04, -5.12], 8.IV.1974 (1J 1♀ cBor); Fouarat, Rabat, [34.25, -6.52], 1932, leg.A.Théry (1 J NMW); 10 km N Mdiq, Tetouan, 35.7182, -5.3427, brackish wetland under plants in soil, 27.II.2004, leg. D.W. Wrase (1 J cSch); Mohammedia, Oued Mellah, Chefchaouen, 33.71, -7.33, stony slopes and bushes, 4.II.2003, leg. D.W. Wrase (1♀ cSch); Rif Mts. El-Malha, 19 km SE Bab-Taza, Chefchaouen, 34.9801, -5.0791, stony fallen land, ca. 440 m, 29.II.2004, leg. D.W.Wrase (1 ♀ cSch); 2 km W Tetouan, [35.55, -5.45], 16.II.1999, leg. Wrase (1 ♀ cSch). TUNISIA: El Besbassia, 10 km SW Hammamet,Sousee, [36.35, 10.52], 28.XI.-6.XII.1995, leg. D.W. Wrase (1 J cSch); El Kantaoui, 12 km NW Sousee, [35.89, 10.58], 25.-30.XI.1992, leg. D.W. Wrase (1 J cSch); Gabés, [33.88, 10.12], 30.V.- 5.6.1969, leg Palm (1 J MZLU).</p><p>Redescription. Measurements JJ (n = 4): HW = 1.64– 1.73 (1.69); HL = 1.47–1.51 (1.48); HL/HW 0.86–0.89 (0.88); PW = 2.16–2.31 (2.22); PL = 2.04–2.22 (2.11); PL/PW 0.93–0.96 (0.95); EW = 2.33–2.51 (2.39); EL = 2.16–2.36 (2.23); EL/EW 0.97–1.10 (1.06); EL/PL 0.97– 1.10 (1.06); PW/HW 1.47–1.55 (1.50); forebody length 5.69–6.00 (5.83). ♀♀ (n = 6): HW = 1.62–1.69 (1.67); HL = 1.36–1.47 (1.41); HL/HW 0.83–0.87 (0.84); PW = 2.13–2.27 (2.18); PL = 1.98–2.04 (2.03); PL/PW 0.90–0.96 (0.93); EW = 2.33–2.47 (2.41); EL = 2.13–2.29 (2.22); EL/EW 0.90–0.94 (0.92); EL/PL 1.07–1.12 (1.10); PW/ HW 1.45–1.62 (1.54); forebody length 5.49–5.73 (5.66).</p><p>Medium to large sized species; body black to light brown (Fig. 9C).</p><p>Head black, distinctly transverse, with eyes of medium size (EyL/TL: 1.76–2.20 (1.97)), microsculpture of transverse waves, no interocular setiferous punctures between anterior frontal punctures; antennae pale.</p><p>Thorax: pronotum paler dark brown to brown, slightly wider than long, clearly wider than head, with microsculpture of transverse waves, three punctures in dorsal row, sublateral row of pronotum with a cluster of one to three punctures situated slightly anterior of the anterior most puncture of the dorsal row. Scutellum with fine setiferous punctures and with wavy to meshed microsculpture; elytra usually light brown to brown, occasionally with paler suture and posterior margin, rarely completely black, relatively long, uniformly pubescent, slightly wider than long, of equal length or slightly longer than pronotum; legs pale, except inner tibia surface which is darkened.</p><p>Abdomen black to dark brown and slightly iridescent, tergites uniformly punctured, with slight iridescence.</p><p>Male. Aedeagus (Fig. 14D): paramere elongate, narrow, lanceolate with narrowly slightly asymmetrical apical portion exceeding apex of median lobe, with numerous sensory peg setae forming two subapical, lateral longitudinal groups connected apically below apex; median lobe with sides of apex folded into distinct teeth situated at medial part of fold. Internal sac with a pair of distal, very long medial sclerites, and with a pair of globular proximal sclerites.</p><p>Differential diagnosis. Quedius pallipes is distinguished from other Quedius s. str., except Q. hispanicus by a cluster of 1–3 additional setiferous punctures in the posterior part of the sublateral row of pronotum, which are very rarely missing. It can only be distinguished from Q. hispanicus by male genitalia characters; paramere slightly wider and less attenuate in middle portion, median lobe with sides of apex each folded into distinct tooth, tooth situated at about midlength of fold.</p><p>Synonymic notes. See under Quedius hispanicus . Based on the type locality, we here propose to use the name Q. pallipes for the Italian and North African species with the aedeagus type 2 (Figs 14, 19). Our examination of hundreds of specimens of the Q. pallipes -group across its full distribution range showed that specimens from Corse – which matched the original description of Q. simplicifrons – were conspecific with Q. pallipes . Therefore, Q. simplicifrons Fairmaire, 1862, syn. nov. described from Corse, is here synonymized with Quedius pallipes Lucas, 1849 .</p><p>Quedius sardous Gridelli, 1924 from Sardegna was originally described as a variety of Q. picipennis [now Q. molochinus]. Following Cඈൿൿൺංඍ (1961), Q. sardous was considered to be an infrasubspecific form of Q. simplicifrons . Here we move Quedius sardous Gridelli, 1924 syn. rev. to synonymy with Quedius pallipes Lucas, 1849 .</p><p>The original description of Quedius leonhardi Bernhauer, 1914, a species described from Gargano in Italy and hitherto considered a synonym of Q. molochinus (e.g., Hൾඋආൺඇ 2001), matches Q. pallipes, except for the missing puncture in the sublateral row of pronotum, usually present in Q. pallipes . However, we have seen specimens of Q. pallipes (earlier named as Q. pallipoides, see below) with this puncture missing, while neither the described coloration of Q. leonhardi nor its distribution in central Italy match what we know about Q. molochinus . Thanks to S. Ware and M. Turcatel of the FMNH we were able to study images of the type of Q. leonhardi (see supplementary Fig. S4), which confirmed that Q. leonhardi fits the concept of Q. pallipes . Therefore, Q. leonhardi Bernhauer, 1914, syn. rev. is here moved from synonymy with Q. molochinus (Gravenhorst, 1806) to synonymy with Q. pallipes Lucas, 1849 .</p><p>Based on the original description of Cඈංൿൿൺංඍ (1964), and additional data by Bඈඋൽඈඇං (1976) and Cඈංൿൿൺංඍ (1978), Q. levasseuri, Coiffait 1964, known only from the type series from Porto Vecchio on the island of Corse, we suggest that the male holotype have a teratological aedeagus. The shape of its aedeagus, not seen in any other specimen from this relatively well collected region, is the only unique trait of Q. levasseuri, because we observed its other presumably unique characters, the pitchy black elytra with narrow reddish margins, in both Q. hispanicus and Q. pallipes . Based on our hypothesis about the teratological aedeagus, we propose to synonymize Q. levasseuri, syn. nov. with Q. pallipes, which is also known from Corse.</p><p>Bionomics. Quedius pallipes is found at elevations from sea level to ca. 1000 m, in wet debris and under rocks, mainly near water sources. For example, based on examined specimen labels, it has been recorded in a brackish wetland under plants in soil, and on rocky slopes with bushes, both in interior Morocco. Based on records near coastal sites the species is most likely halotolerant.</p><p>Distribution. Quedius pallipes is distributed across northern Africa, the Italian Peninsula and the islands of Sicily, Sardinia and Corse (Fig. 19). In northern Africa it is found in Morocco, Algeria and Tunisia, mainly in the coastal regions of these countries, or near water sources inland. A single record from Tibesti in Northern Chad is doubtful, as it is based on an unclear label, which may refer to the small settlement of Miski in the Emi Koussi District. In the Italian Peninsula it is found everywhere except its northern mountainous parts.</p></div>	https://treatment.plazi.org/id/038987A0FFD64B37B3DCA6981720F9EA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFD44B29B136A69814BAFBCA.text	038987A0FFD44B29B136A69814BAFBCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius molochinoides Smetana 1965	<div><p>Quedius molochinoides -group</p><p>Diagnosis. Species of medium to large size with two interocular punctures between anterior frontal punctures (Fig. 6G). Body dark, elytra and appendages variable in coloration, from completely dark to reddish-brown or light brown. Eyes medium sized, temples relatively large, with temporal puncture separated from posterior margin of eye by distance equivalent to diameter of this puncture. Scutellum punctate and pubescent. Restricted to the Nearctic Region.</p><p>Comments. The Nearctic Q. molochinoides -group currently includes five species of which four ( Q. molochinoides, Q. horni and Q. lanei, now also Q. altanai) are impossible to separate morphologically from each other without dissection of male genitalia. Only the northern Q. labradorensis can be confidently delimited within the group based on its larger size and pale basal three antennomeres. Our thorough examination of the entire group showed that Q. horni is most easily recognized based on the aedeagus. This species is now known from the Pacific Northwest, from Oregon to southern Alaska, including Vancouver Island and Haida Gwaii (most of ecoregion 7.1). Most specimens of this species were taken on coastal sites or inland near rivers and creeks to mid elevaton in the Cascades and Mount Olympia, suggesting its close association with wet habitats. Presumably this species is somewhat halotolerant. When specimens of what was known as Q. lanei, were closely examined morphologically, we found two distinct morphotypes distinguished by differences in male genitalia. Species from the eastern foothills of the Cascades, the Sierra Nevada and the Blue Mountains (ecoregions 6.2.8, 6.2.9, 6.2.12) represent a morphotype with a broader paramere and median lobe, the latter being acuminate near apex (Figs 16D, 17). This includes specimens from the Sierra Nevada (Nevada and California), which were earlier assigned to Q. molochinoides (e.g., Sආൾඍൺඇൺ 1971a). Specimens from the central and southern Rockies as well as western Sierra Madre (ecoregions 6.2.10, 6.2.13, 6.2.14, 6.2.15, 13.1.1) represent a different morphotype with a more slender paramere and median lobe, the latter being non-acuminate (Figs 16C, 17). COI sequences of both morphotypes showed that four specimens corresponding to the first morphotype from Lake Tahoe in Nevada, Lassen National Forest in California, Gearhart Mountains in Oregon, and Strawberry Range in Oregon, formed a group, separate from the fifth specimen corresponding to the second morphotype (Fig. 4). The central-southern Rockies morphotype was only represented by a single incomplete sequence (407 bp) from Red Mountain Pass in Colorado. This sequence was over 6% divergent from all Q. lanei sequences, but only 1.2% from Q. molochinoides -group 3 (see below), from which it is morphologically distinct (Table 2). Based on the discovered congruence between the morphological and molecular groups, as well their obvious allopatry (Fig. 21), we suggest that Q. lanei is a species restricted to the eastern foothills of the Cascades, Sierra Nevada and Blue Mountains, whereas a new species, Q. altanai sp. nov., is described for specimens from the central and southern Rockies, as well as western Sierra Madre (teal in Fig. 21). Quedius molochinoides showed genetic subclustering of COI barcodes into three distinct BINs (Table 2). One BIN ( Q. molochionide s-group 1) was restricted to the Canadian national parks of Banff and Waterton (ecoregion 6.2.4), an area delimited by the Rocky Mountains Trench to the West, where it is replaced by another BIN ( Q. molochinoides -group 2), and the Canadian Plains to the east, which lack Q. molochinoides . In the north, this group is replaced by