taxonID	type	description	language	source
039787E2FFCAFFC8ED9127132D3BF91C.taxon	materials_examined	Material examined. CANADA: ALBERTA: Lusk Creek, Kananaskis Forest Experiment Station, Seebe, 18 June 1968, H. J. Teskey, 1 ♂, 1 ♀ (BYUC). BRITISH COLUMBIA: Chilliwack River, Vedder Crossing, 24 April 1938, S. Spencer, 1 ♀ (BYUC). Preacher Creek, Crawford Creek Rd, 31 May 2010, B. Stark, R. W. Baumann, 1 ♂ (BPSC). Princeton, Whip-saw Creek, 26 May 1968, C. H. Lindroth, 1 ♂ (USNM). Summit Creek, Hwy 3, Blazed Creek Rest Area, 2 June 2010, R. W. Baumann, B. P. Stark, 1 ♀ (BYUC). USA: ALASKA: Juneau, tributary of McGinnis Creek, 8 June 1979, R. N. Vineyard, 1 ♀ (BYUC). Juneau, Granite Creek, 58.3184 ° N, 134.3236 ° W, 25 June 2011, J. P. Hudson, 2 ♀ (BYUC). Taiya River north of Skagway, 1 July 2011, J. P. Hudson, 1 ♂ (BYUC). Prince of Wales Island, Tracadora Creek, 17 June 1996, unknown collector, 1 ♀ (BYUC). CALIFORNIA: Butte Co., Butte Creek, 1 mi above Cherry Hill Campground, 9 May 1987, Stanger, Maxfield, 1 ♂, 4 ♀ (BYUC). El Dorado Co., Long Canyon Creek, 3 mi E Grizzly Flats, 16 May 1998, B. Stark, C. R. Nelson, S. W. Szczytko, R. Bottorff, I. Sivec, 1 ♀ (BPSC). El Dorado Co., North Cosumnes River, Sciaroni Crossing, Grizzly Flats, 21 April 1984, R. Bottorff, 3 ♂, 2 ♀ (RLBC). Humboldt Co., creek, 0.3 mi W Ruby Creek, Hwy 299, 25 April 1987, R. W. Baumann, B. Stark, C. R. Nelson, S. Wells, 1 ♀ (BYUC). IDAHO: Bonner Co., Granite Creek, Pend Oreille Lake, 15 May 1958, S. G. Jewett, Jr. 1 ♀ (BYUC). MONTANA: Broadwater Co., Deep Creek, Hwy 12, 15 May 1969, M. L. Miner, 1 ♂ (BYUC). Cascade Co., Belt Creek, 19 mi SE Monarch, 7 July 1966, J. R. Grierson, 1 ♀ (USNM). Fergus Co., Windsor Creek, Windsor Rock Campground, 3 July 1952, R. Hays, 3 ♂ (BYUC). Flathead Co., McDonald Creek, Glacier National Park, 15 July 1967, A. R. Gaufin, 1 ♀ (BYUC). Gallatin Co., Hyalite Creek, 1 June 1951, R. Hays, J. Bailey, 1 ♂, 1 ♀ (BYUC). Glacier Co., Swift Current Creek, Glacier National Park, 24 July 1967, A. R. Gaufin, 1 ♂ (BYUC). Glacier Co., Waterton River, Glacier National Park, 23 July 1970, C. Yarmoloy, 1 ♀ (BYUC). Lake Co., Yellow Bay Creek, 25 April 1970, R. Haick, 1 ♀ (BYUC), 1 ♂, 3 ♀ (USNM). Lewis and Clark Co., tributary Blackfoot River near Roger’s Pass, Hwy 20, 17 May 1969, A. R. Gaufin, 1 ♀ (BYUC). Lincoln Co., Ross Creek, FR 398, Cedars Scenic Area, 27 April 2008, R. Durfee, 3 ♂, 3 ♀ (CSUC). Mineral Co., Little Joe Creek, Little Joe Campground, 25 May 2010, B. Stark, R. W. Baumann, 2 ♀ (BPSC). Missoula Co., Butler Creek, Snow Bowl, 12 May 1973, R. A. Haick, 9 ♂, 4 ♀ (USNM). Missoula Co., Grant Creek, Snow Bowl Rd, 9 May 1969, D. S. Potter, 1 ♀ (BYUC). Missoula Co., Rattlesnake Creek, Missoula Reservoir, 16 May 1969, R. W. Baumann, 1 ♂ (BYUC). Powell Co., Arrastra Creek, Hwy 200, 10 May 1969, M. L. Miner, 5 ♀ (BYUC). Ravalli Co., Roaring Lion Creek, 18 May 1998, A. L. Sheldon, 5 ♂ (BYUC). Ravalli Co., South Gash Creek, Rd 737, 30 May 1996, A. L. Sheldon, 1 ♀ (BYUC). OREGON: Deschutes Co., Tumalo Creek, below Tumalo Falls, 11 June 2004, R. W. Baumann, B. P. Stark, 2 ♂ (BYUC). Douglas Co., Muir Creek, Hwy 230, 19 May 1982, B. Stark, D. Zeigler, 1 ♂, 1 ♀ (BPSC). Hood