taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
039087E0FFCB4724FF6D5967FCDDFD01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826207/files/figure.png	https://doi.org/10.5281/zenodo.3826207	FIGURE 1. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Chaetoleptophyllum flexum Golovatch, 1979, male from Khulo, Georgia. B: Kubaniulus gracilis Lohmander, 1936, male from near Gelenjik, Krasnodar Prov. C: Kubaniulus lativelatus sp. nov., male paratype from Pasture Abago, Krasnodar Prov. Scale bar: 1 mm.	FIGURE 1. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Chaetoleptophyllum flexum Golovatch, 1979, male from Khulo, Georgia. B: Kubaniulus gracilis Lohmander, 1936, male from near Gelenjik, Krasnodar Prov. C: Kubaniulus lativelatus sp. nov., male paratype from Pasture Abago, Krasnodar Prov. Scale bar: 1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFCB4724FF6D5967FCDDFD01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826209/files/figure.png	https://doi.org/10.5281/zenodo.3826209	FIGURE 2. Chaetoleptophyllum flexum Golovatch, 1979, male from Sataplia Nature Reserve, Georgia. A–C: anterior, middle and posterior parts of body, respectively, lateral views. D–F: same, respectively, ventral views. G–I: same, respectively, dorsal views. J: midbody ring, frontal view. K: seta and limbus on a midbody ring, dorsal view. L: claw on leg 2, lateral view. M: head, ventral view. Abbreviations: ep, epiproct; hp, hypoproct; lg, lamella lingualis of gnathochilarium; mz, metazona; pz, prozona; sg, stipes of gnathoсhilarium. Scale bars: 0.2 mm (A, B, G), 0.1 mm (C–F, H–J, M) or 0.01 mm (K, L).	FIGURE 2. Chaetoleptophyllum flexum Golovatch, 1979, male from Sataplia Nature Reserve, Georgia. A–C: anterior, middle and posterior parts of body, respectively, lateral views. D–F: same, respectively, ventral views. G–I: same, respectively, dorsal views. J: midbody ring, frontal view. K: seta and limbus on a midbody ring, dorsal view. L: claw on leg 2, lateral view. M: head, ventral view. Abbreviations: ep, epiproct; hp, hypoproct; lg, lamella lingualis of gnathochilarium; mz, metazona; pz, prozona; sg, stipes of gnathoсhilarium. Scale bars: 0.2 mm (A, B, G), 0.1 mm (C–F, H–J, M) or 0.01 mm (K, L).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFCB4724FF6D5967FCDDFD01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826211/files/figure.png	https://doi.org/10.5281/zenodo.3826211	FIGURE 3. Chaetoleptophyllum flexum Golovatch, 1979, male from near Tkibuli, Georgia. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	FIGURE 3. Chaetoleptophyllum flexum Golovatch, 1979, male from near Tkibuli, Georgia. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFCB4724FF6D5967FCDDFD01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826215/files/figure.png	https://doi.org/10.5281/zenodo.3826215	FIGURE 4. Chaetoleptophyllum flexum Golovatch, 1979, male from Sataplia Nature Reserve, Georgia. A: segment 7, ventral view. B: leg-pair 1, caudal view. C & D: gonopods, frontoventral and ventrocaudal views, respectively. Abbreviations: ms, mesomere; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.05 mm (A, B, C) or 0.02 mm (D).	FIGURE 4. Chaetoleptophyllum flexum Golovatch, 1979, male from Sataplia Nature Reserve, Georgia. A: segment 7, ventral view. B: leg-pair 1, caudal view. C & D: gonopods, frontoventral and ventrocaudal views, respectively. Abbreviations: ms, mesomere; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.05 mm (A, B, C) or 0.02 mm (D).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFCB4724FF6D5967FCDDFD01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826217/files/figure.png	https://doi.org/10.5281/zenodo.3826217	FIGURE 5. Chaetoleptophyllum flexum Golovatch, 1979, male from near Tkibuli (A) and male paratype (B–D), Georgia. A: gonopod, mesal view. B: promere of gonopod, caudal view. C: mesomere of gonopod, mesal view. D: gonopod, mesal view. B–D: after Golovatch (1979). Abbreviations: ms, mesomere; op, opisthomere; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.2 mm (A) or 0.1 mm (B–D).	FIGURE 5. Chaetoleptophyllum flexum Golovatch, 1979, male from near Tkibuli (A) and male paratype (B–D), Georgia. A: gonopod, mesal view. B: promere of gonopod, caudal view. C: mesomere of gonopod, mesal view. D: gonopod, mesal view. B–D: after Golovatch (1979). Abbreviations: ms, mesomere; op, opisthomere; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.2 mm (A) or 0.1 mm (B–D).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFCB4724FF6D5967FCDDFD01.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826299/files/figure.png	https://doi.org/10.5281/zenodo.3826299	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC4472BFF6D5C19FC55F807.