identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03966709C8367E50FC2FFAD2EA33FD24.text	03966709C8367E50FC2FFAD2EA33FD24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella Muller 1879	<div><p>GRUMICHELLA MÜLLER, 1879</p> <p>Grumichella Müller, 1879: 407 (Type species: Grumichella rostrata Thienemann, 1905, first includ- ed species). – Holzenthal, 1988a:88 [male, female, larva, pupa, revision, phylogeny, distribution]. – Flint, Holzenthal &amp; Harris, 1999: 128 [catalog]. – Morse, 1981: 259 [classification, phylogeny]. – Morse &amp; Holzenthal, 1987: 140 [classification, phylogeny]. – Calor &amp; Holzenthal, 2008: 255 [phylogeny]. – Malm &amp; Johanson, 2011: 3, 7 [classification, phylogeny].</p> <p>Leptocellodes Ulmer, 1911: 21 [Type species: Leptocellodes flaveola Ulmer, 1911, original designation (the designation of G. flaveola Ulmer, 1911 as type species by Ulmer (1955) is not valid (ICZN 1985, Art. 69(a) sensu Holzenthal, 1988a)]; – Ulmer, 1955: 499 [to synonymy].</p> <p>Grumichella Müller, 1879 (= Leptocellodes Ulmer, 1955) comprises 13 species, including the nine new species described here: G. aequiunguis Flint, 1983; G. blahniki sp. nov.; G. boraceia sp. nov.; G. cressae sp. nov.; G. flaveola (Ulmer) Holzenthal, 1988a, G. jureia sp. nov.; G. leccii sp. nov.; G. muelleri sp. nov.; G. paprockii sp. nov.; G. parati sp. nov.; G. pulchella (Banks) Holzenthal, 1988a; G. rostrata Thienemann, 1905; G. trujilloi sp. nov.</p> <p>Müller (1879a, b, 1880a, b, 1888, 1921) illustrated the larvae of an unknown species and discussed its biology. Pupae and cases of two species, G. rostrata and G. aequiunguis, were described by Thienemann (1905). Roback (1966) described and illustrated the larvae of G. flaveola from Peru, but included it under ‘unknown family 2’. Ulmer (1955) also described immature forms of the genus. Morse (1981) erected the tribe Grumichellini to accommodate Grumichella and Atanatolica in his family classification. Holzenthal (1988a) reviewed the genus, redescribed the four valid species, and described the larvae and pupae, including those of eight probable unknown species of larvae (Holzenthal’s ‘species A–H’). He also corroborated the monophyly of Grumichella based on 20 synapomorphies and proposed the phylogenetic relationships among two species groups: G. pulchella and G. rostrata groups. The first species group, composed of G. flaveola and G. pulchella, is characterized by nine spots on the forewings. The G. rostrata group, comprising G. aequiunguis and G. rostrata, bears only two spots on forewings. Holzenthal &amp; Pes (2004), in their description of Amazonatolica, another Grumichellini genus, discussed other characters and the phylogenetic position of Grumichella. Calor &amp; Holzenthal (2008) described another related genus, Osflintia, and proposed a phylogeny of Grumichellini, placing Osflintia as sistergenus of all other grumichelline genera.</p> <p>Biological remarks: Müller (1879a, b) provided the first records of the habitat of larvae, including their occurrence in small waterfalls and rock surfaces in fast flowing waters of small mountain streams. These observations were corroborated by others authors (e.g. Holzenthal, 1988a; Flint et al., 1999). In our fieldwork, the larvae (and pupae) were often collected from aggregations of more than 50 individuals positioned side by side or in a small clump (e.g. Parque Estadual de Campos do Jordão, São Paulo State, Brazil). Müller (1879a, b) also observed the use of the posterior silken projection on the larval cases to help to hold the larva in the current. The ability of larvae to maintain position in fast flow is aided by the stout legs with modified tarsal claws, as emphasized by Holzenthal (1988a) and by other authors observing larvae of other grumichelline genera (St. Clair, 1994; Ward, 2001). Müller (1879a, b) ‘speculated’ that the water current prevented the pupae from crawling out of the case, and that the anterior attachment pedicle of the case needed to be severed by the emerging pupae. The loose case then drifted to slack current, where the adult could emerge (Holzenthal, 1988a). In laboratory rearings and rarely in natural habitats (e.g. G. boraceia sp. nov.), a behaviour of flotation and drift was observed in the fifth instar larvae, with air bubbles inside the anterior portions of cases (our pers. observ.).</p> <p>The use of abandoned larval cases by other genera of caddisflies is not uncommon, especially in Triplectides (Holzenthal, 1988b; Flint et al., 1999; Crisci-Bispo, Bispo &amp; Froehlich, 2004; Calor &amp; Froehlich, 2008), and includes observations by A.R.C. and by P.A. Rueda- Martín (pers. commun.) in Brazil and Argentina, respectively, of Marilia (Odontoceridae) occupying Grumichella cases.</p> </div>	https://treatment.plazi.org/id/03966709C8367E50FC2FFAD2EA33FD24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C8377E50FCCCFD24EB84F8CC.text	03966709C8377E50FCCCFD24EB84F8CC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella aequiunguis Flint 1983	<div><p>GRUMICHELLA AEQUIUNGUIS FLINT, 1983 (FIG. 2)</p> <p>Grumichella aequiunguis Flint, 1983: 68, figs 246– 248 [Type locality: Argentina: Misiones, Arroyo Piray Mini, rt. 17, W Dos Hermanas, 23.xi.1973, O. S. Flint, Jr, type 100540; NMNH; male]. – Holzenthal, 1988a:91 [male, female, larva, pupa, case, distribution]. – Flint, Holzenthal &amp; Harris, 1999: 128 [catalog]. – Blahnik, Paprocki &amp; Holzenthal, 2004: 5 [distribution]. – Paprocki, Holzenthal &amp; Blahnik, 2004: 12 [distribution]. – Dumas et al., 2009: 368 [distribution]. – Paprocki &amp; França, 2014: 57 [distribution].</p> <p>Diagnosis: Grumichella aequiunguis can be diagnosed by the following characters: 2 brown spots on male forewing; male genitalia bearing long, wide, rhomboid 2nd article of inferior appendage, with an excavate mesal surface, subequal in length to apicodorsal portion of 1st article; larval abdominal sternum I with 25–30 long setae. It is most similar to G. boraceia sp. nov., but differs in having a pointed rather than blunt apex of the 2nd article of the inferior appendage, which is not excavate.</p> <p>Adult: Head and body yellowish.</p> <p>Male: Forewing length: 8–9 mm. Forewing light golden brown, with 2 small brown spots, 1 at midlength, 1 at apex of anal area (arculus). Abdominal segment IX annular; in dorsal view, with anterior margin nearly straight, posterior margin broadly rounded; pleural region sparsely setose; tergum IX without protuberances dorsally. Preanal appendage setose, at least 4× as long as wide; in lateral view, clavate, apex rounded. Segment X saddle-shaped; in lateral view not flattened, dorsal margin slightly concave, ventral margin abruptly convex; segment X in dorsal view, rectangular, with posterolateral borders well developed, lateral margins subparallel, with apicodorsal shelf, shelf almost rectangular, 1/3 as wide as apex of segment X, without</p></div> 	https://treatment.plazi.org/id/03966709C8377E50FCCCFD24EB84F8CC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C8317E55FF2DFD24EBA8F8CF.text	03966709C8317E55FF2DFD24EBA8F8CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella flaveola	<div><p>GRUMICHELLA FLAVEOLA (ULMER, 1911) (FIGS 3, 4)</p> <p>Leptocellodes flaveola Ulmer, 1911: 22 [Type locality: Bolivia: Yungas, Bogota; ZMHU; male]. – Jörgensen, 1919: 397 [male]. – Mosely, 1949: 41 [redescription, male]. – Fisher, 1966: 9 [catalog]. – Holzenthal, 1988a:91 [male, female, larva, pupa, case, distribution]. – Flint, Holzenthal &amp; Harris, 1999: 128 [catalog]. – Munõz- Quesada, 2000: 279 [distribution].</p> <p>Notanatolica poujadei Navás, 1927: 73 [Type locality: Ecuador, Loja; MNHNP; male]. – Mosely, 1936: 107 [to synonym]. – Fisher, 1966: 9 [catalogue].</p> <p>Unknown family 2, Roback, 1966: 256, 303, figs 254– 266 [larva].