taxonID	type	description	language	source
0394879B435A103DCABBF9CCA28FFC96.taxon	description	quadrate; anterior margin bisinuate, not deeply bilobed; seta present in each anterior angle; punctate laterally and basally. Scutellum: elongate, triangular. Elytron: margined laterally; apices conjointly rounded; with 10 rows of punctures; non-costate; male without apical appendage. Venter: mesocoxae separated by the diameter of a coxa; prosternal process projecting between mesocoxae; pro- and mesotibiae expanded at apex. Male genitalia: aedeagus curved, apex acute; apical orifice nearly as wide as apex of aedeagus. Species of Chelobasis are extremely variable in coloration and markings. There have been subspecies and varieties proposed but these are merely color forms and are herein treated as synonyms under each species. The male and female genitalia were examined and found not to have any taxonomic value. Sexual dimorphism is absent.	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B435A103DCABBF9CCA28FFC96.taxon	biology_ecology	Biology (from Strong 1983). Chelobasis bicolor Gray and C. perplexa Baly develop in the rolled leaves of various species of Zingiberales. Eggs are laid on wet, tender tissue of the host plant and hatch in about 20 days. Larvae begin feeding in rolled leaves immediately after hatching. Development is slow, requiring at least eight months until pupation. Larvae require more than one leaf-roll to complete development and move from maturing leaf-rolls to more tender leaf-rolls at night. Larvae are water penny-like, but much larger and more elongate than Cephaloleia Chevrolat, 1837. Adults are polymorphic and long-lived. In marked recapture studies, adult beetles were found up to 18 months after marking. It is assumed that other species in this genus will be found on Zingiberales.	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B435B103ECABBFA80A762F9D4.taxon	description	(Figures 1, 5)	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B435B103ECABBFA80A762F9D4.taxon	description	Larva. Unknown. Larval host plant. Unknown. Comparative notes. This species can be easily distinguished by the rounded elytral apices, the angulate lateral margin of the pronotum, and by antennomere 2 being transverse.	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B435B103ECABBFA80A762F9D4.taxon	distribution	Distribution. Ecuador. Specimens examined. Ecuador. Carchi: Gabriel, 750 m, 12. x. 1970 (BPBM). Napo: Sarayacu (BMNH). Pinchincha: Estacion Orongo Palmitopomba, 23. vii. 2001 (USNM). Total: 8.	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B4358103ACABBF915A0C5FDBE.taxon	description	(Figures 2, 6, 9)	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B4358103ACABBF915A0C5FDBE.taxon	description	Arescus labiatus Baly (not Perty) 1858: 84 (misidentification).	