identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039687B4C10C476BB082204C5A26FC5B.text	039687B4C10C476BB082204C5A26FC5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon Verma 1967	<div><p>Aspidophorodon Verma, 1967</p><p>Aspidophorodon Verma, 1967: 507 . Type species: Aspidophorodon harvensis Verma, 1967; by monotypy. Aspidophorodon Verma: Miyazaki, 1971: 183; Eastop &amp; Hille Ris Lambers. 1976: 95; Blackman &amp; Eastop, 1994: 569; Remaudière &amp; Remaudière, 1997: 73; Zhang et al., 1999: 349; Blackman &amp; Eastop, 2006: 1098; Stekolshchikov &amp; Novgorodova, 2010: 44; Nieto Nafría et al., 2011: 145.</p><p>Diagnosis. Body elliptical, dorsum corrugated, reticulate, or with different shaped markings in apterae. Abdominal tergites in alatae with segmental marginal sclerites and spino-pleural sclerotic bands, sometimes with a large sclerotic patch extending on several segments. In apterae, both spinal and marginal processes on body present or absent, or marginal processes absent while spinal ones present at least on abdominal tergite VIII, or spinal processes absent while marginal ones present on thoracic notum and abdominal tergites. In alatae, dorsal processes on body absent, or single spinal process present on abdominal tergite VIII, and sometimes marginal processes present on tergites I–IV as well. Dorsal setae of body sparse, short, blunt, capitate, or pointed. Median frontal tubercle distinctly protuberant, especially in apterae, hemispherical or rectangular, sometimes with a depression at the middle. Antennal tubercles in apterae protuberant or developed in the form of cylindrical, finger-like, or long horn-shaped projections. Antennae in apterae 5- or 6-segmented, rarely 4-segmented, in alatae 5- or 6-segmented. Ant.I usually slightly or distinctly projected at inner apex. Secondary rhinaria absent in apterae, small round or oval in alatae. URS wedge-shaped. First tarsal segment with 2–3 setae in all legs. SIPH long cylindrical, broad at base, thin at middle, slight swollen at distal, obliquely truncated at tip, without flange, siphuncular pore small. Cauda tongue-shaped, slightly constricted near the middle, sometimes with a constriction at base, or elongate triangular, with 3–10 setae. Wings with normal venation.</p><p>Distribution. Canada, China, India, Japan, the Altai Republic, Sakhalin and the Kuril Islands.</p><p>Host plants. Polygonaceae (Polygonum), Rosaceae ( Cotoneaster, Potentilla, Sorbus, Spiraea), and Salicaceae (Salix) .</p><p>Comments. Genus Aspidophorodon includes two subgenera: Aspidophorodon Verma, 1967 mostly feeding on Salix (Salicaceae) and Eoessigia David, Rajasingh &amp; Narayanan, 1972 mainly infesting Rosaceae . Species of Aspidophorodon are distributed in Asia and North America. The life cycles of most species are unknown.</p></div>	https://treatment.plazi.org/id/039687B4C10C476BB082204C5A26FC5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C10F476BB08224835A8FFB5C.text	039687B4C10F476BB08224835A8FFB5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon	<div><p>Key to subgenera of Aspidophorodon (based on apterous viviparous females)</p><p>1. Spinal processes on body absent; antennal tubercles well developed, cylindrical, finger-like, or long horn-shaped; antennae 4- or 5-segmented; cauda with 3–5 setae; mainly on Salix (Salicaceae) ............................ Aspidophorodon Verma</p><p>- Spinal processes on body present; antennal tubercles protuberant or developed in the form of finger-like projections; antennae 5- or 6-segmented; cauda with 4–10 setae; mainly on Rosaceae ................ Eoessigia David, Rajasingh &amp; Narayanan</p></div>	https://treatment.plazi.org/id/039687B4C10F476BB08224835A8FFB5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C10F476BB082239A5C49FA10.text	039687B4C10F476BB082239A5C49FA10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon Verma 1967	<div><p>Subgenus Aspidophorodon Verma, 1967</p><p>Aspidophorodon Verma, 1967: 507 . Type species: Aspidophorodon harvensis Verma, 1967; by monotypy. Aspidophorodon (Aspidophorodon) Verma: Remaudière &amp; Remaudière, 1997: 73; Stekolshchikov &amp; Novgorodova, 2010: 44; Nieto Nafría et al., 2011: 145.</p><p>Comments. The nominate subgenus contains five species, including three new species from China described herein. Species of this subgenus are distributed in eastern Asia.