identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03975B72FFC2760896FDC8D95568FA4A.text	03975B72FFC2760896FDC8D95568FA4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister bengalensis Aube 1838	<div><p>Cybister bengalensis Aubé, 1838</p> <p>(Figs 1A–E)</p> <p>Cybister bengalensis Aubé, 1838: 61 (orig. descr.).</p> <p>Material examined. CHINA: JIANGXI: 1 ♀, Pingshiang [Pingxiang], coll.A.Zimmermann (ZSMG). TAIWAN: 2JJ, Formosa,coll.A.Zimmermann (ZSMG); 1 J, Ku Sia, coll. A. Zimmermann (ZSMG). WITHOUT ඉRECISE DATA: 1 ♀, China, coll. A. Zimmermann (ZSMG).</p></div> 	https://treatment.plazi.org/id/03975B72FFC2760896FDC8D95568FA4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFC2760896F9CEEB5536FB7B.text	03975B72FFC2760896F9CEEB5536FB7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister Curtis 1827	<div><p>Subgenus Cybister Curtis, 1827</p> <p>Type species. Dytiscus lateralis Fabricius, 1798.</p> <p>Diagnosis. With distinctive yellow margins laterally on the pronotum and elytron. Males with a single claw per metatarsus; females usually with single claws, except in some North and Central American species.</p> <p>Included (Chinese) species. Cybister bengalensis Aubé, 1838; C. chinensis Motschulsky, 1854; C. fumatus Sharp, 1882; C. guerini Aubé, 1838; C. laevis Falkenström, 1936; C. lateralimarginalis torquatus (Fischer von Waldheim, 1829); C. lewisianus Sharp, 1873; C. limbatus (Fabricius, 1775); C. rugosus (W. S. Macleay, 1825); C. tripunctatus lateralis (Fabricius, 1798); C. ventralis Sharp, 1882.</p> </div>	https://treatment.plazi.org/id/03975B72FFC2760896F9CEEB5536FB7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFC1760D95B2CD6D55C7FF2F.text	03975B72FFC1760D95B2CD6D55C7FF2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister chinensis Motschulsky 1854	<div><p>Cybister chinensis Motschulsky, 1854</p> <p>(Figs 2A–E; 18A–C; 19A; 20A)</p> <p>Cybister chinensis Motschulsky, 1854: 44 (orig. descr.).</p> <p>Cybister japonicus Sharp, 1873: 45 (orig. descr.), synonymy by ZAITZEV (1915): 284 and confirmed by NILSSON &amp; PETROV (2007): 43; SHARP (1882):748 (Manchuria, Shandong, Taiwan);RḖGIMBART (1899): 348</p> <p>(Manchuria, Taiwan); FENG (1932): 36 (Beijing, Fujian: Fuzhou and Xiamen, Guangdong, Hainan, Manchuria, Shandong:Yantai, Taiwan); FALKENSTRÖM (1936): 244 (Sichuan); FENG (1936): 13 (Liaoning: Shenyang, Shaanxi); GSCHWENDTNER (1938): 76 (Manchuria, Taiwan); BRINCK (1946): 151 (Heilongjiang, Liaoning); ZAITZEV (1972): 360 (Taiwan); ZHAO (1981): 110 (Fujian); ZENG (1989): 90 (NE China, Fujian, Guangdong, Hainan, Hebei, Ningxia: Yinchuan, Shandong, Taiwan); LI (1992): 36 (Jilin, Liaoning); MORI &amp; KITAYAMA (1993): 145 (Taiwan).</p> <p>Material examined. CHINA: GUANGDONG: 3 JJ, Canton, 1.–21.</p> <p>iii.2001, L. Hojný leg. (NMPC). NINGXIA: 1 ♀, Yinchuan, under light,</p> <p>6.vi.1960 (SYSU); 1 J, Yinchuan, under light, 15.vii.1962 (SYSU); 1</p> <p>J, Yinchuan, Gujiaqiao, 23.vi.2011 (SYSU); 2 ♀♀, Yinchuan (SYSU). TIANJIN: 1 ♀, Tsing tou, 17.viii.1933 (SYSU); 1 J, Tientsin, 10.iv.1935 (SYSU); 1♀, Tianjin, Balitai, 4.v.1955,Y.Y.Liu leg. (SYSU). WITHOUT ඉRECISE DATA: 15 spec., donated by Xing-min Wang, purchased from market, without data (SYSU); 1 J, China (LHCM).</p> <p>Diagnosis. TL 35–40 mm, elongate, oblong-oval (Fig. 2A). Head dark green, clypeus yellow. Pronotum and elytra dark green to black, with distinct yellow margins laterally. Ventral surface yellow; prosternal process, middle of metaventrite and metacoxae dark green; metacoxae and abdominal ventrites narrowly margined (Fig. 2B). Dorsal surface dimorphic: smooth with sparse fine punctures in male; in female with isolated short, irregular striae on pronotum, and with dense longitudinal striae of various lengths on elytra except at its apex. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite (ʻmetasternal wingsʼ) tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs yellow, midlegs and hindlegs yellow to brown; protarsomeres 1–3 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; metatarsal claws single in both sexes. Median lobe of aedeagus ʻCʼ- shaped in lateral view, moderately curved (Fig. 2C); ridged in middle in ventral view, lateral margin strongly curved near apex, apex obtuse-angled (Fig. 2D). Parameres moderately broad, strap-like apically (Fig. 2E).</p> <p>Habitat. This species inhabits ponds, paddy fields and wetlands with aquatic plants (MORI &amp; KITAYAMA 1993); specimens from Shanxi were found in a natural lake (Figs 18A–C).</p> <p>Distribution. China (Beijing, Fujian, Guangdong, Hainan, Hebei, Heilongjiang, Jilin, Liaoning, Ningxia, Sichuan, Shaanxi, Shandong, Shanxi, Taiwan, Tianjin) (Fig. 20A), Japan, North Korea, Russia, South Korea (RḖGIMBART 1899; ZIMMERMANN 1920; FENG 1932, 1933b;</p> <p>GSCHWENDTNER 1938; ZAITZEV 1972; ZENG 1989; MORI &amp; KITAYAMA 1993; NILSSON 1995; NILSSON &amp; HáJEK 2022b).</p></div> 	https://treatment.plazi.org/id/03975B72FFC1760D95B2CD6D55C7FF2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFC7760C9682CD0D5126FA2C.text	03975B72FFC7760C9682CD0D5126FA2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister fumatus Sharp 1882	<div><p>Cybister fumatus Sharp, 1882</p> <p>(Figs 3A–E; 20A)</p> <p>Cybister fumatus Sharp, 1882: 731 (orig. descr.); FENG (1936): 13 (Shanghai).</p> <p>Material examined. None from China but 9 specimens from LAOS: 4 spec., Kham Mouan prov., Ban Khoun Ngeun, 18°07′N 104°29′E, 200 m, 19.–31.v.2001, P. Pachol á tko leg. (NHMB, LHCM); 1 spec., Vientiane prov., Phou Khao Khouay, 18°20.369′N 102°48.523′E, nr. strongly disturbed primary rainforest on light, 25.–30.v.2008, A. Solodovnikov &amp; J. Pedersen leg. (ZMUC); 1 spec., Sekong Prov., Tad Faek waterfalls, 15°14.7′N 106°42.1′E, 118 m, light trap, 8.v.2010, J. Hájek leg.(NMPC); 1 spec., Bolikhamsay prov., Nam Kading NPA research center near Ban Phone Kham, 18°20′N 104°08′E, 250 m, 23.–29.v.2011, NHMB Basel Laos 2011 Expedition M. Brancucci, M. Geiser, D. Hauck, Z. Kraus, A. Phantala &amp; E. Vongphachan (NHMB); 1 spec., Bolikhamsay prov., Pakkading Ban Phone Kham env. 18°19′N 104°08′E 200–300 m, 23.–29.v.2011, NHMB Basel Laos 2011 Expedition M. Brancucci, M. Geiser, D. Hauck, Z. Kraus, A. Phantala &amp; E. Vongphachan (NHMB); 1 spec., Savannakhet prov., 10 E of Savannakhet, Dong Natad forest, 180 m, 4.xii.2011, J. Šťastný leg. (JSCL).