identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03893F32FFA8FFBDFF46FF70FE8A8830.text	03893F32FFA8FFBDFF46FF70FE8A8830.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus Enderlein 1938	<div><p>Dasysyrphus Enderlein 1938</p><p>Figures: 1 (field photos), 5–19 (lab photos), 20–22 (range maps)</p><p>Synonyms and References:</p><p>Conosyrphus Matsumura 1918: 11 (name preoccupied by Conosyrphus Frey 1915). Type species Conosyrphus okunii Matsumura 1918</p><p>Dasysyrphus Enderlein 1938: 208 . Type species Scaeva albostriata Fallén 1817</p><p>Syrphella Goffe 1944: 129 . Type species Scaeva tricincta Fallén 1817</p><p>Dendrosyrphus Dusek &amp; Laska 1967: 365 . Type species Syrphus lunulatus Meigen 1822</p><p>Diagnosis: Eye pilose; metasternum bare; at least at apical ⅓ of wing densely microtrichose; abdomen oval with abdominal margin present, dark with light maculae present on tergites 2, 3 &amp; 4.</p><p>Description:</p><p>Body Length: 5.0– 11.7mm; Wing Length: 4.5–11.4mm</p><p>Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in females it is variable and can be diagnostic); face light, typically with dark vitta ¼–½ the width of the face, reaching antennal socket or not; face with medial tubercle, more prominent in some species than others; gena dark or light; occiput dark, covered in light pollen, pile typically light; eyes covered in dense pile, male holoptic, female dichoptic; scape, pedicel and flagellomere light to dark, arista bare.</p><p>Thorax: Scutum dark, usually with shiny or metallic sheen (lighting can enhance sheen); scutellum variable in colour (light to dark); wings densely microtrichose on apical ⅓, location and extent of bare areas on basal ⅔ varies between species; posterior anepisternum, katepisternum, anepimeron, katepimeron with dense pile, usually light, sometimes dark or mixture (colour variable within species); anterior anepisternum, meron, metasternum bare; pterostigma elongate and dark (brown); haltere light, sometimes knob or capitulum darkened; calypter light with long light pile around edge and very short light pile on both dorsal and ventral surfaces; legs cylindrical, femora typically basally dark, apically light, tibiae light, metatibia sometimes with dark band close to apex, tarsi light usually with darker anterior side.</p><p>Abdomen: Oval; tergites dark with light maculae, maculae on tergite 2 typically oval, while maculae on tergites 3&amp;4 are typically longer, arcuate, lunulate, transverse or oblique, sometimes meeting medially, maculae of some species cross abdominal margin (Figs 7, 9 &amp; 17), in others they do not (Figs 5, 6, 8, 10–16); sternites light with dark maculae or fasciae, sometimes fascia absent on sternite 2; terminalia dark.</p><p>Male Genitalia: Hypandrium roughly trapezoid in shape, smooth, without pile; epandrium rounded ventrally with posterior projection (see Figs 6–17 &amp; 19), smooth, without pile; surstylus more or less triangular in shape in lateral view, flattened posteriorly or anteroposteriorly, long pile on dorsal, laterodorsal and posterodorsal sides, row of spines on posteromedial edge; cercus oval with long pile over entire surface; gonostylus variably shaped; basiphallus variable in shape, but roughly shaped like a faucet, posterodorsal edge bare or covered in pile or spines; distiphallus elongate, basal end flared out into a horn in most species, in D. albostriatus and D. eggeri it tapers to a point (Fig. 19), it is mostly sclerotized but membranous at apex of the horn, with either setae, pile or bare dorsally (Figs 6–17).</p><p>Etymology: From the Greek, dasys, meaning thick with hair, hairy, shaggy; and syrphos (masculine), meaning a kind of fly.</p><p>Distribution: Throughout Nearctic and Palaearctic (excluding North Africa). Throughout the Neotropical and Oriental regions, however there is low diversity in these regions. Not recorded from the Afrotropical or Australasian regions (Figs 20–22).</p><p>Key to Species of Nearctic Dasysyrphus Enderlein 1938</p><p>1. Tergites 3 &amp; 4 with maculae that extend over the abdominal margin, usually reaching the edge of the abdomen (cf. Fig. 7D).. .................................................................................................. 11.</p><p>1'. Tergites 3 &amp; 4 with maculae that do not reach the abdominal margin (cf Fig. 8D).................................. 2.</p><p>2. Gena dark in colour (dark brown to black) (cf Fig. 8D)....................................................... 5.</p><p>2'. Gena light in colour (yellow to light brown), sometimes with a black vitta between face and gena (cf Fig. 16C).......... 3.</p><p>3. Tergites 3 &amp; 4 with maculae arcuate, medial edge flattened, usually broadly contacting the anterior edge of the tergite (Figs</p><p>6A–D)................................................................................... creper (Snow) Western (southern AB &amp; BC, south to CA &amp; NM; Fig. 20A); Snow 1895: 234.</p><p>3'. Tergites 3 &amp; 4 with maculae oblique, more straight, with medial edge rounded and usually meeting medially, sometimes contacting anterior edge of tergite, but not as broadly (cf Fig. 16A)................................................ 4.</p><p>4. Gena separated from face by black vitta; wing cells br and bm almost entirely bare (Figs 16A–C)............................................................................................ richardi Locke &amp; Skevington sp. nov. Southwestern (NM; Fig. 20C); Locke &amp; Skevington, here described.</p><p>4'. Gena and face continuously yellow, not separated by black vitta; wing cells br and bm partly microtrichose (Figs 10A–D).......................................................................................... lotus (Williston) Southwestern (CA &amp; AZ, south to Mexico; Fig. 21A); Williston 1887: 75.</p><p>5. Sternite 2 with dark oval macula (Figs 15A–D, 18C)............................................ pinastri (DeGeer) Greenland (Fig. 21D); De Geer 1776: 113, Pl. 7, Figs 1–7.</p><p>5'. Sternite 2 with dark fascia (Fig. 18D)..................................................................... 6.</p><p>6. Tergites 3 &amp; 4 with maculae not greatly constricted, more or less uniform in thickness (cf Fig. 13A)................... 8.</p><p>6'. Tergites 3 &amp; 4 with maculae greatly constricted medially, sometimes completely divided (Figs 5A, 12A &amp; B)........... 7.</p><p>7. Male surstylus with a prominent posterolateral edge. Cell br usually with bare areas above base of spurious vein (sometimes very small); tergites 3 &amp; 4 with maculae usually divided (Figs 12A–E)........... occidualis Locke &amp; Skevington sp. nov. Western (AK, south to CO; Fig. 21B); Locke &amp; Skevington, here described.</p><p>7'. Male surstylus without a prominent posterolateral edge. Cell br densely microtrichose; tergites 3 &amp; 4 with maculae either divided or not (Figs 5A–D).......................................................... amalopis (Osten Sacken) Northern (YK, east to northern NL; NH; Fig. 20A); Osten Sacken 1875: 148.</p><p>8. Tergite 2 with anterolateral corner of maculae extending anterolaterally into a point, in females reaching the edge of the abdomen; profemur light, usually entirely so but some dark basally (Figs 8A–E)....................... laticaudus (Curran) Eastern and boreal (AK, east to NL, south to NH, Fig. 20C); Curran 1925: 175, Pl. XI, Fig. 151.</p><p>8'. Tergite 2 with anterolateral corner of maculae not extending anterolaterally into a point, lateral edges of maculae usually rounded (cf Fig. 13B); profemur dark basally, light apically to entirely dark (cf Fig. 14B)............................ 9.</p><p>9. Size 5.5–7.7 mm, dark flies; profemur ½–entirely dark; pubescent band on female frons complete but sometimes not well defined; males with tergite 2 maculae typically reduced and positioned toward the lateral edge; tergites 3 &amp; 4 uniform in width and slightly constricted medially; gonostylus boot-shaped with projection on posterodorsal surface that projects posterodorsally (Figs 11A–E, 18B)................................................................ nigricornis (Verrall) Northern (AK, east to northern NL; Greenland; Fig. 21A); Verrall 1873: 251.</p><p>9'. Combination of characters not as above. Size 5.0– 9.7 mm, profemur never entirely dark (¼–⅔ dark basially); pubescent band on female frons interrupted; males with tergite 2 maculae large (¼–½ width of tergite); tergites 3 &amp; 4 uniform in width and slightly constricted medially; gonostylus boot-shaped with projection on posterodorsal surface that projects more dorsally (Fig. 14D) or is shaped differently (Fig. 13D).................................................................. 10.</p><p>10. Male gonostylus with narrow, wavy projection on posterodorsal surface. Wing typically largely bare basally, with cells br and bm with large bare areas (Figs 13A–D)...................................................... pacificus (Lovett) Western, (southern BC, south to CA; Sierra Nevada, Cascades and Costal Mountain Ranges; Fig. 21C); Lovett (in Cole &amp; Lovett) 1919: 245.</p><p>10'. Male gonostylus with projection on posterodorsal surface straight; wing typically densely microtrichose, sometimes with bare areas in cells r 1, br, bm and/or c (Figs 14A–D)............................................... pauxillus (Williston) Western (AK, south to NM; Rocky Mountain Range; Fig. 21D); Williston 1887: 74.</p><p>11. Tergite 2 with maculae very small, &lt;¼ width of tergite; tergites 3 &amp; 4 with maculae transverse, usually narrow, with slight swelling on medial edge and narrowing towards margin (Figs 9A–D)................................. limatus (Hine) Widespread (AK, east to NS, south to CO and MA; Fig. 20D); Hine 1922: 146.</p><p>11'. Tergite 2 with maculae large,&gt; ¼ width of tergite; tergites 3 &amp; 4 with maculae distinctly arcuate, lunulate or oblique, with at least slight swelling on medial edge, sometimes constricted medially (Figs 7A, C, E &amp; 17A)........................ 12.</p><p>12. Tergites 3 &amp; 4 with maculae distinctly arcuate or lunulate, with anterior edge forming a distinct U or V shape, maculae are distinctly constricted, sometimes dividing in half with a large swelling on the medial edge (Figs 7A–F)............................................................................................. intrudens complex (Osten Sacken) Widespread (AK, east to NL, south to CA, NM and TN; Fig. 20B); Osten Sacken 1877: 326.</p><p>12'. Tergites 3 &amp; 4 with maculae are more oblique, no distinctive constriction in the centre, never divided and not greatly swollen on medial edge (Figs 17A–D)............................................................. venustus (Meigen) Widespread (AK, east to NS, south to NM &amp; NC; Fig. 22); Meigen 1822: 299.</p><p>Taxonomy</p></div>	https://treatment.plazi.org/id/03893F32FFA8FFBDFF46FF70FE8A8830	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FFAFFFBEFF46F8BEFA0C8DF3.text	03893F32FFAFFFBEFF46F8BEFA0C8DF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus amalopis (Osten Sacken 1875)	<div><p>Dasysyrphus amalopis (Osten Sacken)</p><p>Figures: 5A–D, 20A</p><p>Synonyms and References:</p><p>Syrphus amalopis Osten Sacken 1875: 148 (original description)</p><p>Curran 1925: 179 (discussion, amalopis s.l.)</p><p>Metasyrphus amalopis (Osten Sacken):</p><p>Fluke 1933: 110 (in part; key, redescription and transfer to Metasyrphus)</p><p>Dasysyrphus amalopis (Osten Sacken):</p><p>Stone et al. 1965 (in part): 563 (catalogue and transfer to Dasysyrphus)</p><p>Vockeroth 1986: 202 (lectotype designation and discussion)</p><p>Vockeroth 1992: 63 (in part; redescription)</p><p>Diagnosis: Abdominal maculae do not meet abdominal margin; maculae on tergites 3&amp;4 are constricted medially and usually not divided (Figs 5A &amp; B). Wing cells are densely microtrichose (Fig. 5D). Surstylus without a prominent posterolateral edge (Fig. 5C). Similar to D. occidualis, but D. occidualis has maculae on tergites 3&amp;4 either divided or undivided (Figs 12A &amp; B), cell br usually with a small bare area above base of spurious vein and a prominent posterolateral edge on the surstylus (Fig. 12D).</p><p>Redescription:</p><p>Body Length: 7.8–10mm; Wing Length: 7.7–8.7mm</p><p>Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in females it is continuous, ¼–½ the length between the antennae and ocelli) and dark pile; face light with dark vitta ¼–⅓ the width of the face sometimes reaching antennal socket, face with dark pile; gena dark with pile light to dark; occiput dark, covered in light pollen, pile light; scape, pedicel and flagellomere dark.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile varying from all dark to mixed to all light, more light anteriorly and posteriorly; scutellum light (dark yellow) to dark with darker lateral edges, light pile along anterior edge and dark pile elsewhere; wing completely densely microtrichose (Fig. 5D); haltere light; femora ⅔–¾ dark basially, ¼–⅓ light apically, tibiae dark with very basal end light, tarsii dark.</p><p>Abdomen: Maculae on tergite 2 large and oval, usually with slight narrowing medially, ⅓–&lt;½ the width of tergite 2, maculae on tergites 3 &amp; 4 are oblique, constricted medially and usually not divided, posterior edge of maculae straight, maculae do not reach abdominal margin (Fig. 5B); sternite 2 light with a dark triangular macula, usually not connecting with the lateral edges of the sternite, sternites 3 &amp; 4 are light with dark fasciae.</p><p>Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened posteriorly, long pile on dorsal, laterodorsal and posterodorsal sides, row of spines on posteromedial edge; cercus oval with long pile over entire surface; gonostylus claw-shaped, pile on dorsal surface; basiphallus elongate, basally bent at about a 90 degree angle towards the anterodorsum, apical end curved ventrally with a widened apex and is partly membranous dorsally, with spines on posterodorsal side, apex; distiphallus elongate, apical end flared out into horn, which is mostly sclerotized but membranous at apex with minute setulae dorsally, basally bent towards dorsum and slightly enlarged ventrally at bend (Fig. 5C).</p><p>Intraspecific variation: From the specimens we have had the chance to observe the maculae on tergites 3 &amp; 4 are usually not divided (Fig. 5A). Sometimes only the maculae on tergite 3 are divided and sometimes they are divided on both tergites 3 &amp; 4. The lectotype male has entire maculae, whereas the paralectotype female has divided maculae.</p><p>Etymology: From the Greek, amalos meaning soft, tender, weak and – opis, meaning having the appearance of, like.</p><p>Distribution: Northern Canada (MB, NL, QC, YK) and Northeastern USA (NH) (Fig. 20A)</p><p>Ecology: Collected from June–July at elevations from sea level– 1006m.</p><p>Discussion: In addition to the two specimens selected as lectotype and paralectotype by Vockeroth (1986), Osten Sacken (1875) discussed a third specimen and included it in his description of the species. This third specimen has abdominal maculae that cross the abdominal margin. This third specimen is not a specimen of D. amalopis but a specimen of the D. intrudens complex, and has caused much confusion over the name D. amalopis . Vockeroth (1986) pointed out that Fluke (1933) discussed many specimens of D. amalopis with maculae reaching the margin. Vockeroth stated that he believed those specimens belonged to his concept of D. venustus . We argue that specimens described as D. amalopis that have maculae that cross the abdominal margin belong to the D. intrudens complex. Fluke (1933) does have D. amalopis key out in two places in his key. The D. amalopis keyed out in couplet 54 pertains to D. amalopis s.s. and D. occidualis, while the D. amalopis keyed out in couplet 56 pertains to the D. intrudens complex.</p><p>The lectotype and paralectotype are the only known specimens of this species from eastern North America south of northern Quebec and Labrador. Our new species, Dasysyrphus occidualis, has been confused with D. amalopis . However, the COI sequences and male genitalia characters provide evidence for these being two distinct species with an identical habitus. Dasysyrphus amalopis ranges from southern Yukon to northern Labrador (Fig. 20A), while D. occidualis ranges from Alaska, south to Colorado (Fig. 21B). Specimens referred to as D. amalopis from western Canada or the US are most likely that of D. occidualis and in the overlap zone in southern Yukon male genitalia and/or COI sequences should be used to confidently identify specimens. Despite the long standing use of the name D. amalopis, the similar new species, D. occidualis, has been much more frequently collected.</p><p>It is not known why no other specimens are known from as far south as the lectotype and paralectotype. They were both collected in the White Mountains, in New Hampshire. The date of their collection is unknown, however their description by Osten Sacken was published in 1875, so they would have been collected previous to that. It is possible that they are rare and have thus not been collected again or that their habitat or climatic conditions have changed resulting in their extirpation. More collecting should be done in the Appalachians at high elevation to see if this species still exists that far south.</p><p>Type Material: “White Mts./ Austin”, “Osten/ Sacken/ Coll.”, red label “Type/ 4092”, “ Syrphus / amalopis/ Type Species O.S.”, yellow label “ LECTOTYPE / Syrphus / amalopsis O.S./ Desig. Thompson 1971”, “ Jan.–July 2003 / MCZ Image/ Database”, J. Skevington/ Specimen #/ 23785”, ♂, deposited in the Museum of Comparative Zoology, Harvard University (MCZ). Missing head. (examined)</p><p>Material Examined:</p><p>Paralectotype: United States of America: New Hampshire: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.17313&amp;materialsCitation.latitude=44.387787" title="Search Plazi for locations around (long -71.17313/lat 44.387787)">Gorham</a>, [44.387788, -71.17313], ♀, JSS 23786 (MCZ) . Other Material Examined: Canada: Manitoba: CHURCHILL REGION: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-94.129&amp;materialsCitation.latitude=58.4026" title="Search Plazi for locations around (long -94.129/lat 58.4026)">Weir</a>, [58.4026, -94.129], ♂, 25.vii.2007, ♀, 07 PROBE 4723 , L.E. Roscoe (CNC); Burn site <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.82889&amp;materialsCitation.latitude=58.61861" title="Search Plazi for locations around (long -93.82889/lat 58.61861)">North of Twin</a> lakes, [58.61861111, -93.82888889], 5.vii.2007, ♂, CNCD 9607 , J. Skevington (CNC); Twin Lakes, [58.63, -93.819], 14m, 19.vii.2009, ♀, 09 PROBE 176, (CNC), 26.vi.2010 – 2.vii.2010, ♂, 10 PROBE 13673 , 5.vii.2010 – 9.vii.2010, ♀, 10 PROBE 13805, J. Wang (USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-94.193&amp;materialsCitation.latitude=58.649" title="Search Plazi for locations around (long -94.193/lat 58.649)">Goose Creek</a>, [58.649, -94.193 W], 19.vii.2009, ♀, 09 PROBE 25 , A. Thielman (CNC); Farnworth Lake (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-94.0525&amp;materialsCitation.latitude=58.7045" title="Search Plazi for locations around (long -94.0525/lat 58.7045)">Landing</a> L), [58.7045, -94.0525], 14m, 14.vii.2010, ♀, 10 PROBE 14269 , 14.vii.2010 – 19.vii.2010, ♂, 10 PROBE 15042, 28.vi.2010, ♂, 10 PROBE 10909, 28.vi.2010 – 6.vii.2010, ♀, 10 PROBE 15283, J. Wang (CNC) . Newfoundland and Labrador: Nain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.689987&amp;materialsCitation.latitude=56.5429" title="Search Plazi for locations around (long -61.689987/lat 56.5429)">Anatalak Bay</a>, [56.5429, -61.689989], 16.vii.1928, ♂, JSS 19248, A.C. Weed (FMNH) . Quebec: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-64.72085&amp;materialsCitation.latitude=56.328484" title="Search Plazi for locations around (long -64.72085/lat 56.328484)">Indian House Lake</a>, [56.328482, -64.720845], 19.vii.1954; ♀, CNCD 13616 , 25.vii.1954, ♀, CNCD 13618,W. R. Richards (CNC); 22.vii.1954, ♀, CNCD 13617, R . Coyles (CNC) . Yukon Territory: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-132.33333&amp;materialsCitation.latitude=62.683334" title="Search Plazi for locations around (long -132.33333/lat 62.683334)">La Force Lake</a>, [62.683333, -132.333333], 1006m, 26.vi.1960, ♂, CNCD 13587 , J.E.H. Martin (CNC) .</p></div>	https://treatment.plazi.org/id/03893F32FFAFFFBEFF46F8BEFA0C8DF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FFACFF82FF46FCA3FF7A8A25.text	03893F32FFACFF82FF46FCA3FF7A8A25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus creper (Snow 1895)	<div><p>Dasysyrphus creper (Snow)</p><p>Figures: 6A–D, 20A</p><p>Synonyms and References:</p><p>? Syrphus lotus Williston 1887: 75 var. (short description of a variety of lotus)</p><p>Syrphus pauxillus Snow 1892: 37 (nec. Williston; short description)</p><p>Syrphus creper Snow 1895: 234 (original description)</p><p>Syrphus lotus Curran 1925: 92 Pl. VI Fig. 51 (nec. Williston; redescription)</p><p>Syrphus lotus var. creper Curran 1925: 180 (discussion)</p><p>Metasyrphus creper (Snow):</p><p>Fluke 1933: 107 (key, redescription and transfer to Metasyrphus)</p><p>Dasysyrphus creper (Snow):</p><p>Stone et al. 1965: 563 (catalogue and transfer to Dasysyrphus)</p><p>Vockeroth 1986: 202 (lectotype designation)</p><p>Vockeroth 1992: 65 (key and redescription)</p><p>Diagnosis: Abdominal maculae do not meet abdominal margin; maculae on tergites 3&amp;4 arcuate with medial edge flattened and usually broadly contacting anterior edge; markings unlike any other species (Figs 6A &amp; C).</p><p>Redescription:</p><p>Body Length: 6.7–10.5mm; Wing Length: 6.3–9.2mm</p><p>Head: Frons dark with light pollenose fascia (in male it runs along ventral edge of where eyes meet, in females it is ½ the length between the antennae and ocelli) and dark pile; face light with dark vitta ¼–⅓ the width of the face, usually not reaching antennal socket, face with light pile, dark pile lateral to the eye; gena light (Fig. 6C) with light pile and usually some dark pile below eye; occiput dark, covered in light pollen, pile light; scape and pedicel light to dark, flagellomere dark.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile light; scutellum light to dark with darker lateral edges, light pile along anterior edge and dark pile elsewhere; wing cells br and bm entirely bare, cell c bare from base to ¾, r 1 bare at base and cup bare at base and along vein CuP (Fig. 6D); haltere light; pro- and mesofemora ⅓–½ dark basially, ½–⅔ light apically, metafemur ⅔–¾ dark basially, ¼–⅓ light apically, tibiae light, metatibia with dark band ¼ of the way from the apex, tarsii light, sometime dark anteriorly.</p><p>Abdomen: Maculae on tergite 2 large and oval, maculae on tergites 3&amp;4 are arcuate, with medial edge widened, flattened and usually broadly contacting the anterior edge of tergite, sometimes meeting medially, maculae do not reach abdominal margin (Figs 6A &amp; C); sternites light with dark fasciae.</p><p>Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened posteriorly, long pile on dorsal, laterodorsal and posterodorsal sides, row of spines on posteromedial edge; cercus oval with long pile over entire surface; gonostylus boot-shaped with projection on posterodorsal surface that projects dorsally, pile on dorsal surface; basiphallus elongate, basally bent at 90 degree angle towards dorsum, apical end curved ventrally and is partly membranous with pile on dorsal side; distiphallus elongate, apical end flared out into horn, which is mostly sclerotized but membranous at apex with minute setulae dorsally, basally bent towards dorsum and slightly enlarged at bend (Fig. 6B).</p><p>Intraspecific variation: Scutellum colour varies from light to dark; maculae on tergites 3&amp;4 sometimes meet medially and sometimes are separated, medial edge usually broadly contacts anterior edge of tergite but if not then it comes close to anterior edge and edge of maculae are rounded; dark fasciae on sternites vary in width.</p><p>Etymology: The word creper is Latin, meaning dark or obscure.</p><p>Distribution: Southwestern Canada (AB, BC) and western USA (AZ, CA, CO, ID, MT, NM, OR, SD, UT, WA, WY). (Fig. 20A)</p><p>Ecology: Collected on ponderosa pine ( Pinus ponderosa Douglas ex Lawson &amp; C. Lawson, Pinaceae), sierra willow ( Salix orestera C.K. Schneid., Salicaceae), willow ( Salix Linnaeus, Salicaceae), chokecherry ( Prunus virginiana, Rosaceae), Ross’ avens ( Geum rossii (R. Brown) Seringe, Rosaceae), avens ( Geum Linnaeus, Rosaceae), tall ragwort ( Senecio serra Hooker, Asteraceae), threenerve goldenrod ( Solidago velutina DC., Asteraceae), rabbitbush ( Chrysothamnus Nuttall, Asteraceae). Habitats listed on labels include “along timberline”, “creek margin” and “on tundra”. The specimen collected on the “tundra” was collected in Colorado at 3505m, so this is in reference to an alpine tundra habitat. Collected April to October, although much more frequently collected in July and August. Collected from elevations of 305m – 3688m, although most frequently collected from elevations above 2000m.</p><p>Discussion: Presumably Williston (1887) was referring to a specimen of D. creper when he described the female from New Mexico as a second specimen in his description of lotus . He noted there were differences but that it was “perhaps allied” with the female from Arizona that he described as lotus . He describes the abdominal maculae of this second specimen as “more oblique, and strongly concave in front” and “more convex behind” and the gena as having a “black stripe”, which are characters more befitting of Snow’s description of creper, which came some seven years later.</p><p>Three years before Snow formally described creper in 1895 he described specimens as pauxillus (Snow 1892) . In his description of creper he synonymizes his 1892 pauxillus description with creper . Snow is also the one who suggests Williston’s (1887) variety of lotus is probably creper .</p><p>In Curran’s monograph (Curran 1925) he demotes S. creper from the rank of species to a variety of S. lotus . He claimed that there were no differences between the two except for the amount of interruption in the second and third abdominal bands. He believed it was just an extreme in coloration and did not warrant the rank of species. However, in the same paper he provides a redescription of lotus and based on the characters discussed and illustrated (Plate VI: 51) it is clear that his concept of lotus is not the same as Williston’s and he is actually describing creper . In 1933, Fluke re-elevated creper to the rank of species. Fluke states that at that time lotus was only known from the type and that published records of lotus probably refer to creper . Fluke also notes that Curran was probably looking at specimens of creper, not lotus .</p><p>Fluke (1933) clearly distinguishes between these two species based on the abdominal markings, with lotus having more straight bands and those of creper being more arcuate. Fluke also places both species into Metasyrphus and into the amalopis group with other species currently recognized as Dasysyrphus .