taxonID	type	description	language	source
039687B0FFA2FFF6C8C6F94D397CAC05.taxon	materials_examined	Type species: Dima elateroides Charpentier, 1825: 191. = Celox dima Schaufuss, 1862: 202.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA2FFF6C8C6F94D397CAC05.taxon	diagnosis	Diagnosis. The current generic concept of Dima includes morphologically diverse species from the Iberian Peninsula, the Balkan Peninsula, the Himalayas, East and South East Asia. Species from the Balkan Peninsula, including the type species Dima elateroides, can be recognized by the following combination of characters: body medium-sized, robust, more or less oval, 8.0 – 17.1 mm long, 2.3 – 2.6 times as long as wide, pale brown to blackishbrown (Figs 1 – 118), antennae weakly serrate, surpassing the pronotal posterior angles by about 1 – 4 antennomeres, antennomere III shortest (or subequal to antennomere II), antennomeres IV – X 1.5 – 2.1 times as long as wide, pronotum 1.3 – 1.6 times as wide as long, moderately convex dorsally, with anterior angles obtuse and posterior angles prominent, acute, more or less distinctly produced outward, tarsomeres III – IV lobate ventrally, and a typically elongate aedeagus with narrow paramera, and apical lobe bearing a more or less developed tooth (Figs 172 – 221). Species of Dima from the Iberian Peninsula differ in having the antennae with short, bulbous, subtriangular antennomeres, and the Asian species in having much longer antennae, often reaching beyond metacoxae, and in the differently shaped male genitalia (Schimmel 1996, Schimmel & Platia 2008, and personal observations).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA3FFF5C8C6FD5D396AAFB5.taxon	materials_examined	Type depositories. Holotype, ♂ (NME), 2 paratypes: ♂, ♀ (NME, PCGP). Type locality. Greece: Peloponnese, Langada Pass. Redetermined material. GREECE: Lakonia, Peloponnese [Peloponnesus], Taigetos Mts., 30 km west of Sparti, 12. – 20. V. 1989, 1 ♂, 2 ♀♀, Persohn leg. (as D. isabellae in Schimmel & Platia 2008) (PCRS); Peloponnese [Pelopones], Taigetos Mts., Tripi, 4. V. 1994, 1 ♂, Ďulík et Jeniš leg. (as D. isabellae in Schimmel & Platia 2008) (PCJM); Peloponnese [Peloponésos], Taigetos, 1. VI. 1991, 1 ♂, 1 ♀, Jan Schneider leg. (as D. isabellae in Schimmel & Platia 2008) (PCJM); Taygetos, 26. V. 1990, 1 ♀, Mička leg. (as D. isabellae in Schimmel & Platia 2008) (PCJM); Peloponnese [Peloponnes], Taigetos Mts., Artemisia env. [Agios Petros env. written erroneously on the label; I. Marvan, pers. comm.], 22. – 30. VI. 2005, 3 ♀♀, Ivan Marvan leg. (as D. isabellae in Schimmel & Platia 2008) (PCJM); Taigetos Mts., 1. – 9. VI. 1996, 1 ♂, 1 ♀ (of 4 ex.), Padovani & Malmusi leg. (as D. hladilorum Schimmel & Platia 2008) (PCGP); Taigetos Mts., 1300 m, VI. 1982, 1 ♂, V. Monzini leg. (as D. hladilorum in Schimmel & Platia 2008) (PCGP). New material. GREECE: Peloponnese [Pelopones], Taygetos, 1935, 1 ♂, Mařan et Štěpánek leg., coll. Bartoň (NMPC); Peloponnese [Pelopones], Taygetos, V. 1935, 1 ♂, Dr. Mařan leg., coll. Bartoň (NMPC); Morea, K. Taigetos, 1. – 9. VI. 1996, 1 ♂, Padovani & Malmusi leg. (PCJM); Peloponnese [Pelopones], Taigetos Mts., h – 1300 m, without date, 1 ♀, B. Lassale leg. (PCVD); distr. Kalamáta, Taigetos Mts., Langada pass env., pinewood, 1300 m (37 ° 3 ' 50.92 " N, 22 ° 15 ' 58.82 " E), 1. VI. 2013, 6 ♂♂, 5 ♀♀, J. Mertlik leg. (PCJM); dtto, 18 ♂♂, 11 ♀♀, V. Dušánek leg. (PCVD); dtto, 15 ♂♂, 8 ♀♀, P. Brůha leg. (PCPB); Taigetos Mts., Kalamata — Sparti, Abzw. Neochori (37 ° 8 ' 46.00 " N, 22 ° 16 ' 12.16 " E), 14. VI. 2006, 4 ♂♂, 1 ♀, wood with Pinus, Manfred Egger leg. (PCME); Taigetos Mts., between Kalamata — Sparti, 19. VI. 2007, 1 ♀, Manfred Egger leg. (PCME); Taigetos Mts., Kalamata — Sparti, Neochori, Agios Nikolaos, 10. – 20. V. 2014, 5 ♂♂, 6 ♀♀, Manfred Egger leg. (PCME).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA3FFF5C8C6FD5D396AAFB5.taxon	diagnosis	Diagnosis. Dima arndti is 11.8 – 14.3 mm long, moderately shiny with pronotum densely punctate (Fig. 119) with short decumbent pubescence, longer only at hind angles, and elytra shiny, covered with sparse short pubescence (Figs 1 – 4). It differs from the similar D. parnonensis Schimmel & Platia, 2008 (Figs 81 – 84) in having a less shiny body, relatively wider pronotum, widest at 2 / 3, with sides abruptly constricted before posterior angles (pronotum widest after middle, with sides almost evenly rounded in D. parnonensis; Figs 119, 149), and the apical lobe of paramera elongate, with inner apical part of paramera almost straight (the apical lobe of paramera shorter, more robust, with inner apical part more curved in D. parnonensis; Figs 172 – 173, 204 – 205). The sclerotized spines of bursa copulatrix as in Fig. 222. Intraspecific variability. Body coloration is variable, dark brown to light brown (Figs 1 – 4).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA3FFF5C8C6FD5D396AAFB5.taxon	distribution	Distribution. Greece (Peloponnese: Taygetos Mts.; Fig. 246).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA3FFF5C8C6FD5D396AAFB5.taxon	discussion	Remark. There is a single male specimen of Dima sp. collected in Crete (Fig. 5), with the following data: " Crete [Kreta Is.], Theriso, VI. 1985, 1 ♂, Dr. P. Schurmann leg. " (as D. hladilorum Schimmel, 1987 in Schimmel & Platia 2008) (PCGP). This specimen is morphologically like D. arndti and D. parnonensis from the Peloponnese, but it has shorter antennae surpassing the posterior angles of pronotum by about three antennomeres, thicker antennal segments, longer pronotum, more emarginate basal part of scutellum, and the inner apical part of paramera only slightly curved (Figs 5, 174). This 30 - year-old record must be confirmed by new material in Crete, and more specimens are needed for further investigation. Therefore, we herein refrain from making any taxonomic conclusions.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA0FFF4C8C6FD8339BDAF25.taxon	materials_examined	Type depositories. Holotype of D. assingi, ♂ (PCRS); paratype of D. assingi, ♀ (PCRS); holotype of D. peloponnensis, ♂ (PCRS); 2 paratypes of D. peloponnensis, ♀♀ (PCRS, PCGP). Type locality. Greece: Peloponnese, Aroania Mts., 1450 m, 10 km E of Kalavrita (for both D. assingi and D. peloponnensis). Redetermined material. GREECE: Morea, Trikala, Kilini Mts., 1. – 9. VI. 1996, 1 ♂, 1 ♀, Padovani & Malmusi leg. (as D. hladilorum in Schimmel & Platia 2008) (PCGP); Peloponnese, Kilini Mts., Trikala, 1100 m, 29. V. – 7. VI. 1998, 1 ♀, L. Saltini leg. (as D. isabellae Dajoz, 1973 in Schimmel & Platia 2 008) (PCRS); Peloponnese, Korinthos: Stilia, 700 m (= Kilini Mts.), 30. V. – 6. VI. 1998, 1 ♂, leg. L. Saltini leg. (as D. raineri Wurst, 1997 in Schimmel & Platia 2008) (PCRS). New material. GREECE: North Peloponnese [Nordpeloponnes], Xylokastro, Zemeno env., Driodasos Mongostou (ca. 37 ° 59 ' 47.42 " N, 22 ° 35 ' 43.23 " E), 28. V. – 7. VI. 1997, 1 ♀, Manfred Egger leg. (PCME).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA0FFF4C8C6FD8339BDAF25.taxon	diagnosis	Diagnosis. Dima assingi is a large, moderately convex species (13.5 – 16.3 mm long), with pronotum with coarse punctation, long straight semi-erect to erect pubescence on the sides (Fig. 120), and elytra with decumbent to semi-erect pubescence basally. Dima assingi is very like D. neumanni Platia, 2013 (Fig. 67 – 74) and D. schnitteri Platia, 2013 (Fig. 105 – 108), which also occur in the northern Peloponnese. They are characteristic by the long straight semi-erect to erect setae on the pronotal sides and semi-erect pubescence on elytra. Together they form a complex of morphologically variable species with unclear specific limits. Dima assingi differs from two above mentioned species in elytra with shorter decumbent to semi-erect pubescence in basal third or half, and in semierect setae in the first half of pronotal sides (D. neumanni and D. schnitteri have longer semi-erect to erect setae in basal third or half of elytra and erect setae in the anterior half of pronotal lateral margin; Figs 120, 146, 157). In contrast to the typical D. neumanni and D. schnitteri, all known specimens of D. assingi have relatively long pronotum, however, some specimens of D. neumanni (mainly from Aroania Mts.) have similarly shaped pronota. Intraspecific variability. There is a variability in the coloration of body and pubescence (the specimens from Aroania Mts. have paler pubescence), pronotal pubescence orientation, and in the shape of apical lobe of paramera (Figs 6 – 9, 175 – 177).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA0FFF4C8C6FD8339BDAF25.taxon	distribution	Distribution. Greece (Peloponnese: Aroania Mts., Kilini Mts.; Fig. 246).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA0FFF4C8C6FD8339BDAF25.taxon	discussion	Remarks. The type specimens of D. assingi and D. peloponnensis were collected at the same time and on the same locality in Aroania Mts. (Schimmel & Platia 2008). They differ only in the body coloration (D. peloponnensis is slightly darker; Figs 6 – 7), surface of pronotum (wrinkled, with different orientation of setae in D. peloponnensis), and the shape of an apical lobe of paramera (shorter in D. peloponnensis; Fig. 177). All these characters are variable in Dima species. The specimens of D. assingi from the Kilini Mts. are similarly colored as D. peloponnensis and some of them vary slightly in the orientation of setae in the pronotum. The wrinkled pronotum is probably a molting defect known also in other species (see e. g. D. fialai sp. nov. or D. fthiotidensis Schimmel & Platia, 2008). Therefore, we herein synonymize Dima peloponnensis Schimmel & Platia, 2008 under Dima assingi Schimmel & Platia, 2008. There are two species of Dima reported from the Aroania Mts., i. e. D. assingi and D. neumanni (Fig. 246). Both probably differ in their habitat preference. Specimens of D. assingi from the Kilini Mts (Zemeno, Stilia, Trikala) were found near seasonal streams in agricultural lands, whilst Dima neumanni was collected in the dark wet gorges in the fir forests of the Aroania and Erimanthos Mts. It must be also noted, that the occurrence of D. assingi in Aroania Mts. was not confirmed by the recent field research by JM and colleagues — only D. neumanni were found in the vicinity of the type locality of D. assingi. We cannot exclude a possibility that the type specimens of D. assingi were mislabelled.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA1FFFBC8C6FE333A0FAD70.taxon	materials_examined	Type material. Holotype: GREECE: distr. Kavála, Pangéo Mts., 2 km S of Paleochori, beechwood, 1080 m (40 ° 55 ' 20.25 " N, 24 ° 10 ' 45.56 " E), 3. VI. 2014, ♂, J. Mertlik leg. (PCJM). Paratypes: GREECE: distr. Kavála, Mt. Pangeo, 1050 m (ca. 40 ° 55 ' 14.15 " N, 24 ° 10 ' 30.76 " E), 5. VI. 2006, 4 ♂♂, 2 ♀♀, beechwood, in trap, A. Müller leg. (PCME); Pangeo Mts., 1375 m, 15. VI. 2007, 1 ♂, 1 ♀, S. Peslier leg. (PCGP); distr. Kavála, Pangéo Mts., 2 km S of Paleochori, beechwood, 1080 m (40 ° 55 ' 20.25 " N, 24 ° 10 ' 45.56 " E), 3. VI. 2014, 27 ♂♂, 10 ♀♀, J. Mertlik leg. (PCJM); distr. Kavála, Pangéo Mts., 2.5 km S of Paleochori, beechwood (40 ° 55 ' 13.79 " N, 24 ° 11 ' 1.79 " E), 3. VI. 2014, 3 ♂♂, 2 ♀♀, J. Mertlik leg. (PCJM); dtto, 3 ♂♂ (PCRK); distr. Kavála, Pangéo Mts., 2 km S of Paleochori, beechwood, 1080 m (40 ° 55 ' 20.25 " N, 24 ° 10 ' 45.56 " E), 6. VI. 2015, 28 ♂♂, 11 ♀♀, J. Mertlik leg. (PCJM); dtto, 33 ♂♂, 4 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 20 ♂♂, 11 ♀♀, P. Brůha leg. (PCPB); dtto, 6 ♂♂, 2 ♀♀, T. Németh leg. (HNHM); dtto, 14 ♂♂, 6 ♀♀ (PCRK); 1. – 10. VII. 2015, 1 ♂, 4 ♀♀, Manfred Egger leg. (PCME).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA1FFFBC8C6FE333A0FAD70.taxon	diagnosis	Diagnosis. Dima bruhai sp. nov. is a larger species characterized by having the pubescence of the pronotal sides semi-erect along the whole length, gradually longer towards posterior angles, and moderately matt elytra with dense, moderately long pubescence (Figs 10 – 13, 121). The other Dima spp. in the surrounding area differ from D. bruhai sp. nov. by different pubescence of pronotum and elytra. Dima schimmeli Platia & Gudenzi, 2009 has body surface covered with the long, straight, semi-erect pubescence (Figs 102 – 104, 156). Dima orientalis sp. nov. has pubescence of the pronotal sides almost decumbent, with longer semi-erect hairs only at hind angles, and elytral pubescence much sparser (Figs 77 – 79, 148). Dima kozufensis sp. nov. has the more compact body shape, pubescence of the pronotal sides decumbent anteriorly, pubescence of elytra slightly sparser and shorter, scutellum basally emarginate and more coarsely punctate, and apical lobe of paramera slightly shorter (Figs 50 – 52, 138, 191). Dima lebenbaueri Schimmel & Platia, 2008 has less matt body surface, longer antennomeres II and III, longer pronotal pubescence, and sparser elytral pubescence (much sparser in the larger specimens from the type locality; Figs 53 – 56, 139 – 140).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA1FFFBC8C6FE333A0FAD70.taxon	description	Description. Holotype, male. Body rather large, 14.8 mm long, 5.9 mm wide, matt. Body dark brown, head paler, antennae (except darker antennomeres II – III), pronotal angles, legs, elytral sutura and margins red-brown; pubescence yellowish (Fig. 10). Head including eyes 0.5 times as wide as pronotum, frons with shallow v-shaped depression in the middle; punctation moderately dense and coarse, intervals between punctures shiny posterad, rugose anterad; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Antennae surpassing the posterior angles of pronotum of about four segments; antennomeres II – III short, antennomere II about 1.5 times longer than wide, antennomere III about 1.3 times longer than wide, length ratio of antennomeres II – IV 1.2: 1.0: 2.0, antennomeres IV – X gradually widened apically, less than 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semi-erect setae. Pronotum 1.4 times wider than long, widest behind middle, moderately convex dorsally; sides evenly convex, straight in lateral view; anterior angles obtuse; posterior angles prominent, sharp, distinctly produced outward; punctation dense, coarse, moderately deep, punctures on disc equally distributed, of about the same size; intervals between punctures about two times a diameter of puncture, moderately shiny. Pubescence on pronotal disc short, decumbent to semi-erect; that on pronotal sides semi-erect along the whole length, gradually longer towards posterior angles (Fig. 121). Prosternum finely, densely punctate, with decumbent to semi-ercet setae, denser at margins; prosternal lobe coarsely punctate, with longer pubescence. Scutellum sub-pentagonal, wide, flattened, sides sub-parallel, in lateral view under the plane of elytra (Fig. 163), frontal margin widely rounded, gradually declined; apex rounded, punctation coarse, dense; pubescence fine, moderately long, semi-erect. Elytra sub-parallel, 3.3 times longer than pronotum, moderately convex, widest at posterior two thirds; striae well developed along the whole elytral length, irregularly interrupted by coarse sparse punctures; surface of interstices flattened, moderately matt, densely and coarsely punctate; punctures distinct, equally distributed, suboval; pubescence dense, moderately long. Aedeagus elongate; paramera long, narrow, its apical lobe long, narrow, subapical tooth minute, subacute (Fig. 178). Female (Fig. 11). Like male but with more robust, suboval body, antennae shorter (about 2.5 segments surpassing posterior angles of pronotum), with segments IV – X shorter (about 1.7 times longer than wide), pronotum slightly wider and more convex, widest at middle. The sclerotized spines of bursa copulatrix as in Fig. 223. Intraspecific variability. Body length: 11.8 – 17.1 mm. Some specimens are paler and have more elongate body shape (2.6 times longer than wide) with longer pronotum, including the largest known specimen (17.1 mm), which is also slightly asymmetrical (Figs 10 – 13). Additionally, some specimens have antennomeres I – III or II – III differently colored than remaining antennomeres (either paler or darker, the latter situation is present in the holotype).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA1FFFBC8C6FE333A0FAD70.taxon	distribution	Distribution. Greece (Pangeo Mts; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA1FFFBC8C6FE333A0FAD70.taxon	etymology	Etymology. The specific epithet is a patronym in honor of Mr. Petr Brůha (Ústí nad Labem, Czech Republic), who collected a part of the type series.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAEFFFAC8C6FCC03972AC59.taxon	materials_examined	Type depository. Syntype, male (BMNH). Other type material unknown. Type locality. Croatia: Omiš [Dalmatien: Almissa]. New material. BOSNIA AND HERZEGOVINA: Prenj planina, no date, 1 ex. [without head] (PCGP); CROATIA: Zaostorog, Rilič [vápencové škrapy Riliče], 20. III. 2000, 1 ♂, on the ground between stones [skalní lesostep, na zemi mezi kameny], R. Gabzdil leg. (PCRG); Zaostorog, 22. – 24. III. 2001, 1 ♀, P. Arpáš leg. (PCJM); MONTENEGRO: Jugoslavia, Petrovac, 16. VI. 1974, 1 ♀, J. Hladil leg. (PCRS); Bar distr., Mt .. Vrsouta, Sutorman, mixed forest (42 ° 9 ' 18.32 " N, 19 ° 5 ' 52.81 " E), 1. – 2. VI. 2014, 2 ♀♀, singled at night, swept and beaten, V. Dušánek leg. (PCVD); dtto, 1 ♂, 2 ♀♀, T. Németh leg. (PCRK), dtto, 2 ♀♀, Kundrata lgt. (PCRK); Dalmatien, Budva, coll. Reitter, 1 ♀ (HNHM); Dalmatia, coll. E. Frivaldszky, 1 ♂ (HNHM); Dalmatia, coll. Lichtn., 1 ♀ (HNHM).