Q. molochinoides -group 3 from Jasper National Park, indicating a divide between groups 1 and 3 around Snow Dome peak and the Arctic Divide. One potential split is the Athabasca River Valley, that may have acted as a post-glacial distribution barrier combined with latitudinal climatic gradients. The extent of the distribution of group 1 southwards is still unknown, but it may very well extend into northern-central Montana and Idaho (Figs 4, 22). The Q. molochionides -group 2 is confined to the Canadian Rockies west of the Rocky Mountains Trench (ecoregion 6.2.3) and towards the North continuing to the Pacific Coast in Alaska, to the Kenai Peninsula northwards (ecoregion 6.1 and northern part of 7.1). The Q. molochinoides -group 3 is present from Becharof Peninsula in the west across the North American Taiga to glacial refugia of New England, Quebec and Newfoundland in the East (ecoregions 4, 5, and partly 6.1 and 7.1). Our results show that apparently these three molecular BINs within Q. molochinoides have distinctly allopatric distributions and thus potentially have limited gene flow among each other. However, since we have been unable to find morphological characters clearly separating them, for now we refrain from describing them as species. If future work shows that these are in fact distinct, then they may be described as separate species. Our results add further information to the complex diversification patters found in the Canadian Rocky Mountains as illuminated through a series of studies on the Spruce Budworm complex ( Lepidoptera: Tortricidae: Choristoneura) (Lඎආඅൾඒ &amp; SඉൾඋඅංඇG 2010, 2011; Bඋඎඇൾඍ et al. 2017, Dඎඉඎංඌ et al. 2017). Similar to these studies, genome-wide sequencing may be needed to confidently resolve the population/species boundaries and distribution patterns in the Q. molochinoides group.</p></div>	https://treatment.plazi.org/id/038987A0FFD44B29B136A69814BAFBCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFCA4B28B0C5A4781772FAEA.text	038987A0FFCA4B28B0C5A4781772FAEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius altanai Hansen & Brunke & Simonsen & Solodovnikov 2022	<div><p>Quedius altanai Hansen &amp; Brunke, sp. nov.</p><p>(Figs 1, 4, 10C, 16C, 17, 21)</p><p>Quedius lanei (misidentified): KඈඋGൾ (1962b): 333 (characters, notes); Sආൾඍൺඇൺ (1971a): 144 (characters and distribution); Sආൾඍൺඇൺ (1971b): 1844 (collecting notes and distribution); Sආൾඍൺඇൺ (1976a): 174 (distribution).</p><p>Type material. Hඈඅඈඍඒඉൾ J (CNC), labelled: “ Cache Co., Logan Cn., 5300’, Spring Hollow, 15.VII.1981, J.M. Campbell / sifting pile of squirrel midden / CNC 1017210 / Holotype Quedius altanai Hansen &amp; Brunke, sp. Nov des. A. Hansen &amp; A. Brunke 2021 ”. Pൺඋൺඍඒඉൾඌ: USA: Aඋංඓඈඇൺ: Apache Co., Chuska Mountains, Wagonwheel Campground, [36.45, -109.17], 2500 m, pine and fir 25.-27.VIII.1977, leg. S. Peck (1 CNC); Chuska Mountains, Wagonwheel Campground, [36.45, -109.17], 2250 m, sifted from moss, 12.VII.1976, leg. J.M. Campbell (1 CNC). Cඈඅඈඋൺൽඈ: 6 mi S Molas Pass, [37.66, -107.7], 2895 m, 1.VIII.1973, leg. J.M. Campbell (2 CNC); Red Mountain Pass near Silverton, [37.89, -107.71], 3505 m, 28.VII.1973, leg. J.M. Campbell (1 CNC). IൽൺHඈ: Bloomington Lake Wasatch Mts, [42.14, -111.57], 2500 m, 8.VII.1952, leg.B. Malkin (1J FMNH); Boise County, Sawtooth Region, Bull Trout Lake, 3.2km SW Highway 21, [44.3, -115.25], 2130 m, 6.VI.1989, leg.A. Smetana (3 CNC); Custer County, Sawtooth Range Highway, 21 Banner Summit, [44.3, -115.23], 2190 m, 6.VI.1989, leg. A. Smetana (1 CNC); Custer County, Sawtooth Range, Stanley Lake, [44.24, -115.05], 1900 m, 8.VI.1989, leg. A. Smetana (5 CNC). Nൾඐ Mൾඑංർඈ: Otero, 19. Mi S of Cloudcroft, [32.69, -105.72], IV.22.1972, C.W. &amp; L.B. O’Brien (1 J CNC). UඍൺH: Ashley National Forest, 12 km S Manila, Deep Creek, 40.8550, -109.7297, 2340 m, sifting litter and mushrooms, 30.VI.2015, leg. M. Schülke (1 J ZMHB); Cache County, Logan Canyon, Spring Hollow, [41.75, -111.71], 1615 m, sifting, pile of squirrel midden, 15.VII.1981, leg. J.M. Campbell (3 CNC, 2 NHMD); Cache County, Logan Canyon, Spring Hollow, [41.75, -111.71], 1615 m, 15.VII.1981, leg. J.M. Campbell (4 CNC); Cache County,Logan Canyon, 2km N Wood Camp, [41.75, -111.71], 1706 m, sifting, moss, 14.VII.1981, leg. J.M. Campbell (1 CNC); Lonesome Beaver, Henry Mountain, [38.1, -110.81], 25-26.VII.1968, leg. H.F.Howden (1 CNC);Mount Timpanegos,Wasatch Mountains,[40.39, -111.64], 5.VII.1935, leg.Van Dyke (2 CNC); Summit County, Bald Mount Pass, [40.68, -110.89], 3261 m, 11.VII.1981, leg. J.M.Campbell (2 CNC); Summit County, Whiskey Creek, Mile 39 Highway 150, Sulphur Campground, [40.79, -110.88], 2743 m, 11.VII.1981, leg. J.M. Campbell (2 CNC); Summit County, Wasatch National Forest, Beth Lake, [40.65, -110.96], 2895 m, 12.VII.1981, leg. J.M. Campbell (1 CNC). WඒඈආංඇG: [Yellowstone] National Park, [44.63, -110.87], IV.1908, leg.Hubbard &amp; Schwarz (1 CNC). Wyoming USA, 26.VI.1956, leg. E. Suenson (2 NHMD).</p><p>Description. Measurements J (n = 1): HW = 1.67; HL = 1.49; HL/HW 0.89; PW = 2.20; PL = 2.07; PL/PW 0.94; EW = 2.24; EL = 2.13; EL/EW 0.95; EL/PL 1.03; PW/HW 1.48; forebody length 5.69. ♀♀ (n = 6): HW = 1.56–1.67 (1.60); HL = 1.20–1.42 (1.34); HL/HW 0.75–0.89 (0.84); PW = 1.96–2.24 (2.07); PL = 1.78–1.96 (1.86); PL/PW 0.87–0.95 (0.90); EW = 2.09–2.24 (2.18); EL = 1.96–2.16 (2.05); EL/EW 0.87–0.99 (0.94); EL/PL 1.04–1.14 (1.10); PW/HW 1.27–1.74 (1.51); forebody length 5.04–5.53 (5.25).</p><p>Medium sized species; body brown to black (Fig. 10C).</p><p>Head black, distinctly transverse, with eyes medium sized (EyL/TL = 1.61–2.07 (1.90)), microsculpture of transverse waves, with two interocular punctures between anterior frontal punctures (Fig. 6G); antennae and palpi dark with joints pale reddish, all antennomeres elongate, except the last and penultimate antennomere almost transverse.</p><p>Thorax: pronotum dark brown to black, slightly wider than long, clearly wider than head, with microsculpture of transverse waves, three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond first puncture of dorsal row; scutellum sparsely punctured and pubescent; elytra variable in coloration, from reddish brown, to almost yellowish pale, occasionally dark with lighter sutural margins, uniformly pubescent, with punctures clearly separated, slightly wi-der than long, slightly longer than pronotum; legs reddish brown with inner face of tibia and inner surface of femur darkened and tarsi paler.</p><p>Abdomen brown to black, tergites uniformly punctured, with clear iridescence.</p><p>Male. Aedeagus (Figs 16C, 17): Paramere lanceolate, rather slender, without clear basal attenuation, reaching apex of median lobe, with small sensory peg setae forming two irregular rows meeting in irregular cluster below apex; median lobe lanceolate, slender, continuously constricting towards pointed apex, on parameral side with two small teeth simply protruding outwards and slightly basad; internal sac with two larger oval and two smaller rounded internal sclerites.</p><p>Differential diagnosis. Quedius altanai can be distinguished from Q. laticollis and Q. strenuus by the presence of two additional setiferous punctures between the anterior frontal punctures. It can be distinguished from Q. labradorensis by the darker body and the clearly darkened basal three antennomeres. It can be distinguished from other members of the molochinoides -group ( Q. horni, Q. lanei and Q. molochinoides) only by the male genitalia: by the slender paramere that is evenly narrowed from base to apex, and median lobe with an evenly narrowed apex and two small subapical teeth pointing basad and often appearing as short carinae. However, the clearly allopatric distribution of Q. altanai with respect to all other species of the molochinoides -group makes such detailed examination for diagnostic purposes within the group largely unnecessary.</p><p>Bionomics. Quedius altanai prefers mountainous areas and occurs only at high elevations between 1600 and 3500 m. The habitat seems to be mainly moist debris, and under rocks in meadows or coniferous forests. Specimens from Red Mountain Pass were collected from under rocks near snowfields (Sආൾඍൺඇൺ 1976) and those from Lincoln National Forest, New Mexico were found in a meadow-like biotope, amidst coniferous forest under a piece of wood at a small creek (Sආൾඍൺඇൺ 1971). Other specimens have been reported from litter, moss, and squirrel middens mainly in coniferous forests.</p><p>Etymology. This species is dedicated to our great friend and colleagueAleš Smetana who has greatly advanced the study of the complex genus Quedius globally and who reviewed this paper.We are glad that Aleš saw it. Sadly, he recently passed away while this paper was still unpublished. We foresee large future taxonomic changes, when this huge genus will be split, inevitably following recent phylogenetic findings. So, this becomes a way to capture and eternalize his name within the core of the genus. The name is a combination the first part of the first name (al-) and the last part of the last name (-tana) given as a masculin eponym (-i).</p><p>Distribution. Quedius altanai is known from high elevation sites of the central and southern Rockies, as well as western Sierra Madre (Fig. 21). Its northernmost limit of the distribution is from middle Rockies, i.e. Central- -Southern Idaho and Wyoming (ecoregion 6.2.10, 6.2.15). In the south it extends into the highest elevation sites of the Rockies (ecoregion 6.2.13 and 6.2.14) and interestingly into western Sierra Madre that often is represented by a different fauna (ecoregion 13.1.1). Westwards Q. altanai does not occur in the Cascades, Sierra Nevada or the Blue Mountains. There it is replaced by Q. lanei .</p></div>	https://treatment.plazi.org/id/038987A0FFCA4B28B0C5A4781772FAEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFCB4B2BB126A59817E3FC6B.text	038987A0FFCB4B2BB126A59817E3FC6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius horni Hatch 1957	<div><p>Quedius horni Hatch, 1957</p><p>(Figs 1, 4, 5, 10A, 16B, 17, 21)</p><p>Quedius horni Hatch, 1957:216 [Type locality: Washington: Snohomish Co.: L. Ballinger]</p><p>References. KඈඋGൾ (1962b): 333 (characters, notes); Sආൾඍൺඇൺ (1965a): 39 (characters); (1971a): 146 (characters and distribution).</p><p>Material examined. CANADA: BඋංඍංඌH Cඈඅඎආൻංൺ: 20 mi. E. Hope, Manning Pk., [49.18, -120.97], ex river debris, 21.VI.1968, leg. Campbell &amp; Smetana (1 CNC); Tsawwassen, [49.01, -123.08], 19.V.1968, leg. Campbell &amp; Smetana (1 CNC). Hൺංൽൺ Gඐൺංං: Queen Charlotte Island [Haida Gwaii], Skowkona Creek, 7.9km NW O. C.City, [53.33, -132.13], ex dung and carrion traps, 19-29.VII.1983, leg. J.M. Campbell (1 CNC); 7.9km NW Queen Charlotte City, [53.31, -132.14], dung trap, 19-29. VII.1983, leg. J.M. Campbell (1 CNC). Vൺඇർඈඎඏൾඋ Iඌඅൺඇൽ: Mesachie Lake, Forest Experiment Station, [48.82, -124.13], 15-31.VII.1979, leg. I. Smith (2 CNC); Vancouver Island, [49.76, -125.46] (4 CNC). USA: AඅൺඌKൺ: Auke Bay, 16.2km N Juneau, [58.38, -134.65], sifting moss, 9.VI.1981, leg, Bright (1 CNC). OඋൾGඈඇ: Blodgett, [44.59, -123.52], VI.1927, leg. Darlington (1 MCZ); Hood River County, 10 mi S Mount Hood at Clinger Springs,[45.4, -121.55], 1310 m, 8.VII.1978, leg.L.