</p><p>Type Material: “Magdalena,/ New Mexico ”, “Type”, blank red label, red label “ Syrphus / creper Snow / LECTOTYPE / Des. Vockeroth/ 1984”, “Jeff Skevington/ Specimen #/ 23782”; ♂; deposited in the Snow Entomological Museum, University of Kansas. (examined)</p><p>Material Examined:</p><p>Paralectotypes: United States of America: Colorado: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.52137&amp;materialsCitation.latitude=40.376877" title="Search Plazi for locations around (long -105.52137/lat 40.376877)">Estes Park</a>, [40.376877, -105.52137], viii.1892, ♂, JSS 20203, F.H. Snow (FMNH), ♂, OSUC 319691 , F.H. Snow (OSU) . Other Material Examined: Canada: Alberta: Elkwater, Hwy 41, [49.65, -110.2666667], 7.vii.1982, ♀, JSS 18293, B. V . Peterson (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.67352&amp;materialsCitation.latitude=50.04048" title="Search Plazi for locations around (long -110.67352/lat 50.04048)">Medicine Hat</a>, Division No. 1, [50.040483, -110.673523] , 15.v.1930, ♂, JSS 19477, F.S. Carr (CNC). Kananaskis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.08769&amp;materialsCitation.latitude=51.10088" title="Search Plazi for locations around (long -115.08769/lat 51.10088)">Forest Experimental Station</a> Seebe, [51.100879, -115.087692] , 15.vi.1968, ♂, CNCD 13634, H.J. Teskey, 20.vi.1950, ♀, CNCD 13643 (CNC). BANFF NATIONAL PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.58348&amp;materialsCitation.latitude=51.149437" title="Search Plazi for locations around (long -115.58348/lat 51.149437)">Sulphur Mountain</a>, [51.149438, -115.583484], 2195m, 28.vii.1967, 2♂, CNCD 13635, 13636 , J. R. Vockeroth (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.568436&amp;materialsCitation.latitude=51.180275" title="Search Plazi for locations around (long -115.568436/lat 51.180275)">Banff</a>, [51.180275, -115.568433] , 27.vi.1925, ♀, JSS 19442, O. Bryant (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.80862&amp;materialsCitation.latitude=51.60524" title="Search Plazi for locations around (long -115.80862/lat 51.60524)">Snow Creek Pass</a>, [51.60524, -115.808617], 2256m , 28.vii.1962, ♀, CNCD 13642, 13.viii.1962, ♀, CNCD 13644, K.C. Herrmann (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.50892&amp;materialsCitation.latitude=51.717632" title="Search Plazi for locations around (long -116.50892/lat 51.717632)">Peyto Lookout</a>, [51.717631, -116.508919], 2438m , 23.vii.1962, ♂, CNCD 13630, ♀, CNCD 13641, K.C. Herrmann (CNC) . British Columbia: CATHEDRAL PROVINCIAL PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.2073&amp;materialsCitation.latitude=49.055786" title="Search Plazi for locations around (long -120.2073/lat 49.055786)">Glacier Lake</a>, [49.055785, -120.207299] , 6.vii.1986, ♂, CNCD 13631, R . A. Cannings (CNC); 7.vii.1986, ♂, RBCM ENT 9881939, ♀, RBCM ENT 9881942, R . A. Cannings (RBCM); Quiniscoe Mtn. summit, [49.08, -120.18] , 26.vii.1988, ♂, JSS 20170, S.G. Cannings (UBCZ). OKANAGAN- SIMILKAMEEN R . D.: Manning Provincial Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.766846&amp;materialsCitation.latitude=49.07995" title="Search Plazi for locations around (long -120.766846/lat 49.07995)">Valleyview</a>, [49.07995, -120.766842], 1830m , 14.vii.1986, ♂, JSS 18368, S.G. Cannings (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.76692&amp;materialsCitation.latitude=49.098667" title="Search Plazi for locations around (long -120.76692/lat 49.098667)">Blackwall Mountain</a>, [49.098666, -120.766925], 1828m , 7.viii.1953, ♂, CNCD 13632, D.F. Hardwick (CNC); Osoyoos, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.6647&amp;materialsCitation.latitude=49.111595" title="Search Plazi for locations around (long -119.6647/lat 49.111595)">Mount Kobau</a>, [49.111594, -119.664702] , 9.vii.1968, ♂, CNCD 13627, ♀, CNCD 13640, D. Allen, 13.viii.1967, ♂, CNCD 13628 , J. R. Vockeroth (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.550446&amp;materialsCitation.latitude=49.18234" title="Search Plazi for locations around (long -119.550446/lat 49.18234)">Oliver</a>, [49.182338, -119.550442], 305m , 23.viii.1953, ♀, CNCD 13637, J.E.H. Martin, 14.ix.1923, ♀, CNCD 13638 , C.B. Garrett (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.07596&amp;materialsCitation.latitude=49.357784" title="Search Plazi for locations around (long -120.07596/lat 49.357784)">Hedley</a>, [49.357784, -120.075962] , 7.viii.1953, ♂, CNCD 13633, D.F. Hardwick (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.934456&amp;materialsCitation.latitude=49.482487" title="Search Plazi for locations around (long -119.934456/lat 49.482487)">Apex Mountain</a>, [49.482486, -119.934458], 1900m , 15.vii.1988, ♂, RBCM ENT 9885672, R . A. Cannings (RBCM). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.69783&amp;materialsCitation.latitude=49.34203" title="Search Plazi for locations around (long -117.69783/lat 49.34203)">Robson</a>, [49.342029, -117.697831] , 22.vi.1959, ♀, CNCD 13639, H. R. Foxlee (CNC). Cranbrook, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.79115&amp;materialsCitation.latitude=49.497196" title="Search Plazi for locations around (long -115.79115/lat 49.497196)">Elizabeth Lake</a>, [49.497195, -115.791152] , 6.vii.1980, CNCD 60067, R . A. Cannings (CNC). MOUNT REVELSTOKE NATIONAL PARK: Mount Harry, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.14251&amp;materialsCitation.latitude=51.03598" title="Search Plazi for locations around (long -118.14251/lat 51.03598)">First Summit</a>, [51.035979, -118.142509], 2316m, 20.viii.1952, ♂, CNCD 13629 , G.P. Holland (CNC) . United States of America: Arizona: Rustler Park: w. Portal, [31.90556, -109.27917], 2560m, 2.viii.1977, ♀, USNM ENT 258093 . COCHISE CO.: [31.906851, - 109.278543], 2560m, 13.ix.1965, ♀, JSS 19475, C.W. Sabrosky (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-112.40389&amp;materialsCitation.latitude=34.414722" title="Search Plazi for locations around (long -112.40389/lat 34.414722)">Bradshaw Mountains</a>, [34.41472222, - 112.4038889], 1951m , 7.v.1999, ♂, JSS 20683, R . S. Beal (CSUC). COCONINO CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.753044&amp;materialsCitation.latitude=34.961452" title="Search Plazi for locations around (long -111.753044/lat 34.961452)">Oak Creek Canyon</a>, [34.961452, -111.753046], ♀, CNCD 26378 , F.H. Snow (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.52876&amp;materialsCitation.latitude=34.973633" title="Search Plazi for locations around (long -111.52876/lat 34.973633)">Weimer Spring</a>, [34.973632, -111.528763] , 4.vi.1963, ♂, JSS 19445, C.H. Spitzer (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.67088&amp;materialsCitation.latitude=35.340305" title="Search Plazi for locations around (long -111.67088/lat 35.340305)">Flagstaff</a>, [35.340306, -111.670881], 3505m , 14.viii.1934, ♂, JSS 19435, ♀, JSS 19436, E.L. Bell (CNC). GRAND CANYON: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-113.226776&amp;materialsCitation.latitude=36.17849" title="Search Plazi for locations around (long -113.226776/lat 36.17849)">North Rim</a> [36.17849, -113.226775] , 17.viii.1966, JSS 25081, W.J. Hanson (EMUS). South Arizona , viii.1902, 2♀, CNCD 26379, JSS 19474, C.W. Johnson (CNC); viii.1902, ♂, JSS 21342, ♀, JSS 21340, F.H. Snow (MCZ). California: RIVERSIDE CO.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.16378&amp;materialsCitation.latitude=33.69554" title="Search Plazi for locations around (long -117.16378/lat 33.69554)">Quaking Aspen</a>, [33.69554, -117.163778] , 5.vii.1984, ♂, JSS 18296, D.J. Burdick (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.6792&amp;materialsCitation.latitude=33.81449" title="Search Plazi for locations around (long -116.6792/lat 33.81449)">San Jacinto Mountains</a>, [33.814491, - 116.679201], ♀, JSS 19459, J.M. Aldrich (CNC). LOS ANGELES CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.43063&amp;materialsCitation.latitude=34.05612" title="Search Plazi for locations around (long -118.43063/lat 34.05612)">Westwood</a>, [34.056121, -118.430635], 17.vi.1959, ♀, CNCD 26374 , Kelton &amp; Madge (CNC). SAN BERNARDINO CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.02093&amp;materialsCitation.latitude=34.1003" title="Search Plazi for locations around (long -117.02093/lat 34.1003)">Upper Santa Ana River</a>, [34.1003, - 117.020931] , 7.vii.1957, 3♂, JSS 19466–19468, 2♀, JSS 19438, 19465, A.L. Melander (CNC). YOSEMITE NATIONAL PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.637695&amp;materialsCitation.latitude=37.673916" title="Search Plazi for locations around (long -119.637695/lat 37.673916)">Glacier Point Road</a>, [37.673917, -119.637698] , 1.vii.1947, ♀, JSS 19464, A.L. Melander (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.57777&amp;materialsCitation.latitude=37.74195" title="Search Plazi for locations around (long -119.57777/lat 37.74195)">Yosemite</a>, [37.741951, -119.577767], 1183m , 26.v.1931, ♂, JSS 20584, (EMEC). TUOLUMNE CO .: Mather Site, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.85698&amp;materialsCitation.latitude=37.880173" title="Search Plazi for locations around (long -119.85698/lat 37.880173)">Buck Meadows</a>, [37.880173, -119.856977], 1970, ♂, CNCD 26352 , ♀, CNCD 26375, A. R. Moldenke (CNC). MONO CO .: Tioga Pass, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.25507&amp;materialsCitation.latitude=37.91939" title="Search Plazi for locations around (long -119.25507/lat 37.91939)">Hall Area</a> site, [37.919391, -119.255072], 26.vii.1969, 2♀, CNCD 26376, 26377 , A. R. Moldenke (CNC). CALAVERAS BIG TREES SP: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.257774&amp;materialsCitation.latitude=38.261665" title="Search Plazi for locations around (long -120.257774/lat 38.261665)">Beaver Creek</a>, [38.26166667, -120.2577778], 1375m , 22.v.– 11.vi.2007, 3♂, JSS 20954–20956, ♀, JSS 20951, A. R . Cline &amp; P.H. Kerr (MHPC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.226944&amp;materialsCitation.latitude=38.849445" title="Search Plazi for locations around (long -120.226944/lat 38.849445)">Wright’s Lake</a>, [38.84944444, -120.2269444], 2120m , 20.vi.2006, ♂, JSS 20962, M. Hauser, S. Winterton, &amp; E. Fisher (MHPC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.59917&amp;materialsCitation.latitude=39.32306" title="Search Plazi for locations around (long -120.59917/lat 39.32306)">Yuba Pass</a>, [39.323059, -120.599169], 2042m , 9.vii.1954, ♀, JSS 20589, R . H. Goodwin (EMEC). WHITE MOUNTAINS: Blanco’s Corral, 3048m , 30.vi.1953, ♀, JSS 20590, J.W. MacSwain (EMEC). Truckee, 25.vi.1943, 3♂, JSS 23769, 23771, 23772, ♀, JSS 23770, P.H. Arnaud (USNM). S. CA ., v.1908, 4♂, JSS 20585–20588, R . Woglum (EMEC) . Colorado: SAN JUAN CO .: 10 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-107.48503&amp;materialsCitation.latitude=37.809566" title="Search Plazi for locations around (long -107.48503/lat 37.809566)">Miles East of Silverton</a>, [37.809565, -107.485032], 2743m, 13.viii, ♂, CNCD 26351 , F.M. Carpenter (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-108.309525&amp;materialsCitation.latitude=37.941105" title="Search Plazi for locations around (long -108.309525/lat 37.941105)">Bennet Crk.</a>, [37.941103, -108.309527] , 7.ix.1933, ♀, JSS 20707 (CSUC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.29212&amp;materialsCitation.latitude=38.55185" title="Search Plazi for locations around (long -106.29212/lat 38.55185)">Garfield</a>, [38.55185, -106.292119], 3658m , 5.viii.1971, ♀, USNM ENT 258077, W.E. Steiner (USNM). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-107.592&amp;materialsCitation.latitude=38.86832" title="Search Plazi for locations around (long -107.592/lat 38.86832)">Paonia</a>, [38.86832, - 107.592002] , 16.ix.1912, JSS 20711, Chas. L. Fluke (CSUC). LAKE CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.55732&amp;materialsCitation.latitude=39.10472" title="Search Plazi for locations around (long -106.55732/lat 39.10472)">Independence Pass</a>, [39.104722, - 106.557321], 3688m, 31.vii.1961, ♂, CNCD 26354 , B.H. Poole (CNC); Tennessee Pass, [39.357246, - 106.313889], 3131m, 7.vii, ♀, JSS 19458, J.M. Aldrich (CNC). Tenmile Range, unamed <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.111&amp;materialsCitation.latitude=39.425" title="Search Plazi for locations around (long -106.111/lat 39.425)">Cr.</a> avalanche chute W side, [39.425, -106.111], 3200m , 15.vii.–17.vii.1998, ♂, JSS 20682, S. Fitzgerald &amp; A. Foley (CSUC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.61778&amp;materialsCitation.latitude=39.61582" title="Search Plazi for locations around (long -105.61778/lat 39.61582)">Rogers Pk.</a>, [39.615821, -105.61778], 3660m , 12.vii.1997, ♂, JSS 20805, 10.viii.2002, ♀, JSS 20806, C. Slater (CSUC). ARAPAHO NATIONAL FOREST, MT. EVANS: Mount Evans, [39.25, -106.1666667], 3658m, 3.viii.1961, ♂, CNCD 26355 , ♀, CNCD 26368, S.M. Clark (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.594894&amp;materialsCitation.latitude=39.645576" title="Search Plazi for locations around (long -105.594894/lat 39.645576)">Timberline</a>, [39.645575, -105.594897], 3566m , 22.vii.1961, ♀, CNCD 26367, B.H. Poole (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.60477&amp;materialsCitation.latitude=39.659767" title="Search Plazi for locations around (long -105.60477/lat 39.659767)">Echo Lake</a>, [39.659767, -105.604767], 3176m , 29.vi.2010, 2♂, CNCD 60033– 60034, ♀, JSS 24626, M.M. Locke (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.60135&amp;materialsCitation.latitude=39.67574" title="Search Plazi for locations around (long -105.60135/lat 39.67574)">Doolittle Ranch</a>, [39.675739, -105.601348], 2987m , 10.vii.1961, ♀, CNCD 26364, 3.viii.1961, ♀, CNCD 26365, W. R. M. Mason (CNC). BOULDER CO .: Nederland, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.53389&amp;materialsCitation.latitude=40.031647" title="Search Plazi for locations around (long -105.53389/lat 40.031647)">Science Lodge</a>, [40.031645, -105.533886], 2896m, 1.vii.1961, ♂, CNCD 26357 , A.A. Bucknell, 28.vi.1961, ♀, CNCD 26361 , W. R. M. Mason, 6.vii.1961, ♀, CNCD 26362 , 1.vii.1961, ♀, CNCD 26363, B.H. Poole (CNC); Niwot Ridge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.6074&amp;materialsCitation.latitude=40.069935" title="Search Plazi for locations around (long -105.6074/lat 40.069935)">Near Ward</a>, [40.069933, -105.607397], 3505m , 4.vii.1961, ♂, CNCD 26356, ♀, CNCD 26366, C.H. Mann, 7.vii.1965, ♀, JSS 19446, B. Petersen (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.50833&amp;materialsCitation.latitude=40.07221" title="Search Plazi for locations around (long -105.50833/lat 40.07221)">Ward</a>, [40.072208, -105.508332] , 9.viii.1913, ♂, JSS 20712, (CSUC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.52137&amp;materialsCitation.latitude=40.376877" title="Search Plazi for locations around (long -105.52137/lat 40.376877)">Estes Park</a>, [40.376877, -105.52137] , 8.viii.1931, ♀, JSS 20669, 20.v.1933, ♀, JSS 20671, (CSUC), viii.1892, ♂, JSS 21335, F.H. Snow (MCZ). ROCKY MOUNTAIN NATIONAL PARK, Fall R . entrance, [40.40344, -105.589203], 4.vi.1999, ♀, JSS 20684, D. Leatherman, 1.viii.1997, ♀, JSS 20685, D. Leatherman, Manhattan, W. Rustic (CSUC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.21005&amp;materialsCitation.latitude=40.48717" title="Search Plazi for locations around (long -105.21005/lat 40.48717)">Masonville</a>, [40.487171, -105.210056] , 5.ix.1933, ♀, JSS 20674, (CSUC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.22767&amp;materialsCitation.latitude=40.520817" title="Search Plazi for locations around (long -118.22767/lat 40.520817)">Cameron Pass</a>, [40.520816, - 118.227668] , 7.vii.1931, ♀, JSS 20709, (CSUC); 3048m, vii, ♂, JSS 19981, (FMNH). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.59778&amp;materialsCitation.latitude=40.561092" title="Search Plazi for locations around (long -105.59778/lat 40.561092)">Pingree Park</a>, [40.561094, - 105.597778] , 20.viii.1924, ♀, JSS 20710, C.L. Fluke, 19.viii.1932, ♂, JSS 20668, C.M. Drage, (CSUC), 14.viii.1934, ♂, JSS 23764, C.W. Sabrosky (USNM). LORY STATE PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.191666&amp;materialsCitation.latitude=40.572994" title="Search Plazi for locations around (long -105.191666/lat 40.572994)">Well Gulch Trail</a>, [40.572996, - 105.191669] , 18.vi.1995, ♀, JSS 20809, D. Leatherman (CSUC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.083336&amp;materialsCitation.latitude=40.583332" title="Search Plazi for locations around (long -105.083336/lat 40.583332)">Fort Collins</a>, [40.58333333, -105.0833333] , 19.ix.1942, ♀, JSS 20743, R .F. duChanois, 5.x.1921, ♀, JSS 20676, 19.ix.1932, ♀, JSS 20680, 21.ix.1932, ♀, JSS 20697, 20.ix.1932, ♀, JSS 20698, 8.ix.1913, ♀, JSS 20742, 30.ix.1942, ♀, JSS 20703, (CSUC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.853645&amp;materialsCitation.latitude=40.603664" title="Search Plazi for locations around (long -105.853645/lat 40.603664)">Chambers Lake</a>, [40.603663, -105.853645] , 25.vi.1932, ♂, JSS 20672, 2♀, JSS 20675, 20701, C. R . Jones (CSUC). North Park, [40.73, -106.18] , 1.vii.1932, 3♂, JSS 20695, 20696, 20705, 6♀, JSS 20670, 20673, 20677, 20678, 20699, 20700, (CSUC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.9355&amp;materialsCitation.latitude=40.8078" title="Search Plazi for locations around (long -105.9355/lat 40.8078)">Glendevey</a>, [40.8078, -105.935501] , 7.vii.1932, ♀, JSS 20679, 3.vii.1932, ♂, JSS 20704, C. R . Jones, iv.1931, ♀, JSS 20708, (CSUC). HOMER, 7.ix.1934, ♀, JSS 20706, (CSUC). Boulder Canyon, 2377m , 8.viii.1960, ♀, JSS 21113, R .&amp; K. Dreisbach (MSUC). Colorado State, 4♂, JSS 19450–19453, 2♀, JSS 19454, 19455 (CNC). Idaho: CUSTER CO .: 4th of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-114.67202&amp;materialsCitation.latitude=44.39658" title="Search Plazi for locations around (long -114.67202/lat 44.39658)">July Creek</a>, [44.39658, -114.672022], 27.vii.1975, JSS 25139, T . Griswold (EMUS). LATAH CO: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.86897&amp;materialsCitation.latitude=46.80726" title="Search Plazi for locations around (long -116.86897/lat 46.80726)">Mountains Near Moscow</a>, [46.807258, -116.868969], 9.vii.1911, ♂, CNCD 26358 , ♀, CNCD 26373, F.M. Hull, 8.vii.1911, ♀, JSS 19470, 5.vii.1912, ♂, JSS 19472, 6.vii.1912, ♀, JSS 19471, (CNC). Craig’s Mountain, 29.iv.1916, ♀, JSS 21330, (MCZ). Holbrook, 4 mi. N.W ., 16.v.1969, JSS 25080, G.F. Knowlton &amp; G.E. Bohart (EMUS). Montana: MISSOULA CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-113.99427&amp;materialsCitation.latitude=46.871403" title="Search Plazi for locations around (long -113.99427/lat 46.871403)">Missoula</a>, [46.871403, -113.994267] , 22.viii.1904, ♀, JSS 19473, (CNC). New Mexico: DONA ANA CO.: Las Cruces, [32.3, -106.7666667], 2♂, CNCD 26359, 26360 , 2♀, CNCD 26380, 26381, F.M. Hull (CNC). LINCOLN CO .: White Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.67296&amp;materialsCitation.latitude=33.33165" title="Search Plazi for locations around (long -105.67296/lat 33.33165)">North Fork Ruidoso</a>, [33.331649, -105.672958], 2499m , viii.1920, ♂, JSS 19440, Townsend (CNC). SAN MIGUEL CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.34231&amp;materialsCitation.latitude=35.659687" title="Search Plazi for locations around (long -105.34231/lat 35.659687)">Trout Spring Canyon</a>, [35.659688, - 105.34231] , iv.1927, ♀, JSS 19449, (CNC). Truchos Peak, [35.974028, -105.636663], 4.viii, ♀, JSS 19456, W.P. Cockerell (CNC). Embudo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.96113&amp;materialsCitation.latitude=36.207523" title="Search Plazi for locations around (long -105.96113/lat 36.207523)">Rio Arriba</a>, [36.207523, -105.961132] , 25.ix.1897, ♀, JSS 19441, (CNC). Las Valles, Gallinas River 3505m, 20.iv, ♀, JSS 19437, E.L. Bell (CNC). Oregon: KALMOTH CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.17272&amp;materialsCitation.latitude=42.92532" title="Search Plazi for locations around (long -122.17272/lat 42.92532)">Crater Lake National Park</a>, [42.925321, -122.172722], 10.vii.1968, ♀, JSS 18297, B. V . Peterson (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.54227&amp;materialsCitation.latitude=44.060387" title="Search Plazi for locations around (long -121.54227/lat 44.060387)">Sisters</a>: 15 mi S, [44.060385, - 121.542268] , 20.vii.1976, ♂, USNM ENT 258064, George Steyskal (USNM). WHITMAN NATIONL FOREST: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.23057&amp;materialsCitation.latitude=44.958805" title="Search Plazi for locations around (long -118.23057/lat 44.958805)">Anthony Lake</a>, [44.958805, -118.23057], 2164m, 28.vii.1954, ♀, OSAC 133387 , J.H. Baker (OSAC), ♂, JSS 23767, 3♀, JSS 23765, 23766, 23768, J.H. Baker (USNM), 4.viii.1929, ♀, OSAC 133388, H.A. Scullen (OSAC). South Dakota: LAWRENCE CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-103.8866&amp;materialsCitation.latitude=44.589397" title="Search Plazi for locations around (long -103.8866/lat 44.589397)">Spearfish Creek</a>, [44.589399, -103.886595] , 14.vi.1969, ♂, JSS 19476, W.W. Wirth (CNC). HARDY W.C.: T3 N, R1 E, S30, 3.vii.1965, ♂, CNCD 5445, R . W. Hodges (USNM). Utah: GARFIELD CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.71268&amp;materialsCitation.latitude=38.028416" title="Search Plazi for locations around (long -110.71268/lat 38.028416)">Henry Mountains</a>, [38.028415, -110.712679], 2286m , 29.vii.1968, ♂, JSS 18295, J.E.H. Martin (CNC). WAYNE CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.176956&amp;materialsCitation.latitude=38.177814" title="Search Plazi for locations around (long -111.176956/lat 38.177814)">Capitol Reef National Park</a>, [38.177815, -111.176958] , 3.x.1984, JSS 25137, W.P. Nye (EMUS). DUCHESNE CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-112.36749&amp;materialsCitation.latitude=38.31073" title="Search Plazi for locations around (long -112.36749/lat 38.31073)">Mirror Lake</a>, [38.31073, -112.367492], 2743m , 21.vii.–9.viii.1977, JSS 25135, 2.viii.1971, JSS 25138, G.F. Knowlton (EMUS). BOX ELDER CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.97376&amp;materialsCitation.latitude=41.3941" title="Search Plazi for locations around (long -111.97376/lat 41.3941)">Willard Basin</a>, [41.394101, -111.973766], 2743m , 26.viii.1964, JSS 25134, W.J Hanson (EMUS). CACHE CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.5641&amp;materialsCitation.latitude=41.8866" title="Search Plazi for locations around (long -111.5641/lat 41.8866)">Tony Grove Creek</a>, [41.886599, -111.564102] , 10.viii.1976, JSS 25136, (EMUS); Green Canyon M. T ., 31.viii.2003 – 25.viii.2003, ♀, JSS 24611, W.J. Hanson &amp; Brammer (EMUS). DAGGETT CO .: Rt. 44, 2438m, 10.vii.1962, ♀, JSS 21110, R .&amp; K. Dreisbach (MSUC). SUMMIT CO .: Henry’s Fork Pk., 2926m, 1.viii.1979 – 10.viii.1979, ♀, CNCD 5807 , S. &amp; J. Peck (CNC). Washington: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.493904&amp;materialsCitation.latitude=46.204174" title="Search Plazi for locations around (long -121.493904/lat 46.204174)">Mount Adams</a>, [46.204174, -121.493902] , 3.viii.1930, ♀, JSS 19460, F.P. Dean (CNC). Mt. Ranier, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.72795&amp;materialsCitation.latitude=46.782707" title="Search Plazi for locations around (long -121.72795/lat 46.782707)">White River</a>, [46.782709, - 121.727952] , 20.vii.1924, ♂, JSS 23763, A.L. Melander (USNM). SPOKANE CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.37438&amp;materialsCitation.latitude=47.842392" title="Search Plazi for locations around (long -117.37438/lat 47.842392)">Buckeye</a>, [47.842393, - 117.374385] , 21.vi.1930, ♂, JSS 19444, J.M. Aldrich (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.11165&amp;materialsCitation.latitude=47.92371" title="Search Plazi for locations around (long -117.11165/lat 47.92371)">Mount Spokane</a>, [47.923711, -117.111646] , 22.vi.1930, ♀, JSS 19457, 27.vi.1930, ♀, JSS 19448, 2.vii.1930, ♀, JSS 19447, J.M. Aldrich (CNC). Wawawai, 28.v.1922, ♂, JSS 23762, A.L. Melander (USNM). Wyoming: ALBANY CO ., [41.727896, -105.684712], 2.viii.1927, ♂, CNCD 26353, C.L. Corkins (CNC). Camp Guernsey, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-104.72862&amp;materialsCitation.latitude=42.260113" title="Search Plazi for locations around (long -104.72862/lat 42.260113)">Ryan Springs</a>, [42.260113, -104.728624] , 31.v.2005, ♂, JSS 20686, B. Kondratieff, J. Schmidt, J. Owens (CSUC). YELLOWSTONE NATIONAL PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.80875&amp;materialsCitation.latitude=44.550877" title="Search Plazi for locations around (long -110.80875/lat 44.550877)">Jet Lake</a>, [44.550878, -110.808744] , 16.viii.1927, ♀, JSS 19439, J.M. Aldrich (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.537674&amp;materialsCitation.latitude=44.58772" title="Search Plazi for locations around (long -110.537674/lat 44.58772)">Canyon Village</a>, [44.587719, - 110.537673] , 21.vii.1971, ♂, CNCD 5444, G.C. Steyskal (USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.48622&amp;materialsCitation.latitude=44.73326" title="Search Plazi for locations around (long -110.48622/lat 44.73326)">Canyon Campground</a>, [44.733262, - 110.486225] , 12.viii.1918, ♂, JSS 19461, ♀, JSS 19462, A.L. Melander (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.434364&amp;materialsCitation.latitude=44.797714" title="Search Plazi for locations around (long -110.434364/lat 44.797714)">Mount Washburn</a> [44.797716, - 110.434368], ♂, JSS 21343, C. T . Brues (MCZ). Centennial: 23.viii.1951, 2♀, JSS 21109, 21111, R. R . Dreisbach (MSUC); Snowy Range, 3048m , 8.ix.1950, 2♀, JSS 21105, 21114, R. R . D. &amp; R.K. Schwab, 23.viii.1951, ♂, JSS 21108, 2♀, JSS 21106-21107, R. R . Dreisbach (MSUC). Wyoming, 1952, 4♀, CNCD 26369–26372 , F.M. Hull (CNC) .</p></div>	https://treatment.plazi.org/id/03893F32FFACFF82FF46FCA3FF7A8A25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FF90FF87FF46FA6BFF6189A0.text	03893F32FF90FF87FF46FA6BFF6189A0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus intrudens (Osten Sacken 1877)	<div><p>Dasysyrphus intrudens species complex (Osten Sacken)</p><p>Figures: 7A–F, 20B</p><p>Synonyms and References:</p><p>Syrphus amalopis Osten Sacken 1875: 148 (in part, description)</p><p>Syrphus intrudens Osten Sacken 1877: 326 (original description)</p><p>Syrphus amalopis Osten Sacken:</p><p>Williston 1887: 69 (in part; redescription)</p><p>Syrphus disgregus Snow 1895: 233 (original description; comb. nov.)</p><p>Curran 1925: 180 (discussion)</p><p>Syrphus laticaudatus Curran 1925: 176, Pl. XI Fig. 152 (original description; comb. nov.)</p><p>Syrphus osburni Curran 1925: 177, Pl. XI Fig. 153 (original description; comb. nov.)