taxonID	type	description	language	source
039287BCFF9AFFF3FF66F8B8079C75C3.taxon	materials_examined	Type species. Cancer bernhardus Linnaeus, 1758.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9AFFF3FF66F8B8079C75C3.taxon	diagnosis	Diagnosis. Eleven pairs of biserial gills. Rostrum variable. Crista dentata well developed, with 1 or more accessory teeth. Sternite of third maxilliped armed or unarmed. Chelipeds generally very unequal, right usually larger. Dactyls of ambulatory legs commonly with spiniform bristles on ventral margins. Fourth pereopods usually semichelate; propodal rasp with 1 to several rows of corneous scales. Males usually without, rarely with slight papilla protruded from gonopore on one or both coxae of fifth pereopod; usually with unpaired pleopods 2 – 5 or 3 – 5, rarely without unpaired pleopods. Females usually with paired, rarely with single left gonopore on coxa of third pereopod; without paired first pleopod, usually with unpaired pleopods 2 – 5, rarely 2 – 4. Pleon usually spirally twisted, occasionally straight. Uropods asymmetrical, occasionally symmetrical. Telson with terminal margins rounded, oblique or straight, usually with median cleft.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9AFFF3FF66F8B8079C75C3.taxon	distribution	Distribution. Worldwide.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9BFFF0FF66FA1804E272B3.taxon	description	Pagurus annulipes. — Sánchez & Campos, 1978: 41, fig. 13.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9BFFF0FF66FA1804E272B3.taxon	diagnosis	Diagnosis. Shield longer than broad. Rostrum short, triangular or rounded, unarmed; lateral projections with or without small marginal or submarginal spine. Ocular peduncles long, slender, with corneae slightly dilated. Ocular acicles broad, terminal margins with 3 – 6 acute spines. Antennular peduncles long, as long as or slightly longer than ocular peduncles. Antennal peduncles equaling or slightly exceeding length of ocular peduncles; setae short, less than 1 article in length. Right cheliped larger than left; palm with single or double row of spines or spinulose tubercles on dorsomesial margin, dorsal surface slightly convex, with short setae and irregular rows of strong acute spines (females and small males), or irregular rows of small spines or tubercles (large males). Dorsomesial and dorsolateral margins of carpus with row of strong spines. Left cheliped reaching to base of dactyl of right cheliped, often longest in females; dactyl usually exceeding length of palm, dorsal surface with row of moderately strong spines, all surfaces with scattered long setae, particularly dense along cutting edge; palm one-third to one-half length of carpus, with irregular double row of strong spines extending onto fixed finger as single row; carpus equaling or slightly exceeding length of merus, dorsolateral margin with row of moderately strong spines, distal margin with 1 or 2 prominent acute spines; merus subtriangular. Second and third pereopods with dactyls shorter than propodi, terminating in strong corneous claws; ventral margins with row of strong corneous spines increasing in size distally; dorsal surfaces of propodi each with row of tufts of long setae, ventral surfaces each with row of tufts of long setae and few corneous spines or spinules distally; distal margins of carpi each with moderately strong spine. Telson somewhat asymmetrical with posterior lobes subtriangular, separated by median cleft, terminal margins armed with several strong spines and few small spines.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9BFFF0FF66FA1804E272B3.taxon	distribution	Distribution. Western Atlantic — Bermuda, Florida, Gulf of Mexico, Antilles, Central America, northern South America and Brazil (Fernando de Noronha and from Pernambuco to Santa Catarina). Habitat. Shallow waters to 50 metres, on sandy and rocky substrates, and algae banks.