taxonID	type	description	language	source
03926A1E9E557039FF58F8E4FE829082.taxon	description	(Figs 1 – 6)	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E557039FF58F8E4FE829082.taxon	materials_examined	Type material. Holotype male, CHINA: Sichuan Province, Kangding County, Wasi Gully, 15. vi. 1996, light trap, X. Wang (BDN). Paratypes: 8 males, CHINA: Sichuan Province, Kangding County, Zheduo River, 8. vi. 1996, X. Wang; 2 males, Yunnan Province, Dali City, Wucang Mountain, Qingbi Brook, 23. v. 1996, X. Wang; 2 males, Yunnan Province, Lijiang Country, Tiger Leaping Gorge, 26. v. 1996, Y. Du.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E557039FF58F8E4FE829082.taxon	etymology	Etymology. From Latin alula, posterior lobe of wing and setosa, with setae. Diagnostic characters. The species can be separated from other species of the genus on the inflated gonostylus, presence of many small spinules in apical part of the aedeagal lobe, and by having 13 flagellomeres and alula with 3 – 4 setae. Male (n = 14 – 15) Total length 2.3 – 3.0 mm. Wing length 1.43 – 2.08 mm. Total length / wing length 1.40 – 1.58. Coloration. Head brown to dark brown. Antenna yellowish to brown. Thorax brown to dark brown. Wing dark grey. Abdomen and legs yellowish to dark brown; femur and tibia of all legs with rings, basal and apical parts light brown, median part white. Head. Frontal tubercles triangular to suboval, 15 – 28 µm long; with 1 – 2 setae, 8 – 58 µm long. Temporal setae includes 9 – 20 frontals, 5 – 17 outer verticals, no inner verticals, 4 – 8 postorbitals. Clypeus with 7 – 16 setae. Antenna with 13 flagellomeres, antennal plume reduced, ultimate flagellomere with few small white spots (Figs 4, 6); pedicel with 2 – 4 setae. Lengths (in µm) of flagellomeres 2 – 13: 18 – 26, 21 – 30, 30 – 50, 52 – 68, 52 – 69, 48 – 66, 40 – 63, 38 – 50, 33 – 48, 28 – 45, 24 – 35, 34 – 63. Number of setae on flagellomeres 2 – 13 as: 3 – 4, 4, 4 – 5, 4 – 5, 4 – 5, 4 – 5, 4, 4, 3 – 4, 3 – 4, 3 – 4, 1. Subapical seta of terminal flagellomere 13 – 28 µm long. AR 0.07 – 0.11. Lengths (in µm) of palpomeres 1 – 5: 20 – 38, 42 – 73, 63 – 95, 83 – 125, 144 – 188. Head width / palp length 0.98 – 1.32. Thorax. Antepronotum with 2 – 7 lateral setae. Acrostichals 14 – 38; dorsocentrals 7 – 24, often in 2 groups and sometimes with 1 – 5 setae between the groups; prealars 9 – 23 in 2 groups. Scutellum with 24 – 61 setae. Longest and shortest acrostichals 49 – 103 µm and 18 – 50 µm long. Wing. Costal extension 50 – 88 µm long. Anal lobe well developed, fully fringed. Alula with 1 – 8 setae. Squama with 13 – 34 setae (Fig. 5). R with 10 – 26 setae, R 1 with 0 – 3 setae, R 4 + 5 with 3 – 8 setae, R 2 + 3 reduced (Fig. 3). RM / MCu 1.67 – 2.40. Legs. Spur of foretibia 30 – 45 µm long; spurs of midtibia 26 – 38 µm and 28 – 44 µm long; of hind tibia 33 – 50 µm and 45 – 75 µm long. Hind tibial comb with 11 – 16 setae. Mid ta 1 with 1 – 2 pseudospurs, 23 – 30 µm long; hind ta 1 with 1 – 2 pseudospurs, 22 – 30 µm long. Lengths and proportions of legs as in Table 1. Hypopygium (Figs 1 – 2). Tergite IX with 9 – 24 setae, 60 – 110 µm long. Laterosternite IX with 3 – 11 setae. Transverse sternapodeme 33 – 73 µm wide. Aedeagal lobe weakly sclerotized, apical part with many small spinules. Gonocoxite 155 – 218 µm long. Gonostylus 68 – 103 µm long and inflated, with wide median part and short narrow apical part, ending in short megaseta. HR 2.04 – 2.76.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E557039FF58F8E4FE829082.taxon	distribution	Distribution. The species is known only from mountain areas in Sichuan and Yunnan Provinces in Oriental China.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E57703FFF58F9F8FAFD91EB.taxon	description	(Figs 7 – 10)	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E57703FFF58F9F8FAFD91EB.taxon	materials_examined	Type material. Holotype male, CHINA: Sichuan Province, Kangding County, Wasi Gully, 15. vi. 1996, light trap, X. Wang (BDN).	