</p><p>Metasyrphus amalopis (Osten Sacken):</p><p>Fluke 1933: 110 (in part, includes disgregus and intrudens; key, redescription and transfer to Metasyrphus)</p><p>Metasyrphus laticaudatus (Curran):</p><p>Fluke 1933: 111 (key, redescription and transfer to Metasyrphus)</p><p>Metasyrphus osburni (Curran):</p><p>Fluke 1933: 112 (key, redescription and transfer to Metasyrphus)</p><p>Dasysyrphus amalopis (Osten Sacken):</p><p>Stone et al. 1965: 563 (in part; includes intrudens, catalogue and move to Dasysyrphus)</p><p>Dasysyrphus disgregus (Snow):</p><p>Stone et al. 1965: 563 (catalogue and move to Dasysyrphus)</p><p>Dasysyrphus laticaudatus (Curran):</p><p>Stone et al. 1965: 563 (catalogue and move to Dasysyrphus)</p><p>Dasysyrphus osburni (Curran):</p><p>Stone et al. 1965: 564 (catalogue and move to Dasysyrphus)</p><p>Dasysyrphus venustus (Meigen):</p><p>Vockeroth 1986: 203 (in part, includes disgregus, intrudens, laticaudatus and osburni; lectotype designation for disgregus and intrudens and discussion)</p><p>Vockeroth 1992: 70 (in part, includes intrudens, disgregus, laticaudatus and osburni; key and redescription)</p><p>Diagnosis: Abdominal maculae cross abdominal margin; maculae on tergite 2 large (&gt;⅓ width of tergite) ovalshaped; maculae on tergites 3&amp;4 arcuate or lunulate, usually strongly constricted medially, sometimes dividing maculae in half, with medial edge swollen, never meeting medially, maculae reach abdominal margin, usually broadly contacting edge of abdomen (Figs 7A–E). Has previously been confused with D. venustus, however, species in this complex have maculae that are arcuate, lunulate and/or greatly constricted medially, whereas in D. venustus they are more transverse and much less constricted medially (Fig. 17A).</p><p>Redescription:</p><p>Body Length: 7.0– 11.7mm; Wing Length: 7.0– 10.7mm</p><p>Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in females it is ¼–½ the length between the antennae and ocelli and narrowly to widely separated medially) and dark pile; face light with dark vitta ¼–½ the width of the face, usually reaching antennal socket, face with dark or light pile, sometimes with a both light and dark pile; gena dark with light pile; occiput dark, covered in light pollen, pile light; scape and pedicel light to dark, flagellomere usually light basally and darker apically, sometimes mostly dark.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile light; scutellum light with darker anterolateral edges, light pile some with varying amounts of dark pile posteriorly; cell c usually densely microtrichose but sometimes bare basally, r 1 is usually bare at the base, but can be entirely microtrichose, br and bm range from densely microtrichose to extensively bare, but often with some bare areas, cup either densely microtrichose or bare at base and/or along CuP; haltere light; pro- and mesofemora ⅓–½ dark basally, ½–⅔ light apically, metafemur ⅔–¾ dark basally, ¼–⅓ light apically, tibiae light, metatibia and rarely pro- and/or mesotibiae with dark band ¼ of the way from the apex, tarsi light, sometimes dark anteriorly.</p><p>Abdomen: Maculae on tergite 2 large and oval, sometimes with anterolateral edge extending to edge of abdomen, maculae on tergites 3&amp;4 arcuate or lunulate, usually strongly constricted medially, sometimes dividing maculae in half, with medial edge swollen, never meeting medially, maculae reach abdominal margin, usually broadly contacting edge of abdomen (Figs 7A–E); sternite 2 either all light, with dark oval macula or with dark fascia (usually distinct, sometimes faint), sternites 3&amp;4 light with dark fasciae.</p><p>Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened anteroposteriorly with long pile on posterior side and spines on anteroventral side, similar to D. venustus (Fig. 17D) and D. limatus (Fig. 9D); cercus oval with long pile over entire surface; gonostylus is pointed ventrally, dorsally flattened at base then it expands dorsally towards apex, oblique ridge near apex, with ends of ridge projecting into points; pile on dorsal surface; basiphallus is basally bent at about a 90 degree angle towards the dorsum, at apical end it is curved ventrally, fully sclerotized all the way around with no pile or spines; distiphallus is long with no enlarged area basally, shaft sometimes with small bump dorsally near base, apical end flared out into horn with the dorsal side flattened slightly, which is mostly sclerotized but membranous at apex with setulae dorsally (Fig. 7F).</p><p>Intraspecific variation: The abdominal maculae are highly variable (Figs 7A–E), likely because multiple, unresolved species lie within this complex. Tergites 3 &amp; 4 have maculae that can be arcuate or lunulate, usually strongly constricted medially, sometimes dividing maculae in half, medial edge swollen but degree of swelling varies, maculae usually broadly contact edge of abdomen, but in few specimens they end just past margin and do not contact edge; sternite 2 colouring variable, either all light, with dark oval macula or with dark fascia (usually distinct, sometimes faint); wing cells vary in microtrichia density with c usually densely microtrichose but sometimes bare basally, r 1 is usually bare at the base, but can be entirely microtrichose, br and bm range from densely microtrichose to extensively bare, but often with some bare areas, cup either densely microtrichose or bare at base and/or along CuP; distiphallus shaft sometimes with small bump dorsally near base.</p><p>Etymology: Presumably comes from the Latin, intrudo, meaning enter without invitation or permission.</p><p>Distribution: Canada (AB, BC, NF, NS, ON QC, YT) and US (AK, AZ, CA, CO, ID, MA, MD, ME, MI, NC, NH, NM, NY, OR, PA, TN, UT, VA, WA, WV, WY), also widespread over Europe and Asia (Fig. 20B)</p><p>Ecology: Collected on great camas ( Camassia leichtlinii (Baker) S. Watson, Liliaceae), common camas ( Camassia quamash (Pursh) Greene, Liliaceae), Carolina spring beauty ( Claytonia caroliniana Michaux, Portulacaceae), small-flowered woodland-star ( Lithophragma parviflorum (Hooker) Nuttall ex Torrey &amp; A. Gray, Saxifragaceae), heart-leaved foamflower ( Tiarella cordifolia Linnaeus, Saxifragaceae), birch ( Betula Linnaeus, Betulaceae), common cowparsnip ( Heracleum maximum W. Bartram, Apiaceae), cowparsnip ( Heracleum Linnaeus, Apiaceae), blackberry ( Rubus Linnaeus, Rosaceae), Pacific blackberry ( Rubus ursinus Chamisso &amp; Schlechtendal, Rasaceae), Pacific silver fir ( Abies amabilis (Douglas ex Loudon) Douglas ex J. Forbes, Pinaceae), Labrador tea ( Ledum Linnaeus, Ericaceae), spotted-water hemlock ( Cicuta maculata Linnaeus, Apiaceae), blueberry ( Vaccinium Linnaeus, Ericaceae), dogwood ( Cornus Linnaeus, Cornaceae), jack pine ( Pinus banksiana Lambert, Pinaceae), mountain maple ( Acer spicatum Lamarck, Aceraceae), nannyberry ( Viburnum lentago Linnaeus, Caprifoliaceae), beachgrass ( Ammophila Host, Poaceae), herb-Robert ( Geranium robertianum Linnaeus, Geraniaceae), slender phlox ( Microsteris gracilis (Hooker) Greene, Polemoniaceae), spreading groundsmoke ( Gayophytum diffusum subsp. parviflorum F.H. Lewis &amp; Szweykowski, Onagraceae), chokecherry ( Prunus virginiana, Rosaceae), bristly bearberry ( Arctostaphylos columbiana Piper, Ericaceae), slender stitchwort ( Minuartia tenella (J. Gay) Mattfeld, Caryophyllaceae), hairy cat’s ear ( Hypochaeris radicata Linnaeus, Asteraceae). Habitats listed on labels include “ Ainus– Populus – Picea ” or “Alder–Poplar–Spruce”, “along creek in forested ravine”, “along roadside”, “ Betula woods”, “birch &amp; fir”, “black spruce forest”, “canopy gap in hardwood forest”, “dry sand beach ridge”, “gap in Mh/Be forest”, “hardwood forest”, “hemlock forest”, “hot springs”, “in clearing near river”, “ Ledum - Kalmia bog”, “low vegetation and leaf litter at campsite”, “marshy meadow”, “on tide flats”, “ Picea forest”, “pine forest”, “ Pinus - Salix forest”, “riverbank scrub”, “ Salix-Picea ”, “scrub poplar and low vegetation”, “seepage area”, “fen”, “sphagnum bog”, “ Vaccinium wood” and “wet, scrubby clearing”. This species complex is also collected from hilltops. Collected January to November, although much more frequently collected in May to August. Collected from elevations from sea level– 4000m.</p><p>Discussion: Dasysyrphus intrudens is most certainly a complex of multiple species. However, the boundaries between these species are still unclear. The COI clusters produced in the NJ tree (Fig. 4) yielded no clues to the number of species existing in this clade. When sorting the specimens by barcodes, no consistant morphological patterns were visible. Sometimes characters were similar between two or more clusters and sometimes within one cluster there were multiple character states. As the barcoding region of COI appears to be of little help to sort this complex into species, other markers should be sequenced to look for insight into species boundaries. Fresh material collected into absolute alcohol will be required to sequence the necessary genes.</p><p>Note that this group has been referred to as the friuliensis group in conference abstracts, but never in a refereed publication. The oldest group name, intrudens, best serves as the name for the complex.</p><p>Type Material: “Lagunitas Creek, Cal./ April 15. O. Sacken ”, “Osten/ Sacken/ Coll.”, red label “Type/ 875”, yellow label “ LECTOTYPE / Syrphus / intrudens O.S./ Desig. Thompson 1977”, “ Syrphus / intrudens O.S.”, J. Skevington / Specimen #/ 23783”, ♂, deposited in the Museum of Comparative Zoology, Harvard University (MCZ). Both basoflagellomeres and right metaleg missing. Genitalia have been removed at some point and are not associated with the specimen (presumed to be lost). (examined)</p><p>Material Examined: There were 2138 specimens from throughout the Nearctic (Fig. 20B) that were examined for this study. Because this is a complex and species boundaries need to be resolved, it is of little use to list all information for material examined. Database records for each specimen examined may be obtained from the authors.</p></div>	https://treatment.plazi.org/id/03893F32FF90FF87FF46FA6BFF6189A0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FF95FF8BFF46F8E8FE1E8CF0.text	03893F32FF95FF8BFF46F8E8FE1E8CF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus laticaudus (Curran 1925)	<div><p>Dasysyrphus laticaudus (Curran)</p><p>Figures: 8A–E, 20C</p><p>Synonyms and References:</p><p>Syrphus laticaudus Curran 1925: 175, Pl. XI Fig. 151 (original description)</p><p>Metasyrphus laticaudus:</p><p>Fluke 1933: 117 (key, redescription and transfer to Metasyrphus)</p><p>Dasysyrphus laticaudus:</p><p>Stone et al. 1965: 563 (catalogue and transfer to Dasysyrphus)</p><p>Vockeroth 1986: 203 (synonymized under pauxillus)</p><p>Vockeroth 1992: 68 (key and redescription)</p><p>Diagnosis: Maculae oblique and fairly uniform in thickness. Like D. pauxillus, D. pacificus and D. nigricornis, maculae on tergites 3&amp;4 do not reach edge of abdominal margin (Figs 8A, B &amp; D). Dasysyrphus laticaudus differs from the other three in that the markings on tergite 2 in the female extend anterolaterally to the edge of the abdomen (Fig. 8B) and in the male they extend similarly, sometimes only slightly, but usually do not reach the edge (Fig. 8D). Maculae on tergites 3&amp;4 usually come very close to the margin, if not narrowly touching it, but never crossing it (Figs 8A, B &amp; D).</p><p>Redescription:</p><p>Body Length: 5.0– 8.2mm; Wing Length: 4.8–7.8mm</p><p>Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in females it is ⅓–½ the length between the antennae and ocelli with a separation medially and edges triangular to rounded) and dark pile, some females with light pile; face light with dark vitta ¼–⅓ the width of the face, usually coming close but not reaching antennal socket, face with dark pile lateral to the eye, shorter in females; gena dark with light to dark pile; occiput dark, covered in light pollen, pile light; scape and pedicel light to dark, flagellomere usually dark with light base.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile variable, from all light to variable to all dark; scutellum light to dark with darker lateral edges, pile on female mostly light with some dark, male typically with dark pile, but sometimes with some light pile anteriorly; wings densely microtrichose, some with small bare areas in br above the base of the spurious vein (Fig. 8E); haltere light; profemur ¼–⅓ dark basially, ⅔–¾ light apically, mesofemur ⅓–½ dark basially, ½–⅔ light apically in males, pro-and mseofemora of females are light, base may be slightly darkened (Fig. 8B), metafemur ⅔–¾ dark basially, ¼–⅓ light apically, tibiae light, metatibia with dark band ¼ of the way from the apex, sometimes extending to apex, tarsii light, sometimes dark anteriorly.</p><p>Abdomen: Maculae on tergite 2 large and oval, in female the anterolateral edge reaching the edge of the tergite (Fig. 8B), in males there is a small projection but it terminates before the edge (Fig. 8D), maculae on tergites 3&amp;4 are oblique fairly straight and of uniform thickness, maculae do not reach abdominal margin (Figs 8A, B &amp; D); sternites light with dark fasciae of varying thickness.</p><p>Male Genitalia: Surstylus more or less triangular in lateral view, flattened posteriorly, long pile on dorsal, laterodorsal and posterodorsal sides, row of spines on posteromedial edge; cercus oval with long pile over entire surface; gonostylus boot-shaped but slightly more narrow apically than basally, with projection on posterodorsal surface, projecting dorsally, pile on dorsal surface; basiphallus elongate, basally bent at a slightly more than 90 degree angle towards dorsum, apical end curved ventrally, but does not greatly extend ventrally and is partly membranous dorsally, with spines on posterodorsal side; distiphallus elongate, apical end flared out into horn, which is mostly sclerotized but membranous at apex, with no setulae dorsally, basally bent towards dorsum and usually greatly enlarged at bend with a triangular projection, projecting ventrally (Fig. 8C). Extremely similar to D. pauxillus (Fig. 14D).</p><p>Intraspecific Variation: Female maculae on tergite 2 tend to be similar across individuals, however on males, the extent of projection on the anterolateral edge can vary. Male femora are always dark basally and light apically. Many females have predominantly light femora (especially the profemur; Fig. 8B). Females from the northwestern part of the range (Yukon) have femora that are dark basally and light apically. Wing cells are usually densely microtrichose, but sometimes have very small bare areas in cell br, above the spurious vein. Basal horn of distiphallus occasionally has few minute setulae dorsally.</p><p>Etymology: Comes from the Latin words latus, meaning broad and cauda, meaning tail.</p><p>Distribution: Eastern and northern Canada (MB, NB, NT, NS, NU, ON, QC, YK) and northeastern and northern USA (AK, CT, ME, MA, MI, NH, NY). (Fig. 20C)</p><p>Ecology: Collected from April–July, but more commonly in June and July. Collected at elevations of 792– 1067m. Habitats this species has been collected from, according to label data, are “Mh/Be forest” (mixed hardwood/beech), “hardwood forest”, “mixed forest”, “pine forest”, “coniferous woods”, “at top of limestone bluff” and “top of open S-facing bluff”. It has been recorded as being collected on pine trees ( Pinus Linnaeus; Pinaceae).</p><p>Discussion: Dasysyrphus laticaudus is hereby resurrected from synonymy with Dasysyrphus pauxillus (synonymized by Vockeroth (1986)). Vockeroth (1986) considered them to be the same species, presumably with only slight morphological differences between the species being mistaken for intraspecific variation. Dasysyrphus laticaudus is very similar to pauxillus (Figs 14A &amp; B) and pacificus (Figs 13A &amp; B), with the later two being extremely difficult to distinguish from each other. There are however, morphological and genetic characters that distinguish laticaudus from pauxillus and pacificus . Their distributions are also predictable and largely nonoverlapping.</p><p>Type Material: “Orillia Ont/ 5.v.1921 ”, “Collector/ H. Curran”, red label “ HoloTYPE / Syrphus / laticaudus/ CNC No. 497 Curr.”, “CNC Diptera / #15382”; ♂, deposited in the Canadian Collection of Insects, Arachnids and Nematodes (CNC), Ottawa, ON. (examined)</p><p>Material Examined:</p><p>Paratypes: Canada: Ontario: SIMCOE CO.: Orillia, [44.609505, -79.42068], 4.v.1921, ♀, CNCD 13851, 5.v.1921, ♂, CNCD 13885, H. Curran (CNC). Other Material Examined: Canada: Manitoba: Winnipeg, [49.88333333, -97.13333333], 20.vii.1948, ♂, JBWM 206845, W.H. Fell, 24.v.1949, ♀, JBWM 206517, C.F. Barrett (JBWM). New Brunswick: CHARLOTTE CO., [45.23102, -66.925304], 9.vi.1955, ♀, CNCD 13909, W. T.A. Neilson (CNC). YORK CO.: Fredericton, [45.959225, -66.640351], 3.v.1913, ♂, CNCD 13900, 5.vi.1958, ♂, CNCD 13901, (CNC). Sackville, [45.998, -64.3591], 7.v.2010, ♂, JK 969, J. Klymko, S.L. Robinson, D.M.Mazerolle (NBMB). KENT CO.: Kouchibouguac National Park, [46.819201, -64.96788], 19.v.1977, ♂, JSS 18365, 26.v.1977, ♀, CNCD 13908, W.P. Hanley, 20.v.1977, ♂, CNCD 13897, 24.v.1977, ♂, CNCD 13898, ♀, CNCD 13907, B. E. Cooper, 22.v.1977, ♂, CNCD 13899, 2♀, CNCD 13905, 13906, 23.v.1977, ♂, CNCD 106793, Hanley &amp; Cooper, 23.v.1977, ♂, JSS 18364, J.D. Lafontaine (CNC). Northwest Territories: Wholdaia Lake, [60.680702, -104.293128], 2.vii.–4.vii.1966, ♀, CNCD 13779, J.G. Chillcott (CNC). 21 miles east Tuktoyaktuk, [69.426844, -132.172475], 20.vi.1971 – 25.vi.1971, ♀, CNCD 13773, D.M. Wood (CNC). Nova Scotia: INVERNESS CO., Cape Breton Highlands National Park, Pleasant Bay, [46.822961, -60.799065], 9.vi.1984, CNCD 60066, 12.vi.1984, ♀, CNCD 13904, B.E. Cooper (CNC). Nunavut: Ford Lake, M. T.S. Gravity Survey Camp, [63.133386, -107.416561], 28.vi.1966, 3♂, CNCD 13716–13718, G.E. Shewell (CNC). Ontario: KAWARTHA LAKES DIV., Fenolon Falls, [44.538149, -78.735433], 27.v.1927, ♀, CNCD 13856, F.P. Ide (CNC). Chaffeys Locks, [44.56666667, -76.31666667], 15.v.1975, ♂, ROMEnt Spec. No. 115136, J.C.E. Riotte (ROME). HASTINGS CO., Maynooth, [45.229716, -77.940949], 25.v.1951, ♀, CNCD 13857, J.F. McAlpine, 22.v.1970, 2♂, CNCD 13883, 13884, D.M. Wood, 10.v.1987, CNCD 60346, D. Bell &amp; M. Wood (CNC). Griffith, 7 miles East, [45.243279, -77.031193], 31.v.1983, ♀, CNCD 13858, B.E. Cooper (CNC). ALGONQUIN PROVINCIAL PARK: Cranjelly Lake, [45.28719, -78.28012], 7.v.–21.v.2009, JSS 22433, 22435, 21.v.–4.vi.2009, JSS 22436, E. Proctor (CNC); Madawaska Lake, [45.3261, -78.3056], 12.v.–27.v.2007, 2♂, JSS 20747, 21358, 9.vi.–24.vi.2008, JSS 22430, 24717, 24718, 15.v.–29.v.2009, JSS 22434, 7.vi.–21.vi.2008, JSS 24719, 1.v.–15.v.2009, JSS 22431, E. Proctor (CNC); Crossbar Lake, [45.32686, -78.29997], 15.v.–29.v.2009, ♂, JSS 24614, 22432, E. Proctor (CNC); Florence Lake [45.44371, -78.49012], 13.v.–29.v.2007, ♂, JSS 21353, 19.v.–3.vi.2009, JSS 22437, E. Proctor (CNC); Sitting Duck Lake, [45.44985, -78.46831], 21.v.–21.v.2008, JSS 24643, E. Proctor (CNC); Cecil Lake, [45.4511, -78.4956], 6.vi.–20.vi.2008, JSS 25250, E. Proctor (CNC); Pondweed Lake, [45.46488, -78.43066], 6.vi.–19.vi.2008, JSS 23709, 26.v.–9.vi.2009, JSS 22441, 16.v.–28.v.2007, 2♀, JSS 21354, 21355, 9.v.–16.v.2007, 2♂, JSS 21356, 21357, E. Proctor (CNC); Lake of Two Rivers, [45.57177, -78.46515], 11.v.–25.v.2009, ♂, JSS 24613, 11.v.2009, JSS 22440, 11.v.2009 – 25.v.2009, JSS 22438, 22439, E. Proctor (CNC); Kearney Lake Campround, [45.57267, -78.439397], 11.vi.2005, ♀, JSS 19761, L. Bartels (CNC); Lake Sasajewun, [45.58, - 78.36], 19.v.1965, ♂, ROMEnt Spec. No. 115125, S.M. Smith (ROME); Brewer Lake, [45.58551, -78.31044], 18.v.2009 – 2.vi.2009, JSS 22442, E. Proctor (CNC); Algonquin Park, 11.vi.2003 – 18.vi.2003, ♂, ROMEnt Spec. No. 98519, ♀, ROMEnt Spec. No. 98518, Mark Vanderwel (ROME). Ottawa, Stack Road, [45.320852, - 75.736188], 7.v.2007, 6♂, CNCD 30882–30887, J. R. Vockeroth (CNC). Constance Bay, [45.495933, - 76.08694444], 18.iv.2010, ♂, JSS 19757, J. Skevington (CNC). COCHRANE DIST,: Low Bush, Lake Abitibi, [48.91608, -80.139852], 7.vi.1925, ♀, CNCD 13852, 9.vi.1925, ♀, CNCD 13853, 14.vi.1925, ♀, CNCD 13854, 16.vi.1925, ♀, CNCD 13855, N.K. Bigelow (CNC); 9.vi.1923, UASM # 149023, N.K. Bigelow (UASM); 6.vi.1925, BOC 34683, 7.vi.1925, ♀, BOC 34684, 9.vi.1925, 2♀, BOC 34685, 34686, 13.vi.1925, ♀, BOC 34687, 16.vi.1925, 2♀, BOC 34688, 34689, 24.vi.1925, ♀, BOC 34690, N.K. Bigelow (ROME); 14.vi.1925, ♀, JSS 21337, 15.vi.1925, ♀, JSS 21338, N.K. Bigelow (MCZ). Almonte, 18.v.1951, ♂, CNCD 106791, J.F. McAlpine (CNC). Quebec: OUTAOUAIS: Aylmer, [45.400224, -75.817137], 28.v.1923, 2♀, CNCD 13866, 13867, 29.v.1923, ♀, CNCD 13868, 16.v.1924, 5♂, CNCD 13887–13890, 106792, 2♀, CNCD 13864, 13865, 10.v.1925, ♂, CNCD 13886, C.H. Curran (CNC); Hull, [45.447639, -75.733192], 26.v.1923, 4♀, CNCD 13859–13862, 24.v.1923, ♀, CNCD 13863, 14.v.1924, 2♂, CNCD 13891, 13892, C.H. Curran (CNC); 27.v.1923, ♀, BOC 34691, C.H. Curran (ROME); Beechgrove, [45.64508, -76.123602], 15.v.1951, ♀, CNCD 13874, 3.v.1968, ♂, CNCD 13894, ♀, CNCD 13875, J.F. McAlpine (CNC); 27.v.1963, ♀, CNCD 13876, 15.v.1961, ♂, CNCD 13895, ♀, CNCD 13877, J. R. Vockeroth (CNC). VAUDREUIL- SOULANGES: Mount Rigaud, [45.466534, -74.325875], 13.v.2008, ♂, JSS 17647, J. Skevington, J. Gibson, A. Sugarman (CNC); 1.v.1986, ♂, JSS 18342, B.E. Cooper (CNC); 10.v.1986, 3♂, JSS 20763–20765, B.E. Cooper (CNC). Duncan Lake, near Rupert, [45.68138889, -76.05027778], 5.vi.1971, ♀, CNCD 13881, J.F. McAlpine (CNC). LE CENTRE- DE- LA- MAURICE: Lac Mondor, Near Ste. Flore, [46.625254, - 72.768842], 15.v.1951, ♀, CNCD 13869, 16.v.1951, ♂, CNCD 13893, ♀, CNCD 13870, 21.v.1951, 2♀, CNCD 13871, 13872, E.G. Munroe (CNC). LA HAUTE- GASPESIE: Mont Albert, [49.131813, -66.464367], 1067m, 1.vii.1954, ♂, CNCD 13896, W.J. Brown (CNC). Rupert River, [51.483333, -78.766667], 19.vi.1956, ♀, CNCD 13878, J. R. Lonsway (CNC). SEPT- RIVIERES- CANIAPISCAU: 37 km North West of Schefferville, [55.038335, - 66.424521], 12.vi.1981, ♀, CNCD 13880, F. Brodo (CNC). Great Whale River, [55.116816, -76.405554], 7.vii.1949, ♀, CNCD 13882, J. R. Vockeroth (CNC). Yukon Territory: 10 km North of Carcross on Highway 2, [60.2596, -134.749909], 5.vi.1981, ♀, JSS 18373, C.S. Guppy (CNC). Bluefish Ridge, [67.15, -140.6166667], 792m, 2.vii.1983, 2♀, JSS 20049, 20050, 7.vii.1983, ♀, JSS 20069, R.J. Cannings (UBCZ). 6 km East of Old Crow, [67.566667, -139.683333], 13.vii.1981, ♀, JSS 18366, C.S. Guppy (CNC). Old Crow, [67.571219, - 139.833877], 4.vii.1983, JSS 20071, R.A. Cannings (UBCZ); 13.vii.1981, CNCD 60075, C.S. Guppy (CNC). Firth River, [69.207056, -140.071033], 22.vi.1984, ♀, JSS 18375, S.G. Cannings (CNC). United States of America: Alaska: MATANUSKA- SUSITNA COUNTY: Matanuska, [61.541945, -149.229674], 12.vi.1944, ♀, JSS 19515, J. Chamberlin (CNC). Connecticut: NEW LONDON COUNTY: Lyme, [41.400831, -72.342951], 4.v.1918, ♂, JSS 19491, C. T. Greene (CNC). Maine: AROOSTOOK COUNTY: Crystal, [45.95953, -68.361018], vi.1980, ♂, CNCD 26441, S.N. Pease (CNC). Maryland: PRINCE GEORGE' S COUNTY: Laurel, [39.101761, -76.860489], 25.v.1965, ♂, CNCD 26728, (CNC). Massachusetts: WORCESTER COUNTY: Sunderland, [42.244545, -71.77175], 15.v.1965, ♂, JSS 19492, F.C. Thompson (CNC). HAMPSHIRE COUNTY: Amherst, [42.34038, -72.496818], 3.v.1963, 4♂, JSS 19493–19496, 7.v.1963, ♂, JSS 19497, 14.v.1963, ♂, JSS 19498, F.C. Thompson (CNC). Michigan: CRAWFORD CO.: 27.v.1958, ♀, JSS 21151, 21152, R. &amp; K. Dreisbach (MSUC). New Hampshire: CHESHIRE CO.: Jaffrey, [42.814957, -72.022399], 4.vi.1919, ♀, CNCD 26424, C.W. Johnson (CNC). Glen House, [44.289163, - 71.225764], 14.vi.1911, ♀, JSS 21339, N.K. Bigelow (MCZ). Mt. Monadnock, 11.vi.1932, ♀, JSS 23773, A.L. Melander (USNM). New York: TOMPKINS CO.: Ithaca, [42.444589, -76.499268], 11.v.1898, ♀, CNCD 26425, C.W. Johnson (CNC); ESSEX CO.: Corlear Bay, Lake Champlain, [44.483961, -73.418207], vi.1939, ♀, JSS 19499, R.C. Shannon (CNC).</p></div>	https://treatment.plazi.org/id/03893F32FF95FF8BFF46F8E8FE1E8CF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FF99FF8CFF46FBD8FC6F8A6B.text	03893F32FF99FF8CFF46FBD8FC6F8A6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus limatus (Hine 1922) Widespread	<div><p>Dasysyrphus limatus (Hine)</p><p>Figures: 9A–D, 20D</p><p>Synonyms and References:</p><p>Syrphus limatus Hine 1922: 146 (original description)</p><p>Curran 1925: 178, Pl. XI Fig. 154 (redescription)</p><p>Metasyrphus limatus:</p><p>Fluke 1933: 114 (key, redescription and transfer to Metasyrphus)</p><p>Dasysyrphus limatus:</p><p>Stone et al. 1965: 563 (catalogue and transfer to Dasysyrphus)</p><p>Vockeroth 1986: 202 (lectotype designation)</p><p>Vockeroth 1992: 66 (key and redescription)</p><p>Diagnosis: Abdominal maculae cross abdominal margin; maculae on tergite 2 very small oval-shaped; maculae on tergites 3&amp;4 transverse, narrow and slightly widened medially, narrow slightly towards margin (Fig. 9A). Similar to D. venustus (Fig. 17A) however the maculae on tergite 2 are always small in D. limatus, less than ¼ the width of the tergite, and are much larger in D. venustus, greater than ⅓ the width of the tergite.</p><p>Redescription:</p><p>Body Length: 7.5–10.9mm; Wing Length: 6.8–10.4mm</p><p>Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in female it is ½ or less the length between the antennae and ocelli, with a distinct separation medially) and dark pile; face light with dark vitta ⅓–½ the width of the face and reaches the antennal socket, face with dark pile, bare medially; gena dark with some light pollen ventral to eye, light and dark pile; occiput dark, covered in light pollen, pile light, some specimens with some dark pile; scape, pedicel and flagellomere dark.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile dark in males and light in female; scutellum light to dark with darker lateral edges, pile dark with some light pile along lateral edges to light with some dark on posterior edge; wing cell br bare basally and above spurious vein, bm bare on anterior ½–¾, r 1 bare at base and some with cup bare posterior to vein CuP (Fig. 9C); haltere light; pro- and mesofemora dark on basal half, light on apical half, metafemur dark with light apical end, pro- and mesotibiae light, metatibia dark with light basal end, tarsi dark, pro- and mesotarsi with light ventral side.</p><p>Abdomen: Maculae on tergite 2 small, slender and oval, &lt;¼ width of tergite, maculae on tergites 3&amp;4 are transverse, slender and widen medially, narrowed toward margin, positioned anteriorly on the tergite, maculae cross abdominal margin (Fig. 9A); sternites light with thick, dark fasciae.</p><p>Male Genitalia: Surstylus more or less oval in shape in lateral view, flattened with long pile on posterior side and spines on anteroventral side; cercus oval with long pile over entire surface; gonostylus triangular-shaped in lateral view, wide at base and elongated posterodorsal with small pointed projection on posterodorsal edge, posterior edge flares laterally, making it difficult to see in lateral view; pile on dorsal surface; basiphallus is greatly enlarged dorsally, fully sclerotized all the way around with no pile or spines; distiphallus is short with no enlarged area basally, apical end flared out into horn, which is mostly sclerotized but membranous at apex with no setulae dorsally (Fig. 9D).</p><p>Intraspecific variation: Maculae on abdomen can be of varying thickness, however they are usually quite narrow.</p><p>Etymology: The word limatus is Latin, meaning polished, refined.