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9BFFF0FF66FA1804E272B3.taxon	materials_examined	Material examined. Brazil: Pernambuco — Fernando de Noronha, Ponta da Sapata, 1 male: 1.8 mm, 1 female: 1.7 mm (MZUSP – 9440). Bahia — Alcobaça, Parcel das Paredes, 2 males: 1.4, 1.5 mm (MZUSP – 13911). Rio de Janeiro — Angra dos Reis, 5 males: 2.9, 2.3, 2.1, 2.9, 2.1 mm, 5 females: 2.5, 2.6, 2.5, 2.8, 2.7 mm (MZUSP – 4600). São Paulo — São Sebastião, Saco Grande, 1 male: 2.3 mm, 1 female: 2.1 mm (MZUSP – 13006); Guarujá, Praia de Pernambuco, 1 female: 1.8 mm (MZUSP – 7291); 1 male: 2.3 mm (MZUSP – 7290).	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9BFFF0FF66FA1804E272B3.taxon	discussion	Remarks. Pagurus brevidactylus is very close to P. provenzanoi, but the two species can be quickly distinguished. Pagurus provenzanoi has short dense setae on the dorsal face of the chelipeds, whereas P. b re v i - dactylus has one longitudinal row of strong spines next to the dorsolateral margin of the left cheliped, and very short setae on the antennal flagellum.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF98FFF1FF66F9EF04377238.taxon	diagnosis	Diagnosis. Shield slightly longer than broad. Rostrum obtuse, slightly overreaching lateral projections. Ocular peduncles slender and shorter than shield width, with corneae slightly dilated. Ocular acicles with anterior margins rounded, with 1 strong submarginal spine; occasionally accessory marginal spinule on mesial margin. Antennular peduncles overreaching corneae; antennal peduncles usually not reaching distal margins of corneae; flagella long, usually overreaching right cheliped. Right cheliped with palm approximately twice longer than broad; dorsal face with 2 rows of spinose tubercles; carpus with rows of spines usually well developed on dorsomesial and dorsolateral margins. Left cheliped smaller than right, dorsal surface of carpi with 2 longitudinal rows of 4 or 5 spinose teeth; dorsal face of palm with 2 rows of spinose teeth. Surface of chelipeds covered by long setae which sometimes can hide the ornamentation. First pair of ambulatory legs with dactyls slightly longer than propodi; ventral margins each with row of corneous spines. Propodi unarmed. Telson with left lobe longer than right, both with 4 strong spines and intermediate spinules.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF98FFF1FF66F9EF04377238.taxon	distribution	Distribution. Western Atlantic — Gulf of Mexico, Antilles, northern South America, Brazil (St. Peter and St. Paul Rocks and from Pernambuco to Rio Grande do Sul), Uruguay and Argentina. Habitat. From intertidal to 60 metres, on sandy and muddy substrates.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF98FFF1FF66F9EF04377238.taxon	materials_examined	Material examined. Brazil: St. Peter and St. Paul Rocks, 4 males: 2.5, 1.2, 1.1, 1.6 mm, 2 females: 1.9, 2.0 mm (MZUSP – 9609). Espírito Santo – Vitória, Ilha Galheta, 1 male: 2.8 mm (MZUSP – 12911). Rio de Janeiro — Proj. Ilha Grande, st. 45, 1 male: 1.5 mm (MZUSP – 13376); st. 304, 1 female: 1.6 mm (MZUSP – 13872); st. 212 - F, 1 male: 2.0 mm (MZUSP – 13362); st. 235, 1 male: 1.8 mm (MZUSP – 13377); Manguinhos, 3 females: 2.2, 1.5, 1.4 mm, 1 male: 2.1 mm (MZUSP – 4712). São Paulo – Ilha de São Sebastião, Praia Garapocaia, 4 males: 2.5, 2.4, 2.2, 1.8 mm, 1 female: 2.8 mm (MZUSP – 13352); Praia do Veloso, 9 females: 2.8, 2.7, 2.8, 3.0, 2.0, 2.5, 2.2, 2.0, 1.7 mm, 4 males: 3.6, 1.7, 2.1, 1.4 mm (MZUSP – 13358); São Sebastião, Praia Araçá, 7 males: 3.2, 2.7, 2.8, 2.4, 2.7, 2.5, 2.6 mm, 1 female: 2.5 mm (MZUSP – 12990); Saco Grande, 1 male: 2.4 mm (MZUSP – 12993); Guarujá, Praia Branca, 1 male: 2.5 mm (MZUSP – 13357); Santos, 1 male: 1.8 mm (MZUSP – 13356); Proj. REVIZEE, st. 6658 (near the border of Paraná), 1 male: 1.4 mm (MZUSP – 13909).	