taxonID	type	description	language	source
03948799F957FFE1FF2F7EF5FCF5FA22.taxon	materials_examined	Material examined: Sultanate of Oman: Dhofar Region, Salalah, 16 ° 59 ’ N, 54 ° 64 ’ E, 10 m, 19 – 20. VII- 1995, B. Skule (1 ♀ paratype NHMB); Saudi Arabia: Wadi Marwani, 7 - II- 1985, W. Buttiker, (1 paratype, NHMB); Makkah, Wadi Shuque, 20 ° 44 ’ N, 41 ° 09 ’ E, 1380 m. 12 - IX- 1984 W. Buttiker, (1 paratype, NHMB); Muzahimiyah, AlKhararah 24 ° 24 ’ 24 ” N, 46 ° 14 ’ 40 ” E, 26 - IV- 2012 light trap, Y. Drayhim, H. Al Dhafer, A. Al Gharbawy & M. Al Harbi, 1 ♀ (KSMA); Wadi Qounonah, 19 ° 24 ’ 673 ” N, 41 ° 36 ’ 388 ” E, 348 m, 11 III 2012, light trap, H. Al Dhafer & H. Fadl, 2 ♂, 2 ♀ (KSMA); AlMajardah, Wadi AlTalalie, 19 ° 05 ’ 192 ” N, 41 ° 47 ’ 784 ” E, 286 m, 1 VI 2012, 1 ♂, 2 ♀ (KSMA), sweeping, H. Al Dhafer & B. Kondratieff; 1 ♀ (KSMA); same locality, A. AlAnsi, 1 ♀ (KSMA); Rawdhat Khorim, 25 ° 25 ’ 943 ” N, 47 ° 13 ’ 863 ” E, 572 m, 26 - IV- 2011 light trap, 1 ♀ (KSMA); Jizan, Aiban, Sabia Road, Wadi Shahdan, 17 ° 28 ’ 266 ” N, 42 ° 51 ’ 191 ” E, altitude 433 m, 13 - XI- 2012, light trap, H. Al Dhafer, 1 ♂ (KSMA).	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F957FFE1FF2F7EF5FCF5FA22.taxon	description	Description. Head: Black with dense small punctures mixed with larger punctures (Fig. 1), covered with dense erect pale setae; frons convex becoming flat toward anterior margin with well-developed apical carina; antennae not reaching the hind angles by about one antennomere, antennomere 2 subconical, slightly shorter than antennomere 3; antennomeres 3 – 10 feebly dilated at apex, longer than wide; antennomere 11 subellipsoidal, narrowed at apex. Pronotum: as long as wide, convex, widest at middle, black, dense and fine round punctures alternating with larger round punctures, lateral margins arcuate; hind angles sharply truncate, carinate dorsally. Prosternum with impressed V-shaped lines (Fig. 2). Scutellum: Black, slightly wider than long, heart shaped, emarginated at base. Elytra: Bicolored, yellowish brown, medially black (Fig. 3); striae finely punctuate, interstriae deeply punctuate. Male genitalia: Parameres of aedeagus with a distinct produced subapical rounded lateral lobe, apical expansion triangular, two setae below expansion, distal setae long; median lobe simple, clublike, subparallel margins beyond middle, apically rounded, not bulbous (Fig. 4). Female genitalia: Proximal sclerite of bursa copulatrix (Fig. 5) with 11 – 13 large marginal teeth and 14 – 16 smaller teeth including one below large marginal terminal tooth.	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F957FFE1FF2F7EF5FCF5FA22.taxon	discussion	Remarks. The proximal sclerite of the bursa copulatrix (Fig. 5) in D. brancuccii easily distinguishes it from the two new species described below. The male can be distinguished by the parameres possessing a distinctly produced subapical rounded lateral lobe with the apical expansion triangular in shape (Fig. 4). Additionally, the median lobe of the aedeagus has a clublike apex (Fig. 4).	