</p><p>Distribution: Canada (AB, BC, NT, NS, QC, YT), USA (AK, CO, MA, ME, NY, UT) (Fig. 20D)</p><p>Ecology: Collected on common cow parsnip ( Heracleum maximum). Collected from “sand dunes” and “outlet marsh” habitats. Collected April to September, although much more frequently collected from June through August. Collected from elevations of 320m – 3002m.</p><p>Discussion: The concept of limatus has been applied consistently since Hine erected the name in 1922.</p><p>Type Material: “Savonoski/ Naknek Lake / Alaska / July ‘19”, “ Jas. S. Hine / Collector”, red label “ Allotype ”, yellow label “ PARALECTOTYPE / Syrphus / limatus/ Hine”, barcode label “ OSUC 0373263 ”; ♂ ; deposited in the Ohio State University Collection (OSU). (examined)</p><p>Material Examined:</p><p>Paralectotype: United States of America: Alaska: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-155.2668&amp;materialsCitation.latitude=58.497288" title="Search Plazi for locations around (long -155.2668/lat 58.497288)">Savonoski. Naknek Lake</a>, [58.497288, -155.266794], vii.1919, ♀, OSUC 319689 , Jas. S. Hine (OSU); vii.1919, ♂, CNCD 87874, J. S. Hine (CNC) . Other Material Examined: Canada: Alberta: BANFF NATIONAL PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.58348&amp;materialsCitation.latitude=51.149437" title="Search Plazi for locations around (long -115.58348/lat 51.149437)">Sulphur Mountain</a>, [51.149438, -115.583484], 2195m, 28.vii.1967, ♀, CNCD 13699 , J. R. Vockeroth (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.568436&amp;materialsCitation.latitude=51.180275" title="Search Plazi for locations around (long -115.568436/lat 51.180275)">Banff</a>, [51.180275, -115.568433], 9.vi.1922, ♂, CNCD 13647 , C.B.D. Garrett (CNC); Mile 14 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.83851&amp;materialsCitation.latitude=51.231247" title="Search Plazi for locations around (long -115.83851/lat 51.231247)">Banff-Jasper Highway</a>, [51.231247, -115.838509], 3.viii.1955, ♀, CNCD 13686, R . Coyles (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.80862&amp;materialsCitation.latitude=51.60524" title="Search Plazi for locations around (long -115.80862/lat 51.60524)">Snow Creek Pass</a>, [51.60524, -115.808617], 2256m, 28.vii.1962, ♀, CNCD 13687 , W. R. M. Mason (CNC) . British Columbia: Vaseux L. Hills to E, [49.316685, -119.523082], 2.v.1983, ♂, JSS 20038, S.G. Cannings (UBCZ). CENTRAL KOOTENAY L.D.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.69287&amp;materialsCitation.latitude=49.333015" title="Search Plazi for locations around (long -117.69287/lat 49.333015)">Robson</a>, [49.333015, -117.692871], 6.v.1958, ♂, RBCM ENT 99188122 , 11.v.1966, ♂, Royal British Columbia M ENT 99188124, H. R . Foxlee (RBCM); 12.v.1956, ♂, JSS 20040, 17.vii.1955, ♀, JSS 20041, 28.iv.1958, ♀, JSS 20043, 12.ix.1952, ♀, JSS 20044, 13.v.1967, ♂, JSS 20088, 22.v.1956, ♂, JSS 20108, 27.vii.1959, ♂, JSS 20129, H. R . Foxlee (UBCZ); 15.v.1948, ♂, CNCD 13656, 11.v.1948, 2♂, CNCD 13657, 14473, H. R. Foxlee (CNC). Squamish, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.07434&amp;materialsCitation.latitude=49.811127" title="Search Plazi for locations around (long -123.07434/lat 49.811127)">Diamond Head Trail</a>, [49.811126, -123.07434], 975m, 9.viii.1953, ♀, CNCD 13683 , G.J. Spencer (CNC). MOUNT REVELSTOKE NATIONAL PARK: Mt. Revelstoke, [50.98333333, -118.2], 1.viii.1952, ♀, CNCD 533 , G.P. Holland (CNC); 13.viii.1952, ♀, CNCD 48545, G. J. Spencer (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.14251&amp;materialsCitation.latitude=51.03598" title="Search Plazi for locations around (long -118.14251/lat 51.03598)">Mount Harry</a>, [51.035979, -118.142509], 1890m, 20.vii.1952, ♀, CNCD 13682 , G.J. Spencer (CNC). CASSIAR L.D.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-132.12694&amp;materialsCitation.latitude=58.16401" title="Search Plazi for locations around (long -132.12694/lat 58.16401)">Moosehorn Lake</a>, [58.164009, -132.12694], 1372m, 1.viii.1960, ♀, CNCD 13678 , W.W. Moss (CNC); 1372m, 30.vii.1960, ♀, CNCD 13679, R . Pilfrey (CNC). NORTHERN ROCKIES R . D.: Summit Lake, Mile 392 of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.66691&amp;materialsCitation.latitude=58.6482" title="Search Plazi for locations around (long -124.66691/lat 58.6482)">Alaska Highway</a>, [58.648201, -124.666912], 1280m, 21.vii.1959, ♀, CNCD 13680 , 15.vii.1959, ♀, CNCD 13681, R . E. Leech (CNC). Liard R . hotsprings, [59.425991, -126.096398], 23.vii.1987, ♀, JSS 20046, S.G. Cannings (UBCZ). Atlin, [59.5775, -133.69236],.viii., ♀, CNCD 13684, (CNC). Campbell Mtn., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-108.24928&amp;materialsCitation.latitude=64.63313" title="Search Plazi for locations around (long -108.24928/lat 64.63313)">Pentiction</a>, 880m, 13.v.1983, ♂, JSS 20039, S.G. Cannings (UBCZ). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-108.24928&amp;materialsCitation.latitude=64.63313" title="Search Plazi for locations around (long -108.24928/lat 64.63313)">Northwest Territories</a>: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-108.24928&amp;materialsCitation.latitude=64.63313" title="Search Plazi for locations around (long -108.24928/lat 64.63313)">Muskox Lake</a>, [64.633135, -108.249285], 20.vii.1953, ♂, CNCD 13645 , J.G. Chillcott (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-132.17247&amp;materialsCitation.latitude=69.42684" title="Search Plazi for locations around (long -132.17247/lat 69.42684)">21 miles east Tuktoyaktuk</a>, [69.426844, -132.172475], 2–5.vii.1971, ♂, CNCD 13646 , 20–25.vi.1971, ♀, CNCD 13694, D.M. Wood (CNC). Nova Scotia: CAPE BRETON HIGHLANDS NATIONAL PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.81877&amp;materialsCitation.latitude=46.773975" title="Search Plazi for locations around (long -60.81877/lat 46.773975)">Mackenzie Fire Tower</a>, [46.773977, -60.818772], 8.vii.1983, ♂, CNCD 13665 , D.M. Wood (CNC). Quebec: Cté Missisquoi summit <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.51477&amp;materialsCitation.latitude=45.216717" title="Search Plazi for locations around (long -72.51477/lat 45.216717)">Mt Pinacle</a>, [45.018184, -72.855064], 700m, 20.vi.1991, ♂, JSS 20795, D.M. Wood (CNC). BROME- MISSISQUOI: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.51477&amp;materialsCitation.latitude=45.216717" title="Search Plazi for locations around (long -72.51477/lat 45.216717)">Knowlton</a>, [45.216716, -72.514769], 12.vi.1928, ♀, CNCD 13651 , W.J. Brown (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.326385&amp;materialsCitation.latitude=45.46639" title="Search Plazi for locations around (long -74.326385/lat 45.46639)">Summit of Mt. Rigaud</a>, [45.46638889, -74.32638889], 4.vi.1978, 4♂, CNCD 5809, 13659–13661 , D.M. Wood (CNC). LA VALLEE- DU- RICHELIEU: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.15527&amp;materialsCitation.latitude=45.552948" title="Search Plazi for locations around (long -73.15527/lat 45.552948)">Mont Saint-Hilaire</a>, [45.552948, -73.155276], 17.vi.1965, ♂, CNCD 13667 , 27.vi.1965, 4♂, CNCD 13668–13671, J.W. Boyes (CNC). TEMISCAMINGUE: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.26898&amp;materialsCitation.latitude=47.04583" title="Search Plazi for locations around (long -79.26898/lat 47.04583)">Laniel</a>, [47.045828, -79.268979], 10.vi.1931, ♀, CNCD 13688 , H.S. Fleming (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.066666&amp;materialsCitation.latitude=53.716667" title="Search Plazi for locations around (long -76.066666/lat 53.716667)">Sakami</a> (LG3), [53.71666667, -76.06666667], 13–18.vii.1977, 2♂, CNCD 5808, 13658 , D.M. Wood (CNC). Lac Duplanter, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.78209&amp;materialsCitation.latitude=54.883533" title="Search Plazi for locations around (long -69.78209/lat 54.883533)">Lac Delorme</a>, [54.883534, -69.78209], 7–12.vii.1977, 3♂, CNCD 13662–13664 , D.M. Wood (CNC). Kuujjuaq (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.40618&amp;materialsCitation.latitude=58.100075" title="Search Plazi for locations around (long -68.40618/lat 58.100075)">Fort Chimo</a>), [58.100076, -68.406179], 28.vii.1948, 2♀, CNCD 13689, 13690, R . H. MacLeod (CNC). Yukon Territory: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-134.6927&amp;materialsCitation.latitude=60.18669" title="Search Plazi for locations around (long -134.6927/lat 60.18669)">Carcross</a>, [60.186691, -134.692703], 16–18.vi.1982, ♂, CNCD 13666 , G. &amp; M. Wood (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-135.08209&amp;materialsCitation.latitude=60.733402" title="Search Plazi for locations around (long -135.08209/lat 60.733402)">Whitehorse</a>, [60.733402, -135.082092], 22.vii.1959, ♀, CNCD 13698, R . Madge (CNC). Haines Jct., [60.752472, -137.510006], 29.vi.1974, ♀, JSS 20181, G.G.E. Scudder (UBCZ). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-130.41667&amp;materialsCitation.latitude=62.486816" title="Search Plazi for locations around (long -130.41667/lat 62.486816)">Otter Lake</a>, [62.486817, -130.4166667], 1219m, 28.vii.1960, ♀, CNCD 13650 , 29.vii.1960, ♀, CNCD 13696, E.W. Rockburne (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-131.2666&amp;materialsCitation.latitude=62.616764" title="Search Plazi for locations around (long -131.2666/lat 62.616764)">Sheldon Lake</a>, [62.616763, -131.266603], 1067m, 3.vii.1960, ♂, CNCD 13654 , E.W. Rockburne (CNC); 3.vii.1960, ♀, CNCD 13915, 7.vii.1960, ♂, CNCD 13655, ♀, CNCD 13695, J.E.H. Martin (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-132.33333&amp;materialsCitation.latitude=62.683334" title="Search Plazi for locations around (long -132.33333/lat 62.683334)">La Force Lake</a>, [62.683333, -132.333333], 1006m, 27.vi.1960, ♂, CNCD 14010 , J.E.H. Martin (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-138.21666&amp;materialsCitation.latitude=64.51667" title="Search Plazi for locations around (long -138.21666/lat 64.51667)">Borth Fork Pass</a>, [64.51666667, -138.2166667], 1067m, 27.vii.1982, ♀, JSS 20045, S.G. Cannings, L. Vasington, R . A. Moore (UBCZ). North Fork Crossing, Mile 43 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-138.25064&amp;materialsCitation.latitude=64.56599" title="Search Plazi for locations around (long -138.25064/lat 64.56599)">Peel Pit Road</a>, [64.565983, -138.250648], 1067m, 6.vii.1962, ♂, CNCD 13652, R . E. Leech (CNC); 3.vii.1962, ♂, CNCD 13653, 7.vii.1962, ♀, CNCD 13697, P.J. Skitsko (CNC). DEMPSTER HIGHWAY: km 82, [64.581259, -138.261721], 1300m, 20.vii.1981 21.vii.1981, ♀, CNCD 13685, Lafontaine and G. &amp; M. Wood (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-138.12833&amp;materialsCitation.latitude=65.05472" title="Search Plazi for locations around (long -138.12833/lat 65.05472)">Mile</a> 87, [65.054721, -138.128324], 27–30.vi.1973, ♂, CNCD 13649 , 5–7.vii.1973, ♀, CNCD 13691, 16–17.vii.1973, ♀, CNCD 13693, 4–8.viii.1973, ♀, CNCD 13692, G. &amp; D.M. Wood (CNC); km 416, [66.644581, -136.324471], 750m, 22–28.vi.1980, 3♂, CNCD 13675–13677, Wood &amp; Lafontaine (CNC); km 465, [67.046392, -136.209668], 800m, 5–7.vii.1980, ♂, CNCD 13674, Wood &amp; Lafontaine (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-135.53334&amp;materialsCitation.latitude=66.041664" title="Search Plazi for locations around (long -135.53334/lat 66.041664)">Richardson Mts</a>, [66.04166667, -135.5333333], 1219m, 6.vii.1982, ♂, CNCD 5810, 762m, 7.vii.1982, 3♂, CNCD 532, CNCD 13672, 13673, D.M. Wood (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-140.08333&amp;materialsCitation.latitude=69.21667" title="Search Plazi for locations around (long -140.08333/lat 69.21667)">British Mts.</a>, [69.21666667, -140.0833333], 320m, 21.vi.1984 26.vi.1984, ♀, JSS 20774, G. &amp; M. Wood &amp; D. Lafontaine (CNC) . United States of America: Alaska: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-154.87694&amp;materialsCitation.latitude=58.03389" title="Search Plazi for locations around (long -154.87694/lat 58.03389)">Katmai</a>, [58.033889, -154.876944], viii.1917, ♀, CNCD 112302 , J. S. Hine (CNC). BRISTOL BAY CO.: Savonoski, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-156.86537&amp;materialsCitation.latitude=58.716812" title="Search Plazi for locations around (long -156.86537/lat 58.716812)">Naknek River</a>, [58.716814, -156.865366], vii.1919, ♂, JSS 19479, ♀, JSS 19478, J.S. Hine (CNC). MATANUSKA- SUSITNA CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-149.22968&amp;materialsCitation.latitude=61.541946" title="Search Plazi for locations around (long -149.22968/lat 61.541946)">Matanuska</a>, [61.541945, -149.229674], vii.1944, 3♀, JSS 19483, 19484, 19489, J. Chamberlin (CNC); vii.1944, ♀, OSAC 133385 , J. Chamberlin (OSAC). DENALI CO .: Healy, [63.845424, -149.013681], 26.vi.1921, ♀, JSS 19485, 27.vi.1921, ♂, JSS 19524, J.M. Aldrich (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-160.78334&amp;materialsCitation.latitude=63.866665" title="Search Plazi for locations around (long -160.78334/lat 63.866665)">Unalakleet</a>, [63.86666667, - 160.7833333], 8.viii.1961, ♀, CNCD 112193 , 9.viii.1961, ♀, CNCD 112194, 15.viii.1961, ♀, CNCD 26574, B.S. Heming (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-152.13333&amp;materialsCitation.latitude=69.36667" title="Search Plazi for locations around (long -152.13333/lat 69.36667)">Umiat</a>, [69.36666667, -152.1333333], 6.vii.1959, ♀, CNCD 26579 , J.E.H. Martin (CNC) . Colorado: LAKE COUNTY: Tennessee Pass, [39.357246, -106.313889], 24.vii.1917, ♀, JSS 19490, J.M. Aldrich (CNC). Loveland Pass, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.31389&amp;materialsCitation.latitude=39.357246" title="Search Plazi for locations around (long -106.31389/lat 39.357246)">Western Slope</a>, [39.65, -105.8666667], 3002m, 28.vii.1961, 2♂, CNCD 26385, 26386 , J.G. Chillcott (CNC). CLEAR CREEK CO .: Doolittle Ranch, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.60135&amp;materialsCitation.latitude=39.67574" title="Search Plazi for locations around (long -105.60135/lat 39.67574)">Mt. Evans</a>, [39.675739, -105.601348], 2987m, 23.vii.1961, ♂, CNCD 26383 , B.H. Poole (CNC); 8.vii.1961, ♀, CNCD 111622, 22.vii.1961, ♂, CNCD 26384, 11.viii.1961, ♀, CNCD 112192, W. R. M. Mason (CNC); 23.vii.1961, ♀, CNCD 35315, J.G. Chillcott (CNC). Chambers Lake, [40.603663, -105.853645], 3.ix.1932, ♂, JSS 20739, (CSUC). North Park, [40.73, -106.18], 1.vii.1932, ♀, JSS 20733, (CSUC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.9355&amp;materialsCitation.latitude=40.8078" title="Search Plazi for locations around (long -105.9355/lat 40.8078)">Glendevey</a>, [40.8078, -105.935501], 7.vii.1931, ♀, JSS 20741, C. R . Jones (CSUC) . Maine: PISCATAQUIS CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.92102&amp;materialsCitation.latitude=45.90497" title="Search Plazi for locations around (long -68.92102/lat 45.90497)">Mount Katahdin</a>, [45.904968, -68.921017], 5.vii.1958, ♂, JSS 19481, (CNC) . Massachusetts: NORFOLK CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.278694&amp;materialsCitation.latitude=42.308426" title="Search Plazi for locations around (long -71.278694/lat 42.308426)">Wellesley Hills</a>, [42.308427, -71.278693], 1963, ♀, JSS 19482, D. Meissner (CNC). HAMPSHIRE CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.49682&amp;materialsCitation.latitude=42.34038" title="Search Plazi for locations around (long -72.49682/lat 42.34038)">Amherst</a>, [42.34038, -72.496818], 15.v.1963, ♀, JSS 19486, S. T . William (CNC); 8.v.1965, ♀, JSS 19487, Mansfield (CNC); 8.v.1965, ♂, JSS 19488, B. Scott (CNC); 6.v.1973, ♂, USNM ENT 258065, A. Lavallee (USNM) . New York: ESSEX CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.00738&amp;materialsCitation.latitude=44.132435" title="Search Plazi for locations around (long -74.00738/lat 44.132435)">Mount MacIntyre Summit</a>, [44.132434, -74.007381], 26.vi.1941, ♂, CNCD 26387 , H. Dietrich (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.708725&amp;materialsCitation.latitude=44.2336" title="Search Plazi for locations around (long -73.708725/lat 44.2336)">Mount Hurricane</a>, [44.233601, -73.708728], vi.1939, ♂, JSS 19480, R . C. Shannon (CNC) . Utah: CACHE CO .: Cowley Canyon, [41.772669, -111.60407], 10–17.viii.1989, JSS 25132, W.J. Hanson (EMUS); Logan Cyn., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.42277&amp;materialsCitation.latitude=41.910305" title="Search Plazi for locations around (long -111.42277/lat 41.910305)">Turner Campground</a>, [41.85, -111.5666667], 2.vii.2003 6.vii.2003, ♀, JSS 24607, W.J. Hanson &amp; Brammer (EMUS); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.42277&amp;materialsCitation.latitude=41.910305" title="Search Plazi for locations around (long -111.42277/lat 41.910305)">Tony Grove Creek</a>, [41.886599, -111.564102], 29.vi.–8.vii.1994, ♂, JSS 24610, 8– 15.vii.1994, JSS 24608, 25121–25124, 25127, 15–26.vii.1994, JSS 25128, 4–12.viii.1994, JSS 25131, 12– 19.viii.1994, JSS 24609, 25125, 25126, 25129, 25130, W.J. Hanson (EMUS); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.42277&amp;materialsCitation.latitude=41.910305" title="Search Plazi for locations around (long -111.42277/lat 41.910305)">West Hodges Canyon</a>, [41.910305, - 111.42277], 8.viii.1978 11.viii.1978, JSS 25133, Hanson, Knowlton (EMUS) .</p></div>	https://treatment.plazi.org/id/03893F32FF99FF8CFF46FBD8FC6F8A6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FF9EFF8EFF46FA2BFF7A898B.text	03893F32FF9EFF8EFF46FA2BFF7A898B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus lotus (Williston 1887)	<div><p>Dasysyrphus lotus (Williston)</p><p>Figures: 10A–D, 21A</p><p>Synonyms and References:</p><p>Syrphus lotus Williston 1887: 75 (original description)</p><p>Metasyrphus lotus</p><p>Fluke 1933: 108 (key, redescription and transfer to Dasysyrphus)</p><p>Dasysyrphus lotus</p><p>Stone et al. 1965: 564 (catalogue and transfer to Dasysyrphus)</p><p>Vockeroth 1992: 67 (key and redescription)</p><p>Diagnosis: Abdominal maculae do not cross abdominal margin; maculae on tergites 3&amp;4 oblique and usually meet in the middle; face and gena continuously light (Figs 10A &amp; B).</p><p>Redescription:</p><p>Body Length: 9.4–11.7mm; Wing Length: 8.7–11.4mm</p><p>Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in females it is complete and&gt;½ the length between the antennae and ocelli) and dark pile; face light with dark vitta ¼–⅓ the width of the face and does not reach the antennal socket, face with light pile lateral to the eye, sometimes dark pile along fascia or dorsally, near frons; gena light (Fig. 10B) with light pile; occiput dark, covered in dense, light pollen, pile light, some with some dark; scape, pedicel and flagellomere dark.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile light; scutellum light with light pile along anterior edge, dark posteriorly; wing cell c and r 1 bare at base, br bare above spurious vein and small to large areas below, bm bare on most of anterior half, and cup bare under CuP (Fig. 10C); haltere light; pro- and mesofemora ⅓–½ dark basally ½–⅔ light apically, metafemur ⅔–¾ dark basally, ¼–⅓ light apically, tibiae light, tarsi light, metatarsi with dark anterior side.</p><p>Abdomen: Maculae on tergite 2 large and triangular (lateral edge flattened and medial edge with rounded point), ⅓–&lt;½ width of tergite, maculae on tergites 3&amp;4 are oblique and fairly straight, usually meeting medially, fairly uniform in thickness, narrowing toward medial edge slightly, maculae do not reach abdominal margin (Figs 10A &amp; B); sternites light with dark fasciae, anterior edge of fascia on sternite two usually angled forming a point in the centre.</p><p>Male Genitalia: Surstylus more or less triangular in shape in lateral view, with posterodorsal edge rounded, long pile on posterolateral surface, spines on anteroventral edge; cercus oval with long pile over entire surface; gonostylus hook-shaped with few pile on dorsal surface; basiphallus elongate, basally bent at about a 90 degree angle towards dorsum, apical end curved ventrally and fully sclerotized all the way around without pile or spines; distiphallus elongate, apical end flared out into small horn, which is mostly sclerotized but membranous at apex with minute setulae dorsally, basally curved towards dorsum and not enlarged (Fig. 10D).</p><p>Intraspecific variation: Abdominal maculae on tergites 3&amp;4 usually meet medially, but sometimes they are completely separated from each other.</p><p>Etymology: The word lotus is Latin, meaning washed.</p><p>Distribution: Southwestern USA (AZ, CA) and Mexico (Fig. 21A).</p><p>Ecology: Collected from March to November at elevations from 1524–2743m.</p><p>Discussion: Williston (1887) discusses two different specimens of D. lotus in his species description. The first, the type, and another that he speculates is “allied” with the first. This second specimen is reported to have several differences, mainly the shape of the abdominal maculae and gena colour. Williston (1887) describes this second specimen as having a “black stripe on the cheeks” and he describes the abdominal maculae as being “more oblique, and strongly concave in front” and “more convex behind”. In 1895 Snow describes this lotus variation as D. creper . Fluke (1933) states that most published records of lotus probably refer to creper and that most of the specimens identified as lotus agree with Williston’s (1887) variation (the female from New Mexico). Fluke postulates that the specimen might be teneral due to the light head, but then goes on to say that it does not explain the differently shaped maculae. Curran’s (1925) redescription of lotus also refers to creper . It is clear from the description of the abdominal maculae and the figure that he was not looking at Williston’s (1887) lotus .</p><p>Dasysyrphus lotus COI data has a high mean intraspecific variation of 3.09% (Table 2). In the NJ tree (Fig. 4), the two Nearctic specimens cluster closely together (1.34% different), while the Neotropical specimen is 3.29– 4.38% different from the others. This is highly suggestive that lotus is a complex as currently defined. With an extremely similar habitus among these three specimens as well as the 21 specimens where it was not possible to extract DNA, it can be concluded that lotus s.l. occurs from the southwestern Nearctic through to the Neotropical region, but that it likely includes at least two species. Further investigation needs to be conducted to help discern whether the Neotropical specimens are indeed a new species.</p><p>Type Material: “ Arizona,/ C.U. Lot 35”, “Acc. 19,702/ Williston”, “ Syrphus / lotus/ Will.”, red label “ Type / No. 129/ U.S. N.M.”, “Jeff Skevington/ Specimen #/ 23780”; ♀; deposited in the United States National Museum of Natural History, Washington, D.C (USNM). (examined)</p><p>Material Examined:</p><p>Mexico: 10 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.49364&amp;materialsCitation.latitude=23.727592" title="Search Plazi for locations around (long -105.49364/lat 23.727592)">Miles West of El Salto</a>, [23.727593, -105.49364], 2743m, 10.vii.1964, 2♂, CNCD 26391, 26392 , J.F. McAlpine (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-98.764595&amp;materialsCitation.latitude=19.119148" title="Search Plazi for locations around (long -98.764595/lat 19.119148)">Amecameca</a>, [19.119148, -98.764597], ix.1900, ♂, USNM ENT 35111 , O.W. Barrett (CNC). Atlacomulco, 8.x.1954, ♂, JSS 21093, R . Dreisbach (MSUC) . United States of America: Arizona: COCHISE CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.28964&amp;materialsCitation.latitude=31.440159" title="Search Plazi for locations around (long -110.28964/lat 31.440159)">Carr Canyon</a>, [31.440158, -110.289641], 1524m, 29.x.1967, ♂, CNCD 26388 , 2♀, CNCD 26393, 26394, R . F. Sternitzky (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.30771&amp;materialsCitation.latitude=31.447697" title="Search Plazi for locations around (long -110.30771/lat 31.447697)">Ramsey Canyon</a>, [31.447697, -110.307712], 10.iv.1967, ♀, CNCD 26397 , 12.vi.1967, 2♀, CNCD 26395, 26396, 28.vii.1967, CNCD 60065, 1-15.ix.1967, ♂, CNCD 26389, 1–8.x.1967, ♀, CNCD 26398, 15.xi.1967, ♂, CNCD 106795, R . F. Sternitzky (CNC); 23.ix.1994, ♂, JSS 18303, J.E. O’Hara (CNC); Miller Can., 3.v.1948, ♂, JSS 23760, A.L. Melander (USNM). SANTA CRUZ CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.51448&amp;materialsCitation.latitude=31.542591" title="Search Plazi for locations around (long -110.51448/lat 31.542591)">Canelo</a>, [31.542591, - 110.514478], 1372m, 17.iii.1967, ♂, CNCD 26390, R . F. Sternitzky (CNC). PIMA CO .: Mount Lemmon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.65907&amp;materialsCitation.latitude=38.87882" title="Search Plazi for locations around (long -120.65907/lat 38.87882)">San Catalina Mountains</a>, [32.443131, -110.788429], 2377m, 29.vii.1917, ♀, CNCD 26399, (CNC). California: ELDORADO CO., Blodgett Experimental Forest, [38.87882, -120.659071], 7.vi.1975, ♂, JSS 18302, D. Voeatlia (CNC) .</p></div>	https://treatment.plazi.org/id/03893F32FF9EFF8EFF46FA2BFF7A898B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FF9CFF90FF46F90BFB8D88A4.text	03893F32FF9CFF90FF46F90BFB8D88A4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus nigricornis (Verrall 1873)	<div><p>Dasysyrphus nigricornis (Verrall)</p><p>Figures: 11A–E, 18B, 21A</p><p>Synonyms and References:</p><p>Scaeva obscura Zetterstedt 1838: 601 (original description, name preoccupied)</p><p>Bicik &amp; Laska 1996 (lectotype designation and discussion)</p><p>Syrphus nigricornis Verrall 1873, 251 (new name given, short description and transfer to Syrphus)</p><p>Verrall 1901: 353 (discussion)</p><p>Becker 1921: 44 (redescription)</p><p>Syrphus obscuratus Ringdahl 1928: 19 (original description)</p><p>Syrphus nigricornis var. obscuraticeps Frey 1950: 10 (original description)</p><p>Dasysyrphus nigricornis Verrall:</p><p>Peck 1974: 128 (key and transfer to Dasysyrphus)</p><p>Peck 1988: 15 (catalogue)</p><p>Barkalov 2007: 283 (key and redescription)</p><p>Speight 2011: 70 (discussion)</p><p>Diagnosis: Abdominal maculae of tergites 3 &amp; 4 oblique, fairly uniform in thickness with slight constriction in the middle and do not cross abdominal margin (Figs 11A–C). Maculae on tergite 2 of male small (&lt;¼ width of tergite), triangular to oval and close to lateral edges (Fig. 11A), in female are larger (¼–⅓ width of tergite) and narrowly oval (Fig. 11B); usually darker in appearance due to reduced maculae. Wing densely microtrichose (Fig. 11D). Gonostylus boot-shaped with projection on posterodorsal surface that projects posterodorsally (Fig. 11D). Similar in appearance to D. pauxillus (Fig. 14A), pacificus (Fig. 13A) and laticaudus (Fig. 8A).</p><p>Redescription:</p><p>Body Length: 5.5–7.7mm; Wing Length: 4.5–7.7mm</p><p>Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in females it is complete and ⅓–½ the length between the antennae and ocelli (Fig. 18B)) and dark pile; face light with dark vitta ¼–½ the width of the face, usually not reaching antennal socket, face with dark pile lateral to the eye; gena dark, typically dark pile, but sometimes light; occiput dark, covered in light pollen, pile light; scape and pedicel dark, flagellomere dark.