taxonID	type	description	language	source
039487A8FFCD443EFE59381F8C7AFC76.taxon	description	Species included Dasyhelea dellapei Díaz and Spinelli sp. nov. Dasyhelea fueguina Díaz and Spinelli sp. nov. Dasyhelea gargola Díaz and Spinelli sp. nov. Dasyhelea ona Díaz and Spinelli sp. nov. Dasyhelea patagonica Ingram and Macfie Dasyhelea reynoldsi Ingram and Macfie Dasyhelea shannoni Ingram and Macfie Taxonomic discussion The patagonica group is herein proposed on the basis of the combination of characters listed in the diagnosis, all of them poor for consideration as good synapomorphies. If our knowledge of the immatures of the genus is improved then we will hopefully be able to find new, valuable characters to properly propose a phylogenetic classification. Remm (1971) described the subgenus Sebessia which includes the cincta group, with species in all the mayor biogeographic zones of the World, and the holosericea group, represented by Palaearctic species. The characteristic features of the subgenus are the presence of two well-developed spermathecae, the symmetrical parameres and the ventral surface of the aedeagus covered by a hyaline envelope; the last character being a very good synapomorphy (A. Borkent, pers. comm.; P. Dominiak, pers. comm.). As the Patagonian species lack the aedeagal hyaline envelope, we prefer not to include them within Sebessia.	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
039487A8FFCB443DFEFD3F758B08F990.taxon	description	(Figures 1, 2) Diagnosis The only species of the patagonica group in which the posteromedian margin of the sternite 9 is broadly rounded, not excavated or with distal process or projections. The female subgenital plate is subtriangular, the posterolateral arms are slender, and their inner margins are concave with pointed, mesally directed tips. Both sexes with the largest number (approximately 50) of scutellar setae. Male. Similar to female with usual sexual differences. Flagellum as in Figure 1 A. Palpus (Figure 1 B) with third segment bearing scattered sensilla; PR 4.94 (4.50 – 5.13, n = 4). Scutellum with 28 – 30 long setae and 22 – 28 shorter ones. Wing (Figure 1 C) length 1.91 mm (1.82 – 1.98 mm, n = 4), width 0.54 mm (0.48 – 0.58 mm, n = 4), CR 0.52 (0.51 – 0.54, n = 4). Genitalia (Figure 1 D): tergite 9 tapering distally, extending to level of apex of gonocoxite, apicolateral process slender, with apical seta; cercus with two or three setae; sternite 9 (Figure 1 E) 0.45 times longer than greatest width, posteromedian margin broadly rounded (folded in the available specimens). Gonocoxite stout, 1.8 times longer than greatest width; gonostylus as long as gonocoxite, narrow base, nearly straight, two basal setae, blunt apex. Parameres (Figure 1 F) with basal arms slender, curved; posteromedian projection triangular, extending shortly beyond basal arch of aedeagus, tip pointed. Aedeagus well sclerotized with rounded tip; 0.65 times longer than greatest width, basal arch extending to 0.55 of total length; basal arms stout, slender, directed anterolaterad; posterolateral arms short, recurved laterad. Female. Head dark brown. Eyes contiguous by width of one ommatidia. Antenna with flagellum (Figure 1 G) dark brown; AR 0.96 (0.91 – 1.09, n = 6). Clypeus (Figure 1 H) with 12 – 17 setae. Palpus (Figure 1 I) dark brown; third segment with four sub-basal capitate sensillae; PR 4.49 (4.17 – 4.86, n = 6). Thorax. Scutum dark brown, scutellum with 25 – 30 long setae and 22 – 26 shorter ones. Legs dark brown, including tarsi; hind tibial comb with four spines; prothoracic TR 1.95 (1.70 – 2.15, n = 6), mesothoracic TR 1.95 (1.78 – 2.12, n = 6), metathoracic TR 1.71 (1.57 – 1.89, n = 6). Wing (Figure 1 J), length 1.84 mm (1.76 – 1.92 mm, n = 5), width 0.72 mm (0.68 – 0.76 mm, n = 5), CR 0.53 (0.52 – 0.56, n = 5); membrane slightly infuscated, densely covered with macrotrichiae; second radial cell open; cubital fork at same level of anterior portion of second radial cell. Halter dark brown. Abdomen. Dark brown. Subgenital