taxonID	type	description	language	source
03978787FFB1FFC3FF6BB139FD61DA6A.taxon	materials_examined	TYPE: — MEXICO: Jalisco: [Amatitán Municipality], below presa de Santa Rosa, in the barranca of the río Grande, north of Amatitán, 750 – 800 m, Sep 1, 1960, McVaugh 18530 fruits (holotype, MICH!; isotype, US! )). EPITYPE (here designated): — MEXICO: Jalisco: Municipio San Cristóbal de la Barranca, ca. rancho Jacalitos, a 1 km al SE de Santa Cruz de Atístique, bosque tropical caducifolio con Pilosocereus, Stenocereus queretaroensis, Ceiba sp., Bursera sp., Pithecellobium dulce, 21 ° 06 ’ 37.9 ’’ N, 103 ° 26´34.1 ’’ W, 920 m, originally collected by I. Ramírez, P. Carrillo-Reyes, W. Cetzal & J. L. Tapia, May 2015; flowering in Mérida, Yucatán, May 2018, I. Ramírez 2432 ♂ (CICY!).	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFB1FFC3FF6BB139FD61DA6A.taxon	description	Plants lithophytic, 1.3 – 1.4 m height when blooming rosettes cespitose, generally circular, 30 – 40 cm tall, 60 – 70 cm in diameter, forming dense small clusters of 2 – 5 rosettes. Leaves 25 – 34 in number, reflexed; sheath broadly ovate, 3.5 – 4 × 5 – 6.8 cm, light brown, margins entire at the base, distally serrate, lustrous and glabrous at the base, lepidote distally on both surfaces; blade narrowly triangular, acuminate to long attenuate, 50 – 75 × 3 – 3.5 cm, green, reddish toward the margin when sun exposed, glabrous and shiny above, and white lepidote below; spines antrorse or divaricate, triangular, ca. 3 mm long, 8 – 30 mm apart, green to light brown. Inflorescence central, erect, emerging from a fully grown rosette (strict sympodium growth pattern, type SPP sensu Ramírez-Morillo et al. 2014). Staminate inflorescence a once-divided panicle, cylindrical in general shape, erect to slightly curved, 1.3 – 1.4 m long; peduncle terete, green, sparsely white lepidote, 40 – 50 cm long, 8 – 10 mm in diameter at the base, about as long as rosette height but shorter than rachis of the inflorescence; internodes 2.5 – 3.0 cm long; peduncle bracts triangular, 10 – 13 × 0.5 – 1.0 cm, margins of its sheath sparsely spiny, the blade long attenuate, acute, entire, multinerved, reddish at the base, already brownish and dry at anthesis, densely white lepidote on both surfaces, usually twice as long as internodes; rachis (main axis) in a zig-zag fashion, 90 – 100 cm long, 8 – 10 mm in diameter at the base, terete, green with brown stripes, the surface sparsely white lepidote, the internodes 5 – 5.5 cm long; primary bracts triangular, attenuate, apex ending in a spine, 2.5 – 3 × 0.6 – 0.7 cm, margins sparsely spiny, brownish, densely white lepidote abaxially, almost glabrous adaxially, multi-nerved, much shorter than the branches; branches ca. 22 in number, forming an angle of 70 – 90 ° relative to the rachis, 9 – 16 cm long, 2 – 2.5 cm in diameter, (37 –) 78 – 90 flowered; rachis (branch) terete, longitudinally sulcate, 5 – 6 mm in diameter, densely white lepidote, stipe nearly none; floral bracts triangular, short-acuminate, concave, 6 – 9 × 4 mm, green basally, apex light brown, sparsely white lepidote only on the basal abaxial surface, margins hyaline, erose, 7 - nerved, mid nerved and apiculus conspicuous on abaxial surface, dark brown, equaling the sepals in anthesis. Flowers sessile, verticillate, ca. 1 cm long, 4 – 5 mm in diameter at anthesis, fragrant (sweet smell) during the morning; pedicel (a constriction of the floral receptacle) obconic, 0.5 – 1.0 mm long, 0.8 – 1 mm in diameter, light brown, sparsely lepidote; sepals oblong, acute, cucullate, 4 – 6 × 4 mm, green at the base, brownish at the apex, entire or slightly serrate at the apex, glabrous on both sides, or with some sparse trichomes on the basal abaxial surface, multi-nerved; petals oblong, rounded, cucullate, 5.5 – 6 × 4 mm, white, glabrous, multi-nerved, petal apices convergent in anthesis and stamen filaments emerging between them; filaments triangular, flattened, 6 – 9 mm long, 5 mm wide, white; anthers narrowly oblong, 1.5 – 3 mm long, dorsifixed, green, pollen yellow; pistillode reduced, 1.5 – 2.5 mm long, ca. 2 mm in diameter, green, stigmatic lobes much reduced. Pistillate inflorescences a once-divided panicle, in general shape cylindrical, erect, 1 – 1.4 m long; peduncle terete, ca. 66 cm long, 1 – 1.4 cm in diameter at the base, longer than the rosette; internodes 2 – 3.5 cm long; peduncle bracts triangular, long attenuate, acute, 10 – 13 × 0.5 – 1.5 cm, multi-nerved, reddish at the base, brownish and dry even when inflorescence is in anthesis, densely white lepidote on both surfaces; margins of its sheath sparsely spiny, the blade entire; rachis (main axis) ca. 