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAEFFFAC8C6FCC03972AC59.taxon	diagnosis	Diagnosis. Dima dalmatina is a medium-sized species (body length: 11.5 – 13.5 mm) easily recognizable by its elongate body, relatively long antennae (surpassing hind angles by about 3.5 segments in males), shiny pronotum with sides with the short semi-erect setae along the whole length, elytra with short dense decumbent pubescence, and elytral interstices matt, microreticulated (Figs 14 – 19, 122). This species is very like D. marvani Mertlik & Dusanek, 2006 (Figs 62 – 65). The typical representatives of D. dalmatina differ from the typical D. marvani by more distinctly defined keel between frons and clypeus, shorter antennae in males (about 55 % of body length in D. dalmatina, more than 60 % in D. marvani), wider pronotum (1.4 times wider than long in D. dalmatina, 1.3 times wider than long in D. marvani), pronotal pubescence denser, near sides directed towards posterior angles (Figs 122, 144), and apical lobe of paramera shorter (Figs 180, 198). Intraspecific variability. There are two distinct Dima populations in Rumija Mts. in Montenegro; one resembles more D. dalmatina and the second is more like D. marvani. However, they both differ from the typical representatives of respective species (Figs 18 – 19, 64 – 65). Both of these local forms have a darker body and slightly more convex and longer pronotum than most of the typical specimens. The specimens similar in most aspects to D. dalmatina have longer apical lobes of paramera like those found in D. marvani (Fig. 179). Unfortunatelly, without larger series of both species collected from more localities we are unable to make any taxonomical conclusions and we keep the populations from Rumija Mts. in D. dalmatina and D. marvani.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAEFFFAC8C6FCC03972AC59.taxon	distribution	Distribution. Bosnia and Herzegovina, Croatia, Montenegro (Fig. 243). First record for Bosnia and Herzegovina.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAEFFFAC8C6FCC03972AC59.taxon	discussion	Remarks. The syntype of D. dalmatina deposited in the BMNH, London, is clearly a male, but without genitalia neither inside the abdomen nor glued on a label (Figs 14 – 15); there is a label " Abdomen seems to be empty, R. Schimmel 19 85 ". A specimen from Greece (Florina, 1100 m, 11. VII. 1981, Osella leg.), which is mentioned under D. dalmatina by Guglielmi & Platia (1985: 194), is probably an erroneously identified specimen. That specimen was identified by Dr. Guglielmi and should be deposited in the Verona museum (G. Platia, pers. comm.), however it is probably misplaced in the collection (R. Salmaso, pers. comm.). It might be conspecific with D. florinensis Platia, 2012, which occurs in that area (Platia 2012). One specimen from Rumija Mts. recorded under D. dalmatina in Schimmel & Platia (2008) was redetermined as D. marvani.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAFFFF9C8C6FD203A0FAC95.taxon	materials_examined	Type material. Holotype: MACEDONIA: distr. Kruševo, Kruševo env., beechwood 1 km NW of city, 1340 m (41 ° 22 ' 36.94 " N, 21 ° 14 ' 8.74 " E), 25. V. 2014, ♂, J. Mertlik leg. (PCJM). Paratypes: MACEDONIA: same data as for the holotype, 42 ♂♂, 10 ♀♀, J. Mertlik leg. (PCJM), dtto, 2 ♂♂, 1 ♀ (PCRK); 32 ♂♂, 9 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 34 ♂♂, 12 ♀♀, P. Brůha leg. (PCPB); Pelagonia reg., Krusevo, Fagus forest, singled at night, 41 ° 22 ' 38.01 " N, 21 ° 14 ' 0.25 " E, 5. VI. 2014, 8 ♂♂, 4 ♀♀, leg. V. Dušánek, R. Kundrata & T. Németh (HNHM); dtto, 39 ♂♂, 25 ♀♀ (PCRK); dtto, 17 ♂♂, 17 ♀♀, V. Dušánek leg. (PCVD); prov. Krusevo, Krusevo, Crn Viv, 1350 m, 1997. V. 5. – 6., 1 ♂, leg. A. Podlussány & I. Rozner (HNHM). Other material examined. MACEDONIA: prov. Skopje, Skopska Crna Gora, 500 – 600 m, Mon. Sv. Jovan, 8. VI. 1998, 2 ♀♀, leg. I. Rozner (HNHM). These specimens were probably mislabelled (I. Rozner, pers. comm.).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAFFFF9C8C6FD203A0FAC95.taxon	diagnosis	Diagnosis. Dima dusaneki sp. nov. is a medium-sized to large species with more convex, suboval body and shiny elytral intervals with very sparsely distributed thin setae (Figs 2 0 – 21). It shares the similar body coloration, pronotal punctation and the pubescence of the pronotal sides with D. raineri (shorter decumbent at anterior 1 / 2 or 2 / 3, longer semi-erect posteriorly; Figs 123, 154 – 155). However, D. dusaneki sp. nov. differs from D. raineri (and also other Dima species in the region) in having a robust body and wide, laterally evenly rounded pronotum, which is widest medially (Figs 20 – 21). Dima macedonica Schimmel, 1993 and D. florinensis differ from D. dusaneki sp. nov. in the more distinctly double-sized punctation of pronotum and the pubescence of the pronotal sides which is semi-erect along the whole margin length (shorter anteriorly in D. macedonica, long in D. florinensis; Figs. 129, 141 – 143). Dima fialai sp. nov. has the pronotum widest at posterior half, narrowed toward anterior margin, with sparser punctation (Figs 29 – 30), and the pubescence of the pronotal sides short and almost decumbent at anterior 3 / 4 of the margin length and long semi-erect setae only near hind angles (Fig. 128).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAFFFF9C8C6FD203A0FAC95.taxon	description	Description. Holotype, male. Body medium-sized, 12.5 mm long, 5.2 mm wide, shiny, covered with short dense pubescence. Body blackish-brown, with paler, reddish-brown legs, antennae, elytral sutura and margins; pubescence yellowish (Fig. 20). Head including eyes 0.5 times as wide as pronotum, frons slightly depressed; punctation sparse and coarse, moderately deep, intervals between punctures shiny; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina short, indistinct. Antennae moderately long, surpassing the posterior angles of pronotum of about 3.5 segments; antennomeres II and III subtriangular, antennomere II longer than antennomere III, both less than 1.5 times longer than wide, together 1.3 times longer than antennomere IV, length ratio of antennomeres II – IV 1.2: 1.0: 1.7, antennomeres IV to X gradually widened apically, about 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semi-erect setae. Pronotum 1.4 times wider than long, widest just behind middle, convex dorsally; sides convex; gradually narrowed towards frontal margin; anterior angles obtuse; posterior angles prominent, sharp, slightly produced outward; punctation sparse, moderately fine, punctures on disc equally distributed, of double size; intervals shiny. Pubescence on pronotal disc short, decumbent; that on pronotal sides at anterior 1 / 2 or 3 / 5 short and decumbent, pubescence longer and semi-erect to erect posteriorly, long and erect at hind angles (Fig. 123). Prosternum finely, sparsely punctate, with short semi-erect setae, sparse medially, denser at margins; prosternal lobe moderately coarsely punctate, with longer semi-erect pubescence. Scutellum sub-pentagonal, with sides rounded, with shallow median depression, in lateral view slightly raised above the plane of elytra, widest at middle; frontal margin rounded, gradually declined; apex widely rounded, punctation sparse, coarse; pubescence fine, moderately long, semi-erect. Elytra sub-parallel, 3.3 times longer than pronotum, convex, widest at posterior two thirds; striae well developed along the whole elytral length, irregularly interrupted by coarse sparse punctures; surface of interstices flattened, shiny, densely and finely punctate; punctures suboval, equally distributed, of the same size; pubescence short, sparse. Aedeagus elongate; paramera long, narrow, its apical lobe moderately long, subapical tooth subacute (Fig. 181). Female (Fig. 21). Like male, but more robust, with shorter antennae (surpassing the posterior angles of pronotum about one segment), pronotum distinctly wider than in male (1.6 times wider than long), scutellum distinctly wider. The sclerotized spines of bursa copulatrix as in Fig. 224. Intraspecific variability. Body length: 10.6 – 15.2 mm.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAFFFF9C8C6FD203A0FAC95.taxon	distribution	Distribution. Macedonia (Kruševo; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAFFFF9C8C6FD203A0FAC95.taxon	etymology	Etymology. The specific epithet is a patronym in honor of Mr. Václav Dušánek (Zábřeh, Czech Republic), who collected a part of the type series.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFACFFFFC8C6FCE33A61AB1D.taxon	materials_examined	Type depository. Lectotype (SMTD), paralectotype (SMTD), sex undetermined (Schimmel 1996). Type locality. Austria: Styria [Styriae alpibus]. Redetermined material. CROATIA: Dalmatia, Dubrovnik, V. – VII. 1987, 1 ♂, no further data (as D. macedonica in Schimmel & Platia 2008) (PCJM); MONTENEGRO: Durmitor Mts., Stuoc env., 1850 m, VII. 2001, 1 ♂, 3 ♀♀, traps, Ivan Marvan leg. (as D. macedonica in Schimmel & Platia 2008) (PCJM); Durmitor, Žabljak, 1450 m, VI. 1982, 1 ♀, no further data (as D. macedonica in Schimmel & Platia 2008) (PCJM). New material. AUSTRIA: Styria, no further data, 1 ♀ (NMPC); Karawanken, Karnten Waidisch, 10. V. 1985, 1 ex., M. Zeising leg. (PCGP); Kärnten, Plöckenpass, 10. – 28. VI. 2011, 3 ♀♀, M. Egger leg. (PCME); Stiria, D. Ghiliani, 1 ♂ (MCNG); BOSNIA AND HERZEGOVINA: Jablanica, no further data, ♂, (NMPC); Peričica, 1500 m, 2. VI. 1967, 1 ♀, Mařan leg. (NMPC); Šator lake (44 ° 9 ' 56.68 " N, 16 ° 36 ' 13.04 " E), 1. VII. 2013, 17 ♂♂, 2 ♀♀, V. Dušánek leg. (PCVD); dtto, 1 ♂, 1 ♀ (PCJM); Bjelašnica Mts., Umoljani, h – 1300 m, 8.7.2010, 1 ♀, V. Major leg. (PCVD); Bosnia, Stanski Most, Hrustovo, Hrustovača pečina cave, 5. IV. 2015, 1 ♀, R. Lohaj leg. (PCJM); Bosnia, Preslica pl., 1 ♂ (HNHM); Bosnia, Reitter, 1 ♂ (HNHM); Herzegowina, Reitter, 1 ♂ (HNHM); Bosnia, Zepce, 1 ♂ (HNHM); Bosnia, coll. Lichtneck, 1 ♂ (HNHM); Ivan Bosnien Gerger, Coll. Dr. R. Streda, 1 ♀ (HNHM); Trebevic, Coll. Apfelbeck, 3 ♂♂ (HNHM); Ivan – pl., Coll. Apfelbeck, 3 ♀♀ (HNHM); Ivan – pl., Coll. Apfelbeck, 1 ♂ (HNHM); Bosn, Ivan, Coll. Apfelbeck, 1 ♂, 1 ♀ (HNHM); Krug – pl., Coll. Apfelbeck, 1 ♀ (HNHM); Apfelbeck, Pazarić, Umgb. Saraj, 1 ♂ (MCNG); Apfelbeck, Umgb. Saraj, 1 ♂ (MCNG); Ivan, 1 ♀ (MCNG); Bosnai, Koricna, O Leonhard, 3 ♂♂ (MCNG); BULGARIA: 90 km from Sofia, from under a rock, daytime, oak forest, 1987. VII. 9., 1 ♂, leg. Nikola Rahmé (PCBS); CROATIA: Dalmatia, D. Candéze, 1 ♂ (MCNG); Dalmatia, no further data, 1 ♀ (MCNG); Cro. Bernh, Fuzine, 97, 1 ♂ (MCNG); 1 ex., Europe, Sharp Coll., 1905 - 313, abdomen missing (BMNH); 1 ex., Croatia, Janson coll., 1903 - 130, Dr. Dohrn (BMNH); 1 ex., Croatia, further data unreadable, Janson coll., 1903 - 130 (BMNH); 2 ex., Croatia, Janson coll. ex Schaum., 1903 - 130 (BMNH); Lokve, no further data, 1 ♀, Ing. Machulka leg. (NMPC); Velebit, 1 km SW Markoviči, 1010 m, 10. – 14. VI. 2011, 2 ♂♂, swept singled, Tamás Németh leg. (PCJM); Velebit, Veliki Zavižan, 1600 m, 2. – 10. VII. 2015, 1 ♂, R. Udržal leg. (PCRU); Velebit, Karlobag / Ostarjie, 8. VI. 1984, Padovani & Malmusi leg. (PCGP); prov. Karlobag, 2 km E Crni Dabar, 9. – 10. VI. 2014, 1 ♂, M. Šárovec leg. (PCPB); Velebit, Rizvanusa, 8. VI. 1984, Padovani & Malmusi leg. (PCGP); Mt. Novi Vin., 700 – 900 m, 10. VI. 1995, Padovani & Malmusi leg. (PCGP); Mt. Platak, 17. VII. 1977, 1 ex., Perdisa leg. (PCGP); Dalmatia, without date, 1 ex., Schreitter leg. (PCGP); Velebit, Apatisan, mixed forest, 1000 m, sifted, 21. – 25. VI. 2008, 1 ♀, leg. T. Németh & N. Rahmé (HNHM), dtto, 1 ♀ (PCRK); Velebit, 1 km SW Markovici, 1010 m, singled at night, 10. – 14. VI. 2011, 3 ♂♂, 4 ♀♀, leg. Gy. Németh, T. Németh & B. Szelenczey (HNHM); dtto, 16 ♂♂, 32 ♀♀ (PCBS); Lika-Senj county, Velebit Mts., 3 km SE Oltari, Fagus forest, singled at night, 44 ° 51 ' 32.70 " N, 15 ° 0 ' 24.20 " E, 30. V. 2014, 8 ♂♂, 12 ♀♀, leg. V. Dušánek, R. Kundrata & T. Németh (PCRK); dtto, V. Dušánek leg., 9 ♂♂, 9 ♀♀ (PCVD); Croatia, coll. Reitter, 1 ♂, 1 ♀ (HNHM); Velebit, Coll. Reitter, 1 ♂ (HNHM); Lokve (Croatia), Victor Stiller coll., 1 ♂ (HNHM); Delnice (Croatia), Victor Stiller coll., 1 ♂ (HNHM); Dalmatien, 1 ♀ (HNHM); Dalmatia, Coll. K. Fuss, 1 ♀ (HNHM); Delnice, (1) 904, Bíró, 17. maj., 2 ♂♂ (HNHM); Fuzine, Pavel, 1 ♂, 2 ♀♀ (HNHM); MACEDONIA: Sar Pl., 1935. VII. 7., leg. Dr. J. Fodor, 1 ♂, 1 ♀ (HNHM); MONTENEGRO: Murino, 15. VI. 1984, 1 ex., M. & J. Hladilovi leg. (PCGP); Titograd, 19. V. 1986, 1 ex., G. Sama leg. (PCGP); dtto, 1 ♂ (PCVD); Žabljak Mt., 1500 m, 28 – 29. VI. 1958, 1 ♂, Mařan et Hoberlant leg. (NMPC); Durmitor, Zmijinje jezero, 1450 m, 27. VI. 1958, 1 ♀, Mařan et Hoberlant leg. (NMPC); Biogradska Gora, Biogradsko lake, 5. – 6. VII. 2010, 1 ♀, D. Čagánek leg. (PCTS); dtto, 1 ♂, 1 ♀, D. Čagánek leg. (PCPB); dtto, 1150 – 1290 m, 28. – 29. V. 2014, 5 ex., P. Jansa leg. (PCPB); dtto, 2. VI. 2014, 2 ♂♂, 3 ♀♀, R. Škoda leg. (PCJM); Mojkovac, Bjelasica Mts., Bjelogradsko jezero, 15. VI. 2006, 1 ♂, L. Blažej leg. (PCPB); Durmitor Mts., Žabljak env., 12. VI. 2012, 1 ♂, 1 ♀, J. Stanovský leg. (PCTS); Komovi Mts., Třešněvik, 6. VI. 2012, 2 ♂♂, J. Stanovský leg. (PCTS); Třešnjevik, Komovi Mts., lesní pásmo pod Vasolijevičky Kom, 1500 – 1750 m, 11. – 12. VI. 2006, 1 ♂, 2 ♀♀, L. Blažej leg. (PCPB); Vojnik Mts., Pružine env., 4 km of Gornja Brezna (N 42 ° 58 ' 57 '', E 18 ° 55 ' 55 ''), 1142 m, 23. – 24. V. 2014, 4 ♂♂, 1 ♀, J. Pelikán et R. Škoda leg., 4 ♂♂ (PCJM), dtto, 1 ♀ (PCLS); dtto, Prokletije, Gusinje, VI. 1937, 1 ♀, Dr. Purkyně leg. (NMPC); Čakor, svahy, 1800 – 1900 m, 11. VI. 1967, 1 ♀, Mařan leg. (NMPC); Ternova (?), Coll. Reitter, 1 ♂ (HNHM); Han Garncic, 1983. VII. 19. – 20., leg. Dr. J. Fodor, 1 ♂, 3 ♀♀ (HNHM); Niksic, 1939. VII. 11., leg. Dr. J. Fodor, 1 ♂ (HNHM); Donja Polja, Vrenicki – potok, (Zolski – ljevak) 2003.05.0 6., Juhász P., Kovács T., Sevola, P., 1 ♂ (HNHM); Biogradska Gora, env. jezero, 28. – 29. V. 2014, 2 ♂♂, 2 ♀♀, P. Hubený leg. (PCPH); Prokletje Mts., Gusinje-Grbaja valley, 26. VII. 2014, 1 ♂, 1 ♀, P. Hubený leg. (PCPH); Durmitor, road Žabljak – Šavnik, Bukovica env., ca. 1500 m, 15. – 30.6.2015, 17 ♂♂, 15 ♀♀, old beech forest, traps, I. Marvan leg. (PCJM); Durmitor, road Žabljak – Plužine, Trsa env., ca. 1600 m, 15. – 30.6.2015, 1 ♂, 1 ♀, beech forest, singled at night, I. Marvan leg. (PCJM); Turjak Mts., Rožaje env., ca. 1400 m, 15. – 30.6.2015, 8 ♂♂, 4 ♀♀, spruce / fir forest, traps, I. Marvan leg. (PCJM); Durmitor Mont., 3. VII. 1958, Zmijinje Jezero, 1400 m, leg. Kaszab & Székessy, 2 ♂♂ (HNHM); Durmitor Mont., 27. VI. 1958, Crna Jezero, 1400 m, leg. Kaszab & Székessy, 1 ♂ (HNHM); ITALY: Fusine Laghi Tarvisio Udine, 1.15. VII. 1956, 1 ♂, G. Binaghi (MCNG); ROMANIA: Banat, Bocşa [Süd Ungarn, Neu Bogsán], no further data, 1 ♀, E. Merkl leg. (NMPC); Banat, Semenic [Hungaria, Szemenik], no further data, 2 ♀♀ (NMPC); Mţii Poiana Ruscă, Luncanii de Jos env., 7. – 8. V. 2012, 1 ♂, Vladimír Zieris leg. (PCJM); Gorj county, Topenia Valley, 760 m, VI. 2015, 1 ♂, 1 ♀, C. Busi leg. (PCGP); Hunedoara, Câmpu lui Neag, 900 m, VI. 2015, 1 ♂, C. Busi leg. (PCGP); S of Sibiu, Brezoi env., 10. VI. 2007, Jan Hron leg. (PCBZ); Jud. Caras – Severin, 5 km SW Cerna – Sat, Cerna valley, sifted from among stones, 23. V. 2009, 1 ♂, leg. A. Kotán, A. Márkus & T. Németh (PCAK); Jud. Caraş – Severin, Munţii Locvei, 3.5 km NNE Sfânta Helena, Peştera cu apa din Valea Ceucâ, pond, 320 m wet, humusly soil with stones at rock wall, soil – washing (420), 44 ° 42 ' 14 '' N, 21 ° 43 ' 44 '' E, 08. X. 2009, 1 ♂, Gy. Makranczy (HNHM); Baile Herculane, 1988. IV. 5., leg. Matus G., 2 ♀♀ (HNHM); Mons Risnyak, Coll. Mihók, 1 ♀ (HNHM); Herkulesfürdô, Hungaria, coll. R. Streda, 2 ♀♀ (HNHM); Herkulesf., 2 ♀♀ (HNHM); Fuss, 3016, Ferencfalva, Friv & Pavel, 1 ♂ (HNHM); Mehadia, 1 ♀ (HNHM); Herkulesfürdô, Pável, 1 ♂ (HNHM); Ferencfalva, Friv & Pavel, 1 ♂, 1 ♀ (HNHM); Herkulesfürdô, Cserna – völgy, leg. Pavel, 1885, 1 ♂ (HNHM); Herkulesbad (Hungaria), Victor Stiller coll., 1 ♀ (HNHM); Transsylv., Krassó – Szörény m., Mehádia, 1915. IV. 30., leg. Dr. J. Fodor, 1 ♂ (HNHM); Ferenczfalva, Kanabé Dezsô, Coll. Kanabé, 20 ♂♂, 7 ♀♀ (HNHM); Herkulesfürdô, Hungaria, coll. Dr. R. Streda, 1 ♀ (HNHM); Transsylv., Krassó – Szörény m., Mehádia, 1915. IV. 25., leg. Dr. J. Fodor, 1 ♂ (HNHM); dtto, 1915. IV. 20., 2 ♀♀ (HNHM); Vulkán, sz., Strázsa Csik, 2 ♂♂ (HNHM); SERBIA: Majdanpek, 8. V. 2015, 1 ♂, R. Udržal leg. (PCRU); Javor, Kumanica env., 22. V. 1991, 1 ♀, P. Moravec leg. (PCTS); Golija Mts., Ivanjica env. – pass, 21. V. 1991, 1 ♂, P. Moravec leg. (PCTS); Maljen planina, road Bukovi – Razana, pass, beechwood, 770 m (44 ° 7 ' 2.67 " N, 19 ° 55 ' 5.13 " E), 25. V. 2013, 4 ♂♂, 3 ♀♀, J. Mertlik leg. (PCJM); dtto, 5 ♂♂, 2 ♀♀, V. Dušánek leg. (PCVD); dtto, 2 ♀♀, P. Brůha leg. (PCPB); Valjevo — Požega road, Maljen planina, Divči Bare env., ca. 900 m, 15. – 30.6.2015, 3 ♂♂, edge of pine forest, I. Marvan leg. (PCJM); Jošanička Banja — Belo Brdo road, Kopaonik, ca. 1800 m, 15. – 30.6.2015, 1 ♂, 1 ♀, traps, I. Marvan leg. (PCJM); Užica — Nova Vároš road, Kokin Brod env., ca. 800 m, 15. – 30.6.2015, 1 ♂, edge of beech forest, trap, I. Marvan leg. (PCJM); Srbsko, prov. Kosovo, Šar planina, Ljuboten [Serb. Albania, Ljuboten – Šara pl.], VI. 1930, 1 ♀, Dr. Purkyně leg. (PCVD); SLOVENIA: Carniolia, Reitter, 257, G. C. Champion Coll., B. M. 1927 - 409, 1 ex. (BMNH); Planšarsko jezero [Zg. Jezero], no further data, 1 ♀, Coll. Štěrba (NMPC); Kočevje [SHS, CARN, Kočevjo], no further data, 1 ♂, Ing. Machulka leg. (NMPC); Kočevje, Ledenik SW – Hang, 880 m (Stojna), 19. VI. 1994, 2 ♂♂, 2 ♀♀, Manfred Egger leg. (PCME); Kočevje, Rajhenovski Rog Süd, 850 m, Pragost Kočevski Rog, 27. VI. 1994, 1 ♂, 3 ♀♀, Manfred Egger leg. (PCME); Ljubljana, 25 km S Cerkniško lake, 14. V. 2014, 2 ♂♂, 1 ♀, E. & P. Hajdaj leg. (PCJM); Postumia, VII. 1995, 1 ex., C. Sola leg. (PCGP); Trnovski gozd, Selovec, 7. VII. 1999, 1 ♂, 1 ♀, T. Sitek leg. (PCTS); Trnovski gozd, Krnica, 8. VII. 1999, 1 ♀, T. Sitek leg. (PCTS); Trgovski Gvozd, Čaven, 11.7.1994, 1 ♂, Šárovec leg. (PCVD); Mangart, no further data, 1 ♂ (MCNG); Dobrovlje, 9. – 13. VI. 1998, 1 ♀, T. Růžička leg. (PCBZ); dtto, 9. – 10. VII. 1998, 1 ♂, 1 ♀, Kovařík leg. (PCVD); Vene Giulia, Selva Tarnova, Loqua, VI., 1934, C. Alzona, 1 ♂, 1 ♀ (MCNG); Trnovsky Gora Goliak, 25. VI. 67, M. E. Franciscio, 6 ♂♂, 6 ♀♀ (MCNG); Stojna Mts., Gozdna Koča, 900 m, 8. V. 2001, 1 ♂, J. Vávra leg. (PCJV); dtto, 1 ♂ (PCPB); dtto, 23. – 24. VI. 2001, 2 ♂♂, 4 ♀♀, T. Sitek leg. (PCTS); dtto, 6. – 7. VII. 2002, 1 ♂, T. Sitek leg. (PCTS); Mt. V. Snežnik, 1450 – 1600 m, 4. VII. 2002, 1 ♂, T. Sitek leg. (PCTS); dtto, 26. – 27. VI. 2003, 2 ♂♂, T. Sitek leg. (PCTS); Sněžnik, Okroglina (45 ° 34 ' N, 14 ° 21 ' E), 20. VI. 2001, 1 ♂, P. Kabátek leg. (PCBZ); Illirska Bistrica, Mt. Snežnik, 900 m, 1 ♀, Pierer Fritz leg. (PCME); Illirska Bistrica, Snežnik, Svicaki, 15. – 28. VI. 2009, 6 ♂♂, 8 ♀♀, Manfred Egger leg. (PCME); Lokve – Predmeja, LF, 13. VI. 2009, 3 ♂♂, 5 ♀♀, Manfred Egger leg. (PCME); Brovnica, Koselevka Jama env., 20. V. 2004, 1 ♂, Manfred Egger leg. (PCME); Jezersko, VII. 1936, 1 ♀, Hájek leg. (PCBZ); Tisovec, Mala Rib. Gora, 30. IV. 72, 1 ex. (JPMP).