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E57703FFF58F9F8FAFD91EB.taxon	etymology	Etymology. From Latin ambigua, meaning ambiguous or obscure, as the systematic position of this species in Boreoheptagyia is unclear. Diagnostic characters. B. ambigua sp. n. is described based on a single male only and is closely related to B. xinglongiensis sp. n. Both species have dorsocentrals only in single anterior group. B. ambigua sp. n., however, can easily be separated from the latter by the shape of the gonostylus with wide median part and a short narrow apical part, and by the aedeagal lobe having some small teeth on the inner edge. Male (n = 1) Total length 2.4 mm. Wing length 1.64 mm. Total length / wing length 1.45. Coloration. Head, thorax and abdomen brown. Antenna and legs yellowish; femur and tibia of all legs with rings, basal and apical parts light brown, median part white (Fig. 9). Head. Frontal tubercles damaged. Temporal setae includes 25 frontals, 12 outer verticals, no inner verticals. Clypeus with 9 setae. Antenna with 13 flagellomeres, antennal plume reduced, ultimate flagellomere with some large white spots (Fig. 10); pedicel with 3 setae. Lengths (in µm) of flagellomeres 2 – 13: 24, 24, 40, 46, 50, 46, 42, 42, 40, 40, 32, 72. Number of setae on flagellomeres 2 – 13 as: 3, 3, 5, 5, 4, 4, 4, 3, 3, 4, 4, 3. Subapical seta of terminal flagellomere 23 µm long. AR 0.15. Lengths (in µm) of palpomeres 1 – 5: 30, 44, 72, 86, 160. Head width / palp length 1.16. Thorax. Antepronotum with 6 lateral setae. Acrostichals 30; dorsocentrals 5, in single anterior group; prealars 12; scutellars 21. Longest and shortest acrostichals 76 µm and 27 µm long. Wing. Costal extension 80 µm long. Anal lobe well developed, fully fringed. Alula without setae. Squama with 20 setae. R with 13 setae, R 1 with 2 setae, R 4 + 5 with 2 setae, R 2 + 3 reduced. RM / MCu 2.67. Legs. Spur of foretibia 32 µm long; spurs of midtibia 32 µm and 34 µm long; of hind tibia 30 µm and 48 µm long. Hind tibial comb with 13 setae. Mid ta 1 with 4 pseudospurs, ta 2 with 2 pseudospurs, 23 µm long; hind ta 1 with 4 pseudospurs, ta 2 with 2 pseudospurs, 27 µm long. Lengths and proportions of legs as in Table 2. Hypopygium (Figs 7 – 8). Tergite IX with 11 setae, 104 µm long. Laterosternite IX with 4 setae. Transverse sternapodeme 56 µm wide. Aedeagal lobe weakly sclerotized, apical part smooth, inner edge with some small teeth (Fig. 8). Gonocoxite 196 µm long. Gonostylus 94 µm long, with wide median part and short narrow apical part, ending in short megaseta. HR 2.09.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E57703FFF58F9F8FAFD91EB.taxon	distribution	Distribution. The species is known only from the type locality in Sichuan Province in Oriental China.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E51703DFF58FB23FCA994C1.taxon	description	(Figs 14 – 20)	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E51703DFF58FB23FCA994C1.taxon	materials_examined	Material examined. JAPAN: Honshu, Kyoto Prefecture, Kibune, 2 males, 16. x. 1934, M. Tokunaga (KPU); Gifu Prefecture, Kamioka, Futatsuya, Takahara River, 1 male, 28. v. 2001, K. Endo; Niigata Prefecture, Yuzawa, Tsuchitaru, Kedo-sawa, 1 male, 26. vi. 2001, K. Endo; Hokkaido, Shintoku, Yutomuraushi River, 700 m a. s. l., Malaise trap, 2 males, 11 – 21. vi. 1999, K. Endo; Kiushu, Fukuoka Prefecture, Hoshino, Kumado River, 1 male, 26. iii. 2002, K. Endo. Male (n = 3) Total length 2.27 – 2.50 mm. Wing length 1.7 – 2.16 mm. Total length / wing length 1.16 – 1.27. Coloration. Body dark brown. Legs brown, tibiae of all legs white medially. Head. Frontal tubercles 24 – 28 µm long, with 2 – 3 setae basally. Temporal setae includes 7 – 10 frontals, 7 – 9 inner