plate (Figure 1 K) subtriangular; posterolateral arms slender, inner margin concave with pointed, mesally directed tip. Two ovoid spermathecae (Figure 1 L), subequal, with straight necks, measuring 0.056 by 0.054 mm and 0.052 by 0.046 mm, necks 0.006 mm. Distribution Argentina (Santa Cruz). Type material Holotype male, Argentina, Santa Cruz prov., Lago del Desierto, 9 December 1996, G. Spinelli, red. Paratypes 3 males, 6 females, as follows: same data as holotype 1 male, 5 females (1 male, 4 female in MLP; 1 female in NHM); Parque Nacional Perito Moreno, Lago Burmeister, 24 November 1999, G. Spinelli - P. Marino, 2 males, 1 female, red (1 male, 1 female in MLP, 1 male in NHM). Etymology This species is named after Dr Pablo M. Dellapé, entomologist of the Museo de La Plata. Discussion This very dark brown species is the largest within the patagonica group, and can be easily distinguished from the other species by the numerous scutellar setae. The male genitalia are similar to those of D. patagonica, but in the latter species the aedeagus is distinctly broader and its distal portion is not rounded.	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
039487A8FFC74430FEF93BD38A13FD18.taxon	description	(Figures 2, 3) Diagnosis The only species of the patagonica group with the posteromedian projection of parameres bearing a stout, distally directed process with broad, bifid tip, and the spermathecae with hyaline necks. Male. Similar to female with usual sexual differences. Flagellum as in Figure 3 A. Palpus (Figure 3 B) with third segment bearing scattered sensilla; PR 4.06 (3.64 – 4.28, n = 3). Scutellum with 19 – 22 long setae and eight shorter ones. Wing (Figure 3 C) length 1.50 mm (1.42 – 1.54 mm, n = 3), width 0.43 mm (0.42 – 0.44 mm, n = 3), anal lobe narrow, CR 0.53 (0.52 – 0.54, n = 3). Genitalia (Figure 3 D): tergite 9 tapering distally, extending nearly to level of apex of gonocoxite, apicolateral process slender, with apical seta; cercus with two or three setae; sternite 9 (Figure 3 E) 0.45 times longer than greatest width, posteromedian margin folded, slightly excavated. Gonocoxite stout, 1.8 times longer than greatest width, with elongate, blunt anteromedian process; gonostylus slender, as long as gonocoxite, narrow base, nearly straight, tip pointed. Parameres (Figure 3 F) with basal arms stout, the fusion to posteromedian projection slightly sclerotized; posteromedian projection short, well sclerotized anteriorly, bearing a stout, distally directed process with broad, bifid tip. Aedeagus stout, well sclerotized, 0.7 times longer than greatest width, basal arch extending to 0.5 of total length, with rounded tip; basal arms short, recurved; posterolateral arms short, divergent, tips recurved mesad. Female. Head dark brown. Eyes contiguous by width of one or two ommatidia. Antenna with flagellum (Figure 3 G) dark brown; AR 1.05 (0.95 – 1.13, n = 5). Clypeus (Figure 3 H) with 8 – 10 setae. Palpus (Figure 3 I) dark brown; third segment with three or four sub-basal capitate sensillae; PR 3.70 (3.07 – 4.07, n = 5). Thorax. Scutum dark brown, scutellum with 16 – 21 long setae and 8 – 12 shorter ones. Legs dark brown, including tarsi; hind tibial comb with four spines; prothoracic TR 2.14 (1.66 – 2.71, n = 5), mesothoracic TR 2.01 (1.71 – 2.28, n = 5), metathoracic TR 1.87 (1.71 – 2.00, n = 5). Wing (Figure 3 J), length 1.35 mm (1.10 – 1.52 mm, n = 5), width 0.48 mm (0.44 – 0.52 mm, n = 5), CR 0.53 (0.52 – 0.55, n = 5); membrane slightly infuscated, densely covered with macrotrichiae; second radial cell open; cubital fork at same level of distal portion of first radial cell. Halter dark brown. Abdomen. Dark brown. Subgenital plate (Figure 3 K) subtriangular, anterior margin somewhat rounded, irregular; posterolateral arms slender, inner margin concave with pointed, mesally directed tip. Two ovoid spermathecae (Figure 3 L), unequal, with straight, hyaline necks, measuring 0.070 by 0.046 mm and 0.058 by 0.042 mm, necks 0.006 mm. Distribution