taxonID	type	description	language	source
03926F62FFEEFFE5D2EDFD50FCA73CF8.taxon	distribution	Distribution. South America: Brazil, Espírito Santo, Minas Gerais, Rio de Janeiro, São Paulo. Included species. H. bicornutus (Mello-Leitão, 1940), H. nigripalpis Roewer, 1910, H. dissimilis sp. nov., H. ibitirama sp. nov., H. uaisoh sp. nov.	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEEFFE5D2EDFD50FCA73CF8.taxon	discussion	Remark: Males of the four species of Holcobunus have very similar external morphology, although they can be separated by examining their color pattern. There are three yellow species included in Holcobunus. However only H. ibitirama sp. nov. is completely yellow without stains or stripes of other colors, while H. nigripalpis have anterior border of carapace with black spots. The median posterior region of the dorsal scute and anterior border of the coxae are covered by a faint black spot. Anterior median portion of scute and middle of second prosomatic tergite (metapeltidium) and carapace behind eye mound with longitudinal short stripe of same color. Holcobunus dissimilis sp. nov has a larger and longer black stripe extending from the anterior to the posterior dorsal scute. Females of H. nigripalpis have one lateral spot on each side of the posterior dorsal scute, while the female of H. dissimilis sp. nov. having a brown dorsal background with a white arch surrounding the anterior prosoma and several white spots and stripes distributed on the dorsal scute.	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEDFFE0D2EDFC4DFCCA3E6A.taxon	etymology	Etymology. From Latin dissimilis (dis- + similis = different, unlike or dissimilar) is given in reference to the highly different color pattern between males and females of this species.	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEDFFE0D2EDFC4DFCCA3E6A.taxon	materials_examined	Type locality. Brasil, Espírito Santo, Santa Teresa, Estação Biológica Santa Lúcia (19 º 58 ’ 36 ” S – 40 º 32 ’ 06 ” W, 867 m), Type material. Brasil, Espírito Santo, Santa Teresa, Estação Biológica Santa Lúcia, D. Mota leg., 25. VIII. 2006; male holotype (MZSP- 29514); idem, 1 male paratype (MZSP- 29516), REBIO Augusto Ruschi, R. Pinto-da-Rocha, C. Bragagnolo & M. B daSilva leg., 11. IV. 2009; 1 male paratype (MZSP 57151); idem, 2 males, 6 females paratypes (MNRJ- 17968), T. G. Souza leg., 26. VIII. 2006; 2 males, 4 females paratypes (MNRJ- 18627), C. O. Azevedo et al. leg, 10. VI. 2001.	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEDFFE0D2EDFC4DFCCA3E6A.taxon	distribution	Distribution. Known only from mountains of Espirito Santo State in the municipality of Sanat Tereza (Fig. 7).	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEDFFE0D2EDFC4DFCCA3E6A.taxon	diagnosis	Diagnosis. Male: specimens preserved in ethanol have yellow body, eye mound black, abdomen with large dark median stripe along its full extent (Fig. 1 A, C), legs black, coxae I – II dorsally with pale yellow spots. Femur, patella and proximal half of pedipalpal tibia black, proximal half of tibia and tarsus cream, chelicerae black (Fig. 1 A – C). Eye mound armed with sharp granules (Fig. 1 C). Femoral nodule formula: 2 / 4 - 5 / 2 - 3 / 3 - 4. Dorsal orifice of anterior shaft of penis lacking (Fig. 2 A, see also Tourinho & Kury, 2001, Fig. 37), glans with three pairs of subapical setae (Fig. 2 C, E).	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEDFFE0D2EDFC4DFCCA3E6A.taxon	description	Description. Lengths: body: 2.75 mm, carapace: 0.85 mm, chelicera: 0.6 mm, pedipalps: 2.82 mm. Femora: 10.5 / 18.7 / 10.2 / 14.4 mm. Dorsal surface: body surface reticulate. Supracheliceral lamina armed distally with one triangular and crenulated tubercle on each lobe. Eye mound armed with two rows of 10 or more sharp granules. Ventral surface reticulate. Lateral margins of genital operculum unarmed, arculi genitales II unarmed. Chelicerae: ventrobasal spine of basichelicerite blunt. Pedipalps: femur armed with ventral row of sharp pointed tubercles. Patella, including inner apophysis, armed with dorsolateral sharp-pointed tubercles, inner apical apophysis longer than wide. Tibia armed with sharp tubercles. Tarsus unarmed. Legs: femoral pseudoarticular formula: 2 / 5 / 2 / 2 - 3. Penis (Fig. 2): morphology very similar to H. nigripalpis. Winglets longer than wide, narrower than shaft, rift depressions (see Tourinho & Kury, 2001, Fig. 35 – 36) of