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFACFFFFC8C6FCE33A61AB1D.taxon	diagnosis	Diagnosis. This species is easily recognizable from all its congeners by the oval body shape, shiny pronotum with sparse and shallow punctation, with the disc glabrous and the outer parts covered with sparse pubescence, and elytra with glabrous surface and individual setae only at margins, with shallow striae, missing at apical third of abdomen in some cases (Figs 22 – 24, 124). The sclerotized spines of bursa copulatrix as in Fig. 225. Intraspecific variability. Body length: 8.8 – 12.5 mm. Dima elateroides is variable in the striae on elytra. There are specimens with no, or almost no, striae in the apical third of the elytra (mostly found in the populations from Austria, Italy, Slovenia, Croatia, Bosnia and Herzegovina; Figs 23 – 24), and specimens with distinct striae along the whole length of the elytra (especially in the populations from Romania and Serbia; Fig. 22). In the southern part of the distributional area of D. elateroides (Montenegro, Kosovo), these two forms occur together equally often.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFACFFFFC8C6FCE33A61AB1D.taxon	distribution	Distribution. Austria, Bosnia and Herzegovina, Bulgaria, Croatia, Italy, Macedonia, Montenegro, Romania, Slovenia, Serbia (Fig. 243). First records for Bulgaria and Macedonia. Tesař (1927) reported one specimen from Ukraine (Chust), however, this record must be confirmed by further data. Dajoz (1973) listed also southern part of Hungary for D. elateroides, but this record is probably based on a mislabelled material or specimen (s) collected outside of the present-day Hungary.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAAFFFEC8C6FA6B3ABCAF78.taxon	materials_examined	Type depository. Holotype, ♀ (PCGP). Type locality. Greece: Etolia – Akarnania, Panetolikó Mts., Nerosirtis, 850 m.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAAFFFEC8C6FA6B3ABCAF78.taxon	diagnosis	Diagnosis. This species is known only from the large (14.3 mm) female holotype specimen (Fig. 25). It differs from D. evritaniensis Schimmel & Platia, 2008 and D. fthiotidensis by the shiny pronotum with sparser punctation (Figs 125, 126, 130) and the scutellar frontal margin flattened, declined gradually to the plane of elytra (frontal margin of scutellum convex, declined steeply in both other species). Dima pelionensis sp. nov. has also a flattened scutellum and shiny pronotum with sparse punctation, but it differs from D. etoliensis in the sides of pronotum with decumbent setae in the anterior two thirds (D. etoliensis has the pronotal sides with semi-erect setae along its whole length; Figs 125, 152). Dima zbuzeki sp. nov. is smaller, suboval, and has shorter antennae (in females) with shorter antennomeres II and III (Figs 116 – 118). The most similar species to D. etoliensis is D. hladilorum, but D. etoliensis is larger, with a flattened scutellum, more parallel-sided elytra, distinct striae along the whole length of elytra, and antennomeres IV – X narrow, slightly more than two times longer than wide (Figs 25, 47 – 49). The sclerotized spines of bursa copulatrix as in Fig. 226.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFAAFFFEC8C6FA6B3ABCAF78.taxon	distribution	Distribution. Greece (Panetolikó Mts.; Fig. 245).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFABFFFEC8C6FECD3898A990.taxon	materials_examined	Type depositories. Holotype, ♂ (MRSN), 3 paratypes: ♀♀ (MCSB, PCGP, PCRS). Type locality. Greece: Evritania province, Panetoliko Mts., 620 m, Proussos env. Redetermined material. One of the paratypes of D. evritaniensis, a male from Mt. Ossa (PCGP; Fig. 87), belongs under D. pecoudi Fleutiaux, 1944. New material. GREECE: distr. Evritania, Timfristos Mts. [Veluchi], 1 ♀, without further data (PCVD); distr. Evritania, Timfristos Mts., Karpeníssi env., firry wood, 1446 m (38 ° 55 ' 25.22 " N, 21 ° 48 ' 33.22 " E), 13. VI. 2012, 4 ♂♂, 14 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♂ (PCRK); dtto, 8 ♂♂, 28 ♀♀, V. Dušánek leg. (PCVD); dtto, 3 ♂♂, 11 ♀♀, P. Brůha leg. (PCPB); dtto, 4 ♂♂, 8 ♀♀, B. Zbuzek leg. (PCBZ); distr. Evritania, Timfristos Mts., Karpeníssi env., firry wood, 1446 m (38 ° 55 ' 25.22 " N, 21 ° 48 ' 33.22 " E), 8. VI. 2015, 8 ♂♂, 9 ♀♀, J. Mertlik leg. (PCJM); dtto, 4 ♂♂, 7 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 2 ♂♂, 14 ♀♀, P. Brůha leg. (PCPB); dtto, 6 ♂♂, 1 ♀, T. Németh leg. (PCRK); dtto, 1 ♂ (HNHM); dtto, 9. VI. 2015, 1 ♀, J. Mertlik leg (PCJM).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFABFFFEC8C6FECD3898A990.taxon	diagnosis	Diagnosis. Dima evritaniensis is a medium-sized species (11.3 – 13.5 mm) with dense pronotal punctuation, semi-erect setae on the pronotal lateral margin, and elytra with short, dense, decumbent pubescence (Figs 26 – 28, 126 – 127). This species is morphologically very like D. fthiotidensis, based on the body shape, size, and matt pronotum with dense, fine punctation (Figs 35 – 41). Dima fthiotidensis has slightly coarser punctation of pronotum (Figs 130 – 134), shorter hairs on pronotum and elytra, decumbent to semi-erect pubescence on the pronotal sides (in most cases), and shorter apical lobe of paramera (Figs 186 – 187). Dima evritaniensis differs from the remaining species in the central Greece by its matt body surface and pronotum with denser, finer punctation (Figs 26 – 28, 126 – 127). Additionally, D. pelionensis sp. nov. has the sides of pronotum with decumbent setae in the anterior two thirds (semi-erect setae along its whole length in D. evritaniensis; Figs 126 – 127, 152 – 153), D. etoliensis has the scutellar frontal margin flattened, declined gradually to the plane of elytra (convex, steeply declined in D. evritaniensis; visible from the lateral view), D. hladilorum has more robust apical lobe of paramera (Fig. 190), broader scutellum, which is less convex basally (in most cases; scutellum longer and more convex basally in D. evritaniensis), and D. zbuzeki sp. nov. is a smaller species with much longer apical parameral lobe (Fig. 221). Intraspecific variability. In some cases, elytra or whole body is paler than in a typical form.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFABFFFEC8C6FECD3898A990.taxon	distribution	Distribution. Greece (Evritania: Panetoliko Mts., Timfristos Mts.; Fig. 245).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFABFFFEC8C6FECD3898A990.taxon	discussion	Remarks. We did not have an opportunity to study the holotype of D. evritaniensis, which is deposited in the Natural History Museum in Torino (MRSN). The collection is closed and not available for study (M. Garsena, pers. comm.). We studied the female paratype with the same locality data as the holotype (i. e. Panetoliko Mts., Proussos env.; from MCSB; Fig. 26) and this is conspecific with the specimens from Timfristos Mts. (Figs 27 – 28). Based on the comparison of the high-resolution holotype male photograph (kindly provided us by R. Schimmel) and fresh male specimens collected in Timfristos Mts. in 2012 and 2015, these populations share similar body size and shape, pronotal punctation and pubescence, and length of antennae. According to the original description of D. evritaniensis (Schimmel & Platia 2008), the holotype has short antennomeres II and III, with antennomere III slightly longer than II, and both combined as long as antennomere IV. However, all specimens which we examined here (including the paratype) have antennomere II slightly elongate, subtriangular, narrower than antennomere III, antennomeres II and III of subequal length, both combined longer than antennomere IV. Additionally, the parameral apical lobes of D. evritaniensis figured in the original description (Schimmel & Platia 2 008) are shorter than in the specimens from Timfristos, however, it might be caused by the different angle and / or the low-quality photograph. Therefore, the real shape of paramera might differ from the one originally figured (compare e. g. the shape of paramera of D. assingi holotype in the original description and herein; Schimmel & Platia (2008); Fig. 175).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA8FFFCC8C6FF3039A4AF90.taxon	materials_examined	Type material. Holotype: MACEDONIA: distr. Tetovo, Šar planina, Shipkovica env., beechwood, 1610 m (42 ° 1 ' 35.68 " N, 20 ° 53 ' 52.41 " E), 7. VI. 2013, ♂, J. Mertlik leg. (PCJM). Paratypes: MACEDONIA: without locality, 22. VII. 1999, 1 ♀, pitfall, no further data, ex coll. Penev (PCJM); distr. Tetovo, cca 1500 m, Popova Šapka env., 8. – 10. VII. 2010, 1 ♂, M. Fiala leg. (PCJM); Šar planina Mts., Gajre env., 1190 m, beechwood (42 ° 0 ' 58.02 " N, 20 ° 55 ' 28.91 " E), 26. V. 2013, 10 ♂♂, 8 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♂, 2 ♀♀ (PCRK); dtto, 13 ♂♂, 16 ♀♀, V. Dušánek leg. (PCVD); dtto, 5 ♂♂, P. Brůha leg. (PCPB); distr. Tetovo, Šar planina Mts., Gajre env., beechwood, 1190 m (42 ° 0 ' 58.02 " N, 20 ° 55 ' 28.91 " E), 7. VI. 2013, 4 ♂♂, 3 ♀♀, J. Mertlik leg. (PCJM); dtto, 9 ♂♂, 12 ♀♀, V. Dušánek leg. (PCVD); dtto, 2 ♂♂, 3 ♀♀, P. Brůha leg. (PCPB); distr. Tetovo, Šar planina, Shipkovica env., beechwood, 1610 m (42 ° 1 ' 35.68 " N, 20 ° 53 ' 52.41 " E), 7. VI. 2013, 3 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♀, V. Dušánek leg. (PCVD); dtto, 2 ♂♂, P. Brůha leg. (PCPB); distr. Tetovo, Šar planina Mts., Gajre env., 1190 m, beechwood (42 ° 0 ' 58.02 " N, 20 ° 55 ' 28.91 " E), 7. VI. 2014, 5 ♂♂, 4 ♀♀, V. Dušánek leg. (PCVD); Mavrovo env., 16. VI. 2011, 1 ♀, under stones, P. Hlaváč leg. (PCJM); distr. Tetovo, NP Mavrovo, Lazaropole env., 8. VI. 2012, 1 ♂, V. Zieris leg. (PCJM); Tetovo, Mt. Šar planina, 3 km NE of Popova Šapka [Šap], 2 ♂♂, 1 ♀, Richard Škoda leg. (PCPB); Tetovo reg., Tetovo, Popova Sapka, Fagus forest, singled at night, 42 ° 1 ' 13.00 " N, 20 ° 53 ' 52.54 " E, 7. VI. 2014., 1 ♂, 1 ♀, leg. V. Dušánek, R. Kundrata & T. Németh (HNHM); dtto, 9 ♂♂, 10 ♀♀ (PCRK).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA8FFFCC8C6FF3039A4AF90.taxon	diagnosis	Diagnosis. Dima fialai sp. nov. belongs to the group of species with more convex, suboval-shaped body and shiny elytral intervals with very sparsely distributed thin setae (Figs 29 – 30). It differs from D. macedonica, D. florinensis and D. dusaneki sp. nov. by the pubescence of the pronotal sides, which is short and almost decumbent at anterior 3 / 4 of the margin length, with longer semi-erect setae only posteriorly near hind angles (Fig. 128). Additionally, D. dusaneki sp. nov. differs from D. fialai sp. nov. by the more robust body, shape of pronotum (not so narrowed toward anterior margin, with evenly rounded sides in D. dusaneki sp. nov.; Figs 20 – 21) and slightly denser pronotal punctation (Fig. 123). Dima vailatii Schimmel & Platia, 2008, which has a similar type of pubescence at the pronotal sides, differs from D. fialai sp. nov. in different shape of pronotum (widest medially; Figs 109 – 113), matt elytra with denser and longer pubescence, and larger subapical tooth at apex of paramera (Figs 217 – 219). Dima elateroides, which occurs north of D. fialai sp. nov., has a similar shape of pronotum, but differs apparently in the smooth body surface, longer antennomeres II and III, much sparser pronotal punctation, wider, basally emarginate scutellum, and narrower apical lobe of paramera (Figs 22 – 24, 182).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA8FFFCC8C6FF3039A4AF90.taxon	description	Description. Holotype, male. Body medium-sized, 10.9 mm long, 4.6 mm wide, shiny, covered with short dense pubescence on pronotum and very sparse on elytra. Body blackish-brown, with paler, reddish-brown legs and antennae, elytral margins and suture, and sublateral bends posterad of elytra; pubescence yellowish (Fig. 29). Head including eyes 0.6 times as wide as pronotum, frons very slightly depressed; punctation sparse and coarse, denser apically, intervals between punctures shiny; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina short, indistinct. Antennae moderately long, surpassing the posterior angles of pronotum by more than three segments; antennomeres II and III subtriangular, about 1.5 times longer than wide, subequal, together 1.3 times longer than antennomere IV, length ratio of antennomeres II – IV 1.0: 1.0: 1.5, antennomeres IV to X gradually widened apically, about 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semi-erect setae. Pronotum 1.4 times wider than long, widest at two thirds, moderately convex dorsally; sides convex, moderately gradually narrowed toward frontal margin; anterior angles obtuse, posterior angles short, prominent, sharp, only slightly produced outward; punctation sparse, coarse, punctures on disc equally distributed, intervals flat, shiny. Pubescence on pronotal disc moderately long, decumbent, that on pronotal sides decumbent at anterior 3 / 4, semi-erect posteriorly, with longer setae at posterior angles (Fig. 128). Prosternum finely, moderately sparsely punctate, with short semi-erect setae; prosternal lobe moderately coarsely punctate, with longer semi-erect pubescence. Scutellum sub-pentagonal, slightly convex, in lateral view in the plane of elytra, anterior part slightly raised, widest at two thirds; frontal margin straight, gradually declined; apex rounded, punctation fine, dense; pubescence fine, moderately long, semi-erect. Elytra suboval, 3.1 times longer than pronotum, convex, widest at two thirds; striae well developed along the whole elytral length, some irregularly interrupted by coarse sparse punctures; surface of interstices flattened, shiny, densely punctate; punctures shallow, equally distributed, of the same size; pubescence very short, sparsely distributed. Aedeagus elongate; paramera long, narrow, its apical lobe robust, short, subapical tooth short, obtuse (Fig. 184). Female (Fig. 30). Like male, but more robust, with shorter antennae (surpassing the posterior angles of pronotum about 1.5 segments), pronotum distinctly wider than in male (1.6 times wider than long), scutellum wider, with frontal margin rounded. The sclerotized spines of bursa copulatrix as in Fig. 227. Intraspecific variability. Body length: 10.7 – 12.6 mm. As in many other Dima spp., some specimens have denser pronotal punctation with intervals wrinkled, matt.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA8FFFCC8C6FF3039A4AF90.taxon	distribution	Distribution. Macedonia (Šar planina: Tetovo; Mavrovo; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA8FFFCC8C6FF3039A4AF90.taxon	etymology	Etymology. The specific epithet is a patronym in honor of Mr. Martin Fiala (Trutnov, Czech Republic), who collected one of the paratypes.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA9FFE3C8C6FDE039D8AC58.taxon	materials_examined	Type depository. Holotype, ♀ (PCGP). Type locality. Greece: Flórina, Varnous Mts., A. Triados (40 ° 47 ′ 32.1 '' N, 21 ° 13 ′ 35.4 '' E), 1285 m. New material. GREECE: distr. Flórina, Varnoús Mts., Pisoderi, Agia Triada env., 1410 m (40 ° 47 ' 32.1 " N, 21 ° 13 ' 35.4 " E), 27. V. 2013, 2 ♂♂, J. Mertlik et V. Dušánek leg (PCJM, PCVD); distr. Flórina, Varnoús Mts., Vigla pass, beechwood, 1565 m (40 ° 46 ' 20.80 " N, 21 ° 15 ' 55.25 " E), 27. V. 2013, 4 ♂♂, 4 ♀♀, J. Mertlik leg. (PCJM); dtto, 3 ♂♂, 6 ♀♀, V. Dušánek leg. (PCVD); dtto, 4 ♂♂, 2 ♀♀, P. Brůha leg. (PCPB); Makedonia, pref. Florina, Mt. Vernon, 2 km W Alona, beech forest, 40 ° 46 ' 52.97 " N, 21 ° 16 ' 33.12 " E, 1400 m, singled at night, 24. VI. 2012, 2 ♂♂, 5 ♀♀, leg. A. Kotán, A. Márkus, P. Nemes & T. Németh (PCRK); dtto, 1 ♂, 1 ♀ (HNHM); dtto, 18. V. 2013, 2 ♀♀, leg. A. Kotán, P. Nemes, Gy. Németh, T. Németh & V. Szénási (HNHM); dtto, 2 ♂♂, 5 ♀♀ (PCRK); prov. Makedonia, distr. Florina, Mt. Vernon, 2 km W Pisoderi ski center, 40 ° 47 ' 20.16 " N, 21 ° 13 ' 29.51 " E, 1330 m, beech forest, singled at night, 18. V. 2013, 2 ♂♂, 2 ♀♀, leg. Kotán, P. Nemes, Gy. Németh, T. Németh & V. Szénási (HNHM); dtto, 3 ♂♂, 3 ♀♀ (PCRK); MACEDONIA: Galičica, northern part [sev. str.], IX. 1926, 1 ♀, Dr. Rambousek leg. (NMPC); Baba planina, Niže Pole – Golemo jezero, 1500 – 2000 m, 18. VII. 2006, 1 ♀, V. Zieris leg. (PCJM); dtto, 14. – 15. VII. 2010, 1 ♀, V. Zieris leg. (PCTS); Baba Mts., Malovište, Šemnica valley, 1600 – 1800 m, 11. – 12. VI. 2008, 1 ♂, L. Blažej leg. (PCPB); distr. Bitola, Baba plateau, 1.7 km S of Nižepole, beechwood, 1400 m (40 ° 58 ' 47.54 " N, 21 ° 14 ' 42.92 " E), 26. V. 2014, 3 ♂♂, 1 ♀, J. Mertlik leg. (PCJM); dtto, 2 ♂♂, 3 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 3 ♂♂, 1 ♀, P. Brůha leg. (PCPB); distr. Bitola, Baba plateau, Nižepole – ski centre env., beechwood, 1375 m (40 ° 58 ' 52.22 " N, 21 ° 15 ' 13.82 " E), 26. V. 2014, 1 ♂, 2 ♀♀, J. Mertlik leg. (PCJM); dtto, 8 ♂♂, 2 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 3 ♂♂, P. Brůha leg. (PCPB); Nizhepole reg., NP Pelister, 2.5 km S Nizhepole, mixed forest, swept and singled at night, 40 ° 59 ' 3.65 " N, 21 ° 14 ' 37.24 " E, 6. VI. 2014, 2 ♂♂, 4 ♀♀, leg. V. Dušánek, R. Kundrata & T. Németh (HNHM); dtto, 27 ♂♂, 11 ♀♀ (PCRK); dtto, 18 ♂♂, 15 ♀♀, V. Dušánek leg. (PCVD); Nizhepole reg., NP Pelister, 3 km S Nizhepole, 40 ° 58 ' 52.5 " N, 21 ° 15 ' 16.2 " E, 6. VI. 2014, 1 ♂, leg. V. Dušánek, R. Kundrata & T. Németh (PCRK); Pelagonia region, Pelister Mts., Nižepole, forest brook below the ski station, N 40 ° 58.889 ', E 21 ° 15.246 ', 1370 m, 25. VI. 2014, 1 ♂, leg. Tibor Kovács (MMGY).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA9FFE3C8C6FDE039D8AC58.taxon	description	Description of male. Like female, but with relatively longer antennae, surpassing the hind angles of pronotum by about three antennomeres (about one in females), and pronotum slightly narrower than in females (Fig. 32). Aedeagus as in Fig. 185.