identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039587C6FFA7FFA00898FED74BF05DC0.text	039587C6FFA7FFA00898FED74BF05DC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maera Leach 1814	<div><p>Maera -clade</p><p>Diagnosis and included genera. See Ariyama (2018).</p></div>	https://treatment.plazi.org/id/039587C6FFA7FFA00898FED74BF05DC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ariyama, Hiroyuki;Kodama, Masafumi;Tomikawa, Ko	Ariyama, Hiroyuki, Kodama, Masafumi, Tomikawa, Ko (2020): Species of the Maera-clade collected from Japan. Part 4: addenda to genera Maera Leach, 1814 and Quadrimaera Krapp-Schickel & Ruffo, 2000, with revised keys to Japanese species of the clade (Crustacea: Amphipoda: Maeridae). Zootaxa 4885 (3): 336-352, DOI: 10.11646/zootaxa.4885.3.2
039587C6FFA7FFA00898FD9B4A7D5EB8.text	039587C6FFA7FFA00898FD9B4A7D5EB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maera Leach 1814	<div><p>Maera Leach, 1814</p><p>[Japanese name: Hon-sunnariyokoebi-zoku]</p><p>Synonym, diagnosis and included species. See Ariyama (2020).</p></div>	https://treatment.plazi.org/id/039587C6FFA7FFA00898FD9B4A7D5EB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ariyama, Hiroyuki;Kodama, Masafumi;Tomikawa, Ko	Ariyama, Hiroyuki, Kodama, Masafumi, Tomikawa, Ko (2020): Species of the Maera-clade collected from Japan. Part 4: addenda to genera Maera Leach, 1814 and Quadrimaera Krapp-Schickel & Ruffo, 2000, with revised keys to Japanese species of the clade (Crustacea: Amphipoda: Maeridae). Zootaxa 4885 (3): 336-352, DOI: 10.11646/zootaxa.4885.3.2
039587C6FFA7FFA00898FF674B3E5D0C.text	039587C6FFA7FFA00898FF674B3E5D0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maeridae Krapp-Schickel 2008	<div><p>Maeridae Krapp-Schickel, 2008</p><p>[Japanese name: Sunnariyokoebi-ka]</p></div>	https://treatment.plazi.org/id/039587C6FFA7FFA00898FF674B3E5D0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ariyama, Hiroyuki;Kodama, Masafumi;Tomikawa, Ko	Ariyama, Hiroyuki, Kodama, Masafumi, Tomikawa, Ko (2020): Species of the Maera-clade collected from Japan. Part 4: addenda to genera Maera Leach, 1814 and Quadrimaera Krapp-Schickel & Ruffo, 2000, with revised keys to Japanese species of the clade (Crustacea: Amphipoda: Maeridae). Zootaxa 4885 (3): 336-352, DOI: 10.11646/zootaxa.4885.3.2
039587C6FFA7FFAA0898FD4349EE5DBF.text	039587C6FFA7FFAA0898FD4349EE5DBF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maera denticoxa Ariyama & Kodama & Tomikawa 2020	<div><p>Maera denticoxa sp. nov.</p><p>[Japanese name: Kita-sunnariyokoebi, new]</p><p>(Figs 2–7)</p><p>Maera danae .— Tomikawa et al., 2010: table 1, figs 1–3. [not Maera danae (Stimpson, 1853)]</p><p>Type material. Holotype: male, 14.7 mm (OMNH-Ar-11724), Nagane, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.95972&amp;materialsCitation.latitude=39.3625" title="Search Plazi for locations around (long 141.95972/lat 39.3625)">Otsuchi Bay</a> in Iwate Prefecture, 39°21'45"N, 141°57'35"E (Fig. 1B), 3–4 m depth, under stones, by SCUBA, 18 February 2019, coll. M. Kodama. Paratypes: female, 15.6 mm (OMNH-Ar-11725), same locality, 7–8 m depth, under stones, by SCUBA, 11 July 2018, coll. M. Kodama; female, 15.5 mm (OMNH-Ar-11734, not dissected), same locality, 7–8 m depth, on coarse sand under stones, by SCUBA, 19 May 2020, coll. M. Kodama; ovigerous female, 13.2 mm (OMNH-Ar-11726), Nagoshi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.94722&amp;materialsCitation.latitude=39.35111" title="Search Plazi for locations around (long 141.94722/lat 39.35111)">Otsuchi Bay</a> in Iwate Prefecture, 39°21'04"N, 141°56'50"E (Fig. 1B), 4.3 m depth, by SCUBA and airlift, 14 November 2018, coll. N. Ohtsuchi ; male, 16.2 mm and female, 16.3 mm (OMNH-Ar-11727, 11728), Daikoku Island, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=144.87195&amp;materialsCitation.latitude=42.963055" title="Search Plazi for locations around (long 144.87195/lat 42.963055)">Akkeshi Town</a>, Hokkaido, 42°57'47"N, 144°52'19"E (Fig. 1A), intertidal, using hand net, fixed by 99% Ethanol, 24 June 2005, coll. K. Tomikawa.