identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03923C218625FF8FFF45FB28025CF993.text	03923C218625FF8FFF45FB28025CF993.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bruaylogus	<div><p>Genus Bruaylogus gen. nov.</p><p>Type species. Bruaylogus magnificus sp. nov.</p><p>Etymology. Named after the outcrop of Bruay-en-Artois and for the similarities with the genus Heterologus. Gender masculine.</p><p>Diagnosis. Forewing characters only. Only two anal veins reaching posterior wing margin; first anal vein 1A fused with second 2A; second visible anal vein (3A) very long and parallel with posterior wing margin; CuPaβ, CuPb and anal veins simple; CuPa differentiated into two branches (CuPaα and CuPaβ) just basal of fusion of anterior one (CuPaα) with CuA; CuA + CuPaα ramified with five posterior branches; a regular space between posterior branches of CuA + CuPaα; basal fork of M rather close to point of separation between M and CuA; MP divided into four distal branches; MA simply forked into MA1 and MA2; RP and MA1 not approximate; RA simple; ScP reaching anterior wing margin and not RA.</p></div>	https://treatment.plazi.org/id/03923C218625FF8FFF45FB28025CF993	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Coty, David;Háva, Jiří;Prokop, Jakub;Roques, Patrick;Nel, André	Coty, David, Háva, Jiří, Prokop, Jakub, Roques, Patrick, Nel, André (2014): New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera). Zootaxa 3878 (5): 462-470, DOI: 10.11646/zootaxa.3878.5.4
03923C218625FF8FFF45FF30031CFEC7.text	03923C218625FF8FFF45FF30031CFEC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cnemidolestodea Handlirsch 1937	<div><p>Subclade Cnemidolestodea Handlirsch, 1937 (sensu Béthoux, 2005) Family Tococladidae Carpenter, 1966</p><p>Genus Tococladus Carpenter, 1966</p></div>	https://treatment.plazi.org/id/03923C218625FF8FFF45FF30031CFEC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Coty, David;Háva, Jiří;Prokop, Jakub;Roques, Patrick;Nel, André	Coty, David, Háva, Jiří, Prokop, Jakub, Roques, Patrick, Nel, André (2014): New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera). Zootaxa 3878 (5): 462-470, DOI: 10.11646/zootaxa.3878.5.4
03923C218625FF8FFF45FE8400A1FBD7.text	03923C218625FF8FFF45FE8400A1FBD7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tococladus	<div><p>Cf. Tococladus species undetermined</p><p>(Fig. 1)</p><p>Material studied: Specimen 0 0 13 (print and counterprint), coll. Patrick Roques, to be deposited in the Musée Géologique Pierre Vetter, Decazeville, France.</p><p>Age and outcrop. Type locality and horizon. Moscovian (Westphalian C/D equivalent to Bolsovian/Asturian), ‘Terril N°7’, Avion, Pas-de-Calais, France.</p><p>Description. A fragment of wing corresponding to the anterior part of the apex, fragment 19.4 mm long, 11.7 mm wide; convex RA with a series of sigmoidal anterior veinlets, RA distally fused with concave main branch of RP, near wing apex, common branch 2.6 mm long; a series of straight crossveins in area between RA and RP; RP with seven to nine parallel posterior branches with a series of simple straight crossveins between them; apical parts of branches of M and possibly CuA also preserved, parallel to the branches of RP.</p><p>Discussion. Although very incomplete, this wing fragment fits quite well with the equivalent portion of a forewing of Tococladus ralus Carpenter, 1966, type species of the Permian genus Tococladus . It is especially remarkable in the distal fusion of RA with RP near wing apex and the numerous parallel posterior branches of RP with short crossveins between them (see Béthoux et al., 2003: fig. 1). Of course an attribution to the genus Tococladus sensu stricto is impossible to establish. The main interest of this fossil is that it represents the second Cnemidolestodea from the outcrop of Avion, the first one being Aviocladus pectinatus Prokop, Roques and Nel, 2014, from which it differs in the more numerous branches of RP and veinlets between RA and costal margin more sigmoidal (Prokop et al., 2014).</p></div>	https://treatment.plazi.org/id/03923C218625FF8FFF45FE8400A1FBD7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Coty, David;Háva, Jiří;Prokop, Jakub;Roques, Patrick;Nel, André	Coty, David, Háva, Jiří, Prokop, Jakub, Roques, Patrick, Nel, André (2014): New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera). Zootaxa 3878 (5): 462-470, DOI: 10.11646/zootaxa.3878.5.4