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA9FFE3C8C6FDE039D8AC58.taxon	diagnosis	Diagnosis. This species belongs to the group of species with a more convex, suboval-shaped body and shiny elytral intervals with very sparsely distributed thin setae (Figs 31 – 34). Dima florinensis is a medium-sized species (body length: 11.0 – 13.0 mm) with long, almost tangled, semi-erect pubescence on the head and pronotal sides (typical for this species; Fig. 129), short antennae and by the distinct double punctation of pronotum (Fig. 129). It is very like D. macedonica by the habitus, double punctation of pronotum, and shape and length of antennae. These species differ in the scutellum (D. florinensis has a scutellum on the same plane as the elytra in lateral view; D. macedonica has a scutellum raised above the plane of the elytra; Fig. 164) and pubescence of the pronotal sides (D. macedonica has pubescence in anterior third of the pronotal sides about 0.5 times as long as in the posterior half; Figs 141 – 143). D. raineri differs from D. florinensis by the decumbent pubescence in the first half of pronotal sides and double punctation of pronotum being less distinct, with punctures more similar in size to each other (Figs 154 – 155), and D. vailatii by the denser pubescence of the elytra and the shorter pubescence of pronotal sides (Figs 109 – 113, 158 – 160). The sclerotized spines of bursa copulatrix as in Fig. 228. Intraspecific variability. There is an intraspecific variability in the length of the apical lobe of paramera in D. florinensis (similarly as in D. macedonica). The holotype is darker than remaining specimens, which are reddishbrown (Figs 31 – 34). There is a variability in the punctation density on the pronotum; most specimens have densely punctate pronotum, but some specimens from Nizhepole region have sparser punctures. The female collected in the northern slope of Galičica is paler and has much sparser pronotal punctation than the remaining specimens of this species (Fig. 34).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFA9FFE3C8C6FDE039D8AC58.taxon	distribution	Distribution. Greece (Varnous Mts., Mt. Verno), Macedonia (Baba Mts., Galičica Mts.) (Fig. 244). First record for Macedonia.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB6FFE2C8C6FD283D0BAFED.taxon	materials_examined	Type depositories. Holotype of D. fthiotidensis, ♂ (MRSN); 7 paratypes of D. fthiotidensis, ♀♀ (MCSB, MRSN, PCGP, PCRS); holotype of D. fokidensis, ♀ (MCSB); holotype of D. scutellaris, ♀ (PCGP). Type locality. Greece: Phthiotis, Kallidromo Mt., 990 m (D. fthiotidensis); Greece: Fokida province, O. Iti, Pira, 1150 m (D. fokidensis); Greece: Phocis, Giona Mts., vers. N str. Kaloskopi Stromi, 1170 m (D. scutellaris). New material. GREECE: distr. Fthiótida, Vardoússia Mts., road Plátanes – Gramméni Oxiá, beechwood, 1447 m (38 ° 46 ' 57.47 " N, 21 ° 58 ' 06.86 " E), 14. VI. 2012, 12 ♂♂, 11 ♀♀ J. Mertlik leg. (PCJM); dtto, 1 ♀ (PCRK); dtto, 32 ♂♂, 32 ♀♀, V. Dušánek leg. (PCVD); dtto, 9 ♂♂, 12 ♀♀, P. Brůha leg. (PCPB); dtto, 18 ♂♂, 26 ♀♀, B. Zbuzek leg. (PCBZ); dtto 11 ♂♂, 8 ♀♀, M. Samek leg. (PCMS); Fthiotis env., Mt. Kallidromo, 3 km SW Pavliani, oakpine forest, singled at night, 23. V. 2011, 2 ♂♂, 2 ♀♀, leg. A. Kotán, A. Márkus & T. Németh (PCRK); dtto, 1 ♀ (HNHM); Ftiotida, Iti Mts., Kastania, 1500 m, 4. VI. 2006, 1 ♂, 1 ♀, Pavel Štěpánek leg. (PCJM).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB6FFE2C8C6FD283D0BAFED.taxon	diagnosis	Diagnosis. Dima fthiotidensis is a medium-sized species (body length: 10.5 – 14.5 mm) with matt, densely punctate pronotum (Figs 130 – 134), more or less decumbent pubescence on the pronotal lateral margin, moderately densely pubescent pronotum and elytra, and the tongue-like, convex scutellum with steeply declined frontal margin (Fig. 166). This species is similar in the above mentioned characters to D. evritaniensis, however the latter differs in the slightly finer punctation of pronotum, longer hairs on pronotum and elytra, semi-erect pubescence on the pronotal sides (Figs 26 – 28, 126 – 127), and longer apical lobe of paramera (Fig. 183). The remaining species occurring in the surrounding mountains have a shiny pronotum with sparser punctation. Additionally, D. pelionensis sp. nov. and D. etoliensis have a flattened scutellum, and D. hladilorum and D. zbuzeki sp. nov. have a more robust and longer apical lobe of paramera, respectively (Figs 190, 221). The sclerotized spines of bursa copulatrix as in Fig. 229. Intraspecific variability. The specimens from Mt. Iti have a darker body. Several specimens (including the holotype of D. scutellaris) have slightly wrinkled to rugose pronotum with denser punctation (Fig. 132). The scutellum is more or less convex, in some cases with a longitudinal shiny keel running from the base towards apex. Most specimens have pubescence near anterior pronotal angles oriented latero-posteriorly, but a female from Mt. Iti has pubescence oriented medio-posteriorly.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB6FFE2C8C6FD283D0BAFED.taxon	distribution	Distribution. Greece (Mt. Kallidromo, Mt. Iti, Vardousia Mts., Giona Mts., Oxia Mts.; Fig. 245).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB6FFE2C8C6FD283D0BAFED.taxon	discussion	Remark. We were not able to study the holotype of D. fthiotidensis (deposited in Torino, Italy), however we examined four paratypes, recently collected specimens from the type locality (Mt. Kallidromo; Fig. 38), and the specimens from the surrounding mountain ranges to investigate the limits of this species (Figs 39 – 41). Platia (2010) described D. scutellaris from a single female from Mt. Giona, with a raised scutellum and wrinkled pronotum with denser punctation. This specimen, however, fits the definiton of D. fthiotidensis in all aspects, and what is more, we found similarly wrinkled (teratological?) female specimens in the Vardousia Mts. (see the similar cases in D. assingi / D. peloponnensis and D. fialai). Therefore, we herein synonymize Dima scutellaris Platia, 2010 under Dima fthiotidensis Schimmel & Platia, 2008. Schimmel & Platia (2008) described D. fokidensis from three female specimens. The holotype comes from Mt. Iti and paratypes from Mt. Pelion, which is located more than 80 km away from the holotype locality. This holotype was found to be conspecific with D. fthiotidensis (described in the same paper as D. fokidensis; Fig. 36) and both paratypes represent a new species, described here as D. pelionensis sp. nov. (Fig. 95). Therefore, we herein synonymize Dima fokidensis Schimmel & Platia, 2 0 0 8 under Dima fthiotidensis Schimmel & Platia, 2008.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB7FFE2C8C6FE7B3BA8AA13.taxon	materials_examined	Type depositories. Holotype, ♂ (MRSN), 3 paratypes, ♀♀ (MCSB, PCGP, PCRS). Type locality. Greece: Évia prov., Dirfi str., south of Dirfios – A., Irini, 1080 m. New material. GREECE: distr. Évia, Dirfys Mts., Mt. Xerovouni env., 1020 m, firrywood (38 ° 35 ' 32.50 " N, 23 ° 52 ' 32.79 " E), 29. V. 2013, 3 ♀♀, V. Dušánek leg. (PCVD, PCJM, PCPB); dtto, 29. V. 2014, 2 ♂♂, J. Mertlik leg. (PCJM); dtto, 4 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 1 ♂, 3 ♀♀, P. Brůha leg. (PCPB).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB7FFE2C8C6FE7B3BA8AA13.taxon	diagnosis	Diagnosis. Dima giachinoi is a medium-sized species (body length: 11.3 – 14.2 mm) easily recognizable by the relatively short antennae (Figs 42 – 44), shiny pronotum with moderately large sparse punctures on disc and denser on margins (Fig. 135), flattened scutellum and long, semi-erect body pubescence. The sclerotized spines of bursa copulatrix as in Fig. 230.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB7FFE2C8C6FE7B3BA8AA13.taxon	distribution	Distribution. Greece (Euboea: Dirphys Mts.; Fig. 245).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB7FFE2C8C6FE7B3BA8AA13.taxon	discussion	Remark. We did not have an opportunity to study the holotype of D. giachinoi (deposited in the MRSN, Torino), but we studied all paratypes, which have the same locality data as the holotype. Due to the short antennae and type of body pubescence, it is possible that this species is conspecific with D. isabellae, which was described only from a single female specimen and occurs in the same area (Fig. 245).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB7FFE1C8C6FB6E39FBAE9D.taxon	materials_examined	Type depository. Holotype, ♀ (PCGP). Type locality. Greece: nom. Lakonia, Taigetos Mts., Profitis Ilias [Prof. Elias], 1940 m.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB7FFE1C8C6FB6E39FBAE9D.taxon	diagnosis	Diagnosis. Dima hirtipennis is an easily recognizable species by its very long semi-erect to erect pubescence all over the body (Figs 45, 136, 167). The similar pubescence is known only in D. olympica Meschnigg, 1934 from Mt. Olympus, but that species has two types of hairs on the elytra (long erect and short decumbent; Fig. 169). Several Dima species from the Peloponnese have also erect pubescence on the pronotum, but shorter (Figs 120, 146, 157). Additionally, they have decumbent to semi-erect short pubescence on the elytra (Figs 6 – 9, 67 – 74, 105 – 108). Dima hirtipennis has tiny antennae with long antennal segments (more than 2 times longer than wide in antennomeres IV – X), and the sclerites of bursa copulatrix which are distinctly thinner than in other known Dima species (Fig. 231).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB7FFE1C8C6FB6E39FBAE9D.taxon	distribution	Distribution. Greece (Peloponnese: Taygetos Mts.; Fig. 246).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB7FFE1C8C6FB6E39FBAE9D.taxon	discussion	Remark. This species is known only from the female holotype (Fig. 45). There is a single male specimen collected in the Mani Peninsula, Peloponnese, with the following data: " Peloponnese [Peloponnesus], Mani peninsula, nr. Langada village, 350 m, 16. II. 1991, ♂, G. Kakiopoulos leg. " (PCGK) (Fig. 45; male genitalia as in Fig. 189). This specimen resembles D. hirtipennis by its long, semi-erect to erect body vestiture and elongate antennomeres IV – X (Figs 45 – 46), but can be distinguished by having a darker body (light brown in D. hirtipennis), different shape of pronotum, with slightly longer pubescence on the pronotal sides, and straight frontal margin of scutellum (distinctly emarginate in D. hirtipennis). Since we have only two specimens from different sexes available for the study, we cannot assess what is the the sexual dimorphism and what is the interspecific difference. Therefore, it seems the best approach is to postpone any taxonomic decision until more material from the southern Peloponnese is available.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB4FFE1C8C6FEEB3AA5A845.taxon	materials_examined	Type depositories. Holotype, ♂ (PCRS), allotype, ♀ (PCJH), 4 paratypes: 2 ♂♂, 2 ♀♀ (PCRS, PCGP, PCJM, PCJH). Type locality. Greece, Parnassos Mt., 1700 m. Redetermined material. Three paratypes of D. hladilorum from Macedonia (Galičica Mts.; Schimmel 1987) belong to D. macedonica (described later than D. hladilorum; Schimmel 1993) (Fig. 61). Some specimens recorded under D. hladilorum in Schimmel & Platia (2008) were redetermined as D. arndti (Taygetos Mts., Crete; 4 ex.), D. assingi (Kilini Mts.; 1 ex.), D. pelionensis sp. nov. (Mt. Pilio; 2 ex.), and D. vailatii (Kajmakčalan Mts.; 2 ex.). New material. Greece: distr. Viotía, Parnassos Mts., Ski center env., firrywood, 1700 m (38 ° 32 ' 54.89 " N, 22 ° 34 ' 25.84 " E), 28. V. 2013, 9 ♂♂, 10 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♂, 2 ♀♀ (PCRK); dtto, 6 ♂♂, 8 ♀♀, V. Dušánek leg. (PCVD); dtto, 6 ♂♂, 8 ♀♀, P. Brůha leg. (PCPB); distr. Viotía, Parnassos Mts., Ski center env., firrywood, 1700 m (38 ° 32 ' 54.89 " N, 22 ° 34 ' 25.84 " E), 30. V. 2014, 41 ♂♂, 15 ♀♀, J. Mertlik leg. (PCJM); dtto, 32 ♂♂, 10 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 12 ♂♂, 14 ♀♀, P. Brůha leg. (PCPB); prov. Central Gr., distr. Phthiotis, Mt. Parnassos, 4 km NE Kalivia Arachova, spruce forest, 38 ° 32 ' 22.65 " N, 22 ° 34 ' 12.45 " E, 1700 m, singled at night, 25. – 26. V. 2015, leg. A. Kotán, Gy. Németh, B. Szelenczey & V. Szénási, 9 ♂♂, 6 ♀♀ (PCBS); dtto, 10 ♂♂, 3 ♀♀ (PCRK); dtto, 2 ♂♂, 2 ♀♀ (HNHM).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB4FFE1C8C6FEEB3AA5A845.taxon	diagnosis	Diagnosis. Dima hladilorum is a smaller species (body length: 9.4 – 10.0 mm; Figs 47 – 49) with pronotum with sparse punctuation and shiny intervals, pubescence of the pronotal sides semi-erect along its whole length (Fig. 137), scutellum convex with its frontal margin steeply declined, and paramera robust, elongate (Fig. 190). This species is like D. zbuzeki sp. nov. from Agrafa Mts., however it differs from the latter by the less distinct pubescence on the posterior angles of pronotum, convex scutellum with medially emarginate frontal margin, elytral striae often indistinct or missing in the apical part, and by the shape of the apical lobe of paramera (Figs 47 - 49, 116 – 118, 137, 162, 190, 221). Dima fthiotidensis and D. evritaniensis differ from D. hladilorum in the matt, densely punctate pronotum (Figs 26 – 28, 35 – 41) and smaller apical lobe of paramera (Figs 183, 186 – 187); D. etoliensis by the shape of antennae (narrower, with longer antennomeres), flattened scutellum with gradually declined frontal margin, larger body with more parallel-sided elytra, and the distinct striae along the whole length of elytra (Fig. 25). Dima pelionensis sp. nov. has a flattened scutellum, pubescence of the pronotal sides decumbent in apical two thirds (Figs 152 – 153), and the apical lobe of paramera shorter (Fig. 209). Intraspecific variability. Some specimens have more or less indistinct elytral striae.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB4FFE1C8C6FEEB3AA5A845.taxon	distribution	Distribution. Greece (Parnassos Mts.; Fig. 245).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB4FFE0C8C6F9D33A78AC7D.taxon	materials_examined	Type depository. Holotype, ♀ (MHNP?). Type locality. Greece: Euboea, Mt. Dirphys [Ile d'Eubée, mont Dirphys vers 1400 m].	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB4FFE0C8C6F9D33A78AC7D.taxon	diagnosis	Diagnosis. Dima isabellae is known only from the female holotype, which is not available for study (Schimmel 1996, pers. comm.). Therefore, we must rely only on the original description (Dajoz 1973). Dima isabellae is a medium-sized species (body length: 12.5 mm) with relatively short antennae, moderately long pubescence, and dense punctation on the pronotum (Dajoz 1973). This species occurs in Dirphys Mts. together with D. giachinoi, which has also short antennae and dense pubescence of pronotum (Figs 42 – 44, 135). It is possible, that D. isabellae was described from the specimen with denser, non-typical pronotal punctation (see similar situations in D. assingi / D. peloponnensis and D. fthiotidensis / D. scutellaris) and that the specimens determined later as D. giachinoi are conspecific with D. isabellae. However, until we find specimens with very dense pronotal punctation, as in D. isabellae, within a population of D. giachinoi, we refrain from making any taxonomic conclusions.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB4FFE0C8C6F9D33A78AC7D.taxon	distribution	Distribution. Greece (Euboea: Dirphys Mts.; Fig. 245).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB4FFE0C8C6F9D33A78AC7D.taxon	discussion	Remark. The type material has not been located. The material mentioned under D. isabellae by Schimmel & Platia (2008) was redetermined and belongs to D. arndti (Taygetos Mts.; 4 ex.), D. assingi (Kilini Mts.; 1 ex.) and D. parnonensis (Parnon Mts.; 1 ex.).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB5FFE7C8C6FDCB3DD5AD5D.taxon	materials_examined	Type material. Holotype: MACEDONIA: distr. Kavadarci, Kožuf Mts., Mt. Keči Kaj, beechwood, 1600 – 1630 m (41 ° 11 ' 55.25 " N, 22 ° 14 ' 26.58 " E), 9. – 10. VI. 2012, ♂, J. Mertlik leg. (PCJM). Paratypes: MACEDONIA: distr. Kavadarci, Kožuf Mts., 1800 m, Smrdliva Voda, 13. VI. 2011, 1 ♀, Vladimír Skoupý leg. (PCJM); distr. Kavadarci, Kožuf Mts., Mt. Keči Kaj, beechwood, 1600 – 1630 m (41 ° 11 ' 55.25 " N, 22 ° 14 ' 26.58 " E), 9. – 10. VI. 2012, 6 ♂♂, 12 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♀ (PCRK); dtto, 7 ♂♂, 4 ♀♀, V. Dušánek leg. (PCVD); dtto, 2 ♂♂, 2 ♀♀, P. Brůha leg. (PCPB); dtto, 8 ♂♂, 2 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 16. VI. 2012, 2 ♂♂, 2 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 2 ♂♂, P. Brůha leg. (PCPB); dtto, (41 ° 12 ' 13 " N, 22 ° 14 ' 33 " E), 16. VI. 2012, 3 ♂♂, 1 ♀, P. Brůha leg. (PCPB); Kožuf Mts., Smrdliva Voda, 1800 m, 3 ♂♂, Pavel Štěpánek leg. (PCJM); Jugoslavia, Kožuf Mts., Mt. Čiči Kaja, 12. VII. 1990, 1 ♀, leg. I. Karaman (HNHM). Other material examined. GREECE: Kožuf Mts., Vorino env., 19. VI. 2003, 1 ♀, Vladimír Skoupý leg. (as a paratype of D. lebenbaueri in Schimmel & Platia 2008; Fig. 52) (PCJM).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB5FFE7C8C6FDCB3DD5AD5D.taxon	diagnosis	Diagnosis. Dima kozufensis sp. nov. is like D. lebenbaueri (especially to its smaller form from the higher elevations; Figs 55 – 56) but it can be distinguished by the pubescence of the sides of pronotum (slightly shorter, in anterior third or half more decumbent in D. kozufensis sp. nov.; longer, semi-erect along the whole margin in D. lebenbaueri; Figs 138 – 140) and the shape of the scutellum (heart-shaped, basally emarginate in D. kozufensis sp. nov.; basally straight or rounded in D. lebenbaueri). The apical lobe of paramera is in most cases robust and short as in Fig. 191, but we found a specimen with longer parameral lobe, broadly like those in D. lebenbaueri (Figs 192 – 193). Dima vailatii from Kaimakchalan (Fig. 244) differs from D. kozufensis sp. nov. by the denser pronotal punctation and short, curved, decumbent pubescence of pronotal sides (Figs 158 – 159). Additionally, Dima florinensis has long semi-erect setae along the whole length of the pronotal sides (Figs 31 – 34).