70 cm long, ca. 0.8 – 1 cm in diameter at the base, terete; internodes 1.5 – 3.5 cm long; primary bracts narrow-triangular, acuminate, 2.5 – 4 × 0.7 – 0.9 cm, brown-reddish when fresh, brown when dry, sparsely lepidote abaxially, glabrous adaxially, densely serrate, multi-nerved, much shorter than the branches; branches 20 – 22 in number, in an angle of ca. 45 ° with the rachis, 13 – 25 cm long, 20 – 34 flowered; rachis (branch) with stipe 10 – 12 mm long, terete, channeled, glabrous; floral bracts lanceolate-ovate, acuminate, 7 – 9 mm long, 3.5 – 5.5 mm wide, brown, glabrous except the sparsely white lepidote base adaxially, brown, margin minutely denticulate, about equaling the sepals. Flowers sessile, fragrant (sweet smell) during the morning, 13 – 15 mm long, 4.5 – 5.5 mm in diameter at anthesis; sepals broadly ovate, acute, 3.3 – 4 × 3.3 – 3.6 mm, glabrous on both sides, except some sparsely trichomes on the basal abaxial surface, greenish basally, apically brown, multi-nerved; petals triangular to ovate, acute, 3.5 – 4.3 × 1.5 – 2.1 mm, glabrous on both surfaces, with a few minute trichomes on basal abaxial surface, white, multi-nerved, during anthesis adnate to the ovary and apical portion erect; staminodes six in number, narrowly triangular, laminar, 2 – 2.4 mm long, ca. 0.5 mm at the base; ovary oblongoid, 3.9 – 4.9 mm long, 1.8 – 2 mm in diameter, glabrous, greenish, stigmatic lobes recurved, 1.3 – 1.7 mm long, white. Fruits narrowly ovoid, 11 – 15 mm long, 6 – 8 mm in diameter, base shrinks and resembles a pedicel, glabrous, brown when dry; seeds fusiform, 5.5 – 7.5 mm long, 0.8 – 1.4 mm in diameter, brown, reticulate, caudate, wings 2 – 3.5 mm long, brown reddish.	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFB1FFC3FF6BB139FD61DA6A.taxon	materials_examined	Additional specimens examined: — MEXICO. Jalisco: Municipio Amatitán, barranca de Santa Rita, 1000 m, July 13, 1969, L. M. Villarreal de Puga 3423 ♂ (GUADA!, IBUG!); Municipio San Cristóbal de la Barranca, 3 km al sur de Techaluta, 780 m, July 25, 1995, P. Carrillo-Reyes & R. Ramírez-Delgadillo 22 ♂ (IBUG!, IEB!); arroyo El Tamarindo, 3.5 km al NNO de San Cristóbal de la Barranca, por el Río Cuixtla, 21 ° 04 ’ 00 ” N, 103 ° 25 ’ 00 ” W, 880 m, February 26, 2000, P. Carrillo-Reyes & R. Bello 1136 ♂ (IBUG!, UAMIZ!); Santa Cruz de Enmedio, 0.5 km al NNO de Santa Cruz de Atístique y 8.5 km en línea recta al NNO de San Cristóbal de la Barranca, 21 ° 07 ’ 08 ” N, 103 ° 27 ’ 10 ” W, 940 m, March 25, 2013, P. Carrillo-Reyes & A. Castro-Castro 6937 ♂ (IBUG!); La Mesa de los Caballos, ca. 8 km al NW de San Cristóbal de la Barranca, 1565 m, July 17, 2013, P. Carrillo-Reyes et al. 7001 ♀ (CICY!, IBUG!); 0.5 km al N de Ixcamilpa por el camino a Sta. Cruz, 21 ° 06 ’ 07 ” N, 103 ° 26 ’ 27 ” W, 910 m, September 9, 2014, P. Carrillo-Reyes et al. 7506 ♀ (CICY!, IBUG!); 1 km en línea recta al N de El Escalón, 21 ° 00 ’ 37 ” N, 103 ° 25 ’ 24 ” W, 1050 m, March 26, 2015, P. Carrillo-Reyes & C. J. Ramírez-Díaz 7714 ♀ (IBUG!); 0.5 km en línea recta al N de El Escalón, km 30 carretera Guadalajara-Colotlán, 1040 m, January 22, 2016, P. Carrillo-Reyes et al. 8221 ♀ (IBUG!); Municipio Zapopan, cerca de la ranchería La Coronilla del Ocote, llegando por Tesistán y Santa Lucía, 20 ° 56 ’ 07 ” N, 103 ° 36 ’ 03 ” W, 1260 m, June 19, 2000, J. A. Lomelí et al. 3265 fruits (IBUG!, UAMIZ!). Nayarit: Municipio Amatlán de Cañas, Cerca de Los Barrenitos, ca. 2 km en línea recta al NE de Amatlán de Cañas, 20 ° 49 ’ 29 ” N, 104 ° 23 ’ 29 ” W, 805 m, March 28, 2018, P. Carrillo-Reyes & D. Cabrera-Toledo 8880 - A ♀ (IBUG!); 8880 - B ♂ (IBUG!). Zacatecas: Municipio Trinidad García de la Cadena, El Abra, 5 km en línea recta al SW de García de la Cadena, 21 ° 10 ’ 03 ” N, 103 ° 29 ’ 22 ” W, 1760 m, August 24, 2014, P. Carrillo-Reyes et al. 7470 fruits (CICY!, IBUG!).	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFB1FFC3FF6BB139FD61DA6A.taxon	biology_ecology	Phenology: — Staminate plants have been collected in January, March and July; female flowers are documented during January-March and July-September, while fruits have been collected during June-August.	