verticals, 8 – 18 outer verticals, and 4 – 7 postorbitals. Clypeus with 10 – 14 setae. Antenna with 13 flagellomeres. AR 0.11 – 0.16. Lengths (in µm) of palpomeres 1 – 5: 28 – 32, 36 – 46, 72 – 86, 96 – 100,148 – 160. Head width / palp length 1.16 – 1.20. Thorax. Antepronotum with 7 lateral setae. Acrostichals 30 – 37, dorsocentrals 17 – 23 in anterior and posterior groups, prealars 17 – 28, supraalars 1 – 2. Scutellum with 61 – 70 setae. Wing. Costal extension 65 – 73.8 µm long. Anal lobe well developed, fully fringed. Alula without setae. Squama with 27 – 48 setae. R and R 1 with 8 – 11 setae, R 4 + 5 with 2 – 4 setae, R 2 + 3 reduced and visible only in basal part. RM / MCu 2.5 – 2.6. Legs. Spur of foretibia 32 – 36 µm long, spurs of midtibia 28 – 36 µm and 32 – 36 µm long, of hind tibia 32 – 38 µm and 48 – 56 µm long. Hind tibial comb with 11 – 12 setae. Lengths and proportions of legs as in Table 3. Hypopygium (Figs 15 – 20). Tergite IX with concave posterior margin and 29 – 36 setae. Laterosternite IX with 3 – 4 setae. Transverse sternapodeme 60 – 64 µm wide. Aedeagal lobe sharply triangular, phallapodeme not clearly discernable, sometimes visible as narrow appendage (Figs 19 – 20). Gonocoxite 180 µm long; basal lobe widely triangular with rounded apex and 1 – 2 setae. Gonostylus 88 µm long, wrinkled, slightly curved, ending in short megaseta.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E51703DFF58FB23FCA994C1.taxon	discussion	Remarks. We redescribe the male of B. brevitarsis as K. Endo compared specimens from the Kolyma River Basin of the Magadan Region in Russia and from Hokkaido of Japan with the type material of B. brevitarsis deposited in Kyushu University, Japan and concluded that all males from Kolyma River and some males from Hokkaido clearly differs from the type material in several features of the thorax and hypopygium. We therefore describe these specimens as a separate species, B. sasai sp. n., below.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E51703DFF58FB23FCA994C1.taxon	distribution	Distribution. The species is now for certain only known from Japan. All other records based on larvae from China and the Russian Far East should be revised.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E537032FF58FBEBFA179727.taxon	description	(Fig. 21)	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E537032FF58FBEBFA179727.taxon	discussion	Remarks. The male can be separated from all other known species of Boreoheptagyia by the extremely short antenna with 5 flagellomeres and by the high AR. The hypopygium of the holotype is squeezed and additional material is needed to study this interesting species in detail.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E537032FF58FBEBFA179727.taxon	distribution	Distribution. The species is known only from the type locality, the Kurobe River Basin in Honshu, Japan.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5C7032FF58FD64FE3792A3.taxon	description	(Figs 22 – 25)	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5C7032FF58FD64FE3792A3.taxon	materials_examined	Type material. Holotype male, NEPAL: Langtang Region, pond near Khyimjung Glacier, 4.175 m a. s. l., 4. viii. 1999, K. Endo (LEOU).	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5C7032FF58FD64FE3792A3.taxon	etymology	Etymology. The name refers to the type country. Diagnostic characters. The male is well separated from other species of the genus by the short antenna with 8 flagellomeres, presence of some setae on alula, absence of megaseta on the gonostylus, and by the shape of the gonostylus and the sternapodeme. Male (n = 1) Total length 2.9 mm. Wing length 2.49 mm. Total length / wing length 1.17. Coloration. Brown with grayish abdomen. Legs brown, without light spots or rings. Head. Temporal setae includes 43 frontals and 16 outer verticals, no inner