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB5FFE7C8C6FDCB3DD5AD5D.taxon	description	Description. Holotype, male. Body medium-sized, 10.0 mm long, 4.3 mm wide, with moderately shiny pronotum and matt elytra covered with short dense pubescence. Body brown, with head and pronotum darker, blackish-brown; elytral and pronotal margins and elytral sutura paler; pubescence yellowish (Fig. 50). Head including eyes 0.6 times as wide as pronotum, frons slightly depressed in the middle; punctation dense and coarse, intervals between punctures shiny; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina weakly defined, short. Antennae longer, surpassing the posterior angles of pronotum of about four segments; antennomeres II and III short, subtriangular, more than 1.5 times longer than wide, subequal, together about 1.3 times longer than antennomere IV, length ratio of antennomeres II – IV 1.0: 1.0: 1.5, antennomeres IV to X gradually widened apically, less than 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semi-erect setae. Pronotum 1.4 times wider than long, widest just behind middle, moderately convex dorsally; sides convex, gradually narrowed towards anterior margin, more steeply narrowed towards posterior margin; straight in lateral view; anterior angles obtuse; posterior angles prominent, sharp, slightly produced outward; punctation sparse, coarse, moderately deep, punctures on disc equally distributed; of about the same size; intervals between punctures flattened, moderately shiny. Pubescence on pronotal disc moderately long, decumbent to semi-erect; that on pronotal sides decumbent to semi-erect at anterior half and semi-erect at posterior half, with long semi-erect to erect setae at posterior angles (Fig. 138). Prosternum finely, sparsely punctate, with short decumbent to semi-erect setae; prosternal lobe moderately coarsely punctate, with longer semi-erect pubescence. Scutellum heart-shaped, widest at posterior two thirds, slightly convex, in lateral view slightly above the plane of elytra, frontal margin emarginate medially, gradually declined; apex narrowed, rounded, punctation shallow, coarse, sparse; pubescence sparse, fine, moderately long, decumbent to semi-erect. Elytra sub-parallel, 3.2 times longer than pronotum, moderately convex, widest at middle; striae shallow, developed along the whole elytral length; surface of interstices flattened, matt, moderately densely and finely punctate; punctures suboval, equally distributed, of the same size; pubescence short, dense. Aedeagus elongate; paramera long, narrow, its apical lobe robust, short, subapical tooth short (Fig. 191). Female (Figs 51 – 52). Like male, but suboval, with shorter antennae (surpassing the posterior angles of pronotum about more than two segments), wider pronotum (1.5 times wider than long), widest at middle, with evenly convex sides. The sclerotized spines of bursa copulatrix as in Fig. 232. Intraspecific variability. Body length: 10.0 – 12.2 mm. A single specimen from the Greek part of Kožuf Mts. (paratype of D. lebenbaueri; Fig. 52) is paler (not bicolored as specimens from Macedonia), has more elongate body shape (2.6 times longer than wide; specimens from Macedonia 2.2 – 2.3 times longer than wide) and pubescence of pronotal sides with less semi-erect setae in posterior half. More material from Greece is needed for further conclusions.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB5FFE7C8C6FDCB3DD5AD5D.taxon	distribution	Distribution. Macedonia (Kožuf Mts.), Greece (Kožuf Mts.) (Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB5FFE7C8C6FDCB3DD5AD5D.taxon	etymology	Etymology. The specific epithet refers to Kožuf Mts., Macedonia / Greece, the type locality of the new species.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB2FFE7C8C6FC2B3AF6A9D7.taxon	materials_examined	Type depositories. Holotype, ♂ (SMNS), 22 paratypes: 13 ♂♂, 9 ♀♀ (SMNS, PCJM, PCRS). Type locality. Greece: Pieria Mts., Elatohori, 1000 m. Redetermined material. One of the paratypes (Greece: Kozhuf Mts., more than 80 km far from the type locality; Fig. 52) differs from the specimens of D. lebenbaueri from Pieria Mts. in the pubescence of pronotal sides and shape of scutellum, and is transferred herein under D. kozufensis sp. nov. New material. GREECE: distr. Pieria, Pieria Mts., Elatohóri env., 1515 m, beechwood (40 ° 18 ' 48.19 " N, 22 ° 12 ' 23.83 " E), 5. VI. 2013, 2 ♂♂, 1 ♀, J. Mertlik leg. (PCJM); dtto, 1 ♂, 1 ♀, V. Dušánek leg. (PCVD), dtto, 1 ♂, 1 ♀, P. Brůha leg. (PCPB).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB2FFE7C8C6FC2B3AF6A9D7.taxon	diagnosis	Diagnosis. Dima lebenbaueri is a medium-sized to large species (body length: 10.0 – 14.0 mm), with pronotum with short straight pubescence, sides with straight semi-erect setae along the whole length, pronotal punctation fine, intervals moderately shiny, elytra with short, fine, more or less dense pubescence and interstices moderately shiny (Figs 53 – 56, 139 – 140). Dima lebenbaueri shares similar pubescence of pronotal sides and relatively long subapical parameral tooth with D. macedonica, however these species differ in the pronotal punctation (denser, double-sized punctures in D. macedonica; Figs 141 – 143), body surface (more shiny in D. macedonica; Figs 57 – 61) and pubescence of elytra (sparser in D. macedonica; Fig. 168). Dima olympica differs from D. lebenbaueri in the long erect pubescence on pronotum and two types of hairs on elytra (Figs 147, 169), D. vailatii differs in the short curved decumbent pubescence of pronotal sides and surface of elytra (Figs 109 – 113, 158 – 160), and D. kozufensis sp. nov. differs in the more or less decumbent pubescence in anterior part of pronotal sides and emarginate frontal part of scutellum (Fig. 138). Intraspecific variability. The specimens from the type series (1000 m a. s. l.) are larger, more robust (13.0 – 14.0 mm), have shorter antennae (surpassing hind angles of pronotum by about three antennomeres), and finely and sparsely pubescent elytra (Figs 53 – 54). The specimens from the higher elevations (1515 m a. s. l.) are smaller (10.0 – 11.5 mm), have longer antennae (surpassing hind angles of pronotum by about four antennomeres), and denser elytral pubescence (Figs 55 – 56).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB2FFE7C8C6FC2B3AF6A9D7.taxon	distribution	Distribution. Greece (Pieria Mts.; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB3FFE6C8C6FF303B2AA82F.taxon	materials_examined	Type depositories. Holotype, ♂ (PCPC), 2 paratypes, ♂♂ (PCPC, PCRS). Type locality. Macedonia, Galičica, 1600 m. [YU: Makedonija]. Redetermined material. A male paratype of D. macedonica from Albania: Mt. Tomorr (PCGP) was redetermined as D. raineri (Fig. 100). Some specimens recorded under D. macedonica in Schimmel & Platia (2008) were redetermined as D. elateroides (Croatia, Montenegro; 6 ex.) and D. raineri (Greece; 7 ex.). For a female specimen from Librazhd, Albania (Fig. 101), identified as D. macedonica by Schimmel & Platia (2008), see the remarks under D. raineri. Three paratypes of D. hladilorum from Macedonia (Galičica Mts) belong under D. macedonica (Fig. 61). New material. MACEDONIA: Galičica Mts., Livada pass, Mt. Magaro, 1650 – 1800 m, 9. – 10. VI. 2008, 1 ♀, P. Vonička leg. (PCJM); dtto, 5 ♂♂, 3 ♀♀, L. Blažej leg. (PCPB); NP Galičica – Livada, Trpejca env., 1401 m (40 ° 57 ' 48 " N, 20 ° 48 ' 49 " E), 2. – 3. VI. 2011, 1 ♂, D. Navrátil leg. (PCJM); distr. Ohrid, Galičica plateau, Trpejca env., beechwood, 1540 m (40 ° 57 ' 42.18 " N, 20 ° 48 ' 55.76 " E), 24. V. 2014, 4 ♂♂, 3 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♂, 1 ♀, B. Zbuzek leg. (PCBZ); dtto, 7 ♂♂, 1 ♀, P. Brůha leg. (PCPB); distr. Ohrid, Galičica plateau, beechwood, near pass, 1590 m (40 ° 57 ' 7.72 " N, 20 ° 49 ' 56.16 " E), 24. V. 2014, 28 ♂♂, 4 ♀♀, J. Mertlik leg. (PCJM); dtto, 19 ♂♂, B. Zbuzek leg. (PCBZ); dtto, 22 ♂♂, 2 ♀♀, P. Brůha leg. (PCPB); Ohrid, PN Galičica, 2 km S of Lako Signoj, 1940 m, 40 ° 56.285 ' N, 20 ° 49.407 ' E, 7. VI. 2015, 3 ♀♀, Richard Škoda leg. (PCPB); Pelagonia reg., 1.5 km SW Ehloec (41 ° 27 ' 3.88 " N, 20 ° 44 ' 30.19 " E), 6. VI. 2014, Fagus forest, swept & beaten, 3 ♂♂, leg. Dusanek, Kundrata & Németh (PCRK); dtto, 1 ♀ (HNHM); dtto, 4 ♂♂, 4 ♀♀, V. Dušánek leg. (PCVD).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB3FFE6C8C6FF303B2AA82F.taxon	diagnosis	Diagnosis. Dima macedonica is a medium-sized species (body length: 9.8 – 13.2 mm) which belongs to the group of species with more convex, suboval-shaped body and shiny elytral intervals with very sparsely distributed thin setae, similarly as e. g. D. florinensis and D. raineri (Figs 57 – 61). Dima macedonica is characterized by the scutellum raised above the plane of the elytra (clearly visible in lateral view; Fig. 164), distinctly double-sized punctation of pronotum, and the pubescence of the pronotal sides which is semi-erect along the whole margin length but in anterior half about 0.5 times as long as in posterior half (Figs 141 – 143). All these characters can be used for the separation of D. macedonica and D. raineri, which has the scutellum on the same plane as the elytra, double punctation of pronotum less distinct, with punctures more similar in size to each other (Figs 154 – 155, 165), and pubescence on the anterior halfs of pronotal sides shorter and decumbent (Figs 154 – 155). Dima florinensis shares with D. macedonica the double punctation of pronotum, but differs in the scutellum, which is on the same plane as the elytra, longer pubescence of pronotal sides, which is of about the same length along the whole margin length (Fig. 129), and by the shorter apical lobe of paramera (Figs 185, 194 – 197). Intraspecific variability. The shape of antennomeres II and III is variable. Most specimens of D. macedonica have antennomeres II and III subtriangular, but in some specimens they are more elongate, as in most specimens of D. raineri (see Wurst 1997). Similar variability occurs also in D. raineri, when some specimens have similar antennomeres II and III as the typical representatives of D. macedonica. Specimens of the northern population of D. macedonica (near Ehloec) have more convex base of scutellum.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB3FFE6C8C6FF303B2AA82F.taxon	distribution	Distribution. Macedonia (Galičica Mts., Ehloec; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB3FFE5C8C6F93A3BECAC95.taxon	materials_examined	Type depositories. Holotype, ♂ (PCJM); allotype, ♀ (PCJM); 3 paratypes, 1 ♂, 2 ♀♀ (PCRM, PCRS, PCRU). Type locality. Montenegro: Lovčen Mts. [Kotor Mts.], Njeguš – Rajčeviči (42 ° 25 ' 28.87 " N, 18 ° 49 ' 5.43 " E), 880 m. Redetermined material. MONTENEGRO: Rumija Mts., Virpazar [Virpozar], 20. VI. 1984, 1 ♀, M. & J. Hladilovi leg. (as D. dalmatina in Schimmel & Platia 2008) (PCRS). New material. MONTENEGRO: Njeguši vill. env., 1000 – 1200 m (42 ° 25 ' 30.09 " N, 18 ° 50 ' 45.00 " E), 7. – 10. V. 2014, 1 ♂, fagetum, Jan Habarta leg. (PCJM).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB3FFE5C8C6F93A3BECAC95.taxon	diagnosis	Diagnosis. Dima marvani is a medium-sized species (body length: 9.5 – 12.8 mm) with an elongate body shape, relatively long antennae (surpassing hind angles by 3.5 – 4.5 segments), shiny pronotum with sides with the short semi-erect setae along the whole length, elytra with short dense decumbent pubescence, and the elytral interstices matt, microreticulated. This species can be easily distinguished from D. elateroides by the denser body pubescence and matt elytra (Figs 22 – 24, 62 – 65), from D. moraveci by the more elongate body shape and microreticulated surface of elytra (Fig. 66), and from D. pelikani sp. nov. by the shiny pronotum (Figs 144, 151). Dima marvani is very like D. dalmatina (Figs 14 – 19, 62 – 65). The specimens from the type locality (Lovčen Mts.) differ from the typical D. dalmatina by the less distinct keel between the frons and clypeus, longer antennae in males (more than 60 % of the body length), longer pronotum (1.3 times wider than long), pronotal pubescence sparser, near sides of pronotum directed towards sides (Figs 62 – 63), and apical lobe of paramera longer (Fig. 198). Intraspecific variability. The specimens from Rumija Mts. have a darker body, almost no keel between the frons and clypeus, slightly longer antennae, and more convex and shorter pronotum than the specimens from Lovčen Mts. (Figs 64 – 65). Similar variability in Rumija Mts. is known also for the very similar D. dalmatina (Figs 18 – 19). More material of both species from different collecting sites is needed for further conclusions.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB3FFE5C8C6F93A3BECAC95.taxon	distribution	Distribution. Montenegro (Lovčen Mts., Rumija Mts.; Fig. 243).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB0FFE5C8C6FCE33C54AB2B.taxon	materials_examined	Type depository. Holotype, ♀ (PCJM). Type locality. Albania: Skhodër, Mt. Cukalit, 1300 m.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB0FFE5C8C6FCE33C54AB2B.taxon	diagnosis	Diagnosis. This species belongs to the group of species with more convex, suboval-shaped body and shiny elytral intervals with very sparsely distributed thin setae (Fig. 66). It differs from other Dima species in the region by the combination of following characters: paler and wider, more robust body, wide, shiny pronotum with very shallow fine punctures and short decumbent pubescence, and wide angulate scutellum (Figs 66, 145). By the general habitus and shapes of pronotum and scutellum, this species is like D. elateroides, but they differ by the body surface and pubescence.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB0FFE5C8C6FCE33C54AB2B.taxon	distribution	Distribution. Albania (Skhodër: Mt. Cukalit; Fig. 243).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB0FFE5C8C6FCE33C54AB2B.taxon	discussion	Remark. Dima moraveci is known only from the female holotype (Fig. 66).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB0FFE4C8C6FA063D4CAD88.taxon	materials_examined	Type depository. Holotype, ♂ (NME). Type locality. Greece: Peloponnese, Mt. Erimanthos, SE Kalentzi, 1190 m. Redetermined material. GREECE: nom. Ahaia, O. Aroania canalino, 1330 m, 1. VI. 1995 – 4. VI. 1996, 1 ♀, Giachino & Vailati leg. (as D. hladilorum in Schimmel & Platia 2008) (PCGP). New material. GREECE: distr. Ahaïa, Erímanthos Mts., Kalentzi, firrywood, 1240 m (37 ° 56 ' 52.21 " N, 21 ° 46 ' 15.90 " E), 3. VI. 2013, 1 ♂, 2 ♀♀, J. Mertlik leg. (PCJM); dtto, 2 ♀♀, V. Dušánek leg. (PCVD); dtto, 2 ♀♀, P. Brůha leg. (PCPB); distr. Ahaïa, Panachaiko Mts., Kalanistra, 20. IV. – 2. V. 2014, 2 ♀♀, Manfred Egger leg. (PCME); Erímanthos Mts., Kalentzi env. (ca. 37 ° 56 ' 49.41 " N, 21 ° 46 ' 11.26 " E), 21. V. 2015, 3 ♀♀, Manfred Egger leg. (PCME); Erímanthos Mts., Kalentzi env., 1265 m (37 ° 57 ' 2.22 " N, 21 ° 46 ' 35.46 " E), 10. VI. 2015, 8 ♂♂, 2 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♀, B. Zbuzek leg. (PCBZ); dtto, 1 ♂, P. Brůha leg. (PCPB); dtto, 2 ♂♂, T. Németh leg. (PCRK); dtto, 1 ♀ (HNHM); distr. Ahaïa, Panachaiko Mts., Kalanistra env., Agios Athanasios (ca. 38 ° 4 ' 44.65 " N, 21 ° 51 ' 35.45 " E), 10. V. 2015, 2 ♂♂, 3 ♀♀, Manfred Egger leg. (PCME); Peloponnese, Mt. Chelmos, Kalavrita env., 28. IV. 1993, 1 ♂, J. Macek leg. (PCVK); distr. Ahaïa, Aroánia Óri Mts., firrywood, 1420 m (38 ° 1 ' 14.26 " N, 22 ° 10 ' 21.28 " E), 30. V. 2013, 1 ♂, 1 ♀, J. Mertlik leg.; dtto, 30. V. 2013, 1 ♂, P. Brůha leg. (PCVD); Mt. Chelmos (38 ° 0 ' 54.31 " N, 22 ° 11 ' 24.35 " E), 12. V. 2015, 1 ♀, Manfred Egger leg. (PCME).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB0FFE4C8C6FA063D4CAD88.taxon	description	Description of female. Like male, but more robust body, shorter antennae, surpassing the posterior angles of pronotum about two segments (more than three in males), and pronotum wider (Figs 69, 71 – 74). The sclerotized spines of bursa copulatrix as in Fig. 233.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB0FFE4C8C6FA063D4CAD88.taxon	diagnosis	Diagnosis. Dima neumanni is a medium-sized to large species (body length: 11.5 – 14.7 mm) with the coarse pronotal punctation, the long straight erect setae on the pronotal sides (Fig. 146), and long straight semi-erect to erect pubescence of elytra (Figs 67 – 74). The species limits of D. neumanni and D. schnitteri are unclear and it is possible, that they are conspecific. Dima schnitteri is known only from the lower elevations of the southern part of the Erimanthos Mts. (south from the distributional range of D. neumanni; Fig. 246), is mostly larger, more flattened and paler (Figs 105 – 108). The third species in the region, D. assingi, is also morphologically very like D. neumanni, but differs in the elytra with shorter decumbent to semi-erect pubescence in basal third or half, and in not erect (only semi-erect) setae in the first halfs of the pronotal sides (Figs 6 – 9, 120). Intraspecific variability. Dima neumanni is rather a morphologically variable species, with the long erect pubescence of elytra as the only stable character. The apical part of the scutellum is either more or less rounded or subacute. The specimens from Aroania Mts. are paler, and one of the males has more elongate apical lobe of paramera. All known specimens from Aroania Mts. and some from Panachaiko Mts. have more parallel-sided and elongate pronotum than the specimens from the type locality in Erimanthos Mts. (Figs 67 – 74). It must be noted, that all specimens of D. assingi (which differs only slightly from D. neumanni and occurs also in Aroania Mts.) are lightly colored and have also more elongate pronotum (Figs 6 – 9). The limits of the Dima species distributed in the northern Peloponnese must be confirmed by future studies including much more extensive sampling and the DNAbased methods of species identification.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB0FFE4C8C6FA063D4CAD88.taxon	distribution	Distribution. Greece (Peloponnese: Erimanthos Mts., Panachaiko Mts., Aroania Mts.; Fig. 246).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB1FFE4C8C6FB983A90A979.taxon	materials_examined	Type depository. Type material and type depository unknown (Schimmel 1996). Type locality. Greece: Thessaly, Mt. Olympus, 2200 m. New material. GREECE: Olympos – Litochoro, 26. – 28. V. 1989, 1 ex., J. Turna leg. (PCGP); Olympus Mts., chata A, cca 2000 m, 9. – 10. VII. 1996, 1 ♂, R. Chodura leg. (PCJM); Litochoro – Mt. Olympus, 2000 m, 3. VI. 2008, 1 ♂, R. Reitmaier leg. (PCRU); distr. Pieriá, Olimbos Mts., Spilios Agapitos refuge env., 2000 m (40 ° 4 ' 57.01 " N, 22 ° 22 ' 21.24 " E), 12. VI. 2015, 21 ♂♂, J. Mertlik leg. (PCJM); dtto, 16 ♂♂, 4 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 22 ♂♂, P. Brůha leg. (PCPB); dtto, 2 ♂♂, T. Németh leg. (HNHM); dtto, 8 ♂♂ (PCRK).