</p> <p>Diagnosis. TL 22–24 mm, elongate, oblong-oval (Fig. 3A). Head dark green, clypeus yellow. Pronotum and elytra dark green, with distinct yellow margins laterally; apex of yellow margin on elytra slightly hooked. Ventral surface black; with lateral part of metaventrite and metacoxae yellow; abdominal ventrites III–V with a pale spot on each side (Fig. 3B). Dorsal surface smooth in both sexes. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs reddish-brown, midlegs and hindlegs reddish-brown to black; protarsomeres 1–3 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; metatarsal claws single in both sexes. Median lobe of aedeagus ʻCʼ- shaped in lateral view, moderately curved (Fig. 3C); gradually narrowed from base to near apex in ventral view, apex furcate, with a deep acute-angled notch medially, lateral margins of apex straight and parallel (Fig. 3D). Parameres moderately broad, approximately the same width apically (Fig. 3E).</p> <p>Habitat. In Laos all specimens were collected at lower altitudes between 180 and 300 m above sea level. The specimen from Sekong was attracted by a light trap placed near the border of a lowland river. The single specimen from Dong Natad was collected in a shallow swamp in lowland monsoon primary forest, associated with Hyphydrus lyratus (Swartz, 1808), Hyphydrus sp. and Lacconectus sp. For habitat photos see HENDRICH &amp; BRANCUCCI (2013).</p> <p>Distribution. China (Shanghai) (Fig. 20A), Indonesia, Laos, Malaysia, Thailand, Vietnam (RḖGIMBART 1899, ZIMMERMANN 1920, FENG 1936, NILSSON 1995, HENDRICH &amp; BRANCUCCI 2013).</p></div> 	https://treatment.plazi.org/id/03975B72FFC7760C9682CD0D5126FA2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFC6760C945FCA0E5506FACF.text	03975B72FFC6760C945FCA0E5506FACF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister guerini Aube 1838	<div><p>Cybister cf. guerini Aubé, 1838</p> <p>(Figs 4A–E; 20A)</p> <p>Cybister guerini Aubé, 1838:57 (orig.descr.); SHARP (1882): 741 (China,</p> <p>Manchuria); RḖGIMBART (1899): 342 (Guangdong); FENG (1932): 35</p> <p>(Guangdong); ZENG (1989): 91 (Guangdong).</p> <p>Material examined. CHINA: GUANGDONG: 76 spec., Guangzhou, 14.xi.1964, T.X.Peng got from market (SYSU); 1J 1♀, Nanhai District, Xiqiaoshan Mt., 11.x.1973, S. T. Lu leg. (SYSU); 1 J 2♀♀, Guangzhou (SYSU). HAINAN: 1 J, Lingshui County, Diaoluoshan Mt., Xiaomei Village, 29.xii.1963, L. F. Huang leg. (SYSU).</p> <p>Diagnosis. TL 31–36 mm, elongate, oblong-oval (Fig. 4A). Head dark green, clypeus yellow. Pronotum and elytra dark green to black, with distinct yellow margins laterally; apex of yellow margin on elytra hooked. Ventral surface black; abdominal ventrites III–V with a yellow spot on each side (Fig. 4B). Dorsal surface dimorphic: smooth sometimes with inconspicuous small granules in male, in female with isolated short, irregular striae on pronotum, and with dense longitudinal striae of various lengths on elytra except at its apex. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs reddish-brown, midlegs and hindlegs reddish-brown to black; protarsomeres 1–3 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; metatarsal claws single in both sexes. Median lobe of aedeagus ʻCʼ- shaped in lateral view, moderately curved (Fig. 4C); slightly narrow near apex in ventral view, apex furcate, with a notch medially, lateral margins of apex sinuate (Fig. 4D). Parameres moderately broad, strap-like apically (Fig. 4E).</p> <p>Comments to classification. As already mentioned in HENDRICH &amp; BRANCUCCI (2013), it will be necessary to study the type material of C. guerini and C. limbatus carefully before identifying freshly collected material. In many collections, historical specimens of the same species can be found under the two different names.</p> <p>Habitat. The knowledge of habitat of Cybister guerini is very poor, all specimens examined here were collected in 1960s and 70s without habitat information. Some specimens from Guangzhou were obtained from a market.</p> <p>Distribution. China (Guangdong, Hainan, Hong Kong, “Manchuria”) (Fig. 20A), India, Indonesia, Laos, Thailand, Vietnam (RḖGIMBART 1899; ZIMMERMANN 1920; FENG 1932, 1933b; GSCHWENDTNER 1938; VAZIRANI 1969; ZENG 1989; NILSSON 1995, NILSSON &amp; HÁJEK 2022b). The record of C. guerini from Northeast China by SHARP (1882) is outside the distribution area of this Oriental species and needs to be confirmed. First record from Hainan.</p> </div>	https://treatment.plazi.org/id/03975B72FFC6760C945FCA0E5506FACF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFC6760C96EBC92D557BF82F.text	03975B72FFC6760C96EBC92D557BF82F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister laevis Falkenstrom 1936	<div><p>Cybister laevis Falkenström, 1936</p> <p>(Figs 5A–C; 20A)</p> <p>Cybister laevis Falkenström, 1936: 243 (orig. descr.).</p> <p>Cybister tripunctatus var. laevis: GSCHWENDTNER (1938: 64).</p> <p>Type material. HOLOTYPE: ♀ (NHRS), labelled: “Giufu-Shan [Chongqing, Jinfoshan Mt.] / Szechuan / Em. Reitter [p] // Cybister ♀ / laevis / Falk. [hw] / det. Falkenström [p] // Typus [red label with black frame, p] // 7533 / E91 + [blue label, p] // NHRS-JLKB / 000073974 [p]”.</p> <p>Diagnosis. See Cybister tripunctatus lateralis. Ventral side yellowish.</p> <p>Comments to classification. The female type specimen has smooth elytra without any striolae – a condition which can be observed also in all females of C. tripuncatus lateralis. Since it is an odd, old museum specimen, it has a yellowish ventral side, which is not the case for C. tripunctatus lateralis from China, where the ventral side is always black or ferruginous in freshly hatched specimens. The size (26 mm) and the coloration of the dorsal side also indicate that C. laevis is conspecific with the variable and widespread C. tripunctatus lateralis, as already suggested by GSCHWENDTNER (1938).</p> <p>Distribution. China (Chongqing) (Fig. 20A).</p></div> 	https://treatment.plazi.org/id/03975B72FFC6760C96EBC92D557BF82F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFC976039453CCAC51ACFA20.text	03975B72FFC976039453CCAC51ACFA20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister lewisianus Sharp 1873	<div><p>Cybister lewisianus Sharp, 1873</p> <p>(Figs 7A–E; 21A)</p> <p>Cybister lewisianus Sharp, 1873: 46 (orig. descr.); SHARP (1882): 732 (Shanghai); RḖGIMBART (1899): 350 (Shanghai, Jiangxi); FENG (1932): 36 (Anhui:Wuhu, Beijing, Hubei:Wuchang, Jiangsu:Suzhou, Zhejiang: Hangzhou and Jiaxing); ZHAO (1981): 110 (Fujian); ZENG (1989): 92 (Anhui, Beijing, Hubei, Jiangsu, Zhejiang); LI (1992): 36 (Liaoning).