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F954FFE4FF2F7B70FDA2FCA9.taxon	materials_examined	Material examined: Holotype: ♀ Saudi Arabia, Rawdhat Khorim, 25 ° 25 ’ 943 ” N, 47 ° 13 ’ 863 E, ” 572 m, 29 - IV- 2012, Al Dhafer (KSMA). Paratypes: (197): 4 ♂, 7 ♀ same data as holotype, 69 specimens same data as holotype, 19 V 2012, 2 ♂, 4 ♀ and 30 other specimens (KSMA); 15 - IV- 2012, 4 ♂, 3 ♀ and 5 other specimens (KSMA); 27 - V- 2012, 3 ♀ and 2 other specimens (KSMA); 28 - IV- 2012, 1 ♂ and 11 other specimens (KSMA); 10 - VI- 2012, 3 specimens (KSMA); 14 - IV- 2012, light trap, 1 specimen (KSMA); Wasiel, 45 km W Riyadh, Abdurahman AlThonian Farm, 4 - V- 2008, light trap, M. Otybi, 1 ♀ (KSMA); Wasiel, 45 km W Riyadh, Mazin AlQwrashi farm, 13 - V- 2008, light trap, M. Otybi, 1 ♀ (KSMA); Al Muzahimiyah, AlKhararah, 24 ° 24 ’ 24 ” N, 46 ° 14 ’ 40 ” E, 26 - IV- 2011, light trap, Y. Drayhim, H. Al Dhafer, A. Al Gharbawy & M. Al Harbi. 1 ♂ (KSMA); AlMuzahimiyah, AlKhararah, 24 ° 24 ’ 24 ” N, 46 ° 14 ’ 40 ” E, 17 - V- 2011, light trap, Y. Drayhim, H. Al Dhafer & A. AlGharbawy, 1 ♂ (KSMA); same locality, H. Setyaningrum, 1 ♂, 1 ♀ (KSMA); Hotat Bani Tamim, Riyadh, 23 ° 49 ’ 35 ” N, 46 ° 81 ’ 58 ”, 16 - V- 2010, sweeping, A. AlOthman, 1 ♂ (KSMA); AlMajardah, Wadi Baqrah, 18 ° 55 ’ 642 ” N, 42 ° 02 ’ 829 ” E, 433 m, 10 - XI- 2012, hand collecting, H. Al Dhafer, 1 ♀ (KSMA).	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F954FFE4FF2F7B70FDA2FCA9.taxon	description	Description: Head: Dark brown, fine punctures with scattered larger round punctures, covered with decumbent pale hair; frons convex becoming flat toward anterior margin with well-developed apical carina; antennae just reaching the hind angles in males, not reaching the hind angles by about one antennomere in females; antennomere 2 subconical, slightly shorter than antennomere 3; antennomeres 3 – 10 feebly dilated at apex, longer than wide; antennomere 11 subellipsoidal, narrowed at apex. Pronotum: As long as wide, convex, widest at middle, dark brown to black, densely and finely round punctures with scattered larger round punctures, lateral margins arcuate; hind angles sharply truncate, carinate dorsally. Scutellum: Dark brown medially, more blackish along suture, slightly wider than long, heart- shaped, emarginate at base. Elytra: Yellowish-brown, medially shaded with black (Fig. 6), covered with dense erect yellow setae, striae densely and finely punctate with scattered large rounded punctures, interstriae deeply punctate. Male genitalia: Parameres without a distinctly produced subapical rounded lobe, apicolateral expansion reduced, teardrop-like, two setae below expansion, distal setae long; median lobe simple, subparallel margins beyond middle apically narrowly rounded (Fig. 7). Female genitalia: Proximal sclerite of the bursa copulatrix with 9 – 11 large marginal teeth, terminal teeth largest, and 11 – 13 smaller internal discal teeth including one just below large marginal terminal tooth (Fig. 8). Etymology. The patronym honors Latifa A. Almutlaq.	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F954FFE4FF2F7B70FDA2FCA9.taxon	diagnosis	Diagnosis. Dicronychus latifae is most similar to D. brancuccii, but can be distinguished in the female by fewer teeth of the proximal sclerite of the bursa copulatrix (Fig. 8). The parameres of the male genitalia lack the distinctly produced subapical rounded lateral lobe with the triangular expansions (Fig. 7), as present in D. brancuccii (Fig. 4). Additionally the apex of the median lobe is more narrowly rounded than either D. brancuccii (Fig. 4) or D. bushrae n. sp. (Fig. 10).	