</p></div>	https://treatment.plazi.org/id/039687B4C10F476BB082239A5C49FA10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C10E476AB08227C05A7AF8D8.text	039687B4C10E476AB08227C05A7AF8D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon (Aspidophorodon) cornuatus Qiao	<div><p>Aspidophorodon (Aspidophorodon) cornuatus Qiao sp. nov.</p><p>(Figs 1–19, Table 1)</p><p>Specimens examined. Holotype: apterous viviparous female, CHINA: Tibet (Yadong County, 27.52°N, 88.97°E, altitude 2800 m), 15.viii.2010, No. 25908-2-3-1, on Salix cupularis, coll. Y. Wang (NZMC). Paratypes: 5 apterous viviparous females (slides), and 1 apterous viviparous female (COI: KJ374724; Buchnera 16S rRNA: KT 221035), with the same collection data as holotype (NZMC); 2 apterous viviparous females, No. 25908-2-1, with the same collection data as holotype (BMNH).</p><p>Etymology. The new species is named for its long horn-shaped antennal tubercles. The Latin word “ cornuatus ” means “horn-shaped”.</p><p>Description. Apterous viviparous females: Color of living specimens: yellow. General measurements see Table 1.</p><p>Color of mounted specimens: body pale (Fig. 9).</p><p>Body. Elliptical.</p><p>Head. Dorsum of head covered with many small oval or semicircular sculptures (Figs 1, 10), venter smooth with slight wrinkles. Median frontal tubercle distinctly protuberant, hemispherical (Figs 1, 10), with 1 pair of long pointed setae on venter. Antennal tubercles long horn-shaped, strongly imbricated, each with a short capitate seta at apex (Figs 1, 10). Dorsal setae of head short and capitate. Head with 1 pair of cephalic setae, 1 pair of dorsal setae between antennae, and 2 pairs of dorsal setae between compound eyes arranged transversely (Fig. 1). Antennae 5- segmented, Ant.I–II smooth, with weak wrinkles, Ant.IV–V imbricated; Ant.I with a distinct projection at inner apex (Figs 2, 11). Antennal setae short, capitate or blunt at apices; Ant.I–V each with 4, 3, 3, 1, 1–2+1 setae, respectively; PT with 2–3 apical setae. Primary rhinaria not ciliated, secondary rhinaria absent. Rostrum reaching between fore and mid-coxae, with apex dark brown. URS thin and long wedge-shaped (Figs 3, 12), with 2 pairs of primary setae and 2 pairs of accessory setae.</p><p>Thorax. Thoracic nota with irregular polygonal reticulations. Pro-, meso-, and metanotum each with 1 pair of marginal processes, each process with a short capitate seta at apex (Fig. 4); processes on pronotum very small, conical (Figs 4, 13); processes on meso- and metanotum long horn-shaped, strongly imbricated, sometimes also with a short capitate seta near the middle (Figs 4, 14). Thoracic dorsal setae sparse, very short, and capitate. Legs normal, distal part of tibiae with weak spinulose stripes. Setae on hind tibia stiff and pointed. First tarsal chaetotaxy: 3, 3, 3.</p><p>Abdomen. Abdominal tergites I–VI with irregular polygonal reticulations (Fig. 15), tergites VII–VIII with irregular scaly sculptures. Abdominal tergites I–IV each with 1 pair of long horn-shaped and strongly imbricated marginal processes, each process with a short capitate seta at apex (Figs 4, 14). Abdominal dorsal setae similar to those on thorax. Abdominal tergite VIII with 4 long, thick, and capitate setae. SIPH long cylindrical, broad at base, thin at middle, slightly swollen at distal, basal half with distinct imbrications, distal part smooth, obliquely truncated at tip, without flange (Figs 5, 16). Cauda, anal plate, and genital plate with spinulose imbrications. Cauda long tongue-shaped, slightly constricted near the middle (Figs 6, 17), with 4–5 setae. Anal plate semicircular, with 11–14 setae (Figs 7, 18). Genital plate broad round, with 2–3 anterior setae and 4–5 setae along the posterior margin (Figs 8, 19).</p><p>Distribution. China (Tibet).</p><p>Host plant. Salix cupularis .</p><p>Biology. Mostly colonized on undersides of leaves of host plant. The life cycle is unknown.</p><p>Comments. The new species resembles A. (A.) harvensis Verma, but differs from it as follows: body 1.190– 1.478 mm long (in harvensis: body larger, 1.500–1.850 mm long); marginal processes on thoracic nota and abdominal tergites present (in harvensis: absent); antennal tubercles long horn-shaped (in harvensis: finger-like, obtuse at apices); PT 1.27–1.55 times as long as Ant.Vb (in harvensis: equal to or a little longer than Ant.Vb).