</p> <p>Material examined. CHINA: GUANGDONG: 2 JJ 1 ♀, Nanhai District, Jiujiang Township, 16.xi.1957, D. X. Gu leg. (SYSU); 6JJ 2♀♀,</p> <p>Guangzhou, 14.xi.1964, T. X. Peng got from market (SYSU). HAINAN:</p> <p>1 ♀, Mt. Wuzhishan at Wuzhishan city, 15–24.vii.2011, Jingke Li leg.</p> <p>(SYSU). HUBEI: 2JJ 2♀♀, Wuchang, vii.1956, Q. Y. Hu leg.(SYSU).</p> <p>Diagnosis. TL 23–27 mm, elongate, oblong-oval (Fig. 7A). Head dark green, clypeus reddish-brown. Pronotum and elytra dark green to black, with distinct yellow margins laterally. Ventral surface black, with lateral part of metaventrite and metacoxae reddish-brown; abdominal ventrites III–V with a yellow spot on each side (Fig. 7B). Dorsal surface smooth in both sexes. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs yellow, midlegs and hindlegs yellow to reddish-brown; protarsomeres 1–3 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; both sexes with a single metatarsal claw. Median lobe of aedeagus ʻCʼ- shaped in lateral view, moderately curved (Fig. 7C); slender in ventral view, apex rounded (Fig. 7D). Parameres moderately broad, strap-like apically (Fig. 7E).</p> <p>Habitat. This species inhabits large ponds with aquatic plants (MORI &amp; KITAYAMA 1993).</p> <p>Distribution. China (Anhui, Beijing, Guangdong, Hainan, Hebei, Hubei, Jiangsu, Jiangxi, Liaoning, Shanghai, Zhejiang) (Fig. 21A), India, Indonesia, Japan, Vietnam (RḖGIMBART 1899; ZIMMERMANN 1920; FENG 1932, 1933b; GSCHWENDTNER 1938; ZAITZEV 1972; ZENG 1989; MORI &amp; KITAYAMA 1993; NILSSON 1995; GHOSH &amp; NILSSON 2012; NILSSON &amp; HÁJEK 2022b). First record from Guangdong and Hainan.</p></div> 	https://treatment.plazi.org/id/03975B72FFC976039453CCAC51ACFA20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFC9760395ABCA325496F8EF.text	03975B72FFC9760395ABCA325496F8EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister limbatus (Fabricius 1775) : SHARP 1882	<div><p>Cybister cf. limbatus (Fabricius, 1775)</p> <p>(Figs 8A–E; 18F; 21A)</p> <p>Dytiscus limbatus Fabricius, 1775: 230 (orig. descr.).</p> <p>Dytiscus aciculatus Herbst, 1784: 123 (orig. descr.), synonymy by SCHÖNHERR (1808: 10).</p> <p>Cybister limbatus: SHARP (1882: 739) (Manchuria); ZIMMERMANN (1919: 77) (Taiwan); KANO (1931: 176) (Taiwan); MORI &amp; KITAYAMA (1993: 144) (Taiwan).</p> <p>Material examined. CHINA: GUANGDONG: 1 ♀, Canton, Honam</p> <p>Island, P’an-yu District. 25.vii.1934, William E. Hoffmann leg. (SYSU);</p> <p>1 J, Lung-tau Shan,Yiu vill., above Taokokwan, 600 m, 7.vi.1947, PM,</p> <p>Gressitt &amp; Lam (SYSU); 13 JJ 3 ♀♀, Guangzhou, 14.xi.1964, T. X.</p> <p>Peng got from market (SYSU). GUANGXI: 1 J, Nanning, 19.vi.1977,</p> <p>Z. H. Huang leg. (SYSU). HAINAN: 1 J, Sanya, 28.xii.1963, T. X. Peng leg. (SYSU); 1 J, Jianfengling Mt., 1.xi.1983, P. L. Chen leg. (SYSU);</p> <p>1 ♀, Limushan Mt., 28.v.1984, G. R. Lin leg. (SYSU). YUNNAN: 1 ♀,</p> <p>Longling County, 1060 m, 24.vi.1956, B. S. Zhou leg. (SYSU).</p> <p>Diagnosis. TL 33–39 mm, elongate, oblong-oval (Fig. 8A). Head dark green, clypeus yellow. Pronotum and elytra dark green to black, with distinct yellow margins laterally, apex of yellow margin on elytra hooked. Ventral surface black; abdominal ventrites III–V with a reddish-brown spot on each side (Fig. 8B). Dorsal surface dimorphic: smooth sometimes with inconspicuous small granules in male, in female with isolated short, irregular striae on pronotum and dense longitudinal striae of various length on elytra except at its apex. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs reddish-brown, midlegs and hindlegs reddish-brown to black; protarsomeres 1–3 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; both sexes with a single metatarsal claw. Median lobe of aedeagus ʻCʼ- shaped in lateral view, moderately curved (Fig. 8C); gradually narrowed from base to near apex in ventral view, apex furcate, with a deep obtuse-angled notch medially, lateral margins of apex concave (Fig. 8D). Parameres moderately broad, strap-like apically (Fig. 8E).</p> <p>Comments to classification. As mentioned under C. guerini, it will be necessary to study the type material of C. limbatus and C. guerini to clarify the status of both taxa. Specimens of the same morphospecies can be found under the two different names in many historical collections (HENDRICH &amp; BRANCUCCI 2013).</p> <p>Habitat. This species inhabits deep ponds and abandoned paddy fields with aquatic plants (MORI &amp; KITAYAMA 1993, HENDRICH &amp; BRANCUCCI 2013); specimens from Danxiashan Mt. (Guangdong) were found in a temporary pond in a valley (Fig. 18F). In Thailand, the adults prey on larvae and pupae of Aedes aegypti (Linnaeus, 1762) and Culex quinquefasciatus Say, 1823 (YUKALAND 2005). The species is sold as food in Thailand (CHEN et al. 1998) and Laos (HENDRICH &amp; BRANCUCCI 2013).</p> <p>Distribution. China (Guangdong, Guangxi, Hainan, Taiwan, Xizang, Yunnan, “Manchuria”) (Fig. 21A), Afghanistan, India, Japan, Laos, Myanmar, Nepal, Pakistan, Philippines, Thailand, Vietnam (RḖGIMBART 1899; ZIMMERMANN 1920; FENG 1932, 1933b; GSCHWENDTNER 1938; VAZIRANI 1969; ZAITZEV 1972; MORI &amp; KITAYAMA 1993; NILSSON 1995; GHOSH &amp; NILSSON 2012; HENDRICH &amp; BRANCUCCI 2013; NILSSON &amp; HÁJEK 2022b). The record of C. limbatus from Northeast China by SHARP (1882) is outside the distribution area of this Oriental species and needs to be confirmed. First records from Guangdong, Guangxi and Hainan.</p> </div>	https://treatment.plazi.org/id/03975B72FFC9760395ABCA325496F8EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFC9760796D7CB4D5385FE4F.text	03975B72FFC9760796D7CB4D5385FE4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister rugosus (W. S. Macleay 1825)	<div><p>Cybister rugosus (W. S. Macleay, 1825)</p> <p>(Figs 9A–E; 18D–E, G; 19C; 21A)</p> <p>Dytiscus rugosus W. S. Macleay, 1825: 32 (orig. descr.).</p> <p>Cybister indicus Aubé, 1838: 62 (orig. descr.), synonymy by SHARP (1882: 992).</p> <p>Cybister rugosus: NILSSON (1995): 78 (Taiwan).