</p> <p>Grumichella flaveola (Ulmer). – Holzenthal, 1988a: 91 [new combination, male, female, larva, pupa, case, distribution]. – Flint, 1991: 93 [male]; – Flint, 1996: 417 [distribution]. – Flint, Holzenthal &amp; Harris, 1999: 128 [catalogue].</p> <p>Diagnosis: Grumichella flaveola can be diagnosed by the following characters: male forewings with 9 brown spots; male genitalia with 1st article of inferior appendage with small basomesal protuberance; 2nd article 1/2 to 1/3 of length of apicodorsal portion of 1st article; apex of 2nd article acuminate, curved mesal; larval abdominal sternum I with 2 pairs of short setae. It is most similar to G. pulchella, except for the quadrate apicodorsal portion of 1st article of the inferior appendage.</p> <p>Adult: Head and body yellowish.</p> <p>Male: Forewing length: 11–12 mm (N = 5). Forewing light golden brown, with 9 small brown spots (Fig. 3). Abdominal segment IX annular; in dorsal view, with anterior margin nearly straight, posterior margin broadly rounded; in lateral view, pleural region setose; tergum IX without protuberances dorsally. Preanal appendage setose, at least 4× as long as wide; in lateral view, clavate, apex rounded. Segment X saddle-shaped; in lateral view not flattened, dorsal margin abruptly concave at 2/3 length, with apex upcurved, ventral margin slightly convex; segment X in dorsal view, subtriangular, with posterolateral borders not developed, lateral margins sinuate, narrowing apically, with apicodorsal shelf, shelf almost rectangular, 1/3 as wide as apex of segment X, without sulcus apically. Inferi- or appendage nearly straight in ventral view, with basal region tuboid, not enlarged; slightly rounded basally, setose, bearing a small basomesal protuberance; middle region of inferior appendage without constriction before apicodorsal region; 2nd article shorter than apicodorsal portion of 1st article, broad basally, tapering throughout length, mesal surface not excavate, apex pointed, curved mesally; apicordorsal portion of 1st article digitate, setose. Phallic apparatus simple, phallicata sharply down-curved; with pair of subequal, wide, posterodorsally directed lateral flanges, apices not acuminate; phallotremal sclerites small, U-shaped in dorsal view.</p> <p>Female: Head and body yellowish. Forewing length: 7–8 mm. Forewing yellowish to brown, without dark setae on posterodorsal region. Genitalia inseparable from congeners (Fig. 4). Abdominal segment IX lightly sclerotized, with dorsal plate, bearing pair of setose protuberances above appendages of segment X, appendages of segment X small; valves flat, thin, lightly sclerotized, with very short setae.</p> <p>Larva: Head light brown, thoracic sclerites light brown, legs yellowish to brownish, with dorsal borders of middle and hind femora black. Length of larva 8 mm. Abdominal sternum I with 2 pairs of short, thin setae; lateral hump sclerite very small.</p> <p>Larval case: Length 9 mm; constructed of alternating rings of clear and dark brown silk, nearly straight, tapered, wide, with 2 distinct diagonal sutures, 1 towards posterior end and 1 more or less at midlength, posterior end with ventral, dorsally directed projection, visible externally, posterior aperture C-shaped.</p> <p>Pupa: Segment IX slender; anal process short, roughly trapezoidal in general shape, tapering posteriorly to narrow apex, with basodorsal, pigmented area shaped like a parallelogram, apically upturned, hooked.</p> <p>Pupal case: With anteroventral silken pedicel, pedicel short; anterior silken membrane with 3 small openings arranged in a triangle.</p> <p>Distribution: Argentina, Bolivia, Colombia, Ecuador, Peru, and Venezuela.</p> <p>Material examined: ARGENTINA: Salta: Rt. 59, Km 23.5, Cañada La Gotera, 16–17.x.1973, O.S. Flint Jr, 34 males, 6 females (NMNH). BOLIVIA: La Paz: Rio Coroico, el. 1200 m, 23–26.ix.1984, L.E. Pena G., 5 males (NMNH). COLOMBIA: Antioquia: Rio Medellin, 6 km,</p> <p>S. Caldas, 24.ii.1984, C.M. &amp; O.S. Flint Jr; 1 male, 12 females (NMNH). ECUADOR: Pastaza: Mera, el. 1300 m, xii.1992, V.O. Becker col., 1 female, det. O.S. Flint Jr (NMNH); Puyo (12 km W), 9.v.1977, P.J. Spangler &amp; D.R. Givens, 2 males (NMNH). PERU:</p> </div>	https://treatment.plazi.org/id/03966709C8317E55FF2DFD24EBA8F8CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C8337E54FF24F9C1EB84F8CC.text	03966709C8337E54FF24F9C1EB84F8CC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella pulchella (Banks 1910) Holzenthal 1988	<div><p>GRUMICHELLA PULCHELLA (BANKS, 1910) (FIG. 5)</p> <p>Leptocella pulchella Banks, 1910: 160 [Type locality: Colombia: Tolima, Canõn de Monte, Dec., type 11783; MCZ; male]. – Fisher, 1966: 60 [catalogue].</p> <p>Leptocellodes pulchellus Ulmer, 1955: 499 [to synonym]. – Flint, 1967: 22 [male].</p> <p>Grumichella pulchella Holzenthal, 1988a:93 [new combination]. – Flint, Holzenthal &amp; Harris, 1999: 128 [catalogue]. – Munõz-Quesada, 2000: 279 [distribution].</p> <p>Diagnosis: Grumichella pulchella can be diagnosed by the following characters: male forewings with 9 brown spots; male genitalia bearing apicodorsal portion of 1st article of inferior appendage quadrate; segment X acuminated, dorsal view. As discussed before, it is most similar to G. flaveola, but differs in the apicodorsal portion of the 1st article of the inferior appendage, which is not digitate.</p> <p>Adult: Head and body yellowish.</p> <p>Male: Forewing length: 11–12 mm. Forewing light golden brown, with 9 small brown spots. Abdominal segment IX annular; in dorsal view, with anterior margin straight, posterior margin broadly rounded; in lateral view, pleural region setose; tergum IX without dorsal</p></div> 	https://treatment.plazi.org/id/03966709C8337E54FF24F9C1EB84F8CC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C83C7E59FF0DF96EEC46FED6.text	03966709C83C7E59FF0DF96EEC46FED6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella rostrata Thienemann 1905	<div><p>GRUMICHELLA ROSTRATA THIENEMANN, 1905 (FIG. 6)</p> <p>Grumichella rostrata Thienemann, 1905: 49 (Type locality: no type nor type depository designated, but name based on material probably from Brazil, Santa Catarina, Gruta dos Macacos, near Blumenau according to Holzenthal, 1986; case, pupa). – Thienemann, 1909: 41, 42, 125 [larva, pupa]. – Fisher, 1966: 10 [catalogue]. – Holzenthal, 1988a:93–95 [male, female, larva, case, pupa, distribution]. – Flint, Holzenthal &amp; Harris, 1999: 128 [catalogue]. – Paprocki, Holzenthal &amp; Blahnik, 2004: 12 [distribution]. – Calor, 2011: 322 [distribution]. – Paprocki &amp; França, 2014: 57 [distribution].</p> <p>Diagnosis: Grumichella rostrata can be diagnosed by the following characters: male forewing with 2 or 3 brown spots; male genitalia bearing segment X flattened, lateral view; 2nd article of inferior appendage subequal to apicodorsal portion of 1st article; apex of 2nd article acuminated, curved mesad. Grumichella rostrata is most similar to G. lecci sp. nov., but differs in having segment X without sulcus apically, and the subequal lengths of the 2nd article of inferior appendage and the apicodorsal portion of the 1st article.</p> <p>Adult: Head and body yellowish.