</p></div>	https://treatment.plazi.org/id/039687B4C10E476AB08227C05A7AF8D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C10E4765B08220015E9CFEBE.text	039687B4C10E4765B08220015E9CFEBE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon (Aspidophorodon) harvensis Verma 1967	<div><p>Aspidophorodon (Aspidophorodon) harvensis Verma, 1967</p><p>Aspidophorodon harvensis Verma, 1967: 507; Eastop &amp; Hille Ris Lambers, 1976: 96; Blackman &amp; Eastop, 1994: 569. Aspidophorodon (Aspidophorodon) harvensis Verma: Remaudière &amp; Remaudière, 1997: 73; Stekolshchikov &amp; Novgorodova, 2010: 44.</p><p>Specimens examined. Paratype: apterous viviparous female, INDIA: Kashmir (Harven), 14.v.1964, on Salix sp., coll. K. D. Verma (BMNH).</p><p>Distribution. India.</p><p>Host plant. Salix sp.</p><p>Biology. This species feeds on undersides of leaves of Salix, sitting tightly along the veins (Verma, 1967). The life cycle is unknown.</p></div>	https://treatment.plazi.org/id/039687B4C10E4765B08220015E9CFEBE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C1004761B08220685B64FB1E.text	039687B4C1004761B08220685B64FB1E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon (Aspidophorodon) musaicus Qiao	<div><p>Aspidophorodon (Aspidophorodon) musaicus Qiao sp. nov.</p><p>(Figs 20–38, Table 1)</p><p>Specimens examined. Holotype: apterous viviparous female, CHINA: Sichuan (Meigu County, altitude 2600 m), 4.v.2005, No. 17257-1-4, host plant unknown, coll. X. L. Huang (NZMC). Paratypes: 1 apterous viviparous female (slide) and 1 apterous viviparous female (COI: KJ374722; Buchnera 16S rRNA: KT 221036), with the same collection data as holotype (NZMC); 1 apterous viviparous female, No. 17257-1-3, with the same collection data as holotype (BMNH).</p><p>Etymology. The new species is named for its mosaic-like markings on thoracic nota. The Latin word “ musaicus ” means “mosaic”.</p><p>Description. Apterous viviparous females: Color of living specimens: green. General measurements see Table 1.</p><p>Color of mounted specimens: body pale (Fig. 28).</p><p>Body. Elongated oval.</p><p>Head. Dorsum of head with distinct wavy sculptures (Figs 20, 29). Median frontal tubercle distinctly protuberant (Figs 20, 29). Antennal tubercles developed, short cylindrical and wrinkled, higher than median frontal tubercle, each with a short blunt seta at apex (Figs 20, 29). Dorsal setae of head short and capitate. Head with 1 pair of cephalic setae, 1 pair of dorsal setae between antennae, and 2 pairs of dorsal setae between compound eyes arranged transversely (Fig. 20). Antennae 5-segmented, Ant.I–II smooth, with weak wrinkles, Ant.III–V with imbrications, Ant.I slightly projected at inner apex (Figs 21, 30). Antennal setae short, weakly capitate or blunt at apices; Ant.I–V each with 3, 3, 2, 1, 1+1 setae, respectively; PT with 2 apical setae. Primary rhinaria ciliated, secondary rhinaria absent. Rostrum reaching mid-coxae, with apex dark brown. URS thin and long wedge-shaped (Figs. 22, 31), with 2 pairs of primary setae and 4–5 accessory setae.</p><p>Thorax. Thoracic nota with irregular polygonal mosaic-like markings (Fig. 32). Pro-, meso-, and metanotum each with 1 pair of strongly imbricated marginal processes, each process with a short capitate seta at apex, sometimes also with a short capitate seta near the middle (Fig. 23); processes on pronotum small, conical or short cylindrical (Figs 23, 33); processes on meso- and metanotum long taper-shaped (Figs 23, 34). Thoracic dorsal setae sparse, very short, fine, and capitate. Legs normal, distal part of tibiae with sparse spinulose short stripes. Setae on hind tibia short, stiff, and pointed. First tarsal chaetotaxy: 3, 3, 3.</p><p>Abdomen. Abdominal tergites I–VI with irregular polygonal mosaic-like markings, those on marginal area small and sparse (Fig. 34), tergites VII–VIII with irregular sculptures, abdominal venter with spinulose transverse stripes. Abdominal tergites I–IV each with 1 pair of long taper-shaped and strongly imbricated marginal processes, each process with a short capitate seta at apex (Figs 23, 34). Abdominal dorsal setae sparse, similar to those on thorax. Abdominal tergite VIII with 6–7 long, thick, blunt or weakly capitate setae. SIPH long cylindrical, broad at base, distal 1/5 slightly expanded and smooth, other parts with distinct imbrications, obliquely truncated at tip, without flange (Figs 24, 35). Cauda long tongue-shaped, slightly constricted near the middle, with spinulose imbrications (Figs 25, 36), with 4 setae. Anal plate semicircular, spinulose, with 14–16 setae (Figs 26, 37). Genital plate broad round, with dense spinulose transverse stripes, with 2–3 anterior setae and 6–9 setae along the posterior margin (Figs 27, 38).