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFB1FFC3FF6BB139FD61DA6A.taxon	distribution	Distribution and habitat: — Hechtia jaliscana has been collected in the municipality of Amatlán de Cañas, in the State of Nayarit, municipalities of Amatitán, San Cristóbal de la Barranca, and Zapopan in the State of Jalisco, and municipality of Trinidad García de la Cadena, in Zacatecas, Mexico. Most of the localities are in the basin of the Santiago River, except for those of Amatlán de Cañas, Nayarit, which is in the Ameca river basin, all of them are situated in the Pacific Lowlands biogeographic province according to Morrone et al. (2017), in the low elevation valleys of the southern portion of the Sierra Madre Occidental province (Figure 1). Individuals of this species grow as lithophytes, mostly in northern exposure rocky slopes, inside tropical deciduous forest vegetation (Rzedowski 1978), at elevations between 750 – 1750 m, frequently accompanied by individuals of Bursera fagaroides (Kunth) Engler, Casearia corymbosa Kunth, Ceiba aesculifolia (Kunth) Britten & Baker f., Celtis caudata Planch., Haematoxylum brasiletto H. Karst., Opuntia fuliginosa Griffiths, Pachycereus pecten-aboriginum Britton & Rose, Pseudobombax palmeri (S. Watson) Dugand, Stenocereus queretaroensis (F. A. C. Weber) Buxb. In some localities, H. jaliscana grows in pine-oak forest dominated by Quercus resinosa Liebm., Q. praineana Trel., and Pinus oocarpa Scheid., inhabiting rocky slopes alongside species as Agave guadalajarana Trel., A. rzedowskiana P. Carrillo, Vega & R. Delgad., A. schidigera Lem., Mammillaria scrippsiana (Britton & Rose) Orcutt, species of the genera Muhlenbergia Schreb., Opuntia Mill., and Selaginella P. Beauv. (Figure 2).	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFB1FFC3FF6BB139FD61DA6A.taxon	discussion	Discussion: — Hechtia jaliscana (Figures 2 and 3) is recognized by the cespitose rosettes, thick succulent leaves, glabrous and shiny above, and white lepidote below, armed with coarse spines. The inflorescence, a once-divided panicle in both sexes (Figure 3 A), erect to slightly pending, is produced by young, newly formed rosettes. Flowers are fragrant during morning hours, with a sweet scent that attract bees (Figure 3 B). The rachis of the pistillate inflorescence is sinuose to almost fractiflex (Figure 2 A) while it is slightly sinuose in staminate inflorescences (Figure 2 B), with branches in almost 90 ° angle with the rachis (main axis); flowers open, but the petal apices converge and form a closed corolla with the stamens protruding in between (Figure 3 D, F), while petals on female flowers remain adnate to the ovary with the three stigmatic lobes exerted (Figure 3 E). The rachis of pistillate branches is longitudinally sulcate (Figure 3 G) with nearly sessile flowers, with a pedicel (rather the floral receptacle that shrinks and looks like a pedicel) when in fruit (Figure 3 H), a common phenomenon in species of Hechtia. IUCN Conservation assessment: — The EOO (1434 km 2) and AOO (44 km 2) calculated from 13 collections corresponding to 12 localities with the GeoCAT (Bachman et al. 2011), suggest that H. jaliscana is an Endangered (EN) species, according to the IUCN B set of criteria.	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBAFFCBFF6BB0B3FCEFD32A.taxon	materials_examined	TYPE: — MEXICO: Durango: Municipio Nombre de Dios, adelante de Nombre de Dios, camino a El Venado, 11 km del desvío, matorral de encino-pino, sobre laderas en suelo volcánico, 23 ° 47 ’ 55 ’’ N, 104 ° 17 ’ 40 ’’ W, 1745 m, originally collected by I. Ramírez, A. Espejo, A. López-Ferrari, A. Mendoza & J. Ceja, September 2005, flowering under cultivation, June 2009, I. Ramírez & G. Carnevali 1627 ♂ (Holotype, CICY!).	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBAFFCBFF6BB0B3FCEFD32A.taxon	diagnosis	Diagnosis: — Hechtia marthae is similar to H. subalata but differs in its pistillate branches much shorter than the primary bracts (vs. conspicuously larger than the primary bract), with shorter branches (2 – 5 vs. (2 –) 5 – 15 cm long), floral bracts narrowly triangular, 2 – 5 × 1 mm (vs. broadly ovate, 3 – 4.5 × 3 – 4 mm), ovary 1 / 3 inferior (vs. completely superior), with longer petals (5 – 7 vs. 3 – 5 cm long).	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBAFFCBFF6BB0B3FCEFD32A.taxon	description	Plants lithophytic or terrestrial, cespitose, 1.1 – 1.75 m height when flowering; rosettes generally round, 30 – 40 cm tall, 35 – 45 cm in diameter, generally forming dense, colonies of 5 – 21 rosettes. Leaves 20 – 30 in number, central ones erect to slightly reflexed in basal ones; sheath ovate to broadly ovate, 2.8 – 6.5 × 2 – 6.3 cm, light brown, margins entire and erose distally, lustrous and glabrous at the base, slightly lepidote distally; blade narrowly triangular, acuminate to long attenuate, 20 – 33 (– 50) × 1.3 – 4 cm, green, reddish toward the apex and margin when sun exposed, densely white lepidote abaxially, white lepidote adaxially; marginal spines antrorse and retrorse, triangular, 2 – 5 mm long, 6 – 20 mm apart, brown, with a short tuff of white trichomes in the concave portion of the spine. Inflorescence