</p> <p>Male: Forewing length: 8–11 mm. Forewing light golden brown, with 2 or 3 small brown spots, bearing dark setae on posterodorsal margin. Abdominal segment IX annular; in dorsal view, with anterior margin slightly convex, posterior margin nearly straight; in lateral view, pleural region sparsely setose; tergum IX without protuberances dorsally. Preanal appendage setose, at least 4× as long as wide; in lateral view, clavate, apex subacute. Segment X saddle-shaped; in lateral view flattened, dorsal margin abruptly concave at 2/3 length, with apex upcurved, ventral margin abruptly convex; segment X in dorsal view, rectangular, with posterolateral borders developed, lateral margins sinuate, with apicodorsal shelf, shelf rounded, 1/3 as wide as apex of segment X, without sulcus apically. Inferior appendage slightly concave in ventral view, with basal region tuboid, not enlarged; slightly rounded basally, setose, bearing a small basomesal protuberance; middle region of inferior appendage without constriction before apicodorsal region; 2nd article subequal in length to apicodorsal portion of 1st article, broad basally, tapering throughout length, mesal surface not excavate, apex pointed, curved mesally; apicordorsal portion of 1st article digitate, setose. Phallic apparatus simple, phallicata slightly down-curved; with pair of subequal, wide, posterodorsally directed lateral flanges, apices not acuminate; phallotremal sclerites small, U-shaped in dorsal view.</p> <p>Female: Head and body yellowish. Forewing length: 6–9 mm. Forewing yellowish to brown or dark brown, without dark setae on posterodorsal region. Genitalia inseparable from congeners. Abdominal segment IX lightly sclerotized, with dorsal plate, bearing pair of setose protuberances above appendages of segment X, appendages of segment X small; valves flat, thin, lightly sclerotized, with very short setae.</p> <p>Larva: Head dark brown, almost black, thoracic sclerites dark brown, almost black, legs dark brown, almost black. Length of larva 6–7 mm. Abdominal sternum I with 25–30 long setae; lateral hump sclerite very small.</p> <p>Larval case: Length 7–9 mm; constructed entirely of rings of dark brown silk, gently curved, tapered, wide, with 2 distinct diagonal sutures, 1 towards posterior end and 1 more or less at midlength, posterior end with ventral, dorsally directed projection, visible externally, posterior aperture C-shaped.</p> <p>Pupa: Segment IX very slender; anal process long, narrow, slender, unpigmented or slightly yellowish, apically upturned, hooked.</p> <p>Pupal case: With anteroventral silken pedicel, pedicel very long, thin; anterior silken membrane with an eccentric, horizontal, slit-like opening.</p> <p>Distribution: Brazil.</p> <p>Material examined: BRAZIL: Bahia: Elisio Medrado, Serra da Jibóia, Faz. Jequitibá, GAMBA, Córrego Caranguejo, 12°52′127″S, 39°28′324″W, el. 510m, 28.iii.2012, Quinteiro F.B., Duarte T. &amp; Garcia I., 2 males (UFBA); Varzedo, Serra da Jibóia, Faz. Baixa da Areia Cai Camarão, 12°57′45″S, 39°27′12″W, Prop. Sr. Getulio, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-39.453335&amp;materialsCitation.latitude=-12.9625" title="Search Plazi for locations around (long -39.453335/lat -12.9625)">Córrego Cai Camarão</a>, 24.x.2012, Gomes V., Campos R. &amp; Vilarino A., 3 males (UFBA); Minas Gerais: Rio Tanque, c. 12 km (rd) from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.448055&amp;materialsCitation.latitude=-19.536667" title="Search Plazi for locations around (long -43.448055/lat -19.536667)">Ipoema</a>, 16.v.1998, UV light, Holzenthal &amp; Paprocki, 19°32′12″S, 43°26′53″W, el. 750 m, 1 male, 2 females (UMSP); Parque Estadual Itacolomi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.427223&amp;materialsCitation.latitude=-20.417221" title="Search Plazi for locations around (long -43.427223/lat -20.417221)">Rio Belchior</a>, 20.ii.1999, UV light, Paprocki &amp; Amarante, 20°25′02″S, 43°25′38″W, el. 725 m, 2 males (UMSP); Córrego Pitanga, upstream of confluence with <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.665&amp;materialsCitation.latitude=-19.094444" title="Search Plazi for locations around (long -42.665/lat -19.094444)">Rio Santo Antônio</a>, 19.x.2000, UV light, Paprocki &amp; Ferreira, 19°05′40″S, 42°39′54″W, el. 238 m, 3 males, 1 female (UMSP); Parque Estadual do Ibitipoca, Corrego dos <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.893333&amp;materialsCitation.latitude=-21.709167" title="Search Plazi for locations around (long -44.893333/lat -21.709167)">Macacos</a>, 19.ix.2001, UV light, Holzenthal, Blahnik, Neto &amp; Paprocki, 21°42′33″S, 44°53′36″W, el. 1360 m, 45 males, 142 females (UMSP); Córrego das <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.637222&amp;materialsCitation.latitude=-22.061666" title="Search Plazi for locations around (long -44.637222/lat -22.061666)">Aguas Pretas</a> &amp; tribs., c. 15 km S Aiuruoca, 21.xi.2001, UV light, Holzenthal, Blahnik, Neto &amp; Paprocki, 22°03′42″S, 44°38′14″W, el. 1386 m, 5 males (UMSP); Rio de Janeiro: Parque Nacional do Itatiaia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.61361&amp;materialsCitation.latitude=-22.450556" title="Search Plazi for locations around (long -44.61361/lat -22.450556)">Rio Campo Belo</a>, 23.xi.2001, UV light, Holzenthal, Blahnik, Neto &amp; Paprocki, 22°27′02″S, 44°36′49″W, el. 1300 m, 1 male, 1 female (UMSP); Parque Nacional do Itatiaia, Lago Azul, 09.vii.1965, Froehlich C.G., 8 larvae (MZSP); Parque Nacional do Itatiaia, Rio Campo Belo, trail to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.619442&amp;materialsCitation.latitude=-22.428333" title="Search Plazi for locations around (long -44.619442/lat -22.428333)">Veu da Noiva</a>, 24.xi.2001, UV light, Holzenthal, Blahnik, Neto &amp; Paprocki, 22°25′42″S, 044°37′10″W, el. 1310 m, 2 males (UMSP); Santa Catarina: Nova Teutonia, 27°11′S, 52°23′S, 29.x.1939, Fritz Plauman, 1 male (NMNH); São Paulo: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.582222&amp;materialsCitation.latitude=-22.762222" title="Search Plazi for locations around (long -45.582222/lat -22.762222)">Campos do Jordão</a>, 28.i.1959 (Hotel Umuarama, riacho a jusante do lago, 22°45′44″S, 45°34′56″W, el. 1700 m), Froehliuch C.G., 1 pupa (MZUSP); Parque Estadual de Campos do Jordão, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.463055&amp;materialsCitation.latitude=-22.694443" title="Search Plazi for locations around (long -45.463055/lat -22.694443)">Rio Galharada</a>, 4.iii.1996, UV light &amp; sweeping, Holzenthal &amp; Guahyba, 22°41′40″S, 45°27′47″W, el. 1530 m, 1 larva, 3 females (UMSP); Parque Estadual de Campos do Jordão, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.490833&amp;materialsCitation.latitude=-22.695833" title="Search Plazi for locations around (long -45.490833/lat -22.695833)">Campo do Meio</a>, 6.iii.1996, UV light, Holzenthal &amp; Guahyba, 22°41′45″S, 45°29′27″W, el. 1500 m, 1 pupa (UMSP); Parque Estadualde Campos do Jordão, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.463055&amp;materialsCitation.latitude=-22.694443" title="Search Plazi for locations around (long -45.463055/lat -22.694443)">Rio Galharada</a>, 22.i.1998, UV light, Holzenthal, Froehlich &amp; Paprocki, 22°41′40″S, 45°27′47″W, el. 1530 m, 5 males, 9 females (UMSP); Parque Estadual de Campos do Jordão, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-45.461945&amp;materialsCitation.latitude=-22.695555" title="Search Plazi for locations around (long -45.461945/lat -22.695555)">Cachoeira Galharada</a>, 15.x.1998, UV light, Paprocki &amp; Froehlich, 22°41′44″S, 45°27′43″W, el. 1620 m, 3 males, 18 females (UMSP); Parque Estadual de Campos do Jordão, Rio Galharada, 10.ix1999, Froehlich C.G., 40 larvae (MZUSP); Cachoeira do Paredão, Lajeado, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.62111&amp;materialsCitation.latitude=-22.725555" title="Search Plazi for locations around (long -44.62111/lat -22.725555)">Serra da Bocaina</a>, 1.iii.2002, UV light, Holzenthal, Blahnik, Paprocki &amp; Prather, 22°43′32″S, 44°37′16″W, el. 1550 m, 17 males, 92 females (UMSP); Cachoeira do Príncipe, Lajeado, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.623608&amp;materialsCitation.latitude=-22.715279" title="Search Plazi for locations around (long -44.623608/lat -22.715279)">Serra da Bocaina</a>, 4.iii.2002, UV light, Holzenthal, Blahnik, Paprocki &amp; Prather, 22°42′55″S, 44°37′25″W, el. 1560 m, 1 female (UMSP); Parque Estadual de Campos do Jordão, córrego Campo do Meio, 6.vi.2005, Calor A.R., c. 100 larvae (UFBA); Parque Estadual de Campos do Jordão, Rio Galharada, acima da cachoeira, 28.iii.2007, luz UV/ branca, Calor A.R., Mariano R. &amp; Lecci L., 4 males, 2 females (UFBA); Parque Estadual de Campos doJordão, Rio Galharada, 31.iii.2007, luz UV/branca, Calor A.R., Mariano R. &amp; Lecci L., 1 larva (UFBA).