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF98FFF1FF66F9EF04377238.taxon	discussion	Remarks. According to Forest & Saint Laurent (1968), Pagurus criniticornis shows remarkable individual variations, which can cause doubts in identification. Perhaps one of the most important variations is in the length of the antennal peduncles. Forest & Saint Laurent (1968) observed that specimens from high latitudes (Uruguay and Argentina) had the antennal peduncles distinctly longer than the corneae, whereas in specimens from northern regions, the antennal peduncles reached only the anterior margins of the corneae. We examined specimens from Ilha Grande, Rio de Janeiro, in which the antennal peduncles overreached the corneae. Thus if there exist two populations of Pagurus criniticornis, one with antennal peduncles overreaching the corneae (in the south) and another with antennal peduncles shorter than the corneae (in the north), the distributional limits of these populations are unclear.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF99FFF4FF66FA7002B97663.taxon	diagnosis	Diagnosis. Shield slightly broader than long. Rostrum triangular, with rounded tip. Ocular peduncles with median constriction; corneae dilated. Ocular acicles large, with rounded tips and small submarginal spine; dorsal face remarkably concave. Antennular and antennal peduncles overreaching corneae, approximately 2 / 3 as long as ultimate segment. Right cheliped elongate; palm with several conical or flattened tubercles on dorsal face, more developed near mesial margin. Left cheliped smaller than right; fingers 1.5 length of palm; dorsal face of carpus slightly convex, with midline of strong conical tubercles. Second and third pereopods strong; on dorsal and ventral faces of meri and on propodi transversal granulated grooves with short setae inserted; dactyls slightly turned inside, with median grooves on lateral faces. Telson with posterior lobes short and broad, left lobe more developed than right; posterior margins armed with irregular strong spines. Shield and cephalic region: A, Pagurus brevidactylus; B, P. criniticornis; C, P. exilis; D, P. heblingi; E, P. leptonyx; F, P. limatulus; G, P. longimanus; H, P. provenzanoi.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF99FFF4FF66FA7002B97663.taxon	distribution	Distribution. Western Atlantic — Brazil (from Rio de Janeiro to Rio Grande do Sul), Uruguay and Argentina. Habitat. From shallow waters to 50 metres, on sandy and muddy bottoms.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF99FFF4FF66FA7002B97663.taxon	materials_examined	Material examined. Brazil: Rio de Janeiro – Cabo Frio, 30 m, 1 male: 6.4 mm (MZUSP – 8635); 45 m, 1 male: 4.9 mm (MZUSP – 8637); 30 m, 1 female: 4.8 mm (MZUSP – 8638); Proj. Ilha Grande, st. 335, 1 male: 3.6 mm (MZUSP – 4700); st. 268, 2 males: 4.8, 3.1 mm (MZUSP – 4702); st. 294, 15 m, 1 female: 3.1 mm (MZUSP – 4696); st. 273, 2 males: 4.7, 3.6 mm (MZUSP – 4704). São Paulo – Ubatuba, Proj. Integrado, st. 29, 1 male: 4.5 mm (MZUSP – 8751); st. 15, 3 males: 5.1, 4.6, 4.7 mm (MZUSP – 8745); st. 22, 4 males: 5.5, 5.4, 5.3, 4.2 mm (MZUSP – 8749); st. 26, 3 males: 5.5, 5.0, 4.7 mm (MZUSP – 8750); st. 33, 2 females: 3.7, 3.6 mm (MZUSP – 8753); st. 14, 1 male: 5.3 mm (MZUSP – 8744); st. 20, 4 males: 4.6, 4.8, 5.5, 4.9 mm (MZUSP – 8747); st. 42, 3 males: 5.1, 4.9, 4.5 mm (MZUSP – 8754); fixed st. 4, 3 males: 5.9, 4.8, 5.7 mm (MZUSP – 8757); st. 18, 1 male: 6.5 mm (MZUSP – 8746); Ilha da Moela, 1 male: 4.9 mm, 1 female: 4.5 mm (MZUSP – 7363); 2 males: 5.2, 5.2 mm (MZUSP – 7362); Santos, 36 m, 1 female: 4.6 mm (MZUSP – 7360); 1 male: 5.2 mm (MZUSP – 7361); 3 females: 4.7, 4.4, 3.5 mm, 1 male: 3.9 mm (MZUSP – 13346). Rio Grande do Sul – Proj. GEDIP, st. 409, 32 m, 1 male: 5.1 mm (MZUSP – 4698).	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF99FFF4FF66FA7002B97663.taxon	discussion	Remarks. Pagurus exilis is relatively close to P. longimanus (which is only found in Amapá), P. limatulus and P. heblingi. The four species are similar in the aspect of the cephalic region, but can be quickly distinguished by differences in the shape and ornamentation of the chelipeds and ambulatory legs.