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB1FFE4C8C6FB983A90A979.taxon	description	Description of female. Like male, but with more robust body, shorter antennae surpassing the hind pronotal angles about one segment (about three segments in males), and slightly wider pronotum (Fig. 76). The sclerotized spines of bursa copulatrix as in Fig. 234.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB1FFE4C8C6FB983A90A979.taxon	diagnosis	Diagnosis. Dima olympica is easily recognizable by its small body (body length: 8.0 – 10.3 mm), very long, erect pubescence of head and pronotum, and hairs of two types on elytra: long erect ones and short decumbent ones (Figs 75 – 76, 147, 169). Dima hirtipennis from southern Peloponnese, which has also long erect setae all over its body, is larger and lacks the short decumbent pubescence on elytra (Fig. 167).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB1FFE4C8C6FB983A90A979.taxon	distribution	Distribution. Greece (Mt. Olympus; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB1FFEAC8C6F8C839E9AD38.taxon	materials_examined	Type material. Holotype: BULGARIA: distr. Petrich, Belasitsa Mts., Leshniska river, beechwood, 1060 m (41 ° 21 ' 0.64 " N, 23 ° 10 ' 45.92 " E), 6. VI. 2013, ♂, J. Mertlik leg. (PCJM). Paratypes: BULGARIA: the same data as for holotype; 6 ♂♂, 4 ♀♀ (PCJM); dtto, 2 ♂♂ (PCRK); dtto, 10 ♂♂, 3 ♀♀, V. Dušánek leg. (PCVD); dtto, 7 ♂♂, 2 ♀♀, P. Brůha leg. (PCPB); distr. Petrich, Belasica Mts., 7 km SW Petrich, beechwood, 7. VI. 2006, 1 ♂, 1 ♀, J. Mertlik leg. (PCJM); distr. Petrich, Belasica Mts., Belasica Chalet env., 6. – 8. V. 2009, 1 ♂, T. Kopecký leg. (PCTK); distr. Petrich, Belasitsa Mts., chestnut project, plot No. 4, 600 m, N 41.36560567, E 23.20410224, soil traps, 27. III. – 7. V. 2010, 1 ♂, B. Guéorguiev & C. Deltshev leg. (PCJM).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB1FFEAC8C6F8C839E9AD38.taxon	diagnosis	Diagnosis. This species is like D. lebenbaueri and D. kozufensis sp. nov. in the body shape and coloration, and in the shape of antennae. Dima orientalis sp. nov. differs from the above mentioned species in having pubescence of the pronotal sides almost decumbent, with longer semi-erect hairs only at hind angles, and scutellum subpentagonal (Fig. 148). Dima lebenbaueri has long semi-erect to erect setae at the pronotal sides and the scutellum distinctly wider (Figs 139 – 140). Dima kozufensis sp. nov. has longer semi-erect pubescence at the posterior third of the pronotal sides and a basally emarginate and more coarsely punctate scutellum (Fig. 138). Dima bruhai sp. nov. is more matt, has shorter antennomeres II and III, a wider scutellum, semi-erect pubescence at pronotal sides, much denser and longer pubescence of elytra, and longer apical lobe of paramera (Figs 10 – 13, 121, 178). Dima schimmeli, which occurs near to D. orientalis sp. nov. (Fig. 244), differs in having long, straight, dense, semi-erect pubescence on pronotum and elytra (Fig. 156).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB1FFEAC8C6F8C839E9AD38.taxon	description	Description. Holotype, male. Body medium-sized, 12.0 mm long, 4.8 mm wide, moderately shiny. Body brown, with paler, reddish-brown legs, antennae and elytral sutura and margins, elytral apices slightly paler; pubescence yellowish (Fig. 77). Head including eyes 0.5 times as wide as pronotum, frons shallowly depressed in the middle; punctation dense and coarse, intervals between punctures shiny; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina weakly defined, short. Antennae moderately long, surpassing the posterior angles of pronotum of about three segments; antennomeres II and III short, more than 1.5 times longer than wide, antennomere II slightly longer than antennomere III, together about 1.4 times longer than antennomere IV, length ratio of antennomeres II – IV 1.1: 1.0: 1.6, antennomeres IV to X gradually widened apically, less than 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semi-erect setae. Pronotum 1.4 times wider than long, widest just behind middle, moderately convex dorsally; sides convex, gradually narrowed towards anterior margin, more steeply narrowed towards posterior margin; straight in lateral view; anterior angles obtuse; posterior angles prominent, sharp, slightly produced outward; punctation sparse, coarse, moderately deep, punctures denser and coarser towards anterior margin and sparser and finer towards posterior margin; of about the same size; intervals between punctures flattened, moderately shiny. Pubescence on pronotal disc short, decumbent; that on pronotal sides decumbent at anterior two thirds and semi-erect at posterior third, long semi-erect to erect setae at posterior angles (Fig. 148). Prosternum finely, sparsely punctate, with short decumbent to semi-erect setae; prosternal lobe coarsely punctate, with long, dense semi-erect pubescence. Scutellum sub-pentagonal, sides subparallel-sided, flattened, in lateral view in the plane of elytra; frontal margin rounded, gradually declined; apex narrowed, rounded, punctation fine, sparse; pubescence sparse, fine, moderately long, decumbent. Elytra slightly elongate, sub-parallel, 3.3 times longer than pronotum, moderately convex, widest behind middle; striae well developed along the whole elytral length; surface of interstices flattened, shiny, moderately densely and finely punctate; punctures suboval, equally distributed, of the same size; pubescence sparse, short. Aedeagus elongate; paramera long, narrow, its apical lobe short, subapical tooth moderately long, subacute (Fig. 203). Female (Figs 78 – 79). Like male, but darker, blackish-brown, suboval (elytra less then 3.2 times longer than pronotum), with shorter antennae (surpassing the posterior angles of pronotum about two segments), distinctly wider pronotum (1.5 times wider than long), widest at middle, with sides evenly convex, scutellum wider, with posterior margin widely rounded. The sclerotized spines of bursa copulatrix as in Fig. 235. Intraspecific variability. Body length: 11.5 – 13.4 mm. One male specimen is paler, chestnut brown, with habitus resembling female specimens in e. g. suboval body shape, wider pronotum with evenly convex sides, and wider scutellum.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB1FFEAC8C6F8C839E9AD38.taxon	distribution	Distribution. Bulgaria (Belasica Mts.; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB1FFEAC8C6F8C839E9AD38.taxon	etymology	Etymology. The species name " orientalis " refers to the distribution of this species (eastern part of the Balkan Dima distribution; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB1FFEAC8C6F8C839E9AD38.taxon	discussion	Remark. There are two female specimens of Dima sp. collected near Stavroupoli in Xanthi district, Greece, with the following data: " Greece: distr. Xanthi, road Stavroupoli – Xanthi, ca. 10 km E Stavroupoli, 360 m (41 ° 12 ' 27.45 " N, 24 ° 47 ' 44.43 " E), 25. IV. 2007, broadleaved forest, stream, Jan Schneider leg. " (PCBZ) (Fig. 80). These specimens represent the easternmost European Dima records except D. elateroides (Fig. 244) and display a mixture of characters present in D. orientalis sp. nov. and D. bruhai sp. nov. (Figs 10 – 13, 77 – 79). They share with D. orientalis sp. nov. the sparse, short, decumbent pubescence of elytra (denser and longer in D. bruhai sp. nov.), shape of sides of pronotum (more evenly rounded in females of D. bruhai sp. nov., Fig. 11), shape of scutellum (distinctly wider in D. bruhai sp. nov.; Figs 10 – 13), and shape and size of the antennomeres II and III, which are subequal and more elongate (antennomere III apparently shorter than antennomere II in D. bruhai sp. nov.). The punctation of the pronotum is within the limits of intraspecific variability for both species, but it is denser and coarser than in most specimens of D. orientalis sp. nov. The surface of the pronotum and elytra is more matt than in D. orientalis sp. nov., similar as in D. bruhai sp. nov. (Figs 10 – 13, 77 – 80). The anterior part of the head is rather coarsely punctate and rugose, which is similar as in D. bruhai sp. nov. The sclerotized spines of bursa copulatrix are relatively longer and thinner in D. orientalis sp. nov. than in the Dima sp. from Xanthi and D. bruhai sp. nov. (Figs 223, 235 – 236). These specimens were collected at the river bank in the low elevation area. Similarly, D. orientalis can also be found in wet places near rivers in the gorges of Belasica Mts. The specimens from Xanthi are probably closely related to D. orientalis sp. nov. but more material of both sexes is needed for any reliable taxonomic conclusions.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBFFFE9C8C6FC0839DCAF5A.taxon	materials_examined	Type depositories. Holotype, ♀ (PCJM); paratype, ♀ (PCRS). Type locality. Greece: Peloponnese, Parnon Mts. Redetermined material. Peloponnese [Peloponnesus], 40 km south-east of Tripoli, SW Ag. Petros, 1100 m (37 ° 18 ' 49 '' N, 22 ° 31 ' 10 '' E), 22. III. 1997, 1 ♂, V. Assing leg. (as D. raineri in Schimmel & Platia 2008) (PCRS). New material. GREECE: Argolida, O. Parnon, str. Kosmac – Polidroso, 1070 m, 29. V. 2003 – 13. VI. 2004, 1 ♂, 1 ♀, Giachino & Vailati leg., D. hladilorum, det. Platia 2010 (PCGP); dtto, 2 ♀♀ (MCNG); Peloponnese, Parnon Mts., 22. – 30. VI. 2005, 1 ♂, I. Marvan leg. (PCVD); Parnon Mts., Kastanitsa env., 1100 – 1300 m, 19. V. 2007, 1 ♂, Karel Rébl leg. (PCJM); dtto, 1 ♂, 1 ♀, Rudolf Kmeco leg. (PCTS); dtto, 1 ♂ (PCRK); Parnon Mts., Kastanitsa env., 1100 – 1300 m, 27. – 28. V. 2010, 1 ♀, Rudolf Kmeco leg. (PCJM); distr. Arkadía, Parnon Mts., Agios Petros env., pinewood, 1190 m (37 ° 19 ' 0.99 " N, 22 ° 31 ' 53.66 " E), 31. V. 2013, 22 ♂♂, 13 ♀♀, J. Mertlik leg. (PCJM); dtto, 15 ♂♂, 8 ♀♀, V. Dušánek leg. (PCVD); dtto, 6 ♂♂, 6 ♀♀, P. Brůha leg. PCPB); Lakonia env., Parnon Mts., 3 km N Karies, 1100 m, oak – pine forest, singled at night from rocky ground, 21. V. 2011, 4 ♂♂, 6 ♀♀, leg. A. Kotán, A. Márkus & T. Németh (HNHM); dtto, 6 ♂♂, 5 ♀♀ (PCRK); dtto, 1 ♂, 1 ♀ (PCJM); Greece, Peloponnese, Mt. Parnon, Kastanitsa, 19. V. 2007, 1 ♂, 2 ♀♀, Mikyška leg. (PCAY); Peloponnese, distr. Lakonia, Parnon Mts., 3 km N Karies, oak-pine forest, 37 ° 18 ' 48.80 " N; 22 ° 31 ' 8.95 " E, 1100 m, singled at night from rocky ground, 12. V. 2013, 9 ♂♂, 9 ♀♀, leg. A. Kotán, P. Nemes, Gy. Németh, T. Németh & V. Szénási (PCRK).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBFFFE9C8C6FC0839DCAF5A.taxon	diagnosis	Diagnosis. Dima parnonensis is 12.0 – 14.5 mm long species with a shiny pronotum, moderately densely covered with coarse punctures, and shiny elytra, covered with sparse short pubescence (Figs 81 – 84, 170). It differs from the similar D. arndti by the more shiny body, pronotum widest after middle, with sides almost evenly rounded (pronotum widest at 2 / 3, with sides abruptly constricted before posterior angles in D. arndti; Figs 119, 149), and the apical lobe of paramera shorter, more robust, with the inner apical part of paramera moderately to distinctly curved (Figs 172 – 173, 204 – 205). Intraspecific variability. The typical D. parnonensis are dark brown-blackish but some specimens are paler (Figs 81 – 84).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBFFFE9C8C6FC0839DCAF5A.taxon	distribution	Distribution. Greece (Peloponnese: Parnon Mts.; Fig. 246).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBFFFE9C8C6FC0839DCAF5A.taxon	discussion	Remarks. A specimen with light body coloration (Fig. 84) was determined and figured by Schimmel & Platia (2008) as D. raineri and used erroneously for the redescription of that species, which led to the description of D. pindosensis Schimmel & Platia, 2 0 0 8 based on the typical specimens of D. raineri (Schimmel & Platia 2008). A single male specimen of Dima sp. from Crete, which is similar to both D. arndti and D. parnonensis, is discussed above under D. arndti.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBCFFE9C8C6FE333A02A9C8.taxon	materials_examined	Type depository. Holotype, ♂ (MHNP). Type locality. Greece: Mt. Ossa. Redetermined material. GREECE: Ossa Mts., Larisa, 800 m, V. 1987, 1 ♂, B. Lassalle leg. (as a paratype of D. evritaniensis in Schimmel & Platia 2008) (PCGP) (Figs 87, 206). New material. GREECE: Thessalien, Larissa, Ossageb., 1200 m, 9. – 30. V. 1996, 20 ♂♂, 11 ♀♀, T. Lebenbauer leg. (PCME); Ossa Mts., Kokino Nero, 13. V. 2005, 1 ex., J. Halada leg. (PCGP); Ossa Mts., 1200 – 1500 m, 26. IV. 2004, 1 ex., C. Busi leg. (PCGP); Thessalien, Larissa, Ossageb., 1200 m, 9. – 30. V. 1996, 1 ♂, T. Lebenbauer leg. (PCVD); Ossa (Kissavos) Mts., Spilia env., 800 m, 10. – 13. VI. 2007, 1 ♂, J. Velc leg. (PCBZ); Thessalie, Ossa Óros Mts., Spilia env., 14. – 17. VI. 2010, 1 ♂, 1 ♀, Jiří Plecháč leg. (PCJM); dtto, 2 ♂♂ (PCBZ); Thessalia, Ossa Oros Mts., NE slope W Stomio, 550 m, 30. V. 2012, 7 ♂♂, 3 ♀♀, fagetum, Tomáš Sitek leg. (PCTS); distr. Lárissa, Ossa Mts., 6.3 km NE of Spilia, 953 m (39 ° 50 ' 16.08 " N, 22 ° 41 ' 49.62 " E), beechwood, 16. VI. 2012, 1 ♂, 4 ♀♀, J. Mertlik leg. (PCJM); dtto, 4 ♂♂, 1 ♀♀, V. Dušánek leg. (PCVD); dtto, 1 ♀, M. Samek leg. (PCMS); distr. Lárissa, Ossa Mts., 6 km NE of Spilia, 975 m (39 ° 50 ' 12.89 " N, 22 ° 41 ' 48.04 " E), beechwood, 1. VI. 2014, 10 ♂♂, 12 ♀♀, J. Mertlik leg. (PCJM); dtto, 4 ♂♂, 10 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 4 ♂♂, 11 ♀♀, P. Brůha leg. (PCPB); Greece SE, Greece SE, Mavrovouni Mts., 35 km N Volos, 12. V. 2005, 1 ♀, M. Kadlecová leg. (PCPB); Thessalia, Mavrovouni Mts., 800 m, W of Sklithro, 14. – 15. VI. 2014, 1 ♀, fagetum, T. Sitek leg. (PCTS); Ossa Mts., N slopes (39 ° 50 ' 21.47 '' N, 22 ° 42 ' 34 '' E), ca. 820 m, 17. – 18. V. 2015, 3 ♂♂, 4 ♀♀, David Král leg. (PCBZ); Larisa env., Ossa Mts., 6 km E Spilea, beech forest, singled at night, 1000 m, 14. V. 2011, 5 ♂♂, 5 ♀♀, leg. T. Németh (HNHM); dtto, 18 ♂♂, 5 ♀♀, leg. A. Kotán (PCAK); dtto, 9 ♂♂, 1 ♀, leg. A. Márkus (PCAM); dtto, 2 ♀♀ (PCBS); dtto, 1 ♂, 1 ♀ (PCJM); Larisa env., Ossa Mts., 10 km W Stomio, beech forest, singled at night, 1000 m, 10. VI. 2014, 5 ♂♂, 3 ♀♀, leg. A. Kotán, Gy. Németh, B. Szelenczey & V. Szénási (PCRK); dtto, 16 ♂♂, 16 ♀♀ (PCBS); dtto, 1 ♂, 1 ♀ (PCRK); Larisa env., Ossa Mts., 5 km N Stomio, stream valley, sifted, 150 m, 14. V. 2011, 1 ♂, leg. A. Kotán, A. Márkus & T. Németh (PCRK); Thessalia, Stomio, 18. VI. 2011, 1 ♂, Pavel Štěpánek leg. (PCJM); Ossa Mts., N. slopes, ca. 820 m (39 ° 50 ' 21.47 '' N, 22 ° 42 ' 34 '' E), 17. – 18. V. 2015, 3 ♂♂, 4 ♀♀, David Král leg. (PCBZ); prov. Makedonia, distr. Larissa, Mt. Ossa, 4 km S Stomio, mixed forest, 39 ° 50 ' 15.57 " N; 22 ° 42 ' 50.62 " E, 965 m, swept & singled, 23. – 24. V. 2015, 3 ♂♂, 3 ♀♀, leg. A. Kotán, Gy. Németh, B. Szelenczey & V. Szénási (PCRK); prov. Makedonia, distr. Larissa, Ossa Mts., 6 km E Spilea, mixed forest, 39 ° 50 ' 0.20 " N; 22 ° 41 ' 2.83 " E, 1060 m, singled, 10. V. 2013, 1 ♂, leg. A. Kotán, P. Nemes, Gy. Németh, T. Németh & V. Szénási (RKPC).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBCFFE9C8C6FE333A02A9C8.taxon	diagnosis	Diagnosis. Dima pecoudi is a large species (body length: 13.0 – 15.7 mm), with typical very short, thin and decumbent body pubescence, clinging to the pronotal sides (Fig. 150), very dense pronotal punctation with matt intervals, and elytra matt, with small, dense, fine punctures, with intervals microreticulated (Figs 85 – 88, 171). Male genitalia is as in Figs 206 – 207. Dima pelikani sp. nov. shares with D. pecoudi the similar surface of elytra but differs in the semi-erect pubescence on the sides of pronotum (Fig. 151).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBCFFE9C8C6FE333A02A9C8.taxon	distribution	Distribution. Greece (Ossa Mts., Olympus Mts., Mavrovouni Mts.; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBCFFE9C8C6FE333A02A9C8.taxon	discussion	Remark. Two female specimens in the collection of G. Platia (PCPG) bear labels " Greece, Col de Katara, 15. V. 2000, G. Dubault leg. ". The occurrence of this species near Katara Pass is very improbable and we suppose that these specimens were mislabelled.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBDFFEFC8C6FF303A0FAFED.taxon	materials_examined	Type material. Holotype: MONTENEGRO: Kotor distr., Lovčen Mts., Cetinje env., 4 km of Ivanova Korita (N 42 ° 24 ' 07 '', E 18 ° 49 ' 32 ''), 1364 m, 25. – 26. V. 2014, ♂, J. Pelikán leg. (PCJM). Paratypes: MONTENEGRO: Kotor distr., Lovčen Mts., Cetinje env., 4 km of Ivanova Korita (N 42 ° 24 ' 07 '', E 18 ° 49 ' 32 ''), 1364 m, 25. – 26. V. 2014, 3 ♂♂, J. Pelikán et R. Škoda leg. (PCJM, PCLS); Kotor distr., Lovčen Mts., Ivanova Korita, Fagus forest (42 ° 22 ' 44.81 " N, 18 ° 50 ' 9.90 " E), 31. V. 2014, singled at night, 1 ♂, 1 ♀, leg. Dusanek, Kundrata & Nemeth (HNHM); dtto, 4 ♂♂, 3 ♀♀ (PCRK); dtto, 1 ♂, 3 ♀♀, V. Dušánek leg. (PCVD); Lovčen Mts, Njeguši env., 1000 – 1100 m, 10. IV. 2016 (fagetum), ♂, P. Vonička leg. (PCPV); Lovčen Mts, Njeguši env., 1000 – 1100 m, 25. V. 2016 (fagetum), 2 ♀♀, P. Vonička leg. (PCPV); ALBANIA: Skhoder distr., 1 km E Fushe Okol, Fagus forest (42 ° 23 ' 49.68 " N, 19 ° 42 ' 2.61 " E), 2. VI. 2014, swept & singled, 1 ♂, leg. Dusanek, Kundrata & Nemeth (HNHM); dtto, 3 ♂♂, 1 ♀ (PCRK); dtto, 1 ♂, 1 ♀, V. Dušánek leg. (PCVD). Other material examined. MONTENEGRO: Prekornica Abyss. pr. Konjic (– 150 m), Alexandr Veliký (Abyss), 3. VI. 2006, 1 ♂, Roman Mlejnek leg. (PCJM).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBDFFEFC8C6FF303A0FAFED.taxon	diagnosis	Diagnosis. Dima pelikani sp. nov. is very like D. pecoudi. Both species are easily recognizable by their matt, microreticulate pronotal and elytral intervals, densely punctate pronotum, very short decumbent pubescence of pronotum and elytra, and very short, decumbent, almost clinging pubescence of the pronotal sides (Figs 85 – 91, 150 – 151). They differ in body size (11.0 – 12.5 mm in D. pelikani sp. nov, 13.5 – 16.0 mm in D. pecoudi), shape of pronotum (sides more evenly rounded in D. pelikani sp. nov., longer, more sub-parallel sided in D. pecoudi), elytral striae (less distinct to indistinct in D. pelikani sp. nov, well-developed in D. pecoudi; Figs 85 – 91), and the shape of paramera (shorter apical lobe with smaller subapical lateral tooth in D. pelikani sp. nov, longer lobe with more distinct tooth in D. pecoudi; Figs 206 – 208).