</p><p>Type locality. Nagane, Otsuchi Bay in Iwate Prefecture, Japan .</p><p>Etymology. From the Latin dentatus, -a, -um (= dentate) + coxa, referring to the shapes of coxae 1–6.</p><p>Diagnosis. Eyes present. Mandible, palp article 1 with acute distal tooth. Coxae 1–6, posteroventral margins each with 1–7 notches; coxa 1 anteroventral corner acutely produced. Gnathopod 2 in both sexes, palm strongly oblique, defined by small tooth. Pereopods dactyli simple; bases of pereopods 5–7 rectangular, posterodistal corners square. Pleonal e pimera 1–3 acutely projected posteroventrally. Uropod 3, outer ramus longer than inner ramus, rami about 3 times as long as peduncle. Telson longer than wide, distal ends pointed, lateral margins each with 7–8 robust setae.</p><p>Description. Based on holotype, male, 14.7 mm (OMNH-Ar-11724).</p><p>Head (Figs 2–3). Eyes oval, about 0.2 times length of head. Antenna 1 weakly setose; peduncular articles 1–3 with length ratio of 1: 1.1: 0.3, article 1 with 8 posteromedial robust setae; accessory flagellum with 7 articles; primary flagellum with 29 articles, terminal article minute. Antenna 2 about 0.55 times length of antenna 1, setose; peduncular articles 3–5 with length ratio of 1:3.15:2.3; flagellum with 10 articles, terminal article minute. Upper lip, ventral margin rounded, weakly setose. Mandible, incisor bearing 5 and 4 cusps in left and right, respectively, each lacinia mobilis 4-dentate, with 12–14 accessory setae; palp article 1 with acute distal tooth, articles 1 – 3 length ratio (excluding distal tooth of article 1) 1: 3.6–3.7: 2.2–2.4, articles 2 – 3 setose. Lower lip with inner lobes, distal margins of inner and outer lobes weakly setose, mandibular processes small. Maxilla 1, inner plate with 3 long and 3 short apical setae, medial margin bearing a few feeble setae, outer plate with 11 apical robust setae and medial feeble setae, palp article 1 with a few laterodistal and lateral setae, article 2 bearing many apical setae. Maxilla 2, outer plate larger than inner plate; both plates bearing many apical setae, medial margin of inner plate with 2 setae. Maxilliped, inner plate truncate, distal margin with many thick setae, distomedial corner with 4 robust setae ventrally; outer plate with 27 long-to-short robust setae on distomedial margin; palp 4-articulate, article 2 setose medioventrally, article 3 with many distolateral setae, article 4 with large apical robust seta.</p><p>Pereon (Fig. 4). Gnathopod 1, coxa anteroventral corner acutely produced, ventral margin with 5 small notches posteriorly; basis setose on anteroproximal and posterior margins; carpus swollen posterodistally, posterior margin and medial surface heavily setose; propodus about 80% as long as carpus, anterior and posterior margins setose, palm defined by 3 short robust setae; dactylus slender, slightly curved posteriorly. Gnathopod 2, coxa subquadrate, posteroventral margin with 7 small notches; basis narrow, anteroproximal and posterior margins and anterodistal corner with several setae; merus with acute posterodistal spine; carpus lobate posterodistally, posterior margin heavily setose; propodus slender, length about 1.9 times width, anteromedial surface and posterior margin setose; palm strongly oblique, defined by small triangular tooth with robust seta, palmar margin with 6 minute teeth, bearing 9 lateral and 12 medial robust setae; dactylus strongly curved, anterior margin with 17 setae.