Argentina, only known from its type locality. Type material Holotype male, Argentina, Tierra del Fuego, Parque Nacional Tierra del Fuego, río Ovando y ruta nacional no. 3, 54 ◦ 50 ′ 45.3 ′′ S, 68 ◦ 34 ′ 47.7 ′′ W, 1 December 2008, G. Spinelli, red. Paratypes 2 males, 5 females, same data (1 male, 4 females in MLP; 1 male, 1 female in NHM). Etymology The specific epithet refers to the province of Tierra del Fuego, where the type series was collected. Discussion The aedeagus of D. fueguina is very similar to the one of D. dellapei, especially by its rounded tip. However, the species can be easily distinguished by the parameres bearing a stout process with broad, bifid tip.	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
039487A8FFC54436FEFF385D8BB1FAB2.taxon	description	(Figures 2, 4) Diagnosis The only species of the patagonica group with the posteromedian margin of the male sternite 9 with a deep rounded excavation, and the posterolateral margins of the excavation broad and truncate. The female subgenital plate is rectangular, with irregular anterior margin. Male. Similar to female with usual sexual differences. Flagellum as in Figure 4 A. Palpus (Figure 4 B) with third segment bearing scattered sensilla; PR 4.23, fifth segment missing. Scutellum with 18 long setae and 18 shorter ones. Wing (Figure 4 C) length 1.74 mm, width 0.50 mm, CR 0.51. Genitalia (Figure 4 D): tergite 9 tapering distally, extending to level of apex of gonocoxite, apicolateral process slender, with apical seta; cercus with three or four setae; sternite 9 (Figure 4 E) 0.5 times longer than greatest width, posteromedian margin with deep, rounded anteromesal excavation, posterolateral margins of excavation broad, truncate. Gonocoxite stout, 1.9 times longer than greatest width, with short, blunt anteromedian process; gonostylus slender, as long as gonocoxite, narrow base, nearly straight, tip pointed. Parameres (Figure 4 F) with basal arms stout, with conspicuous laterodistal, pointed processes; anteromesad projection short, blunt, fused to posteromedian projection, the latter slender, slightly produced beyond distal portion of aedeagus body, with lightly recurved tip. Aedeagus well sclerotized, 0.75 times longer than greatest width, basal arch extending to 0.25 of total length; basal arms stout, short, recurved; body stout with triangular pointed, divergent posterolateral projections; posterolateral arms convergent, tips recurved mesad. Female. Head dark brown. Eyes contiguous by width of two to three ommatidia. Antenna with flagellum (Figure 4 G) dark brown; AR 1.13 (1.09 – 1.17, n = 2). Clypeus (Figure 4 H) with 14 – 17 setae. Palpus (Figure 4 I) dark brown; third segment with four sub-basal capitate sensillae; PR 3.82 (3.71 – 3.93, n = 2). Thorax. Scutum dark brown, scutellum with 19 – 20 long setae and 15 – 16 shorter ones. Legs dark brown, including tarsi; hind tibial comb with five spines; prothoracic TR 1.88, mesothoracic TR 1.90, metathoracic TR 1.87 (1.84 – 1.90, n = 2). Wing (Figure 4 J), length 1.46 mm (1.44 – 1.48 mm, n = 2), width 0.58 mm (0.56 – 0.60 mm, n = 2), CR 0.52 (n = 2); membrane slightly infuscated, densely covered with macrotrichiae; second radial cell open; cubital fork at same level of anterior portion of second radial cell. Halter dark brown. Abdomen. Dark brown. Subgenital plate (Figure 4 K) rectangular, anterior margin irregular; posterolateral arms stout, posteromedian projection stout, inner margin straight distally. Two ovoid spermathecae (Figure 4 L), subequal, with straight necks, measuring 0.048 by 0.042 mm and 0.044 by 0.038 mm, necks 0.004 mm. Distribution Argentina, only known from its type locality. Type material Holotype male, Argentina, Rio Negro, Parque Nacional Nahuel Huapi, cerro Catedral, laguna Toncek, 41 ◦ 11 ′ 52.2 ′′ S, 71 ◦ 29 ′ 31.2 ′′ W, 1770 m, 20 February 2007, C. Cazorla – M. Donato, red. Paratypes, 2 females, same data (MLP). Etymology The specimens herein described as a new species were collected in the laguna Toncek located in the cerro Catedral. The specific epithet refers to the gargoyles that are common sculptures of Cathedrals. Discussion The male genitalia of this species are similar to those of D. ona, especially by the aedeagus stout body and the triangular pointed, divergent posterolateral projections. However, it can be distinguished from D. ona by several other characteristics that may be found in the discussion under the description of the later species.	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