</p> <p>Material examined. CHINA: CHONGQING: 1 J, Fengdu County, Shiping Village, 610 m, 5.x.1994, J. Chen leg. (SYSU); 2 JJ, Fengdu County, Shiping Village, 580 m, 6.x.1994, J. Yao leg. (SYSU). GUANG- DONG: 1J 2♀♀, suburbs of Guangzhou, 6.xi.1956 (SYSU); 1♀, Nanhai District, Jiujiang Township, 16.xi.1957, D. X. Gu leg. (SYSU); 69 spec., Guangzhou, 14.xi.1964, T. X. Peng got from market (SYSU); 1 J, Sanshui District, 18.xi.1964, T. X. Peng leg. (SYSU); 2 JJ, Nanhai District, Xiqiaoshan Mt., 11.x.1973, S. T. Lu leg. (SYSU); 1 J, Guangzhou, Sun Yat-sen University, x.1973, S. T. Lu leg. (SYSU); 1 J, Dinghushan Mt., 28.vi.1978, Z. Y. Wang leg. (SYSU); 1 J, Guangzhou, Sun Yat-sen University, in front of Huaishitang, 2.x.1999, L. Z. Hua leg. (SYSU); 1 ♀, Shaoguan, Danxiashan Mt., 30.v.–8.vi.2008, F. L. Jia leg. (SYSU); 1 J, Fengkai County, He’erkou, light trap, 2.xii.2010, L. J. Yang &amp; Y. L. Yu leg. (SYSU). HUBEI: 1 ♀, Hwang-mei &amp; vicinity, Hwang-mei Dist., 9.–11.vii.1935, H. W. Djou (SYSU). HUNAN: 1 J, Tongdao County, 20.viii.1982, Z. H. Huang leg. (SYSU); 1 J, Hengyang, Zhoujiaao, 28.vi.1986, Q.Zou leg. (SYSU). JIANGXI: 1♀, main peak of Jinggangshan Mt., 1.ix.2011, L. J. Yang leg. (SYSU); 1 J, Nanchang (SYSU); 1 ♀, Shusi, H. Y. Cai leg. (SYSU). YUNNAN: 1 ♀, Yingjiang County, 820 m, 25.v.1983, L. Z. Hua leg. (SYSU). ZHEJIANG: 2 ♀♀, Kecheng district, Lankeshan Mt., Quzhou, 28.8781N, 118.9199E, 118 m, 18.iii.2020, Z. Y. Jiang leg. (SYSU).</p> <p>Diagnosis. TL 27–33 mm, elongate, oblong-oval (Fig. 9A). Head dark green, clypeus yellow. Pronotum and elytra dark green to black, with distinct yellow margins laterally, apex of yellow margin on the elytra hooked. Ventral surface black with lateral part of metaventrite and metacoxae yellow; abdominal ventrites I–V with yellow spot on each side (Fig. 9B). Dorsal surface dimorphic: smooth sometimes with inconspicuous small granules in male; in female with isolated short, irregular striae on pronotum and dense longitudinal striae of various length on elytra at its apex. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs yellow, midlegs and hindlegs yellow to black; protarsomeres 1–3 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; both sexes with a single metatarsal claw. Median lobe of aedeagus ʻCʼ- shaped in lateral view, moderately curved (Fig. 9C); slightly narrow near apex in ventral view, apex furcate, with a notch medially, lateral margins of apex concave (Fig. 9D). Parameres moderately broad, strap-like apically (Fig. 9E).</p> <p>Habitat. This species inhabits ponds, abandoned paddy fields and wetlands with aquatic plants (MORI &amp; KITAYA- MA 1993); specimens from Shenzhen (Guangdong) were found in a lowland marsh (Figs 18D–E) and specimens from Lankeshan Mt. (Zhejiang) were found in a natural pond (Fig. 18G).</p> <p>Distribution. China (Chongqing, Guangdong, Hubei, Hunan, Jiangxi, Taiwan, Yunnan, Zhejiang) (Fig. 21A), Cambodia, Indonesia (Java, Kalimantan), Japan, Laos, Malaysia, Singapore, Thailand, Vietnam (RḖGIMBART 1899, ZIMMERMANN 1920, GSCHWENDTNER 1938, MORI &amp; KITAYAMA 1993, NILSSON 1995, HENDRICH et al. 2004, HENDRICH &amp; BRANCUCCI 2013). First record from Chongqing, Guangdong, Hubei, Hunan, Jiangxi, Yunnan and Zhejiang.</p> <p>Cybister tripunctatus lateralis (Fabricius, 1798) (Figs 10A–E; 18D–F; 19D; 21B)</p> <p>Dytiscus lateralis Fabricius, 1798: 64 (orig. descr.).</p> <p>Cybister asiaticus Sharp, 1882:731 (orig. descr.), synonymy by NILSSON (2001: 90).</p> <p>Cybister gotschii Hochhuth, 1846: 214 (orig. descr.), synonymy by NILSSON (2001: 90).</p> <p>Cybister similis Régimbart, 1899: 352 (orig. descr.), synonymy by NILSSON (2001: 90).</p> <p>Cybister tripunctatus orientalis Gschwendtner, 1931: 99 (orig. descr.), synonymy by NILSSON (2001:90); FENG (1932):37 and FENG (1933a): 329 (Sichuan, Hubei:Wuchang, Fujian:Fuzhou and Xiamen, Guangdong, Hainan); BRINCK (1946): 151 (Heilongjiang); ZHAO (1981): 110 (Fujian); ZENG (1989): 91 (Fujian, Guangdong, Hainan, Hubei, Hunan, Jiangsu, Sichuan, Yunnan, Xizang, Zhejiang); LI (1992): 36 (NE China); PU et al. (1992): 484 (Yunnan); MORI &amp; KITAYAMA (1993): 142 (Taiwan).</p> <p>Cybister szechwanensis Falkenström, 1936: 238 (orig. descr.), synonymy by JIA et al. (2010: 258).</p> <p>Cybister tripunctatus lateralis: JIA et al.(2010):258 (NE China, N China, Fujian, Guangdong, Hainan, Hubei, Hunan, Jiangsu, Sichuan, Taiwan, Yunnan, Xizang, Zhejiang).</p> <p>Material examined. CHINA: CHONGQING: 1 ♀, Jinyunshan Mt., 800</p> <p>m, 13.vi.1994, Y. W. Zhang leg. (SYSU). FUJIAN: 1 J, Haitong, NR.,</p> <p>Foot How, iii.1926, H.T. Chen (SYSU); 1J, Fuqing, Xitai, 6.–13.x.1932</p> <p>(SYSU); 1 J, Nan-ping District, Yen-ping, vi.–vii.1933, D. C. Ngu</p> <p>(SYSU). GUANGDONG: 1 J 2 ♀♀, Ch’eng-hai District, K’ei-luk, Swa-</p> <p>tow, viii.1932, Y. C. Ng (SYSU); 1 J, Lingnan University, 10.vi.1948</p> <p>(SYSU); 3 ♀♀, suburbs of Guangzhou, 6.xi.1956 (SYSU); 1 J 2 ♀♀,</p> <p>Guangzhou, Honam Island, 2.–9.xi.1957, D. X. Gu leg. (SYSU); 6 JJ</p> <p>6 ♀♀, Nanhai District, Jiujiang Township, 16.xi.1957, D. X. Gu leg.</p> <p>(SYSU); 1 J 2 ♀♀, Guangzhou, Xinshi, 11.x.1964, C. M. Chen &amp; Z.</p> <p>W. Huang leg. (SYSU); 1 J, Guangzhou, Lianhe, 18.x.1964, Z. Y. Chen leg. (SYSU); 1♀, Guangzhou, Baiyunshan Mt., 2.xi.1964, J. C. Bao leg.</p> <p>(SYSU); 126 spec., Guangzhou, 14.xi.1964, T. X. Peng got from market</p> <p>(SYSU); 1♀, Guangzhou, 18.xi.1964, T. X. Peng leg.(SYSU); 1J, Nanhai District, Shijing Village, 10.x.1973, S. T. Lu leg. (SYSU); 84 spec.,</p> <p>Nanhai District, Xiqiaoshan Mt., 10.–11.x.1973, S. T. Lu leg. (SYSU); 1</p> <p>J 2 ♀♀, suburbs of Guangzhou, Renhe Township, 26.–27.x.1973, S. T.</p> <p>Lu leg. (SYSU); 1 ♀, Nanhai District, 2.xi.1973, S. T. Lu leg. (SYSU); 1</p> <p>J, Shunde District, Xingtan Township, 2.xi.1973, S. T. Lu leg. (SYSU);</p> <p>1 ♀, Guangzhou, Baiyunshan Mt., 6.xi.1973, S. T. Lu leg. (SYSU); 1 ♀, Ruyuan County, Luoyang Commune, 26.v.1974, C. Liao leg. (SYSU); 1 J, Guangzhou, Kangle, 28.iv.1989, F. L. Jia leg. (SYSU); 1 J 1 ♀, Shenzhen, Noi Ling Ding Island, 20.x.1998, H. D. Chen leg. (SYSU); 1 J 1 ♀, Xinhui District, viii.2001, X. L. Tong leg. (SYSU); 4 JJ 11 ♀♀, Gaoming District, Yangmei Township, 20.–27.iv.2006, F. L. Jia et al. leg. (SYSU); 1 J, Lian County, Dadongshan Mt., 3.–9.vii.2008, W. C. Xie leg.