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B4358103ACABBF915A0C5FDBE.taxon	description	Larva. (from Maulik 1931). Total length 9 – 13.5 mm. Total width 5.5 – 7.0 mm. Body flat, upper surface somewhat convex longitudinally; sides expanded into interlacing flaps which have a fringe of fine setae; ovoid. Head: covered from above by two overlapping expansions; epicranial lobes bifurcate; antennae well developed, three segmented; mandible tridentate; maxillary palps with two segments; labial palps with one segment; four ocelli placed near outer margin of each antenna; legs widely separated; spiracles visible from above, spiracle 8 larger than others. See Figure 9. Larval host plant. Wild banana (Maulik 1931); Heliconia sp. (Heliconiaceae); Musa sp. (Musaceae) (Maulik 1937). Adults have been collected on Heliconia sp., H. curtispatha Peterson, H. imbricata (Kuntze) Baker, H. irrasa R. R. Sm., H. latispatha Benth., H. metallica Planch. & Linden, H. platystachys Baker, H. tortuosa Griggs, H. vaginalis Benth., and H. wagneriana Peterson (Strong 1977 a, 1977 b, 1983); H. cathaeta R. R. Sm., Calathea latifolia Klotzsch, (Marantaceae) (Meskins et al. 2008). Comparative notes. This species can be distinguished by the rounded elytral apices, by the lateral margin of the pronotum not angulate in middle, and by antennomere 2 being cylindrical.	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B4358103ACABBF915A0C5FDBE.taxon	distribution	Distribution. Bolivia, Colombia, Costa Rica, Ecuador, French Guyana, Guatemala, Nicaragua, Panama. Specimens examined. Bolivia. Beni: Cosinecho Regn., viii. 1925 (USNM); Isiamas, xii (USNM); Rurrenabaque, x, xi (USNM). Colombia. ??: Pto. Berrie, 8. viii. 1938 (USNM). Valle del Cauca: Lago Calima, 5 miles below dam, 1250 m, 24. iv. 1996 (FSCA). Costa Rica. Alajuela: Angeles, R. B. Alberto Manuel Brenes, 900 – 1000 m (INBio); Dos Rios, P. N. Guanacaste, 600 – 700 m (INBio); 10 mi. N. Varablanca, 14. v. 1967 (FSCA). Cartago: Aquiares nr. Santa Cruz, 9 km NW Turrialba, 16. v. 1985 (EMEC); Hacienda de Movaria de Chirripo, 7 – 10. vii. 1969 (LACM); Orori, P. N. Tapantí-Macizo de la Muerte, 1200 – 1300 m (INBio); Santa Teresita, 1100 – 1200 m (INBio); Tayutic, 1100 – 1200 m (INBio); Turrialba, 13 – 14. viii. 1968 (FSCA), 4 – 13. viii. 1970 (BPBM). Guanacaste: Curubandé, P. N. Rincón de la Vieja, 800 – 900 m (INBio); Mayorga, P. N. Guanacaste, 600 – 700 m (INBio); Santa Cecilia, P. N. Guanacaste, 600 – 700 m (INBio); Santa Elena, P. N. Guanacaste, 100 – 200 m (INBio); Tierras Morenas, Z. P. Tenorio, 900 – 1000 m (INBio); Tilaran, 7. vii. 1972 (FSCA); 3 mi west Tilaran (USNM). Heredia: Horquetas, 600 – 700 m (INBio); Est. Biol. La Selva, various dates (BYUC, FCSA, INBio, USNM); La Virgen, 0 – 100 m (INBio); 11 km south La Virgen, 9. iv. 2003 (USNM). Limón: Bratsi, R. I. Talamanca, 0 – 100 m (INBio); Colorado, Tortuguero, 0 m (INBio); Hdas. La Suerte, Tapezco, 13 – 31. viii. 1979 (LACM); La Amistad Caribe, 0 – 100 m (INBio); 7 miles north Guacimo, 22. ii – 3. iii. 1988 (USNM); Hamburg Farm (USNM); Rita, 0 – 100 m (INBio); Siquirres, 100 – 200 m, 14. viii. 1970 (BPBM); Valle La Estrella, R. B. Hitoy Cerere, 100 – 200 m (INBio). Puntarenas: Barranca near Puntarenas, 6. vii. 1972 (FSCA); Fca. Las Cruces, San Vito de Java, 27. vi. 1969 (USNM); 8.1 mi. W Esparta, 5. vi. 1967 (FSCA); 10.9 mi. east Esparta, 17. vi. 1969 (USNM); Guacimal, 1000 – 1100 m (INBio); Jiménez, P. N. Corcovado, 0 – 100 m (INBio); La Palma (USNM); Los Alturas, 22. v. 1992 (USNM), 23 – 26. v. 1992 (USNM); Finca Las Cruces, 6 km S San Vito, 21 / 25. viii. 1976 (CASC); Los Alturas Biol. Stn., 12. viii. 1995 (BYUC); Monteverde, various dates (INBio, USNM, CASC, EMEC, LACM); Pittier, P. N. La Amistad, 1700 – 1800 m (INBio); 5.4 miles south Palmar Sur, 11. viii. 1969 (USNM); Potrero Grande, P. N. La Amistad, 1400 – 1500 m (INBio); Barranca site, 10 km north Puntarenas, 17. vi. 1969 (USNM); Monteverde Cloud Forest Preserve, 20. vi. 1984 (BYUC); Puerto Cortes, 19. vii. 1972 (FSCA); Rincon, 18. ii. 1966 (PMNH); Rincon, Osa Peninsula, 27. vii. 1972 (FSCA); 3.5 miles south Rincon, 1. iii. 1969 (USNM), 28. ii – 12. iii. 1969 (CASC); Rincon Forest, 26. vi. 1983 (LACM); Rio Piedras, 15. viii. 1969 (USNM); San Vito-Neilly area, 13. viii. 1969 (USNM); Sierpe, R. F. Golfo Dulce, 200 – 300 m (INBio); near Sirena, Osa Peninsula, 0 – 5 m, xii. 1983 (CMNH); Tárcoles, 0 – 100 m (INBio). San José: San Jose, vi. 1927 (USNM); San Pedro, 1200 – 1300 m (INBio). Ecuador. Carchi: San Gabriel, 750 m, 12. x. 1970 (BPBM). Napo: Yasuni, 10 – 13. ciii. 1998 (USNM); road between E. Puyo & Pt. Nano, vii. 1964 (USNM). Pichincha: Chimbo (USNM). Nicaragua. Matagalpa Department: Jinotega, 20. xi. 1994 (USNM), 22. iii. 1995, 9. vi. 1995 (CMN); 6 km north Matagalpa, Selva Negra Hotel (USNM). Panamá. Chiriquí: Barro Colorado Island (USNM); Bugaba (USNM); Caldera, 31. v. 1994 (CMNH); Cerro Azul, 5. vi. 1994 (USNM); Vulcan de Chiriquí (USNM); 11.2 km east Chiriquí, 30. v. 1993 (USNM); Continental Divide Trail (USNM); Fortuna, 17. v. 1978 (BYUC), 19. v. 1978, 26. v. 1993 (USNM), 30. v. 1994 (CMNH); Hartmanns Finca, 4 – 7. vii. 1997 (FSCA, USNM, UGAC, EMEC); Santa Clara, 23 – 25. v. 1980 (USNM). Coclé: El Valle, 14. vi. 1967 (USNM), no date (USNM); Cerro Gaital, 28. v. 1994 (USNM); Gamboa (USNM); Limbo, 24. viii. 1970 (BPBM); Madden Forest (USNM). Colón: Porto Bello (USNM). Panamá: Cerro Azula, Los Altos, 24. v. 1994 (CMNH); Cerro Campana, 27. v. 1994; Old Gamboa Road, 25. vi. 1994 (CMNH); Reserva Soberania, Pipeline Road (USNM); Pipeline Road K 1 - 12, 8 – 9. vii. 1997 (EMEC). San Blas: Nasagandi, 27 – 28. v. 1995 (BYUC). Total: 1073.	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B435C103ACABBFD23A741F9C6.taxon	description	(Figures 3, 7)	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B435C103ACABBFD23A741F9C6.taxon	description	Larva. Unknown. Larval host plant. Unknown. Comparative notes. This species can be distinguished by the angulate elytral apices, by antennomere 3 being subequal in length to 1 and 2 combined, and by antennomere 2 being transverse.	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B435C103ACABBFD23A741F9C6.taxon	distribution	Distribution. Colombia. Specimens examined. Colombia. Magdalena: Medellin (BMNH). Total: 1.	