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile varies between all light and dark anteriorly, light posteriorly; scutellum light to dark with darker lateral edges, light pile sometimes with light and dark pile; wings entirely microtrichose (Fig. 11D); haltere light; pro- and mesofemora dark to dark ½ basally, ½ light apically, metafemur dark, pro- and mesotibiae all dark to all light, meta tibiae all dark to light ⅓ basally, dark ⅔ apically, tarsi dark.</p><p>Abdomen: Maculae on tergite 2 small (&lt;¼ width of tergite), triangular to oval and close to lateral edges in males (Fig. 11A), larger (¼–⅓ width of tergite) and narrowly oval in females (Fig. 11B), maculae on tergites 3&amp;4 are oblique, fairly uniform in width with slight constriction in the middle, maculae do not reach abdominal margin (Figs 11A–C); sternites light with dark fasciae varying in thickness.</p><p>Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened posteriorly, long pile on dorsal, laterodorsal and posterodorsal sides, row of spines on posteromedial edge; cercus oval with long pile over entire surface; gonostylus boot-shaped with projection on posterodorsal surface that projects posterodorsally, pile on dorsal surface; basiphallus elongate, basally bent at 135 degree angle towards anterodorsum, apical end curved ventrally and is partly membranous dorsally, with spines on posterodorsal side; distiphallus elongate, apical end flared out into horn, which is mostly sclerotized but membranous at apex with minute setulae dorsally, basally bent towards dorsum and enlarged ventrally at bend (Fig. 11D).</p><p>Intraspecific variation: Maculae on tergite two can vary in width. In males maculae are typically &lt;¼ the width of the tergite, but in rare occasions may be up to ⅓ of the width. In females hey are typically ¼–⅓ the width. Maculae on tergite 3 &amp; 4 in the male look similar to those of pauxillus, pacificus and laticaudus, however in the female the maculae tend to be slightly thinner than similar species and sometimes are more narrow on the medial edge.</p><p>Etymology: From the Latin niger, meaning black, dark, dusky, and cornus, meaning horn (possibly referring to antennae).</p><p>Distribution: Northern Canada (MB, NL, NT, QC, YT), Northern USA (AK) and Greenland (Fig. 21A). Palaearctic (likely throughout far north but distribution needs to be verified).</p><p>Ecology: Has been collected from late May–July, from elevations of 762–1250m.</p><p>Discussion: Dasysyrphus nigricornis is recognized from the Nearctic for the first time. Long buried within pauxillus in this region, specimens from northern Canada and the United States are morphologically and genetically different from pauxillus and the two other species also split off from it here, laticaudus and pacificus . Male genitalia of these Nearctic specimens match the genitalia of the nigricornis paralectotype figured by Bicik &amp; Laska (1996, Fig. A). Part of the reason that this species has not been recognized in this region before is that much confusion has existed over the use of the name in the Palaearctic. When Verrall provided a new name for obscura in 1873, because the name was preoccupied, he used specimens from Scotland for his description. However, obscura was described by Zetterstedt from Scandinavia and is not known to occur in the United Kingdom (Bicik &amp; Laska 1996; Speight 2011). The material used by Verrall in his redescription is apparently actually pinastri, leading to the obvious confusion. In 1996, Bicik &amp; Laska examined the type series of obscura to help straighten out the nomenclature. Upon their examination of the type series they found three different species, three specimens of nigricornis s.s., two of pauxillus s.l. and one of pinastri . They selected a lectotype to stabilize the name, as well as provided figures of genitalia and characters to differentiate nigricornis from pauxillus s.l. and pinastri .</p><p>Despite being morphologically distinct from D. occidualis, D. nigricornis cannot be separated from occidualis by COI DNA.</p><p>Type Material: “ S. obscura Juckasj. ” (Zetterstedt label), “ S. obscuraticeps Frey ” (Heese label, 1969), number: “185”, ♂, deposited in the Museum of Zoology, Lund University (MZLU), Lund, Sweden (Bicik &amp; Laska 1996). (not examined)</p><p>Material Examined:</p><p>Canada: Manitoba: CHURCHILL AREA: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.99777&amp;materialsCitation.latitude=58.50767" title="Search Plazi for locations around (long -93.99777/lat 58.50767)">Warkworth Creek</a>, [58.507672, -93.997769], 10.vi.1952, ♂, CNCD 13766 , C.D. Bird (CNC); Burn site N[orth] of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.82889&amp;materialsCitation.latitude=58.61861" title="Search Plazi for locations around (long -93.82889/lat 58.61861)">Twin</a> lakes, [58.61861111, -93.82888889], 29.vi.2007, 3♂, CNCD 9204–9206 , J. Skevington (CNC); Bluff D at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.89833&amp;materialsCitation.latitude=58.761665" title="Search Plazi for locations around (long -93.89833/lat 58.761665)">Bird Cove</a>, [58.76166667, -93.89833333], 30.vi.2007, CNCD 9274 , J. Skevington (CNC) . Newfoundland and Labrador: Pinware River Provincial Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.708115&amp;materialsCitation.latitude=51.634266" title="Search Plazi for locations around (long -56.708115/lat 51.634266)">Division No.</a> 9, [51.634267, -56.708114], 29.vi.1983, ♂, CNCD 13902 , A. Borkent (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.010406&amp;materialsCitation.latitude=53.69477" title="Search Plazi for locations around (long -57.010406/lat 53.69477)">Cartwright</a>, [53.694772, -57.010406], 29.vi.1955, ♀, CNCD 13903 , E.E. Sterns (CNC) . Northwest Territories: Resdelta, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.78045&amp;materialsCitation.latitude=66.051735" title="Search Plazi for locations around (long -120.78045/lat 66.051735)">Great Slave Lake</a>, [66.051734, - 120.780449], 22.vi.1944, ♀, CNCD 13777 , J. R. Vockeroth (CNC). Km 491 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-135.79105&amp;materialsCitation.latitude=67.18279" title="Search Plazi for locations around (long -135.79105/lat 67.18279)">Dempster Highway</a>, [67.182795, - 135.791046], 26.vi.1982, ♂, CNCD 13772 , G. &amp; M. Wood (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-135.01071&amp;materialsCitation.latitude=68.219635" title="Search Plazi for locations around (long -135.01071/lat 68.219635)">Aklavik</a>, [68.219638, -135.010707], 30.vi.1956, ♀, CNCD 13776 , E.F. Cashman (CNC). Reindeer Depot, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-134.07158&amp;materialsCitation.latitude=68.668976" title="Search Plazi for locations around (long -134.07158/lat 68.668976)">Mackenzie Delta</a>, [68.668974, -134.071574], 25.vi.1948, ♂, CNCD 13712 , 27.vi.1948, 2♂, CNCD 13714, 15384, 28.vi.1948, ♂, CNCD 13713, 6.vii.1948, ♂, CNCD 13715, J. R. Vockeroth (CNC). Granet Lake, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-125.58813&amp;materialsCitation.latitude=68.70147" title="Search Plazi for locations around (long -125.58813/lat 68.70147)">Franklin</a>, [68.701471, -125.58813], 20.vii.1969, ♀, CNCD 13778 , G.E. Shewell (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-132.17247&amp;materialsCitation.latitude=69.42684" title="Search Plazi for locations around (long -132.17247/lat 69.42684)">21 miles east Tuktoyaktuk</a>, [69.426844, -132.172475], 20–25.vi.1971, 4♂, CNCD 13702–13704, 13707 , 2–5.vii.1971, ♂, CNCD 13705, ♀, CNCD 13774, 8–12.vii.1971, ♀, CNCD 13775, D.M. Wood (CNC); 21.vi.1971, ♂, CNCD 13706, W. R. M. Mason (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-133.01637&amp;materialsCitation.latitude=69.43837" title="Search Plazi for locations around (long -133.01637/lat 69.43837)">Tuktoyaktuk</a>, [69.438369, -133.01637], 17.vi.1971, 3♂, CNCD 13709–13711 , D.M. Wood (CNC) . Quebec: Kuujjuaq (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.40618&amp;materialsCitation.latitude=58.100075" title="Search Plazi for locations around (long -68.40618/lat 58.100075)">Fort Chimo</a>), [58.100076, - 68.406179], 23.vii.1948, ♀, CNCD 13879, R . H. MacLeod (CNC) . Yukon Territory: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-132.80998&amp;materialsCitation.latitude=62.210136" title="Search Plazi for locations around (long -132.80998/lat 62.210136)">Swim Lakes</a>, [62.210135, - 132.809978], 975m, 11.vi.1960, ♀, CNCD 13568 , 15.vi.1960, ♀, CNCD 13733, 25.vi.1960, 2♀, CNCD 13734, 13735, J.E.H. Martin (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-131.2666&amp;materialsCitation.latitude=62.616764" title="Search Plazi for locations around (long -131.2666/lat 62.616764)">Sheldon Lake</a>, [62.616763, -131.266603], 1067m, 7.vii.1960, 2♀, CNCD 13737, 13738 , E.W. Rockburne (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-132.33333&amp;materialsCitation.latitude=62.683334" title="Search Plazi for locations around (long -132.33333/lat 62.683334)">La Force Lake</a>, [62.683333, -132.333333], 1006m, 26.vi.1960, ♀, CNCD 13726,7 . vii.1960, ♀, CNCD 13722, 9.vii.1960, ♀, CNCD 13723, 10.vii.1960, 2♀, CNCD 13724, 13725, J.E.H. Martin (CNC); 25.vi.1960, ♀, CNCD 13732, 7.vii.1960, ♀, CNCD 13727, 11.vii.1960, ♀, CNCD 13729, 13.vii.1960, 2♀, CNCD 13730, E.W. Rockburne (CNC). Hunter Creek, Dawson, [63.779, -138.61], 914m, 3.vii.1949, ♂, CNCD 13701 , ♀, CNCD 13740, P.F. Bruggemann (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-139.41052&amp;materialsCitation.latitude=64.05002" title="Search Plazi for locations around (long -139.41052/lat 64.05002)">Dawson</a>, [64.050017, -139.41052], 16.vi.1949, ♀, CNCD 13741 , W.W. Judd (CNC). North Fork Pass, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-138.25064&amp;materialsCitation.latitude=64.56599" title="Search Plazi for locations around (long -138.25064/lat 64.56599)">Ogilvie Mountains</a>, [64.565983, -138.250648], 1250m, 12.vi.1962, ♂, CNCD 13700, R . E. Leech (CNC). North Fork Crossing, Mile 43 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-138.25064&amp;materialsCitation.latitude=64.56599" title="Search Plazi for locations around (long -138.25064/lat 64.56599)">Peel Pit Road</a>, [64.565983, -138.250648], 1067m, 4.vii.1962, ♀, CNCD 13736 , P.J. Skitsko (CNC). DEMPSTER HIGHWAY: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-138.33789&amp;materialsCitation.latitude=64.605995" title="Search Plazi for locations around (long -138.33789/lat 64.605995)">Mile</a> 51, [64.605992, -138.33789], 17–21.vi.1973, ♂, CNCD 13708 , ♀, CNCD 13739, G. &amp;D.M. Wood (CNC); km 465, [67.046392, -136.209668], 25–26.vi.1982, 2♂, CNCD 5811, 13720, ♂ and ♀ (pinned together), CNCD 13719, G. &amp; M. Wood (CNC); Wright Pass, [67.083297, -136.162751], 9.vii.1985, ♀, JSS 20169, S.G. Cannings (UBCZ). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-136.98886&amp;materialsCitation.latitude=68.16365" title="Search Plazi for locations around (long -136.98886/lat 68.16365)">Richardson Mountains</a>, [68.163651, -136.988868], 762m, 12.vii.1982, ♂ &amp; ♀, CNCD 13721 , M. Wood (CNC) .</p><p>Greenland: Sondrestrom Air Base; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.709167&amp;materialsCitation.latitude=67.01056" title="Search Plazi for locations around (long -50.709167/lat 67.01056)">Near Sugar Loaf Mountain</a>, [67.010556, -50.709167], 11.vii.1963, ♂, JSS 23779, T . J. Curtin (USNM) . United States of America: Alaska: MATANUSKA- SUSITNA CO.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-149.22968&amp;materialsCitation.latitude=61.541946" title="Search Plazi for locations around (long -149.22968/lat 61.541946)">Matanuska</a>, [61.541945, -149.229674] , 28.v.1945, ♀, JSS 19518, 7.vi.1944, ♀, JSS 19519, J.C. Chamberlin (CNC); 7.vi.1944, 2♂, JSS 19520, 19521, J.C. Chamberlin (USNM). Mount Fairplay, Mile 32 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-142.26294&amp;materialsCitation.latitude=63.67904" title="Search Plazi for locations around (long -142.26294/lat 63.67904)">Taylor Highway</a>, [63.679038, - 142.262944], 1097m , 10.vii.1962, ♀, CNCD 26443, P.J. Skitsko (CNC). Nome Area, Mile 17 of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-165.29723&amp;materialsCitation.latitude=64.70611" title="Search Plazi for locations around (long -165.29723/lat 64.70611)">Kougarok Road</a>, [64.70611111, -165.2972222] , 26.vi.2005 28.vi.2005, ♀, CNCD 30896, J.&amp; R. Skevington (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-152.13333&amp;materialsCitation.latitude=69.36667" title="Search Plazi for locations around (long -152.13333/lat 69.36667)">Umiat</a>, [69.36666667, -152.1333333], 24.vi.1947, ♂, JSS 19522, K.L. Knight (USNM) .</p></div>	https://treatment.plazi.org/id/03893F32FF9CFF90FF46F90BFB8D88A4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FF80FF97FF46FF70FF7A8C08.text	03893F32FF80FF97FF46FF70FF7A8C08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus occidualis Locke & Skevington 2013	<div><p>Dasysyrphus occidualis Locke &amp; Skevington sp. nov.</p><p>Figures: 12A–E, 21B</p><p>Synonyms and References:</p><p>Syrphus amalopis Osten Sacken:</p><p>Williston 1887: 69 (in part; description)</p><p>Metasyrphus amalopis:</p><p>Fluke 1933: 110 (in part; key and redescription)</p><p>Dasysyrphus amalopis:</p><p>Stone et al. 1965: 563 (in part; catalogue)</p><p>Vockeroth 1992: 63 (in part; key and redescription)</p><p>Diagnosis: Abdominal maculae do not meet abdominal margin; maculae on tergites 3&amp;4 are constricted medially and are either divided or not (Figs 12A–C). Wings cell br usually with a small bare area above base of spurious vein (Fig. 12E). Surstylus with a prominent posterolateral edge (Fig. 12D). Similar to D. amalopis, but D. amalopis has maculae on tergites 3&amp;4 usually undivided (Fig. 5A), cell br densely microtrichose and does not have a prominent posterolateral edge on the surstylus (Fig. 5C). Male genitalia should be used to reliably differentiate between these two species. There is a range of overlap in southern Yukon where it may not be possible to distinguish between females of these two species due to variations in the aforementioned characters.</p><p>Description:</p><p>Body Length: 8.0– 11mm; Wing Length: 7.2–10.7mm</p><p>Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in females it is continuous, ⅓–½ the length between the antennae and ocelli) with dark pile; face light with dark vitta &lt;¼–⅓ the width of the face, usually not reaching antennal socket, if it does then narrowly so, face with dark pile; gena dark with dark pile ventral to the eye and light pile elsewhere; occiput dark, covered in light pollen, pile light; scape, pedicel and flagellomere dark.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile light anteriorly and dark posteriorly (amount varies); scutellum light to dark with darker lateral edges, some light pile along anterior edge and dark pile elsewhere; wing densely microtrichose, cell br usually with small bare areas at above base of spurious vein, r 1 sometimes bare at base and cup sometimes bare in areas along vein CuP (Fig. 12E); haltere light; pro- and mesofemora ½–¾ dark basially, ¼–½ light apically, metafemur ¾ dark basially, ¼ light apically, tibiae variable from all light to slightly darkened apically to ¾ dark apically, tarsi light with dark anterior side to all dark.</p><p>Abdomen: Maculae on tergite 2 large, ⅓–&lt;½ the width of tergite 2, and oval with constriction medially (sometimes just slight constriction), maculae on tergites 3&amp;4 are oblique and constricted medially, usually completely dividing maculae into four spots, if not completely divided then maculae on tergite 4 usually more constricted that on tergite three and almost divided, maculae do not reach abdominal margin (Figs 12A–C); sternite 2 is light with a dark triangular to straight macula connecting with the lateral edges of the sternite, sternites 3 &amp; 4 are light with dark fasciae of varying thickness.</p><p>Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened posteriorly with prominent posterolateral edge, long pile on dorsal, laterodorsal and posterodorsal sides, row of spines on posteromedial edge; cercus oval with long pile over entire surface; gonostylus boot-shaped with pointed projection on posterodorsal surface that projects dorsally, pile on dorsal surface; basiphallus elongate, basally bent at about a 135 degree angle towards the anterodorsum, apical end curved ventrally with a widened apex and is partly membranous dorsally, with spines on posterodorsal side, apex; distiphallus elongate, apical end flared out into horn, which is mostly sclerotized but membranous at apex with minute setulae dorsally, basally bent towards dorsum and enlarged ventrally at bend (Fig. 12D).</p><p>Intraspecific variation: The maculae on tergites 3 &amp; 4 are often divided (Fig. 12B). If not completely divided then maculae on tergite 4 usually more constricted that on tergite three and almost divided. Can have maculae on tergites 3 &amp; 4 entire, but this less common (Fig. 12A). Wing densely microtrichose, except for cell br, which is usually bare above base of spurious vein, sometimes only in very tiny areas with one or two microtrichia missing, r 1 sometimes bare of microtrichia at base and cup sometimes bare in areas along vein CuP (Fig 12E).</p><p>Etymology: From the Latin occidualis, meaning western.</p><p>Distribution: Western Canada (AB, BC, YT) and western USA (AK, CO, WA, WY) (Fig. 21B).</p><p>Ecology: Collected from May–August, but more frequently collected in July. Collected at elevations of 613– 3658m. Habitats listed on labels include “outlet marsh”, “sedge fen” and “subalpine”.</p><p>Discussion: Until now, D. occidualis has been hidden within D. amalopis . With an absence of distinguishing external morphological characters, additional data were necessary to illuminate cryptic species such as this. In this study, COI DNA first highlighted the existence of two distinctly different genetic clusters within amalopis . Upon further inspection of the specimens, the male genitalia showed a prominent posterolateral edge on the surstylus (Fig. 12D), which amalopis does not have (Fig. 5C). When their geographic ranges were studied it was found that specimens of occidualis were only located in western Canada and the US (from Alaska to Colorado), while amalopis was found only in northern Canada. There is an overlap zone in southern Yukon so specimens in this area must be carefully studied to make the correct species determination.</p><p>Type Material: “ USA: WA: Mt. Baker-/ Snoqualmie NF: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.697914&amp;materialsCitation.latitude=48.90495" title="Search Plazi for locations around (long -121.697914/lat 48.90495)">Mt. Baker</a>:/ SilverFirCampground; 2012ft. / 48.904950°N, 121.697917°W;/ 8.vii.2010; M.M.Locke ”, “ J. Skevington / Specimen #/ 23708”, “ HOLOTYPE / Dasysyrphus / occidualis/ Locke &amp;/ Skevington”, ♂, deposited in the Canadian Collection if Insects, Arachnids and Nematodes (CNC), Ottawa, Ontario , Canada.</p><p>Material Examined:</p><p>Canada: Alberta: Kananaskis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.08769&amp;materialsCitation.latitude=51.10088" title="Search Plazi for locations around (long -115.08769/lat 51.10088)">Forest Experimental Station Seebe</a>, [51.100879, -115.087692], 15.vi.1968, ♀, CNCD 5014 , H.J. Teskey (CNC). BANFF NATIONAL PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.568436&amp;materialsCitation.latitude=51.180275" title="Search Plazi for locations around (long -115.568436/lat 51.180275)">Banff</a>, [51.180275, -115.568433], 15.vii.1922, 3♂, CNCD 13621–13623 , C.B.D. Garrett (CNC); ♀, JSS 19432, N.B. Sanson (CNC); 11.vii.1951, ♀, CNCD 13619, (CNC) . British Columbia: CATHEDRAL PROV. PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.2073&amp;materialsCitation.latitude=49.055786" title="Search Plazi for locations around (long -120.2073/lat 49.055786)">Glacier Lake</a>, [49.055785, -120.207299], 6.vii.1986, ♀, RBCM ENT 9882357, R . A. Cannings (RBCM); fen, Lk. of the Woods trail, [49.08, -120.18], 26.vii.1988, ♀, JSS 20186, S.G. Cannings (UBCZ). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.07434&amp;materialsCitation.latitude=49.811127" title="Search Plazi for locations around (long -123.07434/lat 49.811127)">Squamish</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.07434&amp;materialsCitation.latitude=49.811127" title="Search Plazi for locations around (long -123.07434/lat 49.811127)">Diamond Head Trail</a>, [49.811126, -123.07434], 1219m, 11.viii.1953, ♀, CNCD 13610 , W. R. M. Mason (CNC). Duffey L. Blowdown Cr., lake S. of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.34167&amp;materialsCitation.latitude=50.404167" title="Search Plazi for locations around (long -122.34167/lat 50.404167)">Gott Peak</a>, [50.40416667, - 122.3416667], 21.vii.1988, ♀, RBCM ENT 9918197 , C.S. Guppy (RBCM). Cornwall Hills Lookout, [50.7, - 121.5], 2037m, 14.vi.1989, ♂, RBCM ENT 990626, R . A. Cannings &amp; C.S. Guppy (RBCM). KOOTENAY NATIONAL PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.93692&amp;materialsCitation.latitude=50.77666" title="Search Plazi for locations around (long -115.93692/lat 50.77666)">Daer-Pitts Aspen Control</a> 2, [50.776662, -115.936917], 17.vi.–2.vii.2000, 2♂, CNCD 30908, 30909 , ♀, CNCD 30903, G. Gareau (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.137&amp;materialsCitation.latitude=51.054" title="Search Plazi for locations around (long -116.137/lat 51.054)">Floe Lake</a>, [51.054, -116.137], 1981m, 9.viii.1955, ♀, CNCD 13611 , J. R. W. McGillis (CNC). MOUNT REVELSTOKE NATIONAL PARK: [51.035979, -118.142509], 1828m, 26.vii.1952, ♀, CNCD 13612, G.P. Holland (CNC); 15.viii.1952, ♀, CNCD 13613, G.J. Spencer (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.14251&amp;materialsCitation.latitude=51.03598" title="Search Plazi for locations around (long -118.14251/lat 51.03598)">Mount Harry</a>, [51.035979, -118.142509], 2073m, 8.viii.1952, ♀, CNCD 13614 , G.J. Spencer (CNC); Mt. Revelstoke, [51.05, - 118.1333333], 1829m, 7.viii.1952, ♀, RBCM ENT 99188105 , G.P. Holland (RBCM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.10883&amp;materialsCitation.latitude=51.079906" title="Search Plazi for locations around (long -118.10883/lat 51.079906)">Eva Lake Trail</a>, [51.079905, -118.108837], 1828m, 6.viii.1952, ♀, CNCD 13615 , G.J. Spencer (CNC). SKENNA- QUEEN CHARLOTTE R . D.: Tlell, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-131.93234&amp;materialsCitation.latitude=53.56259" title="Search Plazi for locations around (long -131.93234/lat 53.56259)">Queen Charlotte Islands</a>, [53.562591, -131.93235], 25.v.1957, ♂, CNCD 13620 , E.E. MacDougall (CNC). NORTHERN ROCKIES R . D.: Summit Lake, Mile 392 of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-124.66691&amp;materialsCitation.latitude=58.6482" title="Search Plazi for locations around (long -124.66691/lat 58.6482)">Alaska Highway</a>, [58.648201, -124.666912], 1372m, 1.vii.1959, ♀, CNCD 13603, R . E. Leech (CNC); 1.vii.1959, ♀, CNCD 13605, 2–4.vii.1959, 3♀, CNCD 13606– 13608, 6.vii.1959, ♀, CNCD 13604, E.E. MacDougall (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.69743&amp;materialsCitation.latitude=58.804592" title="Search Plazi for locations around (long -122.69743/lat 58.804592)">Ft. Nelson</a>, [58.804594, -122.697436], 4.vii.1949, ♀, JSS 19993, J.K. Morie (UBCZ). Liard R . hotsprings, [59.425991, -126.096398], 23.vii.1987, 3♀, JSS 20176– 20178, S.G. Cannings (UBCZ). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-127.14242&amp;materialsCitation.latitude=59.666676" title="Search Plazi for locations around (long -127.14242/lat 59.666676)">Fireside</a>, [59.666676, -127.14242], 24.v.1980, CNCD 60083 , Bruce Gill (CNC). 7 miles N Lions Bay, 18.v.1979, ♂, CNCD 5463 , G.F. Hevel (CNC). Spanish Hills, Galiano Island, 10.v.1980, CNCD 60094 , G.G.E. Scudder (CNC) . Yukon Territory: Carcross Dunes at <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-134.7181&amp;materialsCitation.latitude=60.17066" title="Search Plazi for locations around (long -134.7181/lat 60.17066)">Bennett</a> L. beach, [60.17066, -134.71809], 666m, 25.vi.2009, ♀, JSS 20819, Lee Mennell (RBCM). Takhini R ., west dunes 6.8km NNE of Kusawa L. outlet, [60.66824, -136.07664], 19.vi.2009, ♀, JSS 20820, S. Cannings (RBCM). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-131.2666&amp;materialsCitation.latitude=62.616764" title="Search Plazi for locations around (long -131.2666/lat 62.616764)">Sheldon Lake</a>, [62.616763, - 131.266603], 914m, 21.vii.1960, ♀, CNCD 13561 , J.E.H. Martin (CNC). DEMPSTER HIGHWAY: North Klondike River, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-138.21835&amp;materialsCitation.latitude=64.45056" title="Search Plazi for locations around (long -138.21835/lat 64.45056)">Km</a> 64 [64.45056, -138.218356], 9.vi.1980, ♂, JSS 18376, R . J. Cannings (CNC) . United States of America: Alaska: MATANUSKA- SUSITNA CO.: Matanuska, [61.541945, -149.229674], 29.v.1944, ♀, JSS 19431, J. Chamberlin (CNC). Nome Area, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-165.29723&amp;materialsCitation.latitude=64.70611" title="Search Plazi for locations around (long -165.29723/lat 64.70611)">Mile</a> 17 of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-165.29723&amp;materialsCitation.latitude=64.70611" title="Search Plazi for locations around (long -165.29723/lat 64.70611)">Kougarok Road</a>, [64.70611111, -165.2972222], 26–28.vi.2005, ♀, CNCD 30899 , J.&amp; R. Skevington (CNC) . Colorado: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.045006&amp;materialsCitation.latitude=38.840343" title="Search Plazi for locations around (long -105.045006/lat 38.840343)">Pike's Peak Summit</a>, [38.840342, -105.045006], 14.viii.1977, ♀, USNM ENT 258091 , A.B. Gurney (USNM). GUNNISON CO.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-107.40768&amp;materialsCitation.latitude=39.118885" title="Search Plazi for locations around (long -107.40768/lat 39.118885)">Ragged Mountain</a>, [39.118885, -107.407678], 20.vi.1950, ♂, CNCD 26345 , C.A. Downing (CNC). CLEAR CREEK CO.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.594894&amp;materialsCitation.latitude=39.645576" title="Search Plazi for locations around (long -105.594894/lat 39.645576)">Mount Evans</a>, [39.645575, -105.594897] 22.vii.1961, ♀, CNCD 26335 , J.G. Chillcott (CNC); 3.viii.1961, 2♀, CNCD 26330, 26331, S.M. Clark (CNC); 3.viii.1961, 2♀, CNCD 26332, 26333, W. R. M. Mason (CNC); 4.viii.1961, ♂, CNCD 26342, J. R. Stainer (CNC); 10.viii.1961, ♀, CNCD 26334, C.H. Mann (CNC); Timberline, Mount Evans, 3566m, 22.vii.1961, ♀, CNCD 26336 , C.H. Mann (CNC); 22.vii.1961, ♀, CNCD 26337, B.H. Poole (CNC); Echo Lake, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.6035&amp;materialsCitation.latitude=39.658085" title="Search Plazi for locations around (long -105.6035/lat 39.658085)">Mt. Evans</a>, [39.658085, - 105.6035], 3231m, 22.vii.1961, ♀, CNCD 26338 , J.G. Chillcott (CNC); Doolittle Ranch, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.60135&amp;materialsCitation.latitude=39.67574" title="Search Plazi for locations around (long -105.60135/lat 39.67574)">Mt. Evans</a>, [39.675739, - 105.601348], 2987m, 12.viii.1961, ♀, CNCD 26339 , J. R. Stainer (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.711105&amp;materialsCitation.latitude=39.594994" title="Search Plazi for locations around (long -105.711105/lat 39.594994)">Guanella Pass</a>, [39.594992, - 105.711109], 3536m, 14.vii.1961, ♀, CNCD 26340 , S.M. Clark (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.58483&amp;materialsCitation.latitude=40.06926" title="Search Plazi for locations around (long -105.58483/lat 40.06926)">Brainard Lake Rec. Area</a> 4mi W of Ward Colorado, [40.069261, -105.584835], 11.viii.1952, ♀, JSS 21103, R. R . Dreisbach (MSUC); 21.viii.1977, 2♂, USNM ENT 258062, 258063, A.G. Lavallee (USNM). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-118.22767&amp;materialsCitation.latitude=40.520817" title="Search Plazi for locations around (long -118.22767/lat 40.520817)">Cameron Pass</a>, [40.520816, -118.227668], 28.vii.1954, ♂, OSAC 133386 , Chas. L. Fluke (OSAC). North Park, [40.73, -106.18], 1.vii.1932, ♀, JSS 20665, (CSUC) . Washington: MT. RANIER NATIONAL PARK: Paradise Valley, [46.782709, -121.727952], 15.vii.1920, ♂, JSS 23880, 18.vii.1920, 7♂, JSS 23722, 23888, 23889, 23893, 23894, 23900, 23904, 19.vii.1920, 2♂, JSS 23886, 23887, ♀, JSS 23718, 25.vii.1920, ♂, JSS 23883, E.C. Van Dyke (CAS); 18.vii.1920, ♂, JSS 20613, E.C. Van Dyke (EMEC); Carbon River near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.33207&amp;materialsCitation.latitude=47.60621" title="Search Plazi for locations around (long -122.33207/lat 47.60621)">Ranger station</a>, [46.85, -121.75], 15.viii.1999, ♀, JSS 20681, B. Kondratieff (CSUC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.33207&amp;materialsCitation.latitude=47.60621" title="Search Plazi for locations around (long -122.33207/lat 47.60621)">Louis Lake</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.33207&amp;materialsCitation.latitude=47.60621" title="Search Plazi for locations around (long -122.33207/lat 47.60621)">Fryingpan Cr.</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.33207&amp;materialsCitation.latitude=47.60621" title="Search Plazi for locations around (long -122.33207/lat 47.60621)">Sunrise Rd.</a>, [46.85, -121.75], 17.vi.2004, ♂, JSS 20804, B. Kondratieff, J. Schmidt, J. Owens (CSUC). Seattle, [47.60621, -122.332071], 16.vi.1901, ♀, OSAC 133444, (OSAC). MT. BAKER- SNOQUALMIE NATIONAL FOREST : Mt. Baker, Picture Lake, [48.866217, -121.676533], 1273m, 7.vii.2010, ♂, JSS 23706, ♀, JSS 24625, A. Young (DEBU); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.697914&amp;materialsCitation.latitude=48.90495" title="Search Plazi for locations around (long 121.697914/lat 48.90495)">Silver Fir Campground</a>, [48.90495, 121.697917], 613m, 8.vii.2010, ♂, JSS 23708, 9.vii.2010, ♀, CNCD 35311, 60043 , M.M. Locke (CNC); 9.vii.2010, CNCD 60046, CNCD 60047, A. Young (CNC) . Wyoming: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-107.55642&amp;materialsCitation.latitude=43.084282" title="Search Plazi for locations around (long -107.55642/lat 43.084282)">Wyoming</a>, [43.084281, -107.556417], 1952, ♀, CNCD 26329 , F.M. Hull (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.537674&amp;materialsCitation.latitude=44.58772" title="Search Plazi for locations around (long -110.537674/lat 44.58772)">Yellowstone National Park</a>, [44.587719, -110.537673], 20.vii.1925, ♂, CNCD 26343 , A.A. Nichol (CNC) .</p></div>	https://treatment.plazi.org/id/03893F32FF80FF97FF46FF70FF7A8C08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FF85FF98FF46FC80FBB18E78.text	03893F32FF85FF98FF46FC80FBB18E78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus pacificus (Cole & Lovett 1919)	<div><p>Dasysyrphus pacificus (Lovett)</p><p>Figures: 13A–D, 21C</p><p>Synonyms and References:</p><p>Syrphus pacifica Lovett 1919: 245 (in (Cole &amp; Lovett 1919); original description)</p><p>Metasyrphus pacifica (Lovett):</p><p>Fluke 1933: 116 (key, redescription and transfer to Metasyrphus)</p><p>Dasysyrphus pacificus (Lovett):</p><p>Stone et al. 1965: 564 (catalogue and transfer to Dasysyrphus)</p><p>Dasysyrphus pauxillus (Williston):</p><p>Vockeroth 1986: 203 (in part; synonymy)</p><p>Vockeroth 1992: 68 (in part; key and redescription)</p><p>Diagnosis: Abdominal maculae of tergites 3&amp;4 slightly arcuate, uniform in thickness and do not cross abdominal margin (Figs 13A &amp; B); very similar in appearance to D. pauxillus (Fig. 14A), D. nigricornis (Figs 11A &amp; B) and D. laticaudus (Fig. 8A). Wing typically much more bare than any of the three aforementioned species, with cells br and bm with large dense areas (Fig. 13C). Male gonostylus with narrow, wavy projection on posterodorsal surface (arrow, Fig. 13D). Due to an extremely similar habitus, it is difficult to distinguish D. pacificus from D. pauxillus . Male genitalia should be used to conclusively identify specimens, especially in the overlap zone of these species’ ranges in southern British Columbia.</p><p>Redescription:</p><p>Body Length: 5.0– 8.5mm; Wing Length: 5.0– 7.7mm</p><p>Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in female it is ½ the length between the antennae and ocelli) and dark pile; face light with dark vitta ¼ the width of the face, reaching antennal socket, face with dark pile; gena dark with some light pollen ventral to eye, dark pile dorsally and light pile ventrally; occiput dark, covered in light pollen, pile light with some dark pile dorsally; scape, pedicel and flagellomere light to dark.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile light; scutellum light to dark with darker anterior and lateral edges, light pile with some dark pile posteriorly; wing cell br bare above spurious vein and usually on basal half, bm bare medially or basal half, cells c and r 1 bare at base or with at least small bare areas at base, cup call has small bare areas along vein CuP, wing less frequently completely microtrichose (Fig. 13C); haltere light to dark; pro- and mesofemora ⅓–½ dark basially, ½–⅔ light apically, metafemur ⅔–¾ dark basially, ¼–⅓ light apically, tibiae light, metatibia with dark band ¼ of the way from the apex, tarsi light, sometime dark anteriorly.</p><p>Abdomen: Maculae on tergite 2 large, slender and oval, ⅓–½ width of tergite, maculae on tergites 3&amp;4 are slightly arcuate, uniform in thickness, with out a distinct swelling medially, maculae do not reach abdominal margin (Figs 13A &amp; B); sternites light with dark fasciae.</p><p>Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened posteriorly, long pile on dorsal, laterodorsal and posterodorsal sides, row of spines on posteromedial edge; cercus oval with long pile over entire surface; gonostylus triangular-shaped with narrow, wavy projection on posterodorsal surface projecting posterodorsally, pile on dorsal surface; basiphallus elongate, basally bent at 90 degree angle towards dorsum, apical end curved ventrally with a widened apex and is partly membranous dorsally, with spines on posterodorsal side; distiphallus elongate, apical end flared out into horn, which is mostly sclerotized but membranous at apex with few minute setulae dorsally, basally bent towards dorsum and usually greatly enlarged at bend with a triangular projection, projecting ventrally (Fig. 13D).</p><p>Intraspecific variation: Distiphallus sometimes without triangular projection basally. Abdominal maculae on tergites 3 &amp; 4 rarely meet medially, and if so, barely touch.</p><p>Etymology: Pacificus is Latin for peaceful or peace-making, but more probably was named so because this species is found along the Pacific coast of North America.</p><p>Distribution: Southwestern USA (CA, NV, OR, WA) and southwestern Canada (BC); Cascades and Sierra Nevada Mountain Ranges (Fig. 21C).</p><p>Ecology:</p><p>Collected from April-July, but more commonly in May and June, at elevations of 475–3100m. Recorded from willow ( Salix Linnaeus, Salicaceae) and bitterbush ( Purshia de Candolle ex Poiret, Rosaceae).</p><p>Discussion: Dasysyrphus pacificus is hereby resurrected from synonymy with Dasysyrphus pauxillus (synonymized by Vockeroth (1986)). The type specimen of this species could not be examined as it is out on loan to another individual. However, based on the written description and distribution of D. pacificus (Cole &amp; Lovett 1919) the authors are confident that the individuals examined were of this species. The range of pacificus is from southern British Columbia, south to California, in the Cascades and Sierra Nevada mountain ranges . The type specimen is from Oregon, where it is the only species of the pauxillus s.l. in this range.</p><p>Type Material: “ Holotype (No. 502), male, in Museum of the California Academy of Sciences, collected at Corvallis, Ore., May 17, by A. L. Lovett” (Cole &amp; Lovett 1919). (not examined)</p><p>Material Examined:</p><p>Canada: British Columbia: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.740555&amp;materialsCitation.latitude=49.300556" title="Search Plazi for locations around (long -121.740555/lat 49.300556)">Bear Mountain</a>, [49.300555, -121.740556], 6.v.1920, ♂, CNCD 13743, (CNC). EAST KOOTENAY R . D.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.761505&amp;materialsCitation.latitude=49.517982" title="Search Plazi for locations around (long -115.761505/lat 49.517982)">Cranbrook</a>, [49.517983, -115.761504], 19.v.1920, ♂, CNCD 13748 , C.B.D. Garrett (CNC); 27.iv. ♂, RBCM ENT 99188108, H. R. Foxlee (RBCM). 7 miles N Lions Bay, North of Vancouver, 18.v.1979, ♂, CNCD 5464 , G.F. Hevel (USNM) . United States of America: California: RIVERSIDE CO.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.71667&amp;materialsCitation.latitude=33.816666" title="Search Plazi for locations around (long -116.71667/lat 33.816666)">Dark Canyon</a> N of Idyllwild, [33.81666667, -116.7166667], 24.v.2003, CNCD 8472 , J. Skevington (CNC). SAN BERNARDINO CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.12074&amp;materialsCitation.latitude=34.09125" title="Search Plazi for locations around (long -117.12074/lat 34.09125)">Upper Santa Ana River</a>, [34.091253, -117.120741], 3.vi.1947, ♀, CNCD 26415 , G.H. &amp; J.L. Sperry (CNC). 1 km NW Wrightwood, Big Pines Hwy, [34.38138889, -117.6716667], 1974m, 21–28.v.2005, 5♂, JSS 20974, 20976–20978, 20980, 2♀, JSS 20975, 20979, A. R . Cline &amp; S.L. Winterton (MHPC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.649315&amp;materialsCitation.latitude=37.328" title="Search Plazi for locations around (long -119.649315/lat 37.328)">Oakhurst</a>, [37.328, -119.649315], 1.vi.1942, ♀, JSS 20598, E.C. Linsley (EMEC). TUOLUMNE CO .: Mather Site, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.85698&amp;materialsCitation.latitude=37.880173" title="Search Plazi for locations around (long -119.85698/lat 37.880173)">Buck Meadows</a>, [37.880173, -119.856977], 1970, ♂, CNCD 26437 , 4♀, CNCD 26404–26407, A. R. Moldenke (CNC). MARIN CO .: Lily Pond, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.63498&amp;materialsCitation.latitude=37.953804" title="Search Plazi for locations around (long -122.63498/lat 37.953804)">Alpine Lake</a>, [37.953803, -122.634977], 457m, 25.v.1970, ♀, CNCD 26414 , 11.iv.– 25.v.1971, ♂, CNCD 26344, D.D. Munroe (CNC). CALAVERAS BIG TREES SP: S. grove, banks of Beaver Creek, MT#2 , [38.26166667, -120.2577778], 1375m, 22.v.–11.vi.2007, 6♀, JSS 20966–20970, 20972, A. R . Cline &amp; P.H. Kerr (MHPC). INDIAN GRINDING ROCK SP: dry wash nr. S. Nature Trail MT #2 , [38.41666667, -120.6333333], 715m, 5.iv.–24.iv.2007, JSS 20981, ♀, JSS 20992, P. Kerr &amp; M. Hauser (MHPC); firebreak nr. envtl camp MT3, [38.41861111, -120.635], 715m, 24.iv.2007 24.v.2007, ♀, JSS 20971, P. Kerr &amp; M. Hauser (MHPC). Sonora Pass, [38.53666667, -119.6855556], 3100m, 21.vi.2007, ♀, JSS 20963, S.M. Blank (MHPC). Wright’s Lake, [38.84913, -120.2318], 2.vii.1948, ♂, JSS 20597, D. Carter (EMEC); 2.vii.1948, ♀, JSS 20601, L.W. Quate (EMEC). Placer County, [39.090423, -120.803069], 1524m, iv, ♀, JSS 19511, W.M. Davidson (CNC). Tahoe City, Alpine Meadow Road Jenkinson Reservoir, [39.42138889, -120.3333333], 2200m, 14.vi.2007, ♂, JSS 20957, ♀, JSS 20958, S.M. Blank (MHPC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-120.333336&amp;materialsCitation.latitude=39.42139" title="Search Plazi for locations around (long -120.333336/lat 39.42139)">Sinkyone Wilderness State Park</a>, [39.83583333, 123], 28m, 23.iii.2003, ♂, JSS 20797, J. &amp; A. Skevington (CNC). PLUMAS CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.185776&amp;materialsCitation.latitude=39.89096" title="Search Plazi for locations around (long -121.185776/lat 39.89096)">Buck’s Lake</a>, [39.890961, -121.185778], 13.v.1949, ♀, CNCD 26409 , P.D. Hurd (CNC), 4♂, CNCD 26432–26435, ♀, CNCD 26410, W.W. Middlekauff (CNC); 23.vi.1949, 2♀, CNCD 26412, D. Cox (CNC), ♀, CNCD 26413, J.W. MacSwain (CNC); 13.v.1949, ♂, JSS 20594, E.O. Essig (EMEC), ♂, JSS 20595, ♀, JSS 20603, P.D. Hurd (EMEC); 23.vi.1949, 2♀, JSS 20599, 20600, J.W. MacSwain (EMEC). TEHAMA CO .: 60 kilometers <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.521385&amp;materialsCitation.latitude=40.19611" title="Search Plazi for locations around (long -121.521385/lat 40.19611)">North East of Chico</a> on Road 32, [40.19611111, -121.5213889], 1100m, 15.v.2010, JSS 25002, 25003, 25005–25009, D. Striley, M. Hauser (CSCA). LASSEN CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.09448&amp;materialsCitation.latitude=40.492317" title="Search Plazi for locations around (long -121.09448/lat 40.492317)">Susan River Campground</a>, [40.492316, - 121.094484], 9.vii.1949, ♀, CNCD 26411 , A. T. McClay (CNC). SHASTA CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.68097&amp;materialsCitation.latitude=40.51792" title="Search Plazi for locations around (long -121.68097/lat 40.51792)">Viola</a>, [40.517921, -121.680971], 19.v.1941, ♂, CNCD 26436 , ♀, CNCD 26408, E.G. Linsley (CNC). LASSEN NATIONAL PARK: Manzanita Lake, [40.534513, 121.56655], 8.vi.1941, ♀, JSS 20596, C.D. Michener (EMEC), ♀, JSS 20602, Arthur J Walz (EMEC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.56655&amp;materialsCitation.latitude=40.53451" title="Search Plazi for locations around (long 121.56655/lat 40.53451)">Ryan Creek</a>, 17.v.1938, ♀, CNCD 106796 , N.F. Hardman (CNC). San Bernardino Mountains; Up Sta. Ana River, 15.vi.1955, ♀, JSS 23775; 1957, ♀, JSS 26926, A.L. Melander (USNM). Nevada: WASHOE CO.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-119.951584&amp;materialsCitation.latitude=39.80685" title="Search Plazi for locations around (long -119.951584/lat 39.80685)">Little Valley</a>, [39.806851, -119.951585], 4.vi.1981, JSS 25100, R . W. Rust (EMUS). Oregon: KLAMATH CO .: Lake of the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.20848&amp;materialsCitation.latitude=42.368633" title="Search Plazi for locations around (long -122.20848/lat 42.368633)">Woods</a>, [42.368634, -122.208482], 22.vi.1975, JSS 25082–25086, 25088–25095, G.F. Knowlton (EMUS), ♀, CNCD 5457 , G.F. Knowlton (USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.11877&amp;materialsCitation.latitude=43.454906" title="Search Plazi for locations around (long -122.11877/lat 43.454906)">Summit Lake</a>, [43.454905, -122.118766], 7.vii.1952, ♂, CNCD 26442 , E. R. Jaycox (CNC). DESCHUTES CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.43545&amp;materialsCitation.latitude=43.874554" title="Search Plazi for locations around (long -121.43545/lat 43.874554)">Camp Abbot</a>, [43.874555, -121.435449], 4.vi.1944, ♀, CNCD 26401 , P.H. Arnaud (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.35528&amp;materialsCitation.latitude=44.36361" title="Search Plazi for locations around (long -123.35528/lat 44.36361)">Bellfountain</a>, [44.363611, -123.355278], 27.v.1922, ♂, OSAC 133389, 133391 , A.L. Lovett (OSAC). BENTON CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.26203&amp;materialsCitation.latitude=44.564564" title="Search Plazi for locations around (long -123.26203/lat 44.564564)">Corvallis</a>, [44.564564, -123.262033], v.1977, ♂, JSS 19507, A.L. Lovett (CNC) .</p><p>Washington: WENATCHE NATIONAL FOREST: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.378784&amp;materialsCitation.latitude=46.64332" title="Search Plazi for locations around (long 121.378784/lat 46.64332)">White Pass Trail</a>, [46.643317, 121.378783], 1345m, 6.vii.2010 , CNCD 60054, A. Young (CNC), ♂, CNCD 35314, ♀, CNCD 35313, M.M. Locke (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-121.75313&amp;materialsCitation.latitude=46.853046" title="Search Plazi for locations around (long -121.75313/lat 46.853046)">Mount Rainier</a>, [46.853048, -121.753127], ♂ , CNCD 26429, ♀, CNCD 26400, 26419, F.M. Hull (CNC). SAN JUAN CO.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.83546&amp;materialsCitation.latitude=48.61955" title="Search Plazi for locations around (long -122.83546/lat 48.61955)">Olga</a>, [48.619548, -122.835459], 18.v.1910, ♀ , JSS 19504, (CNC) . MOUNT BAKER- SNOQUALMIE NATIONAL FOREST, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.67654&amp;materialsCitation.latitude=48.86622" title="Search Plazi for locations around (long 121.67654/lat 48.86622)">Picture Lake</a>, [48.866217, 121.676533], 1255m, 8.vii.2010 , CNCD 60058, A. Young (CNC). Price Co.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-122.70333&amp;materialsCitation.latitude=48.90667" title="Search Plazi for locations around (long -122.70333/lat 48.90667)">Pleasant Valley</a>, [48.90667, -122.70333], 21.v.1977, JSS 25097 , T.L. Whitworth (EMUS) .</p></div>	https://treatment.plazi.org/id/03893F32FF85FF98FF46FC80FBB18E78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FF8AFF9CFF46FE50FE928DF3.text	03893F32FF8AFF9CFF46FE50FE928DF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus pauxillus (Williston 1887)	<div><p>Dasysyrphus pauxillus (Williston)</p><p>Figures: 14A–D, 18A &amp; D, 21D</p><p>Synonyms and References:</p><p>Syrphus pauxillus Williston 1887: 74 (original description)</p><p>Metasyrphus pauxillus:</p><p>Fluke 1933: 115 (key, redescription and transfer to Metasyrphus)</p><p>Dasysyrphus pauxillus:</p><p>Stone et al. 1965: 564 (catalogue and transfer to Dasysyrphus)</p><p>Vockeroth 1986: 203 (discussion)</p><p>Vockeroth 1992: 68 (key and redescription)</p><p>Speight 2011: 70 (discussion)</p><p>Diagnosis: Abdominal maculae of tergites 3&amp;4 slightly arcuate, uniform in thickness and do not cross abdominal margin (Figs 14A &amp; B); very similar in appearance to D. pacificus (Fig. 13A), D. nigricornis (Figs 11A &amp; B) and D. laticaudus (Fig. 8A). Wing typically densely microtrichose, sometimes with bare areas in cells r 1, br, bm and/or c (Fig. 14C). Male gonostylus with projection on posterodorsal surface straight (Fig. 14D). Due to an extremely similar habitus, it is difficult to distinguish D. pacificus from D. pauxillus . Male genitalia should be used to conclusively identify specimens, especially in the overlap zone of these species’ ranges in southern British Columbia.</p><p>Redescription:</p><p>Body Length: 5.3–9.7mm; Wing Length: 5.2–9.5mm</p><p>Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in females it is ⅓–½ the length between the antennae and ocelli with a separation medially and edges triangular to rounded (Fig. 18A)) and light to dark pile; face light with dark vitta ¼–⅓ the width of the face, coming close to antennal socket if not reaching it, face with dark pile lateral to the eye, some females with varying amounts of light pile interspersed with dark pile; gena with light to dark pile; occiput dark, covered in light pollen, pile light; scape and pedicel light to dark, flagellomere dark with lighter base.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile light to dark; scutellum light to dark with darker lateral edges, pile on female mostly light with some dark, male mostly dark pile some with light interspersed throughout; wing cell r 1 with the base bare or very small areas at base bare, br, bm and c may all be entirely microtrichose or they may have bare areas along spurious vein, along the middle of the cell form the base to ¾ or small areas at base, respectively (Fig. 14C); haltere light; pro- and mesofemora ⅓–⅔ dark basally, ⅓–⅔ light apically, metafemur ⅔–¾ dark basally, ¼–⅓ light apically, tibiae light, meso- and metatibiae with dark band ¼ of the way from the apex, tarsi light with dark anterior side.</p><p>Abdomen: Maculae on tergite 2 large and oval, lateral edge tends to be more flattened than medial edge, female usually with very small, pointed, anterolateral projection, but it never reaches the edge of the tergite, maculae on tergites 3&amp;4 are oblique, fairly straight and uniform in width, sometimes with medial edge slightly swollen, maculae do not reach abdominal margin (Figs 14A &amp; B); sternites light with dark fasciae of varying thickness (Fig. 18D).</p><p>Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened posteriorly, long pile on dorsal, laterodorsal and posterodorsal sides, row of spines on posteromedial edge; cercus oval with long pile over entire surface; gonostylus boot-shaped but more narrow apically than basally, with projection on posterodorsal surface, projecting dorsally, pile on dorsal surface; basiphallus elongate, basally bent at a slightly more than 90 degree angle towards dorsum, apical end curved ventrally, but does not greatly extend ventrally and is partly membranous dorsally, with spines on posterodorsal side; distiphallus elongate, apical end flared out into horn, which is mostly sclerotized but membranous at apex, with no setulae dorsally, basally bent towards dorsum and usually greatly enlarged at bend with a triangular projection, projecting ventrally (Fig. 14D). Extremely similar to D. laticaudus (Fig. 8C).</p><p>Intraspecific variation: Wings are typically densely microtrichose with cell r 1 having bare areas at base ranging from entire base bare to very small areas at base being bare, cells br, bm and c may all be entirely microtrichose or they may have bare areas; female usually have a very small, anterolateral projection on maculae of tergite 2, males do not have this projection.</p><p>Etymology: From the Latin pauxillus, meaning small.</p><p>Distribution: Rocky Mountains; Western Canada (AB, BC, MB, YT) and western USA (AK, CO, ID, NM, UT, WY) (Fig. 21D).</p><p>Ecology: Collected from March–July, but more commonly collected in June. Collected at elevations of 14– 3505m. Habitats collected from are listed as “moraine lake”, “grassy lakeshore”, “over stream in fir forest”, “on tundra”, “spruce forest” and “ Pinus-Salix forest”. It has been collected from the following plant species: common cow parsnip ( Heracleum maximum W. Bartram, Apiaceae), blackberry ( Rubus Linnaeus, Rosaceae), willow ( Salix Linnaeus, Salicaceae), Canada gooseberry ( Ribes oxyacanthoides Linnaeus, Grossulariaceae) and spruce ( Picea A. Dietrich, Pinaceae).</p><p>Discussion: Many authors have used the name pauxillus in the Palaearctic. During the course of this revision few specimens from that region were examined and no DNA evidence could be studied. Therefore, we cannot comment on whether the use of this name is appropriate in the Palaearctic. However, based on the geographic breakdown of pauxillus and its allies ( laticaudus, pacificus and nigricornis) in the Nearctic it seems likely that only nigricornis is Holarctic. European and Asian specimens should be studied in greater depth, using both morphology and DNA, to confirm this suspicion.</p><p>Due to the widespread use of the name pauxillus much confusion exists over the concept of this species. Many Holarctic species have been named which have an extremely similar habitus to pauxillus . The main reason for such confusion is that the holotype specimen is lost (Fluke 1933). In Williston’s (1887) original designation he describes one female from New Mexico. He does not indicate where this specimen is deposited. The only other information that is provided is that the specimen was collected by G.F. Gaumer. In Fluke’s (1933) revision of Metasyrphus he indicates that he was unable to locate the type for examination. No record exists in the literature of any author examining Williston’s specimen; either Williston’s species description was used or specimens determined by other researchers were used for comparison. Dasysyrphus pauxillus was described in ‘Synopsis of the North American Syrphidae’, Williston’s PhD dissertation (Beckemeyer 2009). The specimens from his dissertation were donated to the USNM (Flint et al. 1976, F.C. Thompson, pers. comm.). According to Thompson (pers. comm.) the type specimen is not in the USNM collection today and it is not known what happened to it. Williston worked in Kansas and Chicago until his death, after which his collection was donated to the American Museum of Natural History (AMNH). Byers et al. (Byers et al. 1962) do not list pauxillus as a type in the Kansas collection and the specimen is not listed in the type database of the AMNH (The American Museum of Natural History 2005). A neotype is designated here to bring stability to the name pauxillus . This will create a solid concept for this species, allowing for better reconciliation of closely related species. The specimen used in the original designation provided a locality of ‘New Mexico’. The neotype specimen was chosen from a series of males from Utah, as the two specimens from New Mexico were a female and a male in poor condition. This is a locality that is not overlapped by the other species with a similar habitus to pauxillus ( pacificus, laticaudus and nigricornis). The neotype varies from Williston’s description in a few characters that are variable in many Dasysyrphus species. Also, Williston’s description was of a female and the neotype is male, which may account for some of the variation. The dorsum of the scutum in the neotype is covered by mainly dark pile, Williston’s description states the pile is light yellowish. This character is variable within most Dasysyrphus species and often males have more dark pile and females have more light pile on the scutum. The face is also not projecting to the extent described by Williston. The face seems to project slightly more in females than males, but in most specimens the distance between the antennae and the tip of the tubercle is more or less equal to the distance from the antennae to the ocelli. The specimen matches the description in all other characters. The Neotype is a male from Tony Grove Lake, Wasatch-Cache National Forest, UT, USA, collected on 1.vii.2010 by M.M. Locke. It is deposited in the CNC and databased with the number Jeff Skevington Specimen 24616.</p><p>Type Material: “ USA: UT:Wasatch-CacheNF:/ Tony Grove Lake;/ 41.893136°N, 111.641841°W;/ 1.vii.2010; M.M.Locke ”, “ J. Skevington / Specimen #/ 24616”, “ NEOTYPE / Syrphus / pauxillus/ Williston”, “designated by/ Locke &amp;/ Skevington/ 2013”; ♂, deposited in the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.64184&amp;materialsCitation.latitude=41.893135" title="Search Plazi for locations around (long -111.64184/lat 41.893135)">Canadian National Collection of Insects</a>, Arachnids and Nematodes, Ottawa, Ontario , Canada.