(SYSU); 1♀, Lianjiang City, 26. ix.2008 (SYSU); 1J, Fengkai County, Heishiding, 5.x.2011, Y. Li leg. (SYSU). GUANGXI: 2 JJ, Rong County, 10.x.1962, D. X. Gu leg. (SYSU); 4 JJ 2 ♀♀, Nanning, 19.vi.1977, Z. H. Huang leg. (SYSU). HAINAN: 1 J, Jianfengling Mt., 18.xii.1963, T. X. Peng leg. (SYSU); 3 JJ 1♀, Sanya, 24.xii.1963, T. X. Peng leg. (SYSU); 1 J, Dongfang-Huangliu, 25.xii.1963, T. X. Peng leg. (SYSU); 2 JJ, Tongshi, 27.xii.1963, T. X. Peng leg. (SYSU); 1 J, Lingshui County, Diaoluoshan Mt., Xiaomei Village, 29.xii.1963, L. F. Huang leg. (SYSU); 1 ♀, Qionghai, Shangyong Village, light trap, 17.vii.1984, C. L. Wang leg. (SYSU). HUBEI: 1 J, Zigui County, Maoping Township, 80 m, 28.iv.1994, W. Z. Li leg. (SYSU). HUNAN: 2 JJ, Tongdao County, 20.viii.1982, Z. H. Huang leg. (SYSU); 1 ♀, Hengyang, Zhoujiaao, 2.vii.1986, Q. Zou leg. (SYSU). JIANGXI: 1 ♀, Nanchang, iii.1963, X. W. Chen leg. (SYSU); 1 ♀, Nanchang (SYSU). SHANGHAI: 1J, Shanghai, 16.ii.1926, W.E.Hoffmann (SYSU). YUNNAN: 1 ♀, Xishuangbanna, Menglun Township, light trap, 580 m, 13.ix.1993, H. L. Xu leg. (SYSU); 1 ♀, Xishuangbanna Tropical Botanical Garden, 650 m, 9.viii.2003,Y.Zhang leg. (SYSU). XIZANG: 1♀, Gyirong County, Gyirong Commune, 26.vii.1984, Z. X. Yan leg. (SYSU); 5 JJ 10 ♀♀, Linzhi, Mêdog County, Beibeng Township, Didong Village, 840 m, 16.viii.2006, M. Bai leg. (SYSU); 3 JJ 1 ♀, Linzhi, Mêdog County, Beibeng Township, 887 m, 17.viii.2006, M. Bai leg. (SYSU); 3 JJ 2 ♀♀, Linzhi, Mêdog County, Yarang Township, Didong Village, 760 m, 19.viii.2006, M.Bai leg. (SYSU); 20 spec., Linzhi, Mêdog County, 1084 m, 21.viii.2006, M. Bai leg. (SYSU); 20 spec., Linzhi, Mêdog County, 1084 m, 23.viii.2006, M. Bai leg. (SYSU); 1 J, Linzhi, Mêdog County 80K, 2110 m, 24.viii.2006, M. Bai leg. (SYSU).</p> <p>Diagnosis. TL 24–28 mm, elongate, oblong-oval (Fig. 10A). Head dark green, clypeus yellow. Pronotum and elytra dark green to black, with distinct yellow margins laterally, apex of yellow margin on the elytra slightly hooked. Ventral surface black, sometimes with lateral part of metaventrite and metacoxae yellow; abdominal ventrites III–V with a yellow spot on each side (Fig. 10B). Dorsal surface smooth in both sexes. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs reddish-brown, midlegs and hindlegs reddish-brown to black; protarsomeres 1–3 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; metatarsal claws single in both sexes. Median lobe of aedeagus ʻCʼ- shaped in lateral view, moderately curved (Fig. 10C); slightly narrow near apex in ventral view, apex furcate, with a deep acute-angled notch medially, lateral margins of apex convex (Fig. 10D). Parameres moderately broad, tapering apically (Fig. 10E). Habitat. The most common species of the genus in Southeast Asia. An eurytopic species which is widespread in artificial habitats such as irrigation channels, flooded paddy fields, open swamps, fish and ornamental ponds and larger ditches, even in urban and suburban areas (MORI &amp; KITAYAMA 1993, HENDRICH et al. 2004). Specimens from Shenzhen (Guangdong) were found in a lowland marsh(Figs 18D–E) and specimens from Danxiashan Mt. (Guangdong) were found in a temporary pond in a valley (Fig. 18F).</p> <p>Distribution. China (Chongqing, Fujian, Guangdong, Guangxi, Hainan, Hebei, Heilongjiang, Hubei, Hunan, Jiangsu, Jiangxi, Macao, Shanghai, Sichuan, Taiwan, Yunnan, Xizang, Zhejiang) (Fig. 21B), Afghanistan, United Arab Emirates, Azerbaijan, Bhutan, Cyprus, India, Iran, Iraq, Japan, Kyrgyzstan, Laos, Mongolia, Nepal, Oman, Pakistan, Russia, Singapore, Syria, Thailand, Tajikistan,</p> <p>Turkmenistan, Turkey, Uzbekistan (RḖGIMBART 1899; ZIMMERMANN 1920; FENG 1932, 1933a,b; GSCHWENDTNER 1938; VAZIRANI 1969; ZAITZEV 1972; ZENG 1989; MORI &amp; KITAYAMA 1993; HENDRICH et al. 2004; GHOSH &amp; NILSSON 2012; HENDRICH &amp; BRANCUCCI 2013; NILSSON &amp; HÁJEK 2022b). First record from Chongqing, Guangxi, Jiangxi and Shanghai.</p></div> 	https://treatment.plazi.org/id/03975B72FFC9760796D7CB4D5385FE4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFCC76199465CCAC5208FDCF.text	03975B72FFCC76199465CCAC5208FDCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister ventralis Sharp 1882	<div><p>Cybister ventralis Sharp, 1882</p> <p>(Figs 11A–E; 22A)</p> <p>Cybister ventralis Sharp, 1882: 742 (orig. descr.); KANO (1931): 177</p> <p>(Taiwan); FENG (1933a): 329 (Sichuan, Shandong: Qingdao).</p> <p>Material examined. None from China. INDIA: 1 J 2 ♀♀, Behar, India or., Coll.A. Zimmermann (ZSMG). SRI LANKA: 1♀, Negombo, 1899, W. Horn, Coll. A. Zimmermann (ZSMG).</p> <p>Diagnosis. TL 28–32 mm, broad, oblong-oval (Fig. 11A). Head dark green, clypeus yellow. Pronotum dark green to black with lateral yellow margins projecting a little inwards along anterior margin behind eyes; elytra dark green to black, with distinct yellow margin laterally, apex of yellow margin on elytra hooked. Ventral surface yellow with central part of metaventrite and metacoxae black; abdominal ventrites III–V yellowish with a dark brown band anteriorly (Fig. 11B). Dorsal surface dimorphic in females: smooth or with isolated short, irregular striae on pronotum and dense longitudinal striae of three-fourths of elytral length from base. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs and midlegs yellow, tarsi darker; metafemur and metatibia yellow with broad dark brown marking; female with a single metatarsal claw. Median lobe of aedeagus ʻCʼ- shaped in lateral view, moderately curved (Fig. 11C); slightly narrow near apex in ventral view, apex furcate, with a notch medially, lateral margins of apex concave (Fig. 11D). Parameres more elongate, strap-like apically (Fig. 11E).</p> <p>Distribution. China (Macao, Sichuan, Shandong, Taiwan) (Fig. 22A), Bangladesh, India, Myanmar, Pakistan, Sri Lanka (RḖGIMBART 1899, ZIMMERMANN 1920, KANO 1931, FENG 1933a, VAZIRANI 1969, NILSSON 1995, GHOSH &amp; NILSSON 2012, NILSSON &amp; HÁJEK 2022b).</p></div> 	https://treatment.plazi.org/id/03975B72FFCC76199465CCAC5208FDCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFD376199446CE2E556CFE8E.text	03975B72FFD376199446CE2E556CFE8E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister (Melanectes) Brinck 1945	<div><p>Subgenus Melanectes Brinck, 1945</p> <p>Type species. Cybister owas Laporte, 1835.</p> <p>Diagnosis. Without distinctive yellow or pale margins on the elytron and pronotum. Males with a single metatarsal claw; females with two claws, the posterior claw small.</p> <p>Included (Chinese) species. Cybister brevis Aubé, 1838; C. convexus Sharp, 1882; C. danxiaensis sp. nov.; C. posticus Aubé, 1838; C. sugillatus Erichson, 1834.