</p><p>Pereopods 3–4, shapes subequal, pereopod 3 about 1.2 times as long as pereopod 4; coxa trapezoidal, posteroventral margins with 4 and 2 small notches, respectively, gill large, shorter than basis length; basis rectilinear, anterior and posterior margins each with a few long proximal setae and many small robust setae; propodus lined with bundle of robust setae posteriorly; dactylus short, about 0.4 times length of propodus, simple. Pereopods 5–7, shapes subsimilar, about 1.1, 1.5, 1.6 times lengths of pereopod 4, respectively; coxae 5–6, posteroventral corner with single small notch, coxa 7 lacking notch, coxae 6–7 anteroventral margin bearing 3 robust setae; basis rectangular, lengths about 2.0, 1.95, 1.8 times widths respectively, posteroproximal corner roundly projected, anterior margin with many short robust setae, posterior margin serrate, posterodistal corner square; carpus and merus lined with bundle of robust setae; dactylus short, 0.3–0.4 times length of propodus, simple.</p><p>Pleon (Fig. 5). Epimera 1–3 acutely projected posteroventrally, each with lateral ridge, ventral margins of epimera 1–3 with 7, 7, 5 robust setae, respectively. Pleopods 1–3 slender, pleopods 2 – 3 shorter than pleopod 1; peduncles with 6 (4 lost), 8 (1 lost), 7 coupling hooks respectively; both rami each with 12–14 articles. Uropod 1, peduncle with large basofacial and distolateral robust setae and 17 dorsolateral and 17 dorsomedial robust setae; outer ramus subequal length of inner ramus, about 0.7 times as long as peduncle, with 4 dorsolateral, 6 dorsomedial and 4 terminal robust setae; inner ramus with 6 dorsolateral, 12 dorsomedial, 1 ventrolateral and 5 terminal robust setae. Uropod 2 short, about 0.55 times length of uropod 1; peduncle laterodistal corner triangularly projected, dorsolateral margin, dorsomedial margin and distomedial corner with 11, 6 and 3 robust setae, respectively; outer ramus about 0.9 times length of inner ramus, almost same length as peduncle, with 8 dorsolateral, 5 dorsomedial and 5 terminal robust setae; inner ramus with 5 dorsolateral, 14 dorsomedial, 1 ventrolateral and 5 terminal robust setae. Uropod 3 long, about 195% as long as uropod 2; peduncle, lateral and medial surfaces with 1 and 8 robust setae, respectively, laterodistal corner bearing 8 robust setae; both rami rectilinear, outer ramus about 1.1 times length of inner ramus, about 3.1 times length of peduncle, with vestigial second article; lateral and medial margins of both rami lined with many robust setae, distal margins bearing several robust setae and a few slender setae, longest seta about 0.2 times length of outer ramus. Telson about 1.15 times longer than wide, each lateral margin with 7 – 8 robust setae, tips acutely projected, dorsal surfaces of each lobe with a robust seta and 3 sensory setae.</p><p>Description of female (paratype, 15.6 mm, OMNH-Ar-11725). Gnathopod 2 (Fig. 5 F-G2) almost same as male, holotype, 14.7 mm (OMNH-Ar-11724) except for oostegite.</p><p>Variation. Notches on posteroventral margins of coxae 1 – 6 present in all specimens. Their numbers vary from 1 to 7 (Table 1).</p><p>Coloration in life (Fig. 6). Eyes dark brown, other parts pale pink.</p><p>Remarks. Maera denticoxa sp. nov. has small notches on the coxae 1 – 6. Only three species having such notches were recorded in the genus (Ariyama 2020): Maera loveni (Bruzelius, 1859) [Japanese specimens], M. pachytelson Karaman &amp; Ruffo, 1971, and M. sagamiensis Ariyama, 2020 . However, the notches are present on the coxa 2, the coxae 1–2 and the coxae 1–3 in these species, respectively. These three species also differ from the new species in the shape of the gnathopod 2 palm, the shape of the telson, and the number of the coupling hooks, respectively.</p><p>Habitat. On coarse sand bottom under stones (Fig. 7), intertidal to 8 m depth (present study).</p><p>Distribution. Coast of Daikoku Island in Hokkaido and Otsuchi Bay in Iwate Prefecture, Japan (present study).</p></div>	https://treatment.plazi.org/id/039587C6FFA7FFAA0898FD4349EE5DBF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ariyama, Hiroyuki;Kodama, Masafumi;Tomikawa, Ko	Ariyama, Hiroyuki, Kodama, Masafumi, Tomikawa, Ko (2020): Species of the Maera-clade collected from Japan. Part 4: addenda to genera Maera Leach, 1814 and Quadrimaera Krapp-Schickel & Ruffo, 2000, with revised keys to Japanese species of the clade (Crustacea: Amphipoda: Maeridae). Zootaxa 4885 (3): 336-352, DOI: 10.11646/zootaxa.4885.3.2