039487A8FFC34435FE043ECE8B96FD3E.taxon	description	(Figures 2, 5) Diagnosis The only species of the patagonica group with posteromedian margin of the sternite 9 slightly concave and bearing a crescent-shaped sclerite followed by two reniform, stout processes directed caudad, the parameres lacking posteromedial protection. The female subgenital plate has two slender processes arising from laterodistal margins of gonopore, and the spermathecae are distinctly pyriform. Male. Similar to female with usual sexual differences. Flagellum as in Figure 5 A. Palpus (Figure 5 B) with third segment bearing scattered sensilla; PR 4.18 (3.57 – 4.83, n = 8). Scutellum with 16 – 20 long setae and 6 – 14 shorter ones. Wing (Figure 5 C) length 1.45 mm (1.40 – 1.52 mm, n = 7), width 0.44 mm (0.42 – 0.48 mm, n = 7), CR 0.49 (0.47 – 0.51, n = 7). Genitalia (Figure 5 D): tergite 9 tapering distally, extending to level of apex of gonocoxite, apicolateral process slender, with apical seta; cercus with two or three setae; sternite 9 (Figure 5 E) 0.6 times longer than greatest width, posteromedian margin slightly concave, with crescent-shaped sclerite followed by two reniform, stout processes directed caudad. Gonocoxite stout, 2.2 times longer than greatest width, with short, blunt anteromedian process; gonostylus slender, 0.9 times longer than gonocoxite, narrow base, nearly straight, tip pointed. Parameres (Figure 5 F) without posteromedian projection; basal arms slender, fused distally, with blunt tip. Aedeagus well sclerotized, 0.55 times longer than greatest width, basal arch extending to 0.3 of total length; basal arms stout, elongated, recurved; posterolateral arms triangular, distal margin slightly concave, with posteriorly directed, recurved filament with pointed tip. Female. Head dark brown. Eyes contiguous by width of two ommatidia. Antenna with flagellum (Figure 5 G) pale brown; AR 0.94 (0.88 – 1.05, n = 9). Clypeus (Figure 5 H) with eight or nine setae. Palpus (Figure 5 I) pale brown; third segment with two subbasal capitate sensillae; PR 3.66 (3.00 – 4.25, n = 9). Thorax. Scutum dark brown, scutellum with 14 – 20 long setae and 10 – 22 shorter ones. Legs dark brown, tarsi slightly paler; hind tibial comb with five spines; prothoracic TR 1.86 (1.61 – 2.06, n = 9), mesothoracic TR 1.86 (1.57 – 2.25, n = 8), metathoracic TR 1.99 (1.75 – 2.11, n = 9). Wing (Figure 5 J), length 1.28 mm (1.12 – 1.42 mm, n = 9), width 0.52 mm (0.46 – 0.60 mm, n = 9), CR 0.49 (0.47 – 0.54, n = 9); membrane slightly infuscated, densely covered with macrotrichiae; second radial cell open; cubital fork at same level of distal portion of first radial cell. Halter pale brown. Abdomen. Dark brown. Subgenital plate (Figure 5 K) rectangular with rounded anteromesal margin, posterolateral arms stout; two short, slender processes arising from laterodistal margin of gonopore. Two pyriform spermathecae (Figure 5 L), subequal, with straight, stout necks, measuring 0.056 by 0.042 mm and 0.052 by 0.038 mm, necks 0.006 mm. Distribution Argentina (Tierra del Fuego). Type material Holotype male, Argentina, Tierra del Fuego prov., Parque Nacional Tierra del Fuego, laguna Negra (turbera), 54 ◦ 50 ′ 42.1 ′′ S, 68 ◦ 35 ′ 17.9 ′′ W, 15 m, 4 – 7 December 2008, M. Donato - G. Spinelli, Malaise trap. Paratypes 7 males, 9 females, as follows: same data as holotype 6 males, 5 females (5 males, 4 females in MLP; 1 male, 1 female in NHM); same data except 1 female, red. Tierra del Fuego prov., ruta complementaria a (turbera), 54 ◦ 