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9DFFF5FF66FF6A02607275.taxon	diagnosis	Diagnosis. Shield slightly broader than long, anterior margins concave between rostrum and lateral projections. Rostrum obtuse, rounded. Lateral projections slightly overreaching rostrum, each with small spine turned outwards. Ocular peduncles short and stout, corneae dilated. Ocular acicles each with strong submarginal spine, mesial margins straight. Right cheliped longer and stronger than left; dactyl approximately twothirds length of palm; palm slightly convex, approximately as long as carpus, dorsolateral margin with low serrulate teeth; dorsomesial margin of carpus with row of spines increasing in size from median to distal region; merus with several transverse rows of setae on dorsal, dorsomesial, and mesial faces. Left cheliped approximately as long as right, but much weaker and more slender; dactyl and fixed finger slender, each longer than palm; carpus as long as merus, with row of irregular spines on ventromesial and ventrolateral margins; merus with several scattered transverse row of setae. Second pereopods long and slender; dactyls longer than propodi, with setae on dorsal face, curved ventrally; propodi longer than carpi, unarmed; dorsal face of carpi with tufts of setae and strong distal spine; meri twice longer than carpi and with 4 or 5 spines on each ventral margin; meri of third pereopods totally unarmed. Telson with posterior lobes almost symmetrical, separated by median cleft; terminal margins each armed with strong spines; lateral margins each with tufts of long setae.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9DFFF5FF66FF6A02607275.taxon	distribution	Distribution. Known only from type locality, Cabo Frio, Rio de Janeiro, in depths between 300 and 400 metres. Types. Holotype — male, Cabo Frio, RJ, Carlo M. da Cunha col. (MZUSP – 13993); Paratypes — 8 males, Cabo Frio, RJ, Carlo M. da Cunha col. (MZUSP – 13994).	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9DFFF5FF66FF6A02607275.taxon	materials_examined	Material examined. Brazil: Rio de Janeiro — Cabo Frio, 9 males with shield lengths from 3.2 to 4.4 mm (MZUSP – 13993 and MZUSP – 13994).	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9DFFF5FF66FF6A02607275.taxon	discussion	Remarks. Pagurus heblingi, P. exilis, P. longimanus and P. limatulus are similar in the general aspect of the cephalic region. These species can be easily distinguished by means of several characters: the ocular acicles of P. heblingi have straight mesial margins, whereas in P. exilis, P. longimanus and P. limatulus they are rounded. The right cheliped of P. heblingi has the palm ovate and almost without tubercles and granules; in P. exilis, the palm is elongated and covered by tubercles and granules and in P. longimanus, the palm is longer than the palms of the other three species. In P. heblingi, the dactyls of the ambulatory legs are longer and / or more slender than in P. exilis, P. longimanus and P. limatulus. P. heblingi also has the antennular flagellum longer than in all other species of the genus Pagurus which occur in Brazil.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9DFFFAFF66FA320330757D.taxon	diagnosis	Diagnosis. Shield as broad as long. Rostrum obtuse, triangular, smaller than lateral projections. Ocular peduncles elongate and slender, corneae slightly dilated. Ocular acicles each with 3 or 4 submarginal spines and sometimes 1 accessory marginal spinule. Antennular and antennal peduncles overreaching corneae; flagellum of antennal peduncle with long setae inserted in pairs. Right cheliped flattened on mesial face, palm longer than carpus, length of fingers half that of palm; dorsomesial margin of carpus with single or double row of spines or spinulose tubercles. Left cheliped much smaller than right; carpus elongate, with row of strong teeth on dorsomesial region. Second and third pereopods with dactyls curved along entire length; ventral margins each with row of corneous spinules; propodi unarmed; carpi each with dorsodistal spine. Telson with posterior margins strongly oblique, armed with large and small spines; lateral margins with corneous spinules.