039587C6FFADFFAB0898F8874B7E5CE8.text	039587C6FFADFFAB0898F8874B7E5CE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quadrimaera Krapp-Schickel & Ruffo 2000	<div><p>Quadrimaera Krapp-Schickel &amp; Ruffo, 2000</p><p>[Japanese name: Kaku-sunnariyokoebi-zoku]</p><p>Synonym, diagnosis and included species. See Ariyama (2019). However, on the mandibular palp in the diagnosis, “article 1 lacking distal tooth” is modified to “article 1 usually lacking distal tooth” (see Remarks of Quadrimaera angulata sp. nov.). In addition, the following four species are added to the included species: Quadrimaera anhi Dang &amp; Le, 2011; Q. miramirandella Alves, Neves &amp; Johnsson, 2018; Q. vietnamica Dang &amp; Le, 2011; and Q. yemanjae Alves, Neves &amp; Johnsson, 2018 .</p></div>	https://treatment.plazi.org/id/039587C6FFADFFAB0898F8874B7E5CE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ariyama, Hiroyuki;Kodama, Masafumi;Tomikawa, Ko	Ariyama, Hiroyuki, Kodama, Masafumi, Tomikawa, Ko (2020): Species of the Maera-clade collected from Japan. Part 4: addenda to genera Maera Leach, 1814 and Quadrimaera Krapp-Schickel & Ruffo, 2000, with revised keys to Japanese species of the clade (Crustacea: Amphipoda: Maeridae). Zootaxa 4885 (3): 336-352, DOI: 10.11646/zootaxa.4885.3.2
039587C6FFACFFAC0898FEF24DC65EE0.text	039587C6FFACFFAC0898FEF24DC65EE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quadrimaera angulata Ariyama & Kodama & Tomikawa 2020	<div><p>Quadrimaera angulata sp. nov.</p><p>[Japanese name: Maru-sunnariyokoebi, new]</p><p>(Figs 8–11)</p><p>Type material. Holotype: female, 10.1 mm (OMNH-Ar-11729), north of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=131.03284&amp;materialsCitation.latitude=30.910833" title="Search Plazi for locations around (long 131.03284/lat 30.910833)">Tanegashima Island</a> in Kagoshima Prefecture, 30°54.07'N, 131°02.32'E – 30°54.65'N, 131°01.97'E (Fig. 1C), 140–152 m depth, using beam trawl, 29 May 2006, coll. K. Tomikawa. Paratypes: females, 9.2 and 6.6 mm (OMNH-Ar-11730, 11731), same data as holotype.</p><p>Type locality. North of Tanegashima Island in Kagoshima Prefecture, Japan .</p><p>Etymology. From the Latin angulatus, -a, -um (=angular, angulate), referring to the shape of the pereopod 7 basis.</p><p>Diagnosis. Mandible, palp article 1 with distal tooth. Gnathopod 1, coxa anteroventral corner pointed. Female gnathopod 2, anterior margin of basis without robust setae, anterodistal corners of basis and ischium lobate on lateral and medial surfaces, propodus subrectangular, palm slightly oblique and rounded, defined by small tooth, palmar margin with small rounded excavation in middle, posterior excavation absent. Pereopods 5–7, bases subovate, posterodistal corners roundly lobate in pereopods 5–6 and angular in pereopod 7, posterior margins not setose. Uropod 3, outer ramus longer than inner ramus, about 1.7 times length of peduncle. Telson longer than wide, tips of both lobes incised, each with 2 robust setae, longest robust seta about 0.8 times as long as telson.</p><p>Description. Based on holotype, female, 10.1 mm (OMNH-Ar-11729) and paratype, female, 9.2 mm (OMNHAr-11730) for lower lip.</p><p>Head (Figs 8–9). Anteroventral corner produced acutely, eyes subrounded, about 0.2 times as long as head. Antenna 1 slender, poorly setose; peduncular articles 1–3 with length ratio of 1: 1.15: 0.25, article 1 with 4 robust setae posteriorly; accessory flagellum long, with&gt;12 articles, tip lost; primary flagellum with&gt;28 articles, tip lost. Antenna 2 about 35% length of body, poorly setose; peduncular articles 3–5 with length ratio of 1: 2.9: 2.65; flagellum with 8 articles, terminal article minute. Upper lip, ventral margin rounded, with short setae. Mandible, incisor bearing 5 cusps and 8 accessory setae, left lacinia mobilis 4-dentate, right lacinia mobilis 5-dentate; palp article 1 with distal tooth, articles 1–3 length ratio (excluding distal tooth