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBDFFEFC8C6FF303A0FAFED.taxon	description	Description. Holotype, male. Body medium-sized, 12.1 mm long, 4.8 mm wide, matt, covered with very short dense pubescence. Body brown, with paler, reddish-brown legs and antennae; pubescence yellowish (Fig. 89). Head including eyes slightly more than 0.5 times as wide as pronotum, frons with almost v-shaped depression; punctation dense and coarse, intervals between punctures rugose; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina present, sharply defined. Antennae moderately long, surpassing the posterior angles of pronotum by more than three segments; antennomeres II and III subtriangular, about 1.5 times longer than wide, subequal, together 1.2 times longer than antennomere IV, length ratio of antennomeres II – IV 1.0: 1.0: 1.6, antennomeres IV to X gradually widened apically, less than 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semierect setae. Pronotum 1.4 times wider than long, widest at middle, moderately convex dorsally; sides evenly convex, gradually slightly curved anteriorly in lateral view; anterior angles obtuse; posterior angles prominent, sharp, distinctly produced outward; punctation dense, fine, shallow, punctures on disc equally distributed, of about the same size; intervals between punctures shorter or of the same length as a diameter of puncture, microreticulate on most of pronotum, more flattened posteriorly. Pubescence on pronotal disc very short, decumbent; that on pronotal sides decumbent to almost clinging at anterior two thirds, slightly semi-erect at posterior third, longer setae at posterior angles (Fig. 151). Prosternum finely, sparsely punctate, with moderately long semi-erect setae; prosternal lobe moderately coarsely punctate, with longer semi-erect pubescence. Scutellum sub-trapezoidal, flattened, in lateral view under the plane of elytra, widest at two thirds; frontal margin rounded, gradually declined; apex widely rounded, punctation fine, dense; pubescence fine, moderately long, semi-erect. Elytra sub-parallel, 3.1 times longer than pronotum, moderately convex, widest at middle, from middle gradually narrowing towards apex; striae well developed along the whole elytral length, irregularly interrupted by coarse sparse punctures; surface of interstices flattened, matt, microreticulate, densely and finely punctate; punctures equally distributed, of about the same size; pubescence very short, very dense. Aedeagus elongate; paramera long, narrow, its apical lobe robust, subapical tooth short (Fig. 208). Female (Fig. 90). Like male, but with suboval body, shorter antennae (surpassing the posterior angles of pronotum about two segments), head less than 0.5 times as wide as pronotum, pronotum more convex and distinctly wider than in male (1.5 times wider than long), scutellum wider. The sclerotized spines of bursa copulatrix as in Fig. 237. Intraspecific variability. Body length: 11.2 – 14.0 mm. Some specimens are dark brown. The frontal depression is variable in size, shape and depth. The specimen from Prekornica, which was found in the deep abyss, differs from the remaining specimens in the smaller size, light brown body coloration, and paramera with smaller and shorter lateral subapical tooth (Fig. 91).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBDFFEFC8C6FF303A0FAFED.taxon	distribution	Distribution. Monte Negro (Lovčen Mts; Prekornica Abyss.), Albania (Skhoder: Boge) (Fig. 243). Etymology. The specific epithet is a patronym in honor of Mr. Jan Pelikán (Hradec Králové, Czech Republic), who collected a part of the type series.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBAFFEEC8C6FE7B3929ACCD.taxon	materials_examined	Type material. Holotype: GREECE: distr. Magnissia, Mt. Pilio, Chania env., 39 ° 23 ' 52.96 '', 23 ° 3 ' 24.54 '' E, 1263 m, singled at night from ground, 11. VI. 2015, ♂, leg. P. Brůha, J. Mertlik, T. Németh & B. Zbuzek (HNHM). Paratypes: GREECE: same data as holotype, 8 ♂♂, 6 ♀♀ (HNHM); dtto, 20 ♂♂, 9 ♀♀ (PCRK); dtto, 19 ♂♂, 18 ♀♀, J. Mertlik leg. (PCJM); dtto, 6 ♂♂, 5 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 2 ♂♂, 3 ♀♀, P. Brůha leg. (PCPB); Agios Onoufrios (Volos), 10. VI. 2013, 1 ♂, 1 ♀, M. Malavasi leg. (PCGP); Mt. Pelio, Chania env., 1133 m (N 39 ° 23 ' 37 ", E 23 ° 02 ' 39 "), 3. VI. 2011, 1 ♂, J. Krátký leg. (PCJM); distr. Magnissia, Mt. Pilio, Chania village env., 1263 m (39 ° 23 ' 52.96 " N, 23 ° 03 ' 24.54 " E), 15. VI. 2012, 1 ♂, 1 ♀, M. Samek leg. (PCMS); dtto, 35 ♂♂, 30 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♀ (PCRK); dtto, 10 ♂♂, 24 ♀♀, V. Dušánek leg. (PCVD); dtto, 17 ♂♂, 17 ♀♀, P. Brůha leg. (PCPB); dtto, 15 ♂♂, 17 ♀♀, B. Zbuzek leg. (PCBZ); Thessalia, Volos, Mt. Pilion, 1100 m, 7. VI. 1985, 1 ♂, 1 ♀, G. Etonti leg. (as D. hladilorum in Schimmel & Platia 2008) (PCGP); Magnissia, O. Pilio, 900 m, str. Hánia-Makriráhi, 13. VI. 1991 / 7. VI. 1992, 2 ♀♀, leg. P. M. Giachino & D. Vailati (paratypes of D. fokidensis) (PCGP, PCRS).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBAFFEEC8C6FE7B3929ACCD.taxon	diagnosis	Diagnosis. Dima pelionensis is a medium-sized species (body length: 10.5 – 14.0 mm), with sparsely punctate shiny pronotum with sides bearing decumbent pubescence in the anterior two thirds and semi-erect setae in the posterior third (Figs 152 – 153). Dima fthiotidensis and D. evritaniensis differ from D. pelionensis by the denser elytral pubescence (Figs 26 – 28, 35 – 41), pronotum matt, with denser and finer punctation (Figs 126 – 127, 130 – 134), and more or less convex scutellum (in lateral view) with more steeply declined frontal margin. Additionally, D. evritaniensis, D. etoliensis and D. zbuzeki sp. nov. have the semi-erect pubescence along the whole length of the pronotal sides (Figs 125 – 127, 162). Dima hladilorum has a more convex scutellum and more robust apical lobe of paramera (Fig. 190) and D. zbuzeki sp. nov. is smaller and has distinctly longer apical lobe of paramera (Fig. 221).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBAFFEEC8C6FE7B3929ACCD.taxon	description	Description. Holotype, male. Body medium-sized, 12.8 mm long, 5.1 mm wide, matt, covered with very short dense pubescence. Body dark brown, with paler, reddish-brown antennae, legs, elytral outer margins and suture; pubescence yellowish (Fig. 92). Head including eyes 0.6 times as wide as pronotum, frons with shallow depression; punctation dense and coarse, intervals between punctures dull; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina weakly defined. Antennae moderately long, surpassing the posterior angles of pronotum by more than three segments; antennomeres II – III subtriangular, about 1.5 times longer than wide, II slightly longer than III, together 1.2 times longer than antennomere IV, length ratio of antennomeres II – IV 1.1: 1.0: 1.9, antennomeres IV – X gradually widened apically, slightly more than 2 times longer than wide, apical antennomere subacute apically; surface of antennomeres covered with moderately long, semi-erect setae. Pronotum 1.4 times wider than long, widest at middle, moderately convex dorsally; sides evenly convex, gradually slightly curved anteriorly in lateral view; anterior angles obtuse; posterior angles prominent, sharp, distinctly produced outward; punctation dense, fine, shallow, punctures on disc equally distributed, of about the same size; intervals between punctures shorter or of the same length as a diameter of puncture, shiny, more flattened posteriorly. Pubescence on pronotal disc very short, decumbent; that on sides decumbent at the anterior two thirds and semi-erect at the posterior third, longer setae at hind angles (Fig. 152). Prosternum finely punctate, with short decumbent to semi-erect setae; prosternal lobe moderately coarsely punctate, with longer pubescence. Scutellum heart-shaped, flattened, in lateral view slightly raised above the plane of elytra, widest at two thirds; frontal margin rounded in lateral view, gradually declined; apex widely rounded, punctation fine, dense; pubescence fine, moderately long, semi-erect. Elytra sub-parallel, 3.3 times longer than pronotum, moderately convex, widest at middle, from middle gradually narrowing towards apex; striae well developed along the whole elytral length; surface of interstices flattened, moderately shiny, densely and finely punctate; punctures suboval, equally distributed, of the same size; pubescence short, dense. Aedeagus elongate; paramera long, narrow, its apical lobe robust, subapical tooth short, blunt (Fig. 209). Female (Figs 93 – 95). Like male, but with suboval body, shorter antennae (surpassing the posterior angles of pronotum about two segments), head 0.5 times as wide as pronotum, pronotum wider than in male (1.5 times wider than long), scutellum wider. The sclerotized spines of bursa copulatrix as in Fig. 238. Intraspecific variability. Some specimens have rugose, more convex and more densely punctate pronotum (Fig. 94), similarly as in some other Dima species (e. g. D. fthiotidensis, D. scutellaris). Body coloration is variable; some specimens have paler, reddish-brown some parts of body, e. g. antennae, pronotum, legs.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBAFFEEC8C6FE7B3929ACCD.taxon	distribution	Distribution. Greece (Thesally: Mt. Pelion; Fig. 245).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBAFFEEC8C6FE7B3929ACCD.taxon	etymology	Etymology. The name " pelionensis " refers to Mt. Pelion, Greece, the type locality of the new species.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBAFFEEC8C6FE7B3929ACCD.taxon	discussion	Remark. Schimmel & Platia (2008) described D. fokidensis with the holotype from Mt. Iti and paratypes from Mt. Pelion. That holotype is conspecific with D. fthiotidensis and the paratypes are here assigned to D. pelionensis sp. nov. (Fig. 95).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBBFFEDC8C6FD5B3D21A94D.taxon	materials_examined	Type depositories. Holotype of D. raineri, ♂ (SMNS); paratype of D. raineri, ♀ (SMNS); holotype of D. pindosensis, ♂ (PCJM); 2 paratypes of D. pindosensis, ♂♂ (PCJM, PCRS). Type locality. GREECE, Macedonia, Kozáni, Pass 8 km W Pendálofos, 1400 m (D. raineri); Greece, Pindos Mts., Metsovo to Anilio, 950 – 1500 m (D. pindosensis). Redetermined material. ALBANIA: Tomor, 1 ♂ (paratype of D. macedonica) (PCGP); GREECE: Epiro, Pentalofos, 1400 m, 22. VII. 1990, 1 ♂, 1 ♀, P. Cavazzuti leg. (as D. macedonica in Schimmel & Platia 2008) (PCGP); Passo Katara, 10. VII. 2001, 2 ♂♂, 1 ♀, leg. L. Saltini (as D. macedonica in Schimmel & Platia 2008) (PCGP); Pindus, Col de Katara, m 1600, VI. 1990, 2 ♂♂, B. Lassalle leg. (as D. macedonica in Schimmel & Platia 2008) (PCGP). Two specimens recorded under D. raineri in Schimmel & Platia (2008) were redetermined as D. assingi (Stilia) and D. parnonensis (Ag. Petros). New material. GREECE: distr. Kozáni, Vório Bóïo Mts., 4.5 km WNW of Pentálofos, beechwood, 1350 m (40 ° 12 ' 42.01 " N, 21 ° 5 ' 24.59 " E), 28. V. 2014, 26 ♂♂, 16 ♀♀, J. Mertlik leg. (PCJM); dtto, 2 ♂♂, 1 ♀ (PCRK); dtto, 16 ♂♂, 16 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 19 ♂♂, 14 ♀♀, P. Brůha leg. (PCPB); Grammos, Aetomilitsa env., 1300 m, 14. VI. 1996, 1 ♀, under Fagus bark, G. Kakiopoulos leg. (PCGK); Grammos, Aetomilitsa env., 1700 m, 6. – 7. VII. 2005, 1 ♀, Heinz leg. (PCGP); Grammos Mts., Aetomilitsa env., 1800 m, 7. VI. 2008, 1 ♂, Orszulík leg. (PCTS); Epirus, Smolikas Mts., 5 km NW Samarina, 1974 m, 8. VI. 2011, 1 ♂, 1 ♀, Tomáš Staněk leg. (PCJM); Pindos Mts., Kiatra Broastra pass, 1400 m, 3 km SW Amarandos vill., 3. VI. 2012, 13 ♂♂, 11 ♀♀, Tomáš Sitek leg. (PCTS); distr. Ioánina, Smolikas Mts., 4.5 km SE of Foúrka env., beechwood, 1728 m (40 ° 07 ' 46.06 " N, 20 ° 58 ' 52.94 " E), 11. VI. 2012, 90 ♂♂, 21 ♀♀, J. Mertlik leg. (PCJM); dtto, 2 ♂♂ (PCRK); dtto, 16 ♂♂, 49 ♀♀, V. Dušánek leg. (PCVD); dtto, 57 ex., P. Brůha leg. (PCPB); 47 ♂♂, 10 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 24 ♂♂, 6 ♀♀, M. Samek leg. (PCMS); distr. Ioánina, Notía Pindos Mts., 3.15 km NW of Metsovo env., beechwood, 1400 m (39 ° 47 ' 16.86 " N, 21 ° 08 ' 49.31 " E), 12. VI. 2012, 25 ♂♂, 15 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♂ (PCRK); dtto, 57 ♂♂, 11 ♀♀, V. Dušánek leg. (PCVD); dtto, 22 ♂♂, 16 ♀♀, P. Brůha leg. (PCPB); dtto, 40 ♂♂, 25 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 8 ♂♂, 12 ♀♀, M. Samek leg. (PCMS); Epirus, Pindos Mts., Milia, 1360 m, 20. VI. 2014, 1 ♂, 3 ♀♀, fagetum, Tomáš Sitek leg. (PCTS); Trikala, Kastania [dint. Castanea], 10 – 15. VII. 2010, 1 ♀, Padovani & Malmusi leg. (PCGP); Pindo, 7 km Est Kastania, 1200 m, VI. 2011, 2 ♀♀, C. Sola leg. (PCGP); Ioanina, Metsovo, 1400 m, VI. 1997, 22. VI. 1997, C. Busi leg. (PCGP); Col de Katara, 15. V. 2000, 1 ♂, G. Dubault leg. (PCGP); Pindos, Col Kathara, 15. V. 2000, 1 ♂, no further data (PCGP); Ipiros, pref. Joannina, 2.5 km NE Metsovo, 39 ° 46 ' 58.94 " N, 21 ° 9 ' 7.27 " E, 1430 m, beech forest, singled at night, 3. VII. 2012, 2 ♂♂, 2 ♀♀, leg. A. Kotán, A. Márkus, P. Nemes & T. Németh (HNHM); dtto, 3 ♂♂, 6 ♀♀ (PCRK); dtto, 14. VI. 2014, 2 ♂♂, 1 ♀, leg. A. Kotán, Gy. Németh, B. Szelenczey & V. Szénási (PCRK); dtto, 10 ♂♂, 4 ♀♀ (PCBS); Ipiros, pref. Joannina, 7 km NW Metsovo, edge of a beech forest, singled at daytime, 39 ° 48 ' 51.53 " N, 21 ° 8 ' 28.16 " E, 1360 m, 3. VII. 2012, 2 ♂♂, leg. A. Kotán, A. Márkus, P. Nemes & T. Németh Németh (PCRK); Ipiros, pref. Joannina, 2 km Fourka, 40 ° 8 ' 11.60 " N, 20 ° 57 ' 37.40 " E, 1450 m, beech forest, singled at night, 4. VII. 2012, 2 ♀♀, leg. A. Kotán, A. Márkus, P. Nemes & T. Németh (HNHM); dtto, 7 ♀♀ (PCRK).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBBFFEDC8C6FD5B3D21A94D.taxon	diagnosis	Diagnosis. Dima raineri is a medium-sized species (body length: 9.0 – 13.5 mm) with a more convex, suboval body and shiny elytral intervals with very sparsely distributed thin setae (Figs 9 6 – 100). This species is characterized by the pubescence of the pronotal sides, which is short decumbent in anterior third or half and longer semi-erect posteriorly (Figs 154 – 155). This type of pubescence is present also in D. dusaneki sp. nov. but it differs in the more robust body and the shape of pronotum (wide, evenly convex, more rounded laterally; Figs 20 – 21). Similar species, D. macedonica, has the scutellum raised above the plane of the elytra (Figs 164 – 165), more distinctly double-sized pronotal punctation (punctures more like each other in size in D. raineri, larger differences between the size of punctures in D. macedonica; Figs 141 – 143, 154 – 155), head and pronotum surface in most cases more densely covered with pubescence, and the pubescence on the pronotal sides semi-erect along the whole margin length (although shorter at anterior half; Figs 141 – 143), and D. florinensis has also the punctures on pronotum of two distinct sizes and longer pubescence on the pronotal sides, which is of about the same length along the whole margin length (Fig. 129). Male genitalia is as in Figs 210 – 213. Intraspecific variability. The striae on the elytra are variable in this species. The holotype and some specimens are distinctly striate, but usually the striae are weakly developed and in some cases they are replaced by the rows of punctures, mainly in the posterior half. Some smaller specimens with the rather smooth elytra with sparse punctures (i. e. those from Grammos) may resemble D. elateroides but they can be easily distinguished by the almost smooth pronotum and oval scutellum. Additionally, the shape of antennomeres II and III is variable. Most specimens of D. raineri have antennomeres II and III elongate, but in some specimens they are more subtriangular (or at least antennomere III), similarly as in typical D. macedonica (see Wurst 1997). This variability led to the several misidentifications between these species (Schimmel & Platia 2008).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBBFFEDC8C6FD5B3D21A94D.taxon	distribution	Distribution. Albania (Mt. Tomorr), Greece (Vório Bóïo Mts., Grammos Mts., Pindos Mts.) (Fig. 244). First record for Albania.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFBBFFEDC8C6FD5B3D21A94D.taxon	discussion	Remarks. Schimmel & Platia (2008) did not study the holotype of D. raineri described from the northern part of Pindos Mts. (deposited in the SMNS). Instead, they used a pale specimen of the another (but Peloponnesian) species as a representative of D. raineri and the diagnosis and all conclusions are based on that specimen (Fig. 84). Unfortunatelly, the specimen from the Peloponnese, which Schimmel & Platia (2008) thought was D. raineri, actually represented a species described newly in the same study from several other specimens as D. parnonensis (Schimmel & Platia 2008). What is more, as a result of the above mentioned confusion, specimens of the real D. raineri were newly described in the same study as D. pindosensis (Schimmel & Platia 2008; Fig. 99). Therefore, we herein synonymize Dima pindosensis Schimmel & Platia, 2008 under Dima raineri Wurst, 1997. The specimen from Mt. Tomorr, Albania, which was originally described as a part of the type series in D. macedonica, is definitely more like D. raineri than to D. macedonica (e. g. in the shape of the scutellum and in the pronotal punctation; Fig. 100), however, its pubescence of the pronotal sides is almost missing (Fig. 100). This specimen represents the northernmost known population of this species (Fig. 244). Additionally, we studied a dark brown female from the Elbasan county, Albania (locality label as follows: Albania, Librazhd, Shebenikut Qarishte env., 1500 m, 3. VII. 2001, leg. P. Moravec; PCJM), which was identified as D. macedonica by Schimmel & Platia (2008) (Fig. 101). This specimen is much more like D. raineri in the pronotal punctation and pubescence of pronotal sides (Fig. 155). However, it is about 70 km farther north from the known D. raineri distributional range, closer to the area of D. macedonica (Fig. 244). This female might be a teratological specimen; it has the strongly transversely convex pronotum in the posterior half, distinctly convex basal and central parts of elytra and only weakly developed elytral striae (Fig. 101). Unfortunately, we are not able to properly determine this specimen to a species level until more material from that area is available for study.