&amp; N. Herman (1 CNC); Mt.Hood, South Side, [45.29, -121.7], 7.VI.1927, leg. Darlington (1 MCZ). WൺඌHංඇGඍඈඇ: Olympic National Park, Hoh Ranger Station, [47.85, -123.93], 182 m, 13.V.1968, leg. Campbell &amp; Smetana (1 CNC); Olympic National Park, 4-6 mi SE Soleduck Campground, [47.93, -123.77], 944 m, 15.VIII.1979, leg. J.M. &amp; B.A. Campbell (1 CNC).</p><p>Redescription. Measurements JJ (n = 4): HW = 1.64– 1.69 (1.67); HL = 1.36–1.40 (1.39); HL/HW 0.82–0.84 (0.83); PW = 2.02–2.22 (2.13); PL = 1.82–2.00 (1.92); PL/PW 0.88–0.91 (0.90); EW = 2.11–2.31 (2.21); EL = 2.04–2.29 (2.11); EL/EW 0.88–1.01 (0.96); EL/PL 1.02– 1.14 (1.10); PW/HW 1.44–1.59 (1.54); forebody length 5.24–5.69 (5.42). ♀♀ (n = 2): HW = 1.62–1.71 (1.67); HL = 1.36–1.42 (1.39); HL/HW 0.83–0.84 (0.83); PW = 2.02–2.16 (2.09); PL = 1.78–2.00 (1.89); PL/PW 0.88–0.93 (0.90); EW = 2.13–2.29 (2.21); EL = 2.04–2.16 (2.10); EL/EW 0.94–0.96 (0.95); EL/PL 1.08–1.15 (1.11); PW/ HW 1.49–1.52 (1.50); forebody length 5.18–5.58 (5.38).</p><p>Medium sized, robust species; body black (Fig. 10A).</p><p>Head black, distinctly transverse, with eyes medium sized (EyL/TL = 1.76–2.06 (1.91)), microsculpture of transverse waves, with two interocular punctures between anterior frontal punctures (Fig. 6G); antennae and palpi pale reddish with most of antennomeres 1–3 clearly darkened; all antennomeres elongate, except apical antennomere almost transverse.</p><p>Thorax: pronotum black, slightly wider than long, clearly wider than head, with microsculpture of transverse waves, three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond first puncture of dorsal row; scutellum sparsely punctured and pubescent; elytra most often reddish brown, but occasionally fully darkened or dark with reddish sutural margins, uniformly pubescent, with punctures clearly separated, quadrate or slightly wider than long, slightly longer than pronotum; legs reddish brown with inner face of tibia and inner surface femur darkened and tarsi paler.</p><p>Abdomen black, tergites uniformly punctured, with clear iridescence.</p><p>Male. Aedeagus (Figs 16B, 17): paramere rather broad, with clear basal attenuation and pointed apex, reaching to apex of median lobe, with small sensory peg setae forming thin row in a semi-circle near edge of paramere, except at pointed apex; median lobe broad with sudden constriction to a point at apex, on parameral side with two fang-like subapical teeth basad; internal sac with elongate and broad sclerites.</p><p>Differential diagnosis. Quedius horni can be distinguished from Q. laticollis and Q. strenuus by the presence of two additional setiferous punctures between the anterior frontal punctures. It can be distinguished from Q. labradorensis by the darker body and the clearly darkened antennomeres 1–3. Quedius horni can be clearly distinguished from other members of the molochinoides -group ( Q. altanai, Q. lanei and Q. molochinoides) only by the spoon-like shape of the paramere, which has a clear basal attenuation and broadening towards pointed apex, with its peg setae situated in a semi-circle near the edge of the of the paramere below its pointed apex. However, this species co-occurs with other species of the molochinoides -group only in Alaska, which makes such detailed examination for diagnostic purposes within the group less necessary anywhere else.</p><p>Bionomics. Little is known about the habitat requirements of Q. horni . It seems to prefer wet environments, as it has been collected in moist habitats from sea level to an elevation slightly above 1000 meters. Most specimens were collected in coastal sites or inland near rivers and creeks suggesting its close association with wet habitats near water bodies. For example, the species was found in river flood debris and on beaches under driftwood in British Columbia, in moss at Auke Bay, Alaska, in a dung trap on Haida Gwaii, and in forest litter around large firs at Mary’s Peak, Oregon (Sආൾඍൺඇൺ 1971a). Immature specimens were collected in late May at Tsawwassen, British Columbia and in early July on Mt. Hood, Oregon (Sආൾඍൺඇൺ 1971a). Based on the records near coastal sites the species is most likely somewhat halotolerant.</p><p>Distribution. Quedius horni is now known from the Pacific Northwest, from Oregon to southern Alaska, including Vancouver Island and Haida Gwaii (most of ecoregion 7.1; Fig. 21). It can be found at coastal and riverine sites and to mid elevation of the Cascades and Mount Olympia. This revision, including barcoding, confirmed its first record from southern Alaska, and thus extended the species range significantly to the north. It is still uncertain how far south the species occurs, with the current southernmost record coming from central Oregon. Based on the extent of ecoregion 7.1, this may extent along the coast into California.</p></div>	https://treatment.plazi.org/id/038987A0FFCB4B2BB126A59817E3FC6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFC84B2AB0E3A4191140FC0A.text	038987A0FFC84B2AB0E3A4191140FC0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius labradorensis Smetana 1965	<div><p>Quedius labradorensis Smetana, 1965</p><p>(Figs 1, 4, 10F, 16A, 17, 21)</p><p>Quedius labradorensis Smetana, 1965a: 37 [Type locality:Südost-Labrador: Forteau]</p><p>Quedius labradorensis insularis Smetana, 1971a: 140 [Type locality: Alaska: Anton Larsen Bay, Kodiak]. Preoccupied by Quedius insularis Bernhauer &amp; Schubert, 1916; currently a junior synonym of Philonthus umbratilis (Gravenhorst, 1802) .</p><p>Quedius labradorensis kodiakensis Smetana 1999: 79 (as a replacement name for insularis Smetana), syn. nov.</p><p>References. Sආൾඍൺඇൺ (1964): 37, (1965a): 37, (1965b): 12 (distribution; misidentified as Q. lanei according to Sආൾඍൺඇൺ 1971a: 138); (1971a): 138 (characters and distribution); (1973): 1428, (1976a): 172, (1978a): 826, (1981): 640 (distribution); Dඈඐඇංൾ &amp; Aඋඇൾඍඍ (1996): 391 (characters and distribution); PඈHඅ et al. (2007): 299 (biology); Wൾൻඌඍൾඋ et al. (2012): 315 (distribution).</p><p>Type material examined. Quedius labradorensis: Hඈඅඈඍඒඉൾ J (CNC), labelled: “SE Labrador Forteau N:o 285.13-15.VII.51 Lindroth / Holotype Quedius labradorensis m. / 1963 Smetana det. / Quedius labradorensis m. 63 Smetana det. 19 / Holotype CNC No. 9473 Quedius labradorensis Smet. ”.</p><p>Quedius labradorensis insularis: Hඈඅඈඍඒඉൾ J (CNC), labelled: “ Alaska, Anton Larsen Bay, Kodiak, 445, 24.VIII.1958, Lindroth / Holotype Quedius labradorensis insularis Smetana 1968 CNC No. 10866”. Pൺඋൺඍඒඉൾ: same collection data as holotype (1 ♀ CNC).</p><p>Additional material examined. CANADA: Aඅൻൾඋඍൺ: George Lake, [53.95, -114.1], 10.IX.1966, leg. R. E. Leech (18 CNC); George Lake, [53.95, -114.1], 1.VI.1969, leg. H. Goulet (3 CNC); Wagner Natural Area, Edmonton 6km W, [53.57, -113.82], pan trap, 12-25.VI.1985, leg. Finnamore &amp; Thormin (4 CNC). BඋංඍංඌH Cඈඅඎආൻංൺ: 10 mi E of East Pine, [55.71, -120.97], 16.VI.1958, leg. Lindroth (1 CNC). Mൺඇංඍඈൻൺ: Onah, [49.81, -99.52], 20.VIII.1921, leg. N. Criddle (1 CNC); Riding Mountain National Park, [50.83, -100.20], pan traps, Bison enclosure, 4.VIII.1979, leg. D. B. Lyons (1 CNC); Riding Mountain National Park, Edward Creek at Highway 10, [50.92, -100.05], 17.IX.1979, leg. A. Smetana (1 CNC); Riding Mountain National Park, 1km SE Long Lake, [50.75, -100.32], 16.IX.1979, leg. A. Smetana (1 CNC). NඈඋඍHඐൾඌඍ Tൾඋඋංඍඈඋංൾඌ: Highway 2 mi SE Fort Simpson, [61.82, -121.3], 23.VI-3.VII.1972, leg. A. Smetana (3 CNC). Nඈඏൺ Sർඈඍංൺ: Cape Breton Highland National Park, near Benjies Lake, [46.73, -60.8], 17.VI.1984, leg. A. Smetana (1 CNC); Cape Breton Highlands National Park, Fishing Cove Trail, [46.79, -60.87], 305 m, sifting, moss at seepage, 25.IX.1984, leg. J. M. Campbell &amp; A. Davies (1 CNC); Cape Breton Highlands National Parks, Lone Shieling, [46.81, -60.73], pan trap, forest, 19-24.VII.1983, leg. L. Masner (7 CNC); Cape Breton Highland National Park, MacKenzie Mountain, [46.81, -60.81], pan trap, 2.VIII.1983, leg. D. E. &amp; J. E. Bright (7 CNC); Cape Breton Highland National Park, North Mountain, [46.82, -60.65], fern bog, 11-19. VII.1983, leg. L. Masner (3 CNC); Cape Breton Highland National Park, Paquette Lake, [46.81, -60.44], pan trap, 1.VII.1983, leg. R. Vockeroth (1 CNC); Cape Breton Highlands National Park near Sammy’s Barren, [46.81, -60.72], 440 m, sifting litter, Birch litter, 24.IX.1984, leg. J. M. Campbell &amp; A. Davies (4 CNC); Cape Breton Highland National Park, [46.81, -60.76], 400 m, pan trap, forest, 11-13.VI.1983, leg.Y.Bousquet (1 CNC). Oඇඍൺඋංඈ: Cochrane, [49.14, -81.02], 22.VIII.1918, leg. H. Notman (1 CNC); Moosonee, [51.29, -80.71], 30.VI.1973, leg. J. M. Campbell &amp; R. Parry (1 CNC); 22 mi S Pickle Lake, [51.15, -90.21], 20.VI.1973, leg. J. M. Campbell &amp; R. Parry (1 CNC); 36 mi S Pickle Lane, [50.76, -90.28], 22.VI.1973, leg. J. M. Campbell &amp; R. Parry (2 CNC); Thunder Bay, Mount McKay, [48.34, -89.28], 487 m, 15.VI.1973, leg. J.M. Campbell &amp; R. Parry (3 CNC); Thunder Bay, Powell Lakes, 10km NE Lake Superior, [49.02, -82.98], forest FIT, 31.V.1980, leg. M Kaulbars (1 CNC). Qඎൾൻൾർ: Charlevoix-East, Ste-Mathilde, [47.68, -70.11], 28.VIII.1975, leg. J. F. Landry (2 CNC); Duparquet, [48.47, -79.17], 30.V.1936, leg. G. Stace Smith (2 CNC); Lac Duparquet, [48.49, -79.26], cedar fir forest, 4.-11.VIII.1996, leg. Paquin (5 CNC); Lac Cascapedia, Parc Gaspesie, [48.92, -66.32], 518 m, pan trap, 14.VII.1972, leg. J. M. &amp; B. A. Campbell (1 CNC); Lac Pointe, [46.33, -72.7], 22.IX.1980, leg.C. Chantal (3 CNC); Longue-Pointe-de-Mingan, [50.28, -64.19], undergrowth vegetation, 16.VII.2008, leg. LeSage &amp; Lussier (1 J CNC); Montreal, [45.5, -73.62], 9.VI.1908, leg. E. J. Kiteley (2 CNC). Iඅൾ ൽ’ Aඇඍංർඈඌඍං: Pte Sud-Ouest, [49.39, -63.59], 15.VII.1977, leg. J. F. Landry (1 CNC). SൺඌKൺඍർHൾඐൺඇ: Flotten Lake, [54.62, -108.50], 6.-13.VI.1998, leg. T. Cobb (3m SMNH); Lac La Rouge, Stueck Island, [55.19, -105.01], 19.VII.1979, leg. E. J. Kiteley, (1 CNC). USA: AඅൺඌKൺ: Denali State Park, Byers Lake Campground, [62.74, -150.11], 23.VI.1978, leg. Smetana &amp; Becker (2 CNC); Denali NP, Wickersham Dorne, 63.348, -150.97749, 977m, alpine tundra, pitfall traps, 29.VI.2017, leg. A. Haberski et al. (3 UAM); Denali NP, Wickersham Dorne, 63.348, -150.97749, 977m, shrubs, pitfall traps, 23.- 27.VII.2017, leg.A. Haberski et al. (19 UAM); Denali NP, Wickersham Dorne, 63.548, -150.97749, 811m, shrubs, pitfall traps, 3.-19.VII.2018, leg. D. Sikes et al. (4 UAM); Denali NP, Wonder Lake, 63.48822, -150.86237, 645m, spruce, pitfall traps, 7.-21.VI.2018, leg. D. Sikes et al. (1 UAM); Fairbanks Park, 64.7924, -148.1511, 440 m, S-facing hill, forest edge Populus tremuloides, pitfall traps, 17.