shaft present (Fig. 2 A – B), ventral collar of winglet present but not marked (Fig. 2 B and D), shaft almost straight. Dorsal and ventral faces of glans not depressed, with three pairs of subapical setae (Fig. 2 C – F). Stylus straight not bent in ventral or lateral view (Fig. 2 C). Female paratype (Fig. 1 D – E): length: Body: 4.1 mm, carapace: 0.85 mm, abdominal dorsal scute: 2.9 mm, chelicera: 0.6 mm, pedipalps: 3.0 mm. Color: dorsal background brown with a white arch surrounding anterior prosoma and several white spots and stripes distributed on scute, eye mound darkish brown with a white stripe surrounding the contour of the eye (Fig. 1 D, F). Trochanter I – IV cream with darkish lateral spots, coxae with darkish brown spots (Fig. 1 D). Legs: femoral nodule formula: 2 / 4 - 5 / 3 / 3 - 4, femur, patella and proximal half of pedipalpal tibia black, proximal half of tibia and tarsus cream, chelicera cream with a dorsal darkish spot on fixed finger. Ventral surface cream (Fig. 1. F, E). Dorsal and ventral surfaces: as in the male, except the body swollen dorsoventrally and by having an articular membrane isolating all sclerites. Membrane much evident, widely surpassing sclerites.	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEDFFE0D2EDFC4DFCCA3E6A.taxon	discussion	Remarks. This species has external and genital morphology very similar to H. nigripalpis, but is distinguished by the large black stain present on male dorsal scute and the lack of dorsal orifice on anterior portion of penis shaft present in H. nigripalpis (Tourinho & Kury, 2001. Fig. 37)	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEBFFE1D2EDFA02FE223A1E.taxon	etymology	Etymology. After type locality, Ibitirama, a word of the indigenous Tupi people of Brazil that means " region with many mountains. "	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEBFFE1D2EDFA02FE223A1E.taxon	materials_examined	Type locality. Brazil, Espírito Santo, Ibitirama, Santa Marta. Type Material: Brazil, Espírito Santo, Ibitirama, Santa Marta, (20 o 29 ’ 52 ” S – 41 o 43 ’ 47 ” W) M. B. da Silva & G. E. Kaneto leg., III. 2006, MZSP 57264, male holotype and two male paratypes; 3 immatures.	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEBFFE1D2EDFA02FE223A1E.taxon	distribution	Distribution. only known from type locality (Fig. 7).	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEBFFE1D2EDFA02FE223A1E.taxon	diagnosis	Diagnosis. Body color orange, legs trochanter and eye mound black (Fig. 3 A – C). Femur, patella and proximal half of pedipalpal tibia brownish, distal half of tibia and tarsus cream (Fig. 3 A, C), chelicerae cream. Eye mound armed with sharp granules (Fig. 3 C). Femoral nodule formula: 2 / 4 / 2 - 3 / 3 - 4.	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEBFFE1D2EDFA02FE223A1E.taxon	description	Description. Male paratype: lengths: body: 2.55 mm, carapace: 0.71 mm, chelicera: 0.80 mm, pedipalps: 2.75 mm. Femora: 9.6 / 15.4 / 8.5 / 11.9 mm. Dorsal and ventral body surface reticulate. Supracheliceral lamina armed distally with one triangular and crenulated tubercle on each lobe. Eye mound armed with two rows of 10 or more sharp granules. Lateral margins of genital operculum armed with three-pointed, sharp granules, arculi genitales II unarmed. Chelicerae: ventrobasal spine of basichelicerite blunt. Pedipalps: femur armed with ventral row of sharp pointed tubercles. Patella, including inner apophysis, armed with dorsolateral sharp-pointed tubercles, inner apical apophysis as long as wide. Tibia armed with sharp tubercles. Tarsus unarmed. Legs: femoral nodule formula: 2 / 4 / 2 - 3 / 3 - 4. Penis (Fig. 4): Winglets not projected laterally, longer than wide, shorter than in other species of Holcobunus (Fig. 4 A, B). Dorsal surface of glans not depressed (Fig. 4 C, D). Stylus bent in ventral view, with two pairs of lateroapical setae (Fig. 4 D).	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEAFFE2D2EDF955FD223D9C.taxon	etymology	Etymology. “ uai sô ” is a popular expression from people of Minas Gerais State of Brazil, that has English origin, meaning and pronunciation of “ Why so? ”	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEAFFE2D2EDF955FD223D9C.taxon	materials_examined	Type locality. Brasil, Minas Gerais, Fervedouro, Parque Estadual Serra do Brigadeiro. Type Material. Male holotype, 4 females paratypes, 3 immatures (MZSP 69117), M. B. daSilva & G. E. Kaneto leg., III. 2006.	