central, erect, emerging from a mature rosette (strict sympodium growth pattern, type SPP sensu Ramírez-Morillo et al. 2014). Staminate inflorescences a once-divided panicle, cylindrical in general shape, erect, 0.7 – 1.35 m long; peduncle terete, green to light brown, slightly white lepidote, 33 – 79 cm long, 8 – 17 mm in diameter at the base, longer than the rosette; internodes 0.8 – 2.5 cm long; peduncle bracts the basal ones, with ovate sheath, 2 – 3 × 1.5 – 2 cm, the blades narrow triangular, 9 – 24 × 0.9 – 1.3 cm, the distal bracts with a sheath rectangular to triangular, 4 – 7 × 0.8 – 1.4 cm, margins entire then erose and spiny toward the blade, this long attenuate to acuminate, multi-nerved, brownish, densely white lepidote abaxially, glabrate or slightly lepidote adaxially, longer than internodes, clasping the peduncle; rachis (main axis) 33 – 62 cm long, 3 – 9 mm in diameter at the base, terete, green to brownish; internodes 0.8 – 3.6 cm long, white lepidote; primary bracts rectangular to triangular, abruptly acuminate, 4 – 5 × 1.2 – 1.4 cm, entire at the base to erose and spiny toward the apex, brown when dry, sparsely white-lepidote abaxially, glabrous to slightly lepidote adaxially, multi-nerved, about the length of the branches; branches 18 – 27 in number, forming an angle of 15 ° or less with the rachis, (1.8 –) 4 – 7.5 cm long, ca. 5 mm in diameter, 16 – 29 flowered; rachis (branch) flattened at its base, 0.2 – 1 cm long, glabrous, stipe; floral bracts inconspicuous, narrowly-triangular, acuminate, 1.5 – 2 (– 4) × 0.1 mm, green, brownish at the apex, slightly lepidote, inconspicuously nerved, equaling or slightly exceeding the pedicel. Flowers pedicellate, polystichous, 7 – 9 mm long, 3 – 5 mm in diameter, fragrant during the morning (sweet smell); pedicel obconic, 2 – 3 (– 4) mm long, ca. 1 mm in diameter, light green, slightly lepidote; sepals ovate to triangular, acute, 3 – 4 × 1.8 – 3 mm, entire, green at the base, apically brownish, slightly lepidote at the base, 3 - nerved, shorter than petals; petals elliptic, acute, 5 – 6 × 2 – 3.5 mm, green with brownish spots, multi-nerved; filaments triangular, terete, 4 – 5 mm long, ca. 1 mm in diameter, white to greenish; anthers oblong, 2 – 3 mm long, dorsifixed, green, pollen yellow; pistillode reduced, 0.5 – 0.8 mm long × 1.5 mm in diameter, light green, stigmatic lobes much reduced. Pistillate inflorescences a once-divided panicle, in general shape cylindrical, erect, 0.83 – 1.12 m long; peduncle terete, 40 – 55 cm long, 0.8 – 1.9 cm in diameter at the base, longer than the rosette; internodes 1 – 2.5 cm long; peduncle bracts without clear distinction between sheath and blade, basal ones with ovate sheath, blade triangular and long acuminate, 5 – 8.5 × 1.8 – 2.6 cm, margins entire at the base to erose and spiny distally, brownish, densely white lepidote abaxially, glabrate or slightly lepidote adaxially, multi-nerved; longer than internodes, clasping the peduncle; rachis (main axis) 30 – 57 cm long, ca. 0.7 – 1.1 cm in diameter at the base, terete, green, internodes 1 – 3 cm long; primary bracts rectangular to ovate-triangular, acuminate, 2 – 4 × 0.8 – 2 cm, brown when dry, sparsely white-lepidote abaxially, glabrous to slightly lepidote adaxially, entire at the base to erose and spiny toward the apex, multi-nerved, as long or shorter than the branches; branches 15 – 25 in number, in an angle of ca. 45 ° with the rachis, 2 – 5 cm long, ca. 1.3 – 1.8 cm in diameter (up to 4 cm when in fruit) 5 mm in diameter, 10 – 20 flowered; rachis (branch) terete, white lepidote, glabrous in the infructescence, stipe nearly none; floral bracts inconspicuous, narrowly-triangular, acuminate, 2 – 5 × 1 mm, green, white lepidote, margins serrate. Flowers pedicellate, verticillate, fragrant during the morning (sweet smell), 10 – 12 mm long, ca. 5 – 7 mm in diameter; pedicels which are a constriction of the base of the floral receptacle resembling a pedicel, obconic, 2 – 3 mm long, 1 – 2 mm in diameter, tomentose; sepals ovate to triangular, acute, 4 – 5 × 3 – 4 mm, green, slightly lepidote abaxially, entire, 3 - nerved; petals elliptic, acute, 5 – 7 × 3 – 4 mm, green, glabrous, entire, multi-nerved; staminodes six in number, narrowly triangular, laminar, 2.5 – 2.9 mm long, ca. 1 mm at the base; ovary 1 / 3 inferior, oblongoid, 7 – 9 mm long, 3 – 4 mm in diameter, glabrous, green, stigmatic lobes recurved, ca. 1 mm long, pale green. Fruits pedicellate (rather the base of the floral receptacle shrinks and resembles a pedicel), ellipsoid, 10 – 14 mm long, 5 – 8 mm in diameter, glabrous, green when mature, brown when dry; seeds fusiform, 3.5 – 5 mm long, 1 – 1.5 mm in diameter, brown, reticulate, caudate, wings ca. 1 mm long, brownish.	