taxonID	type	description	language	source
03961979FFAEFF8DF7D8FB1EEC0C78DB.taxon	description	C o m m e n t: The known distribution of this endogean species is confined to Mallorca (HERMAN 2001).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFAEFF8EF7D8FA2BEA387C51.taxon	description	C o m m e n t: This adventive species was first recorded from Europe and is now widespread in West Europe (including the Atlantic Islands) eastwards to Central Europe 1107 (HERMAN 2001, SMETANA 2004). According to SMETANA (2004), it has been reported also from mainland Spain, but primary records are unknown to me.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFADFF8EF7D8FE51EA4B7F7E.taxon	description	C o m m e n t: This rare species is widespread in Europe, but was previously unknown from Spain and the whole of the Iberian Peninsula (SMETANA 2004).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFADFF8FF7D8FD46ECB77C89.taxon	description	D e s c r i p t i o n: Body length 4.5 mm. Habitus as in Fig. 1. Coloration: forebody reddish; abdomen dark-brown, with the posterior margins of the segments reddish; legs and antennae reddish. Head strongly transverse, 1.25 times as wide as long, distinctly widened behind eyes (Fig. 2); punctation relatively fine, dense, and granulose; interstices with microsculpture and on average approximately as wide as average diameter of punctures; eyes small, not projecting from lateral contours of head, little more than half as long as postocular region in dorsal view. Antennae short; antennomeres V-VIII only indistinctly oblong (Fig. 3). Pronotum strongly transverse, 1.35 times as wide as long and 1.1 times as wide as head (Fig. 2); antero-laterally with rather extensive sub-circular impression on either side; anterior and posterior angles each with very long black seta; punctation granulose, denser and more distinctly granulose in lateral than in median portions; interstices without distinct microreticulation and distinctly more glossy than those of head. Elytra extremely short, approximately 0.5 times as long as pronotum (Fig. 2), posterolaterally with rather extensive and deep impression; lateral margins with numerous long dark setae; punctation coarser than that of pronotum, somewhat granulose; interstices without microsculpture. Hind wings absent. Legs short; tibiae flattened; metatibia approximately 0.45 mm long; metatarsus approximately 0.7 times as long as metatibia. Abdomen approximately as wide as elytra; punctation dense and moderately fine; interstices without distinct microsculpture and glossy; posterior margin of tergite VII without palisade fringe.: sternite VIII with the usual posterior excision, otherwise unmodified; aedeagus shaped as in Figs 4 - 5; apex of ventral process rounded in ventral view. E t y m o l o g y: The species is dedicated to Alexey Shavrin, one of the collectors of the holotype. C o m p a r a t i v e n o t e s: In external appearance, A. shavrini somewhat resembles A. latus (ROSENHAUER) from southern Andalucía. It is distinguished from this species by the distinct microsculpture and the different punctation on the head (A. latus: shallowly areolate), the much more transverse pronotum, the presence of antero-lateral impressions on the pronotum, the even shorter and more transverse elytra, and by the shape of the aedeagus (A. latus: apex of ventral process concave in ventral view). 1108 D i s t r i b u t i o n a n d b i o n o m i c s: The type locality is situated in Jaén, Andalucía, southern Spain, at an altitude of approximately 2000 m. It seems likely that, like other Eurysunius species, A. shavrini is associated with ants of the genus Tetramorium MAYR (ASSING 2003 b).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFACFF8FF7D8FAD8EB257B39.taxon	description	C o m m e n t: According to SMETANA (2004), this rarely found species is distributed from the Balkans to the Western Mediterranean, but was previously unknown from Spain. It was only recently reported from Portugal for the first time (SILVA et al. 2006).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFACFF8FF7D8FC9CEDB679C8.taxon	description	C o m m e n t: Stenistoderus nothus is widespread in the Western Mediterranean, but was previously unknown from Portugal (SMETANA 2004).