</p><p>Distribution. China (Sichuan).</p><p>Host plant. Unknown.</p><p>Biology. Colonized on undersides of leaves of host plant. The life cycle is unknown.</p><p>Comments. The new species resembles A. (A.) harvensis Verma, but differs from it as follows: marginal processes on thoracic nota and abdominal tergites present (in harvensis: absent); antennal tubercles short cylindrical, 1.50–1.70 times as long as median frontal tubercle (in harvensis: long finger-like, about 2.00 times); Ant.I slightly projected at inner apex (in harvensis: with distinct finger-like projections).</p></div>	https://treatment.plazi.org/id/039687B4C1004761B08220685B64FB1E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C105477DB08222405DD2FB33.text	039687B4C105477DB08222405DD2FB33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon (Aspidophorodon) obtusus Qiao	<div><p>Aspidophorodon (Aspidophorodon) obtusus Qiao sp. nov.</p><p>(Figs 39–70, Table 1)</p><p>Specimens examined. Holotype: apterous viviparous female, CHINA: Sichuan (Luding County, Minya Konka), 16.v.2009, No. 22562-1-3-1, on Salix sp., coll. X. M. Su (NZMC). Paratypes: 1 apterous viviparous female and 4 fundatrices (slides), and 1 apterous viviparous female (COI: KJ374723; Buchnera 16S rRNA: KT 221037), with the same collection data as holotype (NZMC); 1 apterous viviparous female and 2 fundatrices, No. 22562-1-1, with the same collection data as holotype (BMNH).</p><p>Etymology. The new species is named for the marginal processes on body having obtuse apices in apterae. The Latin word “ obtusus ” means “dull, blunt”.</p><p>Description. Apterous viviparous females: Color of living specimens: green. General measurements see Table 1.</p><p>Color of mounted specimens: body pale (Fig. 51).</p><p>Body: Elongated elliptical.</p><p>Head. Dorsum of head with distinct wavy sculptures on marginal area, median area with weak wrinkles (Figs 39, 52). Median frontal tubercle distinctly protuberant, hemispherical (Figs 39, 52), with 1 pair of pointed setae on venter. Antennal tubercles developed, short cylindrical and wrinkled, with obtuse apices, higher than median frontal tubercle, each with a stiff, blunt or weakly capitate seta at apex (Figs 39, 52). Dorsal setae of head short and weakly capitate. Head with 1 pair of cephalic setae, 1 pair of dorsal setae between antennae, and 2 pairs of dorsal setae between compound eyes arranged transversely (Fig. 39). Antennae 5-segmented, Ant.I–IV pale in color, Ant.V light brown; Ant.I–II with weak wrinkles, distal part of Ant.III and Ant.IV–V with imbrications; Ant.I slightly projected at inner apex (Figs 40, 53). Antennal setae short and blunt; Ant.I–V each with 4, 4, 2–3, 1, 1–2+1 setae, respectively; PT with 2–3 apical setae. Primary rhinaria ciliated, secondary rhinaria absent. Rostrum reaching mid-coxae, with apex dark brown. URS long wedge-shaped (Figs 41, 54), with 2 pairs of primary setae and 2 pairs of accessory setae.</p><p>Thorax. Pronotum with distinct irregular wavy and small oval sculptures (Fig. 55), meso- and metanotum with wrinkles and small irregular oval markings. Pro-, meso-, and metanotum each with 1 pair of cylindrical, imbricated, and strongly wrinkled marginal processes, each process with a short blunt or weakly capitate seta at the obtuse apex (Fig. 42); processes on pronotum very short (Figs 42, 55); processes on meso- and metanotum sometimes also with a seta near the apical quarter or the middle (Figs 42, 56). Thoracic dorsal setae sparse, short, and weakly capitate. Legs normal, tarsi light brown, claws brown. Distal part of tibiae with spinulose short stripes, 2HT with short imbrications. Setae on legs short, stiff, and pointed. First tarsal chaetotaxy: 3, 2–3, 2.