21 ′ 42.3 ′′ S, 66 ◦ 39 ′ 42.4 ′′ W, 40 m, 3 December 2008, G. Spinelli, 3 females, red (MLP); Lago Yehuin, 54 ◦ 21 ′ 39.6 ′′ S, 67 ◦ 46 ′ 44.4 ′′ W, 45 m, 5 December 2008, G. Spinelli, 1 male, red (MLP). Etymology The species is named after the Ona indians, early inhabitants of Tierra del Fuego island. Discussion The male genitalia of this new species are similar to D. gargola. However, the latter species lacks the posteromedian crescent-shaped sclerite and reniform processes of the sternite 9, and the parameres show a distinct slender posteromedian projection that is produced beyond the aedeagus body. The female is clearly distinguished from other species by the pyriform spermatheca and by the presence in the subgenital plate of two slender processes arising from laterodistal margins of gonopore.	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
039487A8FFC04429FE22386089CFFE38.taxon	description	(Figures 6, 7)	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
039487A8FFC04429FE22386089CFFE38.taxon	diagnosis	Diagnosis The only species of the patagonica group with at most 28 scutellar setae and the tarsi paler than femora. The posteromedian margin of the male sternite 9 has a deep rounded excavation and the posterolateral margins of the excavation are pointed. Male. Similar to female with usual sexual differences. Flagellum as in Figure 6 A. Palpus (Figure 6 B) with third segment narrowed at midportion, with scattered sensilla; PR 4.18 (3.84 – 4.54, n = 3). Scutellum with 14 – 18 long setae and 10 shorter ones. Wing (Figure 6 C) length 1.25 mm (1.18 – 1.34 mm, n = 3), width 0.37 mm (0.36 – 0.40 mm, n = 3), CR 0.47 (0.45 – 0.50, n = 3). Genitalia (Figure 6 D): tergite 9 rounded distally, extending to level of apex of gonocoxite, apicolateral process slender, with apical seta; cercus with three or four setae; sternite 9 (Figure 6 E) 0.45 times longer than greatest width; posteromedian margin with deep rounded excavation, posterolateral margins of excavation pointed. Gonocoxite stout, 2.0 times longer than greatest width, with conspicuous anteromedian process, directed anteromesad; gonostylus 0.9 times longer than gonocoxite, narrow base, slender, nearly straight, tip pointed. Parameres (Figure 6 F) with basal arms slender; posteromedian projection slender, reaching distal portion of aedeagus, tip blunt. Aedeagus strongly sclerotized, 0.4 times longer than greatest width, basal arch extending to 0.4 of total length; basal arms slender, directed laterad; posterolateral arms with recurved, ventrally directed tips. Female. Head dark brown. Eyes contiguous by width of one or two ommatidia. Antenna with flagellum (Figure 6 G) dark brown, AR 0.96 (0.88 – 1.00, n = 8). Clypeus (Figure 6 H) with seven to nine setae. Palpus (Figure 6 I) dark brown; third segment with four sub-basal capitate sensillae; PR 3.79 (3.23 – 4.16, n = 10). Thorax. Scutum dark brown, scutellum with 13 – 16 long setae and six to eight shorter ones. Legs dark brown, tarsi paler; hind tibial comb with five spines; prothoracic TR 1.82 (1.73 – 2.00, n = 10), mesothoracic TR 1.90 (1.76 – 2.06, n = 10), metathoracic TR 1.91 (1.75 – 2.23, n = 10). Wing (Figure 6 J), length 1.12 mm (1.00 – 1.42 mm, n = 10), width 0.47 mm (0.42 – 0.58 mm, n = 10), CR 0.49 (0.48 – 0.52, n = 10); membrane slightly infuscated, densely covered with macrotrichiae; cubital fork at same level of middle length of second radial cell. Halter dark brown. Abdomen. Dark brown. Subgenital plate (Figure 6 K) rectangular, anterior margin rounded mesally; posterolateral arms straight, with pointed, mesally directed tip. Two ovoid spermathecae (Figure 6 L), subequal, with straight necks, measuring 0.050 by 0.044 mm and 0.046 by 0.034 mm, necks 0.040 mm. Distribution Argentina (Neuquen, Río Negro), Chile (Llanquihue, Osorno, Valdivia). Type material Dasyhelea patagonica. Holotype male, Argentina, Río Negro prov., Lago Gutiérrez; 3 – 14 November 1926, F. and M. Edwards; allotype female, same data except Bariloche, 1 December 1926, F. and M. Edwards (NHM, examined).	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