-14.VII.2007, leg. B. Mortensen (1 UAM). KඈൽංൺK Iඌඅൺඇൽ: 30 mi S Kodiak, [57.54, -153.51], 23.VIII.1958, leg. G.E. Ball (1 CNC). MංർHංGൺඇ: Isle Royale, Daisy Farm, [48.09-88.60], 17.VIII.1965, leg. Bixler (1 J FMNH). Nൾඐ HൺආඉඌHංඋൾ: Carter Notch, [44.26, -71.19], 6.VIII.1908, leg F. Blanchard (1 CNC). Nൾඐ YඈඋK: Indian Falls, Mount Marcy, [44.11, -73.92], 16.VI.1942, leg. H. Dietrich (1 CNC).</p><p>Redescription. Measurements JJ (n = 3): HW = 1.76– 1.87 (1.80); HL = 1.49–1.56 (1.52); HL/HW 0.83–0.85 (0.84); PW = 2.20–2.44 (2.28); PL = 2.07–2.29 (2.15); PL/PW 0.94–0.95(0.94); EW = 2.29–2.40 (2.35); EL = 1.93–2.18 (2.07); EL/EW 0.82–0.95 (0.88); EL/PL 0.91– 1.04 (0.96); PW/HW 1.46–1.57 (1.50); forebody length 5.49–5.93 (5.73). ♀♀ (n = 2): HW = 1.89–1.93 (1.91); HL = 1.58–1.62 (1.60); HL/HW 0.84; PW = 2.33–2.38 (2.36); PL = 2.13–2.16 (2.14); PL/PW 0.90–0.92 (0.91); EW = 2.36–2.40 (2.38); EL = 2.09–2.13 (2.11); EL/EW 0.87–0.91 (0.89); EL/PL 0.98–0.99 (0.98); PW/HW 1.47–1.48 (1.47); forebody length 5.84–5.87 (5.86).</p><p>Large species; body brown to black (Fig. 10F).</p><p>Head black, transverse, with eyes medium sized (EyL/ TL = 1.72–1.89 (1.82)), microsculpture of transverse waves, with two interocular punctures between anterior frontal punctures (Fig. 6G); antennae and palpi reddish with basal three antennomeres clearly pale, all antennomeres elongate.</p><p>Thorax: pronotum brown to black, slightly wider than long, clearly wider than head, with microsculpture of transverse waves, three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching almost to level of second puncture of dorsal row; scutellum sparsely punctured and pubescent; elytra most often pale orange to reddish brown, rarely fully darkened, clearly wider than long, of similar length as pronotum, with rather short uniform pubescence, with punctures clearly separated from each other; legs reddish with inner face of tibia darkened and tarsi paler.</p><p>Addomen brown to black, tergites uniformly punctured, without clear iridescence.</p><p>Male. Aedeagus (Figs 16A, 17): paramere rather broad, with slight basal attenuation and extending into softly rounded apex, reaching to the apex of median lobe, with small sensory peg setae forming two broad irregular rows near the edge of the of the paramere, except at very apex which is slightly folded; median lobe continuously slimming toward pointed apex, on parameral side with two folded teeth which almost meet to form a cavity with a small slit towards apex; internal sac with elongate and internal sclerites.</p><p>Differential diagnosis. Quedius labradorensis can be distinguished from Q. laticollis and Q. molochinus by the presence of two additional setiferous punctures between the anterior frontal punctures. It can be distinguished from other members of the molochinoides -group ( Q. lanei, Q. molochinoides, and Q. horni) by the larger body size and pale basal three antennomeres.</p><p>Synonymic notes. The subspecies Q. l. kodiakensis was described as a form restricted to Kodiak Island, which differs from the nominotypical subspecies by a slightly larger body and more spatulate paramere with larger peg setae (Sආൾඍൺඇൺ 1971a). Generally, species of Quedius s. str. display some variation in body size and shape of the paramere. With more material available, the observed variation between the putative subspecies falls within the variation observed in the widely distributed nominal subspecies. Thus, we consider Quedius l. kodiakensis Smetana, 1999 syn. nov. a junior subjective synonym of Quedius labradorensis Smetana, 1965 .</p><p>Bionomics. As a northern species, Q. labradorensis prefers wet boggy habitats with specimens collected in litter and moss of mixed forests, mainly poplar and birch, in Ontario (Sආൾඍൺඇൺ 1971a), in deep moss in Saskatchewan (Sආൾ-ඍൺඇൺ 1990), fern bogs in Nova Scotia, margins of creeks on Kodiak Island (Sආൾඍൺඇൺ 1971a), and shrubby alpine tundra in Alaska. Sආൾඍൺඇൺ (1976a) reported this species from moss, mushrooms, and deciduous leaf litter (birch and poplar stands) near streams. In the Alberta foothills forests, Q. labradorensis is considered to be an open-ground specialist after forest logging, which only spread short distances into adjacent pristine forests, with numbers of this species also increased after logging (PඈHඅ et al. 2007). In New Brunswick, most adults have been found from April to May in old-growth eastern white cedar swamps in moss usually near small streams, with the exception of one adult found under a patch of white-tailed deer dung, and one collected at a mercury-vapor light (Wൾൻඌඍൾඋ et. Al 2012). The species is found mainly at higher elevation (750–1500 m) in the southeastern part of its range and at lower elevations up to 800 m in the northern part of its range. Several specimens found in pan traps across Canada indicate that the species may be actively flying or hunting in the vegetation.</p><p>Distribution. Quedius labradorensis is a northern species distributed transcontinentally across boreal North America from Labrador and Newfoundland to Alaska where it extends to Kodiak Island (ecoregions 4, 5, and northern part of 6 and 7, Fig. 21). To the south it is found in some of the large remaining wetlands, as well as at higher elevation sites, e.g. in Northern Rockies and Cypress Hills in the West, and at Mount Washington and the MacIntyre Range in the East. The northernmost records from Fairbanks Park in Alaska, and Fort Simpson in the Northwest Territories suggest that the species extends throughout the boreal ecoregion of the continent.</p></div>	https://treatment.plazi.org/id/038987A0FFC84B2AB0E3A4191140FC0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFC94B2CB3CEA3A917E9F8CA.text	038987A0FFC94B2CB3CEA3A917E9F8CA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius lanei Hatch 1957	<div><p>Quedius lanei Hatch, 1957</p><p>(Fig. 1, 4, 5, 10B, 16D, 17, 21)</p><p>Quedius lanei Hatch, 1957: 216 [Type locality: Washington: Blue Mts.]</p><p>References. KඈඋGൾ (1962b): 333 (characters and notes); Sආൾඍൺඇൺ (1971a): 144 (characters and distribution); (1976a): 174 (distribution).</p><p>Material examined. USA: Cൺඅංൿඈඋඇංൺ: Fresno County, Huntington Lake, [37.24, -119.18], 25.IV.1960, leg. L. G. Ingles (2 CNC); Lassen National Park,Summit Lake, [40.49, -121.42], 2042 m, 15.VII.1979, J. M. &amp; B.A. Campbell (3 CNC). Nൾඏൺൽൺ: Lake Tahoe, Zephyr Cove, [39.01, -119.94], 1889 m, 9.XII.1986, leg.A.Smetana (1 CNC). OඋൾGඈඇ: 16 mi NE Bly Deming Creek Road, [42.55, -120.76], 1828 m, 21.VII.1979, leg. J. M. Campbell &amp; J. Schuh (3 CNC); Grant County, Strawberry Range, Strawberry Lake, [44.3, -118.68], 1920 m, 4.VI.1989, leg. A. Smetana (20 CNC); Klamath County, Gearhart Mountain, [42.51, -120.86], 2194 m, 24.VI.1974, leg.A. &amp; D. Smetana (1 CNC); Klamath County, Mare’s Egg Springs, 7.3 mi SW Fort Klamath, [42.66, -122.08], 25.VI.1974, leg. A. &amp; D. Smetana (2 CNC); Union County, Blue Mountain Road 62, Jarboe Creek, [45.80, -117.85], 1200 m, 29.V.1989, leg. A. Smetana (1 CNC). WൺඌHංඇGඍඈඇ: Blue Mountain, [45.5, -118], 914 m, 29.VIII.1923, leg. M. C. Lane (1 CNC).</p><p>Redescription. Measurements JJ (n = 3): HW = 1.56– 1.78 (1.63); HL = 1.33–1.49 (1.39); HL/HW 0.84–0.87 (0.86); PW = 2.04–2.22 (2.11); PL = 1.80–2.00 (1.87); PL/PW 0.88–0.90 (0.89); EW = 2.16–2.29 (2.20); EL = 2.02–2.11 (2.06); EL/EW 0.92–0.95 (0.94); EL/PL 1.06– 1.14 (1.10); PW/HW 1.49–1.53 (1.52); forebody length 5.18–5.60 (5.33). ♀♀ (n = 6): HW = 1.53–1.73 (1.64); HL = 1.27–1.40 (1.35); HL/HW 0.77–0.86 (0.82); PW = 1.96–2.29 (2.09); PL = 1.73–2.00 (1.85); PL/PW 0.87–0.91 (0.89); EW = 2.00–2.44 (2.18); EL = 1.93–2.29 (2.07); EL/EW 0.92–1.02 (0.95); EL/PL 1.09–1.18 (1.12); PW/ HW 1.46–1.63 (1.55); forebody length 4.98–5.69 (5.27).</p><p>Medium sized species; body dark brown to black (Fig. 10B).</p><p>Head black, distinctly transverse with eyes medium sized (EyL/TL = 1.81–2.14 (1.99)), microsculpture of transverse waves, with two interocular punctures between anterior frontal punctures (Fig. 6G); antennae and palpi dark with joints pale reddish, all antennomeres elongate, except last and penultimate antennomere almost transverse.</p><p>Thorax: pronotum black, slightly wider than long, clearly wider than head, with microsculpture of transverse waves, three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond level of first puncture of dorsal row; scutellum sparsely punctured and pubescent; elytra variable in coloration, from all black or reddish brown to occasionally dark with reddish sutural margins, uniformly pubescent, punctures clearly separated, quadrate or slightly wider than long, slightly longer than pronotum; legs reddish brown with inner face of tibia and inner surface of femur darkened and tarsi paler.</p><p>Abdomen dark brown to black, tergites uniformly punctured, with clear iridescence.</p><p>Male. Aedeagus (Figs 16D, 17): paramere lanceolate, rather broad, without clear basal attenuation, extending to apex of median lobe, with small sensory peg setae forming two irregular rows meeting in an irregular cluster below apex; median lobe lanceolate, broad, with sides slightly constricting until abrupt constriction near pointed apex, on parameral side with two blunt teeth simply protruding inward and slightly basad; internal sac with two larger oval and two smaller rounder sclerites.</p><p>Differential diagnosis. Quedius lanei can be distinguished from Q. laticollis and Q. strenuus by the presence of two additional setiferous punctures between the anterior frontal punctures. It can be distinguished from Q. labradorensis by the darker body and the clearly darkened basal three antennomeres. It can be distinguished from other members of the molochinoides -group ( Q. altanai, Q. horni, and Q. molochinoides) only by the male genitalia, specifically by the paramere, which is parallel-sided to constricted at base, and the broad median lobe with a constricted apex and two blunt teeth pointing in and downwards. However, Q. lanei does not seem to co-occur with any other species of the molochinoides- group, this fact reducing the need of genitalia investigation in all cases.</p><p>Comments. A series of specimens in Sආൾඍൺඇൺ (1971) including one Hatch paratype from Fish Lake in Summerland, British Columbia ascribed to Q. lanei was found to belong to Q. molochinoides . Quedius lanei does not occur in Canada. Paratypes of Q. lanei from Idaho were found to belong to Q. altanai and have thus been moved to this species.</p><p>Bionomics. Quedius lanei prefers mountainous areas and occurs only at high elevations with records from roughly 1000 m in the northern part of its range to up to 2200 m in the southern part. Little is known about the prefered habitat of the species, but based on the examined records it seems to be found mainly in wet debris near creeks or mountain lakes. For example, the Oregon specimens from Klamath County were collected by sifting wet moss and deciduous leaf litter (willow) at the edges of fast-running creeks or by sifting wet debris on the bottom of a dried-out pond in a coniferous forest (Sආൾඍൺඇൺ 1976).