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBAFFCBFF6BB0B3FCEFD32A.taxon	materials_examined	Additional specimens examined: — MEXICO. Durango: Municipio Mezquital, Temoaya, a 4.5 km por el camino a Agua Zarca, 23 ° 20 ’ N, 104 ° 30 ’ W, matorral subtropical, 1650 m, August 18, 1983, M. González E. s. n. fruits (CIIDIR!, HUAA!, UAMIZ!, UJAT!); Temoaya, 2.5 km al noreste por la carretera a Mezquital, 23 ° 18 ’ 59 ’’ N, 104 ° 29 ’ 19 ’’ W, matorral subtropical, 1704 m, May 25, 2020 (cultivated), R. Quirino-Olvera 143 ♀ (CICY!); Municipio Nombre de Dios, Chachacuaxtle, 33 km al SSW de Nombre de Dios, 22 ° 40 ’ N, 104 ° 16 ’ W, matorral subtropical, 1740 m, March 21, 1988, S. González E. 4086 fruits (CIIDIR!, HUAA!, UAMIZ!, UJAT!); ca. 10 km sobre la brecha a El Venado, a partir de la carretera Nombre de Dios-Durango, 23 ° 47 ’ 55 ’’ N, 104 ° 17 ’ 40 ’’ W, matorral espinoso, 1739 m, July 25, 2003, A. R. López-Ferrari et al. 2989 fruits (UAMIZ!, UJAT!); 2993 ♂ (UAMIZ!, UJAT!); ca. 10 km en la brecha que va de la carretera Durango-Nombre de Dios hacia El Venado, 23 ° 47 ’ 54.3 ’’ N, 104 ° 17 ’ 38.8 ’’ W, cañada con matorral xerófilo, 1751 m, June 18, 2009, I. Ramírez 1627 ♂ (CICY!, WU!); same locality, June 3, 2013, I. Ramírez 2205 ♂ (CICY!); same locality, October 31, 2019, I. Ramírez et al. 2442 fruits (CICY!); Municipio Topia, canyon below Topia, just downstream from Los Molinos, on cliff, 3800 feet alt., June 1978, M. Kimnach & H. Sánchez-Mejorada 1777 ♂ (HNT!, US!).	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBAFFCBFF6BB0B3FCEFD32A.taxon	biology_ecology	Phenology: — Staminate plants have been collected in June, pistillate plants only reported for May, while fruiting specimens have been collected from March to October.	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBAFFCBFF6BB0B3FCEFD32A.taxon	distribution	Distribution and habitat: — Populations of Hechtia marthae has been found thus far only on four localities in the western slope of the Sierra Madre Occidental, in the state of Durango, in Mezquital, Nombre de Dios, and Topia municipalities. Its distribution matches partially with the extensions of the Pacific Lowlands province that penetrate the lowlands of the Sierra Madre Occidental province (Morrone et al. 2017). It grows at elevation between 1158 – 1751 m, on xerophytic shrublands and subtropical scrub along with species of Agave L., Dasylirion Zucc., Fouquieria Kunth, Opuntia, Prosopis L., and Vachellia Wight & Arn. (Figure 5).	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBAFFCBFF6BB0B3FCEFD32A.taxon	etymology	Eponomy: — The species is named after Martha González-Elizondo in recognition for her outstanding contribution to the knowledge on the Flora of Durango and for collecting material of the species. IUCN Conservation assessment: — The GeoCAT (Bachman et al. 2011) indicates that, based upon the EOO (7953 km 2) and AOO (28 km 2), H. marthae is classified as Vulnerable (VU) and Endangered (EN), respectively, according to the IUCN B set of criteria. In order to better understand the conservation status of this species robustly, it is necessary to evaluate demographic aspects of its populations and possible threats.	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBAFFCBFF6BB0B3FCEFD32A.taxon	discussion	Discussion: — Populations included here in Hechtia marthae were misidentified previously as Hechtia subalata or H. podantha Mez (1896: 549), due to a holotype that only represents an apical portion of an immature infructescence (see Table 1 for comparisons of main differences among the three species). As many species of Hechtia, blooming periods are very short and flowers in an inflorescence are open for 1 – 3 days, making difficult to document floral structures which show more accurately differences among species (such as colors, shapes, fragrances, etc.). This species shows wide and relatively long primary bracts, lepidote, with condense pistillate branches resembling species of the Hechtia podantha complex (Martínez-Correa 2008), probably a case of convergent evolution grouping species from different lineages (Ramírez-Morillo et al. 2018 a) invading high elevation dry habitats occasionally exposed to frosts. Hechtia marthae, H. podantha and H. subalata are very similar in several features: they share the central inflorescence that emerges from a fully formed rosette, inflorescences in both sexes have their primary branches subtended by a conspicuous primary bract, branches are usually condensed, and flowers are usually green, as well as the fact that inflorescences are in bloom for 2 – 3 days, a common feature of Hechtioideae but with few exceptions. Main differences among the three species are depicted on Table 1.	