039487A8FFC04429FE22386089CFFE38.taxon	description	Dasyhelea chilensis. Holotype female, Chile, Llanquihue prov., Casa Pangue, 4 – 10 December 1926, F. and M. Edwards (NHM, examined). Other specimens examined Argentina: Neuquen prov., lago Curruhe, 20 November 1994, G. Spinelli, 1 male, 2 females (MLP). Río Negro prov., Parque Nacional “ Nahuel Huapi ”, lago Villarino, 5 December 1992, G. Spinelli, 2 males, 1 female (MLP); Parque Nacional Nahuel Huapi, Manso Medio, La Cantera, 41 ◦ 21 ′ 16 ′′ S, 71 ◦ 42 ′ 27.3 ′′ W, 764 m, 15 January to 7 February 2007, Garré- Montes de Oca, 2 females, Malaise trap (MLP); same date except 11 – 30 December 2007, Garré- Montes de Oca, 2 females, Malaise trap (MLP). Chile: Valdivia prov., Mehuin, 27 December 1992, G. Spinelli, 1 female, red. Osorno prov., camino a Pucatrihue, 29 November 1992, G. Spinelli, 2 females, red (MLP). Discussion This species, restricted to northern Patagonia, is the smallest of the patagonica group and is the one with fewer scutellar setae (at most 28). The female subgenital plate is similar to that of D. gargola, another north patagonian species. Characters to distinguish the male genitalia from those of D. dellapei may be found in the key and in the discussion under the description of the later species. The synonymy of D. chilensis under D. patagonica is based on the examination of the types of both species. They only differ in the tarsi colouration, very slightly darker in the female holotype of D. chilensis, something in itself that is insufficient to recognize another species.	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
039487A8FFDC442FFE3D39698B11FA50.taxon	description	(Figures 7, 8)	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
039487A8FFDC442FFE3D39698B11FA50.taxon	diagnosis	Diagnosis The only species of the patagonica group with the sternite 9 of male bearing a posterior trefoil-shaped projection and the gonostylus with a double curve. Male. Similar to female with usual sexual differences. Flagellum as in Figure 8 A brown. Palpus (Figure 8 B) brown; with third segment bearing scattered sensilla; PR 5.23 (4.53 – 6.08, n = 10). Scutellum with 24 – 34 long setae and 12 – 16 shorter ones. Wing (Figure 8 C), length 1.81 mm (1.72 – 1.94 mm, n = 9), width 0.52 mm (0.50 – 0.56 mm, n = 9), CR 0.50 (0.49 – 0.51, n = 9). Genitalia (Figure 8 D): tergite 9 not reaching to level of apex of the gonocoxite, tip broad, truncate, apicolateral process slender, with apical seta; cercus with four or five short setae; sternite 9 (Figure 8 E) 0.65 times longer than greatest width, posteromedian margin narrow, slightly concave, with posterior projection trefoil-shaped slightly produced beyond aedeagus tip. Gonocoxite stout, 2.2 times longer than greatest width, with conspicuous anteromedian process directed anteromesad; gonostylus as long as gonocoxite, slender, with a double curve, tip pointed. Parameres (Figure 8 F) with basal arms slender, curved; posteromedian projection triangular with blunt tip, reaching midlength of aedeagus. Aedeagus strongly sclerotized, 0.5 times longer than greatest width, basal arch extending to 0.4 of total length; basal arms slender, curved directed laterad; posterolateral arms stout recurved ventrad. Female. Head dark brown. Eyes contiguous by width of two ommatidia. Antenna with flagellum (Figure 8 G) very dark brown, AR 1.01 (1.00 – 1.08, n = 10). Clypeus (Figure 8 H) with seven or eight setae. Palpus (Figure 8 I) dark brown; third segment with two sub-basal capitate sensillae; PR 4.21 (3.33 – 4.57, n = 10). Thorax. Scutum very dark brown, scutellum with 20 – 26 long setae and 14 – 22 shorter ones. Legs very dark brown, except tarsis pale brown; hind tibial comb with six spines; prothoracic TR 1.80 (1.59 – 2.00, n = 10), mesothoracic TR 1.90 (1.76 – 2.27, n = 10), metathoracic TR 1.79 (1.71 – 1.85, n = 10). Wing (Figure 8 J), length 1.68 mm (1.52 – 1.78 mm, n = 10), width 