</p> </div>	https://treatment.plazi.org/id/03966709C83C7E59FF0DF96EEC46FED6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C83E7E58FF35FE29EAEAFDF6.text	03966709C83E7E58FF35FE29EAEAFDF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella blahniki	<div><p>GRUMICHELLA BLAHNIKI CALOR &amp; HOLZENTHAL SP. NOV. (FIG. 7)</p> <p>Diagnosis: Grumichella blahniki can be diagnosed by the following characters: male forewings with 12 brown spots; male genitalia bearing segment X not flattened, lateral view; 1st article of inferior appendage with well-developed basomesal protuberance; 2nd article of inferior appendage shorter than the apicodorsal portion of 1st first article; apex of 2nd article pointed, curved mesad; Cu1a and M3+4 veins fused before the border in male hind wings. It is most similar to G. cressae sp. nov., except for having tergum IX without protuberances dorsally, and segment X with the posterolateral borders not developed.</p> <p>Adult: Head and body yellowish brown.</p> <p>Male: Forewing length: 10–12 mm. Forewing light golden brown, with 12 small brown spots. Male genitalia as in Figure 7. Abdominal segment IX annular; in dorsal view, with anterior margin nearly straight, posterior margin broadly rounded; in lateral view, pleural region setose; tergum IX without protuberances dorsally. Preanal appendage setose, at least 4× as long as wide; in lateral view, digitate, apex subacute. Segment X saddle-shaped; in lateral view not flattened, dorsal margin slightly concave, apex upcurved, ventral margin abruptly convex; with posterolateral borders not developed, lateral margins subparallel after 1/3 length, with apicodorsal shelf, shelf rounded or almost rectangular, almost as wide as apex of segment X, with ventral sulcus apically. Inferior appendage slightly concave in ventral view, with basal region tuboid, not enlarged; rounded basally, heavily setose, bearing a small basomesal protuberance; middle region of inferior appendage with constriction before apicodorsal region; 2nd article shorter than apicodorsal portion of 1st article, broad, tapering throughout end, mesal surface not excavate, apex pointed, slightly curved mesally; apicordorsal portion of 1st article digitate, setose. Phallic apparatus simple, phallicata slightly down-curved; with pair of subequal, wide, posterodorsally directed lateral flanges, apices not acuminate; phallotremal sclerites small, U-shaped in dorsal view.</p> <p>Female: Head and body yellowish. Forewing length: 6–8 mm. Forewing yellowish to brown, without dark setae on posterodorsal region. Genitalia inseparable from congeners. Abdominal segment IX lightly sclerotized, with dorsal plate, bearing pair of setose protuberances above appendages of segment X, ap- pendages of segment X small; valves flat, thin, lightly sclerotized, with very short setae.</p> <p>Etymology: The specific epithet is in honor of Dr Roger Blahnik for his dedication to Neotropical caddisflies.</p> <p>Distribution: Peru.</p> <p>Holotype, male: PERU: Paucartambo: Pte. San Pedro, c. 50 km NW Pilcopata, el. 1600 m, 2–3.ix.1988, O.S. Flint Jr and N. Adams (NMNH).</p> <p>Paratypes: same data as holotype, 6 males (NMNH), 2 males (UFBA), 2 males (UMSP).</p> </div>	https://treatment.plazi.org/id/03966709C83E7E58FF35FE29EAEAFDF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C83F7E5FFCCDFD11EBA8FDC9.text	03966709C83F7E5FFCCDFD11EBA8FDC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella boraceia	<div><p>GRUMICHELLA BORACEIA CALOR &amp; HOLZENTHAL SP. NOV. (FIG. 8)</p> <p>Diagnosis: Grumichella boraceia can be diagnosed by the following characters: male forewings with 3 brown spots; male genitalia bearing segment X not flattened, lateral view; 1st article of inferior appendage with small basomesal protuberance; 2nd article of inferior appendage subequal the apicodorsal portion of 1st article; apex of 2nd article acuminated, curved mesad; larval abdominal sternum I with 16–24 setae.This new species is most similar to G. aequiunguis, but differs in having the 2nd article of the inferior appendage with the mesal surface excavate, and with the apex blunt.</p> <p>Adult: Head and body yellowish brown.</p> <p>Male: Forewing length: 9–10 mm. Forewing light golden brown, with 3 small brown spots, without dark setae on posterodorsal margin. Male genitalia as in Figure 8. Abdominal segment IX annular; in dorsal view, with anterior margin slightly convex, posterior margin broadly rounded, with medial protuberance; in lateral view, pleural region setose; tergum IX without protuberances dorsally. Preanal appendage setose, at least 4× as long as wide; in lateral view, digitate, apex rounded. Segment X saddle-shaped; in lateral view not flattened, dorsal margin slightly concave, apex upcurved, ventral margin abruptly convex, with middle length straight; segment X in dorsal view, rectangular, with posterolateral borders well developed, lateral margins subparallel after 1/3 length, with apicodorsal shelf, shelf almost rectangular, but sometimes varying to rounded, 1/2 as wide as apex of segment X, without sulcus apically. Inferior appendage slightly concave in ventral view, with basal region enlarged; rounded basally, heavily setose, bearing a small basomesal protuberance; middle region of inferior appendage with constriction before apicodorsal region; 2nd article subequal in length to apicodorsal portion of 1st article, narrow, tapering throughout length, mesal surface not excavate, apex pointed, curved mesally; apicordorsal portion of 1st article digitate, setose. Phallic apparatus simple, phallicata slightly down-curved; with pair of subequal, wide, posterodorsally directed lateral flanges, apices not acuminate; phallotremal sclerites small, U-shaped in dorsal view.</p> <p>Female: Head and body yellowish. Forewing length: 7–9 mm. Forewing yellowish to brown, without dark setae on posterodorsal region. Genitalia inseparable from congeners. Abdominal segment IX lightly sclerotized, with dorsal plate, bearing pair of setose protuberances above appendages of segment X, appendages of segment X small; valves flat, thin, lightly sclerotized, with very short setae.</p> <p>Larva: Length of larva 6–9 mm. Abdominal sternum I with 16–24 setae. Lateral hump sclerite small.</p> <p>Larval case: Length 7–11 mm; constructed entirely of rings of dark brown silk, gently curved, tapered, wide, with 2 distinct diagonal sutures, 1 towards posterior end and 1 more or less at midlength, posterior end with ventral, dorsally directed projection, visible externally, posterior aperture C-shaped.</p> <p>Pupa: Segment IX very slender; anal process long, narrow, slender, unpigmented or slightly yellowish. With dark setae. Apically upturned, hooked.</p> <p>Pupal case: With anteroventral silken pedicel, pedicel long; anterior silken membrane with an eccentric, horizontal, slit-like opening.</p> <p>Etymology: The species name refers to ‘Estação Biológica de Boracéia’, Salesópolis, São Paulo, the research station of the Museu de Zoologia, Universidade de São Paulo. The word ‘boracéia’ means ‘agglomeration of people’ or, by derivation, ‘party, dance’. It is treated as a noun in apposition.</p> <p>Distribution: Brazil.