verticals. Clypeus with 20 setae. Antenna with 8 flagellomeres, antennal plume reduced (Fig. 22), ultimate flagellomere with a few small white spots, pedicel with 5 setae. Lengths (in µm) of flagellomeres 2 – 8: 29, 32, 34, 34, 32, 34, 122. Number of setae on flagellomeres 2 – 8 as: 6, 5, 6, 4, 4, 3, 5. Subapical seta of terminal flagellomere 23 µm long. AR 0.15. Lengths (in µm) of palpomeres 1 – 5: 27, 61, 103, 108, 186. Head width / palp length 0.92. Thorax. Antepronotum with 9 lateral setae. Acrostichals 40; dorsocentrals 14 in 2 groups, anterior group with 8 setae, posterior group with 6 setae; prealars 21, in 2 groups, anterior group with 6 setae in single row, posterior group with 15 setae in 3 rows; supraalar 1 (Fig. 23). Scutellum with 86 setae. Longest and shortest acrostichals 57 µm and 23 µm long. Wing 0.87 mm wide. Costal extension 64 µm long. Anal lobe well developed, fully fringed. Alula with 2 – 4 setae (Fig. 24). Squama with 45 setae. R with 20 setae, R 1 with 6 setae, R 4 + 5 with 6 setae, R 2 + 3 clearly visible only in basal half. RM / MCu 2.29. Legs. Spur of foretibia 38 µm long; spurs of midtibia 38 µm and 42 µm long; of hind tibia 46 µm and 61 µm long. Hind tibial comb with 11 setae. Mid ta 1 with 8 pseudospurs, ta 2 with 2 pseudospurs, 19 µm long; hind ta 1 with 10 pseudospurs, ta 2 with 2 pseudospurs, 15 µm and 23 µm long. Lengths and proportions of legs as in Table 4. Hypopygium (Fig. 25). Tergite IX with 16 setae, 89 µm long. Laterosternite IX with 10 setae. Transverse sternapodeme 34 µm wide. Aedeagal lobe weakly sclerotized, apical part smooth. Gonocoxite 201 µm long. Gonostylus 84 µm long, narrow, slightly curved, without megaseta. HR 2.41.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5C7032FF58FD64FE3792A3.taxon	distribution	Distribution. The species is known only from the type locality in the high mountains of the Khyimjung Glacier Region in Nepal.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5D7033FF58F99DFEE99290.taxon	distribution	Distribution. The species is known only from the type locality, a high mountain area not far from Mount Everest in Nepal.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5E7030FF58FF51FE2F92D6.taxon	description	(Figs 26 – 33)	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5E7030FF58FF51FE2F92D6.taxon	materials_examined	Type material. Holotype male, RUSSIA: North-Eastern Russia, Ten’kinsky County, Magadan Region, Kolyma River Basin, Vlastnyi Peak not far from Sibit-Tyellah Village, Olen’ Stream, 1.100 – 1.300 m a. s. l., 11 – 27. vii. 1977, E. Makarchenko & S. Kocharina (IBSS FEBRAS). Paratypes: 11 males, 4 females as holotype. 2 males, JAPAN: Hokkaido, Shintoku, Yutomuraushi River, 700 m a. s. l., Malaise trap, 11 – 21. vi. 1999, K. Endo; 1 male, same as previous except 21. vi – 2. vii. 1999, Malaise trap, K. Endo; 1 male, 1 female, Hokkaido, Obihiro, Opirineppu River, 620 m a. s. l., 5 – 15. vi. 2000, Malaise trap, K. Endo; 2 males, same as previous except 13. ix – 1. x. 1999, K. Endo; 1 male, same as previous except 5 – 15. vi. 2000, Malaise trap, K. Endo.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5E7030FF58FF51FE2F92D6.taxon	etymology	Etymology. Named in honor and memory of the famous Japanese chironomidologist and colleague Prof. Manabu Sasa. Diagnostic characters. The male is well separated from other species of the genus by the presence of setae on mesanepisternum II and preepisternum, and by the field of small spinules distal of transverse sternapodeme (Figs 30, 32). Male (n = 9) Total length 2.4 – 2.7 mm. Wing length 1.61 – 2.13 mm. Total length / wing length 1.17 – 1.29. Coloration. Head, thorax and legs brown to dark brown, abdomen grayish to yellowish. Head. Temporal setae includes 6 – 7 postorbitals, 30 – 31 outer verticals, and 26 – 31 inner verticals. Clypeus with 11 – 12 setae. Antenna with 13 