03923C218625FF89FF45F9510541F9D6.text	03923C218625FF89FF45F9510541F9D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bruaylogus magnificus	<div><p>Bruaylogus magnificus sp. nov.</p><p>(Fig. 2)</p><p>Material studied: Holotype specimen Bruay 00058a (counterpart), coll. Patrick Roques, to be deposited in the Musée Géologique Pierre Vetter, Decazeville, France.</p><p>Age and outcrop. Upper Carboniferous, Bashkirian, Westphalian C or D, Bruay-en-Artois, coal heap n° 26, Department of North, France.</p><p>Etymology. Named after the wonderful state of preservation of the fossil.</p><p>Diagnosis. As for the genus.</p><p>Description. Counterpart of a nearly complete forewing perfectly preserved with only anterior part of apex missing. Preserved length 37.5 mm, width 10.1 mm; no defined ScA; ScP anteriorly pectinate, ending on anterior wing margin, with long branches forked and sigmoidal; area between costal margin and ScP 2.0 mm wide; base of RP 11.5 mm distal of wing base; ScP and R (+ M + CuA) basally separated; presence of a common stem R + (M + CuA); RA convex, simple; area between RA and RP with straight cross-veins; RP rather concave; first fork of RP 6.8 mm from its base, anterior branch with at least one more fork distally, posterior branch anteriorly pectinate; CuA separated from M in basal third of wing; M dichotomously ramified into neutral MA and concave MP; MA well distant from RP, with only one distal fork; MP dichotomously ramified two times (four branches reaching posterior wing margin); CuA fused with CuPaα at base of M; CuA + CuPaα posteriorly pectinate, with its two first posterior branches concave; CuPaα short basal of its fusion with CuA, 0.8 mm long; CuPaβ simple; a series of short crossveins in area between CuA + CuPaα and CuPaβ; division of CuP into CuPa and CuPb 4.0 mm from wing base; CuPb simple; area between CuPa and CuPb narrow and narrowed near posterior wing margin (largest width opposite origin of CuPaα / CuPaβ); three anal veins, all simple; anterior one 1A very short and fused to 2A well separated from CuP at wing base.</p><p>Discussion. This forewing corresponds to that of a clade Panorthoptera Crampton 1928 sensu Béthoux &amp; Nel (2002: 27) because of the following synapomorphies: CuPa differentiated into two branches (CuPaα and CuPaβ) just basal of the fusion of the anterior one (CuPaα) with CuA (forewing autapomorphy of Panorthoptera); ScP reaching anterior wing margin and not RA; MA1 and MA2 can be differentiated; CuA + CuPaα ramified with at least three branches; CuPb simple. Bruaylogus gen. nov. shares with the Orthoptera Olivier, 1789 (sensu Béthoux &amp; Nel, 2002: 38-39) the MA simply forked into MA1 and MA2; and CuPaβ, CuPb and 1A simple. But its MP is divided into four distal branches instead of being simple or with few distal branches and it is not especially very concave. Thus if it certainly belongs to the stem lineage of the Orthoptera, it cannot be included in the crown group of this clade.</p><p>A pattern of venation rather similar to that of Bruaylogus can be found in the Middle Permian Permostridulidae Béthoux et al., 2003 [a family of the panorthopteran order Caloneurodea Handlirsch, 1937 (sensu Béthoux et al., 2004)]: presence of a forked MA, MP with two dichotomies, and CuPaβ, CuPb and 1A simple. The crucial differences between Bruaylogus and Permostridulus Béthoux et al., 2003 are as follows: the vein CuA + CuPaα posteriorly pectinate in the former while it is simple in the latter taxon; CuPaβ and CuA + CuPaα strongly approximate in Permostridulus (apomorphy of the Caloneurodea), while they are less close in Bruaylogus (Béthoux et al., 2003).