</p> <p>Holotype, male: BRAZIL: São Paulo: Salesópolis, Rio</p></div> 	https://treatment.plazi.org/id/03966709C83F7E5FFCCDFD11EBA8FDC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C8387E5FFCDAFBABEBA8F8CF.text	03966709C8387E5FFCDAFBABEBA8F8CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella cressae	<div><p>GRUMICHELLA CRESSAE CALOR &amp; HOLZENTHAL SP. NOV. (FIG. 9)</p> <p>Diagnosis: Grumichella cressae can be diagnosed by the following characters: male forewings with 12 brown spots; male genitalia bearing segment X not flattened, lateral view; 1st article of inferior appendage with well-developed basomesal protuberance; 2nd article of inferior appendage shorter than apicodorsal portion of 1st article; apex of 2nd article acuminated and curved mesad. It is most similar to G. blahniki sp. nov., but differs in having segment X with posterolateral borders developed, and tergum IX bearing 1–2 small protuberances posterodorsally, which are absent in G. blahniki sp. nov.</p> <p>Adult: Head and body yellowish brown.</p> <p>Male: Forewing length: 9–10 mm. Forewing light golden brown, with 12 small brown spots. Male genitalia as in Figure 9. Abdominal segment IX annular; in dorsal view, with anterior margin nearly straight, posterior margin nearly straight; in lateral view, pleural region heavily setose; tergum IX bearing a pair of small probuberances posterodorsally. Preanal appendage setose, at least 4× as long as wide; in lateral view, digitate, apex rounded. Segment X saddle-shaped; in lateral view not flattened, dorsal margin slightly concave, apex</p></div> 	https://treatment.plazi.org/id/03966709C8387E5FFCDAFBABEBA8F8CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C8397E5EFCFDF9E2EB84F8CC.text	03966709C8397E5EFCFDF9E2EB84F8CC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella jureia	<div><p>GRUMICHELLA JUREIA CALOR &amp; HOLZENTHAL SP. NOV. (FIG. 10)</p> <p>Diagnosis: Grumichella jureia can be diagnosed by the following characters: male forewings with 3 brown spots, and dark setae in posterodorsal border; male genitalia bearing segment X with apicodorsal expansion, almost as wide as apical region, and dorsal sulcus on the segment X apex; 2nd article of inferior</p> </div>	https://treatment.plazi.org/id/03966709C8397E5EFCFDF9E2EB84F8CC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C83B7E43FCF9FF5FEBABFC78.text	03966709C83B7E43FCF9FF5FEBABFC78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella leccii	<div><p>GRUMICHELLA LECCII CALOR &amp; HOLZENTHAL SP. NOV. (FIG. 11)</p> <p>Diagnosis: Grumichella leccii can be diagnosed by the following characters: male forewings with 3 brown spots; male genitalia bearing segment X flattened, lateral view; 1st article of inferior appendage bearing a small basomesal protuberance; 2nd article longer than apicodorsal portion of 1st article, narrow, tapering throughout length, apex pointed, curved mesally; larval abdominal sternum I with 16–24 setae. It is most similar to G. rostrata, except for 2nd article of inferior appendage which is longer than the apicodorsal portion of 1st article, and segment X with small sulcus apically.</p> <p>Adult: Head and body yellowish brown.</p> <p>Male: Forewing length: 7–9 mm. Forewing light golden brown, with 3 small brown spots, without dark setae on posterodorsal margin. Male genitalia as in Figure 10. Abdominal segment IX annular; in dorsal view, with anterior margin slightly concave, posterior margin sinuate; in lateral view, pleural region sparsely setose; tergum IX without protuberances dorsally. Preanal appendage setose, at least 4× as long as wide; in lateral view, clavate, apex rounded. Segment X saddleshaped; in lateral view flattened, dorsal margin slightly concave, apex upcurved, ventral margin abruptly convex; segment X in dorsal view, subtriangular, with posterolateral borders developed, lateral margins sinuate, narrowing apically, with apicodorsal shelf, shelf almost rectangular, 1/2 as wide as apex of segment X, with ventral sulcus apically. Inferior appendage nearly straight in ventral view, with basal region tuboid, not enlarged; rounded basally, heavily setose, bearing a small basomesal protuberance; middle region of inferior appendage with constriction before apicodorsal region; 2nd article longer than apicodorsal portion of 1st article, narrow, tapering throughout length, mesal surface not excavate, apex pointed, curved mesally; apicordorsal portion of 1st article digitate, setose. Phallic apparatus simple, phallicata slightly down-curved; with pair of subequal, narrow, posteromesally directed lateral flanges, apices not acuminate; phallotremal sclerites small, U-shaped in dorsal view.</p> <p>Female: Head and body yellowish. Forewing length: 6–7 mm. Forewing yellowish to brown, without dark setae on posterodorsal region. Genitalia inseparable from congeners. Abdominal segment IX lightly sclerotized, with dorsal plate, bearing pair of setose protuberances above appendages of segment X, appendages of segment X small; valves flat, thin, lightly sclerotized, with very short setae.</p> <p>Larva: Length of larva 6–8 mm. Abdominal sternum I with 16–24 setae. Lateral hump sclerite relativally large.</p> <p>Larval case: Length 7–10 mm; constructed of rings of transparent silk, nearly straight, tapered, narrow, without diagonal sutures (or sutures indistinct), posterior end with ventral, dorsally directed projection, visible externally, posterior aperture C-shaped.</p> <p>Pupa: Segment IX very slender; anal process long, narrow, slender, unpigmented or slightly yellowish. Without dark setae. Apically upturned, hooked.</p> <p>Pupal case: With anteroventral silken pedicel, pedicel very long, thin; anterior silken membrane with an eccentric, horizontal, slit-like opening.</p> <p>Etymology: Named in honour of Dr Lucas Silveira Lecci (Instituto Federal do Mato Grosso, Brazil), collector of this species, and also for his friendship and collaboration in the field.</p> <p>Distribution: Brazil.</p> <p>Holotype, male: BRAZIL: São Paulo: Jundiai, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.954445&amp;materialsCitation.latitude=-23.241667" title="Search Plazi for locations around (long -46.954445/lat -23.241667)">Serra do Japi</a>, P.A. 11, stream before reservoir, 23°14′30″S, 46°57′16″W, 27.iii.2007, luz UV &amp; branca, Calor A.R., Mariano R. &amp; Lecci L.S. (MZUSP).</p> <p>Paratypes: Same data as holotype, except 5 males (MZUSP), 5 males (UMSP); 27 males (UFBA); same data except <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.933334&amp;materialsCitation.latitude=-23.216667" title="Search Plazi for locations around (long -46.933334/lat -23.216667)">Córrego da Ermida</a> and small dam, 9.xii.1997, Froehlich C.G., 23°13′S, 46°56′W, 2 males (UMSP); same data, except <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.933334&amp;materialsCitation.latitude=-23.233334" title="Search Plazi for locations around (long -46.933334/lat -23.233334)">Córrego</a> da cachoeira Paraíso, 26.iii.2007, luz UV &amp; branca, 23°14′S, 46°56′W, Calor A.R., Mariano R. &amp; Lecci L.S., 1 male (UFBA).