River Co., Cold Spring Creek, nr. Elk Meadows, 23 July 1983, G. R. Fiala, 1 ♀ (BYUC). Linn Co., Puzzle Creek, Marion Forks Rd, 2 June 2000, B. Stark, I. Sivec, M. C. Zúñiga, 1 ♂ (BPSC). Umatilla Co., 11 mi E Milton-Freewater, 16 April 1952, S. G. Jewett, Jr., 2 ♀ (BYUC). Union Co., Five Points Creek, 28 April 1961, S. G. Jewett, Jr., 13 ♂ (USNM). Union Co., Five Points Creek, Grande Ronde River below Hilgard Junction, 18 June 1977, R. W. Baumann, 1 ♀ (BYUC). Union Co., Grande Ronde River, Bird Track Springs Campground, Hwy 244, 45 ° 18.027 ’ N, 118 ° 18.427 ’ W, 15 May 2014, B. Stark, A. B. Harrison, 1 ♀ (BPSC). WASHINGTON: Chelan Co., Bridge Creek, Hwy 20, Rainy Pass, 17 June 2004, B. Stark, R. W. Baumann, 1 ♀ (BPSC). King Co., Green River, about 10 mi abv Lester, 27 May 1969, R. W. Baumann, M. L. Miner, 1 ♀ (BYUC). Lewis Co., Horse Creek, FR 52, Gifford Pinchot National Forest, 15 May 2001, B. C. Kondratieff, 1 ♂, 1 ♀ (CSUC). Lewis Co., Stevens Creek, Hwy 706 E of Louise Lake, 15 June 1969, R. W. Baumann, 1 ♀ (BYUC). Lewis Co., Sunbeam Creek, Hwy 706, 47 ° 19 ’ N, 122 ° 06 ’ W, Mt. Rainier National Park, 21 June 2003, B. C. Kondratieff, R. E. Zuellig, J. Schmidt, 1 ♂ (CSUC). Okanogan Co., Foggy Dew Creek, 8 mi SW Carlton, 18 July 1995, G. MacKenzie, 1 ♀ (BYUC). Pierce Co., Fryingpan Creek, Mt. Rainier National Park, 24 May 2003, B. C. Kondratieff, J. Schmidt, 1 ♂ (CSUC). Pierce Co., White River, Sunrise Rd, Mt. Rainier National Park, 15 June 2001, R. Lechleitner, 1 ♀ (CSUC). Pierce Co., White River, Hwy 410, 14 May 2001, B. C. Kondratieff, R. Lechleitner, 1 ♀ (CSUC). Pierce Co., unnamed stream N of Crystal Creek, Hwy 410, Mt. Rainier National Park, 29 May 1997, B. C. Kondratieff, R. Lechleitner, 2 ♀ (CSUC). Yakima Co., American River, Pleasant Valley Campground, 10 June 2001, D. Strenge, 2 ♀ (CSUC). Egg. Outline oval. Length ca. 390 - 410 μm, equatorial width ca. 263 - 267 μm (Fig. 1). Collar short, wide with sides elaborately dissected; base surrounded by a low, irregularly-multitoothed rim. Anchor mushroom shaped and abundantly covered with globular bodies (Figs. 1, 3). Surface covered with widely-spaced, irregular striae along longitudinal axis; striae composed of linear tubercles ca. 20 - 25 μm long and ca. 5 - 10 μm thick (Figs. 2 - 6). Surface between striae relatively smooth, but bearing a few, scattered globular bodies. Micropyles subequatorial in position, located on modified strial elements (Fig. 2); orifices smaller than width of strial tubercles.	en	Stark, Bill P., Kondratieff, Boris C., Verdone, Chris J. (2015): Kathroperla Siskiyou, A New Stonefly Species From California And Oregon, U. S. A. (Plecoptera: Chloroperlidae). Illiesia 11 (8): 92-103, DOI: 10.5281/zenodo.4760976