</p><p>Abdomen. Abdominal tergites I–V with reticulations formed by small irregular oval markings (Fig. 57), tergites VI–VIII with distinct irregular scaly sculptures. Abdominal tergites I–IV each with 1 pair of cylindrical, imbricated, and strongly wrinkled marginal processes, each process with a short blunt or weakly capitate seta at the obtuse apex (Figs 42, 56). Abdominal dorsal setae similar to those on thorax. Abdominal tergite VIII with 4–5 long, thick, and capitate setae. SIPH long cylindrical, broad at base, gradually thin towards distal part, distal 1/5 slightly expanded and smooth, other parts with imbrications, obliquely truncated at tip, without flange (Figs 43, 58). Cauda light brown, long tongue-shaped, slightly constricted near the middle, with spinulose imbrications (Figs 44, 59), with 4 setae. Anal plate light brown, semicircular, and spinulose, with 13–15 setae (Figs 45, 60). Genital plate broad round, with dense spinulose transverse stripes, with 2 anterior setae and 6 setae along the posterior margin (Figs 46, 61).</p><p>Fundatrices: Color of living specimens: green. General measurements see Table 1.</p><p>Color of mounted specimens: body pale (Fig. 62).</p><p>Body: Broadly elliptical.</p><p>Head. As in apterous viviparous females, except the following: Antennal tubercles distinctly protuberant, about as high as median frontal tubercle (Figs 47, 63). Dorsal setae of head short and pointed. Antennal setae short and pointed (Figs 48, 64); Ant.I–V each with 3–4, 3–4, 1–2, 1, 2+1 setae, respectively; PT with 2–3 apical setae. URS (Fig. 65) with 2 pairs of primary setae and 4–5 accessory setae.</p><p>Thorax. As in apterous viviparous females, except the following: Marginal processes on Pro-, meso-, and metanotum thicker and much shorter than those in apterae, setae on process short and pointed (Figs 49, 66, 67). Thoracic dorsal setae sparse, short, and pointed. First tarsal chaetotaxy: 3, 3, 2–3.</p><p>Abdomen. As in apterous viviparous females, except the following: Marginal processes on abdominal tergites I–IV thicker and much shorter than those in apterae, setae on process short and pointed (Figs 49, 67). Abdominal dorsal setae pointed. Abdominal tergite VIII with 4 long, thick, and pointed setae. Distal part of SIPH brown (Fig. 68). Cauda elongate triangular, sometimes slightly constricted near the middle (Figs 50, 69), with 4–5 setae. Anal plate (Fig. 70) with 12–15 setae. Genital plate with 2 anterior setae and 4–6 setae along the posterior margin.</p><p>Distribution. China (Sichuan).</p><p>Host plant. Salix sp.</p><p>Biology. Colonized on undersides of leaves of host plants. The life cycle is unknown.</p><p>Comments. The new species resembles A. (A.) harvensis Verma, but differs from it as follows: marginal processes on thoracic nota and abdominal tergites present (in harvensis: absent); antennal tubercles short cylindrical, with obtuse apices, 1.30–1.60 times as long as median frontal tubercle (in harvensis: long finger-like, about 2.00 times); Ant.I slightly projected at inner apex (in harvensis: with distinct finger-like projections); first tarsal chaetotaxy: 3, 2–3, 2 (in harvensis: 3, 3, 3).</p></div>	https://treatment.plazi.org/id/039687B4C105477DB08222405DD2FB33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C119477CB082226B5FC5FD17.text	039687B4C119477CB082226B5FC5FD17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon (Aspidophorodon) salicis Miyazaki 1971	<div><p>Aspidophorodon (Aspidophorodon) salicis Miyazaki, 1971</p><p>(Figs 71–75, Table 1)</p><p>Aspidophorodon salicis Miyazaki, 1971: 183; Eastop &amp; Hille Ris Lambers, 1976: 96; Blackman &amp; Eastop, 1994: 569.</p><p>Aspidophorodon sinisalicis Zhang, 1980 in Zhang &amp; Zhong: 58. syn. nov.</p><p>Trichosiphonaphis lijiangensis Zhang, Zhong &amp; Zhang, 1992: 389 . syn. nov.</p><p>Aspidophorodon (Aspidophorodon) salicis Miyazaki: Remaudière &amp; Remaudière, 1997: 74; Stekolshchikov &amp; Novgorodova, 2010: 44.