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F952FFE5FF2F7D85FBF7FDFD.taxon	materials_examined	Material examined: Holotype: ♀ Saudi Arabia, Jizan Aiban, Sabia Road, Wadi Qusai, 17 ° 15 ’ 248 ” N, 42 ° 57 ’ 960 ” E, 257 m, 12 - XI- 2012, light trap, H. Al Dhafer (KSUM). Paratypes: 4 ♀ same data as a holotype, 1 ♀ same data as a holotype, beating, H. Fadl (KSMA); 2 ♂ Jizan, Aiban, Sabia Road, Wadi Shahdan, 17 ° 28 ’ 266 ” N, 42 ° 51 ’ 191 ” E, 433 m, 13 - XI- 2012, light trap, H. Al Dhafer (KSMA); 1 ♀ Jizan Aiban Sabia Road, Wadi Dafa, 17 ° 22 ’ 495 ” N, 43 ° 04 ’ 529 ” E, 870 m, 12 - XI- 2012, hand collecting, H. Al Dhafer (KSMA); 6 ♂, 6 ♀, Jizan Abu Arish Al Redhah road W. Dhamad, 17 ° 12 ’ 324 ” N, 43 ° 01 ’ 580 ”, 258 m, 11 - XI- 2012, light trap, H. Al Dhafer (KSMA); 1 ♀ Wadi Qounonah, 19 ° 24 ’ 673 ” N, 41 ° 36 ’ 388 ”, 348 m, 11 - III- 2012, light trap, H. Al Dhafer & H. Fadl (KSMA); 1 ♂ same data, Abdeldayem and A. El Torkey (KSMA); 1 ♂ AlMajardah, Wadi Baqrah, 18 ° 55 ’ 642 ” N, 42 ° 02 ’ 829 ” E, 433 m, 10 - XI- 2012, hand collecting, H. Al Dhafer (KSMA); 1 ♂ Wadi Targ, 19 ° 37 ’ 365 ” N, 42 ° 18 ’ 020 ” E, 1317 m, 14 - III- 2012, light trap, H. Al Dhafer and H. Fadl (KSMA); 1 ♀, Baha Thee Ain 19 ° 55 ’ 774 ” N, 41 ° 28 ’ 574 ” E, 10 - III- 2012, H. Setyaningrum (KSMA); 1 ♂ Baha, Wadi Bawa, 20 ° 44 ’ 976 ” N, 41 ° 14 ’ 846 ” E, 8 - XI- 2012, beating, H. Fadl (KSMA).	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F952FFE5FF2F7D85FBF7FDFD.taxon	description	Description. Head: black, fine punctures with scattered larger round punctures, covered with decumbent pale hair; frons convex becoming flat toward anterior margin with well-developed apical carina; antennae just reaching the hind angles in males, not reaching the hind angles by about one antennomere in females, antennomere 2 subconical, as long as antennomere 3; antennomeres 3 – 10 dilated at apex, longer than wide; antennomere 11 subellipsoidal, narrowed at apex. Pronotum: Slightly wider than long, convex, widest at middle, dark brown to black, dense and fine round punctures with scattered larger round punctures, lateral margins subparallel; hind angles sharply truncate, carinate dorsally. Scutellum: Brown to dark brown, slightly wider than long, heart shaped, emarginate at base. Elytra: Yellowish brown, medially black (Fig. 9) or completely black, covered with dense yellow setae; striae densely and finely punctate, interstriae deeply punctate. Male genitalia: Parameres without a distinctly produced subapical rounded lobe, apicolateral expansion reduced, teardrop-like, two setae below expansion, distal setae long, medial lobe simple, subparallel margins beyond middle apically rounded (Fig. 10). Female genitalia: Proximal sclerite of bursa copulatrix with 8 large marginal teeth, terminal teeth almost bifid in appearance, and 10 smaller internal discal teeth (Fig. 11).	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F952FFE5FF2F7D85FBF7FDFD.taxon	etymology	Etymology. The patronym honors Bushra H. Al Dhafer.	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F952FFE5FF2F7D85FBF7FDFD.taxon	diagnosis	Diagnosis. Dicronychus bushrae is most similar to both D. brancuccii and D. latifae, but can be distinguished in the female by the sclerite of the bursa copulatrix with eight large teeth (Fig. 11). Additionally, in the male, the median lobe of the aedeagus of D. bushrae (Fig. 10) is not as narrowly rounded as D. latifae (Fig. 7).	