039787E2FFCAFFC8ED9127132D3BF91C.taxon	discussion	Comments. The images (Fig. 13 in Stark & Surdick 1987; Fig. 1.9 in Stark et al. 1998 a) prepared from eggs taken from a Muir Creek, Oregon female collected in May 1982 are consistent with the images presented herein from Montana and Oregon, and also consistent with eggs from California and Washington specimens examined from the sites listed above. One female from Humboldt Co., California had eggs typical of this species, although eggs from other specimens collected in that area were typical of K. takhoma. No eggs were present in specimens examined from Alaska, but one Canadian female from Summit Creek, British Columbia contained eggs which were typical of K. perdita. Scanning electron micrographs (Figs. 7 - 8) are provided to show the relatively straight anterodorsal margin of the K. perdita male epiproct. In this species and K. siskiyou, described below, the smooth, sclerotized anterodorsal margin is completely black, whereas this structure is usually bicolored with a brown band appearing adjacent to the marginal black pigmented area in K. takhoma. The anterodorsal margin of the epiproct of K. perdita is also straighter than either of the other species.	en	Stark, Bill P., Kondratieff, Boris C., Verdone, Chris J. (2015): Kathroperla Siskiyou, A New Stonefly Species From California And Oregon, U. S. A. (Plecoptera: Chloroperlidae). Illiesia 11 (8): 92-103, DOI: 10.5281/zenodo.4760976
039787E2FFCAFFC8ED9127132D3BF91C.taxon	distribution	Distribution. The species is reported from Canada: Alberta and British Columbia, and the United States: Alaska, California, Idaho, Montana, Nevada, Oregon and Washington (Baumann et al. 1977; Kondratieff & Lechleitner 2002; Stark & Surdick 1987; Stewart & Oswood 2006; Stewart & Ricker 1997; Stewart & Stark 2002).	en	Stark, Bill P., Kondratieff, Boris C., Verdone, Chris J. (2015): Kathroperla Siskiyou, A New Stonefly Species From California And Oregon, U. S. A. (Plecoptera: Chloroperlidae). Illiesia 11 (8): 92-103, DOI: 10.5281/zenodo.4760976
039787E2FFCFFFCBEE8C20E82A3BFBA9.taxon	description	(Figs. 9 - 19)	en	Stark, Bill P., Kondratieff, Boris C., Verdone, Chris J. (2015): Kathroperla Siskiyou, A New Stonefly Species From California And Oregon, U. S. A. (Plecoptera: Chloroperlidae). Illiesia 11 (8): 92-103, DOI: 10.5281/zenodo.4760976
039787E2FFCFFFCBEE8C20E82A3BFBA9.taxon	materials_examined	Material examined. Holotype ♀, USA: OREGON, Jackson Co., Split Rock Creek, Wagner Gap Rd, 12 mi S Talent, 42.09480 ° N, 122.77397 ° W, 22 May 2014, B. C. Kondratieff, C. J. Verdone, J. Sandberg, B. Stark (USNM). Paratypes, CALIFORNIA: Sierra Co., North Yuba River, Hwy 49, near Sierra City, 10 May 1983, R. W. Baumann, R. C. Mower, 1 ♀ (BYUC). OREGON: Same data as holotype, 2 ♀ (CSUC). Jackson Co., East Fork Ashland Creek, south of Ashland, 19 May 2000, R. W. Wisseman, 1 ♀ (BYUC). Jackson Co., McDonald Creek, Wagner Gap Rd, 22 May 2014, B. C. Kondratieff, C. J. Verdone, 1 ♂ (CSUC).	en	Stark, Bill P., Kondratieff, Boris C., Verdone, Chris J. (2015): Kathroperla Siskiyou, A New Stonefly Species From California And Oregon, U. S. A. (Plecoptera: Chloroperlidae). Illiesia 11 (8): 92-103, DOI: 10.5281/zenodo.4760976