</p><p>Specimens examined. Syntype: apterous viviparous female, JAPAN: Hokkaido (Sapporo), 15.ix.1967, on Salix udensis, coll. M. Miyazaki (BMNH). Syntypes of Aspidophorodon sinisalicis Zhang, 1980 syn. nov.: 14 apterous viviparous females, CHINA: Beijing (Mt. Baihua), 27.viii.1963, No. Y 437, on Salix sp., coll. F. L. Wang (NZMC). Holotype and paratypes of Trichosiphonaphis lijiangensis Zhang, Zhong &amp; Zhang, 1992 syn. nov.: 3 apterous viviparous females and 1 alate viviparous female, CHINA: Yunnan (Lijiang City, Mt. Yulongxue, altitude 2900 m), 27.v.1980, No. 7165, on Polygonum sp., coll. T. S. Zhong and L. Y. Wang (NZMC). Other materials: 1 apterous viviparous female and 5 apteroid nymphs, CHINA: Gansu (Yuzhong County), 30.vii.1986, No. 8569, on Salix pseudotangii, coll. G. X. Zhang, J. H. Li, and T. S. Zhong (NZMC); 1 apterous viviparous female (slide) and 1 apterous viviparous female (COI: KT 221040; Buchnera 16S rRNA: KT 221038), CHINA: Sichuan (Baoxing County), 14.viii.2003, No. 15038, on Salix sp., coll. K. Guo (NZMC); 4 apterous viviparous females, CHINA: Xinjiang (Burqin County), 23.vii.2007, No. 20604, host plant unknown, coll. D. Zhang (NZMC); 4 apterous viviparous females, CHINA: Ningxia (Jingyuan County, Mt. Liupan, altitude 1984 m), 26.vi.2008, No. 21540, on Salix sp., coll. J. Chen (NZMC); 3 apterous viviparous females and 1 alate viviparous female (slides), and 1 apterous viviparous female (COI: KT 221041; Buchnera 16S rRNA: KT 221039), CHINA: Sichuan (Leshan City), 12.vi.2009, No. 23167, on Salix sp., coll. J. J. Yu and X. Y. Li (NZMC).</p><p>Distribution. China (Beijing, Gansu, Ningxia, Sichuan, Xinjiang, and Yunnan), Japan, Sakhalin and the Kuril Islands.</p><p>Host plants. Polygonum sp., Salix pseudotangii, Salix udensis, Salix sp.</p><p>Biology. Colonized on undersides of leaves of host plant. The life cycle is unknown.</p><p>Comments. Zhang and Zhong (1980) described Aspidophorodon sinisalicis Zhang, 1980 based on apterae from Salix sp. Zhang et al. (1992) described Trichosiphonaphis lijiangensis Zhang, Zhong &amp; Zhang, 1992 based on apterae and alatae infesting Polygonum sp. After reviewing the type specimens of these two species, we found that the apterae of A. sinisalicis (Fig. 72) and T. lijiangensis (Fig. 73) were actually spring morph of apterae of A. (A.) salicis (Fig. 71), and the alatae of T. lijiangensis (Fig. 75) and A. (A.) salicis (Fig. 74) were not significantly different. Therefore, Aspidophorodon sinisalicis Zhang, 1980 and Trichosiphonaphis lijiangensis Zhang, Zhong &amp; Zhang, 1992 should be considered as junior synonyms of Aspidophorodon (Aspidophorodon) salicis Miyazaki, 1971 .</p></div>	https://treatment.plazi.org/id/039687B4C119477CB082226B5FC5FD17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C11B477FB08227CF5C62FD64.text	039687B4C11B477FB08227CF5C62FD64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eoessigia David, Rajasingh & Narayanan 1972	<div><p>Subgenus Eoessigia David, Rajasingh &amp; Narayanan, 1972</p><p>Eoessigia David, Rajasingh &amp; Narayanan, 1972: 35 . Type species: Eoessigia indica David, Rajasingh &amp; Narayanan, 1972; by monotypy.</p><p>Indotuberoaphis Chakrabarti &amp; Maity, 1984: 198 . Type species: Indotuberoaphis sorbi Chakrabarti &amp; Maity, 1984; by monotypy.</p><p>Margituberculatus Zhang, Zhong &amp; Zhang, 1992: 381 . Type species: Margituberculatus longituberculatus Zhang, Zhong &amp; Zhang, 1992; by monotypy. syn. nov.</p><p>Raychaudhuriella Chakrabarti, 1978: 355. Type species: Raychaudhuriella myzaphoides Chakrabarti, 1978: 357; by monotypy.</p><p>Aspidophorodon (Eoessigia) David, Rajasingh &amp; Narayanan: Remaudière &amp; Remaudière, 1997: 74; Stekolshchikov &amp; Novgorodova, 2010: 44; Nieto Nafría et al., 2011: 198.</p><p>Comments. This subgenus is characterized by the presence of spinal processes on body in apterae. It is represented by five species worldwide, including a new combination redescribed herein. The Chinese endemic genus Margituberculatus Zhang, Zhong &amp; Zhang, 1992 is synonymised with Eoessigia in this study (see detailed comments below), Eoessigia is thereby newly recorded from China. Species of Eoessigia mainly infest Rosaceae plants and occur in Canada, China, India, and the Altai Republic.</p></div>	https://treatment.plazi.org/id/039687B4C11B477FB08227CF5C62FD64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C11B477EB08221725FCCFEC7.text	039687B4C11B477EB08221725FCCFEC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon (Eoessigia) indica (David, Rajasingh & Narayanan 1972) David, Rajasingh & Narayanan 1972	<div><p>Aspidophorodon (Eoessigia) indica (David, Rajasingh &amp; Narayanan, 1972)</p><p>Eoessigia indica David, Rajasingh &amp; Narayanan, 1972: 35; Eastop &amp; Hille Ris Lambers, 1976: 188; Chakrabarti &amp; Medda, 1989: 133.</p><p>Raychaudhuriella myzaphoides Chakrabarti, 1978: 357.</p><p>Raychaudhuriella potentillae Chakrabarti &amp; Maity, 1984: 202 .