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F952FFE5FF2F7D85FBF7FDFD.taxon	discussion	Remarks on other species. Platia and Schimmel (1997) indicated in their key to the species of Dicronychus from the Arabian Peninsula, that D. talhouki has “ claw distinctly dilated in their first half, ” but in the description of the species (page 314), the claws are described as “ Feebly dilated in basal half. ” Examination of the holotype of this species confirms that the claws are feebly dilated in the basal half (Fig. 12). Additionally, Platia and Schimmel (1997) illustrated the sclerite of the bursa copulatrix with at least 8 large marginal relatively slender teeth, However, the sclerite of the bursa copulatrix of the Holotype female has 6 – 7 large thick marginal teeth with the distal teeth either bifid or trifid, a condition not illustrated by Platia and Schimmel (1997, fig. 70). All material listed below fits well within the concept of the Holotype of D. talhouki. An undetermined number of paratypes from Saudi Arabia and Yemen were not examined (Platia and Schimmel, 1997 indicate 20 males and 6 females but the number listed is 20 and “ several spms. ”). A redescription is provided below.	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F953FFE6FF2F7979FE3BFC45.taxon	materials_examined	Material examined. Saudi Arabia, Wadi Dhiyan, 19 ° 48 ’ N, 41 ° 36 ’ E, 830 m, 7 - III- 1984, W. Buttiker, Holotype ♀ (NHMB); Riyadh Province, Rawdhat Khorim, 100 km. E. Riyadh, 25 ° 25 ’ 943 ” N, 47 ° 13 ’ 863 ” E, 572 m, 29 - IV- 2011, light trap, 18 - II- 2012; same locality, pit fall trap below Acacia gerrardii, 6 - III- 2012, 5 ♂, 6 ♀, Al Dhafer (KSMA); Jizan, Aiban, Sabia Road, Wadi Shahdan, 17 ° 28 ’ 266 ” N, 42 ° 51 ’ 191 ” E, 433 m, 13 - XI- 2012, 1 ♂, light trap, H. Al Dhafer (KSMA); Jizan, Aiban, Wadi Gowra, 17 ° 17 ’ 569 ” N, 43 ° 04 ’ 211 ” E, 451 m, 11 - XI- 2012, 1 ♀, beating sheet, H. Fadl (KSMA); Jizan, Abu Arish, Al Redhah road, Wadi Dhamad, 17 ° 12 ’ 324 ” N, 43 ° 01 ’ 580 ” E, 258 m, 11 - XI- 2012, 1 ♀, light trap, H. Al Dhafer (KSMA); Al Oiynah Thonayan farm, 3 - V- 2008, 1 ♂, light trap, M. Otybi (KSMA); Al Oiynah, 50 km W. Riyadh, Obikan farm, 2 - VII- 2007, 1 ♂, light trap, H. Al Ayedh and H. Al Dhafer (KSMA).	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
03948799F953FFE6FF2F7979FE3BFC45.taxon	description	Description. Head: Arcuate, black, dense rugosely punctured, covered with decumbent pale hair; frons convex with well-developed apical carina interrupted medially; antennae not reaching the hind angles by about one antennomere, antennomere 2 subconical, slightly shorter than antennomere 3, antennomeres 3 – 10 feebly dilated at apex, longer than wide, antennomere 11 subellipsoidal, narrowed at apex. Pronotum: As long as wide or slightly longer, convex, widest at middle, brown, with dense and fine round punctures with scattered larger round punctures, lateral margins arcuate; hind angles sharply truncate, carinate dorsally. Prosternum without V-shaped lines (Fig. 13). Scutellum: Brown to dark-brown, as long as wide, heart shaped, emarginate at base Elytra: Bicolored, black anteriorly, posteriorly brown (Fig. 14); striae finely punctuate, interstriae deeply punctuate. Male genitalia: Parameres of aedeagus with a distinct expanded subapical rounded lobe, apicolateral expansion triangular, two setae below expansion, distal setae long; medial lobe simple, subparallel margins beyond middle, apically bulbous (Fig. 15). Female genitalia: Proximal sclerite of bursa copulatrix with 21 – 27 large marginal sclerotized teeth, apical tooth bifid or trifid (Fig. 16).	en	Al Dhafer, Hathal M., Platia, Giuseppe (2013): Contribution to the knowledge of the genus Dicronychus (Coleoptera: Elateridae) from the Kingdom of Saudi Arabia. Zootaxa 3734 (1): 15-22, DOI: 10.11646/zootaxa.3734.1.2