</p><p>Distribution. Quedius lanei is currently known from high elevation sites on the eastern slopes and foothills of Cascades, the Sierra Nevada and the Blue Mountains (ecoregions 6.2.8, 6.2.9 and 6.2.12, Fig. 21). It is found from sutheastern Washington through Oregon to much of eastern California and limited areas of western Nevada. In the north it does not seem to extend into the northern part of the Cascades, although based on the ecoregions it may be be present quite far north on the eastern slopes. To the northwest it is replaced by Q. horni . Quedius lanei was formerly considered to also include specimens from central and southern Rockies, but we here show that these belong to a different species Q. altanai described as new herein.</p></div>	https://treatment.plazi.org/id/038987A0FFC94B2CB3CEA3A917E9F8CA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFCF4B2EB0E6A7781752F94A.text	038987A0FFCF4B2EB0E6A7781752F94A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius molochinoides Smetana 1965	<div><p>Quedius molochinoides Smetana, 1965</p><p>(Figs 1, 4, 5, 10D,E, 16E, 17, 21, 22)</p><p>Quedius molochinoides Smetana, 1965a: 35 [Type locality: Newfoundland: Ha-Ha Bay]</p><p>Quedius molochinoides miquelonensis Smetana, 1965a: 36 [Type locality: Miquelon], syn. nov.</p><p>References. Sආൾඍൺඇൺ (1971): 141 (characters and distribution); (1976): 174, (1978): 826, (1981): 640 (distribution); Dඈඐඇංൾ &amp; Aඋඇൾඍඍ (1996): 391 (characters and distribution).</p><p>Type material examined. Quedius molochinoides: Hඈඅඈඍඒඉൾ: J (CNC), labelled: “NW Newfoundland Ha-Ha Bay N: o 103. 18.VII.49 Lindroth / Holotypus Quedius molochinoides m. / Quedius molochinoides m. 63 Smetana det. 19 / Holotype CNC No. 9474 Quedius molochinoides s. s. Smet. / CNC”. Pൺඋൺඍඒඉൾ (labelled as allotype): same collection data as holotype (1 ♀ CNC).</p><p>Quedius molochinoides miquelonensis: Hඈඅඈඍඒඉൾ: J (CNC), labelled: “ Miquelon N:o 353 8-10.VIII.51 Lindroth / Holotypus ”. Pൺඋൺඍඒඉൾ (labelled as allotype): “ Quedius molochinoides miquelonensis m. / 1963 Smetana det. / Quedius molochinoides miquelonensis m. Smetana det. 1963/ Allotype CNC No. 9475 Quedius molochinoides miquelonensis Smet. ” (1 ♀ CNC).</p><p>Additional material examined. CANADA:Aඅൻൾඋඍൺ: Castle river valley, SW corner of township, [49.49, -114.22], along the road, under rocks etc., 23.V.1990, leg. B.F.&amp; J.L.Carr (1 CNC);Cirrus Mountain, Banff National Park, [52.16, -116.97], 1676 m, 16.VI.1968, leg. Campbell &amp; Smetana (1 CNC); Crow’s Nest Pass, Rocky Mount, [49.59, -114.53] (1 CNC); Kananaskis Forest Experiment Station, [51, -115.1], 1.VIII.1971, leg. J. M. &amp; B.A. Campbell (1 CNC); Mile S Seebe, [51.09, -115.06], 26.VII.1968, leg. A. G. Raske (1 CNC); Tp. 5 Rge. 3 W. 5 Mer, [49.39, -114.34], leg. 23.VI.1994, leg. B. F. &amp; J. L. Carr (1 CNC); Waterton, [49.05, -113.9], 10.VII.1930, leg. Washur (1 CNC); Waterton Lks. Nat. Pk., Cameron Lk., 49.0833, -113.9164, 12.VI.1973, leg. J. Redner &amp; C. Starr (1 CNC); Waterton Lakes National Park, picnic area on Cameron Creek near Cameron Lake, [49.05, -113.98], 1630 m, pitfall trap, 7-11.VI.1980, leg.I. M.Smith (5 CNC); Waterton Lake National Park, Cameron Lake, [49, -114.04], berlese trap, grassy tussock, seepage area, 7.VI.1980, leg. I. M. Smith (1 CNC);Waterton Lake National Park, Crandell Lake Trail,[49.06,-113.94], berlese trap, Alder litter, 13.VI.1980, leg. I. M. Smith (1 CNC); Waterton Lakes National Parks, Lower Waterton Lake, 49.09, -113.85, 1370 m, ex moss, 26.VI.1980, leg. J. M. Campbell (7 CNC);Waterton Lakes National Park, Cameron Lake, [49, -114.04], pitfall trap, wet meadow, 8.VI.1980, leg.I. M.Smith (14 CNC); Waterton Lakes National Park, Cameron Creek picnic ground,[49.04, -113.98], pitfall trap, edge meadow, 17-28.VI.1980, leg. I. M. Smith (2 CNC); Waterton Lakes National Park, Maskinonge Lake,[49.1,-113.83], 1310 m, sifting litter, Populus litter, 14.VI.1980, leg. J. M. Campbell (1 CNC); Waterton Lakes National Park, near Highway 6, 4.4 km SE Junction Highway 5, [49.08, -113.79], pitfall trap, edge of beaver pond, 12-19.VI.1980, leg. I. M. Smith (2 CNC); Waterton Lakes National Park, Crypt Lake,[49, -113.84], 1981 m,under rocks, 22.VI.1980, leg. J. M. Campbell (1 CNC); Waterton Lakes National Park, near Lower Rowe Lake, [49.05, -114.05], berlese trap,moss from spring seepage area, 18.VI.1980, leg. I. M. Smith (4 CNC); Waterton Lakes National Park, Lookout Butte Picnic Site, [49.72, -113.56], flight interception trap, 9-14. VI.1980,leg. J. M.Campbell (1 CNC);Waterton Lakes National Park, Km 9 Chief Mountain Highway, [48.91, -113.43], 1371 m, flight interception trap, 7.VII.1980, leg. H. J. Teskey (38 CNC); Waterton Lakes National Park, near Highway 6 4.4 km SE Junction Highway 5, [49.08, -113.79], pitfall trap, in meadow by beaver pond, 7-11.VI.1980, leg. I. M. Smith (3 CNC); Waterton Lake National Park, Rowe Lakes Trail, [49.05, -114.06], 1828 m, under bark, 12.VI.1980, leg. J. M. Campbell (2 CNC); Waterton NP, Rowe Creek, 49.0833, -113.9164, 1664 m, 2.VIII.1976, leg. J. M. Campbell (1 CNC). BඋංඍංඌH Cඈඅඎආൻංൺ: Amiskwi River, Yoho National Park, [51.38, -116.55], 7.VIII.1971, J. M. &amp; B. A. Campbell (1 CNC); Bowron Lake, [53.23, -121.37], 15.X.1953, leg. G. S. Smith (1 J CNC); Creston, [49.09, -116.51], 29.VII.1956, leg. G. S. Smith, (1 CNC); 8 mi W Creston, [49.09, -116.72], ex river debris, 10.VI.1968, leg. Campbell &amp; Smetana, (1 CNC); 21 mi W Creston, [49.09, -117.04], ex river debris, 9.VI.1968, leg. Campbell &amp; Smetana (1 CNC); Fish Lake, Summerland, [49.84, -115.44], 27.X.1931, leg. A. N. Gartell (2 CNC); Highway 37, 30km S Dease Lake, [58.43, -129.99], 2.VII.1987, leg. L. Carr (2 CNC); Highway 37A, Bear Glacier, [56.09, -129.66], 6-7.VII.1987, leg.Larson &amp; Carr (1 CNC); Koutenay National Park near Dog Lake, [50.78, -115.92], 24.V.1981, leg. I. Askevold (1 CNC); Mt. Revelstoke, [51.09, -118.04], 1737 m, under rocks in swamp, 1.VII.1952, leg. G. J. Spencer (1 CNC); Mount Revelstoke, [51.09, -118.04], 10.VII.1952, leg. G. J. Spencer (1 CNC); Radium, [50.62, -116.07], 26.IV.1994, leg. B.F. &amp; J. L. Carr (1 CNC); Sinclair Creek, Kootenay National Park, [50.63, -116.06], 1828 m, 12.VI.1968, leg. Campbell &amp; Smetana (1 CNC); Summerland, [49.6, -119.67], 27.X.1931, leg. A. N. Gartell (1 CNC); Trinity Valley, [50.4, -118.91], under old log, 16.IX.1943, leg. H. B. Leech, (1 CNC); Vernon, [50.31, -119.28], under stone, 18.IX.1939, leg.H.Leech (1 CNC);Vernon, Deep Lake, [50.79, -120.34], rotten log, hibernating, 10.IV.1989, leg. H. Leech, (1 CNC); Vernon, Monashee Mountains, [50.27, -119.27], 1371 m, under stone, 14.X.1934, leg. H. Leech (1 CNC). Oඇඍൺඋංඈ: 12 mi NE Moosonee, [51.44, -80.45], 1.VII.1973, leg. R. Parry &amp; J. M. Campbell (1 CNC); Gregoire Mills, [49.29, -81.95], 22.VI.1973, leg. J. Redner &amp; C. Starr (1 CNC). YඎKඈඇ: Mile 11 Canol River, [61.98, -132.45], 1219 m, 22.VIII.1962, leg. P. J. Skitsko (1 CNC); Slim’s River km 1705 Alaska Highway, [61.01, -138.51], ex marsh, 7.VII.1981, leg. Bright (1 CNC). Qඎൾൻൾർ: Lac Mitis, 48.30, -67.80, 9.-12.VII.1993, leg. Hébert &amp; Jobín (1 LFC); Parc Gaspesie (Gaspe-O), Mont Logan, [48.89, -66.64], 1040 m, Spruce forest clearing with moss and ericaceae,pitfall traps, 2-14.VII.1993, leg. F. Landry (1 CNC). USA: AඅൺඌKൺ: Becharof NWR, Island Arm East, 57.76107, -155.98752, S facing slope, Salix, fern, &amp; cow parsnips, pitfall traps, 29.VI.-5.VII.2007, leg. Savage (4 UAM); Chichagof Island, 57.98256, -135.21811, 970 m, Dryas, Salix, Cladonia portentosa, ground search, 15.VII.2013, leg. Ridling (1 UAM); Chichagof Island, 57.98708, -135.20357, 702 m, Nephrophyllidium crista-galli, Geranium erianthum, Artemisia arctica, pitfall traps, 14.-15.VII.2013, leg. Ridling (7 UAM); S Chilkat Peninsula, 58.42520, -135.15205, 687 m, subalpine, Vaccinum, Cornus, pitfall traps, 11.-12.VII.2010, leg. D. Sikes (1 ♀ UAM); Haines, Flower Mountain, 59.39353, -136.31129,alpine Sanguisorba,Carex, pitfall traps, 15.-16.VII.2010, leg. D. Sikes (3 UAM); Hope, Kenai Peninsula, [60.89, -149.65], 27.VII.1951, leg.W.J. Brown (1 CNC); Kenai Mountain CreekAbove Tern Lake Campground,[60.53, -149.53], 274 m, 13.VI.1978, leg. Smetana &amp; Becker (1 CNC); Kenai Mountain, 2 mi S Moose Pass, [60.44, -149.37], 182 m, 30.V.1978, leg. Smetana &amp; Becker (1 CNC); Kenai Mountain, 22 mi N Seward, [60.44, -149.45], 152 m, 27.V.1978, leg. Smetana &amp; Becker (2 CNC); Lituya Bay, Cenotaph Island, [58.64, -137.57], 182 m, 22 Pitfalls in Alder Woods, 25.-26.V.1977, D. Mann, (1 CNC); Moose Pass, Kenai Peninsula, [60.49, -149.42], 16.VI.1951, leg. W. J. Brown (2 CNC); Mount Hawthorne, 58.23652, -134.23904, 748m, subalpine meadow, low ridge top, baited pitfall traps, 22.-23.VII.2013, leg. Ridling (4 UAM); Palmer, 61.60, -149.10, pasture, muskox dung, 1.VIII.1988, leg. Pilny (1 DEBU); Seward [60.12, -149.39], 14.V.1951, leg.W.J. Brown (2 CNC); Skagway, 59.50, -135.10, moist pasture,cow &amp; horse dung, 15.VII.1988, leg.Pilny (1 DEBU);Valdez, [61.14, -146.35], on snow, 29.IV.1972, leg.Dick (1 UAM);Valdez, 61.1167, -146.2667, Picea &amp; Populus forest, under wood and stones,wet vegetated gravel, 10.VIII.1988, leg.Pilny(2 DEBU). IൽൺHඈ: Moscow,[46.80,-116.85], 14.IV.1916,leg.A. C. Burrill (1 CNC). Nൾඐ HൺආඉඌHංඋൾ: Mount Washington, [44.27,-71.31], 1874, leg. Blanchard (4 MCZ); White Mountains, Peabody River, [44.39, -71.15], 1828 m, 12.VII.1925, leg. Quirsfeld (1 CNC). Nൾඐ YඈඋK: Indian Falls,Mount Marcy,[44.11, -73.92], 2.IX.1942, leg.H.Dietrich (1J CNC); Lake Tear, Mount Marcy, Essex County, [44.11, -73.93], 27.VII.1922, leg. H. Notman (1 CNC).</p><p>Redescription. Measurements JJ (n = 2): HW = 1.56– 1.71 (1.63); HL = 1.36–1.44 (1.40); HL/HW 0.84–0.87 (0.86); PW = 2.00–2.22 (2.11); PL = 1.91–2.00 (1.96); PL/PW 0.90–0.96 (0.93); EW = 2.02–2.36 (2.19); EL = 1.84–2.29 (2.07); EL/EW 0.91–0.97 (0.94); EL/PL 0.97– 1.14 (1.05); PW/HW 1.48–1.54 (1.51); forebody length 5.11–5.73 (5.42). ♀♀ (n = 5): HW = 1.69–1.89 (1.77); HL = 1.38–1.56 (1.45); HL/HW 0.81–0.83 (0.82); PW = 2.20–2.38 (2.26); PL = 2.00–2.13 (2.07); PL/PW 0.89–0.93 (0.92); EW = 2.22–2.49 (2.36); EL = 2.09–2.22 (2.18); EL/EW 0.88–0.96 (0.93); EL/PL 1.02–1.11 (1.06); PW/ HW 1.52–1.63 (1.55); forebody length 5.53–5.87 (5.70).</p><p>Medium sized species; body dark brown to black (Figs 10D,E).</p><p>Head black, distinctly transverse with eyes medium sized (EyL/TL = 1.76–1.94 (1.87)), microsculpture of transverse waves, with two interocular punctures between anterior frontal punctures (Fig. 6G); antennae and palpi dark with joints reddish pale, all antennomeres elongate, except last and penultimate antennomere almost transverse.