</p><p>Material Examined:</p><p>Canada: Alberta: WATERTON LAKES NATIONAL PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-113.911804&amp;materialsCitation.latitude=49.05112" title="Search Plazi for locations around (long -113.911804/lat 49.05112)">Waterton</a>, [49.051122, -113.911807], 6.vi.1962, 2♂, CNCD 13764, 13765 , 4♀, CNCD 13836–13839, W. R. M. Mason (CNC), 2♀, CNCD 13834, 13835, K.C. Herrmann (CNC). KANANASKIS: 25 km WNW Bragg Creek Powderface trail <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-114.901665&amp;materialsCitation.latitude=51.021667" title="Search Plazi for locations around (long -114.901665/lat 51.021667)">Rd.</a>, [51.02166667, -114.9016667], 8.vi.2001, ♂, RBCM ENT 6006051 , C.J. Borkent (RBCM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.08769&amp;materialsCitation.latitude=51.10088" title="Search Plazi for locations around (long -115.08769/lat 51.10088)">Forest Experimental Station Seebe</a>, [51.100879, - 115.087692], 2.vi.1968, ♀, CNCD 13840 , 13.vi.1968, ♀, CNCD 13841, 15.vi.1968, ♀, CNCD 13842, H.J. Teskey (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-113.63133&amp;materialsCitation.latitude=51.04575" title="Search Plazi for locations around (long -113.63133/lat 51.04575)">30 miles West of Calgary</a>, [51.04575, -113.631333], 3.vi.1973, CNCD 60100 , G.G.E. Scudder (CNC). BANFF NATIONAL PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.568436&amp;materialsCitation.latitude=51.180275" title="Search Plazi for locations around (long -115.568436/lat 51.180275)">Banff</a>, [51.180275, -115.568433], 27.v.1925, ♂, CNCD 13762 , E. Hearle (CNC); 2.vi.1922, ♂, CNCD 13763, 3♀, CNCD 13845–13847, 9.vi.1922, ♀, CNCD 13849, 3.vii.1922, ♀, CNCD 13848, C.B.D. Garrett (CNC); 2.vi.1914, ♀, JSS 20605, N.B. Sanson (EMEC); 30.v.1948, ♀, UASM# 149024, 2.vi.1948, ♀, UASM# 149025, E.H. Strickland (UASM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.92572&amp;materialsCitation.latitude=51.298695" title="Search Plazi for locations around (long -115.92572/lat 51.298695)">Mount Eisenhower</a>, [51.298694, -115.925724], 3.vi.1968, 2♂, CNCD 13770, 13771 , 5.vi.1968, ♂, CNCD 13769, 7.vi.1968, ♂, CNCD 13768, Mosquin &amp; Seaborn (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.18442&amp;materialsCitation.latitude=51.324074" title="Search Plazi for locations around (long -116.18442/lat 51.324074)">Moraine Lake</a>, [51.324073, -116.184418], 2256m, 20.vii.1955, ♀, CNCD 13850 , J. R. McGillis (CNC); 8 km West of Lake Louise, [51.413684, -116.306046], 31.v.1992, ♀, JSS 20801, A. Borkent (CNC); Lake Agnes, [51.415101, -116.247314], 2073m, 10.vii.1925, ♂, JSS 19508, O. Bryant (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-115.80862&amp;materialsCitation.latitude=51.60524" title="Search Plazi for locations around (long -115.80862/lat 51.60524)">Snow Creek Pass</a>, [51.60524, -115.808617], 2256m, 20.vii.1962, ♀, CNCD 13844 , K.C. Herrmann (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.50892&amp;materialsCitation.latitude=51.717632" title="Search Plazi for locations around (long -116.50892/lat 51.717632)">Peyto Lookout</a>, [51.717631, -116.508919], 2316m, 23.vii.1962, ♀, CNCD 13843 , W. R. M. Mason (CNC). Edmonton, 12.v.1924, ♂, JSS 23724, Owen Bryant (CAS); Laggen, Lake Agnes, 10.vii.1925, ♀, JSS 23714, O. Bryant (CAS). Edmonton; W. Hawrelak Park, 640m, 3.v.2005, 3♂, JSS 25540–25542, W. van Steenis (WVSC). Pincher Creek; Beauvais Lake, 19.v.1989 23.v.1989, ♀, JSS 25550, C. v. Nidek (WVSC) . British Columbia: Victoria, Mt. Work, [48.53, -123.47], 26.v.1982, ♂, RBCM ENT 99188111, R . A. Cannings (RBCM). Galiano I. North End, [48.92, -123.42], 29.iii.1987, ♂, JSS 20184, G.G.E. Scudder (UBCZ). WATERTON LAKES NATIONAL PARK : <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-114.04924&amp;materialsCitation.latitude=49.011456" title="Search Plazi for locations around (long -114.04924/lat 49.011456)">Cameron Lake</a>, [49.011457, -114.049238], 17.vi.1956, 9♂, CNCD 13754–13761, 13767 , 9♀, CNCD 13822–13830, 18.vi.1956, ♀, CNCD 13831, 19.vi.1956, 2♀, CNCD 13832, 13833, E.E. Sterns (CNC). CENTRAL KOOTENAY L.D.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-117.69287&amp;materialsCitation.latitude=49.333015" title="Search Plazi for locations around (long -117.69287/lat 49.333015)">Robson</a>, [49.333015, -117.692871], 21.iv.1947, ♂, CNCD 13746 , 8.v.1950, ♂, CNCD 13750, H. R. Foxlee (CNC); 25.iv.1958, ♂, JSS 20123, H. R . Foxlee (UBCZ); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.74674&amp;materialsCitation.latitude=49.414406" title="Search Plazi for locations around (long -116.74674/lat 49.414406)">Boswell</a>, [49.414407, -116.746741], 9.v.1958, ♂, CNCD 13742 , H. &amp; A. Howden (CNC). SQUAMISH- LILLOOET R . D.: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-123.48634&amp;materialsCitation.latitude=50.56162" title="Search Plazi for locations around (long -123.48634/lat 50.56162)">Meager Creek Hot Springs</a>, [50.561617, -123.486344], 14.v.1987, ♂, JSS 18370, C.S. Guppy (CNC). KOOTENAY NATIONAL PARK : Daer-Pitts Aspen Control 1, [50.95, -116.0333333], 3.vi.2000 17.vi.2000, CNCD 7969 , G. Gareau (CNC); Daer-Pitts Aspen Control 2, 17.vi.2000 2.vii.2000, ♀, CNCD 30906 , G. Gareau (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-129.14029&amp;materialsCitation.latitude=54.194077" title="Search Plazi for locations around (long -129.14029/lat 54.194077)">32 miles South West of Terrace</a>, [54.194076, -129.140296], 30.5m, 6.vi.1960, ♀, CNCD 13819 , G.E. Shewell (CNC); 6.vi.1960, ♀, CNCD 13820, 11.vi.1960, ♂, CNCD 13753, B. Heming (CNC). Shames, 18 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-128.9353&amp;materialsCitation.latitude=54.40965" title="Search Plazi for locations around (long -128.9353/lat 54.40965)">Miles South West of Terrace</a>, [54.409648, -128.935301], 23.vi.1960, ♀, CNCD 13818 , C. H. Mann (CNC). Kleanza Creek, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-128.38625&amp;materialsCitation.latitude=54.597897" title="Search Plazi for locations around (long -128.38625/lat 54.597897)">Near Terrace</a>, [54.597897, -128.386241], 76.2m, 29.vi.1960, ♀, CNCD 13817 , B. Heming (CNC); 30.vi.1960, ♀, CNCD 13821, J.G. Chillcott (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-133.69237&amp;materialsCitation.latitude=59.5775" title="Search Plazi for locations around (long -133.69237/lat 59.5775)">Atlin</a>, [59.5775, -133.69236], 670.56m, 2.vi.1955, ♂, CNCD 13751 ,. vii.1955, ♀, CNCD 13809, H.J. Huckel (CNC); 20.vi.1955, ♂, CNCD 14358, ♀, CNCD 13808, B.A. Gibbard (CNC) . Manitoba: 26km SE Churchill; Twin Lakes, [58.63, -93.819], 14m, 19.vii.2009, ♀, 09 PROBE 197 , 22.vii.2009, ♂, 09 PROBE 255, Arctic Ecology 2009 (CNC) . Yukon Territory: Snafu Creekl, [60.13333333, -138.8], 22.vi.1982, CNCD 60079 , S.G. Cannings, R . D. Wilkie, L. Vasington (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-135.05222&amp;materialsCitation.latitude=60.71965" title="Search Plazi for locations around (long -135.05222/lat 60.71965)">Whitehorse</a>, [60.71965, -135.052213], 6.vii.1988, ♀, CNCD 5812 , F. Brodo (CNC). Simpson L. 81 km n Watson L., [60.75, -129.25], 15.vi.1984 1.ix.1984, ♀, JSS 20770, S. &amp; J. Peck (CNC). Nines Creek, 3 km West of Alaska Highway, [61.18333333, -139.7], 4.vi.1981, CNCD 60072 , C. Guppy (CNC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-132.33333&amp;materialsCitation.latitude=62.683334" title="Search Plazi for locations around (long -132.33333/lat 62.683334)">La Force Lake</a>, [62.683333, -132.333333], 914m, 26.vi.1960, ♀, CNCD 13585 , 10.vii.1960, ♀, CNCD 13728, E.W. Rockburne (CNC) . United States of America: Alaska: MATANUSKA- SUSITNA CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-149.22968&amp;materialsCitation.latitude=61.541946" title="Search Plazi for locations around (long -149.22968/lat 61.541946)">Matanuska</a>, [61.541945, -149.229674], 12.vi.1944, ♀, JSS 19502, J. Chamberlin (CNC) . Colorado: Rocky Gulch, [37.817035, -107.579606], 3444m, 6.xxii.2002, ♂, JSS 20810, C. Slater (CSUC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.32774&amp;materialsCitation.latitude=38.501434" title="Search Plazi for locations around (long -106.32774/lat 38.501434)">Monarch Pass</a>, 2mi W Garfield, [38.501433, -106.327747], 3048m, 19.vi.1985, ♂, USNM ENT 258074 , David R . Smith (USNM). Rock Cr., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.111&amp;materialsCitation.latitude=39.425" title="Search Plazi for locations around (long -106.111/lat 39.425)">May Mus.</a>, [38.705655, -104.837421], 8.v.1996, ♀, JSS 20808, D. Leatherman (CSUC). <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.111&amp;materialsCitation.latitude=39.425" title="Search Plazi for locations around (long -106.111/lat 39.425)">Tenmile Range</a>, unamed <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.111&amp;materialsCitation.latitude=39.425" title="Search Plazi for locations around (long -106.111/lat 39.425)">Cr.</a> avalanche chute W side, [39.425, -106.111], 3200m, 15.vii.1998 17.vii.1998, ♂, JSS 20694, S. Fitzgerald &amp; A. Foley (CSUC). CLEAR CREEK CO .: Mt. Evans, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.59504&amp;materialsCitation.latitude=39.65622" title="Search Plazi for locations around (long 105.59504/lat 39.65622)">Echo Lake Campground</a>, [39.65622, 105.595042], 329m, 29.vi.2010, CNCD 60035 , M.M. Locke (CNC); Mt. Evans, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.6035&amp;materialsCitation.latitude=39.658085" title="Search Plazi for locations around (long -105.6035/lat 39.658085)">Echo Lake</a>, [39.658085, - 105.6035], 3231m, 13.vii.1961, ♀, CNCD 26416 , C.H. Mann (CNC); 29.vi.2010, ♀, JSS 24618, M.M. Locke (CNC); 29.vi.2010, ♀, CNCD 35312, A. Young (CNC); Mt. Evans, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.60135&amp;materialsCitation.latitude=39.67574" title="Search Plazi for locations around (long -105.60135/lat 39.67574)">Doolittle Ranch</a>, [39.675739, -105.601348], 2987m, 13.vii.1961, ♀, CNCD 26417 , W. R. M. Mason (CNC). BOULDER CO .: Nederland, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.53389&amp;materialsCitation.latitude=40.031647" title="Search Plazi for locations around (long -105.53389/lat 40.031647)">Science Lodge</a>, [40.031645, -105.533886], 2896m, 4.vii.1961, ♀, CNCD 26418 , B.H. Poole (CNC); Niwot Ridge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.6074&amp;materialsCitation.latitude=40.069935" title="Search Plazi for locations around (long -105.6074/lat 40.069935)">Near Ward</a>, [40.069933, -105.607397], 3444m, 4.vi.1961, ♂, CNCD 26440 , 28.vi.1961, ♂, CNCD 26439, C.H. Mann (CNC). North Park, [40.73, -106.18], 1.vii.1932, ♀, JSS 20746, (CSUC). Rawah Wilderness, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.47917&amp;materialsCitation.latitude=40.90972" title="Search Plazi for locations around (long -105.47917/lat 40.90972)">Blue Lake Trail</a> near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.47917&amp;materialsCitation.latitude=40.90972" title="Search Plazi for locations around (long -105.47917/lat 40.90972)">Fall Creek</a>, [40.76944444, -105.9894444], 2926m, 20.vi.1998, ♂, JSS 20693, S. Fitzgerald &amp; A. Foley (CSUC). Cherokee Park, [40.90972, -105.47917], 30.vii.1913, ♀, JSS 20713, Chas L. Fluke (CSUC). MINERAL CO .: Continental Divide, 3292m, 20.vi.1919, ♂, JSS 19512, (CNC). Camera Point, Pike’s Peak, 16.vi.1956, ♀, JSS 21157, R . &amp; K. Dreisbach (MSUC) . Idaho: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.37106&amp;materialsCitation.latitude=44.424362" title="Search Plazi for locations around (long -111.37106/lat 44.424362)">Island Park</a>, [44.424364, -111.371064], JSS 25099, Bernard Knapp (EMUS). NEZ PERCE CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.83701&amp;materialsCitation.latitude=46.20594" title="Search Plazi for locations around (long -116.83701/lat 46.20594)">Lake Waha</a>, [46.205942, -116.837015], 9.vi.1918, ♀, CNCD 26402 , A.L. Melander (CNC); 18.vi.1930, ♂, JSS 19514, W.M. Davidson (CNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.84444&amp;materialsCitation.latitude=46.211437" title="Search Plazi for locations around (long -116.84444/lat 46.211437)">Waha</a>, [46.211439, -116.844439], 18.vi.1909, ♀, JSS 19501, (CNC); 30.v.1924, ♀, JSS 23778, A.L. Melander (USNM). LATAH CO .: Moscow Mt., [46.800972, - 116.866093], 8.vi.1921, ♀, JSS 23776, 26.vi.1922, ♀, JSS 23777, A.L. Melander (USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-116.86897&amp;materialsCitation.latitude=46.80726" title="Search Plazi for locations around (long -116.86897/lat 46.80726)">Mountains Near Moscow</a>, Latah County, [46.807258, -116.868969], 1.vi.1907, ♀, CNCD 26403, (CNC); 17.vi.1918, ♂, JSS 19509, 26.vi.1920, ♂, JSS 19506, 9.vii.1920, ♀, JSS 19505, A.L. Melander (CNC); vii.1910, ♀, JSS 19510, 25.vi.1920, ♂, JSS 19516, R . C. Shannon (CNC). Savary, 29.iv.1916, ♀, JSS 21329, (MCZ) . New Mexico: Las Cruces, Dona Ana, [32.3, -106.7666667], ♀, CNCD 26423 , F.M. Hull (CNC). LINCOLN CO ., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.55291&amp;materialsCitation.latitude=33.478065" title="Search Plazi for locations around (long -105.55291/lat 33.478065)">Sierra Blanca</a>, [33.478065, - 105.552907], 2957m, 10–26.vi.1979, ♂, JSS 18307, S. &amp; J. Peck (CNC). SAN MIGUEL CO ., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-105.54573&amp;materialsCitation.latitude=35.705997" title="Search Plazi for locations around (long -105.54573/lat 35.705997)">Santa Fe National Forest</a>, [35.705996, -105.545731], 27.v.1981, ♀, JSS 18346, B.J. Sinclair (CNC). SANDOVAL CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.69055&amp;materialsCitation.latitude=35.771336" title="Search Plazi for locations around (long -106.69055/lat 35.771336)">Jemez Springs</a>, [35.771336, -106.690555], iv, ♀, CNCD 26422, (CNC) . Utah: WASATCH CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.22368&amp;materialsCitation.latitude=40.280033" title="Search Plazi for locations around (long -111.22368/lat 40.280033)">Daniels Pass</a>, 2 miles South, [40.280034, -111.223675], 2591m, 9.vii.1961, ♀, CNCD 26421 , J.G. Chillcott (CNC). DUSCHENE CO .: 2 miles <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.92851&amp;materialsCitation.latitude=40.70363" title="Search Plazi for locations around (long -110.92851/lat 40.70363)">West of Mirror Lake</a>, [40.70363, -110.928511], 23.vi.1968, ♂, CNCD 26438 , B. V. Peterson (CNC). CACHE CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.66348&amp;materialsCitation.latitude=41.76236" title="Search Plazi for locations around (long -111.66348/lat 41.76236)">Card Canyon</a>, [41.76236, -111.663482], 13.vi.1937, ♀, CNCD 26420 , A.E. Nye (CNC); Cowley Canyon, [41.772669, -111.60407], 13–27.v.1989, ♂, JSS 24612, W.J. Hanson (EMUS); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.64184&amp;materialsCitation.latitude=41.893135" title="Search Plazi for locations around (long -111.64184/lat 41.893135)">Tony Grove Lake</a>, [41.893136, - 111.641841], 1.vii.2010, CNCD 60037–60040 , ♂, JSS 24615, M.M. Locke (CNC); West Hodges Canyon, [41.910305, -111.42277], 13–20.vi.1980, JSS 25096, (EMUS); Left Hand Fork, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.42277&amp;materialsCitation.latitude=41.910305" title="Search Plazi for locations around (long -111.42277/lat 41.910305)">Blacksmith Fork Canyon</a>, 24.v.2004, ♀, JSS 24617, Brammer (EMUS). RICH CO .: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-111.49441&amp;materialsCitation.latitude=41.862736" title="Search Plazi for locations around (long -111.49441/lat 41.862736)">Logan Canyon Summit</a>, [41.862735, -111.494404], 26.vi.–2.vii.1982, JSS 25098, (EMUS) . Wyoming: YELLOWSTONE NATIONAL PARK: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-110.537674&amp;materialsCitation.latitude=44.306126" title="Search Plazi for locations around (long -110.537674/lat 44.306126)">Lewis Lake</a>, [44.306126, - 110.537673], 17.vii.1923, ♀, JSS 19500, A.L. Melander (CNC); 17.vii.1923, ♀, JSS 23774, A.L. Melander (USNM) .</p></div>	https://treatment.plazi.org/id/03893F32FF8AFF9CFF46FE50FE928DF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FF8EFF9EFF46FCDEFDEF8AF8.text	03893F32FF8EFF9EFF46FCDEFDEF8AF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus pinastri (De Geer 1776)	<div><p>Dasysyrphus pinastri (DeGeer)</p><p>Figures: 15A–D, 18C, 21D</p><p>Synonyms and References:</p><p>Musca pinastri De Geer 1776: 113, Pl. 7 Fig. 1–7 (original description)</p><p>Syrphus lunulatus of authors nec. Meigen</p><p>Dasysyrphus pinastri (De Geer):</p><p>Vockeroth 1992: 70 (key and redescription)</p><p>Doczkal 1996: 40 (discussion and character chart)</p><p>Mutin &amp; Barkalov 1999: 398 (key)</p><p>Stubbs &amp; Falk 2002: 88 (key and discussion)</p><p>Reemer 2002: 14 (key and discussion)</p><p>van Veen 2004: 88 (key)</p><p>Barkalov 2007: 288 (key and redescription)</p><p>Bartsch et al. 2009: 199 (key and redescription)</p><p>Krpac et al. 2009: 105 (discussion)</p><p>Speight 2011: 71 (discussion)</p><p>Diagnosis: Abdominal maculae of tergites 3&amp;4 slightly arcuate, uniform in thickness and do not cross abdominal margin (Figs 15A &amp; B); very similar in appearance to D. pauxillus (Fig. 14A), D. pacificus (Fig. 13A), D. nigricornis (Figs 11A &amp; B) and D. laticaudus (Fig. 8A), sternite 2 with dark macula (Fig. 18C), dark marking is fasciate in the other species. Male surstylus with posterolateral corner is much less prominent (Fig. 15D) than in other pauxillus group species (Figs 8C, 11D, 13D &amp; 14D).</p><p>Redescription:</p><p>Body Length: 8.4–10.4mm; Wing Length: 7.8–9.0mm</p><p>Head: Frons dark with light pollenose fascia (in males it runs along ventral edge of where eyes meet, in females it is ⅓ the length between the antennae and ocelli) and dark pile; face light with dark vitta ¼–⅓ the width of the face, not reaching antennal socket, face with light pile; gena dark (sometimes slightly lighter posteroventrally) with light pile and usually some dark pile below eye; occiput dark, covered in light pollen, pile light; scape, pedicel and flagellomere dark, usually uniform in colour, flagellomere sometimes appears slightly lighter.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile light; scutellum light with darker anterolateral edges, dark pile with few light pile along anterior edge; wing cells r 1 and br usually with bare base (in br above spurious vein only), cell c sometimes with very small bare areas, cup with very small bare areas below vein CuP, bm densely microtrichose throughout (Fig. 15C); haltere light but can be slightly darkened in areas; femora ⅓–¾ dark basally, ¼–⅔ light apically, tibiae light, metatibia with dark band ¼ of the way from the apex, sometimes extending to apex, tarsi light, sometimes dark anteriorly.</p><p>Abdomen: Maculae on tergite 2 large and oval, maculae on tergites 3&amp;4 are oblique, anterior edge slightly concave and posterior edge straight, never meeting medially, maculae do not reach abdominal margin (Figs 15A &amp; B); sternite 2 with dark macula, sternites 3&amp;4 light with dark fasciae.</p><p>Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened posteriorly and posterolateral corner is much less prominent than in other pauxillus group species, long pile on dorsal, laterodorsal and posterodorsal sides, row of spines on posteromedial edge; cercus oval with long pile over entire surface; gonostylus boot-shaped with projection on posterodorsal surface, projecting dorsally, pile on dorsal surface; basiphallus elongate, basally bent at a slightly more than 90 degree angle towards dorsum, apical end curved ventrally and partly membranous dorsally, with spines on posterodorsal side; distiphallus elongate, apical end flared out into horn, which is mostly sclerotized but membranous at apex, with few setulae dorsally, basally it is curved towards dorsum and not greatly enlarged at bend (Fig. 15D).</p><p>Intraspecific variation: Gena is dark but in some specimens can be slightly lighter posteroventrally; wing cells r 1 and br usually with bare base, cell c sometimes with very small bare areas.</p><p>Etymology: From the Latin pinus, meaning pine and –aster, a diminutive suffix meaning little.</p><p>Distribution: Greenland and Europe (Fig. 21D).</p><p>Ecology: Collected from May to July. One specimen was collected from an elevation of 2200m.</p><p>Discussion: Within the Nearctic, D. pinastri is only found in Greenland. It is found throughout the Palaearctic from the United Kingdom to Scandinavia, south to Italy. Due to its similar habitus to other species in the Palaearctic there has been widespread confusion about this species and its concept (Doczkal 1996; Reemer 2002; Speight 2011). The main reason for such confusion is that the syntypes of this species are lost (Thompson &amp; Pont 1994). They were not in the De Geer collection when it was donated to the Naturhistoriska Riksmuseet (The Swedish Museum of Natural History) in Stockholm, Sweden, in 1834 (Thompson &amp; Nielsen 1984; Thompson &amp; Pont 1994). Thompson &amp; Pont (1994) designated the specimen illustrated by De Geer (1776, Pl. 7 Fig. 1) as the lectotype for pinastri . Since this lectotype has been missing for over 175 years and because this has lead to confusion about the species concept, a neotype is designated here to bring stability of the name pinastri . This designation will create a solid concept for this species, allowing for better reconciliation with closely related species. Characters are consistent with De Geer’s (1776) vague description of the adult and with Doczkal’s (1996) characters used to distinguish it from pauxillus s.l. and lenensis . De Geer (1776) does not state the locality/ localities of his specimens, however, as he was from Sweden and reared these flies from larvae it is likely that he collected them close to home. Therefore the Neotype chosen is from Sweden. The Neotype is a male from Holmsund Vb., Sweden, collected on 1.vii.1967 by W.J. Boyes. It is deposited in the CNC and databased with the number Jeff Skevington Specimen 18310. The genitalia have been dissected and a leg was removed to sequence the 5´region of COI.</p><p>Type Material: “Holmsund/ Vb., SWEDEN / 1.VII.1967 / J.W. Boyes ”, “Boyes Cytolog./ Coll. # WS101/ To remain/ in C.N.C.”, “ J. Skevington / Specimen #/ 18310”, “ Dasysyrphus / pinastri DeG./ Det. J. R. Vockeroth”, “ NEOTYPE / Musca / pinastri/ De Geer”, “designated by/ Locke &amp;/ Skevington/ 2013”; ♂; right meso- and metalegs missing; genitalia dissected, in vial on specimen pin; deposited in the Canadian National Collection of Insects, Arachnids and Nematodes (CNC), Ottawa, Ontario, Canada .</p><p>Material Examined:</p><p>Denmark: Fellins Skov, 22.v.1967, ♂, CNCD 26450 , E. Torp (CNC). Hytter Kobbel, Sdrj., 24.vi.1967, ♀, CNCD 26456 , E. Torp (CNC). Lillebro, Loso, 25.vi.1966, ♀, CNCD 26457 , E. Torp (CNC). North Denmark Region: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=10.519762&amp;materialsCitation.latitude=57.424683" title="Search Plazi for locations around (long 10.519762/lat 57.424683)">Bangsbo Skole</a>, [57.424683, 10.519762], 2.vi.1967, ♂, CNCD 26448 , E. Torp (CNC). South Denmark: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=8.983375&amp;materialsCitation.latitude=55.01665" title="Search Plazi for locations around (long 8.983375/lat 55.01665)">Draved Skov</a>, [55.016651, 8.983375], 10.v.1966, ♂, CNCD 26447, ♀, CNCD 26455 , E. Torp (CNC) . Greenland: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-50.709167&amp;materialsCitation.latitude=67.01056" title="Search Plazi for locations around (long -50.709167/lat 67.01056)">Sondrestom Air Base</a>, [67.010556, -50.709167], 30.viii.1952, ♀, CNCD 26459 , W.J. Brown (CNC) . Italy: Piedmont: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=7.007859&amp;materialsCitation.latitude=44.57995" title="Search Plazi for locations around (long 7.007859/lat 44.57995)">Celle de Bellino</a>, [44.579948, 7.007859], 11–12.vii.1974, intersex, JSS 18309, W. Baker (CNC) .</p><p>Poland: Szklarska Porpba, distr. Telenia Gora Taka, 9.vii.1958, ♂, CNCD 26449, R . Trojan (CNC) . Sweden: Västerbottens Län: Holmsund, [63.705557, 20.37888], 1.vii.1967, ♀, CNCD 26454,3 . vii.1967, ♂, JSS 18311, ♀, CNCD 26453, J.W. Boyes (CNC) . Switzerland: Valais: Zermatt, Rifolalb, 2200m, 23.vi.1959, ♀, CNCD 26458, F. Keiser (CNC) . Jura: Delemont, [47.365206, 7.343584], 17–24.v.1969, ♂, JSS 18308, Herting &amp; Wood (CNC) .</p><p>United Kingdom: England: Bristol, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-2.601311&amp;materialsCitation.latitude=51.502888" title="Search Plazi for locations around (long -2.601311/lat 51.502888)">Westbury</a>, [51.502888, -2.601311], 18.v.1959, ♂, CNCD 26446 , M. Ackland (CNC). Staffordshire, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-2.231836&amp;materialsCitation.latitude=53.008762" title="Search Plazi for locations around (long -2.231836/lat 53.008762)">Newcastle-under-Lyme</a>, [53.008764, -2.231836], 6.vi.1960, ♀, CNCD 26452 , J. R. Vockeroth (CNC). Cumbria, Lakes District National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-3.092892&amp;materialsCitation.latitude=54.42975" title="Search Plazi for locations around (long -3.092892/lat 54.42975)">Blea Tarn</a>, [54.429748, -3.092892], 5.vi.1941, 2♂, CNCD 26444, 26445 , G.E. Shewell (CNC).</p></div>	https://treatment.plazi.org/id/03893F32FF8EFF9EFF46FCDEFDEF8AF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FF8CFFE0FF46F9D0FC138D98.text	03893F32FF8CFFE0FF46F9D0FC138D98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus richardi Locke & Skevington 2013	<div><p>Dasysyrphus richardi Locke &amp; Skevington sp. nov.</p><p>Figures: 16A–C, 20C</p><p>Diagnosis: Similar in appearance to lotus (Figs 10A &amp; B), with maculae on tergites 3&amp;4 oblique, uniform in thickness, meeting medially and not crossing the abdominal margin (Figs 16A &amp; C). Dasysyrphus richardi has a dark vitta separating the gena from the face (Figs 16A &amp; C) and the wing extensively bare basally (Fig. 16B), where as lotus is continuously light from face to gena (Fig. 10B) and the wing is more microtrichose (Fig. 10C).</p><p>Description:</p><p>Body Length: 7.2–8.0mm; Wing Length: 7.2–7.5mm</p><p>Head: Frons dark with light pollenose fascia (in female it is ½ the length between the antennae and ocelli and is continuous, sometimes notched medially) and dark pile; face light with dark vitta medially, that is &lt;¼ the width of the face, not reaching antennal socket, face with dark pile; gena light with light pile and sometimes interspersed with dark pile, separated from face by dark vitta (Figs 16A &amp; C); occiput dark, covered in light pollen, pile light with some dark pile dorsally; scape and pedicel dark, flagellomere dark, lighter at base.</p><p>Thorax: Scutum dark, may appear shiny or metallic, with two faint pollenose vittae medially, extending just past the transverse suture, pile light; scutellum light with darker lateral edges, light pile with some dark pile posteriorly; wing cells br and bm entirely bare, cell c bare at base and along Sc, r 1 bare at base and cup bare at base and along vein CuP and posterior edge, r 2+3, r 4+5, dm, cua 1 and anal lobe with bare areas at base (Fig. 16B); haltere light; pro- and mesofemur ⅓–½ dark basally, ½–⅔ light apically, metafemur ⅔–¾ dark basally, ¼–⅓ light apically, tibiae light, metatibia slightly darkened medially, tarsi light.</p><p>Abdomen: Maculae on tergite 2 large, oval to triangular, slightly narrowed on medial end, ⅓–&lt;½ width of tergite, maculae on tergites 3&amp;4 are straight and oblique, meeting medially, maculae do not reach abdominal margin and are uniform in thickness (Figs 16A &amp; C); sternites light with dark fasciae of varying thickness.</p><p>Male Genitalia: Male unknown.</p><p>Intraspecific variation: Maculae on tergite 2 varying slightly in size and shape from ⅓–&lt;½ width of tergite and oval to slightly more triangular.</p><p>Etymology: This species is named after the systematist Dr. J. Richard Vockeroth, who recognized this as a unique species and labeled the specimens as so, but did not ever describe them.</p><p>Distribution: Southwestern USA (NM) (Fig. 20C)</p><p>Ecology: Two specimens have been collected in June in the southern Rocky Mountains.</p><p>Discussion: Known from two female specimens only. They were collected during the same collecting event in New Mexico in 1977. DNA was not extracted from these specimens as our techniques to extract from 30 year-old specimens need some improvement so that an attempt to extract DNA will have as great a chance as possible of success. With only two known specimens in existence this is crucial. Unfortunately with no male specimens, genitalia characters were not available for study. However, because richardi shares such a similar habitus to lotus it is likely that they are of close relation. An effort to find more specimens should be undertaken to locate the male of this species and to have more material made available in order to ascertain the full breadth of its molecular and morphological variation.</p><p>Type Material: “ New Mexico, Los Alamos/ Bandelier Nat. Mon./ June 1977 / W. Pippen #D-18”, “ USNM ENT 00258072 ”, “ HOLOTYPE / Dasysyrphus / richardi/ Locke &amp;/ Skevington”; ♀, deposited in the United States National Museum of Natural History, Washington, D.C (USNM) .</p><p>Material Examined:</p><p>Paratype: United States of America: New Mexico: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-106.25619&amp;materialsCitation.latitude=35.783115" title="Search Plazi for locations around (long -106.25619/lat 35.783115)">Los Alamos</a>, Bandelier Nat. Mon., [35.783115, - 106.25619], vi.1977, ♀, USNM ENT 258073 , W. Pippin (USNM) .</p></div>	https://treatment.plazi.org/id/03893F32FF8CFFE0FF46F9D0FC138D98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