of article 1) 1: 2.05: 1.7 in left and 1: 2.25: 1.75 in right, articles 2 – 3 setose. Lower lip with inner lobes, distal margins weakly setose, distal parts of outer lobes each with bundle of setae and dense fine setae, mandibular processes small. Maxilla 1, inner plate bullet-shaped, apically with long plumose seta and several short setae, medial margin bare; distal margin of outer plate with 7 robust setae; palp article 2 with many apical setae. Maxilla 2, outer plate larger than inner plate; medial margin of inner plate with several distal setae and several feeble setae. Maxilliped, distomedial corner of inner plate with 3 distal and 2 ventral robust setae; outer plate with 19 long-to-short robust setae on distomedial margin; palp with 4 articles, article 4 with large apical robust seta.</p><p>Pereon (Fig. 10). Gnathopod 1, coxa anteroventral corner pointed; basis with many long setae on posterior margin and medial surface; carpus with shallow excavation on anterior margin; propodus about 85% length of carpus, palm defined by 5 short robust setae. Gnathopod 2, coxa subquadrate; basis relatively narrow, anterodistal corners on lateral and medial surfaces lobate, anterior margin with several minute setae, posterior margin bearing several long setae; ischium, anterodistal corners on lateral and medial surfaces lobate; merus with acute posterodistal spine; propodus subrectangular, slightly widened distally, length about 1.35 times width; palm slightly oblique, rounded, defined by small tooth with small robust seta, margin with small rounded excavation in middle, posterior excavation absent, anterior lobe with 5 lateral and 4 medial robust setae, posterior lobe bearing 8 lateral and 8 medial robust setae; dactylus, inner margin slightly curved.</p><p>Pereopods 3–4 subequal in shape, pereopod 3 about 1.1 times length of pereopod 4; coxa subrectangular; basis with posterior margin bearing several long setae. Pereopod 5 about 1.2 times length of pereopod 4; coxa with posterior lobe bearing robust seta; basis oblong, length about 2.0 times width, posterodistal lobe rounded, anterior margin with 6 short robust setae; merus swollen in middle. Pereopod 6 about 135% length of pereopod 5; coxa with 2 and 1 ventral robust setae on anterior and posterior lobes, respectively; basis subovate, length about 1.65 times width, posterodistal lobe rounded, anterior margin with 6 short robust setae, posterior margin minutely serrate. Pereopod 7 about 0.9 times length of pereopod 6; coxa subtrapezoidal, anteroventral corner with robust seta; basis subovate, length about 1.5 times width, posterodistal lobe angular, anterior margin with 6 short robust setae, posterior margin minutely serrate.</p><p>Pleon (Fig. 11). Epimera 1–3 slightly projected posteroventrally; epimera 1–2 each with oblique lateral ridge, ventral margins of epimera 1–3 with 2, 3, 5 robust setae, respectively. Pleopods 1–3, inner rami each with 10 articles and outer rami with 14, 13, 12 articles respectively; pleopod 3 shortest, peduncle bearing 3 robust setae distally. Uropod 1, peduncle with 1 basofacial, 11 dorsolateral, 10 dorsomedial and 1 distolateral robust setae; outer ramus about 85% length of inner ramus, about 75% length of peduncle, with 4 lateral, 6 medial and 5 (1 lost) terminal robust setae; inner ramus with 8 lateral, 9 medial and 5 terminal robust setae. Uropod 2 about 0.7 times length of uropod 1; peduncle bearing 5 dorsolateral and 3 dorsomedial robust setae; outer ramus about 85% length of inner ramus, about 115% length of peduncle, with 8 lateral, 3 medial and 5 (1 lost) terminal robust setae; inner ramus with 5 lateral, 8 medial and 5 terminal robust setae. Uropod 3 about 0.9 times length of uropod 2; distolateral and distomedial corners of peduncle with 2 and 3 robust setae, respectively; outer ramus about 1.1 times length of inner ramus, about 1.7 times length of peduncle; outer ramus with single and 3 groups of robust setae laterally and 5 robust setae medially, distal margin with minute second article and 10 setae; inner ramus with 1+1+2 robust setae laterally and 4 robust setae medially, distal margin with 7 setae. Telson longer than wide, each lateral margin with 2 sensory setae subdistally; distal margins of lobes each with 2 processes, 1 sensory seta and 2 long robust setae, longest robust seta about 0.8 times length of telson.