</p><p>The Oedischiidae Handlirsch, 1906, probably the most basal lineage of the Orthoptera (see Béthoux &amp; Nel, 2002), differ from Bruaylogus in the veins RP and MA1 strongly approximate or fused for a short distance and the vein MP simple or with weak distal branches (Carpenter, 1992; Beckemeyer, 2011). Nevertheless Bruaylogus shares with the Oedischiidae the regular space between the posterior branches of CuA + CuPaα and basal fork of M rather close to point of separation between M and CuA. The Geraridae Scudder, 1885 is a panorthopteran group whose monophyly and affinities with the Orthoptera sensu stricto are not clear (see Béthoux &amp; Nel, 2002). All the taxa currently included in this family (see Zessin, 2012) differ from Bruaylogus in the three characters: RA with well defined branches, MA (or its anterior branch) and RP strongly approximate or fused for a short distance, CuPb with numerous branches (this last character is unknown in Hirsutgerarus Zessin, 2012 and Progenentomum Handlirsch, 1906) (Burnham, 1983; Kukalová-Peck &amp; Brauckmann, 1992; Béthoux &amp; Nel, 2002, 2003; Zessin, 2009, 2012). Note that Zessin (2012) listed Anepitedius giraffa Handlirsch, 1911 in the gerarids, but Béthoux &amp; Nel (2002: 32) considered it as a ‘Protorthoptera’ incertae sedis, which means that it is a very poorly known taxon. Lastly Heterologus duyiwuer Béthoux et al., 2012 and Heterologus langfordorum Carpenter, 1944 share with Bruaylogus the median and radial veins well separated, a simple CuPaβ, and M with numerous branches, but both strongly differ from Bruaylogus in the long stem of M before its first fork (Carpenter, 1944; Béthoux et al., 2012). Furthemore, H. duyiwuer differs from Bruaylogus in the distally forked CuPb and first branch of CuA + CuPaα in a basal position. A further difference between Heterologus duyiwuer and Heterologus langfordorum is the presence of three anal veins reaching posterior wing margin in H. duyiwuer while there are only two in H. langfordorum (after the photograph of the type in Carpenter, 1944: pl. 3), as in Bruaylogus . Furthermore Bruaylogus and H. langfordorum share the second visible anal vein (3A) very long and parallel with posterior wing margin. Possibly H. langfordorum has the first anal vein 1A fused with second 2A as in Bruaylogus .</p><p>In conclusion Bruaylogus differs from all described Panorthoptera nec Orthoptera in a series of characters that justify a new genus and species. Bruaylogus could be closer to Orthoptera in some characters (pattern of stem of M, regular branching of CuA + CuPaα) but the character “ RP and M not approximating” suggests a more basal position for Bruaylogus than the Geraridae, maybe close to the Heterologus spp. A new phylogenetic analysis of the basal taxa of the Panorthoptera becomes necessary after the recent discoveries of the last 10 years.</p><p>Remark. Another wing fragment (specimen 00058b, see figure 3) is fossilised 35 mm from the specimen 00058a. It corresponds to the apical part of a wing attributable to a Polyneoptera, and possibly to an Archaeorthoptera. It is probably not the same taxon as specimen 00058a because the distribution and shape of the crossveins are different (denser and more irregular pattern in specimen 00058b).</p></div>	https://treatment.plazi.org/id/03923C218625FF89FF45F9510541F9D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Coty, David;Háva, Jiří;Prokop, Jakub;Roques, Patrick;Nel, André	Coty, David, Háva, Jiří, Prokop, Jakub, Roques, Patrick, Nel, André (2014): New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera). Zootaxa 3878 (5): 462-470, DOI: 10.11646/zootaxa.3878.5.4
03923C218623FF88FF45F96D000BFAD4.text	03923C218623FF88FF45F96D000BFAD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aviologus	<div><p>Genus Aviologus gen. nov.</p><p>Type species. Aviologus duquesnei sp. nov.</p><p>Etymology. Named after the outcrop of Avion, and for the similarities with the genus Heterologus. Gender masculine.</p><p>Diagnosis. ScP ending on RA; RA without well defined branches; fusion of MA with first posterior branch of RP; MP divided into two small distal branches; basal fork of M far from point of separation between M and CuA; CuPaβ and CuA + CuPaα not strongly approximate; CuPaβ, CuPb and 1A simple; three anal veins ending on posterior wing margin.</p><p>FIGURE. 3. Archaeorthoptera gen. et sp. indet., specimen Bruay 00058b stored in MNHN coll. finally deposited in the Musée Géologique Pierre Vetter, Decazeville, France. A, photograph of wing apex venation; B, line drawing of apical part of wing. Scale bar = 5 mm.</p></div>	https://treatment.plazi.org/id/03923C218623FF88FF45F96D000BFAD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Coty, David;Háva, Jiří;Prokop, Jakub;Roques, Patrick;Nel, André	Coty, David, Háva, Jiří, Prokop, Jakub, Roques, Patrick, Nel, André (2014): New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera). Zootaxa 3878 (5): 462-470, DOI: 10.11646/zootaxa.3878.5.4