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFACFF8FF7D8FE1EEFE07E04.taxon	description	C o m m e n t: The distribution of S. subangulatus is confined to the extreme south of Spain and Morocco.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFACFF88F7D8F989ED3F7DD1.taxon	description	C o m m e n t: Gabrius cobosi is endemic to the extreme south of Spain. The species was originally described from material from the Sierra de Ronda, but apparently has never been recorded since. 1109	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFABFF88F7D8FC33ECCB78D8.taxon	description	C o m m e n t: The distribution of this rarely collected species is confined to the Western Mediterranean, from Italy to the Iberian Peninsula and North Africa (Algeria, Tunisia) (HERMAN 2001, SMETANA 2004).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFABFF88F7D8FD1BECC57EB3.taxon	description	C o m m e n t: Quedius truncicola is widespread in Europe, but recorded only rarely (HERMAN 2001, SMETANA 2004). COIFFAIT (1978) reports the species (as Q. ventralis) from " le sud de la Péninsule ibérique ".	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFABFF88F7D8FEDCEF337F8B.taxon	description	C o m m e n t: The distribution of T. herculeanus is confined to the extreme south of Spain.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFABFF89F7D8FA28ED3F7DD1.taxon	description	C o m m e n t: Quedius abietum has a circum-Mediterranean distribution ranging from Morocco and the Iberian Peninsula to Turkey. According to GAMARRA & OUTERELO (2008), it has been reported from four provinces in Spain, but was previously unknown from Murcia. 1110	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFAAFF8AF7D8F93AED027F39.taxon	description	ments IX-X missing]: " Alp. aetol. / aetolicus Kr. Typ von Kiesw. / cruentus Oliv., syn: aetolicus Krtz. / aetolicus K sp. pr. = (subapicalis Joy), det. E. Gridelli / coll. Kraatz / Syntypus / Lectotype Quedius aetolicus Kr., designé par D. Drugmand 1999 / DEI Müncheberg Col - 00008 / Lectotypus Quedius aetolicus Kraatz, desig. V. Assing 2010 " (SDEI). 1111 C o m m e n t: The type material of this species was examined in the course of the identification of Microsaurus material from Spain. The original description is based on " ein Pärchen " collected " in den ätolischen Alpen " by " Herrn v. Kiesenwetter " (KRAATZ 1858). The male syntype was located in the Kraatz collection at the SDEI. The specimen has a lectotype label by D. Drugmand attached to it, but this lectotype designation was never published. Since the identity and whereabouts of the second syntype are uncertain, the above male is here designated as the lectotype. The male primary and secondary sexual characters are illustrated in Figs 6 - 8.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA9FF8AF7D8FD8EEFA47E21.taxon	description	C o m m e n t: The above specimen represents the first record since the original description, which is based on a single specimen from the Sierra de Espuña, Murcia (ASSING 2004).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA9FF8AF7D8FB59ED427BC2.taxon	description	C o m m e n t: This adventive species was originally described from Guadeloupe (Caribbean). In Europe, it has been reported only from Italy, where it was first observed in the vicinity of Vicenca in 1982 and is now known from numerous localities, including Sardinia and Sicily (DE MARZO & ZANETTI 2007). The above specimens represent the first records from Spain and the Iberian Peninsula, suggesting that the species is probably widespread in the Western Mediterranean today.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA9FF8AF7D8FCA6EDF57949.taxon	description	C o m m e n t: The previously known distribution of T. marchii included Central and Northern Europe, Italy, France, and the Canary Islands. The above specimens represent the first record from mainland Spain and the Iberian Peninsula.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA9FF8BF7D8F8C2EDAC7CA5.taxon	description	1112 C o m m e n t: This recently described species was previously recorded only from the type locality in Zaragoza province, northeastern Spain. Since the type specimens were collected by trapping (TRONQUET 2002), the habitat was unknown. The above records suggest that the species may be associated with ants, probably wood-ants (Formica sp.), since the examined specimens were sifted from anthills.