</p> <p>Additional material: Same data as holotype, except 5 males, 5 females (UMSP), 27 males, 49 females (UFBA); same data, except <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.933334&amp;materialsCitation.latitude=-23.233334" title="Search Plazi for locations around (long -46.933334/lat -23.233334)">Córrego da Ermida</a> and small dam, 9.xii.1997, Froehlich C.G., 23°14′S, 46°56′W, 2 females (UMSP); same data, except riacho trilha da cachoeira <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.953613&amp;materialsCitation.latitude=-23.241667" title="Search Plazi for locations around (long -46.953613/lat -23.241667)">Paraíso</a>, 25.i.2007, luz, 23°14′30″S, 46°57′13″W, Lecci L.S., Nascimento E. &amp; Polegatto C., 1 female (MZUSP); same data, except 2° riacho trilha da cachoeira <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.953613&amp;materialsCitation.latitude=-23.241667" title="Search Plazi for locations around (long -46.953613/lat -23.241667)">Paraíso</a>, 22.ii.2007, luz, 23°14′30″S, 46°57′13″W, Lecci L.S. &amp; Nascimento E., 1 female (MZUSP); same data, except 2° riacho trilha da cachoeira <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.953613&amp;materialsCitation.latitude=-23.241667" title="Search Plazi for locations around (long -46.953613/lat -23.241667)">Paraíso</a>, 22.ii.2007, luz, 23°14′30″S, 46°57′13″W, Lecci L.S. &amp; Nascimento E., 1 female (MZUSP); same data, except <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-46.933334&amp;materialsCitation.latitude=-23.233334" title="Search Plazi for locations around (long -46.933334/lat -23.233334)">Córrego</a> da cachoeira Paraíso, 26.iii.2007, luz UV &amp; branca, 23°14′S, 046°56′W, Calor A.R., Mariano R. &amp; Lecci L.S., 3 females (UFBA); same data, except P.A. 11, stream before the reservoir, 23°14′30″S, 46°57′15″W, el. 1022 m, 11.iii.2008, rede D, Lecci L.S. &amp; Nascimento E., 21 pupae (UFBA).</p> </div>	https://treatment.plazi.org/id/03966709C83B7E43FCF9FF5FEBABFC78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C8247E41FCA2FB8AEA45FD9E.text	03966709C8247E41FCA2FB8AEA45FD9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella muelleri	<div><p>GRUMICHELLA MUELLERI CALOR &amp; HOLZENTHAL SP. NOV. (FIGS 12, 13)</p> <p>Diagnosis: Grumichella muelleri can be diagnosed by the following characters: male forewings with 3 brown spots; male genitalia bearing segment X rectangular, with posterolateral border developed, dorsal view; without sulcus on the apex of segment X; 1st article of inferior appendage without basomesal protuberance; 2nd article of inferior appendage shorter than apicodorsal portion of 1st article; apex of 2nd article acuminated, curved mesad. This new species is most similar to G. paprocki sp. nov., but differs in having segment X with the posterolateral borders not developed, and the apicodorsal shelf on segment X from 2/3 to almost as wide as the apex of segment X.</p> <p>Adult: Head and body yellowish brown.</p> <p>Male: Forewing length: 6–7 mm. Forewing light golden brown, with 3 small brown spots, without dark setae on posterodorsal margin. Male genitalia as in Figure 12. Abdominal segment IX annular; in dorsal view, with anterior margin slightly concave, posterior margin slightly angulate; in lateral view, pleural region sparsely setose; tergum IX without protuberances dorsally. Preanal appendage setose, at least 4× as long as wide; in lateral view, clavate, apex acute. Segment X saddleshaped; in lateral view not flattened, dorsal margin slightly concave, ventral margin abruptly convex; segment X in dorsal view, rectangular, with posterolateral borders not developed, lateral margins sinuate, with apicodorsal shelf, shelf almost rectangular, but sometimes varying to rounded, almost as wide as apex of segment X, without sulcus apically. Inferior appendage slightly concave in ventral view, with basal region enlarged; rounded basally, heavily setose, especially apically, not bearing a basomesal pro- tuberance; middle region of inferior appendage without constriction before apicodorsal region; 2nd article shorter than apicodorsal portion of 1st article, broad basally, tapering throughout length, mesal surface not excavate, apex pointed, curved mesally; apicordorsal portion of 1st article digitate, setose. Phallic apparatus simple, phallicata nearly straight; with pair of subequal, wide, posterodorsally directed lateral flanges, apices acuminate; phallotremal sclerites small, C-shaped.</p> <p>Female: Head and body yellowish. Forewing length: 5–7 mm. Forewing yellowish to brown, without dark setae on posterodorsal region. Genitalia as in Figure 13, inseparable among the congeners. Abdominal segment IX lightly sclerotized, with dorsal plate, bearing a pair of setose protuberances above appendages of segment X, appendages of segment X small; valves flat, thin, lightly sclerotized, with very short setae.</p> <p>Etymology: The specific epithet is inhonor of the naturalist Fritz Müller, the ‘prince of observers’, according to Charles Darwin. Müller was a pioneer in studies of Neotropical caddisflies (among other taxa), including the description of the genus Grumichella. He was an important collaborator and supporter of Darwin’s theory of evolution, especially by means of Müller’s book ‘Für Darwin’ (1864) and its translations.</p> <p>Distribution: Brazil.</p> <p>Holotype, male: BRAZIL: Santa Catarina: Parque Ecológico Spitzkopf, confl. Rio Ouro &amp; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.111664&amp;materialsCitation.latitude=-27.005833" title="Search Plazi for locations around (long -49.111664/lat -27.005833)">Rio Caeté</a>, 25.ix.2003, UV light, Holzenthal, Paprocki &amp; Calor, 27°00′21″S, 49°06′42″W, el. 140 m (MZUSP).</p> <p>Paratypes: Same data as holotype, except 1male (UMSP), 1 male (UFBA).</p> <p>Additional material: Same data as holotype, except 1 female (UMSP); same data, except <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.111664&amp;materialsCitation.latitude=-27.005833" title="Search Plazi for locations around (long -49.111664/lat -27.005833)">Rio Caeté</a> above 1st falls, 4.iii.1998, UV light, Holzenthal, Froehlich &amp; Paprocki, 27°00′21″S, 49°06′42″W, el. 170 m, 2 females (UMSP), 1 female (UFBA).</p> </div>	https://treatment.plazi.org/id/03966709C8247E41FCA2FB8AEA45FD9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C8267E40FCA4FDE0EB87FB30.text	03966709C8267E40FCA4FDE0EB87FB30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella paprockii	<div><p>GRUMICHELLA PAPROCKII CALOR &amp; HOLZENTHAL SP. NOV. (FIG. 14)</p> <p>Diagnosis: Grumichella paprockii can be diagnosed by the following characters: male forewing with 3 brown spots; male genitalia bearing segment X not flattened, lateral view; 1st article of inferior appendage without basomesal protuberance; 2nd article of inferior appendage shorter than the apicodorsal portion of 1st article; apex of 2nd article acuminated, curved mesad. This new species is most similar to G. muelleri, but differs in having segment X with the posterolateral borders developed, and the apicodorsal shelf on segment X about 1/3 to 1/2 as wide as the apex of segment X.</p> <p>Adult: Head and body yellowish brown.</p> <p>Male: Forewing length: 8–9 mm. Forewing light golden brown, with 3 small brown spots, without dark setae on posterodorsal margin. Male genitalia as in Figure 14. Abdominal segment IX annular; in dorsal view, with anterior margin slightly concave, posterior margin broadly rounded, with medial protuberance; in lateral view, pleural region sparsely setose; tergum IX bearing 1 small probuberance posterodorsally. Preanal appendage setose, at least 4× as long as wide; in lateral view, digitate, apex subacute. Segment X saddle-shaped; in lateral view not flattened, dorsal margin slightly concave, apex upcurved, ventral margin slightly convex; segment X in dorsal view, rectangular, with posterolateral borders well developed, lateral margins subparallel, with apicodorsal shelf, shelf almost rectangular, but sometimes varying to rounded, 1/3 as wide as apex of segment X, with ventral sulcus apically. Inferior appendage concave in ventral view, with basal region tuboid, not enlarged; slightly rounded basally, setose, not bearing a basomesal protuberance; middle region of inferior appendage without constriction before apicodorsal region; 2nd article shorter than apicodorsal portion of 1st article, broad basally, tapering throughout length, apex pointed, curved mesally; apicordorsal portion of 1st article digitate, setose. Phallic apparatus simple, phallicata sharply down-curved; with pair of subequal, wide, posteromesally directed lateral flanges, apices not acuminate; phallotremal sclerites small, U-shaped in dorsal view.