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B435C103BCABBF90BA260F839.taxon	description	(Figures 4, 8, 10)	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B435C103BCABBF90BA260F839.taxon	description	Larva. (from Maulik 1932). Total length 12 – 19 mm. Width 6.5 – 9.0 mm. Very similar to Ch. bicolor. Finely rugose dorsally, surface slightly shining (Fig. 10). Larval host plants. Heliconia imbricata and Calathea insignis Peterson (Maulik 1932). Adults collected from H. lathispatha (Strong 1977 a, 1977 b, 1983); H. pogonantha Cuford. (McKenna & Farrell 2005); H. irrasa, H. mariae Hook. (Meskins et al. 2008). Comparative notes. This species can be distinguished by the angulate elytral apices, by antennomere 3 being longer than 1 and 2 combined, and by antennomere 2 being cylindrical.	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
0394879B435C103BCABBF90BA260F839.taxon	distribution	Distribution. Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Nicaragua, Panama. Specimens examined. Brazil. Rondonia: 62 km SW Ariquemes, nr. Fzda. Rancho Grande, 20 – 31. x. 1997, 5 – 17. x. 1993, 4 – 16. xi. 1997, 15. iii. 1991, 6 – 15. xii. 1990 (FSCA). Colombia. Valle del Cauca: Lower Anchicaya, 400 m, 5. iii. 1976 (FSCA). Costa Rica. Alajuela: Angeles, R. B. Alberto Manuel Brenes, 900 – 1000 m (INBio); Dos Ríos, P. N. Guanacaste, 600 – 700 m (INBio); R. B. Ramon (USNM); Rio Frio, 16. vii. 1972 (FSCA). Cartago: Orosi (USNM); Orosi, P. N. Tapantí-Macizo de la Muerte, 1200 – 1300 m (INBio); Tayutic, 1100 – 1200 m (INBio); Turrialba (USNM), 4 – 13. viii. 1970 (BPBM). Guanacaste: Mayorga, P. N. Guanacaste, 900 – 1000 m (INBio); Santa Cecilia, P. N. Guanacaste, 600 – 700 m (INBio); Tierras Morenas, Z. P. Tenorio, 900 – 1000 m (INBio); 3 miles west Tilaran (USNM). Heredia: Horquetas, 0 – 100 m (INBio); Est. Biol. La Selva, various dates (INBio, USNM, FSCA, CMNH, BYUC, NCSU, BPBM, PMNH). Limón: Bratsi, R. I. Talamanca, 0 – 100 m (INBio); Colorado, 100 – 200 m (INBio); Hamburg Farm (USNM); Limón, vi. 1972 (FSCA); Santa Clara, Los Mercedes (USNM); Sixaola, R. V. S. Gandoca-Manzanillo, 0 – 100 m (INBio); Valle La Estrella, R. B. Hitoy Cerere, 100 – 200 m (INBio). Puntarenas: Golfito, P. N. Piedras Blancas, 0 – 100 m (INBio); Guacimal, 1000 – 1100 m (INBio); Jiménez, P. N. Cocovado, 0 – 100 m (INBio); La Palma (USNM); Monteverde, 1500 – 1600 m, various dates (INBio, USNM); Pittier, 1200 – 1300 m (INBio); Quepos, P. N. Manuel Antonio, 0 – 100 m (INBio); 2.5 miles southwest Rincon (USNM); Sabalito, 1400 – 1500 m (INBio); Sierpe, R. F. Golfo Dulce, 200 – 300 m (INBio); Tárcoles, P. N. Carara, 0 – 100 m (INBio). Ecuador. Esmeraldas: Cachabé, xii. 96 (BPBM); Fuyo, 200 m, 1 – 8. x. 1970 (BPBM). Guatemala. Chimaltenango: Tecpan (USNM). Nicaragua. Matagalpa Department: Jinotega (SEAN). Panamá. Bocas del Toro: 6 km north Punta Peña (USNM). Chiriquí: Reserva La Fortuna, Continental Divide Trail (USNM), 2.5 km north Santa Clara (USNM). Coclé: Cerro Caital (USNM). Colón: Achiote Road, 10 km south Gatun, 12. vi. 1976 (USNM); Madden Forest (USNM, FSCA); Pipeline Road NW Gamboa, 18. iv. 1976 (CASC); Porto Bello (USNM). Total: 589.	en	Staines, C. L. (2009): A review of the genus Chelobasis Gray, 1832 (Coleoptera: Chrysomelidae: Cassidinae). Zootaxa 2033 (1): 58-68, DOI: 10.11646/zootaxa.2033.1.6, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2033.1.6