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA8FF8BF7D8FA00EF9C7BD4.taxon	description	C o m m e n t: In Spain, GAMARRA & OUTERELO (2005) indicate this species only from Madrid.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA8FF8BF7D8FB80EACE7880.taxon	description	C o m m e n t: Amischa forcipata is not listed for Spain by SMETANA (2004), but it was reported from some Spanish provinces by GAMARRA & OUTERELO (2005).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA8FF8BF7D8FD31EC3B7900.taxon	description	C o m m e n t: SMETANA (2004) does not list this widespread species for Spain, but GAMARRA & OUTERELO (2005) report it (as Atheta, subgenus Philhygra [sic]) from one mainland province and the Baleares. The above specimen from Algarve represents the first record from Portugal.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA8FF8BF7D8FE22EC8E7FB1.taxon	description	C o m m e n t: According to SMETANA (2004), this widespread European species was previously unknown from Spain.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA8FF84F7D8F92DED3F7DD1.taxon	description	C o m m e n t: This species is not listed for Spain by SMETANA (2004). According to GAMARRA & OUTERELO (2005), however, it has been recorded from the northeast. 1113	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA7FF84F7D8FEDCEDE07CD3.taxon	description	C o m m e n t: This species is not listed for Spain by SMETANA (2004). GAMARRA & OUTERELO (2005), however, report it from several provinces.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA7FF84F7D8FD27EB3E7EAB.taxon	description	C o m m e n t: According to SMETANA (2004), this species was previously known only from Spain and Morocco. The above specimens represent the first record from Portugal.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA7FF84F7D8FDD3EB0C7FDF.taxon	description	C o m m e n t: GAMARRA & OUTERELO (2005) report A. pruinosa, which is not listed for Spain by SMETANA (2004), from two mainland provinces and from the Baleares.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA7FF84F7D8FC3BED7979DB.taxon	description	C o m m e n t: This rare species was listed for Spain by SMETANA (2004), but omitted in the catalogue by GAMARRA & OUTERELO (2005).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA7FF84F7D8FB28EF557B11.taxon	description	C o m m e n t: The above specimens represent the first records from Spain and Portugal.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA7FF85F7D8F991EA0D7F77.taxon	description	1114 1500 m, sifted, 16. V. 1994, leg. Schülke & Grünberg (cAss); 1 ex., Corsica, SE Corte, 200 - 900 m, car-net, 8. IV. 1990, leg. Assing (cAss). Italy: 1 ex., Lombardia, Bergamo, M. Alben, 1500 m, 12. VII. 1991, leg. Starke (cAss); 1 ex., Veneto, Monte Grappa, 1700 m, Alnus and Rhododendron litter sifted, 27. VI. 1995, leg. Assing (cAss); 1 ex., Basilicata, Accettura, 950 m, 14. V. 1989, leg. Angelini (cAss); 8 exs., Sicilia, Ficuzza, 700 m, sifted, 1. - 4. V. 2000, leg. Angelini (cAss); 16 exs., Sicilia, Ficuzza, Alpe Ramosa, 800 m, sifted, 3. V. 2000, leg. Angelini (cAss). Morocco: 11 exs., Ar Rif, Chefchaouen region, Bad Bessen env., 34 ° 59 ' N, 4 ° 51 ' W, 1120 m, 3. VI. 2007, leg. Hlaváþ (cAss). C o m m e n t: This species is widespread in the Western Mediterranean, but had not not been reported from Morocco (SMETANA 2004). In Spain, A. reyi was previously known only from the northeast (GAMARRA & OUTERELO 2005).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA6FF85F7D8FC4EEC067940.taxon	description	C o m m e n t: GAMARRA & OUTERELO (2005) recently reported A. cinnamoptera from Spain for the first time.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA6FF85F7D8FD73ED467E7A.taxon	description	C o m m e n t: This species is not listed for Spain by SMETANA (2004), but GAMARRA & OUTERELO (2005) report it from one province.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA6FF85F7D8FB44ECCC7B0D.taxon	description	C o m m e n t: SMETANA (2004) reports A. orcina only from France and Switzerland. According to GAMARRA & OUTERELO (2005), however, it has been recorded from at least two provinces in northern Spain.