039787E2FFCFFFCBEE8C20E82A3BFBA9.taxon	description	Adult habitus. General color brown patterned with dark brown. Lateral margins of head bearing dark brown stripes extending from posterior margin to eye (Fig. 15); occiput pale brown in fully pigmented specimens, but infuscated in some specimens with slightly darker brown pigment. Dark ocellar spot extends forward on frons to level of antennae; antennae dark brown. Pronotum with pale brown median stripe, broad sublateral dark stripes, and an almost complete marginal ring of dark pigment interrupted on lateral margins near midlength (Fig. 15); sublateral stripes sometimes almost broken, or notched on lateral margins. Wings transparent, veins dark brown except pale brown C and Sc. Femora brown except apical black 18 19 ring; tibiae brown but slightly darker distally, proximally at knee and along inner margin. Abdomen pale brown on dorsum and bearing a darker median stripe and a pair of dark lateral stripes on terga 1 - 8; basal cercal segment pale brown but bearing minute dark spots around setal bases, subsequent segments black. Thoracic sterna dark brown; abdominal sterna brown with darker markings forming a broken, median longitudinal stripe from sterna 3 - 7. Male. Forewing length 16 mm. Epiproct typical of genus but dorsolateral extensions diverge posterolaterally from central axis (Fig. 16); anterodorsal margin of epiproct uniformly black. Basal cercal segments ca. 2 - 2.5 X long as wide. Vesicle body almost circular but with moderately wide base forming a short, stalk-like structure (Fig. 17). Aedeagus broad and membranous except for a pair of curved, dark brown lateral bands and a ventromedian pale sclerite (Fig. 18); surface of lateral bands bear fine, short, densely packed setae; distal margins foot-shaped. Aedeagal apex consists of three membranous lobes, median lobe finely divided by a suture-like line along most of its length. Female. Forewing length 17 - 18 mm. Lateral margins of subgenital plate slightly concave beyond midlength, apex notched; apex extends over base of sternum 9 (Fig. 19). Egg. Outline oval. Length ca. 380 - 400 μm, equatorial width ca. 230 - 240 μm (Figs. 9 - 10). Collar short, ca. 22 - 30 μm, wide ca. 70 - 80 μm (Fig. 11), elaborately dissected, and surrounded by three irregular rows of meshes and a conspicuous outer ridge forming a ring ca. 104 μm in diameter (Figs. 11 - 12). Anchor well developed, cushion shaped and covered with small globular bodies (Fig. 10). Chorion bearing multiple rows of longitudinal striae composed of closely grouped, irregularly shaped tubercles ca. 2 - 3 μm wide; interstrial areas bear scattered tubercles similar to those which comprise striae (Figs. 11, 13). Micropyles subequatorial in position and often associated with striae (Fig. 14); orifices slightly larger in diameter than bead-like components of striae. Larva. Unknown.	en	Stark, Bill P., Kondratieff, Boris C., Verdone, Chris J. (2015): Kathroperla Siskiyou, A New Stonefly Species From California And Oregon, U. S. A. (Plecoptera: Chloroperlidae). Illiesia 11 (8): 92-103, DOI: 10.5281/zenodo.4760976
039787E2FFCFFFCBEE8C20E82A3BFBA9.taxon	etymology	Etymology. The species name, used as a noun in apposition, is based on the type locality in the Siskiyou Mountains of southern Oregon. The proposed common name, Wagner Gap Sallfly, is based on the access road into the sites where most of the known specimens were collected. Common names for K. perdita (Longhead Sallfly) and K. takhoma (Slenderhead Sallfly) were previously proposed by Stark et al. (1998 b; 2012).	en	Stark, Bill P., Kondratieff, Boris C., Verdone, Chris J. (2015): Kathroperla Siskiyou, A New Stonefly Species From California And Oregon, U. S. A. (Plecoptera: Chloroperlidae). Illiesia 11 (8): 92-103, DOI: 10.5281/zenodo.4760976