</p><p>Thorax: pronotum dark brown to black, slightly wider than long, clearly wider than head, with microsculpture of transverse waves, three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond level of first puncture of dorsal row; scutellum sparsely punctured and pubescent; elytra variable in coloration, from all black to reddish brown, occasionally dark with reddish sutural margins, uniformly pubescent, with punctures clearly separated, slightly wider than long, same length to slightly longer than pronotum; legs reddish brown with inner face of tibia and inner femur darkened and tarsi paler.</p><p>Abdomen dark brown to black, tergites uniformly punctured, with clear iridescence.</p><p>Male. Aedeagus (Figs 16E, 17): paramere lanceolate, rather broad, parallel-sided or with slight basal attenuation, reaching apex of median lobe, with small sensory peg setae in two irregular rows meeting below apex to form a slightly pointed ‘arrowhead’ shape; median lobe lanceolate broad, sides slightly constricting until abrupt constriction near pointed apex, on parameral side with two large pointed teeth folded from lateral margin to form almost closed cavity; internal sac with two larger oval and two smaller rounder sclerites.</p><p>Differential diagnosis. Quedius molochinoides can be distinguished from Q. laticollis and Q. strenuus by the presence of two additional setiferous punctures between the anterior frontal punctures. It can be distinguished from Q. labradorensis by the darker body including the clearly darkened basal three antennomeres. It can be distinguished from other members of the molochinoides -group ( Q. altanai, Q. horni, and Q. lanei) only by the male genitalia, specifically by the broad median lobe with two large, flattened pointed teeth strongly folded to form an almost closed cavity. Quedius molochinoides can also be accidentally confused with Q. brunnipennis Mannerheim, 1843 from the subgenus Raphirus, but will easily be recognized by the entire labrum and by more parallel-sided abdomen.</p><p>Synonymic notes. The subspecies Q. m. miquelonensis was described from the islands of St. Pierre and Miquelon as a slightly broader and larger form with the male sixth sternite wider and more sharply triangular (Sආൾඍൺඇൺ 1971a) compared to the nominal subspecies. Generally, species within Quedius s. str. display notable intraspecific variation in body size. With the new material examined here the observed variation between the two subspecies falls within the limits of the variation observed within the widely distributed nominal subspecies. Thus, we consider Quedius molochinoides miquelonensis Smetana, 1965 syn. nov. a junior subjective synonym of Quedius molochinoides Smetana, 1965 .</p><p>Comments. One of the paratypes of Q. lanei from Fishlake in Summerland of British Columbia, Canada was found to belong to Q. molochinoides . Specimens from the Okanagan area (Summerland, Copper Mountain and Vernon) are slightly different from the nominal species of Q. molochinoides and others of Q. molochinoides- group 3, specifically their genitalia being more slender. Images of the genitalia for two of these are provided in the supplementary material (Fig. S5). We are hesistant to describe them as a new species as we are lacking genetic barcodes as evidence for this. That said they may very well represent yet another species new for science.</p><p>Bionomics. Quedius molochinoides prefers meadows or open forests where it has mainly been collected in litter, grass tussock or under rocks from low elevations up to 2000 m. Specimens from Newfoundland were found on more or less open ground with rich vegetation or in leaf litter under bushes near river edges (Sආൾඍൺඇൺ 1971a), in British Columbia some specimens were collected in river flood-debris and by sifting wet debris and fallen leaves under bushes at mountain creeks, other specimens were found under rocks near snow in Kootenay National Park (Sආൾඍൺඇൺ 1971a). In Alaska the species was collected using pitfall traps in subalpine meadows and shrublands. Immature specimens were collected in July in Alberta and Idaho and from July to September on Mt. Washington in New Hampshire (Sආൾඍൺඇൺ 1971a).</p><p>Distribution. Quedius molochinoides is a northern species distributed transcontinentally across boreal North America from Labrador and Newfoundland to Alaska (ecoregions 4, 5, 6.1 and partly 6.2 and 7.1, Fig. 21). To the south it is found at higher elevation sites, e.g. in the northern Rockies in the west (north part of ecoregion 6.2) and Mount Washington and Mount Marcy in the East (ecoregion 5.3). The northernmost records are from Valdez in Alaska and Churchill in Manitoba, suggesting that the species extends throughout the boreal ecoregion of the continent. There is a single record from Moscow Mountains in Central Idaho, which represents the southernmost record in the Rockies (ecoregion 6.2.3). COI barcoding showed that Q. molochinoides forms three distinct clusters (Figs 4, 5)). Quedius molochionide s-group 1 is known from the Canadian national parks of Banff and Waterton Lakes and is delimited by the Rocky Mountains Trench to the west and the Canadian plains to the East (part ecoregion 6.2.4, Fig. 22). Quedius molochionides -group 2 is found in the Canadian Rockies, west of the Rocky Mountains trench and towards the north continuing to the Pacific Coast in Alaska all the way to the Kenai Peninsula northwards (ecoregion 6.2.1, 6.2.2, 6.2.3 and partly 6.1 and 7.1). Quedius molochinoides -group 3 is present from the Becharof Peninsula of Alaska in the west across the North American boreal region to glacial refugia of New England, Quebec and Newfoundland in the east (ecoregions 4, 5, and partly 6.1 and 7.1).</p></div>	https://treatment.plazi.org/id/038987A0FFCF4B2EB0E6A7781752F94A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFCD4B2EB157A6F817E3F7BB.text	038987A0FFCD4B2EB157A6F817E3F7BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius laticollis (Gravenhorst 1802)	<div><p>Quedius laticollis -group</p><p>Diagnosis. Species of medium size with variable coloration of elytra and appendages, from completely dark to brown, reddish, or yellowish pale. Eyes smaller, temples larger, with temporal puncture separated from posterior margin of eye by distance equal to diameter of this puncture. Frons without interocular punctures (Fig. 6F). Scutellum punctate and setose. Pronotum without puncture(s) in the posterior part of the sublateral row. Restricted to the Nearctic Region.</p></div>	https://treatment.plazi.org/id/038987A0FFCD4B2EB157A6F817E3F7BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFCD4B21B380A0B91106F78A.text	038987A0FFCD4B21B380A0B91106F78A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius laticollis (Gravenhorst 1802)	<div><p>Quedius laticollis (Gravenhorst, 1802)</p><p>(Figs 1, 4, 9D,E, 14E, 22)</p><p>Quedius laticollis (Gravenhorst, 1802): 173 [Type locality: Baltimore] Quedius neomolochinus Korge, 1962b: 333 [Type locality: Washington, D.C.]</p><p>References. EඋංർHඌඈඇ (1840): 536 (characters); MൺඇඇൾඋHൾංආ (1843): 233 (characters); GඋൺඏൾඇHඈඋඌඍ (1847):227 (characters); Kඋൺൺඍඓ (1857): 501 (characters); GൺඇGඅൻൺඎൾඋ (1895): 403 (characters); Fൺඎඏൾඅ (1902): 115 (characters); Pඈඋඍൺ (1907): 129 (characters); Gඋංൽൾඅඅං (1924): 81 (characters); SർHൾൾඋඉൾඅඍඓ (1933):1457 (characters); MඣඊඎංGඇඈඇ (1937): 3 (characters); Sආൾඍൺඇൺ (1958): 364, (1965a): 39 (characters); (1971a): 136 (characters and distribution); (1973): 1427 (lectotype designation, synonymy); (1978a):826, (1981): 640, (1990):98 (distribution), Dඈඐඇංൾ &amp; Aඋඇൾඍඍ (1996): 391 (characters and distribution), Gඈඅൽൾඇ &amp; Cඋංඌඍ (2000): 532 (biology), TංඌHൾർHĸංඇ &amp; Cඅංඇൾ (2008): 335 (biology).</p><p>Material examined. CANADA: Oඇඍൺඋංඈ: Manitoulin I., Kip Fleming Tract, 8 km SW Gore Bay, 45.8702, -82.5418, under stones oak savannah/alvar, 27.V.2010, leg. S. M. Paiero. (1 DEBU); Pinery Prov. Pk., 43.30, -81.83, oak savannah, ex. Leaf litter, reared from larva, 17.IV.2010, leg. A. Brunke (1 DEBU); Rondeau Provincial Park, Spicebush Trail, [42.30, -81.85], mushroom and litter, 4.VI.1985, leg. J. M. Campbell &amp; A. Davies (3 CNC). Qඎൾൻൾർ: Levis, [46.73, -71.24], 25.VI.1979, leg. C. Chantal (1 CNC). USA: Aඅൺൻൺආൺ: Auburn, [32.59, -84.47], 8.IV.1978, leg. E. J. Kiteley (1 CNC); Ophelika, [32.65, -85.38], 23.I.1974, leg. E. J. Kiteley (1 CNC). AඋKൺඇඌൺඌ: Franklin County, Shores Lake, Ozark National Forest [35.63, -93.95], 20.V.1986, leg. J. M. Campbell (2 CNC); Logan County, Cove Lake, 9 mi SE Paris, [35.23, -93.63], ex debris at edge of lake, 25.V.1986, leg. J. M. Campbell (2 CNC); Logan County, 1km E Lookout Mount. Magazine, [35.16, -93.94], sifting litter, leaf litter along edge of stream, 23.V.1986, leg. J. M. Campbell (2 CNC); Logan County, Mount. Magazine, Lookout, [35.16, -93.64], sifting litter, deciduous leaf litter, 26.V.1986, J. M. Campbell (1 CNC); Mobile, [30.69, -88.06], 22.VI.1919 (1 MCZ); Newton County, Buffalo National Reserve, Steel Creek, [36.03, -93.33], treading clumps of Carex, 17.V.1986, leg. J. M. Campbell (2 CNC); Pulaski County, Pinnacle Mountain State Park, Base Trail, [34.84, -92.49], ex flood debris on river, 11.V.1986, leg. J. M. Campbell (1 CNC); Searcy County, Ozark National Forest, Forest Service Road 1205, Richland Creek, [35.77, -95.99], sifting litter, leaf litter along edge of stream, 15.V.1986, leg. J. M. Campbell (2 CNC); Stone County, Blanchard Springs State Park, [35.96, -92.17], 182 m, 18.V.1973, leg. D. H. Kavanaugh &amp; H. Goulet (2 CNC). Fඅඈඋංൽൺ: Alacha Co., Gainesville, [29.63, -82.33], leaf litter &amp; rotten logs, 5.V.1981, leg. Watrous (1 FMNH); Fort Walton Beach, [30.41, -86.61], 21.III.1975, leg. E.J. Kiteley (1 CNC); Leon Co., Tallahassee, [30.43, -84.29], mixed hardwood, 19.XI.1976, leg. Marshall &amp; Justice (2 JJ 3 ♀♀ FMNH); St. Augustine, [29.89, -81.31], leg. Liebeck (1 CNC); Suwanee Co., 15 mi SW Live Oak Hwy. 51., [30.11, -83.17], pine and hardwood litter, 24.III.1977, leg. O’Brien (1 J FMNH); Volusia Co., 2 mi S Enterprise, [28.85, -81.30], leaf litter, 23.VIII.1965, leg. Suter (1 J FMNH). Iඅඅංඇඈංඌ: Union County, Pine Hills near McCann Springs, [37.58, -89.43], 6.V.1976, leg. A Smetana, (4 CNC); Union County, 5 mi NE Reynoldsville, [37.40, -89.28], 9.V.1976, leg. A Smetana (1 CNC); St Clair County, [38.46, -89.95], 6.X.1901, leg. G. W. Bock (1 CNC); Willow Springs, [41.72, -87.86], 16.VIII.1923 (1 MCZ). Lඈඎංඌංൺඇൺ: Calcasieu Parish, Choupique Bayou, 7 mi N Hackberry, [30.13, -93.36], 16.IV.1976, leg. A. Smetana (27 CNC); Calcasieu Parish, 2 mi W Sulphur, [30.24, -93.45], 13.IV.1976, leg. A. Smetana (1 CNC); Corney Lake, [32.91, -92.74], 26.VI.1983, leg. B. F. &amp; J. L. Carr (2 CNC). Mൺඋඒඅൺඇൽ: Baltimore, [39.31, -76.61], 6.VII.1909, leg. F. E. Blaisdell (1 CNC); Baltimore, [39.28, -76.62] (1 MCZ). MൺඌඌൺർHඎඌൾඍඍඌ: Barnstable Co., Woods Hole, 41.5304, -70.6533, 30 m, near small pond, 30.IX.2010, leg. BIObus (3 CMNH). Mංඌඌංඌඌංඉඉං: Oktibeha County, Craig Springs, [33.31, -88.92], peripheral to cult. Cotton, pitfall trap, 10.X.1979, leg. G.L. Snodgrass (2 CNC). Mංඌඌඈඎඋං: Williamsville, [36.97, -90.54], deciduous duff, 12.VII.1955, leg. E. C. Becker (1 CNC). Nൾඐ Jൾඋඌൾඒ: Woodbury, [39.83, -75.15], VIII.1920, leg. H. A. Wenzel (1 CNC). Nൾඐ YඈඋK: Inlet Valley, Ithaca, [42.44, -76.5], 18.X.1917, leg. H. Dietrich (1 CNC). NඈඋඍH Cൺඋඈඅංඇൺ: Ocracoke Island, Ocracoke, [35.12, -75.93], 10.VIII.1972, leg. A. Smetana (2 CNC). OKඅൺHඈආൺ: Latimer Co, [34.85, -95.25], IV.1994, leg. Stephen (5 TAMU). Pൾඇඇඌඒඅඏൺඇංൺ: Allegheny County, [40.46, -79.98], leg. E. A. Klages (1 CNC), Frankford, [40.01, -75.07], leg. H. W. Wenzel (1 CNC). SඈඎඍH Cൺඋඈඅංඇൺ: Camden, [34.25, -80.60], 23.VI.1929, leg. Darlington (1 MCZ). Tൾඑൺඌ: Anderson Co., 15 mi SSE Palestine, 31.5667, -95.4667, pitfall traps, 1.