03923C218622FF8BFF45FA98009DFCA6.text	03923C218622FF8BFF45FA98009DFCA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aviologus duquesnei	<div><p>Aviologus duquesnei sp. nov.</p><p>(Fig. 4)</p><p>Material studied: Holotype specimen Avion 0 0 0 34 (print and counterprint), coll. Patrick Roques, to be deposited in the Musée Géologique Pierre Vetter, Decazeville, France.</p><p>Age and outcrop. Type locality and horizon. Moscovian (Westphalian C/D equivalent to Bolsovian/Asturian), ‘Terril N°7’, Avion, Pas-de-Calais, France.</p><p>Etymology. Named after Mr. Hervé Duquesne, who is a well known collector of plants and insects in the Carboniferous of North of France.</p><p>Diagnosis. As for the genus.</p><p>Description. A nearly complete forewing perfectly preserved with only fragments of mid part missing. Preserved length 22.6 mm, width 6.9 mm; no defined ScA; ScP anteriorly pectinate, ending on RA, with branches simple or forked and more or less sigmoidal; area between costal margin and ScP 0.8 mm wide; base of RP 5.3 mm distal of wing base; ScP and R (+ M + CuA) basally separated; presence of a common stem R + (M + CuA); RA convex, simple; crossveins in area between RA and RP not preserved, if present; RP rather concave; first fork of RP 6.5 mm from its base, RP with seven branches on wing margin; first branch short before its fusion with MA; CuA separated from M in basal third of wing; M dichotomously ramified into neutral MA and concave MP, far, 5.3 mm, from its base; MA distally fused with first branch of RP, simple; MP with two branches near posterior wing margin; CuA fused with CuPaα at base of M; CuA + CuPaα posteriorly pectinate, with its first posterior branch concave; CuPaα short basal of its fusion with CuA; CuPaβ present, simple; a series of short crossveins in area between CuA + CuPaα and CuPaβ; division of CuP into CuPa and CuPb 2.8 mm from wing base; concave CuPb simple; area between CuPa and CuPb narrow; three anal veins ending on posterior wing margin, all simple and more convex than CuP.</p><p>Discussion. This forewing corresponds to that of a clade Panorthoptera for the same reasons as above (except for the ScP ending on RA). The veins CuPaβ and CuA + CuPaα not strongly approximate exclude affinities with the Caloneurodea. Aviologus gen. nov. shares with the Orthoptera the veins CuPaβ, CuPb and 1A simple, and MP divided into two small distal branches. Nevertheless Aviologus differs from the Oedischiidae in the basal fork of M far from point of separation between M and CuA. Also its fusion of MA with first posterior branch of RP is quite different from the stem of RP and MA strongly approximate or fused for a short distance of the Oedischiidae . Aviologus differs from the Geraridae in the three characters: RA without well defined branches, MA fused with first posterior branch of RP instead of being fused or approximating the stem of RP, and CuPb simple. Lastly Heterologus duyiwuer and Heterologus langfordorum share with Aviologus a simple CuPaβ, and a long stem of M before its first fork; but both strongly differ from Aviologus in the median and radial veins well separated and MP with numerous branches. Bruaylogus differs from Aviologus in median and RP veins well separated, MP with numerous branches, M with a long stem, plus the presence of only two anal veins ending on posterior wing margin instead of three in Aviologus . Aviologus corresponds to a new genus but its family affinities remains undetermined since a new phylogenetic analysis is not performed, as for Bruaylogus . It is even not sure if they belong to the same or to two separate families.</p></div>	https://treatment.plazi.org/id/03923C218622FF8BFF45FA98009DFCA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Coty, David;Háva, Jiří;Prokop, Jakub;Roques, Patrick;Nel, André	Coty, David, Háva, Jiří, Prokop, Jakub, Roques, Patrick, Nel, André (2014): New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera). Zootaxa 3878 (5): 462-470, DOI: 10.11646/zootaxa.3878.5.4