03893F32FFF2FFE6FF46FD30FAB68873.text	03893F32FFF2FFE6FF46FD30FAB68873.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasysyrphus venustus (Meigen 1822) Widespread	<div><p>Dasysyrphus venustus (Meigen)</p><p>Figures: 17A–D, 22</p><p>Synonyms and References:</p><p>Scaeva arcuata Fallén: 1817: 42 (original description)</p><p>Dusek &amp; Laska 1969: 171 (name suppression proposal)</p><p>International Commission on Zoological Nomenclature 1972 1972: 9 (name suppression)</p><p>Syrphus venustus Meigen 1822: 299 (original description)</p><p>Syrphus lunulatus Meigen 1822: 299 (original description)</p><p>Syrphus reflectipennis Curran 1921: 157 (original description)</p><p>Metasyrphus reflectipennis (Curran):</p><p>Fluke 1933: 109 (key, redescription and transfer to Metasyrphus)</p><p>Metasyrphus venustus (Meigen):</p><p>Fluke 1933: 112 (key, redescription and transfer to Metasyrphus)</p><p>Dasysyrphus lunulatus (Meigen):</p><p>Enderlein 1938: 208 (transfer to Dasysyrphus)</p><p>Stone et al. 1965: 564 (catalogue and transfer back to Dasysyrphus)</p><p>Goeldlin de Tiefnau 1974: 208 (lectotype designation)</p><p>Syrphella lunulatus (Meigen):</p><p>Goffe 1944: 129 (transfer to Syrphella)</p><p>Syrphella venustus (Meigen):</p><p>Goffe 1944: 129 (transfer to Syrphella)</p><p>Dasysyrphus arcuatus (Fallén):</p><p>Stone et al. 1965: 563 (catalogue and transfer to Dasysyrphus)</p><p>Peck 1974: 128 (key)</p><p>Dasysyrphus reflectipennis (Curran):</p><p>Stone et al. 1965: 564 (catalogue and transfer to Dasysyrphus)</p><p>Dasysyrphus venustus (Meigen):</p><p>Dusek &amp; Laska 1967: 354 (transfer to Dasysyrphus ?)</p><p>Goeldlin de Tiefnau 1974: 208 (lectotype designation)</p><p>Vockeroth 1986: 203 (discussion)</p><p>Peck 1988: 16 (catalogue)</p><p>Vockeroth 1992: 70 (key and redescription)</p><p>Mutin &amp; Barkalov 1999: 397 (key)</p><p>Stubbs &amp; Falk 2002: 88 (key and discussion)</p><p>van Veen 2004: 87 (key)</p><p>Barkalov 2007: 294 (key and redescription)</p><p>Bartsch et al. 2009: 202 (key and redescription)</p><p>Speight 2011: 72 (discussion)</p><p>Diagnosis: Abdominal maculae cross abdominal margin; maculae on tergite 2 large (&gt;⅓ width of tergite) ovalshaped; maculae on tergites 3&amp;4 are transverse, constricted slightly medially, with medial edge swollen slightly (Figs 17A &amp; B). Similar to D. limatus however maculae on tergite 2 are always small in D. limatus (Fig. 9A), less than ¼ the width of the tergite, and are much larger in D. venustus, greater than ⅓ the width of the tergite. Also often confused with species in D. intrudens complex, however, species in this complex have maculae that are arcuate, lunulate and/or greatly constricted medially.</p><p>Redescription:</p><p>Body Length: 7.9–11.2mm; Wing Length: 7.4–10.4mm</p><p>Head: Frons dark with light pollinose fascia (in males it runs along ventral edge of where eyes meet, in females it is ⅓–½ the length between the antennae and ocelli and widely separated medially) and dark pile; face light with dark vitta ¼–⅓ the width of the face, usually reaching antennal socket, face with light to dark pile or combination of light and dark pile; gena dark with dark to light pile or a combination of light and dark pile; occiput dark, covered in light pollen, pile light; scape and pedicel light, flagellomere usually light basally and darker apically, sometimes mostly dark.</p><p>Thorax: Scutum dark, may appear shiny or metallic, pile light; scutellum light with darker anterolateral edges, light pile some with few dark pile along posterior edge; wings can range from being entirely densely microtrichose to having wing cells r 1, br and bm bare base (in br above spurious vein only) and cell cup with small bare areas below vein CuP (Fig. 17C); haltere light; pro- and mesofemora ¼–½ dark basially, ½–¾ light apically, metafemur ⅔ dark basially, ⅓ light apically, tibiae light, metatibia with dark band ¼ of the way from the apex, tarsi light, metatarsi dark anteriorly.</p><p>Abdomen: Maculae on tergite 2 large and oval with anterolateral edge extending to edge of abdomen, maculae on tergites 3&amp;4 are transverse, constricted slightly medially, with medial edge swollen slightly, almost never meeting medially, maculae cross abdominal margin (Figs 17A &amp; B); sternite 2 either all light or with dark fascia (usually distinct, sometimes faint), sternites 3&amp;4 light with dark fasciae.</p><p>Male Genitalia: Surstylus more or less triangular in shape in lateral view, flattened anteroposteriorly with long pile on posterior side and spines on anteroventral side, similar to D. intrudens complex (Fig. 7F) and D. limatus (Fig. 9D); cercus oval with long pile over entire surface; gonostylus is pointed ventrally, dorsally flattened at base then it expands dorsally at apex, oblique ridge near apex, with ends of ridge projecting into points; pile on dorsal surface; basiphallus is basally bent at about a 90 degree angle towards dorsum, apical end enlarged and curved ventrally, fully sclerotized all the way around with no pile or spines; distiphallus long with no enlarged area basally, apical end flared out into horn with the dorsal side flattened slightly, which is mostly sclerotized but membranous at apex with setulae dorsally (Fig. 17D).</p><p>Intraspecific variation: The abdominal maculae of the male are sometimes more arcuate and with a more swollen medial edge of maculae. Wings cells r 1, br, bm, and cup range from being entirely microtrichose to having bare areas.</p><p>Etymology: From the Latin venustus, meaning charming, lovely, beautiful, graceful.</p><p>Distribution: Canada (AB, BC, MB, NT, NS, ON, QC, YT) and US (AK, ID, ME, MI, NH, NM, NC, PA, UT, VT, VA, WV), also widespread over Europe and Asia (Fig. 22).</p><p>Ecology: Collected from May–August, but more frequently in May and June. Collected at elevations from 260–2957m. Habitats listed on labels include “bog”, “fell-field tundra”, “Mh/Be forest” (mixed hardwood/beech), “pine forest”, “boreal mixed wood forest”, “black spruce forest (25%) with moss”, “Picea-Betula forest”. Specimens have been collected from hilltops.</p><p>Discussion: The species concept for D. venustus has been uncertain for quite some time now, especially in the Nearctic. Fluke’s (1933) concept was much like that of this one, distinguishing venustus as having markings that are more straight than specimens in the intrudens complex, which is treated by Fluke as laticaudatus, amalopis (in part) and osburni . Vockeroth (1986) synonymized the intrudens complex under venustus as he believed it was simply a variable species. While abdominal maculae vary greatly, many of the other characters are similar, including the male genitalia.</p><p>Dasysyrphus osburni, intrudens, disgregus and laticaudatus were all considered synonyms of venustus by Vockeroth (1992), however these four names fall within what we recognize as the intrudens complex.</p><p>The only specimen of D. reflectipennis is that of the holotype (located at the CNC). It is a female with its abdomen broken and glued to a point below the specimen. It was treated as a unique species by Fluke (1933) and by Vockeroth (1986). However, at the time, Vockeroth believed that reflectipennis was a specimen of venustus with aberrant markings. He stated that he was not ready to treat it as a synonym until other specimens were found. Other specimens were not found, but in 1992 he synonymized reflectipennis with venustus . The abdominal maculae of tergite 3 are like that of venustus, however they narrowly meet medially. We have seen no other specimens where the maculae meet, but since the specimen of reflectipennis looks like that of venustus in every other way, it is logical to conclude that Vockeroth was correct in saying that the specimen of reflectipennis is actually a specimen of venustus with aberrant markings.</p><p>There are five specimens that appear to be morphologically identical and sympatric with venustus but differ from it in their COI sequences (CNCD 5458, 30889, 30890, 30902, JSS 20982; Fig. 4). It is possible that this is a cryptic species but until more specimens are available to provide more precise geographical and ecological evidence and/or morphological and DNA characters they are retained under the name venustus .</p><p>Type Material: From Germany, deposited in the Muséum National d'Histoire Naturelle (The National Museum of Natural History), Paris, France (Pape &amp; Thompson 2010b). (not examined; requested but not obtained) Material Examined: Canada: Alberta: Itaska Audubon Reserve, Pigeon Lake, [53, -114], 18.v.2008, ♂, JSS 21073, ♀, JSS 21074, F.A.H. Sperling (FAHS). Drayton, [53.220865, -114.978236], 29.vi.1988, ♀, UASM# 149016, E.H. Strickland (UASM). Edmonton, Division No.11, [53.540941, -113.493698], 5.vi.1948, ♀, CNCD 14537, J.R. Vockeroth (CNC); 20.v.1923, ♀, CNCD 14538, E.H. Strickland (CNC); 12.v.1924, ♀, JSS 19540–19545, 19551–19554, 19561–19564, 13.v.1924, ♀, JSS 19559, 19560, 19565, 16.v.1924, ♀, JSS 19558, O. Bryant (CNC). Elk Island National Park, [53.604578, -112.859291], 19.v.1981, ♀, CNCD 14540, J.E. O’Hara (CNC). Bilby, Division No. 13, [53.698357, -114.10198], 1.vi.1924, ♀, JSS 19546, O. Bryant (CNC). British Columbia: CENTRAL KOOTENAY L.D.: Robson, [49.333015, -117.692871], 14.v.1954, ♂, JSS 20034, 17.v.1954, ♂, JSS 20025, 19.v.1954, JSS 20127, 26.v.1954, ♂, JSS 19999, 20.v.1955, ♂, JSS 19998, 7.v.1956, ♀, JSS 20015, 8.v.1956, ♂, JSS 20019, 14.v.1956, 3♂, JSS 20023, 20164, 15.v.1956, ♂, JSS 20132, ♀, JSS 20130, 6.v.1957, ♂, JSS 20012, 11.v.1957, ♀, JSS 20002, 21.v.1957, ♀, JSS 20095, 7.vi.1965, ♀, JSS 20122, 25.v.1965, ♀, JSS 20092, H.R. Foxlee (UBCZ); 14.v.1947, ♂, CNCD 14510, 11.v.1948, ♀, CNCD 14242, 2.vi.1948, ♀, CNCD 14303, H.R. Foxlee (CNC). Miracle Beach, Near Oyster Bay, [49.854574, -125.098409], 13.vi.1955, ♀, CNCD 14314, J.R. McGillis (CNC). Salmon Arm, [50.703923, -119.273924], 2.v.1930, ♀, JSS 20159, Hugh B. Leech (UBCZ). 32 miles South West of Terrace, [54.194076, -129.140296], 30.5m, 8.vi.1960, ♂, CNCD 14509, G.E. Shewell (CNC); 11.vi.1960, 2♀, CNCD 14217, 14341, B. Heming (CNC). Spring Creek, Terrace, [54.539511, -128.618192], 1.vi.1960, ♀, CNCD 14231, C.H. Mann (CNC). Kitsumkalum Lake, 20 Miles North of Terrace, [54.762473, -128.770968], 16.vi.1960, ♀, CNCD 14227, J.G. Chillcott (CNC). Kispiox, [55.348319, -127.687346], 24.v.1958, ♂, JSS 20152, J. Foster (UBCZ). Atlin, [59.5775, -133.69236], 670.56m, 9.vi.1955, 2♀, CNCD 14251, 14252, 20.vi.1955, ♀, CNCD 14223, 6.vii.1955, ♀, CNCD 14284, B.A. Gibbard (CNC); 20.vi.1955, ♂, CNCD 14360, 7.vii.1955, ♀, CNCD 14289, H.J. Huckel (CNC). Blanchard R. Haines Rd., [11.vi.1979, ♀, JSS 20146, G.J. Spencer (UBCZ). Kiskatinaw Public Campground, Alaska Highway DC-20 (31.4km), [26.vi.1978, ♀, JSS 23719, Paul H. Arnaud, Jr. (CAS). Manitoba: Aweme, [49.708529, -99.602758], 19.v.1926, ♀, CNCD 14550, 27.v.1925, ♀, CNCD 14554, 1.vii.1966, ♀, CNCD 14555, R.D. Bird (CNC); 20.vi.1925, ♀, CNCD 14551, R.M. White (CNC). 5 miles South West of Shilo, [49.763964, -99.717403], 28.v.1958, ♀, CNCD 14552, J.F. McAlpine (CNC); 5.vi.1958, ♀, CNCD 14553, C.D.F. Miller (CNC). Erickson, [50.498283, -99.9078], 17.vi.1983, ♀, JSS 20195, D.H. Pengelly (JBWM). CHURCHILL: Burn site N[orth] of Twin lakes, [58.61861111, -93.82888889], 4.vii.2007, ♀, CNCD 9495, 5.vii.2007, ♀, CNCD 9582, J. Skevington (CNC); Bus Stop, [7.viii.2007, ♀, JSS 24644, P. Kevan (CNC). New Brunswick: CHARLOTTE CO.: Chamcook, [45.126407, -67.071693], 28.vi.1965, 3♀, CNCD 14817-14819, G.E. Shewell (CNC). NORTHUMBERLAND CO.: Doaktown, [46.556585, -66.126203], 6.vii.1971, ♀, CNCD 14820, B.V. Peterson (CNC). KENT CO.: Kouchibouguac National Park, [46.819201, -64.96788], 19.vi.1978, 2♀, CNCD 14815, 14816, S.J. Miller (CNC). Snow Cap Park, Hwy 17, [5.vii.1971, ♀, CNCD 14821, B.V. Peterson (CNC). Northwest Territories: Jean-Marie Creek, 42 Miles South West of Fort Simpson, [61.372555, -122.224182], 13.vi.1972, ♀, CNCD 14047, B.V. Peterson (CNC). Coppermine, [62.562732, -115.092764], 30.vi.1951, ♀, CNCD 14058, S.D. Hicks (CNC). Wrigley, Mackenzie, [63.209367, -123.34594], 15.vi.1969, ♀, CNCD 14056, G.E. Shewell (CNC). Nova Scotia: KINGS CO.: [45.017428, -64.610757], 22.vi.1983, ♀, CNCD 14779, 23.vi.1983, ♀, CNCD 14778, J.R. Vockeroth (CNC). COLCHESTER CO., Truro, [45.364959, -63.279846], 7.vii.1919, ♂, CNCD 14769, (CNC). VICTORIA CO., Baddeck, Cape Breton Island, [46.10001, -60.753507], 15.vi.1931, ♀, CNCD 14780, G. Fairchild (CNC); Dingwall, [46.896638, -60.469402], 23.vi.1983, ♀, CNCD 14777, J.R. Vockeroth (CNC); INVERNESS CO., Cape Breton Highlands National Park: French Lake, [46.728452, -60.86483], 24– 30.vi.1984, 2♀, CNCD 14774, 14775, H.J. Teskey (CNC); North Mtn. Bog, [46.805111, -60.688224], 400m, 7.vi.1984, 2♂, CNCD 14759, 14760, B.E. Cooper (CNC); Mackenzie Mountain, [46.773977, -60.818772], 400m, 9.vi.1984, ♂, CNCD 14762, B.E. Cooper (CNC); Pleasant Bay, [46.822961, -60.799065], 6.vi.1984, ♂, CNCD 14765, 11.vi.1984, ♂, CNCD 14763, 12.vi.1984, 3♂, CNCD 14757, 14758, 14766, 2♀, CNCD 14770, 14771, B.E. Cooper (CNC); 25–29.vi.1984, ♀, CNCD 14776, H.J. Teskey (CNC). Ontario: Crawford, [44.253882, - 80.929867], 7–16.vi.1998, 4♀, ROMEnt Spec. No. 57872, 57887–57889, 8–17.vii.1998, ROMEnt Spec. No. 74031, A. Sugar (ROME). Cranberry Lake, Runtz property, [44.44111111, -76.29055556], 18–24.v.2007, 2♀, CNCD 30889, 30890, J. Skevington, H. Douglas, D. Runtz (CNC). LEEDS CO.: Chaffeys Lock, [44.579098, - 76.319816], 28.v.1972, ♀, CNCD 14593, P. Ward (CNC); SIMCOE CO.: Orillia, [44.609505, -79.42068], 26.v.1913, ♀, CNCD 15386, 30.v.1921, ♀, CNCD 14595, 7.vi.1925, ♀, CNCD 14594, C.H. Curran (CNC); Midland, [44.752113, -79.887253], 26.v.1959, ♀, CNCD 14600, J.G. Chillcott (CNC). Griffith, 7 miles East, [45.243279, - 77.031193], 22.v.1982, ♂, CNCD 14567, B.E. Cooper (CNC). Stittsville, [45.263479, -75.925163], 24.vi.1963, ♀, CNCD 14598, W.R.M. Mason (CNC). Nepean, 24 Gervin Street, [45.31694444, -75.72], 28.v.2006 4.vi.2006, ♀, CNCD 30888, J.E. O’Hara (CNC). Mer Bleu, 5 Miles East of Ottawa, [45.40331, -75.5083], 31.v.1923, ♀, CNCD 14579, C.H. Curran (CNC); 5.vi.1966, 2♀, CNCD 14585, 14586, 13.vi.1966, ♀, CNCD 14587, 15.vi.1966, ♀, CNCD 14588, 17.vi.1966, ♀, CNCD 14589, 19.vi.1966, 2♀, CNCD 14590, 1.vii.1966, ♀, CNCD 14592, D.D. Munroe (CNC). Ottawa, [45.411604, -75.688669], 29.v.1925, ♀, CNCD 14571, C.H. Curran (CNC). Waubamick, Parry Sound District, [45.446658, -80.017278], 14.vi.1915, 2♀, JSS 19533, 19534, E.S. Parish (CNC). Dirleton, [45.494944, -76.142874], 29.iv.1968, ♀, CNCD 14599, J.E.H. Martin (CNC). ALGONQUIN PROVINCIAL PARK: Florence Lake [45.44371, -78.49012], 13.v.2007 29.v.2007, ♂, JSS 21385, E. Proctor (CNC); Cecil Lake, [45.4511, -78.4956], 17.v.2007 29.v.2007, ♂, JSS 21371, E. Proctor (CNC); 2 km E of Pondweed Lake, [45.4654, - 78.4298], 16–28.v.2007, 2♂, JSS 21394, 21397, E. Proctor (CNC); Algonquin Park, [46, -78.08333333], 12– 19.vi.2003, ♀, ROMEnt Spec. No. 98532, 13.–20.vi.2003, 7♀, ROMEnt Spec. No. 98522–98525, 98535, 98536, 98540, 17–24.vi.2003, ♀, ROMEnt Spec. No. 98541, 18–25.vi.2003, 3♀, ROMEnt Spec. No. 98526, 98527, 98528, 98533, 20–27.vi.2003, 2♀, ROMEnt Spec. No. 98530, 98531, 10.vii.2003 17.vii.2003, ♀, ROMEnt Spec. No. 98534, Mark Vanderwel (ROME). Atikokan, 14 miles East on Hwy 11, [48.723518, -91.313476], 3.vii.1978 4.vii.1948, ♀, CNCD 14582, H.J. Teskey (CNC). Iroquois Falls, [48.761811, -80.685262], 20.vi.1987, ♀, JSS 20773, J.R. Vockeroth (CNC). LowBush, Lake Abitibi, [48.91608, -80.139852], 6.vi.1925, ♂, BOC 34653, 7.vi.1925, 2♀, ROMEnt Spec. No. 115126, 115127, 13.vi.1925, 2♀, ROMEnt Spec. No. 115128, 115130, 16.vi.1925, ♀, ROMEnt Spec. No. 115131, 17.vi.1925, ♀, ROMEnt Spec. No. 115132, 23.vi.1925, ♀, ROMEnt Spec. No. 115133, 1.vii.1925, 4♀, BOC 34649–34652, 14.vii.1925, ♀, BOC 34654, 18.vii.1925, 4♀, BOC 34655– 34657, ROMEnt Spec. No. 115129, N.K. Bigelow (ROME); 9.vi.1925, ♀, CNCD 14572, 15.vi.1925, ♀, CNCD 14573, 16.vi.1925, 2♀, CNCD 14574–14576, 14.vii.1925, ♀, CNCD 14578, N.K. Bigelow (CNC); 1.vii.1925, ♀, UASM# 149015, N.K. Bigelow (UASM); 1.vii.1925, ♀, JSS 21349, 14.vii.1925, ♀, JSS 21350, N.K. Bigelow (MCZ). Macdiarnid, Lake Nipigon, [49.438216, -88.12695], 7.vi.1921, ♀, BOC 34659, 24.vi.1923, ♀, BOC 34660, N.K. Bigelow (ROME). Quebec: Covey Hill, Le Haut-Saint-Laurent, [45.019681, -73.745746], 20.vi.1924, ♀, CNCD 14755, G.S. Walley (CNC). Aylmer, [45.400224, -75.817137], 28.v.1923, ♂, CNCD 14653, C.H. Curran (CNC). Ste. Anne de Bellevue, [45.406417, -73.950944], 12.v.1965, ♀, LEM 8806, 1.vi.1966, ♀, LEM 8804, W. Boyle (LEMQ); 13.v.1965, ♀, LEM 8807, G. Jamieson (LEMQ); 15.v.1967, ♂, LEM 8805, A. Campbell (LEMQ). Abbotsford, [45.437637, -72.887923], vii.1935, ♀, CNCD 14711, G.E. Shewell (CNC); 25.vi.1937, ♀, CNCD 14725, G. Shewell (CNC); 14.vi.1972, ♀, CNCD 14689, D.M. Wood (CNC). VAUDREUIL CO.: Mount Rigaud, [45.46638889, -74.32638889], 19.v.2004, ♂, CNCD 68, J. Skevington, L. Bartels, E. St. Louis, J. King (CNC); 24.v.2007, CNCD 8656, 8658, 8675, J. Skevington, A. Bailie, R. Farmahin, A. Kealey, S. Kelso, J. King (CNC). LES COLLINES- DES- L'OUTAOUAIS: Summit of King Mountain, Old Chelsea, [45.498058, -75.864116], 351m, 14.vi.1963, ♂, CNCD 14680, 2♀, CNCD 14737, 14738, 21.vi.1966, ♂, CNCD 14651, ♀, CNCD 14686, 13.vi.1970, 6♂, CNCD 14606, 14608–14612, 11.vi.1971, 2♂, CNCD 14654, 14679, ♀, CNCD 14736, 16.vi.1971, ♂, CNCD 14671, 12.vi.1980, ♀, CNCD 14693, J.R. Vockeroth (CNC); 9.vi.1971, ♂, CNCD 14681, D.M. Wood (CNC); Champlain Lookout, Gatineau Park, [45.508314, -75.913273], 335m, 1.vi.1965, ♂, CNCD 14672, J.R. Vockeroth (CNC). Montreal Area, Communaute-Urbaine-de-Montreal, [45.526155, - 73.702807], 6.vi.1964, 2♀, CNCD 14692, 14694, J.W. Boyes (CNC). Mont Saint-Hilaire, La Vallee-du-Richelieu, [45.552948, -73.155276], 411m, 11.vi.1967, ♂, CNCD 14675, J.W. Boyes (CNC). Lac-Megantic, Le Granit, [45.57781, -70.884105], 18.vi.1923, ♂, CNCD 14646, 20.vi.1923, ♀, CNCD 14753, C.H. Curran (CNC); 30.v.1971, ♂, CNCD 14667, D.M. Wood (CNC). Duncan Lake, near Rupert, [45.68138889, -76.05027778], 21.vi.1969, ♀, CNCD 14745, J.F. McAlpine (CNC); 19.v.1987, ♂, JSS 20767, D.M. Wood (CNC). Lac Cloutier, [46.184459, -73.64769], 30.v.1987 7.vii.1987, ♀, LEM 13620, (LEMQ). Schefferville, Iron Arm Fern, [54.805276, -66.805274], 28.vi.1990, ♀, LEM 13928, 3–4.vii.1990, ♀, LEM 13459, P. McElligott (LEMQ). Great Whale River, [55.116816, -76.405554], 7.vii.1949, ♀, CNCD 14718, 12.vii.1949, 3♀, CNCD 14715–14717, 24.vii.1949, ♀, CNCD 14724, 25.vii.1949, ♀, CNCD 14714, 31.vii.1949, ♀, CNCD 14719, J.R. Vockeroth (CNC). Indian House Lake, [56.328482, -64.720845], 8.vii.1954, ♀, CNCD 14721, 10.vii.1954, ♂, CNCD 14683, 17.vii.1954, ♀, CNCD 14722, W.R. Richards (CNC). Kuujjuaq (Fort Chimo), [58.100076, -68.406179], 29.v.1948, ♂, CNCD 14684, H.N. Smith (CNC); 14.vii.1948, ♀, CNCD 14712, 28.vii.1948, ♀, CNCD 14713, R.H. MacLeod (CNC). Saskatchewan: Great Deer, [52.580321, -107.056629], 21.v.1949, ♀, CNCD 14547, J.R. Vockeroth (CNC). Nipawin, Division No.14, [53.365608, -104.011519], 2.vi.1948, ♀, CNCD 14548, J.R. Vockeroth (CNC). Yukon Territory: Otter Lake, [62.486817, -130.4166667], 1219m, 17.vii.1960, ♀, CNCD 13966, J.E.H. Martin (CNC). Sheldon Lake, [62.616763, -131.266603], 1067m, 3.vii.1960, ♀, CNCD 13967, 7.vii.1960, ♀, CNCD 13968, J.E.H. Martin (CNC); 3.vii.1960, ♀, CNCD 13976, 7.vii.1960, 2♀, CNCD 13974, 13975, E.W. Rockburne (CNC). La Force Lake, [62.683333, -132.333333], 1006m, 28.vi.1960, ♀, CNCD 13955, E.W. Rockburne (CNC); 7.vii.1960, ♀, CNCD 13949, J.E.H. Martin (CNC). DEMPSTER HIGHWAY: Mile 40, [64.448494, -138.223783], 1– 6.vii.1973, ♂, CNCD 14029, G. &amp; D.M. Wood (CNC); Engineer Cr., km 152 [65.070828, -138.2399], 30.vi.1982, ♀, JSS 20140, S.G. Cannings, L. Vasington, R.A. Moore (UBCZ); km 416 [66.644581, -136.324471], 750m, 22– 28.vi.1980, ♀, CNCD 14005, Wood &amp; Lafontaine (CNC). Rampart House, [67.421455, -140.983336], 18.vii.1951, ♀, CNCD 13981, C.C. Loan (CNC). Old Crow, [67.571219, -139.833877], 2.vii.1983, ♀, JSS 20138, R.A. Cannings (UBCZ). British Mountains, Firth River, [68.848549, -140.48328], 24.vii.1956, ♀, CNCD 14053, R.E. Leech (CNC); 24.vi.1984, ♀, JSS 18367, S.G. Cannings (CNC). Kluane; Ruby Ranges; Pika Camp, [16.vi.2004 23.vi.2004, ♀, JSS 25586, E.S. Bakker (WVSC). United States of America: Alaska: ANCHORAGE CO.: Anchorage, [61.175333, -149.861715], 13.vi.1921, ♀, JSS 19547, J.M. Aldrich (CNC). MATANUSKA- SUSITNA CO.: Matanuska, [61.541945, -149.229674], 4.vi.1944, ♀, JSS 19557, 10.vi.1944, ♀, JSS 19537, 11.vi.1944, ♀, JSS 19536, 17.vi.1944, ♀, JSS 19538, 1.vi.1945, 2♀, JSS 19535, 19539, J.C. Chamberlin (CNC). Trapper Creek, 36 miles North of Willow, [62.31666667, -150.2166667], 19.vii.1984, CNCD 60064, S. &amp; J. Peck (CNC). Unalakleet, [63.86666667, -160.7833333], 17.vi.1961, ♀, CNCD 26575, B.S. Heming (CNC); 16.viii.1961, 2♀, CNCD 26577, 26578, R. Madge (CNC). SOUTHEAST FAIRBANKS CO.: Big Delta, Lake Boleo, [64.154432, - 145.836524], 19.v.1951, ♀, CNCD 26553, W.R.M. Mason (CNC). NOME AREA: Mile 55 of Kougarok Road, [65.08972222, -164.6722222], 20–26.vi.2005, ♀, CNCD 30893, J.&amp;R. Skevington (CNC); Nome Area, Mile 17 of Kougarok Road, [64.70611111, -165.2972222], 21–23.vi.2005, 2♀, CNCD 30881, 30902, 26–28.vi.2005, 2♀, CNCD 30895, 30897, 30898, J.&amp;R. Skevington (CNC). Colorado: Chambers Lake, [40.603663, -105.853645], 3.viii.1932, ♀, JSS 20714, Chas L. Fluke (CSUC). Idaho: Keene Valley, [44.18333333, -73.78333333], 25.v.1910, JSS 21351, H. Notman (MCZ). Maine: Bar Harbor, [44.38333333, -68.2], ♀, JSS 21348, (MCZ). FRANKLIN CO.: Dryden, [44.586729, -70.209783], 20.vii.1959, ♂, CNCD 26734, G.H. Heinrich (CNC). Massachusetts: HAMPSHIRE CO.: [42.335786, -72.644117],. vi.1922, ♂, CNCD 26739, P.A. Readio (CNC). MIDDLESEX CO.: Melrose Highlands, [42.470929, -71.061441], 16.vi.1911, ♂, CNCD 26740, J.D. Tothill (CNC). Michigan: Manistee Lake, [44.24749, -86.30471], 22.v.1955, ♂, JSS 21138, R.W. Hodges (MSUC). Sagola, [46.09028, - 88.0775], 6.vi.1983, ♀, JSS 21136, M. Arduser (MSUC). Isle Royale, [48, -88.83333333], 3.vii.1957, ♀, JSS 21142, 15.vii.1957, 2♀, JSS 21144, JSS 21147, 18.vii.1957, ♀, JSS 21143, 11.viii.1957, ♀, JSS 21141, R.W. Hodges (MSUC). KALKASKA CO.: 14.vi.1949, ♀, JSS 21139, R.R. Dreisbach (MSUC); 12.vi., ♀, JSS 21140, R.R. Dreisbach (MSUC). CHIPPEWA CO.: 3.vi.1957, 2♀, JSS 21145, 21146, R. &amp; K. Dreisbach (MSUC). CLARE CO.: 16.v.1951, ♀, JSS 21119, 23.v.1959, 2♀, JSS 21148, 21149, R. Dreisbach (MSUC). New Hampshire: Franconia, Rt.3 Fanconia North, [44.209935, -71.685178], 20–21.vi.1972, ♀, USNM ENT 258066, B.J. &amp; F.C. Thompson (USNM). Pittsburg, Rt.3 Connecticut Lakes, [45.104264, -71.259207], 22–24.vi.1972, 4♀, USNM ENT 258067–258070, B.J. &amp; F.C. Thompson (USNM). CHESHIRE CO.: Jaffrey, [42.814957, -72.022399], 1941, ♂, CNCD 26741, C.W. Johnson (CNC). Bretton Woods, [44.25, -71.43333333], 27.vi., ♂, CNCD 26742, C.W. Johnson (CNC). COOS CO.: Mount Washington, [44.270481, -71.30255], 6.vii., ♂, CNCD 26735, (CNC). Lancaster Mt. Prospect, [44.449317, -71.570888], 625m, 19.vi.1982, ♀, JSS 20780, J.R. Vockeroth (CNC). New Mexico: LINCOLN CO.: Sierra Blanca, [33.478065, -105.552907], 2957m, 10–26.vi.1979, ♀, JSS 18336, S. &amp; J. Peck (CNC). New York: ULSTER CO.: Oliverea, [42.065369, -74.460152], 18.vi.1918, ♂, CNCD 26745, (CNC). Maplecrest, Catskill Mts., [42.275919, -74.186809], 18.vi.1927, JSS 19255, F.M. Schott (FMNH). ESSEX CO.: Heart Lake, Mount Jo, [44.182434, -73.966816], 13.vi.1935, ♂, CNCD 26743, C.W. Johnson (CNC). North Carolina: Shining Rock Wild. Area, [35.355, -82.84388889], 23.v.1993, ♀, JSS 20807, Kondratieff &amp; Kirchner (CSUC). Pisgah National Forest, 10km SE Linville on Road, [36, -81.8], 760m, 25.v.1999, ♀, JSS 20988, M.I. Erwin (MHPC). 13km W Boone on Blue Ridge Pkwy, [36.23333333, -81.51666667], 1050m, 24.v.1999, ♀, JSS 20987, 25.v.1999, ♂, JSS 20983, M.I. Erwin (MHPC). Pennsylvania: Black Moshannon State Park, [40.896041, - 78.044398], 13.v.1990, ♂, USNM ENT 258081, FC Thompspn (USNM). CENTRE CO., Bear Meadows Natural Area, 11.v.1983, ♂, CNCD 5458, F.C. Thompson (USNM). Utah: CACHE CO., Tony Grove Crk, [41.892197, - 111.634869], 8–15.vii.1994, ♀, JSS 24642, W.J. Hanson (EMUS). Vermont: ESSEX CO.: 5 Miles West of Bloomfield, [44.753513, -71.727243], 28.vi.1972, 2♀, JSS 18338, 18339, H.J. Teskey (CNC). Virginia: SMYTH CO., Hurricane Creek, 22.v.1981, ♂, CNCD 5456, F.C. &amp; B.J. Thompson (USNM). West Virginia: Standard, off I- 77, [38.136111, -81.400556], 6.v.2005, ♀, JSS 20692, B. Kondratieff, R. Zuelig, R.F. Kirchner (CSUC).</p></div>	https://treatment.plazi.org/id/03893F32FFF2FFE6FF46FD30FAB68873	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Locke, Michelle M.;Skevington, Jeffrey H.	Locke, Michelle M., Skevington, Jeffrey H. (2013): Revision of Nearctic <i> Dasysyrphus </ i> Enderlein (Diptera: Syrphidae). Zootaxa 3660 (1): 1-80, DOI: 10.11646/zootaxa.3660.1.1, URL: http://dx.doi.org/10.11646/zootaxa.3660.1.1