</p> <p>Female: Head and body yellowish. Forewing length: 6–7 mm. Forewing yellowish to brown, without dark setae on posterodorsal region. Genitalia inseparable from congeners. Abdominal segment IX lightly sclerotized, with dorsal plate, bearing pair of setose protuberances above appendages of segment X, appendages of segment X small; valves flat, thin, lightly sclerotized, with very short setae.</p> <p>Etymology: Named in honour of Dr Henrique Paprocki, collector of this species and a colleague who encouraged this revision.</p> <p>Distribution: Brazil.</p> <p>Holotype, male: BRAZIL: Minas Gerais: Córrego da Serra de Ouro Fino, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.64306&amp;materialsCitation.latitude=-20.206112" title="Search Plazi for locations around (long -43.64306/lat -20.206112)">Valedo Tropeiro</a>, 8.x.2000, UV light, Paprocki, Salgado &amp; Isaac, 20°12′22″S, 43°38′35″W, el. 1000 m (MZUSP).</p> <p>Paratypes: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.636112&amp;materialsCitation.latitude=-20.204445" title="Search Plazi for locations around (long -43.636112/lat -20.204445)">Same</a> data as holotype, except Cachoeira do Abacaxi, Vale do Tropeiro, 7.xi.2001, UV light, Holzenthal, Amarante, Blahnik &amp; Paprocki, 20°12′16″S, 43°38′10″W, el. 1120 m, 2 males (UMSP); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.427223&amp;materialsCitation.latitude=-20.417221" title="Search Plazi for locations around (long -43.427223/lat -20.417221)">Parque Estadual Itacolomi</a>, Rio Belchior, 2.ii.1998, UV light, Holzenthal &amp; Paprocki, 20°25′02″S, 043°25′38″W, el. 725 m, 3 males (UMSP), 2 males (UFBA); same data, except 24.i.1999, UV light, Amarante M.C., 20°25′02″S, 43°25′38″W, el. 725 m, 1 male (UMSP); same data, except 20.ii.1999, UV light, Paprocki &amp; Amarante, 20°25′02″S, 43°25′38″W, el. 725 m, 2 males (UMSP).</p> <p>Additional material: Same data as holotype, except 2 females (UMSP); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.427223&amp;materialsCitation.latitude=-20.417221" title="Search Plazi for locations around (long -43.427223/lat -20.417221)">Parque Estadual Itacolomi</a>, Rio Belchior, 2.ii.1998, UV light, Holzenthal &amp; Paprocki, 20°25′02″S, 043°25′38″W, el. 725m, 30 females (UMSP), 2 females (UFBA); same data, except 8.v.1998, UV light, Holzenthal &amp; Paprocki, 20°25′02″S, 43°25′38″W, el. 725 m, 2 females (UMSP); same data except 20.xi.1998, UV light, Paprocki &amp; Amarante, 20°25′02″S, 43°25′38″W, el. 725 m, 8 females (UMSP); same data except 17.xii.1998, UV light, Paprocki &amp; Amarante, 20°25′02″S, 43°25′38″W, el. 725 m, 4 females (UMSP); same data except 24.i.1999, UV light, Amarante M.C., 20°25′02″S, 43°25′38″W, el. 725 m, 4 females (UMSP); same data except 20.ii.1999, UV light, Paprocki &amp; Amarante, 20°25′02″S, 43°25′38″W, el. 725 m, 10 females (UMSP).</p> </div>	https://treatment.plazi.org/id/03966709C8267E40FCA4FDE0EB87FB30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C8277E46FCFBFB53ECC5FD0C.text	03966709C8277E46FCFBFB53ECC5FD0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella parati	<div><p>GRUMICHELLA PARATI CALOR &amp; HOLZENTHAL SP. NOV. (FIG. 15)</p> <p>Diagnosis: Grumichella parati can be diagnosed by the following characters: male forewing with 3 brown spots; male genitalia with segment X not flattened, lateral view; 1st article of inferior appendage without basomesal protuberance; 2nd article of inferior appendage longer than apicodorsal portion of 1st article; apex of 2nd article acuminated, curved mesad. It is most similar to G. jureia sp. nov., but differs in having the 2nd article of the inferior appendage longer than the apicodorsal portion of the 1st article, and segment X with the posterolateral borders developed.</p> <p>Adult: Head and body yellowish brown.</p> <p>Male: Forewing length: 8–9 mm. Forewing light golden brown, with 3 small brown spots, without dark setae on posterodorsal margin. Male genitalia as in Figure 15. Abdominal segment IX annular; in dorsal view, with anterior margin nearly straight, posterior margin broadly rounded; in lateral view, pleural region setose; tergum IX without protuberances dorsally. Preanal appendage setose, at least 4× as long as wide; in lateral view, clavate, apex rounded. Segment X saddle-shaped; in lateral view not flattened, dorsal margin slightly concave, ventral margin abruptly convex, with middle length straight; segment X in dorsal view, subtriangular, with posterolateral borders developed, lateral margins sinuate, narrowing apically, with apicodorsal shelf, shelf slightly trapezoidal, 1/2 as wide as apex of segment X, without sulcus apically. Inferior appendage slightly concave in ventral view, with basal region tuboid, not enlarged; slightly rounded basally, setose, not bearing a basomesal protuberance; middle region of inferior appendage without constriction before apicodorsal region; 2nd article longer than apicodorsal portion of 1st article, narrow, tapering throughout length, mesal surface not excavate, apex pointed, curved mesally; apicordorsal portion of 1st article digitate, setose. Phallic apparatus simple, phallicata sharply down-curved; with pair of subequal, narrow, posterodorsally directed lateral flanges, apices acuminate; phallotremal sclerites small, U-shaped in dorsal view, with middle region enlarged.</p> <p>Female: Head and body yellowish. Forewing length: 6–8 mm. Forewing yellowish to brown, without dark setae on posterodorsal region. Genitalia inseparable from congeners. Abdominal segment IX lightly sclerotized, with dorsal plate, bearing pair of setose protuberances above appendages of segment X, appendages of segment X small; valves flat, thin, lightly sclerotized, with very short setae.</p> <p>Etymology: Parati’y in the Tupi-guarani language is a composite word formed by parati, a species of fish in the genus Mugil (Mugilidae) and ′ y (river). It is treated as a noun in apposition.</p> <p>Distribution: Brazil.</p> <p>Holotype, male: BRAZIL: Rio de Janeiro: Parati, Riacho Perequê-açu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.79&amp;materialsCitation.latitude=-23.220556" title="Search Plazi for locations around (long -44.79/lat -23.220556)">Sitio Cachoeira Grande</a>, 25.ix.2002, UV light, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.79&amp;materialsCitation.latitude=-23.220556" title="Search Plazi for locations around (long -44.79/lat -23.220556)">Blahnik</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.79&amp;materialsCitation.latitude=-23.220556" title="Search Plazi for locations around (long -44.79/lat -23.220556)">Prather</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.79&amp;materialsCitation.latitude=-23.220556" title="Search Plazi for locations around (long -44.79/lat -23.220556)">Melo</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.79&amp;materialsCitation.latitude=-23.220556" title="Search Plazi for locations around (long -44.79/lat -23.220556)">Froehlich</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.79&amp;materialsCitation.latitude=-23.220556" title="Search Plazi for locations around (long -44.79/lat -23.220556)">Silva</a>, 23°13′14″S, 44°47′24″W, el. 120 m (MZUSP).</p> <p>Paratypes: Same data as holotype, except 1 male (MZUSP), 13 males (UMSP), 2 males (UFBA); same data, except <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.79139&amp;materialsCitation.latitude=-23.21389" title="Search Plazi for locations around (long -44.79139/lat -23.21389)">Trib.