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEAFFE2D2EDF955FD223D9C.taxon	distribution	Distribution. Only known from the type locality (Fig. 7).	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEAFFE2D2EDF955FD223D9C.taxon	diagnosis	Diagnosis. Males with body color black (Fig. 5 A – C), females with light dorsal stripes and spots, eye mound with a pair of yellowish stripes (Fig. 5 D – F). Femur, patella and proximal half of pedipalpal tibia black, distal half of tibia and tarsus cream, chelicerae cream (Fig. 5 C and F). Eye mound armed with sharp granules. Femoral nodule formula: 2 / 5 - 6 / 2 - 3 / 2 - 3. Penis: Dorsal orifice of anterior shaft of penis lacking (Fig. 6 A and C).	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFEAFFE2D2EDF955FD223D9C.taxon	description	Description. Male paratype: lengths: body: 2.65 mm, carapace: 0.81 mm, chelicera: 1.22 mm, pedipalps: 2.95 mm. Femora: 10.1 / 19.2 / 9.5 / 14.2 mm. Dorsal and ventral surface body surface reticulate. Supracheliceral lamina armed distally with one or two blunt tubercles on each lobe. Eye mound armed with two rows of 10 or more sharp granules (Fig. 5 C). Lateral margins of genital operculum armed with three-pointed sharp granules, arculi genitales II unarmed. Chelicerae: ventrobasal spine of basichelicerite blunt. Pedipalps: Femur armed with a ventral row of sharp pointed tubercles. Patella, including inner apophysis, armed with dorsolateral sharp pointed tubercles, inner apical apophysis longer than wide. Tibia armed with sharp tubercles. Tarsus unarmed. Legs: Femoral nodule formula: 2 / 5 - 6 / 2 - 3 / 2 - 3. Penis (Fig. 6) similar to H. nigripalpis and H. dissimilis. Winglets longer than wide, narrower than shaft (Fig. 6 A – B), ventral collar present but not very marked (Fig. 6 B and D), shaft almost straight. Dorsal and ventral faces of glans not depressed, with two pairs of subapical setae (Fig. 6 C – F). Stylus bent in ventral view (Fig. 6 D). Female paratype: lengths: body: 4.12 mm, carapace: 1.15 mm, chelicera: 1.4 mm, pedipalps: 2.9 mm. As in the male, except body swollen dorsoventrally and articular membrane isolating all sclerites (Fig. 5 D – F). Membrane very evident, widely surpassing sclerites (Fig. 5 E).	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFE9FFE3D2EDF891FD3C3BA6.taxon	diagnosis	Diagnosis. Holcobunus bicornutus is know only by the juvenile holotype, although it can be separated from the other species of Holcobunus by the eye mound armed with two anterior blunt spines, longer than in other species (see Tourinho & Kury, 2001, Figs 9 and 11), and by the anterior leg I much shorter than body when compared to other species (see Tourinho & Kury, 2001, Fig. 11).	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFE7FFEDD2EDF9D4FEA73AF2.taxon	materials_examined	Type locality. Brazil, São Paulo, Santos.	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFE7FFEDD2EDF9D4FEA73AF2.taxon	diagnosis	Diagnosis. Living specimens are darkish yellow, museum specimens are pale yellow. Males of H. nigripalpis can be separated from other species of Holcobunus by having the body yellow without a marked strong stripe; in contrast to H. dissimilis males, it does not have a dark well-marked median stripe; it has a faint black spot covering the anterior median portion of dorsal scute instead. Dorsal orifice of anterior shaft of penis present. Females are yellow but with rounded black spot on posterolateral border.	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFE7FFEDD2EDF9D4FEA73AF2.taxon	materials_examined	Material examined (New Record). Brazil. 1 male MNRJ 4967, 1 female MNRJ 4968, Minas Gerais, Rio Preto, Santa Bárbara, Fazenda do Luluca, 29 / 03 / 2002, leg. Amazonas Chagas, Alessandro Giupponi.	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
03926F62FFE7FFEDD2EDF9D4FEA73AF2.taxon	distribution	Distribution. Brazil. Minas Gerais state. Rio Preto. Rio de Janeiro state. Guapimirim, Lumiar, Nova Friburgo, Petrópolis: Itaipava, Rezende: Serrinha do Alambari, Silva Jardim: Aldeia Velha, Teresópolis. São Paulo state. Santos. (Fig. 7).	en	Tourinho, Ana Lúcia, Pinto-Da-Rocha, Ricardo, Bragagnolo, Cibele (2015): Taxonomic notes on Holcobunus Roewer, 1910, with descriptions of three new species, and new records for Holcobunus nigripalpis Roewer, 1910 (Opiliones: Eupnoi: Sclerosomatidae). Zootaxa 4027 (3): 425-436, DOI: 10.11646/zootaxa.4027.3.6