</p> </div>	https://treatment.plazi.org/id/03975B72FFD376199446CE2E556CFE8E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFD3761B969CCD6C52D1FE6F.text	03975B72FFD3761B969CCD6C52D1FE6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister brevis Aube 1838	<div><p>Cybister brevis Aubé, 1838</p> <p>(Figs 12A–E; 18G; 22A)</p> <p>Cybister brevis Aubé, 1838:98 (orig.descr.); KANO (1931): 177 (Taiwan); FALKENSTRÖM (1936): 238 (Sichuan); LI (1992): 36 (Liaoning).</p> <p>Material examined. CHINA: GUIZHOU: 1 J, Jiangkou County, Shengxisi, 19.vi.1984 (SYSU); 1 J, west slope of Fanjingshan Mt., Huguosi, 1.viii.2001, H. Pang leg. (SYSU). HUNAN: 1 J, Longshan County, Tongche, 5.vi.1965, Z. Y. Chen leg. (SYSU). ZHEJIANG: 2 JJ 1 ♀, Quzhou, Kecheng district, Lankeshan Mt., 28.8781N, 118.9199E, 118 m, 18.iii.2020, Z. Y. Jiang leg. (SYSU).</p> <p>Diagnosis. TL 20–25 mm, elongate, oblong-oval (Fig. 12A). Head dark green, clypeus orange. Pronotum and elytra dark green to black, apex of elytra with reddish-brown spots. Ventral surface black; abdominal ventrites III–IV with a yellow spot on each side (Fig. 12B). Head and pronotum densely covered with coarse punctures, elytra sparsely covered with inconspicuous small granules. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs reddish-brown, midlegs and hindlegs reddish-brown to black; protarsomeres 1–3 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; male with a single metatarsal claw; female with two metatarsal claws, posterior claw small and sinuate. Median lobe of aedeagus ʻCʼ- shaped in lateral view, slightly curved, gradually broader from base to apex, then strongly narrowing at apex (Fig. 12C); broad medially in ventral view, apex obtuse-angled (Fig. 12D). Parameres moderately broad, strap-like apically (Fig. 12E).</p> <p>Habitat. This species inhabits ponds, abandoned paddy fields and ditches of paddy fields with aquatic plants (MORI</p> <p>&amp; KITAYAMA 1993); specimens from Lankeshan Mt. (Zhejiang) were found in a natural pond (Fig. 18G).</p> <p>Distribution. China (Guizhou, Hunan, Liaoning, Sichuan, Taiwan, Zhejiang) (Fig. 22A), Japan, South Korea (RḖGIMBART 1899, ZIMMERMANN 1920, FENG 1933a, GSCHWENDTNER 1938, ZAITZEV 1972, MORI &amp; KITAYAMA 1993, NILSSON 1995, NILSSON &amp; HÁJEK 2022b). First record from Guizhou, Hunan and Zhejiang.</p></div> 	https://treatment.plazi.org/id/03975B72FFD3761B969CCD6C52D1FE6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFD1761B9459CDCD521EF8EF.text	03975B72FFD1761B9459CDCD521EF8EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister convexus Sharp 1882	<div><p>Cybister convexus Sharp, 1882</p> <p>(Figs 13A–E; 18H; 19E; 22A)</p> <p>Cybister convexus Sharp, 1882: 718 (orig. descr.); RḖGIMBART (1899): 353 (Yunnan); FENG (1932): 35 (Yunnan); VAZIRANI (1969): 284 (Yunnan); ZENG (1989): 93 (Yunnan).</p> <p>Material examined. CHINA: YUNNAN: 1J, Tia Feng on (Grassy hill 3ml. N Yunnanfu). 25.viii.1934, C. Brownell leg. (SYSU); 1 ♀, Honghe Hani- Yi Autonomous Prefecture, Lvchun County, Niukong Township Section, terrace, 22.9873N, 102.2675E, 1336 m, 30.iv.2021,Z.Y.Jiang, Z.M.Yang, B. P.Huang &amp; Z. Q. Mai leg. (SYSU); 1 ♀, Dehong Dai Aut. Pref. mount. Range 31 km E Luxi, 2280 m, 24°29′31″N 98°52′58″E, 3.vi.2007, D.W. Wrase leg. (LHCM); 1 J, Yunnan, Baoshan Pref. mount. Range, 22 km S Tengchong, 1750m, 24°49′29″N 98°29′27″E (in fishpond), 30.viii.2009, D. W. Wrase leg. (LHCM).</p> <p>Diagnosis. TL 24–30 mm, elongate, oblong-oval (Fig. 13A). Head, pronotum and elytra dark green to black, apex of elytra with reddish-brown spots. Ventral surface black; abdominal ventrites III–IV with a yellow spot on each side (Fig. 13B). Head and pronotum smooth, elytra densely covered with conspicuous small granules. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Legs black; protarsomeres 1–3 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; male with a single metatarsal claw; female with two metatarsal claws, posterior claw small and sinuate. Median lobe of aedeagus ʻCʼ- shaped in lateral view, slightly curved, broadest at middle, strongly narrowing in apical fifth, apex rounded (Fig. 13C); broad medially in ventral view, apical fifth slender like a tubular spine, apex truncated (Fig. 13D). Parameres moderately broad, strap-like apically (Fig. 13E).</p> <p>Habitat. The habitat information is not well known for this species. It can be found from hilly to high mountain regions (ATTHAKOR et al. 2018). In Yunnan, it was collected in a water-filled rut on a forest track from Dehong (HENDRICH 2008) and in a fish pond. In Lvchun County it was collected at night from terraced fields (Fig. 18H).</p> <p>Distribution. China (Yunnan) (Fig. 22A), India, Thailand (RḖGIMBART 1899; ZIMMERMANN 1920; FENG 1932, 1933b; GSCHWENDTNER 1938; VAZIRANI 1969; ZENG 1989; GHOSH &amp; NILSSON 2012; ATTHAKOR et al. 2018).</p></div> 	https://treatment.plazi.org/id/03975B72FFD1761B9459CDCD521EF8EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFD1761D9598CB4D5265FDCF.text	03975B72FFD1761D9598CB4D5265FDCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister (Melanectes) danxiaensis Jiang & Zhao & Mai & Jia & Hendrich 2023	<div><p>Cybister (Melanectes) danxiaensis sp. nov.</p> <p>(Figs 14A–E; 15A–E; 22B)</p> <p>Type locality. China, Guangdong Province, Shaoguan, Renhua County, Danxiashan Mt., ca. 25.0244N, 113.7158E.</p> <p>Type material. HOLOTYPE: J (SYSU), labelled:“tṭ ṯffl ñȇ / 1941. VIII / Hľ [hw] // CHINA: Guangdong Province / Renhua County /</p> <p>Danxiashan Mt. / viii.1941, Ping Lin leg. [p] // HOLOTYPE / CYBISTER / danxiaensis sp. nov. / Jiang, Zhao, Mai &amp; Jia det. 2021 [red label, p]”.</p> <p>Description. Habitus (Fig. 15A) elongate oblong oval, with continuous outline, widest in apical third of elytra, moderately convex. Dorsal surface distinctly matt. TL 21.7 mm; TL-h 19.4 mm. MW 11.3 mm.</p> <p>Colouration. Head black; pronotum black with broad brown sides; elytra blackish brown; appendages black; ventral side blackish brown.</p> <p>Head moderately broad, ca. 0.6× width of pronotum, trapezoidal. Surface densely covered with fine punctures. A pair of depressions present near anterior margin of clypeus. Anterior margin of clypeus concave. Labium and genae with some punctures near gular suture.Antenna with antennomeres long and slender.