039787E2FFCFFFCBEE8C20E82A3BFBA9.taxon	diagnosis	Diagnosis. Although some overlap in subgenital plate shape occurs, females of K. siskiyou are usually distinguished from K. takhoma by a shorter subgenital plate, and by the egg structure which bears a collar and longitudinal striations formed from bead-like tubercles (Figs. 11, 13). The new species and K. perdita completely overlap in subgenital plate structure and in head pattern, but the eggs differ in the form of the chorionic tubercles. In K. perdita these structures are thick and long (Figs. 5 - 6), whereas in the new species they are formed as longitudinal clusters of beadlike structures (Figs. 11, 13). The only known male specimen of this species appears intermediate between K. perdita and K. takhoma. The basal cercal segments and, general head coloration and pigment pattern of the dorsal margin of the epiproct are similar to those of K. perdita, whereas the vesicle shape and dorsal epiproct margin are more similar to K. takhoma. Males of the three Nearctic species have similar aedeagal structure but when a larger sample is available the shape and size of the lateral sclerites may offer another useful character. We offer a preliminary key below to assist in distinguishing these three species.	en	Stark, Bill P., Kondratieff, Boris C., Verdone, Chris J. (2015): Kathroperla Siskiyou, A New Stonefly Species From California And Oregon, U. S. A. (Plecoptera: Chloroperlidae). Illiesia 11 (8): 92-103, DOI: 10.5281/zenodo.4760976
039787E2FFCCFFC5EDCE22042CDBFC3E.taxon	materials_examined	Holotype ♀ (United States National Museum), Falls Creek above Ohanapecosh River, Mt. Rainier National Park, Hwy 706, Washington	en	Stark, Bill P., Kondratieff, Boris C., Verdone, Chris J. (2015): Kathroperla Siskiyou, A New Stonefly Species From California And Oregon, U. S. A. (Plecoptera: Chloroperlidae). Illiesia 11 (8): 92-103, DOI: 10.5281/zenodo.4760976
039787E2FFCCFFC5EDCE22042CDBFC3E.taxon	materials_examined	Material examined. USA: CALIFORNIA: Del Norte Co., 3 mi NE Hiouchi, 1 June 1991, B. Stark, R. W. Baumann, C. Henderson, 1 ♀, 1 larva (BPSC). Humboldt Co., Cedar Creek, Hwy 299, 25 April 1987, B. Stark, R. W. Baumann, C. R. Nelson, 2 ♂, 5 ♀ (BPSC). Humboldt Co., 1 mi E Cedar Creek, Hwy 299, 25 April 1987, B. Stark, R. W. Baumann, C. R. Nelson, 1 ♂ (BPSC). Humboldt Co., East Fork Willow Creek, East Fork Willow Creek Campground, 24 April 1987, B. Stark, R. W. Baumann, C. R. Nelson, 1 ♂, 4 ♀ (BPSC). Humboldt Co., Slide Creek, Hwy 13 south of Fish Lake Campground, 31 May 1991, B. Stark, R. W. Baumann, C. Henderson, 1 ♀ (BPSC), 2 ♀ (BYUC). Mendocino Co., Irish Gulch, 39 ° 01.209 ’ N, 123 ° 41.268 ’ W, 22 April 2005, L. D. Bottorff, 1 ♀ (RLBC). Mendocino Co., Same location, 2 April 2005, L. D. Bottorff, 1 ♂ (RLBC). Trinity Co., Clear Creek and tributaries, Hwy 36, 2.6 mi E of Forest Glen Campground, 40.3743 ° N, 123.3634 ° W, 9 May 2009, J. B. Sandberg, 2 ♂, 2 ♀, 1 larva, 2 exuvium (JBSC). OREGON: Benton Co., Yew Creek, Mary’s Peak Rd, 1 June 