0.67 mm (0.64 – 0.74 mm, n = 10), CR 0.53 (0.51 – 0.55, n = 10); membrane slightly infuscated, densely covered with macrotrichiae; cubital fork at same level of anterior portion of second radial cell. Halter dark brown. Abdomen. Dark brown. Subgenital plate (Figure 8 K), subtriangular; posterolateral arms stout, inner margin with bulbous protuberance. Two ovoid spermathecae (Figure 8 L), subequal, with straight necks, measuring 0.056 by 0.038 mm and 0.054 by 0.038 mm, necks 0.004 mm. Distribution Argentina (Santa Cruz, Tierra del Fuego). Type material Holotype male, allotype female, Argentina, Tierra del Fuego prov., Río Grande, estancia Viamonte, 1930, P. W. Reynolds (NHM, examined). Other specimens examined Argentina, Santa Cruz prov., Lago del Desierto, 9 December 1996, G. Spinelli, 3 males (MLP). Tierra del Fuego prov., 40 km west of Río Grande, 3 December 2005, G. Spinelli, 1 male, red (MLP); Parque Nacional Tierra del Fuego, Río Ovando y ruta nacional no. 3, 54 ◦ 50 ′ 45.3 ′′ S, 68 ◦ 34 ′ 47.7 ′′ W, 1 December 2008, G. Spinelli, 1 male, red (MLP); laguna Negra (turbera), 54 ◦ 50 ′ 2.1 ′′ S, 68 ◦ 35 ′ 17.9 ′′ W, 15 m, 4 – 7 December 2008, M. Donato - G. Spinelli, 4 males, Malaise trap (MLP); Lago Yehuin, 54 ◦ 21 ′ 39.6 ′′ S, 67 ◦ 46 ′ 44.4 ′′ W, 45 m, 5 December 2008, G. Spinelli, 1 female, red (MLP); río San Pablo y ruta complementaria a, 54 ◦ 17 ′ 59.5 ′′ S, 66 ◦ 42 ′ 42.5 ′′ W, 12 m, 3 December 2008, G. Spinelli, 1 male, red (MLP); ruta complementaria a (turbera), 54 ◦ 21 ′ 42.3 ′′ S, 66 ◦ 39 ′ 42.4 ′′ W, 40 m, 3 December 2008, G. Spinelli, 8 females, red (MLP). Discussion This species, restricted to southern Patagonia, is very similar to D. s hannoni. However, the latter species is smaller, the sternite 9 of the male is arrow-shaped, the posterolateral arms of the aedeagus are convergent and the female subgenital plate is higher. The type-locality of D. reynoldsi is located in the Argentinean side of the island of Tierra del Fuego, not in Chile as stated by Wirth (1974) and followed by Borkent and Wirth (1997) and Borkent and Spinelli (2000, 2007). On the other hand, Spinelli (1998) properly recorded the species from Argentina.	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
039487A8FFDA4422FE3D3EBC8B13FADC.taxon	description	(Figures 7, 9)	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
039487A8FFDA4422FE3D3EBC8B13FADC.taxon	diagnosis	Diagnosis The only species of the patagonica group with arrow-shaped posteromedian of male sternite 9, and female subgenital plate triangular and very high. Male. Similar to female with usual sexual differences. Flagellum as in Figure 9 A. Palpus (Figure 9 B) with third segment bearing scattered sensilla; PR 4.36 (4.00 – 5.00, n = 10). Scutellum with 14 – 19 long setae and 10 – 14 shorter ones. Wing (Figure 9 C); length 1.26 mm (1.20 – 1.32 mm, n = 7), width 0.39 mm (0.36 – 0.42 mm, n = 7), CR 0.49 (0.48 – 0.50, n = 7). Genitalia (Figure 9 D): tergite 9 rounded distally, not reaching level of apex of gonocoxite, apicolateral process slender, with apical seta; cercus with four or five setae; sternite 9 (Figure 9 E) 0.5 times longer than greatest width, with posteromedian margin arrow-shaped, reaching aedeagus apex. Gonocoxite stout, 1.5 times longer than greatest width; gonostylus 0.8 longer than gonocoxite, width base, nearly straight, tapering to pointed apex. Parameres (Figure 9 F) with basal arms slightly curved; posteromedian projection triangular, short, reaching aedeagus midlength. Aedeagus very sclerotized, 0.6 times longer than greatest width, basal arch extending to 0.3 of total length; basal arms elongate, directed laterad; posterolateral arms convergent, slender, apex recurved directed ventrad. Female. Head dark brown. Eyes contiguous by width of one or two ommatidia. Antenna with flagellum (Figure 9 G) pale; AR antennal 1.01 (0.84 – 1.07, n = 10). Clypeus (Figure 9 H) with 9 – 13 setae. Palpus (Figure 9 I) brown; third segment with three sub-basal capitate sensillae; PR 3.91 (3.50 – 4.45, n = 