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9DFFFAFF66FA320330757D.taxon	distribution	Distribution. Western Atlantic — Brazil (from Ceará to Santa Catarina). Habitat. On sandy and muddy substrates, from shallow waters to 20 metres.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9DFFFAFF66FA320330757D.taxon	materials_examined	Material examined. Brazil: Alagoas — Maceió, st. 31, 2 females: 1.5, 1.2 mm, 1 male: 1.3 mm (MZUSP – 6472); ES- 1, 2 females: 1.9, 1.0 mm (MZUSP – 6829); S- 3, 1 male: 2.2 mm (MZUSP – 6831); 1 female: 1.6 mm (MZUSP – 6827); EO- 21, 1 male: 1.9 mm (MZUSP – 6835). Sergipe – Aracaju, 2 males: 2.1, 2.3 mm (MZUSP – 9603). São Paulo – Proj. Monitoramento, st. 14, 5 males: 1.6, 2.2, 1.7, 1.3, 1.6 mm, 2 females: 1.3, 1.2 mm (MZUSP – 13347).	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF9DFFFAFF66FA320330757D.taxon	discussion	Remarks. Pagurus leptonyx is very close to P. criniticornis and P. trichocerus. One can quickly distinguish P. leptonyx from P. criniticornis by means of the ocular acicles, which in P. criniticornis have only one spine. Pagurus trichocerus, which occurs in Uruguay, can be distinguished from P. l e p t o n y x basically by the shape and ornamentation of the right cheliped and by the armature of the dactyls of the ambulatory legs.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF93FFFBFF66FF6A0470730D.taxon	diagnosis	Diagnosis. Shield slightly broader than long. Rostrum almost obsolete. Ocular peduncles short, with median constriction and corneae dilated. Ocular acicles rounded, with small ventrodistal spine. Antennular peduncles overreaching corneae, to midlength of third segment. Antennal peduncles long, almost reaching tip of antennular flagellum. Right cheliped long; palm slightly larger than fingers; surfaces of dactyls, palm, carpus and merus granulated; median regions of palm and carpus with each 1 longitudinal row of granules, slightly larger on carpus distally. Left cheliped with lateral face of palm finely granulated, and median longitudinal row of granules little larger; carpus very short. Second and third pairs of pereopods each with sulci on lateral and mesial face of dactyl, with mesial groove more conspicuous; meri with dorsal fringe of setae; dactyls of second pereopods curved ventrally, 1.5 length of propodi Telson with anterior lobes rounded and posterior lobes almost straight; posterior margin with small central cleft and irregular spines.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF93FFFBFF66FF6A0470730D.taxon	distribution	Distribution. Western Atlantic — Brazil (Pará, Ceará and Rio Grande do Norte). Habitat. From very shallow waters to 10 metres on muddy substrates.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF93FFFBFF66FF6A0470730D.taxon	materials_examined	Material examined. Brazil: Pará — Proj. NNE I, st. 1755, 80 m, 5 males: 5.7, 5.8, 5.3, 6.3, 4.1 mm, 4 females: 3.8, 4.2, 3.1, 3.6 mm (MZUSP – 8870).	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF93FFFBFF66FF6A0470730D.taxon	discussion	Remarks. Fausto-Filho (1970 a) described a new Brazilian species, Pagurus limatulus, based on material from Ceará and Rio Grande do Norte. This species is very close to P. exilis, but can be distinguished by means of several differences in the shape and ornamentation of the chelipeds and ambulatory legs. Pagurus limatulus is also close to Pagurus heblingi.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF93FFFBFF66FBAA04907095.taxon	diagnosis	Diagnosis. Shield flattened, broader than long. Rostrum obsolete. Ocular peduncles short and stout, with corneae dilated. Ocular acicles triangular, concave and with minute submarginal spine. Antennular peduncles long, with ultimate segment overreaching completely base of corneae. Antennal peduncles with acicles reaching corneae. Right cheliped long; merus and carpus stout; dorsal surface of carpus with transverse rows of denticles, dorsomesial margin with some irregular denticles, lateral margin granulated; palm longer than carpus; fingers flattened, cutting edge with fine calcareous teeth. Left cheliped with palm smooth, and 1.5 length of carpus; dorsal face of carpus with spines. Second and third pereopods slender; carpi with dorsal crest of spines; dactyls with short and stiff distal sheaf of setae on dorsal margin. Telson with lobes asymmetrical and distal margins bearing irregular spines.