-10.XI.1996, leg. Yantis (2 TAMU); Brazos Co., Koppe’s Bridge, 5 mi SW College Station, [30.57, -96.33], leaf litter, hardwood forest, 15.I.1994, leg. Riley (1 TAMU); Columbus, [29.70, -96.54], VIII.1906, leg. C. V. Riley (1 CNC). Orange County, Bridge City, [30.02, -93.83], 18.IV.1976, leg. A. Smetana (33 CNC); Orange County, 3 mi NW Orange, [30.15, -93.81], 17.IV.1976, leg. A. Smetana (3 CNC); Sabine Co., Sabine Nat. Forest, 3.2 mi E Hemphill, [31.35, -93.79], pine litter, 29.VII.1975, leg. Ashe (1 TAMU); Trinity County, 2.5 mi NW Apple Springs, [31.24, -95.01], 23.IV.1976, leg. A. Smetana (1 CNC); Trinity County, 12 mi SW Lufkin, [31.20, -94.99], 22.IV.1976, leg.A. Smetana (3 CNC). VංඋGංඇංൺ: Arlington Co., Roaches Run, [38.86, -77.04]., 15.IV.2013, leg. J. R. Fisher (1 ♀ FMNH); Lee Co., Young-Fugate Cave, [36.79, -82.97], 25.VI.1963, leg. Holsinger (1 ♀ CMNH). Wൾඌඍ VංඋGංඇංൺ: Glen Dale, [39.94, -80.75], grassy hillside, 29.VII.1973, leg. J. H. Redner (1 CNC).</p><p>Redescription. Measurements JJ (n = 3): HW = 1.67– 1.76 (1.70); HL = 1.33–1.40 (1.37); HL/HW 0.80–0.82 (0.80); PW = 2.20–2.24 (2.21); PL = 1.96–2.04 (1.99); PL/PW 0.89–0.91 (0.90); EW = 2.18–2.29 (2.24); EL = 1.93–2.07 (1.99); EL/EW 0.86–0.92 (0.89); EL/PL 0.99– 1.01 (1.00); PW/HW 1.60–1.65 (1.62); forebody length 5.22–5.51 (5.36). ♀♀ (n = 4): HW = 1.60–1.84 (1.71); HL = 1.29–1.47 (1.38); HL/HW 0.80–0.83 (0.81); PW = 2.04–2.31 (2.19); PL = 1.89–2.22 (2.04); PL/PW 0.91–0.96 (0.93); EW = 2.09–2.44 (2.30); EL = 1.91–2.16 (2.03); EL/EW 0.86–0.91 (0.88); EL/PL 0.97–1.01 (1.00); PW/ HW 1.57–1.60 (1.58); forebody length 5.09–5.84 (5.46).</p><p>Medium sized, robust species; body dark brown to black (Figs 9D,E).</p><p>Head black, distinctly transverse with eyes medium sized (EyL/TL = 1.61–1.88 (1.76)); microsculpture of transverse waves; no additional punctures between anterior frontal punctures (Fig. 6F); antennae and palpi reddish, all antennomeres elongate.</p><p>Thorax: pronotum dark brown to black, slightly wider than long, wider than head, with microsculpture of transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond level of first puncture of dorsal row; scutellum punctured and pubescent; elytra most often dark brown more rarely fully darkened, uniformly pubescent, slightly wider than long, roughly same length as pronotum; legs reddish brown with inner face of tibia and inner femur darkened.</p><p>Abdomen dark brown to black, tergites uniformly punctured, without clear iridescence.</p><p>Male. Aedeagus (Fig. 14E): paramere rather broad lanceolate without clear medial attenuation, reaching just beyond apex of median lobe, with sensory peg setae forming a distinct triangular patch; median lobe broad with gentle constriction to a point at apex, on parameral side with two small teeth protruding slightly basad, positioned at level near lower level of peg setae patch of paramere.</p><p>Differential diagnosis. Quedius laticollis is very similar in North America to the introduced species Q. molochinus, from which it can be confidently distinguished through examination of male genitalia. Other differences include slightly more stout antennae with less elongate, darker middle antennomeres in Q. molochinus . Also, the elytra in Q. laticollis have a finer and denser punctation and are most often brownish-red, yellowish, or dark in coloration, where it is either brick red or dark in Q. molochinus . Quedius laticollis is also similar to Q. strenuus, which replaces Q. laticollis in the Great Plains. However, all examined specimens of Q. strenuus are completely dark with pale appendages, contrary to the rarely fully darkened specimens of Q. laticollis . For confident separation of Q. laticollis and Q. strenuus the position of the peg setae of the paramere are diagnostic as they form two distinct rows in Q. strenuus and a triangular patch in Q. laticollis . Quedius laticollis cannot be confused with any other North American Quedius as they either have interocular punctures on frons or (from other subgenera) have clear medial incision of labrum and abdomen notably tapering apicad.</p><p>Synonymic notes. Quedius neomolochinus Korge, 1962 was originally described from Baltimore most likely without knowledge of Gravenhorst’s description of Q. laticollis, which was at that time a synonym of Q. molochinus . KඈඋGൾ (1962) also described a color variant (black with paler elytral suture and posterior margin) of Q. neomolochinus as Q. marginatus Korge, 1962 which is unavailable name according to the ICZN Article 45.6. Quedius marginatus was thought to be identical with Q. neomolochinus by Sආൾඍൺඇൺ (1971a) when he revised the North American fauna of the tribe Quediini . Later Sආൾඍൺඇൺ (1973) realized that Q. laticollis was identical to Korge’s Q. neomolochinus and established the respective synonymy along with the lectotype designation for Q. laticollis . Our data fully corroborates such established concept of Q. laticollis .</p><p>Bionomics. Quedius laticollis is a rather common generalist found in a large variety of damp lowland habitats. Specimens have been reported from sphagnum in a tamarack swamp (Sආൾඍൺඇൺ 1971a), from sifting damp leaves in a deciduous forest in Florida, from flood-debris along the edges of swamps and marshes, from tufts of swamp grass dumped into water in Louisiana and Texas (Sආൾඍൺඇൺ 1978a), from forest floor litter (hardwood and pine) and from under rocks in Arkansas, and around cotton and soybean fields in Mississippi (Sආൾඍൺඇൺ 1990). One of us (AJB) has collected this species under large boulders in spring in an open scrubby alvar habitat with few oak trees (Manitoulin, Island, Ontario, Canada). This habitat has poorly drained, thin soil on exposed bedrock. TංඌHൾർHĸංඇ &amp; Cඅංඇൾ (2008) recorded a single specimen from a pocket gopher burrow in Louisiana, most likely an accidental occurrence.</p><p>Distribution. Quedius laticollis is found in Eastern North America from the Atlantic Ocean to the Great Plains, where it is replaced by Q. strenuus (ecoregion 8, Fig. 22). In the south it extends to northern Florida and along the Gulf Coast, but not into the subtropical regions of Texas, Florida and Mexico. To the north it extends to the central and north part of Lake Huron (Manitoulin Island) and follows the Saint Lawrence Valley to Quebec City, as the northernmost record.</p></div>	https://treatment.plazi.org/id/038987A0FFCD4B21B380A0B91106F78A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFC74B27B137A5DF14ADF78A.text	038987A0FFC74B27B137A5DF14ADF78A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius strenuus Casey 1915	<div><p>Quedius strenuus Casey, 1915</p><p>(Figs 1, 4, 9F, 14F, 22)</p><p>Quedius strenuus Casey, 1915: 401 [Type locality: Texas]</p><p>References. Sආൾඍൺඇൺ (1971a): 132 (lectotype designation, characters and distribution); (1981): 638, (1990): 98 (distribution).</p><p>Type material examined. Lൾർඍඈඍඒඉൾ: ♀, designated by Sආൾඍൺඇൺ (1971a) (USNM) [not examined], labelled: “Tex / Casey bequest 1925/ Type USNM 48279 / strenuus Csy ”.</p><p>Additional material examined. USA: Kൺඇඌൺඌ: Lawrence, below bridge, Kansas River,[38.96, -95.26],under chunk of mud flat, 16.X.1936, leg. M. W. Sanderson (1 CNC). Tൾඑൺඌ: Braunfels, [29.69, -98.11], 13.VI.1927, leg. Darlington (1 MCZ); Del Rio, [29.36, -100.9], 8.VII.1938, leg. R. L. Sailer (3 CNC); Kerrville, [30.02, -99.13], 11.IV.1998, leg. Chamberlain (3 TAMU); Uvalde County, [29.30, -99.70], 26.V.1938, leg. J. H. Robinson (1 CNC).</p><p>Redescription. Measurements JJ (n = 3): HW = 1.60– 1.82 (1.73); HL = 1.33–1.51 (1.44); HL/HW 0.80–0.85 (0.83); PW = 2.20–2.47 (2.37); PL = 2.02–2.22 (2.14); PL/PW 0.89–0.92 (0.90); EW = 2.33–2.60 (2.50); EL = 2.00–2.24 (2.16); EL/EW 0.85–0.88 (0.86); EL/PL 0.99–1.03 (1.01); PW/HW 1.63–1.67 (1.65); forebody length 5.36–5.96 (5.73). ♀♀ (n = 2): HW = 1.69–1.76 (1.72); HL = 1.44–1.47 (1.46); HL/HW 0.82–0.87 (0.85); PW = 2.33–2.44 (2.39); PL = 2.07–2.09 (2.08); PL/PW 0.85–0.89 (0.87); EW = 2.62–2.64 (2.63); EL = 2.18–2.29 (2.23); EL/EW 0.83–0.87 (0.85); EL/PL 1.05–1.10 (1.07); PW/HW 1.59–1.69 (1.64); forebody length 5.71–5.82 (5.77).</p><p>Medium sized, robust species; body black (Fig, 9F).</p><p>Head black, distinctly transverse, with eyes medium sized (EyL/TL = 1.76–2.00 (1.88)); microsculpture of transverse waves; no additional punctures between anterior frontal punctures (Fig. 6F); all antennomeres elongate, antennae and palpi reddish.</p><p>Thorax: pronotum black, slightly wider than long, wider than head, with microsculpture of transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond first puncture of dorsal row; scutellum punctured and pubescent; elytra black, uniformly pubescent, slightly wider than long, roughly as long as pronotum; legs reddish brown with inner face of tibia and femur darkened.</p><p>Abdomen black, tergites uniformly punctured, with slight iridescence.</p><p>Male. Aedeagus (Fig. 14F): paramere rather broad lanceolate, slightly protruding beyond apex of median lobe, with sensory peg setae forming two distinct short and wide rows of multiple punctures; median lobe broad with gentle constriction towards pointed apex, on parameral side with two small hooked teeth protruding slightly basad, positioned at level near lower level of peg setae patch of the paramere.</p><p>Differential diagnosis. Quedius strenuus is similar to Q. laticollis from which it normally differs by the completely dark body with pale appendages. For confident delimitation of both species from each other the position of the peg setae of the paramere can be used, which form two distinct rows in Q. strenuus and a triangular patch in Q. laticollis . Co-occurrence of the two species is unlikely since Q. strenuus replaces Q. laticollis to the west, in the forested areas of Great Plains of central-eastern and lowland areas of the southwestern United States. Quedius strenuus is also very similar to the introduced species Q. molochinus, which has not yet been found within the distribution range of Q. strenuus . Other differences include that, in Q. molochinus, the antennae are slightly stouter, less elongate, with darker middle antennomeres; as well as elytra are finer and more densely punctate, and either brick red or dark red (always dark in Q. strenuus). Quedius strenuus cannot be confused with other North American Quedius as they either have interocular punctures on frons or (those from other subgenera) have clear medial incision of labrum and abdomen notably tapering apicad.</p><p>Bionomics. Little is known about the bionomics of Q. strenuus . Based on current records, Q. strenuus is most likely restricted to lowland sites. As the region covered by its distribution is rather arid and most specimens were found near creeks, rivers, and lakes, it is likely to be strictly confined to moist microhabitats like wet flood or water-edge debris.</p><p>Distribution. Quedius strenuus is found across the Great Plains (ecoregion 9.4) and potentially a large part of lowland southwest United States (parts of ecoregion 10, Fig. 22). Only a few specimens, which we were not able to examine, were imprecisely recorded from lowland Southwest United State, from Arizona and Southern California. Of these, only a record from Lyman Lake in Apache County, Arizona (Sආൾඍൺඇൺ 1981) is exact and fully reliable. Earlier Sආൾඍൺඇൺ (1971a) mentioned subtle differences between these lowland specimens and ones from the Great Plains, but not clear enough to describe an additional species. The known range of Q. strenuus extends southwards almost to the border between USA and Mexico, northwards to Lawrence in Kansas. To the east Q. strenuus is replaced by Q. laticollis .</p></div>	https://treatment.plazi.org/id/038987A0FFC74B27B137A5DF14ADF78A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