</a> to Riacho Perequê-açu, 26.ix.2002, UV/mercury vapor lights, Blahnik, Prather, Melo, Froehlich, Silva, 23°12′50″S, 44°47′29″W, el. 190 m, 7 males (UMSP); same data except Rio das Flores, Macaéde Cima, 10 km SE Mury, 9.iii.2002, UV light, Holzenthal, Blahnik, Paprocki &amp; Prather, 1 male (UMSP); same data, except <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.311665&amp;materialsCitation.latitude=-22.391388" title="Search Plazi for locations around (long -42.311665/lat -22.391388)">Encontro</a> dos Rios Macaé / Bonito, 6 km S Lumiar, 10.iii.2002, Holzenthal, Blahnik, Paproki &amp; Prather, 22°23′29″S, 42°18′42″W, el. 600 m, 8 males (UMSP), 5 males (UFBA).</p> <p>Aditonal material: Same data as holotype, except 1 female (MZUSP), 2 females (UMSP); same data, except trib. to <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-44.79139&amp;materialsCitation.latitude=-23.21389" title="Search Plazi for locations around (long -44.79139/lat -23.21389)">Riacho Perequê-açu</a>, 26.ix.2002, UV/mercury vapour lights, Blahnik, Prather, Melo, Froehlich, Silva, 23°12′50″S, 44°47′29″W, el. 190 m, 15 females (UMSP); same data, except <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-42.502224&amp;materialsCitation.latitude=-22.394722" title="Search Plazi for locations around (long -42.502224/lat -22.394722)">Macaé</a>, Rio Macaé, Macaé de Cima, 8.iii.2002, UV light, Holzenthal, Blahnik, Paproki &amp; Prather, 22°23′41″S, 42°30′08″W, el. 1000 m, 15 females (UMSP); same data, except Rio das Flores, Macaé de Cima, 10 km SE Mury, 09.iii.2002, UV light, Holzenthal, Blahnik, Paprocki &amp; Prather, 3 females (UMSP).</p> </div>	https://treatment.plazi.org/id/03966709C8277E46FCFBFB53ECC5FD0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
03966709C8217E45FF27FD71EA18FAC5.text	03966709C8217E45FF27FD71EA18FAC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grumichella trujilloi	<div><p>GRUMICHELLA TRUJILLOI CALOR &amp; HOLZENTHAL SP. NOV. (FIG. 16)</p> <p>Diagnosis: Grumichella trujilloi can be diagnosed by the following characters: male forewings with 12 brown spots; male genitalia bearing segment X flattened, lateral view; 1st article of inferior appendage bearing small basomesal protuberance; 2nd article of inferior appendage subequal in length to apicodorsal portion of 1st article; apex of 2nd article acuminated, curved mesad. Grumichella trujilloi is most similar to G. cressae and G. blahniki, but differs in having the 2nd article of inferior appendage shorter than the apicodorsal portion of the 1st article, and in having segment X, in lateral view, not flattened.</p> <p>Adult: Head and body yellowish brown.</p> <p>Male: Forewing length: 9–11 mm. Forewing light golden brown, with 12 small brown spots. Male genitalia as in Figure 16. Abdominal segment IX annular; in dorsal view, with anterior margin nearly straight, posterior margin sinuate; in lateral view, pleural region setose; tergum IX bearing a pair of small probuberances posterodorsally. Preanal appendage setose, at least 4× as long as wide; in lateral view, digitate, apex rounded. Segment X saddle-shaped; in lateral view flattened, dorsal margin slightly concave, apex upcurved, ventral margin convex; segment X in dorsal view, subtriangular, with posterolateral borders not developed, lateral margins sinuate, with apicodorsal shelf, shelf variable, acuminate to rounded, almost as wide as apex of segment X, with ventral sulcus apically. Inferior appendage slightly concave in ventral view, with basal region enlarged; subquadrangular basally, setose, bearing a small basomesal protuberance; 2nd article subequal in length to apicodorsal portion of 1st article, narrow, tapering throughout length, but constricted subapically or broad basally, tapering throughout length, mesal</p> <p>KEY TO MALES OF GRUMICHELLA MÜLLER</p> <p>1. Forewing with 2–3 small brown spots... 2</p> <p>Forewing with 9–12 small brown spots... 9</p> <p>2(1). Inferior appendage basally bearing a basomesal protuberance (Fig 2)... 3</p> <p>Inferior appendage basally not bearing a basomesal protuberance (Fig 14)... 6</p> <p>3(2). Segment X, in lateral view, not flattened (Fig 2)... 4</p> <p>Segment X, in lateral view, flattened (Fig 6)... 5</p> <p>4(3). 2nd article of inferior appendage with mesal surface not excavate; 2nd article with apex pointed (Fig 8)... Grumichella boraceia sp. nov.</p> <p>2nd article of inferior appendage with mesal surface excavate; 2nd article with apex blunt (Fig 2)... Grumichella aequiunguis Flint</p> <p>5(3). 2nd article of inferior appendage subequal in length to apicodorsal portion of 1st article; segment X without sulcus apically (Fig 6)... Grumichella rostrata Thienemann</p> <p>2nd article of inferior appendage longer than apicodorsal portion of 1st article; segment X with small sulcus apically (Fig 11)... Grumichella leccii sp. nov.</p> <p>6(2). 2nd article of inferior appendage shorter than apicodorsal portion of 1st article... 7</p> <p>2nd article of inferior appendage subequal in length or longer than apicodorsal portion of 1st article... 8</p> <p>7(6). Segment X with posterolateral borders developed; apicodorsal shelf on segment X 1/3 to 1/2 as wide as apex of segment X (Fig 14)... Grumichella paprockii sp. nov.</p> <p>Segment X with posterolateral borders not developed; apicodorsal shelf on segment X 2/3 to almost as wide as apex of segment X (Fig 12)... Grumichella muelleri sp. nov.</p> <p>8(6). 2nd article of inferior appendage subequal in length to apicodorsal portion of 1st article; segment X with posterolateral borders not developed (Fig 10)... Grumichella jureia sp. nov.</p> <p>2nd article of inferior appendage longer than apicodorsal portion of 1st article; segment X with posterolateral borders developed (Fig 15)... Grumichella parati sp. nov.</p> <p>9(1). 2nd article of inferior appendage shorter than apicodorsal portion of 1st article; segment X, in lateral view, not flattened... 10</p> <p>2nd article of inferior appendage subequal in length or longer than apicodorsal portion of 1st article; segment X, in lateral view, flattened (Fig 11)... Grumichella trujilloi sp. nov.</p> <p>10(9). Segment X with posterolateral borders developed; tergum IX bearing 1-2 small protuberances posterodorsally (Fig 9)... Grumichella cressae sp. nov.</p> <p>Segment X with posterolateral borders not developed; tergum IX without protuberances dorsally (Fig 7)... 11</p> <p>11(10). Forewing with 12 small brown spots... Grumichella blahniki sp. nov.</p> <p>Forewing with 9 small brown spots... 12</p> <p>12(11). Apicodorsal portion of 1st article of inferior appendage digitate (Fig 3)... Grumichella flaveola Ulmer Apicodorsal portion of 1st article of inferior appendage quadrate (Fig 5)... Grumichella pulchella Banks</p> <p>surface excavate, apex pointed, curved mesally; apicordorsal portion of 1st article digitate, setose. Phallic apparatus simple, phallicata slightly down-curved; with pair of subequal, wide, posterodorsally directed lateral flanges, apices not acuminate; phallotremal sclerites relativelly large, U-shaped in dorsal view.</p> <p>Female: Head and body yellowish. Forewing length: 6–7 mm. Forewing yellowish to brown, without dark setae on posterodorsal region. Genitalia inseparable from congeners. Abdominal segment IX lightly sclerotized, with dorsal plate, bearing pair of setose protuberances above appendages of segment X, appendages of segment X small; valves flat, thin, lightly sclerotized, with very short setae.</p> <p>Etymology: The name of the species refers to the state of Trujillo, Venezuela, where the species was collected.</p> <p>Distribution: Venezuela.</p></div> 	https://treatment.plazi.org/id/03966709C8217E45FF27FD71EA18FAC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Calor, Adolfo R.;Holzenthal, Ralph W.;Froehlich, Claudio G.	Calor, Adolfo R., Holzenthal, Ralph W., Froehlich, Claudio G. (2016): Phylogeny and revision of the Neotropical genus Grumichella Müller (Trichoptera: Leptoceridae), including nine new species and a key. Zoological Journal of the Linnean Society 176 (1): 137-169, DOI: 10.1111/zoj.12310, URL: http://dx.doi.org/10.1111/zoj.12310