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBEFFD7FF6BB15BFCAAD026.taxon	materials_examined	TYPE: — MEXICO: Durango, Sierra Madre, August 13, 1897, J. N. Rose 3467 fruits (holotype, US!; photograph and fruits, GH!). EPITYPE (designated here): — MEXICO: Zacatecas: Municipio Valparaíso, 1 km SE del Puente Tepetatita, 26 km W de Huejuquilla el Alto rumbo a San Juan Capistrano, 22 ° 40 ’ 36.1 ’’ N, 104 ° 03 ’ 03.3 ’’ W, 1078 m, matorral xerófilo con Prosopis, Jatropha, Croton, Myrtillocactus, originally collected May 27, 2015 by P. Carrillo-Reyes, J. L. Tapia, C. J. Ramírez-Díaz and I. Ramírez; flowering in cultivation in 2019, I. Ramírez et al. 2430 ♂ (epitype CICY!; isoepitypes, IBUG!, MEXU!).	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBEFFD7FF6BB15BFCAAD026.taxon	description	Plants lithophytic, cespitose, 1.4 – 1.8 m when flowering; rosettes in general shape globose, 40 – 60 cm tall, 50 – 60 cm in diameter, generally forming dense, small colonies of 2 – 6 rosettes, rarely clumps of 8 – 12 rosettes. Leaves 20 – 35 in number, central ones erect, basal ones slightly reflexed; sheath broadly ovate to widely elliptical, 4 – 5.2 × 4.5 – 7 cm, margins entire to erose toward the apex, lustrous and glabrous at the base adaxially and slightly lepidote at the apex, white lepidote abaxially; blade narrowly triangular, acuminate to long attenuate, 20 – 50 × 2.5 – 4.8 cm, green, sometimes with red spots at the apex and or margins, at the base of every spine, densely white lepidote abaxially, white lepidote at base but soon glabrous and glossy adaxially; marginal spines retrorse rarely antrorse, triangular, 1.5 – 2.5 mm long, 0.8 – 2.5 cm apart, light brown or occasionally with red spots close to the spines. Inflorescence central, erect, emerging from a fully grown rosette (strict sympodium growth pattern, type SPP sensu Ramírez-Morillo et al. 2014). Staminate inflorescences a once-divided panicle, cylindrical in general shape, erect, 0.65 – 1.8 m long; peduncle terete, green to brownish, sparsely lepidote, 20 – 56 cm long, 5 – 14 mm in diameter at the base, surpassing the rosette; internodes (0.8 –) 1.1 – 4.8 cm long; peduncle bracts the basal ones with wide-ovate sheath, 1 – 2.2 × 1.1 – 7 cm, the blade narrow triangular, long attenuate, slightly pungent, (3.5 –) 8.7 – 17 × 0.3 – 0.7 cm, green to brownish, the apical ones without a clear distinction between sheath and blade, ovate to triangular, long attenuate to acuminate, 2.2 – 4.5 × 0.8 – 1.1 cm, margins entire to erose and spiny toward the blade multi-nerved, brownish when dry, white lepidote abaxially, glabrate adaxially, longer than internodes; rachis (main axis) 0.44 – 1.32 m long, 5 – 10 mm in diameter at the base, terete, brownish; internodes 1 – 3.4 cm long; primary bracts ovate-triangular, acuminate, 1.2 – 4 × 0.8 – 1.4 cm, entire to dentate, light brown, densely lepidote abaxially, slightly lepidote adaxially, multi-nerved, shorter than branches; branches 16 – 59 in number, forming an angle of 45 ° or less with the rachis, 2.5 – 10 (– 15) cm long, ca. 1.5 cm in diameter, 28 – 72 flowered; rachis (branch) terete, light brown, 7 – 8 cm long, 0.8 – 1.2 mm in diameter, densely lepidote, stipe nearly none; floral bracts broadly ovate, acute, 4.5 – 6 × 2.5 – 4.5 mm, brown, margins erose to dentate, glabrous on both surfaces, multi-nerved, sometimes with a conspicuous central nerve, equaling or slightly surpassing the sepals. Flowers sessile, polystichous, divaricate, fragrant during the morning (sweet smell); sepals ovate to triangular, acute, cucullate, 3 – 4 × 2 – 2.5 mm, entire to slightly erose, brownish, glabrous on both surfaces, multi-nerved; petals elliptical, acute to rounded, cucullate at the apex, 4 – 5 × 2 – 3 mm, white with brown spots, glabrous on both surfaces, multinerved; stamens exserted, ca. 4.5 mm long; filaments triangular, flattened, ca. 4 mm long, white; anthers oblong, 2 – 3 mm long, dorsifixed, yellow, pollen yellow; pistillode reduced, 1.5 × 2 mm, stigmatic lobes much reduced. Pistillate inflorescences a once-divided panicle, cylindrical in general shape, erect, 1.2 – 1.7 m long; peduncle terete, 0.35 – 1.18 m long, 8 – 20 mm in diameter at the base, surpassing the rosette; internodes (1 –) 2 – 4 cm long; peduncle bracts the basal ones with triangular sheath, 1.5 – 25 × 1.4 – 1.8 cm, the blades narrow triangular, long attenuate, slightly pungent, 10 – 27 × 0.5 – 0.7 cm, foliaceous, green to brownish, the apical ones without a clear distinction between sheaths and