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA6FF86F7D8F998ED3F7DD1.taxon	description	C o m m e n t: GAMARRA & OUTERELO (2005) reported this species from Zaragoza. According to GAMARRA & OUTERELO (2009), however, this record was erroneous and no specified localities of A. hybrida were known from Spain. 1115	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA5FF86F7D8FEDCECB379EA.taxon	description	C o m m e n t: This species was originally attributed to Acrotona THOMSON 1859, at that time a subgenus of Atheta THOMSON 1858 (BRUNDIN 1952). According to SMETANA (2004), it was subsequently moved to the Atheta subgenus Paradilacra BERNHAUER 1909. Based on a study of type material, however, GUSAROV (2003) discovered that Atheta densissima BERNHAUER 1909, the type species of Paradilacra, in fact refers to the subtribe Tachyusina, which today is doubtfully placed in the Oxypodini. A comparison of the sexual characters of Acrotona oxypodoides with the illustrations of Atheta densissima provided by GUSAROV (2003) leaves no doubt that the two species are not congeneric. Like its close relative, Acrotona pellucida (FAUVEL 1878), A. oxypodoides clearly belongs to the Athetini, with which it shares an athetal bridge of the aedeagus (absent in A. densissima) and numerous other characters. Although the sexual characters suggest that A. oxypodoides and A. pellucida may not be congeneric with the type species of Acrotona either, both species are moved from Paradilacra and tentatively placed in Acrotona again. Originally described from Tunisia, A. oxypodoides has subsequently been recorded also from Italy, France, Morocco, and Algeria (SMETANA 2004). The above specimens from Spain represent new country records.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA5FF86F7D8F993EC757A1E.taxon	description	C o m m e n t: In Spain, L. strumosus has been recorded only from Asturias (GAMARRA & OUTERELO 2005).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA5FF86F7D8FAFBEC047B12.taxon	description	C o m m e n t: This rare species has been reported from Spain (MARUYAMA 2006, SMETANA 2004); according to GAMARRA & OUTERELO (2005), however, specified localities were unknown.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA5FF87F7D8F8E6EC757C51.taxon	description	1116 C o m m e n t: In Spain, this rare species was previously known only from Madrid (GAMARRA & OUTERELO 2005).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA4FF87F7D8FDB8ED267991.taxon	description	C o m m e n t: The distribution of this rare myrmecophile is confined to Spain and Morocco; for an additional recent record from southern Spain see ASSING (2008). The above specimens were exclusively found associated with a relatively large blackish Aphaenogaster sp. (Formicidae: Myrmicinae).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA4FF87F7D8FB16ED0E78A2.taxon	description	C o m m e n t: This species is widespread in Europe and North Africa, but was previously unknown from the Iberian Peninsula.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA4FF87F7D8FEAFEDC97F23.taxon	description	C o m m e n t: In Spain, this extremely rare Western Mediterranean species had been known only from Madrid (GAMARRA & OUTERELO 2005).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA4FF80F7D8FA23ED3F7DD1.taxon	description	C o m m e n t: The previously known distribution of H. villosula ranged from France and Great Britain eastwards to China (ASSING 2005, SMETANA 2004). The above specimens represent the first record from Spain, the Iberian Peninsula, and Azerbaijan. 1117	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA3FF80F7D8FEDCEA3A7F82.taxon	description	C o m m e n t: According to SMETANA (2004), O. amicta was previously known only from North Africa (Tunisia, Algeria, Morocco) and Italy (Sardinia, Sicily). The above specimens represent first records from Spain, Portugal, and France.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA3FF80F7D8FD03EC357EB3.taxon	description	C o m m e n t: The known distribution of this recently described species is confined to Andalucía, southern Spain.