038987A0FFC44B58B384A0B817EDFD6A.text	038987A0FFC44B58B384A0B817EDFD6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quedius Stephens 1829	<div><p>Key to species and species groups of Quedius sensu stricto</p><p>1 Presence of 4 or 6 additional setiferous punctures between anterior frontal punctures (Fig. K1A). Palearctic, two species introduced to the Nearctic. Q. fuliginosus -group .................................................. 5</p><p>– Presence of 2 or no additional setiferous punctures between anterior frontal punctures (Fig. K1B). ..... 2</p><p>2 Presence of 2 additional setiferous punctures between anterior frontal punctures (Fig. K2A). Nearctic. Q. molochinoides -group ..................................... 15</p><p>– No additional setiferous punctures between anterior frontal punctures (Fig. K2B). ................................. 3</p><p>3 Additional 1 to 3 punctures in the posterior half of the sublateral row (Fig. K3A). W Palearctic. Q. pallipes - group ...................................................................... 8</p><p>– No additional puncture in the posterior half of the sublateral row (Fig. K3B). ..................................... 4</p><p>4 Palearctic. Q. molochinus -group .......................... 9</p><p>– Nearctic. Q. laticollis -group and Q. molochinus introduced from Palearctic. .................................. 19</p><p>5 Scutellum punctured and pubescent (Fig. K5A). W Palearctic, usually preferring warmer drier habitats. ................................. Quedius levicollis Brullé, 1832</p><p>– Scutellum unpunctured and glabrous (Fig. K5B). ......................................................................................6</p><p>6 Head with 4 additional punctures between posterior frontal punctures, but without additional puncture between anterior and posterior frontal punctures (Fig. K6A). Median lobe without apical ridge. Paleartic. Introduced to multiple Nearctic localities. ............. 7</p><p>– Presence of 4 additional punctures between posterior frontal punctures and one additional puncture on either side between anterior and posterior frontal punctures (Fig. K6B). Median lobe with apical ridge (Fig. 14A). Palearctic, known only from North Africa and the islands of Sardinia and Corse. ...................... .................. Quedius afrofuliginosus Gusarov, 1991</p><p>7 Basal three antennomeres at least slightly darkened. Eyes clearly protruding making the head seem more rectangular wider across (Fig. K7A). Apex of median lobe not laterally flattened (Fig. 14D). Palearctic. Introduced to E Canada. Prefers wetter humus rich habitats. Paleartic. Introduced to multiple Nearctic localities. ................................................................... ................. Quedius fuliginosus (Gravenhorst, 1802)</p><p>– Basal three antennomeres completely pale. Eyes not clearly protruding making the head seem round (Fig. K7B). Apex of median lobe laterally flattened (Fig. 14C). Paleartic. Introduced to multiple Nearctic localities. ..... Quedius curtipennis Bernhauer, 1908</p><p>8 Sides of median lobe apex folded with distinct tooth situated at lower part (Fig. 15C). Western Europe along Atlantic Coast from Denmark to Iberia and inland in Southern France and Iberian Peninsula, also on the islands of Madeira, Azores, Canaries, and Balearic (Fig. K8A). .................................................. ....................... Quedius hispanicus Bernhauer, 1898</p><p>– Sides of median lobe apex folded with distinct tooth situated at medial part (Fig. 15D). North Africa, Italy and islands of Sardinia and Corse (Fig. K8B). .......... ................................... Quedius pallipes Lucas, 1846</p><p>9 Elytra shortened, distinctly shorter than pronotum (Fig. K9A), obviously brachypterous species without whitish apical seam on abdominal tergite VII (Fig. 11F). East Palearctic. ................................................. .......................... Quedius sundukovi Smetana, 2003</p><p>– Elytra normal, not shortened, about as long as pronotum (Fig. K9B), with whitish apical seam on abdominal tergite VII. ............................................................ 10</p><p>10 Basal three antennomeres at least slightly darkened (Fig. K10A). ......................................................... 11</p><p>– Basal three antennomeres pale (Fig. K10B). ....... 13</p><p>11 Elytra most often reddish, very rarely darkened. Appendages only slightly darkened. Tibia, tarsi and apical antennomeres of antennae pale reddish brown (Figs K11A, 11E). Found near water bodies in Central-Eastern Europe and Middle Asia (Fig. 23). .. .................................. Quedius balticus Korge, 1960</p><p>– Elytra always dark. Appendages darkened. Tarsi slightly paler brown to dark brown at most (Figs K11B, 10E–F). Found at higher elevations or latitudes (Figs 25, 27). .......................................... 12</p><p>12 Head disc with fine isodiametric microsculpture (Fig. K12A). Without C-sclerite of the internal sac (Fig. 16A). Found at alpine sites of Central and Southern Europe (Fig. K12C). .................................................. ...................... Quedius unicolor Kiesenwetter, 1847</p><p>– Head disc most often with microsculpture of fine transverse waves, rarely isodiametric (Fig. K12B). With clear C-sclerite of the internal sac (Fig. 15C). Found disjunct in boreal Scandinavia and Russia, as well as alpine in Altai Mountains and Transbaikalia (Fig. K12D). ..... Quedius subunicolor Korge, 1961</p><p>13 Body brown. Elytra yellowish red. Apex of paramere pointing ventral, away from median lobe in form of a small blunt hook (Figs K13B, 15A). Central Anatolia, Middle East, Transcauscasus, Central Asia (Fig. K13A). ..... Quedius vicinus (Ménétriés, 1832)</p><p>– Body most often black, or at most brownish. Elytra brick or brownish red. Apex of paramere straight, not pointing ventrad, away from median lobe (Figs K13C–D, 15D, 15F). Distribution more northern (Figs K14A–B)...................................................... 14</p><p>14 C-shaped sclerite of internal sac without spine-like basal extension (Figs K13C, 15F). SE European distribution (Fig. K14A). ........................................... ........... Quedius meridiocarpathicus Smetana, 1958</p><p>– C-shaped sclerite of internal sac with spine-like basal extension (Figs K13D, 15D). N Palearctic distribution (Fig. K14B). ........................................... ............... Quedius molochinus (Gravenhorst, 1806)</p><p>15 Basal three antennomeres pale and antennae more elongate (antennomere 10 clearly longer than wide) (Fig. K15A). Elytra light to dark brown. Generally larger. Forebody length 5.49–5.93 (5.78) mm (Fig. 13F). Boreal North America (Fig. 28). ...................... ..................... Quedius labradorensis Smetana, 1965</p><p>– Basal three antennomeres darkened and antennae more robust (antennomere 10 more or less transverse) (Fig. K15B). Elytra black or reddish. Generally smaller. Forebody length 4.97–5.86 (5.39) mm. ........................................................... 16</p><p>16 Paramere with basal attenuation and broadening towards apex forming a spoon like shape, peg setae situated in a semi-circle along the edge of the paramere (Figs K16A, 8, 17B). Coastal to mid elevation from Oregon to Southern Alaska including Vancouver and Haida Gwaii. ..................................... ...................................... Quedius horni Hatch, 1957</p><p>– Paramere only with at most slight attenuation forming a lanceolate shape (Figs K16B–D). ...................... 17</p><p>17 Distributed across Boreal North America. South of the Northern Rocky Mountains (Fig. K17A). Teeth of median lobe clearly folded inwards (Figs K16B, 17E). ......... Quedius molochinoides Smetana, 1965</p><p>– Central and Southern Rocky Mountains, Cascades, Blue Mountains and Sierra Nevada (Fig. K17B). Teeth of median lobe either protruding or not clearly formed (Figs K16C–D). ....................................... 19</p><p>18. Found in the Cascades, Blue Mountains and Sierra Nevada (Fig. K18A). Paramere and median lobe clearly broader (Figs K16C, 17D). ............................ ....................................... Quedius lanei Hatch, 1957</p><p>– Found in Central and Southern Rocky Mountains (Fig. K18B). Paramere and median lobe clearly more slender (Figs K16D, 17C). ........................................ ........................................... Quedius altanai sp. nov.</p><p>19 Paramere rather slender and clear C-sclerite of the internal sac (Figs K19A, 15D). Introduced in parts of Eastern North America, seems to be spreading. ........ ............... Quedius molochinus (Gravenhorst, 1806)</p><p>– Paramere rather broad and No C-sclerite of internal sac (Figs K19B–C). Q. laticollis -group .............. 20</p><p>20 Peg setae of paramere forming a solid triangular field, usually with two wide and short extensions posteriorly (Figs K20A, 16E). E North America (Fig. K20B). ........ Quedius laticollis (Gravenhorst, 1802)</p><p>– Peg setae of paramere arranged into two short longitudinal rows, which more or less diverge posteriorly (Figs K20C, 16F). Great American Plains (Fig. K20D). ........... Quedius strenuus Casey, 1915</p></div>	https://treatment.plazi.org/id/038987A0FFC44B58B384A0B817EDFD6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hansen, Aslak Kappel;Brunke, Adam;Simonsen, Thomas;Solodovnikov, Alexey	Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas, Solodovnikov, Alexey (2022): Revision of Quedius sensu stricto (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1): 225-299, DOI: 10.37520/aemnp.2022.017, URL: http://dx.doi.org/10.37520/aemnp.2022.017