blades, ovate to triangular, 2 – 3.5 × 0.9 – 1.2 cm, margins entire to erose and spiny toward the blade, long attenuate to acuminate, brownish when dry, white lepidote abaxially, glabrate adaxially, multi-nerved, longer than internodes; rachis (main axis) 63 – 73 cm long, 6 – 9 (– 14) mm in diameter at the base, terete, green; internodes (1 –) 2 – 3.5 cm long; primary bracts triangular, acuminate, 1.5 – 2.5 × 0.8 – 1.5 cm, light brown, glabrous on both surfaces, entire to dentate, multi-nerved, shorter than branches; branches 22 – 32 in number, forming an angle of 45 ° or less with the rachis, (2 –) 5 – 15 cm long, 4 – 8 mm in diameter, with 10 – 44 flowered, flowers occasionally emerging from the base of the branch; rachis (branch) terete, green, 0.4 – 0.8 cm long, 2 – 4 mm in diameter, glabrescent, stipe nearly none; floral bracts broadly ovate, acute, 3 – 4.5 × 3 – 4 mm, without covering or exceeding the sepals, margins erose to dentate, glabrous on both surfaces, multi-nerved, sometimes with a conspicuous central nerve, brown, membranous. Flowers sessile, polystichous, divaricate, fragrant (sweet smell during the morning); sepals ovate, acute, 2 – 3.5 mm long, ca. 2.5 mm wide at the base, entire, green to brownish, glabrous on both surfaces, multi-nerved; petals triangular, acute, cucullate at apex, 3 – 4.5 mm long, ca. 2 mm wide at the base, entire, white with brown spots, multi-nerved; staminodes six in number, triangular, laminar, 1.5 – 2 mm long, ca. 0.5 mm wide, inserted in the corolla; ovary superior, oblongoid to ellipsoid, ca. 7 mm long, 3 mm in diameter, light green, glabrous, stigmatic lobes recurved, ca. 1 mm long, light green, placentation axillary. Fruits ellipsoid, 9 – 15 mm long, 6 – 7 mm in diameter, glabrous, erect, brown when dry; s eeds narrowly triangular, brown, 3.5 – 4 mm long, 1 mm wide, caudate, apical wing ca. 2 mm long, basal wing reduced, brown.	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBEFFD7FF6BB15BFCAAD026.taxon	materials_examined	Additional specimens examined: — MEXICO. Jalisco: Municipio Huejuquilla el Alto, 3.6 km al NW del poblado Los Arroyos del Agua, km 17.5 de la carretera Huejuquilla el Alto-Ruiz, 22 º 40 ’ 30.7 ’’ N, 103 º 59 ’ 21.2 ’’ W, 1260 m, October 6, 2018, D. Figueroa et al. 235 ♂ (IBUG), 236 ♀ (IBUG!); Municipio Bolaños, Bolaños, orilla del pueblo, 21 ° 49 ’ 48.0 ” N, 103 ° 46 ’ 48.0 ” W, 885 m, November 7, 2018, K. Romero-Soler et al. 1239 fruits (CICY!, IBUG!), 1240 ♂ (CICY!); Municipio Chimaltitán, foothills 8 – 12 km SW of Bolaños, R. McVaugh 25929 ♀ (MICH!); Municipio Mezquitic, 3 km al E de Tetakararo, bajando la barranca, 800 m, November 10, 1985, C. Chávez-Reyes D. 169 ♀ (IBUG!); Municipio San Martín de Bolaños, 1 km al S del Rancho La Joya, brecha San Martín al Platanito, 900 m, May 18, 1989, A. Flores et al. 1583 fruits (XAL!); Municipio Villa Guerrero, 11.2 km al oeste de Villa Guerrero camino a Bolaños, 21 ° 58 ’ 03 ” N, 103 ° 40 ’ 36 ” W, 1780 m, November 6, 2018, K. Romero-Soler et al. 1238 fruits (CICY!); 11.2 km al WSW de Villa Guerrero por la carretera a Bolaños (7.7 km en línea recta), 21 ° 58 ’ 03 ” N, 103 ° 40 ’ 36 ” W, 1780 m, April 4, 2017, P. Carrillo-Reyes et al. 8520 fruits (IBUG!). Nayarit: Municipio El Nayar, arroyo El Fraile, al SE de la Mesa del Nayar, 22 ° 09 ’ N, 104 ° 33 ’ W, 520 m, October 23, 1989, P. Tenorio & G. Flores 16620 ♀ (MEXU!, MO!); Vereda de la Mesa del Nayar al Cangrejo, que cruza por la barranca, 900 – 1400 m, August 5, 1990, R. Ramírez et al. 551 ♀ ♂ (MEXU!). Zacatecas: Municipio Valparaíso, 1 km SE del Puente Tepetatita, 26 km W de Huejuquilla el Alto rumbo a San Juan Capistrano, 22 ° 40 ’ 36.1 ’’ N, 104 ° 03 ’ 03.3 ’’ W, 1078 m, May 27, 2015; flowering under cultivation in 2018, I. Ramírez et al. 1989 fruits (CICY!); I. Ramírez et al. 1989 a ♂ (CICY!, IBUG!, MEXU!); San Juan Capistrano, August 23, 1883, J. Rose 3556 fruits (HUH!, NY!); 16.8 km al W de Huejuquilla rumbo a San Juan Capistrano, alrededores del río Atengo o Chapalagana, 1019 m, March 24, 2010, A. Castro-Castro 2027 ♀ (IBUG!); Valley of río Atenco (Chapalagana), 8 – 15 km northeast of San Juan Capistrano, road to Huejuquilla el Alto, sheltered rocky canyon, and adjoining arid flats, 1000 – 1200 m, January 14, 1975, R. McVaugh 25795 ♀ (MEXU!); 6 miles before San Juan Capistrano on road to Jesus María, 1036 m, January 22, 1976, M. Kimnach 1855 ♀ (MEXU!); aprox. 2 km antes de llegar a Chimaltitán, después de Florencia, 790 m, May 20, 1985, J. A. Lomelí-Sención 85 - 00190 ♂ (GUADA!).	