</p><p>Coloration in life. Unknown.</p><p>Remarks. This new species has a distal tooth on the mandibular palp article 1, although Ariyama (2019) diagnosed the lack of the tooth in the genus Quadrimaera . As a result of reexamining all literature on the genus, the following five species are revealed to possess the distal tooth: Q. frater Myers, 2014 from the Chagos Archipelago; Q. kaiulani (Barnard, 1970) from Hawaii; Q. massavensis (Kossmann, 1880) from the Red Sea (Krapp-Schickel &amp; Ruffo 2006); Q. micheli Appadoo, Myers &amp; Fagoonee, 2002 sensu Hughes (2015) from Cocos Islands (another species different from the true Q. micheli); and Q. vietnamica Dang &amp; Le, 2011 from Vietnam. Quadrimaera angulata sp. nov. and these species resemble one another and share some characters (e.g. slightly oblique and rounded palm of the gnathopod 2 and incised lobes of the telson). However, this new species can be distinguished from the other five species by the shape of the pereopod 7 basis and the longer rami of the uropod 3 (Table 2).</p><p>Habitat. Bottom material unknown, 140–152 m depth. The examined specimens were collected together with Quadrimaera quadrimana (Dana, 1853) [two males, 5.4 and 4.8 mm (OMNH-Ar-11732, 11733)].</p><p>Distribution. North of Tanegashima Island in Kagoshima Prefecture, Japan (present study).</p></div>	https://treatment.plazi.org/id/039587C6FFACFFAC0898FEF24DC65EE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ariyama, Hiroyuki;Kodama, Masafumi;Tomikawa, Ko	Ariyama, Hiroyuki, Kodama, Masafumi, Tomikawa, Ko (2020): Species of the Maera-clade collected from Japan. Part 4: addenda to genera Maera Leach, 1814 and Quadrimaera Krapp-Schickel & Ruffo, 2000, with revised keys to Japanese species of the clade (Crustacea: Amphipoda: Maeridae). Zootaxa 4885 (3): 336-352, DOI: 10.11646/zootaxa.4885.3.2
039587C6FFABFFAC0898FCFA4C975B99.text	039587C6FFABFFAC0898FCFA4C975B99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maera Leach 1814	<div><p>Key to adult males of Japanese species of the Maera -clade (revised).</p><p>Quadrimaera angulata sp. nov. is not included because the male is unknown.</p><p>1. Gnathopod 2, anterior margin of dactylus bearing many setae.................................................. 2</p><p>- Gnathopod 2, anterior margin of dactylus with single seta..................................................... 4</p><p>2. Body very large (maximum BL:&gt; 50 mm), gnathopod 2 palm defined by tooth bearing strong robust seta............................................................................................ Maera loveni (Bruzelius, 1859)</p><p>- Body small (maximum BL: about 10–16mm), gnathopod 2 palm defined by tooth bearing small robust seta.............. 3</p><p>3. Small notches present on coxae 1–6.................................................... Maera denticoxa sp. nov.</p><p>- Small notches present on coxae 1–3............................................ Maera sagamiensis Ariyama, 2020</p><p>4. Pereopods 3–7 dactyli bifid............................................................................. 5</p><p>- Pereopods 3–7 dactyli simple............................................................................ 8</p><p>5. Gnathopod 2 palm with 2 excavations in middle.............................. Quadrimaera quqdrimana (Dana, 1853)</p><p>- Gnathopod 2 palm with single excavation in middle.......................................................... 6</p><p>6. Excavation on gnathopod 2 palm quadrate................................... Quadrimaera gotoensis Ariyama, 2019</p><p>- Excavation on gnathopod 2 palm rounded.................................................................. 