</p> <p>Pronotum transverse (width/length ratio = 2.36), broadest between posterior angles, lateral margins moderately curved, without lateral bead. Punctation double; coarse setigerous puncture row present along anterior margin, with a perpendicular setigerous puncture row laterally that does not reach posterior margin; fine punctures dense throughout whole surface. Centre of disc with a moderately impressed longitudinal line medially.</p> <p>Elytra with dense large protuberances that are arranged almost in rows; each protuberance with a puncture medially in which is a small granule (Fig. 14A). Punctation double; larger punctures sparse between protuberances; finer punctures dense throughout whole surface. Medioapical area of elytra with a field of short, coarse setae (Fig. 14B).</p> <p>Ventral side (Fig. 15B). Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed (Fig. 14C). Lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures (Fig. 14D). Lateral part of abdominal ventrites without spot.</p> <p>Legs. Protarsomeres 1–3 with four rows of adhesive setae on their ventral side, broadly expanded into a palette; claws simple. Mesotibia with setae in a continuous line on posteroapical margin; mesotarsomeres with two separate series of setae present along posteroventral and posterodorsal apical margin; claws simple (Fig. 14E). Hindlegs lost.</p> <p>Male genitalia. Median lobe of aedeagus ʻCʼ- shaped in lateral view, strongly curved, broadest at middle, strongly narrowing in apical fourth, apex rounded (Fig. 15C); in ventral view broad at base, apical fourth slender like a tubular spine, apex rounded (Fig. 15D). Parameres moderately broad, distinctly curved in apical third, strap-like apically (Fig. 15E).</p> <p>Female. Unknown.</p> <p>Differential diagnosis. Based on the pronotum and elytra without distinct lateral yellow margins, the new species can be assigned to subgenus Melanectes (MILLER et al. 2007). Within the subgenus, the new species is similar to Cybister sugillatus but can easily be recognized by the large punctate protuberances on the elytra and the longer apical slender part of the median lobe of the aedeagus (elytra smooth without punctate protuberances, apical slender part of median lobe of aedeagus shorter in C. sugillatus).</p> <p>Etymology. The species is named after the type locality, Danxiashan Mt., Guangdong Province, China; the name is an adjective.</p> <p>Distribution. So far known only from the type locality in northern Guangdong, China (Fig. 22B).</p></div> 	https://treatment.plazi.org/id/03975B72FFD1761D9598CB4D5265FDCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFD7761F9688CCAC52A2FF2F.text	03975B72FFD7761F9688CCAC52A2FF2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister posticus Aube 1838	<div><p>Cybister posticus Aubé, 1838</p> <p>(Figs 16A–E; 22B)</p> <p>Cybister posticus Aubé, 1838: 87 (orig. descr.); JIA et al. (2010): 259 (Xizang: Zayü County).</p> <p>Material examined. CHINA: XIZANG: 1 J, Mêdog County, Beibeng</p> <p>Township, 800 m, 5. ix.1974, F. S. Huang leg. (SYSU); 1 ♀, Bomê</p> <p>County,Yigong, 1450 m, 9.v.1983,Y.H.Han leg.(SYSU); 1J 1♀, Zayü</p> <p>County, Xia Zayü, 28.50N 97.00E, 4.–8.x.2007, F. M. Shi leg. (SYSU).</p> <p>Diagnosis. TL 23–35 mm, elongate, oblong-oval (Fig. 16A). Head dark green, clypeus reddish-brown. Pronotum and elytra dark green to black, apex of elytra with reddish-brown spots.Ventral surface black; abdominal ventrites III–IV with a yellow spot on each side (Fig. 16B). Head and pronotum smooth, elytra densely covered with conspicuous small granules. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs reddish-brown, midlegs and hindlegs reddish-brown to brown; protarsomeres 1–3 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; male with a single metatarsal claw; female with two metatarsal claws, posterior claw small and sinuate. Median lobe of aedeagus ʻCʼ- shaped in lateral view, slightly curved, broadest at middle, strongly narrowing in apical fifth, apex rounded (Fig. 16C); broad medially in ventral view, apical fifth slender like a tubular spine, apex rounded (Fig. 16D). Parameres moderately broad, strap-like apically (Fig. 16E).</p> <p>Habitat. Although this species is widely distributed in India, nothing is known about its habitat.</p> <p>Distribution. China (Xizang) (Fig. 22B), Bangladesh, India, Nepal, Pakistan (RḖGIMBART 1899, ZIMMERMANN 1920, VAZIRANI 1969, JIA et al. 2010, GHOSH &amp; NILSSON 2012).</p></div> 	https://treatment.plazi.org/id/03975B72FFD7761F9688CCAC52A2FF2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFD5761F944FCD0D549CFD4F.text	03975B72FFD5761F944FCD0D549CFD4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister sugillatus Erichson 1834	<div><p>Cybister sugillatus Erichson, 1834</p> <p>(Figs 17A–E; 18D–E, H; 19F; 22B)</p> <p>Cybister sugillatus Erichson, 1834: 227 (orig.descr.); RḖGIMBART (1899): 355 (Xizang); ZIMMERMANN (1919): 77 (Taiwan); KANO (1931): 176 (Taiwan); FENG (1932): 36 (Beijing, Fujian: Fuzhou and Xiamen, Guangdong, Hainan, Hubei: Wuchang, Jiangxi, Sichuan, Xizang, Zhejiang); FALKENSTRÖM (1936): 238 (Sichuan); ZHAO (1981): 110 (Fujian); ZENG (1989): 92 (Beijing, Fujian, Guangdong, Hainan, Hubei, Jiangxi, Sichuan, Xizang, Zhejiang); MORI &amp; KITAYAMA (1993): 142 (Taiwan).</p> <p>Cybister bisignatus Aubé, 1838: 88 (orig. descr.), synonymy by SHARP (1882: 991).</p> <p>Cybister notasicus Aubé, 1838: 90 (orig. descr.), synonymy by SHARP (1882: 717).</p> <p>Cybister olivaceus Boheman, 1858: 21 (orig. descr.), synonymy by SHARP (1882: 717).</p> <p>Material examined. CHINA: GUANGDONG: 8JJ 7♀♀, Guangzhou, 14.xi.1964, T. X. Peng purchased from market (SYSU); 1 ♀, Ruyuan County, Tianjingshan Mt., 8.viii.1980, Q. X. Zhang leg. (SYSU); 1 J, Fengkai County, Heishiding, 10.iv.1985, W. Wu leg. (SYSU); 1 J, Fengkai County, Heishiding, 14.vii.1999, Y. C. Wang leg. (SYSU); 1 ♀, Fengkai County, Heishiding, 12.vii.2000, Y. L. Feng leg. (SYSU); 2 JJ 2 ♀♀, Gaoming District, Yangmei Township, 20–27.iv.2006, D. P. Wu, N. Yu, Q. S. Liu &amp; Y. Y. Niu leg. (SYSU); 1 J, Zhuhai, back mountain of Sun Yat-Sen University, 5.–8.vii.2011, F. L. Jia leg. (SYSU). GUANGXI: 1 ♀, Rong’an County, Dajiang-Yayao, 28.iv.1959 (SYSU); 2 JJ 1 ♀, Nanning, iii.2001, L. Hojný leg. (NMPC); 1 J, Jingxi, Bangliang Nature Reserve, 1.viii.2010, J. H. Huang leg. (SYSU). HAINAN: 1J, Cheung-kon-ts’uen, Ka-luk-kong (18ml. E.of Nam-fung), Kiung-shan Dist., 29.