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB9FFECC8C6FF303B17AD33.taxon	materials_examined	Type depository. Holotype, ♂ (PCGP). Type locality. Greece: Wetland Kerkini, Ramna Site, 41 ° 17 ' 42.5 '' N, 23 ° 11 ' 33.1 '' E [41 ° 17 ' N, 23 ° 11 ' E], 750 m. New material. Greece, distr. Séres, Kerkini Mts., Ano Vironia env., Ramna, 610 m (41 ° 17 ' 37.54 " N, 23 ° 11 ' 42.99 " E), 5. VI. 2015, 2 ♂♂, 1 ♀, J. Mertlik leg. (PCJM); dtto, 1 ♂, 1 ♀, B. Zbuzek leg. (PCBZ); dtto, 3 ♂♂, P. Brůha leg. (PCPB); dtto, 1 ♀, T. Németh leg. (PCRK).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB9FFECC8C6FF303B17AD33.taxon	description	Description of female. Like male, but with wider, more convex pronotum and shorter antennae, surpassing the hind pronotal angles by about 2.5 segments (3.5 in males) (Fig. 104). The sclerotized spines of bursa copulatrix as in Fig. 239.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB9FFECC8C6FF303B17AD33.taxon	diagnosis	Diagnosis. Dima schimmeli is a medium-sized species (body length: 10.3 – 13.0 mm) with pronotal punctures of variable size, sparser on disc and denser near margins, with long, straight, dense, semi-erect pubescence on pronotum and elytra (Figs 102 – 104, 156), and long apical lobes of paramera (Figs 214 – 215). It is the only longhaired species in the region (Fig. 244). Dima schimmeli resembles the species from the north Peloponnese (D. assingi, D. neumanni, D. schnitteri) by the body shape and long straight pubescence, but it differs in the semi-erect (about 45 °) pubescence on pronotum (erect in the Peloponnesian species; Figs 120, 146, 156 – 157).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB9FFECC8C6FF303B17AD33.taxon	distribution	Distribution. Greece (Macedonia: Kerkini Mts.; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB9FFECC8C6FC0E3AB7A892.taxon	materials_examined	Type depositories. Holotype, ♂ (NME), 11 paratypes, 5 ♂♂, 6 ♀♀ (NME, PCGP). Type locality. Greece: Peloponnese, Erimanthos Mts., Panapoulos [Panap]. New material. GREECE: Peloponnese, distr. Illía, Panopoulos env., Kapeli forest, 530 m (37 ° 49 ' 49.91 " E, 21 ° 39 ' 26.02 " N), 11. VI. 2015, 1 ♀, J. Mertlik leg. (PCJM); dtto, 1 ♀, P. Brůha leg. (PCPB); dtto, 1 ♀, T. Németh leg. (HNHM); dtto, 3 ♂♂, 1 ♀ (PCRK); Morea, Cumani, Brenske, Coll. Reitter, 1 ♀ (HNHM).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB9FFECC8C6FC0E3AB7A892.taxon	diagnosis	Diagnosis. Dima schnitteri is a large species (body length: 13.7 – 17.0 mm) with a moderately convex body, coarse pronotal punctation, long straight erect setae on the pronotal lateral margin, and long straight semi-erect to erect pubescence on elytra (Figs 105 – 108, 157). Male genitalia is as in Fig. 216; the sclerotized spines of bursa copulatrix is as in Fig. 240. Dima schnitteri is morphologically very like D. neumanni (Figs 67 – 74). Most specimens of D. schnitteri are larger and paler than typical D. neumanni, however considering geography and the morphological variability in Dima specimens from the northern Peloponnese, we cannot exclude the possibility that the above mentioned species are conspecific. Similar species, D. assingi, differs from D. schnitteri in the more elongate pronotum, semi-erect (not erect) setae in the first half of the pronotal lateral margin, and elytra with shorter decumbent to semi-erect pubescence in basal third or half (Figs 6 – 9, 120). We refrain from making any taxonomic statements until more material from the northern Peloponnese is available for study. Intraspecific variability. Specimens from the type series are light brown (Figs 105 – 106), but the freshly collected specimens (2015; Figs 107 – 108) are dark brown. The light coloration of the type specimens might have been caused by the liquid in the pitfall traps.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB9FFECC8C6FC0E3AB7A892.taxon	distribution	Distribution. Greece (Erimanthos Mts.; Fig. 246).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB9FFD3C8C6F8ED3D7BAB81.taxon	materials_examined	Type depositories. Holotype of D. vailatii, ♂ (MCSB); 13 paratypes of D. vailatii, 7 ♂♂, 6 ♀♀ (MRSN, PCGP, PCRS); holotype of D. riesei, ♂ (MCNG); 2 paratypes of D. riesei, ♂♂ (PCGP, PCSR). Type locality. GREECE: Péla province, Panagitsa, Ag. Georgios, 950 m (D. vailatii); GREECE: Kajmakčalan (Voras – Skicenter), 1500 – 2000 m (D. riesei). Redetermined material. GREECE: Macedonia, Kajmačkalan Mts., 5. VI. 1995, 1 ♂, leg. Vladimír Skoupý (as D. hladilorum in Schimmel & Platia 2008) (PCJM); Maced. occ., distr Péla, Kajmačkalan, 1800 – 2200 m, 5. VI. 1995, 1 ♂, leg. Dr. Fencl & Hosnedl (as D. hladilorum in Schimmel & Platia 2008) (PCJM). New material. GREECE: Macedonia, Kajmakčalan Mts., 21. VI. 2003, 1 ♂, V. Skoupý leg. (PCJB); Macedonia, Aghios Athanasios, Kaimaktsalan Mts., Mt. Voras, 1300 m, 16. VI. 2010, 2 ♂♂, 2 ♀♀, Tomáš Sitek leg. (PCTS); Macedonia, Aghios Athanasios, Mt. Voras, Kaimaktsalan, 1300 m, 7. – 9. VI. 2012, 3 ♂♂, fagetum, Tomáš Sitek leg. (PCTS); Macedonia, Kaimaktsalan Mts., Mt. Voras, 1820 m, 8. VI. 2012, 3 ♂♂, 1 ♀, pinetum, Tomáš Sitek leg. (PCTS); distr. Péla, Kajmakčalan Mts., road from Kerasias, 1563 – 1573 m (40 ° 53 ' 54.42 " N, 21 ° 51 ' 51.69 " E), 10. VI. 2012, beechwood, 48 ♂♂, 48 ♀♀, J. Mertlik leg. (PCJM); dtto, 1 ♂, 1 ♀ (PCRK); 32 ♂♂, 28 ♀♀, V. Dušánek leg. (PCVD); dtto, 19 ♂♂, 15 ♀♀, P. Brůha leg. (PCPB); dtto, 11 ♂♂, 14 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 14 ♂♂, 3 ♀♀, M. Samek leg. (PCMS); dtto, 5. VI. 2015, 1 ♂, 3 ♀♀, Richard Škoda leg. (PCPB); distr. Péla, Kajmakčalan Mts., 1.5 km NW of Agios Georgios, beechwood, 1340 m (40 ° 51 ' 54.07 " N, 21 ° 46 ' 58.81 " E), 27. V. 2014, 5 ♂♂, 4 ♀♀, J. Mertlik leg. (PCJM); dtto, 3 ♂♂, 2 ♀♀, B. Zbuzek leg. (PCBZ); dtto, 2 ♂♂, 3 ♀♀, P. Brůha leg. (PCPB); prov. Makedonia, distr. Péla, Kaimaktsalan Mts., 1550 m, beech forest, 40 ° 53 ' 59.11 " N, 21 ° 51 ' 47.52 " E, 1573 m, singled at night, 17. V. 2013, 10 ♂♂, 5 ♀♀, leg. A. Kotán, P. Nemes, Gy. Németh, T. Németh & V. Szénási (HNHM); dtto, 11 ♂♂, 6 ♀♀ (PCRK); dtto, 28 ♂♂, 7 ♀♀ (PCAK).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB9FFD3C8C6F8ED3D7BAB81.taxon	diagnosis	Diagnosis. Dima vailatii is a medium-sized species (body length: 9.5 – 13.0 mm) with suboval body, pronotum with fine, dense and moderately deep punctures, intervals shiny, with pubescence on the pronotal sides short, decumbent along almost the whole length, with elytral pubescence dense, short, decumbent, with moderately shiny intervals, and with the scutellum slightly raised above the elytral plane (Figs 109 – 113, 158 – 160). Dima vailatii differs from all other species occuring in the surrounding mountain ranges (D. kozufensis sp. nov., D. lebenbaueri and D. florinensis) by the pubescence of pronotal sides, which is short, curved, decumbent and of the same length almost along the whole length; with distinctly longer hairs only at hind angles. Additionally, D. kozufensis sp. nov. and D. lebenbaueri differ from D. vailatii by the sparser pronotal punctation (Figs 138 – 140) and D. florinensis by sparsely pubescent elytra with fine setae (Figs 31 – 34). Male genitalia is as in Figs 217 – 219. Intraspecific variability. There is a variability in body coloration within this species. Paler specimens were collected mainly on the southern slopes of Kajmakčalan in the forested gorges in the surroundings of Agios Georgios (950 – 1340 m), but they are known also from other parts. The darker specimens (Figs 111 – 112) were collected mostly in the beech forests on the southern slopes (1340 m) and in the eastern part of the mountain range (1563 – 1573 m). The highest known locality lies in a pine forest (1820 m), where the smallest specimens were collected.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB9FFD3C8C6F8ED3D7BAB81.taxon	distribution	Distribution. Greece (Voras Mts., Kajmakčalan; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FFB9FFD3C8C6F8ED3D7BAB81.taxon	discussion	Remark. We did not find any morphological differences between D. vailatii and D. riesei (Figs 109, 113). Therefore, we herein synonymize Dima riesei Platia, 2010 under Dima vailatii Schimmel & Platia, 2008.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FF86FFD2C8C6F9903A46AB55.taxon	materials_examined	Type material. Holotype: ALBANIA: Zepës Mts., Lar env., 1600 m, 8. VI. 2014, ♂, P. Moravec leg. (PCJM). Paratypes: ALBANIA: Zepës Mts., Lar env., 1600 m, 8. VI. 2014, 1 ♀, P. Moravec leg. (PCJM); dtto, 1 ♂, 1 ♀, P. Vonička, P. Moravec et L. Blažej leg. (PCPB).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FF86FFD2C8C6F9903A46AB55.taxon	diagnosis	Diagnosis. Dima vonickai sp. nov. belongs to the group of species with more convex, suboval-shaped body and shiny elytral intervals with very sparsely distributed thin setae (Fig. 114 – 115). This species is like D. fialai sp. nov., D. dusaneki sp. nov. and D. raineri based on the pubescence of the pronotal lateral margin, which is short, almost decumbent in anterior part and longer, semi-erect posteriorly (Fig. 161). Dima fialai sp. nov. and D. dusaneki sp. nov. differ from D. vonickai sp. nov. by more robust body, wide and convex pronotum (Figs 20 – 21, 29 – 30), and shorter apical lobe of paramera (Figs 181, 184). Dima raineri has pronotal punctures of two different sizes (almost the same size in D. vonickai sp. nov.; Figs 154 – 155, 161) and differently shaped paramera (Figs 210 – 213, 220).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FF86FFD2C8C6F9903A46AB55.taxon	description	Description. Holotype, male. Body medium-sized, 12.5 mm long, 5.1 mm wide, shiny, covered with short dense pubescence on pronotum and sparse on elytra. Body black, with paler, reddish-brown legs, antennae, elytral sutura and margins; pubescence yellowish (Fig. 114). Head including eyes 0.6 times as wide as pronotum, frons with shallow v-shaped depression; punctation sparse and coarse, intervals between punctures shiny; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina weakly developed, short. Antennae moderately long, surpassing the posterior angles of pronotum of about 3.5 segments; antennomeres II and III subtriangular, more than 1.5 times longer than wide, subequal, together about 1.4 times longer than antennomere IV, length ratio of antennomeres II – IV 1.0: 1.0: 1.4, antennomeres IV to X gradually widened apically, less than 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semi-erect setae. Pronotum 1.4 times wider than long, widest just behind middle, moderately convex dorsally; sides convex; anterior angles obtuse; posterior angles prominent, sharp, slightly produced outward; punctation sparse, moderately fine, punctures on disc equally distributed, of about the same size; intervals between punctures of 2 – 3 times a diameter of puncture, flat, shiny. Pubescence on pronotal disc moderately dense, short, decumbent; that on pronotal sides in anterior two thirds decumbent, in posterior third semi-erect; setae in hind angles long, erect (Fig. 161). Prosternum finely, sparsely punctate, interstices glabrous, with short decumbent to semi-erect setae, denser at margins; prosternal lobe moderately coarsely punctate, with longer semi-erect pubescence. Scutellum sub-pentagonal, with sides sub-parallel, in lateral view slightly raised above the plane of elytra, widest at two thirds; frontal margin rounded, gradually declined, apex rounded; punctation fine, dense; pubescence fine, short, semi-erect. Elytra suboval, 3.3 times longer than pronotum, convex, widest at posterior two thirds; striae well developed along the whole elytral length, some of them irregularly interrupted by coarse sparse punctures; surface of interstices flattened, shiny, sparsely punctate; punctures shallow, suboval, equally distributed, of the same size; pubescence short, very sparse. Aedeagus elongate; paramera long, narrow, its apical lobe elongate, subapical tooth subacute (Fig. 220). Female (Fig. 115). Very like male (in contrast with most other Dima spp.), but more robust, with shorter antennae (surpassing the posterior angles of pronotum about two segments) and slightly wider scutellum. The sclerotized spines of bursa copulatrix as in Fig. 241. Intraspecific variability. Body length: 10.7 – 12.3 mm.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FF86FFD2C8C6F9903A46AB55.taxon	distribution	Distribution. Albania (Zepës Mts.; Fig. 244).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FF86FFD2C8C6F9903A46AB55.taxon	etymology	Etymology. The specific epithet is a patronym in honor of Mr. Pavel Vonička (Liberec, Czech Republic), who collected a part of the type series.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FF87FFD1C8C6FA233A46ABA5.taxon	materials_examined	Type material. Holotype: GREECE: distr. Kardítsa, Agrafa Mts., 2.7 km SW of Neraida, firrywood, 1518 m (39 ° 16 ' 45.92 " N, 21 ° 40 ' 57.60 " E), 31. V. 2014, ♂, J. Mertlik leg. (PCJM). Paratypes: GREECE: distr. Karditsa, Pindos Mts., Neraida env. [Rafina env.] (39 ° 17 ' 31.29 " N, 21 ° 41 ' 58.48 " E), 4. – 8. VI. 2012, 2 ♂♂, 1 ♀, Karel Hodek leg. (PCJM); dtto, 3 ♂♂, 3 ♀♀ (PCPB); dtto, 3 ♂♂, 5 ♀♀ (PCKH); distr. Kardítsa, Agrafa Mts., 2.7 km SW of Neraida, firrywood, 1518 m (39 ° 16 ' 45.92 " N, 21 ° 40 ' 57.60 " E), 31. V. 2014, 15 ♂♂, 5 ♀♀, J. Mertlik leg. (PCJM); dtto, 11 ♂♂, B. Zbuzek leg. (PCBZ); dtto, 6 ♂♂, 3 ♀♀, P. Brůha leg. (PCPB); Thessalia pref., Karditsa, Pindos Mts., Neraida, Abies forest, singled at night from ground, 39 ° 16 ' 45.92 " N, 21 ° 40 ' 57.60 " E, 1500 m, 16. VI. 2014, 1 ♂, 1 ♀, leg. A. Kotán, Gy. Németh, B. Szelenczey & V. Szénási (HNHM); dtto, 5 ♂♂, 2 ♀♀ (PCBS); dtto, 6 ♂♂ (PCRK).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FF87FFD1C8C6FA233A46ABA5.taxon	diagnosis	Diagnosis. Dima zbuzeki sp. nov. is similar in the shape, coloration and pubescence of body, punctation of pronotum and shape of apical lobes of paramera to D. hladilorum (Figs 47 – 49, 116 – 118, 137, 162, 190, 221). These species differ in the scutellum, which is wider, flat, and only slightly emarginate basally in D. zbuzeki sp. nov. and narrower, convex, and strongly emarginate basally in D. hladilorum. This species differs from D. fthiotidensis and D. evritaniensis by the shiny pronotum with sparse punctation (Figs 126 – 127, 130 – 134, 162) and from D. pelionensis by the pubescence of pronotal lateral margin, which is semi-erect along the whole margin length (Figs 152 – 153). Dima etoliensis is larger, has longer antennae with longer segments (in females, as no males are known for D. etoliensis) and more parallel-sided elytra (Fig. 25).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FF87FFD1C8C6FA233A46ABA5.taxon	description	Description. Holotype, male. Body small, 9.9 mm long, 4.0 mm wide, moderately shiny pronotum, more matt elytra. Body brown, with paler, reddish-brown legs, antennae, elytral sutura and margins; pubescence yellowish (Fig. 116). Head including eyes 0.6 times as wide as pronotum, frons with shallow depression medially; punctation dense and coarse, intervals between punctures shiny posterad, rugose in anterior part; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina short, indistinct. Antennae moderately longer surpassing the posterior angles of pronotum of about four segments; antennomeres II and III short, subequal, less than 1.5 times longer than wide, together about 1.2 times longer than antennomere IV, length ratio of antennomeres II – IV 1.1: 1.0: 1.8, antennomeres IV – X gradually widened apically, about 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semi-erect setae. Pronotum 1.4 times wider than long, widest at middle, moderately convex dorsally; sides slightly and evenly convex, anterior angles obtuse; posterior angles prominent, sharp, slightly produced outward; punctation moderately dense, coarse, punctures on disc equally distributed; intervals between punctures moderately shiny. Pubescence on pronotal disc moderately long, semi-erect; that on pronotal sides semi-erect along the whole length, gradually longer towards posterior angles, longest and erect in posterior angles (Fig. 162). Prosternum finely, moderately sparsely punctate, with short semi-erect setae; prosternal lobe moderately coarsely punctate, with long semi-erect pubescence. Scutellum sub-pentagonal, flattened, slightly depressed medially, in lateral view slightly above the plane of elytra, widest at middle; frontal margin slightly emarginate, gradually declined; apex widely rounded, punctation sparse, coarse; pubescence fine, moderately long, semi-erect. Elytra sub-parallel, 3.4 times longer than pronotum, moderately convex, widest behind middle; striae well developed in 4 / 5 of elytral length, interrupted irregularly at apical part with coarse sparse punctures; surface of interstices flattened, slightly matt, densely and finely punctate; punctures suboval, equally distributed, of the same size. Aedeagus elongate; paramera long, narrow, its apical lobe long, narrow, subapical tooth robust, obtuse (Fig. 221). Female (Fig. 117). Like male but slightly paler, suboval, antennae shorter (about 2.5 segments surpassing posterior angles of pronotum), pronotum wider, 1.5 times wider than long, widest at middle. The sclerotized spines of bursa copulatrix as in Fig. 242. Intraspecific variability. Body length: 9.0 – 11.5 mm. Body coloration is light brown in some specimens (Fig. 118).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FF87FFD1C8C6FA233A46ABA5.taxon	distribution	Distribution. Greece (Agrafa Mts.; Fig. 245).	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
039687B0FF87FFD1C8C6FA233A46ABA5.taxon	etymology	Etymology. The specific epithet is a patronym in honor of Mr. Bořivoj Zbuzek (Praha, Czech Republic), who collected a part of the type series.	en	Mertlik, Josef, Németh, Tamás, Kundrata, Robin (2017): Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula. Zootaxa 4220 (1): 1-63, DOI: 10.11646/zootaxa.4220.1.1