2000, B. Stark, I. Sivec, M. Zúñiga, 1 ♀ (BPSC). Benton Co., Lewisburg Saddle, Malaise trap, 44.6423 ° N, 121.4204 ° W, 31 May- 12 June 2013, S. Fitzgerald, 5 ♂, 8 ♀ (CSUC). Same location, Malaise trap, 6 - 31 May 2013, S. Fitzgerald, 2 ♀ (CSUC). Lane Co., Watershed # 10, Anderson Experimental Forest, 20 April 1972, R. W. Wisseman, 1 ♂, 1 ♀ (BYUC). Linn Co., tributary North Santiam River, Hwy 22, 0.2 mi below Minto Rd, 3 June 2000, B. Stark, I. Sivec, M. Zúñiga, 1 ♀ (BPSC). Multnomah Co., Bridal Veil Creek below Bridal Veil Falls, 13 May 2003, B. C. Kondratieff, R. W. Baumann, 1 ♀ (CSUC). Multnomah Co., Columbia River junction with Tanner Creek, 28 March 1987, G. R. Fiala, 1 ♂ (BYUC). WASHINGTON: Clallam Co., Madison Creek, Madison Creek Falls, Elwah River Valley, Olympic National Park, 48 ° 02 ’ N, 123 ° 35 ’ W, 15 June 2005, R. W. Baumann, B. C. Kondratieff, 1 ♀ (BYUC). Jefferson Co., Mineral Creek, tributary Hoh River, N 102 - 103, E 33 - 34, 24 June 1999, D. E. Bowles, 1 ♀ (BYUC). Lewis Co., Faith Creek, Hwy 706, Mt. Rainier National Park, 17 June 2004, B. C. Kondratieff, J. Schmidt, 2 ♀ (CSUC). Lewis Co., St. John Creek, Rt. 22, 13 mi E Rt. 23, 1 July 1985, C. M. Flint, O. S. Flint, Jr., 2 ♂ (USNM). Lewis Co., tributary Ohanapecosh River, Mt. Rainier National Park, 7 July 1999, B. C. Kondratieff, 1 ♀ (CSUC). Mason Co., Brown Creek, FR 2233, 12 mi NW Shelton, 22 April 1995, T. Nash, 1 ♂ (BYUC). Skamania Co., Rock Creek, Millard, tributary Little Salmon River, 15 May 2003, B. C. Kondratieff, R. W. Baumann, 1 ♂ (CSUC). Skamania Co., tributary Little White Salmon River, near Oklahoma Campground, 27 May 1984, G. R. Fiala, 1 ♀ (BYUC). Whatcom Co., Breckenridge Creek, 5 mi E Eversen, 11 May 1967, unknown collector, 1 ♀ (BYUC). Whatcom Co., Canyon Creek, Deming, 21 April 1967, unknown collector, 1 ♂ (BYUC). Whatcom Co., Sygitowicz Creek, 2 mi SW Van Zandt, 5 May 1994, Kraft and Sammeth, 1 ♂ (BYUC). Egg. Outline oval, collar absent (Figs. 20 - 21, 23). Length ca. 431 μm, equatorial width ca. 253 μm. Collar end of egg bearing a dense cluster of small tubercles (Fig. 23). Chorionic surface sparsely covered throughout with small, projecting tubercles of irregular height (Figs. 23 - 25); some tubercles are grouped in poorly organized, short rows but most appear randomly placed. Micropyles subequatorial in position and frequently located in association with tubercles (Fig. 25).	en	Stark, Bill P., Kondratieff, Boris C., Verdone, Chris J. (2015): Kathroperla Siskiyou, A New Stonefly Species From California And Oregon, U. S. A. (Plecoptera: Chloroperlidae). Illiesia 11 (8): 92-103, DOI: 10.5281/zenodo.4760976
039787E2FFCCFFC5EDCE22042CDBFC3E.taxon	distribution	Distribution. This species has previously been reported from California and Washington (Bottorff & Bottorff 2007; Kondratieff & Lechleitner 2002; Nelson & Stark 1987; Stark & Surdick 1987). The Oregon records listed above, though not unexpected, are the first reports from that state.	en	Stark, Bill P., Kondratieff, Boris C., Verdone, Chris J. (2015): Kathroperla Siskiyou, A New Stonefly Species From California And Oregon, U. S. A. (Plecoptera: Chloroperlidae). Illiesia 11 (8): 92-103, DOI: 10.5281/zenodo.4760976