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA3FF82F7D8FC33EFAB7822.taxon	description	D e s c r i p t i o n: Body length 3.0 mm. Habitus as in Fig. 9. Coloration: body probably reddish, with abdominal segment VI infuscate; legs and antennae reddish-yellow. (The holotype is of somewhat dark-brownish coloration, but this is evidently a result of post-mortem darkening.) Head approximately as wide as long; dorsal surface matt due to extremely dense and confluent punctation (Fig. 10). Eyes large (Fig. 11) and bulging, approximately twice as long as postocular portion in dorsal view. Antennae of moderate length and weakly incrassate apically; antennomere I approximately 1.5 times as long as broad; II about as long as I and twice as long as broad; III slightly less than twice as long as broad and slightly shorter than II; IV-X weakly transverse and gradually, but weakly increasing in width; X less than 1.5 times as wide as long; XI slightly longer than the combined length of IX and X. Maxillary palpus slender, preapical palpomere approximately 3.5 times as long as broad. Pronotum 1.45 times as wide as long and 1.53 times as wide as head, maximal width approximately in the middle; posterior margin weakly sinuate between middle and posterior angles; dorsal surface matt due to very dense and confluent punctation (Fig. 10). Elytra slightly narrower than, and 0.75 times as long as pronotum; laterally weakly, but extensively impressed; lateral margin sharply carinate, this carina straight, distinctly elevated, and not depressed in the middle; posterior margin with small and not very deep concavity near posterior angles; punctation similar to that of pronotum (Fig. 10). Hind wings reduced to short rudiments (slightly protruding beyond posterior margin of elytra). Legs slender, metatarsus almost as long as metatibia, metatarsomere I almost as long as the combined length of II-IV. 1118 1119 Abdomen widest at base and gradually tapering posteriad (Fig. 9); tergites III and IV matt due to very dense and coarse punctation; punctation of tergites V-VII increasingly fine and sparse; posterior margin of tergite VII without palisade fringe.: posterior margin of sternite VIII convex, not pointed in the middle; median lobe of aedeagus as in Figs 12 - 13, apex of ventral process bifid, deeply incised in ventral view (Figs 13 - 14); apical lobe of paramere moderately slender (Fig. 15).: unknown. E t y m o l o g y: The species is dedicated to José Luis Lencina (Jumilla, Murcia), who collected the holotype, also in gratitude for the generous gift of numerous Staphylinidae collected with various trapping methods in Spain. C o m p a r a t i v e n o t e s: Based on external and the male sexual characters (habitus, punctation, shape of legs, antennae, and maxillary palpi, large bulging eyes, the depressed and laterally carinate elytra, morphology of the median lobe of the aedeagus, long and slender apical lobe of the paramere), O. lencinai is most closely allied to O. magnicollis FAUVEL 1878 and O. arabs FAUVEL 1904. It is distinguished from both by the shape of the aedeagus, from the former additionally by the darker coloration, the much more transverse pronotum, the shorter antennae, and the straight and more elevated lateral carinae of the elytra. It is separated from O. arabs particularly by much shorter antennae, distinctly shorter, flattened and extensively impressed elytra with pronounced lateral carinae, the shape of the median lobe of the aedeagus, and the less slender apical lobe of the paramere. Oxypoda (Deropoda) andalusiaca ASSING 2003, whose distribution too is confined to southern Spain and which has reduced hind wings, has a much more slender body, distinctly smaller eyes, a weakly transverse pronotum, and longer elytra without impressions and without lateral carinae. For illustrations of O. arabs and O. andalusiaca see ZERCHE (1996) and ASSING (2003 a), respectively. D i s t r i b u t i o n a n d b i o n o m i c s: Thetypelocality is situated in Murcia, southeastern Spain, at an altitude of only 400 m. The reduced wings and the absence of previous records suggest the species has a restricted distribution and may live in a subterranean habitat.