7</p><p>7. Pereopod 6, posterior margin of basis heavily setose............................ Quadrimaera setibasis Ariyama, 2019</p><p>- Pereopod 6, posterior margin of basis not setose........................... Quadrimaera pacifica (Schellenberg, 1938)</p><p>8. Uropod 3, tips of both rami pointed......................................... Austromaera ariakensis Ariyama, 2019</p><p>- Uropod 3, tips of both rami truncate...................................................................... 9</p><p>9. Gnathopod 2 basis wide; uropod 3 rami long, about 3 times length of peduncle........ Meximaera mooreana (Myers, 1989)</p><p>- Gnathopod 2 basis ordinary; uropod 3 rami relatively short, 1 – 2 times length of peduncle........................... 10</p><p>10. Gnathopod 2 palm transverse, without robust setae; mandibular palp article 1 with distal tooth....................... 11</p><p>- Gnathopod 2 palm slightly or moderately oblique, with many robust setae; mandibular palp article 1 lacking distal tooth.. 12</p><p>11. Uropod 3 longer than uropod 2, basis of pereopod 5 subovate.................... Maeropsis okinawensis Ariyama, 2018</p><p>- Uropod 3 shorter than uropod 2, basis of pereopod 5 subrectangular.............. Maeropsis serratipalma (Nagata, 1965)</p><p>12. Palm of gnathopod 2 moderately oblique (angle of palmar corner: 115–130°)..................................... 13</p><p>- Palm of gnathopod 2 slightly oblique (angle of palmar corner: 90–105°)......................................... 14</p><p>13. Coxa 1 anteroventral corner produced acutely, distal robust setae of telson short (ca. 20% of telson length).................................................................................... Orientomaera obliquua Ariyama, 2018</p><p>- Coxa 1 anteroventral corner rounded, distal robust setae of telson long (ca. 45% of telson length)......................................................................................... Orientomaera rotundicoxa Ariyama, 2018</p><p>14. Palm of gnathopod 2 with two distinct teeth.................................... Orientomaera incisa Ariyama, 2020</p><p>- Palm of gnathopod 2 with several blunt teeth.............................................................. 15</p><p>15. Posterior margin of pereopod 6 basis almost straight, body color bluish pale pink................................................................................................... Orientomaera brevispina (Kim &amp; Kim, 1991)</p><p>- Posterior margin of pereopod 6 basis convex, body color light crimson............ Orientomaera decipiens Ariyama, 2018</p></div>	https://treatment.plazi.org/id/039587C6FFABFFAC0898FCFA4C975B99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ariyama, Hiroyuki;Kodama, Masafumi;Tomikawa, Ko	Ariyama, Hiroyuki, Kodama, Masafumi, Tomikawa, Ko (2020): Species of the Maera-clade collected from Japan. Part 4: addenda to genera Maera Leach, 1814 and Quadrimaera Krapp-Schickel & Ruffo, 2000, with revised keys to Japanese species of the clade (Crustacea: Amphipoda: Maeridae). Zootaxa 4885 (3): 336-352, DOI: 10.11646/zootaxa.4885.3.2
039587C6FFAAFFAD0898FF2F4C9758F5.text	039587C6FFAAFFAD0898FF2F4C9758F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maera Leach 1814	<div><p>Key to adult females of Japanese species of the Maera -clade.</p><p>Maera sagamiensis Ariyama, 2020 and Quadrimaera gotoensis Ariyama, 2019 are not included owing to the lack of female information.</p><p>1. Gnathopod 2, anterior margin of dactylus bearing many setae.................................................. 2</p><p>- Gnathopod 2, anterior margin of dactylus with single seta..................................................... 3</p><p>2. Body very large (maximum BL:&gt; 50 mm), gnathopod 2 palm defined by tooth bearing strong robust seta............................................................................................ Maera loveni (Bruzelius, 1859)</p><p>- Body small (maximum BL: about 16mm), gnathopod 2 palm defined by tooth bearing small robust seta.................................................................................................. Maera denticoxa sp. nov.</p><p>3. Pereopods 3–7 dactyli bifid............................................................................. 4</p><p>- Pereopods 3–7 dactyli simple............................................................................ 7</p><p>4. Gnathopod 2 palm rounded, telson lobes incised..................................... Quadrimaera angulata sp. nov.</p><p>- Gnathopod 2 palm almost straight, telson lobes truncate....................................................... 5</p><p>5. Pereopod 5 basis subrectangular, posterodistal corner scarcely lobate............. Quadrimaera quqdrimana (Dana, 1853)</p><p>- Pereopod 5 basis ovate, posterodistal corner roundly lobate.................................................... 6</p><p>6. Pereopod 6, posterior margin of basis setose.................................. Quadrimaera setibasis Ariyama, 2019</p><p>- Pereopod 6, posterior margin of basis not setose........................... Quadrimaera pacifica (Schellenberg, 1938)</p><p>7. Uropod 3, tips of both rami pointed......................................... Austromaera ariakensis Ariyama, 2019</p><p>- Uropod 3, tips of both rami truncate...................................................................... 8</p><p>8. Uropod 3 rami long, about 3 times length of peduncle............................ Meximaera mooreana (Myers, 1989)</p><p>- Uropod 3 rami relatively short, 1 – 2 times length of peduncle................................................... 9</p><p>9. Gnathopod 2 palm transverse, without robust setae; mandibular palp article 1 with distal tooth....................... 10</p><p>- Gnathopod 2 palm slightly or moderately oblique, with many robust setae; mandibular palp article 1 lacking distal tooth.. 11</p><p>10. Uropod 3 longer than uropod 2, basis of pereopod 5 subovate.................... Maeropsis okinawensis Ariyama, 2018</p><p>- Uropod 3 shorter than uropod 2, basis of pereopod 5 subrectangular.............. Maeropsis serratipalma (Nagata, 1965)</p><p>11. Palm of gnathopod 2 moderately oblique (angle of palmar corner: 115–130°)..................................... 12</p><p>- Palm of gnathopod 2 slightly oblique (angle of palmar corner: 90–100°)......................................... 13</p><p>12. Coxa 1 anteroventral corner produced acutely, distal robust setae of telson short (ca. 20% of telson length).................................................................................... Orientomaera obliquua Ariyama, 2018</p><p>- Coxa 1 anteroventral corner rounded, distal robust setae of telson long (ca. 45% of telson length)......................................................................................... Orientomaera rotundicoxa Ariyama, 2018</p><p>13. Palm of gnathopod 2 with two distinct teeth.................................... Orientomaera incisa Ariyama, 2020</p><p>- Palm of gnathopod 2 with several blunt teeth.............................................................. 14</p><p>14. Posterior margin of pereopod 6 basis almost straight, body color bluish pale pink................................................................................................... Orientomaera brevispina (Kim &amp; Kim, 1991)</p><p>- Posterior margin of pereopod 6 basis convex, body color light crimson............ Orientomaera decipiens Ariyama, 2018</p></div>	https://treatment.plazi.org/id/039587C6FFAAFFAD0898FF2F4C9758F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ariyama, Hiroyuki;Kodama, Masafumi;Tomikawa, Ko	Ariyama, Hiroyuki, Kodama, Masafumi, Tomikawa, Ko (2020): Species of the Maera-clade collected from Japan. Part 4: addenda to genera Maera Leach, 1814 and Quadrimaera Krapp-Schickel & Ruffo, 2000, with revised keys to Japanese species of the clade (Crustacea: Amphipoda: Maeridae). Zootaxa 4885 (3): 336-352, DOI: 10.11646/zootaxa.4885.3.2