flagellomeres, antennal plume reduced, pedicel with 4 setae. Lengths (in µm) of flagellomeres 2 – 13: 17 – 29, 25 – 34, 31 – 42, 35 – 46, 39 – 42, 34 – 46, 36 – 46, 38 – 55, 38 – 42, 38 – 46, 34 – 42, 88 – 101. Number of setae on flagellomeres 2 – 13 as: 4 – 5, 6, 8, 9 – 10, 8 – 9, 9 – 10, 8, 8 – 9, 8, 8, 7 – 8, 4. Subapical seta of terminal flagellomere 11 – 15 µm long. AR 0.11 – 0.20. Lengths (in µm) of palpomeres 1 – 5: 27 – 30, 63 – 80, 101 – 109, 92 – 118, 160 – 193. Antennal length / palp length 1.35 – 1.40. Head width / palp length 1.04 – 1.16. Thorax. Antepronotum with 6 – 11 lateral setae. Acrostichals 33 – 36; dorsocentrals 24 – 26 in anterior and posterior groups; prealars 19 – 26 in 2 rows; supraalars 2; preepisternals 6 – 13; median anepisternum II with 4 – 6 setae. Scutellum with 58 – 62 setae. Longest and shortest acrostichals 36 – 42 µm and 11 – 21 µm long. Wing 0.52 – 0.62 mm wide. Costal extension 64 – 80 µm long. Anal lobe well developed, fully fringed. Alula without setae. Squama with 32 – 48 setae. R with 14 setae, R 1 with 13 – 17 setae, R 4 + 5 with 3 – 7 setae, R 2 + 3 usually reduced, often only basal part visible. RM / MCu 1.80 – 2.25. Legs. Spur of foretibia 34 – 42 µm long; spurs of midtibia 34 – 42 µm and 34 – 38 µm long, of hind tibia 34 – 55 µm and 34 – 38 µm long. Hind tibial comb with 10 – 12 setae. Mid ta 1 with 4 – 7 pseudospurs, ta 2 with 1 pseudospur, 23 – 27 µm long; hind ta 1 with 10 – 12 pseudospurs, ta 2 with 1 pseudospur, 23 – 27 µm long. Lengths and proportions of legs of specimens from Kolyma River Basin and from Hokkaido as in Tables 5 and 6, respectively. Hypopygium (Figs 28 – 33). Tergite IX with 20 – 27 setae. Laterosternite IX with 12 – 17 setae. Transverse sternapodeme 46 – 60 µm wide. Aedeagal lobe long, narrow; phallapodeme narrow, well visible. With field of small spinules distal of transverse sternapodeme. Gonocoxite 201 – 205 µm long; basal lobe rounded, with 1 – 3 short setae. Gonostylus 95 – 103 µm long, with wide median part and short narrow apical part, ending in short megaseta. HR 2.20 – 2.73.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5E7030FF58FF51FE2F92D6.taxon	discussion	Remarks. The males from Kolyma River Basin and from Hokkaido differ in the lengths and proportions of the leg segments (see Tables 5 & 6), while in other characters both populations are very similar or identical.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5E7030FF58FF51FE2F92D6.taxon	distribution	Distribution. The species is known from localities in the Kolyma River Basin in Northeastern Russia and from Hokkaido, Japan.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5F7036FF58F93AFBA194AF.taxon	discussion	Remarks. The description and figure of the male hypopygium of B. similis by Chaudhuri and Ghosh (1981) are not informative. A redescription based on the holotype and additional material is needed to enable us to compare this species with other members of Boreoheptagyia.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5F7036FF58F93AFBA194AF.taxon	distribution	Distribution. The species is known only from the type locality in Bhutan.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E587037FF58F90CFF6E9494.taxon	description	(Figs 34 – 36)	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E587037FF58F90CFF6E9494.taxon	discussion	Remarks. The male of B. tibetica can be separated from all other Boreoheptagyia species with femalelike antenna by the features mentioned in the key.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E587037FF58F90CFF6E9494.taxon	distribution	Distribution. The species is known only from a high mountain area (1.700 m a. s. l.) in Tibet Province, China.