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA1FF83F7D8FAA3ED247B9E.taxon	description	D e s c r i p t i o n: Body length 4.8 - 5.3 mm. Habitus conspicuously slender (Fig. 16). Coloration: head dark-brown to blackish-brown; pronotum reddish; elytra reddishyellow; abdomen brown to dark-brown, with the posterior margins of segments III-VI, the posterior third of segment VII, and segments VIII-X reddish; legs dark-yellowish; antennae reddish. Head of suborbicular shape, approximately as wide as long; dorsal surface almost matt due to pronounced microreticulation; punctation sparse and extremely fine, barely noticeable in the microsculpture (Fig. 17). Eyes moderately large and moderately bulging, approximately as long as postocular portion in dorsal view. Antennae long, slender, but rather massive (similar to that of some species of Derocala MULSANT & REY), 1120 weakly incrassate apically; antennomere III slightly longer than II and of distinctly conical shape; IV-X very weakly transverse; XI longer than the combined length of IX and X (Fig. 18). Maxillary palpus conspicuously slender, preapical palpomere approximately 4 times as long as broad. Pronotum slender, only 1.15 - 1.20 times as broad as long and 1.40 - 1.45 times as broad as head, maximal width in or near the middle (Fig. 17); posterior angles obtusely marked; punctation dense, fine, and shallow, but somewhat more distinct than that of head; interstices with pronounced microreticulation and almost matt. Elytra 0.85 - 0.95 times as long as pronotum (Fig. 17); suture posteriorly gaping; posterior margin moderately sinuate near posterior angles; punctation dense, weakly granulose, but not very coarse, much more distinct than that of pronotum; interstices more shiny than those of pronotum. Hind wings present, but of reduced length, extending approximately to posterior margin of abdominal segment IV when unfolded. Legs long and slender; metatarsus approximately 0.9 times as long as metatibia; metatarsomere I approximately as long as the combined length of II-IV. Abdomen with segments III-IV or III-V of subequal width, segments V or VI and following tapering posteriad; punctation of all tergites very fine and very dense; posterior margin of tergite VII with palisade fringe.: unknown.: sternite VIII broadly convex and with long marginal setae; spermatheca as in Fig. 19. E t y m o l o g y: The name is the past participle of the Latin verb defodere (to dig in, to bury) and alludes to the presumably subterranean habitat of this species. C o m m e n t: In general, a description of Oxypoda species based on female alone is not advisable, since many species are reliably identified only based on the male sexual characters. In view of the highly distinctive external morphology of O. defossa, however, an exception from this rule seems justifiable. C o m p a r a t i v e n o t e s: This species is readily distinguished from all other Oxypoda species known from the Western Mediterranean based on external characters alone, above all its conspicuously slender habitus, the long and slender antennae and maxillary palpi, the weakly transverse pronotum, and relatively large size. D i s t r i b u t i o n a n d b i o n o m i c s: Thespeciesis known from three localities in the vicinity of Tarifa, Cádiz, in the extreme south of Spain. As can be inferred from the reduced hind wings, the distribution may be restricted. The type specimens were collected on pastures under stones after very heavy rainfall. This, as well as the slender body with long antennae and legs, and the absence of previous records suggest that O. defossa lives in a cryptic subterranean habitat.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFA0FF9DF7D8F966EC247FC2.taxon	description	D e s c r i p t i o n: Very small species, body length 1.9 mm. Habitus subparallel (Fig. 20). Coloration: head and abdomen dark-brown; pronotum and elytra brown; legs yellowish; antennae dark-brown, with the basal 3 antennomeres yellowish. 