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBEFFD7FF6BB15BFCAAD026.taxon	biology_ecology	Phenology: — Staminate plants have been collected in January, March, August, October and November; female flowers are documented during May, August, October, and November; while fruits have been collected during April, May, August, and November.	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBEFFD7FF6BB15BFCAAD026.taxon	distribution	Distribution and habitat: — Hechtia subalata has been collected in the Mexican States of Durango (Municipality of Mezquital), Jalisco (Bolaños, Chimaltitán, Huejuquilla el Alto, San Martín de Bolaños, and Mezquitic and Villa Guerrero Municipalities), Nayarit (Del Nayar Municipality) and Zacatecas (Valparaíso Municipality). According to the biogeographical provinces of Morrone et al. (2017), this taxon occurs in the Pacific Lowlands province, particularly in canyons that penetrate the Sierra Madre Occidental province (Figure 1), at 800 – 1300 m of elevation, growing on rocky slopes, within tropical dry forest, xerophytic shrubland and ecotones between those vegetation types with species of Agave, Bursera bipinnata (DC.) Engl., B. fagaroides, Coryphantha (Engelm.) Lem., Croton L., Dodonaea viscosa (L.) Jacq., Echinocereus pamanesiorum Lau, Euphorbia schlechtendalii Boiss., Forestiera phillyreoides (Benth.) Torr., Heliocarpus terebinthinaceus (DC.) Hochr., Ipomoea murucoides Roem. & Schult., Leucaena Benth., Myrtillocactus geometrizans (Mart. Ex Pfeiff.) Console, Opuntia, Prosopis laevigata (Humb. & Bonpl. ex Willd.) M. C. Johnst., Stenocereus queretaroensis, and Vachellia pennatula (Schltdl. & Cham.) Seigler & Ebinger (Figure 6).	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBEFFD7FF6BB15BFCAAD026.taxon	conservation	IUCN Conservation assessment: — Hechtia subalata is known from 17 collections (ten localities); most of them by roadsides, and most likely occurs as isolated populations on appropriate microniches, rocky slopes in rugged canyons. It has values of EOO de 6592.7 km ² and an AOO of 44 km ². According to the IUCN, therefore this species should be considered as Vulnerable (VU) to Endangered (EN); nevertheless, because of the lack of population information, we rely mostly upon distributional data, namely, the set of B criteria, geographical distribution.	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
03978787FFBEFFD7FF6BB15BFCAAD026.taxon	discussion	Discussion: — Species of Hechtioideae, in general and those of Hechtia in particular, are still poorly represented in herbaria: nothing is more discouraging for the collector than spiny, usually large rosettes with large inflorescences. Most collectors usually find species in fruit because flowers last only one day and even inflorescences with tens or hundreds of flowers, last but a few days. It is also known that populations of some species do not bloom every year, especially those with strict monocarpic growth pattern (SMP, sensu Ramírez-Morillo et al. 2014) where long-lived, well-developed rosettes take years to produce an inflorescence (for example Hechtia zamudioi Espejo, López-Ferr. & I. Ramírez (2008: 55 )). Species more frequently found in bloom, year after year, are those with sympodial growth, with precocious-flowering pattern (i. e. Hechtia rosea E. Morren ex Baker (1889: 140 )). We still do not know what triggers plants to produce inflorescences. In the case of Hechtia subalata, as for ca. 26 % of the described species, type specimens only have fruits, and few of them included additional information on flowers or rosettes in their protologues. For H. subalata, few features are discernable in the holotype (fruit size, inflorescence branching, branch length, etc.) or the paratype (foliar blade, particularly the margins: sinuose-concave between spines, and spines alternating on both margins). These data, combined with its geographical distribution, allow us to finally complete our understanding of this concept and assign a more precise (other than what is found in type specimens) geographical distribution. It is important to mention that the female flowers of H. subalata (Figure 7 B) are practically identical to those of H. jaliscana (Figure 3 E), but their arrangement on the branches (agglomerated in the first and lax in the second) differentiates these two taxa. It is important to emphasize that this close similarity of female flowers between both species is another reason to propose epitypites based on a staminate specimen.	en	Ramírez-Morillo, Ivón, Carrillo-Reyes, Pablo, Tapia-Muñoz, José Luis, Ramírez-Díaz, Claudia J. (2021): In disentangling two species limits of Hechtia (Bromeliaceae: Hechtioideae) from Sierra Madre Occidental, a new species is discovered from Durango, Mexico. Phytotaxa 523 (1): 55-72, DOI: 10.11646/phytotaxa.523.1.3