–31.iii.1935, F. K. To leg. (SYSU); 1 ♀, Lingshui County, 1.i.1964, T. X. Peng leg. (SYSU); 1 ♀, Jiangfengling, the third partition, 24.xi.1981, J. P. Lu leg. (SYSU); 1 J, Jiangfengling, Heiling, 10.vii.1982, L. Z. Hua leg. (SYSU). HONG KONG: 1 J, Hong Kong, 1934 (SYSU). YUNNAN: 1 J 1 ♀, Chengjiang County, 30.iv.1932, Z. L. Pu leg. (SYSU).</p> <p>Diagnosis. TL 18–23 mm, elongate, oblong-oval (Fig. 17A). Head, pronotum and elytra dark green to black, lateral part of pronotum reddish-brown, apex of elytra with reddish-brown spots. Ventral surface black; abdominal ventrites III–IV with a yellow spot on each side (Fig. 17B). Dorsal surface smooth. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Forelegs reddish-brown, midlegs and hindlegs reddish-brown to black; protarsomeres 1–3 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; male with a single metatarsal claw; female with two metatarsal claws, posterior claw small and sinuate. Medi- an lobe of aedeagus ʻCʼ- shaped in lateral view, strongly curved, broadest at middle, strongly narrowing in apical fifth, apex rounded (Fig. 17C); broad medially in ventral view, apical fifth slender like a tubular spine, apex rounded (Fig. 17D). Parameres moderately broad, strap-like apically (Fig. 17E).</p> <p>Habitat. This species inhabits not only ponds, wetlands, abandoned paddy fields and ditches of paddy fields with aquatic plants, but also eutrophic waters near grazing land</p> <p>(MORI &amp; KITAYAMA 1993). Specimens from Shenzhen (Guangdong) were found in in a lowland marsh (Figs 18D–E) and specimens from Lvchun County (Yunnan) were found in terraced fields (Fig. 18H).</p> <p>Distribution. China (Beijing, Fujian, Guangdong, Guangxi, Hainan, Hebei, Hong Kong, Hubei, Jiangxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang) (Fig. 22B), Afghanistan, Bhutan, India, Indonesia, Japan, Laos, Malaysia, Myanmar, Nepal, Pakistan, Philippines, Singapore, Sri Lanka (RḖGIMBART 1899; ZIMMERMANN 1920; FENG 1932, 1933b; GSCHWENDTNER 1938; VAZIRANI 1969; ZAITZEV 1972; ZENG 1989; MORI &amp; KITAYAMA 1993; NILSSON 1995; HENDRICH et al. 2004; GHOSH &amp; NILSSON 2012; HENDRICH &amp; BRANCUCCI 2013; NILSSON &amp; HÁJEK 2022b). First record from Guangxi and Yunnan.</p></div> 	https://treatment.plazi.org/id/03975B72FFD5761F944FCD0D549CFD4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
03975B72FFD5761196D2CEAE55C7F9A0.text	03975B72FFD5761196D2CEAE55C7F9A0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cybister Curtis 1827	<div><p>Key to the Chinese species of Cybister</p> <p>1 Pronotum and elytra with yellow margins laterally; female with a single metatarsal claw. Subgenus Cybister Curtis, 1827....................................................2</p> <p>– Pronotum and elytra without yellow margins laterally; female with two metatarsal claws. Subgenus Melanectes Brinck, 1945....................................... 11</p> <p>2 Apex of median lobe of aedeagus furcate in ventral view........................................................................ 3</p> <p>– Apex of median lobe of aedeagus not furcate in ventral view.................................................................. 9</p> <p>3 Metaventrite and metacoxae black with lateral parts yellow..................................................................... 4</p> <p>– Metaventrite and metacoxae entirely black............ 7</p> <p>4 Smaller species, TL: 22–28 mm; female with dorsal surface smooth (Fig. 3A)........................................ 5</p> <p>– Larger species, TL: 27–33 mm; female dorsal surface with dense longitudinal striae of various lengths on elytra except at its apex (Fig. 11A)........................ 6</p> <p>5 Smaller species, TL: 22–24 mm; median lobe of aedeagus with apical notch shallower, lateral margins of apex straight and parallel in ventral view (Fig. 3D)..................................... C. fumatus Sharp, 1882</p> <p>– Larger species, TL: 24–28 mm; median lobe of aedeagus with apical notch deeper, lateral margins of apex convex in ventral view (Fig. 10D).................... C. tripunctatus lateralis (Fabricius, 1798) (partim) and C. laevis Falkenström, 1936</p> <p>6 Abdominal ventrites III–V black, with lateral yellow spots (Figs 1B; 9B)....... C. rugosus (W. S. Macleay, 1825) and C. bengalensis Aubé, 1838</p> <p>– Abdominal ventrites III–V yellowish, with a dark brown band anteriorly (Fig. 11B)......................................................................... C. ventralis Sharp, 1882</p> <p>7 Smaller species, TL 24–28 mm; median lobe of aedeagus with acute-angled apico-medial notch (Fig. 10D)...... C. tripunctatus lateralis (Fabricius, 1798) (partim)</p> <p>– Larger species, TL&gt;30.0 mm; median lobe of aedeagus with an obtuse-angled apico-medial notch...... 8</p> <p>8 Median lobe of aedeagus in ventral view with apex and base almost equal in width, lateral margins of apex sinuate, notch of apex shallower (Fig. 4D)..................................................... C. guerini Aubé, 1838</p> <p>– Median lobe of aedeagus in ventral view with apex narrower than base, lateral margins of apex concave, notch of apex deeper (Fig. 8D).................................................................... C. limbatus (Fabricius, 1775)</p> <p>9 Smaller species, TL: 23–27 mm; ventral surface black with lateral parts of metaventrite and metacoxae reddish-brown (Fig. 7B); median lobe of aedeagus slender in ventral view (Fig. 7D); female with dorsal surface smooth.............................................................................. C. lewisianus Sharp, 1873</p> <p>– Larger species, TL&gt; 29 mm; ventral surface yellow with middle parts of metaventrite and metacoxae reddish-brown or dark green; median lobe of aedeagus thick in ventral view; female dorsal surface with isolated short, irregular striae on pronotum, and with dense longitudinal striae of various lengths on elytra except at its apex.................................................. 10</p></div> 	https://treatment.plazi.org/id/03975B72FFD5761196D2CEAE55C7F9A0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Jiang, Zhuo-Yin;Zhao, Shuang;Mai, Zu-Qi;Jia, Feng-Long;Hendrich, Lars	Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long, Hendrich, Lars (2023): Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae 63 (1): 75-102, DOI: 10.37520/aemnp.2023.003, URL: http://dx.doi.org/10.37520/aemnp.2023.003