1121 Head of suborbicular shape (Fig. 21), approximately as wide as long; punctation dense and extremely fine, barely noticeable; interstices with shallow microsculpture. Eyes weakly convex, not distinctly projecting from lateral contours of head, and moderately large, approximately as long as postocular portion from posterior margin of eye to posterior carina in dorsal view. Antennae 0.55 mm long, distinctly incrassate apically; antennomere III weakly oblong; IV distinctly transverse; IV-X gradually increasing in width; X approxiamtely twice as wide as long; XI of ovoid shape, barely as long as combined length of IX and X (Fig. 22). Pronotum small in relation to head, 1.28 times as wide as long and 1.18 times as wide as head, widest approximately in the middle (Fig. 21); posterior angles weakly marked, almost obsolete; punctation and microsculpture similar to those of head. Elytra approximately 0.95 times as long and 1.1 times as wide as pronotum (Fig. 21); punctation fine, but slightly more distinct than that of pronotum; interstices with shallow microsculpture. Legs short; metatarsomere I approximately twice as long as broad and about as long as the combined length of metatarsomeres II and III. Abdomen approximately as wide as elytra, widest at segment VI; punctation fine, dense on anterior tergites and somewhat sparser on posterior tergites; interstices with distinct microsculpture; posterior margin of tergite VII with palisade fringe.: posterior margin of tergite VIII weakly convex; sternite VIII with strongly convex posterior margin; median lobe of aedeagus 0.27 mm long, shaped as in Figs 23; apical lobe of paramere moderately long (Fig. 24).: unknown. 1122 E t y m o l o g y: The specific epithet is the past participle of the Latin verb deminuare (to diminish, to reduce) and refers to the conspicuously small size of the species. C o m p a r a t i v e n o t e s: This species is readily distinguished from all other Cousya species known from the Western Mediterranean by its small size alone. In addition, it is characterized by its parallel habitus, the fine punctation, and by the shape of the aedeagus. According to SMETANA (2004), two species of Cousya MULSANT & REY 1875 have been reported from Spain, C. nigrata (FAIRMAIRE & LABOULBÈNE 1856) and C. nitidiventris FAGEL 1958. The types of the latter have not been examined, but it appears likely that they may be conspecific with C. nigrata. D i s t r i b u t i o n a n d b i o n o m i c s: The type locality is situated in the Sierra del Carche near Jumilla, Murcia, southeastern Spain. The holotype was collected with a pitfall trap in June.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFBEFF9DF7D8FB52EA547865.taxon	description	C o m m e n t: According to TRONQUET (2009), this rare species is probably myrmecophilous and records from southern Spain and Morocco were revised. Additional literature records have become known from Algeria, Gibraltar, and Italy.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFBEFF9DF7D8FA62EB327BFA.taxon	description	C o m m e n t: Aleochara funebris is widespread and not uncommon in the Palaearctic and Oriental regions. It has been recorded from the Canary Islands, but was previously unknown from mainland Spain and the whole of the Iberian Peninsula (SMETANA 2004).	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFBEFF9DF7D8FCC3EFC57952.taxon	description	C o m m e n t: The previously known distribution of S. semirufa ranges from Turkey across the Balkans and Central Europe to Great Britain and France (ASSING in press, SMETANA 2004). The above specimens represent the first records from Spain and the Iberian Peninsula.	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
03961979FFBEFF9FF7D8F8CAED887B3A.taxon	description	1123 Acknowledgements I am most grateful to the colleagues mentioned in the material section for the loan of material, in particular also to José Luis Lencina (Jumilla) for the gift of Staphylinidae collected mostly with flight interception traps in Spain and to Alexey Shavrin for the generous permission to retain the holotype of Astenus shavrini. Jürgen Vogel (Görlitz) helped with the identification of Atheta orcina and A. lindbergiella, Marc Tronquet (Molitg-les-Bains) with that of Aleochara rudella. Raimundo Outerelo (Madrid) made me aware of the erroneous record of Aleochara ruficornis from Spain. Benedikt Feldmann (Münster) proof-read an earlier version of the manuscript. Zusammenfassung	en	Assing, V. (2010): Four new species and additional records of Staphylinidae from Spain, primarily from the south (Insecta: Coleoptera). Linzer biologische Beiträge 42 (2): 1105-1124, DOI: 10.5281/zenodo.10106018