</p><p>Aspidophorodon (Eoessigia) indica (David, Rajasingh &amp; Narayanan): Remaudière &amp; Remaudière, 1997: 74; Blackman &amp;Eastop, 2006: 1098; Stekolshchikov &amp; Novgorodova, 2010: 44.</p><p>Specimens examined. Two apterous viviparous females, INDIA: Himachal Pradesh (Manali), 12.v.1969, on Cotoneaster sp., coll. K. Narayanan (BMNH).</p><p>Distribution. India.</p><p>Host plants. Primary host plants: Cotoneaster obtusus and Cotoneaster sp.; secondary host plant: Potentilla sp.</p><p>Biology. This species is holocyclic and heteroecious, alternating between Cotoneaster and Potentilla (Chakrabarti &amp; Medda, 1989) . It feeds on upper sides of terminal leaves of Cotoneaster near the midrib (David et al. 1972), and infests the under surfaces of leaves of Potentilla without ant-attendance (Chakrabarti &amp; Maity, 1984).</p></div>	https://treatment.plazi.org/id/039687B4C11B477EB08221725FCCFEC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C11A477EB08226385E14FCF9.text	039687B4C11A477EB08226385E14FCF9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon (Eoessigia) longicauda (Richards 1963) Richards 1963	<div><p>Aspidophorodon (Eoessigia) longicauda (Richards, 1963)</p><p>Aspidaphis longicauda Richards, 1963: 297 .</p><p>Eoessigia longicauda Eastop &amp; Hille Ris Lambers, 1976: 95 .</p><p>Aspidophorodon (Eoessigia) longicauda (Richards): Remaudière &amp; Remaudière, 1997: 74; Blackman &amp; Eastop, 2006: 1099; Stekolshchikov &amp; Novgorodova, 2010: 44.</p><p>Specimens examined. Paratype: apterous viviparous female, CANADA: British Columbia (Terrace), 27.viii.1960, on Spiraea sp., coll. W. R. Richards (BMNH).</p><p>Distribution. Canada.</p><p>Host plant. Spiraea sp.</p><p>Biology. This species occurs on the under surfaces of leaves of Spiraea sp. (Richards, 1963). The life cycle is unknown.</p></div>	https://treatment.plazi.org/id/039687B4C11A477EB08226385E14FCF9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C11A477BB08224225B87FEBE.text	039687B4C11A477BB08224225B87FEBE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon (Eoessigia) longituberculatus (Zhang, Zhong & Zhang 1992) Zhang, Zhong & Zhang 1992	<div><p>Aspidophorodon (Eoessigia) longituberculatus (Zhang, Zhong &amp; Zhang, 1992) comb. nov.</p><p>(Figs 76–100, Table 1)</p><p>Margituberculatus longituberculatus Zhang, Zhong &amp; Zhang, 1992: 382 . Margituberculatus longituberculatus Zhang, Zhong &amp; Zhang: Remaudière &amp; Remaudière, 1997: 117; Blackman &amp; Eastop, 2006: 1219.</p><p>Specimens examined. Holotype: alate viviparous female, CHINA: Yunnan (Lijiang City, Mt. Yulongxue, altitude 2900 m), 27.v.1980, No. 7165-1-1-1, on Polygonum sp., coll. T. S. Zhong and L. Y. Wang (NZMC).</p><p>Re-description. Alate viviparous female: Color of living specimens: whitish. General measurements see Table 1.</p><p>Color of mounted specimen: head and thorax dark brown (Fig. 88), abdomen pale in color (Fig. 92).</p><p>Body: Elongated oval.</p><p>Head. Head smooth, with sparse wrinkles dorsally and ventrally. Median frontal tubercle distinctly protuberant; antennal tubercles slightly protuberant, each with a short pointed seta at apex (Figs 76, 88). Dorsal setae of head short and pointed. Head with 1 pair of cephalic setae, 1 pair of dorsal setae between antennae, and 2 pairs of dorsal setae between compound eyes arranged transversely (Fig. 76). Antennae brown, 6-segmented. Ant.I–II smooth, with sparse weak wrinkles, Ant.V–VI imbricated; Ant.I slightly projected at inner apex (Figs 77, 89). Antennal setae short and pointed; Ant.I–VI each with 4, 4, 2, 3, 2, 2+1 setae, respectively; PT with 3 apical setae. Primary rhinaria ciliated. Ant.III–V each with 25–26, 16, and 5 round secondary rhinaria, respectively. Rostrum reaching beyond fore coxae, with apex dark brown. URS long wedge-shaped (Figs 78, 90), with 2 pairs of primary setae and 2 pairs of accessory setae.</p><p>Thorax. Legs brown and long. Distal parts of tibiae dark brown, distinctly spinulous. Setae on legs stiff and pointed. Fore wings with radial sector short and nearly straight, started from the distal end of pterostigma, media twice-branched; hind wings with 2 obliques; wing veins not extending to wing margin (Figs 79, 91).