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF93FFFBFF66FBAA04907095.taxon	distribution	Distribution. Western Atlantic — Colombia, French Guiana and Brazil (Amapá). Habitat. Little is known about this species, only that the individual were collected at 60 metres depth.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF93FFFBFF66FBAA04907095.taxon	materials_examined	Material examined. Brazil: Amapá – “ Alm. Saldanha ”, st. 2012 A, 1 male: 3.7 mm (DOUFPE – 3680).	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF93FFFBFF66FBAA04907095.taxon	discussion	Remarks. Wass (1963) described Pagurus longimanus based on only one incomplete male from French Guiana. Later Sánchez & Campos (1978), studying material from Colombia, examined 20 individuals of P. longimanus and complemented the original description of Wass (1963). Pagurus longimanus is most closely related to P. exilis. The two species can be distinguished by differences in the length, width and ornamentation of the larger cheliped, and in the telson and rostrum.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF90FFF8FF66FF6A052C73DB.taxon	diagnosis	Diagnosis. Shield slightly longer than broad. Rostrum triangular, obtuse or rounded, without terminal spine. Ocular peduncles long, with corneae and bases slightly dilated; ocular acicles each with 5 or 6 denticules. Antennular peduncles overreaching corneae by 1 / 4 length of last segment. Antennal peduncles as long as ocular peduncles. Right cheliped larger than left; palm twice longer than broad, with longitudinal row of acute tubercles; fingers as long as palm; dorsal margin of merus with one distal tooth; carpus with row of spines on dorsomesial margin and several spines and spinules in dorsolateral region; chela covered by dense short setae; dactyl with row of small spines on dorsomesial margin and dorsal midline, also covered by dense setae. Left cheliped with acute spines on each ventral margin of merus; dorsolateral and dorsomesial margins of carpus with row of strong spines; dorsomesial margin of dactyl with row of small spinules and with row of small spines in dorsal midline. Second and third pereopods with dactyls shorter than propodi, each terminating in strong corneous claw, ventral margins each with row of strong corneous spines; propodi unarmed; carpi each with marginal spine dorsodistally. Exopodite of left uropod with ribbon-shaped row of long setae. Distal margins of telson with several strong spines, and lateral margins with corneous spinules or denticles.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF90FFF8FF66FF6A052C73DB.taxon	distribution	Distribution. Western Atlantic - Bermudas, Florida, Antilles, Central America, Colombia, Venezuela, Brazil (Fernando de Noronha and from Pernambuco to Rio Grande do Sul) and Uruguay. Habitat. On several kinds of substrate, including coralline formations, from shallow waters to about 100 metres depth.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF90FFF8FF66FF6A052C73DB.taxon	materials_examined	Material examined. Brazil: Alagoas — Ponta Verde, 2 males: 1.5, 1.2 mm (MZUSP – 13897). Espírito Santo — Vitória, Ilha Galheta, 1 female: 2.6 mm (MZUSP – 12912). Santa Catarina — Porto Belo, 1 males: 2.2 mm (MZUSP – 13808).	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
039287BCFF90FFF8FF66FF6A052C73DB.taxon	discussion	Remarks. Pagurus provenzanoi can be distinguished from its congeners in Brazil by its short setae that form a dense covering on the dorsal faces of the chelipeds.	en	Nucci, Paulo Ricardo, Melo, Gustavo Augusto Schmidt De (2007): Hermit crabs from Brazil. Family Paguridae (Crustacea: Decapoda: Paguroidea): Genus Pagurus. Zootaxa 1406: 47-59, DOI: 10.5281/zenodo.175515