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E597034FF58F989FEC09491.taxon	description	(Figs 37 – 39)	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E597034FF58F989FEC09491.taxon	discussion	Remarks. The male of B. unica can be separated from all known species of Boreoheptagyia with femalelike antenna by the shape of the aedeagal lobe and by the gonostylus being attaches to the gonocoxite twothirds the distance from the apex, and having an apical lobe and two megasetae.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E597034FF58F989FEC09491.taxon	distribution	Distribution. The species is known only from a mountain area (1.450 m a. s. l.) in Nikko National Park in Honshu, Japan.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5A7035FF58FD89FAFD9724.taxon	description	(Figs 11 – 13)	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5A7035FF58FD89FAFD9724.taxon	materials_examined	Type material. Holotype male, CHINA: Sichuan Province, Baoxing County, Xinglong West River, 19. vi. 1996, light trap, X. Wang (BDN). Paratypes: 2 males, as holotype.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5A7035FF58FD89FAFD9724.taxon	etymology	Etymology. Named after the type locality. Diagnostic characters. See key. Male (n = 3) Total length 2.8 – 3.1 mm. Wing length 1.76 – 1.92 mm. Total length / wing length 1.52 – 1.73. Coloration. Head, thorax and abdomen brown. Antenna yellowish. Femur and tibia of all legs with rings, basal and apical parts dark brown, median part white. Head. Frontal tubercles triangular, 18 – 20 µm long, with 1 seta, 50 – 65 µm long. Temporal setae includes 13 – 20 frontals, 6 – 12 inner verticals, and 9 – 14 outer verticals. Clypeus with 11 – 16 setae. Antenna with 13 flagellomeres, flagellomere 12 and 13 not well separated; flagellomere 13 with few median white spots; pedicel with 3 or 4 setae. Lengths (in µm) of flagellomeres 2 – 13: 22 – 26, 26 – 32, 34 – 40, 40 – 46, 40 – 42, 40 – 42, 36 – 40, 38 – 40, 36, 30 – 38, 24, 60. Number of setae of flagellomeres 2 – 13 as: 3 – 4, 3 – 4, 3 – 5, 3 – 5, 3 – 4, 3 – 4, 3 – 4, 3 – 4, 3 – 4, 3 – 4, 3 – 4, 1 – 2. Subapical seta of terminal flagellomere 8 – 28 µm long. AR 0.13 – 0.14. Lengths (in µm) of palpomeres 1 – 5: 30 – 32, 40 – 44, 92 – 96, 86 – 112, 144 – 184. Head width / palp length 1.01 – 1.14. Thorax. Antepronotum with 3 – 6 lateral setae. Acrostichals 20 – 24, dorsocentrals 4 – 6 in anterior group, prealars 1 – 4. Scutellum with 16 – 30 setae. Longest and shortest acrostichals 80 – 114 µm and 38 – 49 µm long. Wing 0.53 – 0.57 mm wide. Costal extension 48 µm long. Anal lobe well developed, fully fringed. Alula without setae. Squama with 14 – 19 setae. R with 14 – 17 setae, R 1 with 1 – 4 setae, R 4 + 5 with 4 – 7 setae, R 2 + 3 very short, easily visible only in basal part; RM / MCu 2.50 – 2.75. Legs. Spur of foretibia 38 – 44 µm long; spurs of midtibia 42 – 44 µm and 44 – 48 µm long; of hind tibia 42 – 46 µm and 53 – 60 µm long. Hind tibial comb with 13 – 15 setae. Mid ta 1 with 4 – 6 pseudospurs, hind ta 1 with 7 – 8 pseudospurs. Lengths and proportions of legs as in Table 7. Hypopygium (Figs 11 – 12). Tergite IX with 18 – 27 setae, 100 – 120 µm long. Laterosternite IX with 0 – 1 seta. Transverse sternapodeme 48 – 52 µm long. Aedeagal lobe weakly sclerotized, distal part without teeth. Gonocoxite 200 – 248 µm long; basal lobe rounded to triangular, with 5 – 6 short setae. Gonostylus 88 – 110 µm long, curved, with single short megaseta. HR 2.00 – 2.41.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
03926A1E9E5A7035FF58FD89FAFD9724.taxon	distribution	Distribution. The species is known only from the type locality in Sichuan Province in Oriental China.	en	Makarchenko, Eugenyi A., Endo, Kazuo, Wu, Jingyang, Wang, Xinhua (2008): A review of Boreoheptagyia Brundin, 1966 (Chironomidae: Diamesinae) from East Asia and bordering territories, with the description of five new species. Zootaxa 1817 (1): 1-17, DOI: 10.11646/zootaxa.1817.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1817.1.1