</p><p>Abdomen. Abdomen smooth, with weak wrinkles dorsally and distinct transverse spinulose imbrications ventrally (Fig. 92). Abdominal tergites I and II with small spinal sclerites, tergites I–IV each with 1 pair of marginal sclerites, tergites III–VI with a large sclerotic patch, tergites VII–VIII each with a faint transverse spinopleural band (Fig. 92). Abdominal tergites I–IV each with 1 pair of marginal processes, each process with a stiff and pointed seta at apex; processes on tergites I (Figs 80, 93) and II (Figs 81, 94) small, conical; processes on tergites III (Figs 82, 95) and IV (Figs 83, 96) finger-like, very slender; tergite VIII with a wart-shaped or conical spinal process, bearing 3 long, thick, stiff, and pointed seta at apex and sloping parts (Figs 84, 97). Abdominal dorsal setae sparse, short, and pointed. Abdominal tergite VIII with 4 setae. SIPH long cylindrical, with weak imbrications, thin at middle, swollen at distal quarter, obliquely truncated at tip, without flange (Figs 85, 98). Cauda and anal plate with spinulose imbrications. Cauda tongue-shaped, slightly constricted near the middle (Figs 86, 99), with 4 setae. Anal plate round at apex, with 11 setae (Figs 87, 100).</p><p>Distribution. China (Yunnan).</p><p>Host plant. Polygonum sp.</p><p>Biology. Colonized on undersides of leaves of host plant. The life cycle is unknown.</p><p>Comments. Zhang et al. (1992) erected the genus Margituberculatus with description of type species Margituberculatus longituberculatus Zhang, Zhong &amp; Zhang, 1992 based on one alata. Afterwards, we collected in the type locality several times, but failed to get any specimens of this species. M. longituberculatus is distinguished by marginal processes on abdominal tergites I–IV, single spinal process on tergite VIII, and radial sector of fore wing short and nearly straight in alatae. The latter two characteristics are shared by alatae of A. (E.) longicauda (Richards) . Apterae of A. (E.) sorbi (Chakrabarti &amp; Maity) and fundatrices of A. (E.) indica (David, Rajasingh &amp; Narayanan) also have spinal and marginal processes on body. Therefore, Margituberculatus Zhang, Zhong &amp; Zhang, 1992 should be regarded as a junior synonym of Aspidophorodon (Eoessigia) David, Rajasingh &amp; Narayanan, 1972, and Margituberculatus longituberculatus Zhang, Zhong &amp; Zhang, 1992 should be referred to as Aspidophorodon (Eoessigia) longituberculatus (Zhang, Zhong &amp; Zhang, 1992) comb. nov.</p></div>	https://treatment.plazi.org/id/039687B4C11A477BB08224225B87FEBE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C11F477BB08226E05EF3FCD8.text	039687B4C11F477BB08226E05EF3FCD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon (Eoessigia) sorbi (Chakrabarti & Maity 1984) Chakrabarti & Maity 1984	<div><p>Aspidophorodon (Eoessigia) sorbi (Chakrabarti &amp; Maity, 1984)</p><p>Indotuberoaphis sorbi Chakrabarti &amp; Maity, 1984: 198; Blackman &amp; Eastop, 1994: 727; Remaudière &amp; Remaudière, 1997: 104.</p><p>Aspidophorodon (Eoessigia) sorbi (Chakrabarti &amp; Maity): Stekolshchikov &amp; Novgorodova, 2010: 43.</p><p>Distribution. India.</p><p>Host plant. Sorbus foliolosa .</p><p>Biology. This species lives on undersides of young leaves of Sorbus foliolosa . No attendance was noticed. (Chakrabarti &amp; Maity, 1984).</p></div>	https://treatment.plazi.org/id/039687B4C11F477BB08226E05EF3FCD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
039687B4C11F477BB08224015E9BFBBF.text	039687B4C11F477BB08224015E9BFBBF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aspidophorodon (Eoessigia) vera Stekolshchikov & Novgorodova 2010	<div><p>Aspidophorodon (Eoessigia) vera Stekolshchikov &amp; Novgorodova, 2010</p><p>Aspidophorodon (Eoessigia) vera Stekolshchikov &amp; Novgorodova, 2010: 39 .</p><p>Distribution. The Altai Republic.</p><p>Host plant. Potentilla fruticosa .</p><p>Biology. This species feeds on undersides of leaves of the host plant along the margins (Stekolshchikov &amp; Novgorodova, 2010).</p></div>	https://treatment.plazi.org/id/039687B4C11F477BB08224015E9BFBBF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chen, Jing;Zhang, Bin;Zhu, Xichao;Jiang, Liyun;Qiao, Gexia	Chen, Jing, Zhang, Bin, Zhu, Xichao, Jiang, Liyun, Qiao, Gexia (2015): Review of the aphid genus Aspidophorodon Verma, 1967 with descriptions of three new species from China (Hemiptera: Aphididae: Aphidinae). Zootaxa 4028 (4): 551-576, DOI: 10.11646/zootaxa.4028.4.6