10). Thorax. Scutum dark brown, scutellum with 16 – 19 long setae and 10 – 18 shorter ones. Legs dark brown; hind tibial comb with five spines; prothoracic TR 1.85 (1.72 – 2.00, n = 10), mesothoracic TR 1.88 (1.71 – 2.22, n = 10), metathoracic TR 1.76 (1.44 – 1.95, n = 10). Wing (Figure 9 J), length 1.23 mm (1.10 – 1.38 mm, n = 10), width 0.50 mm (0.44 – 0.58 mm, n = 10), CR 0.50 (0.49 – 0.52, n = 10); membrane slightly infuscated, densely covered with macrotrichiae; cubital fork at same level of middle length of second radial cell. Halter dark brown. Abdomen. Dark brown. Subgenital plate (Figure 9 K) stout, triangular, extending to anterior margin of eighth segment; posterolateral arms short, slender. Two pyriform spermathecae (Figure 9 L), subequal, with slender, straight necks, measuring 0.042 by 0.032 mm and 0.036 by 0.030 mm, necks 0.004 mm. Distribution Argentina (Neuquen, Santa Cruz); Chile (Arauco, Cautin, Chiloé, Coihaique, Llanquihue, Malleco, Valdivia). Type material Holotype male, Chile, Llanquihue prov., Ensenada, 14 / 15 December 1926, F. and M. Edwards; allotype female, Chile, Chiloe Islands, Ancud, 17 – 19 December 1926, F. and M. Edwards (NHM, examined). Other specimens examined Argentina: Neuquen prov., 20 km north of lago Aluminé, 13 February 1994, G. Spinelli, 1 male (MLP); Hua-Hum, 11 February 1989, G. Spinelli, 1 male, red (MLP); 6 – 7 km south of San Martín de los Andes, 900 m, 26 November 1984, G. Spinelli, 1 female (MLP); Parque Nacional Nahuel Huapi, río Pichitraful, 40 ◦ 42 ′ 46 ′′ S, 71 ◦ 47 ′ 23 ′′ W, 789 m, 4 – 23 February 2008, A. Garre – F. Montes de Oca, 1 male, Malaise trap (MLP). Río Negro prov., Parque Nacional Nahuel Huapi, río Manso superior, 41 ◦ 14 ′ 8.1 ′′ S, 71 ◦ 46 ′ 58.5 ′′ W, 845 m, 7 February to 2 March 2007, A. Garré – F. Montes de Oca, 3 males, 1 female, Malaise trap (MLP). Santa Cruz prov., lago del Desierto, 19 January 1995, I. Garda, 1 male, 1 female, red (MLP), same data except 9 December 1996, G. Spinelli, 1 male (MLP). Tierra del Fuego prov., Parque Nacional Tierra del Fuego, Río Ovando y ruta nacional no. 3, 54 ◦ 50 ′ 45.3 ′′ S, 68 ◦ 34 ′ 47.7 ′′ W, 1 December 2008, G. Spinelli, 4 males, red (MLP). Chile: Arauco prov., Pata de Gallina, 7 December 2003, G. Spinelli, 2 males, red. Malleco prov., Parque Nacional Nahuelbuta (JAD 1682 / 2), 26 December 1984, J. A. Downes, 1 male, red (CNCI). Cautin prov. 10 km E Villarrica, 27 December 1992, G. Spinelli, 1 male; 17 km southwest Villarrica, 6 December 2003, G. Spinelli, 1 male, red. Valdivia prov., Puringue Pobre, comarca Mariquita (JAD 1692 / 1), 8 January 1985, J. A. Downes, 3 males, 1 female, red (CNCI); Fundo San Martín (JAD 1669 / 1), 15 December 1984, J. A. Downes, 1 female, red (CNCI). Llanquihue Prov., Ensenada (JAD 1723 / 1), 8 February 1985, J. A. Downes, 1 females, red (CNCI); same date except 9 February 1985, 1 male; same data except Lago Verde, (JAD 1696 / 5), 12 January 1985, 2 males, 1 female (CNCI); Parque Nacional Vicente Pérez Rosales, río Petrohue (JAD 1695 / 1), 11 January 1985, J. A. Downes, 1 female, red (CNCI); río Peulla, Casa Pangue, 41 ◦ 02 ′ 55.0 ′′ S, 71 ◦ 52 ′ 31.2 ′′ W, 366 m, 18 February 2008, M. Donato, 10 males, Malaise trap; same data except 2 females, red. Chiloé prov., Huillinco (JAD 1690 / 3), 3 January 1985, J. A. Downes, 1 male, red (CNCI); same data except (JAD 1691 / 2), 4 January 1985, 1 female, (CNCI); same data except (JAD 1722 / 1), 8 February 1985, 2 males, (CNCI). Coihaique prov., Coihaique (JAD 1717 / 1), 3 February 1985, J. A. Downes, 1 male (CNCI). Discussion Characters to distinguish D. shannoni from D. reynoldsi may be found in the key and in the discussion under the description of the latter species.	en	Díaz, Florentina, Spinelli, Gustavo R., Ronderos, María M. (2010): A revision of the species allied to Dasyhelea patagonica Ingram and Macfie (Diptera: Ceratopogonidae). Journal of Natural History 44 (45 - 46): 2825-2849, DOI: 10.1080/00222933.2010.512424, URL: http://dx.doi.org/10.1080/00222933.2010.512424