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826207/files/figure.png	https://doi.org/10.5281/zenodo.3826207	FIGURE 1. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Chaetoleptophyllum flexum Golovatch, 1979, male from Khulo, Georgia. B: Kubaniulus gracilis Lohmander, 1936, male from near Gelenjik, Krasnodar Prov. C: Kubaniulus lativelatus sp. nov., male paratype from Pasture Abago, Krasnodar Prov. Scale bar: 1 mm.	FIGURE 1. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Chaetoleptophyllum flexum Golovatch, 1979, male from Khulo, Georgia. B: Kubaniulus gracilis Lohmander, 1936, male from near Gelenjik, Krasnodar Prov. C: Kubaniulus lativelatus sp. nov., male paratype from Pasture Abago, Krasnodar Prov. Scale bar: 1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC4472BFF6D5C19FC55F807.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826219/files/figure.png	https://doi.org/10.5281/zenodo.3826219	FIGURE 6. Kubaniulus gracilis Lohmander, 1936, male from pasture Abago, Krasnodar Prov. A: anterior part of body, ventrolateral view. B: anterior part of body, lateral view, C: middle part of body, lateral view. D: midbody ring, frontal view. E: head, ventral view. F: leg 2, frontolateral view. G: leg-pair 1, caudal view. Abbreviations: lg, lamella lingualis of gnathochilarium; mz, metazona; pz, prozona; sg, stipes of gnathochilarium Scale bars: 0.1 mm (A–E) or 0.03 mm (F–G).	FIGURE 6. Kubaniulus gracilis Lohmander, 1936, male from pasture Abago, Krasnodar Prov. A: anterior part of body, ventrolateral view. B: anterior part of body, lateral view, C: middle part of body, lateral view. D: midbody ring, frontal view. E: head, ventral view. F: leg 2, frontolateral view. G: leg-pair 1, caudal view. Abbreviations: lg, lamella lingualis of gnathochilarium; mz, metazona; pz, prozona; sg, stipes of gnathochilarium Scale bars: 0.1 mm (A–E) or 0.03 mm (F–G).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC4472BFF6D5C19FC55F807.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826221/files/figure.png	https://doi.org/10.5281/zenodo.3826221	FIGURE 7. Kubaniulus gracilis Lohmander, 1936, male from pasture Abago, Krasnodar Prov. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	FIGURE 7. Kubaniulus gracilis Lohmander, 1936, male from pasture Abago, Krasnodar Prov. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC4472BFF6D5C19FC55F807.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5917992/files/figure.png	https://doi.org/10.5281/zenodo.5917992	FIGURE 8. Kubaniulus gracilis Lohmander, 1936, male from pasture Abago, Krasnodar Prov. A: segment 7, ventrolateral view. B: left gonopod, lateral view. C: right gonopod, caudal view. Abbreviations: lp, lamella of promere; ms, mesomere; op, opisthomere; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.03 mm.	FIGURE 8. Kubaniulus gracilis Lohmander, 1936, male from pasture Abago, Krasnodar Prov. A: segment 7, ventrolateral view. B: left gonopod, lateral view. C: right gonopod, caudal view. Abbreviations: lp, lamella of promere; ms, mesomere; op, opisthomere; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.03 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC4472BFF6D5C19FC55F807.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826227/files/figure.png	https://doi.org/10.5281/zenodo.3826227	FIGURE 9. Kubaniulus gracilis Lohmander, 1936, male holotype, after Lohmander (1936). A: promere of gonopod, caudal view. B: right gonopod, mesal view. Abbreviations: lp, lamella of promere; ms, mesomere; op, opisthomere; pr, promere; sl, solenomere; ve, velum. Drawn not to scale.	FIGURE 9. Kubaniulus gracilis Lohmander, 1936, male holotype, after Lohmander (1936). A: promere of gonopod, caudal view. B: right gonopod, mesal view. Abbreviations: lp, lamella of promere; ms, mesomere; op, opisthomere; pr, promere; sl, solenomere; ve, velum. Drawn not to scale.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC4472BFF6D5C19FC55F807.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826299/files/figure.png	https://doi.org/10.5281/zenodo.3826299	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC3472CFF6D5E2FFDF9FA56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826207/files/figure.png	https://doi.org/10.5281/zenodo.3826207	FIGURE 1. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Chaetoleptophyllum flexum Golovatch, 1979, male from Khulo, Georgia. B: Kubaniulus gracilis Lohmander, 1936, male from near Gelenjik, Krasnodar Prov. C: Kubaniulus lativelatus sp. nov., male paratype from Pasture Abago, Krasnodar Prov. Scale bar: 1 mm.	FIGURE 1. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Chaetoleptophyllum flexum Golovatch, 1979, male from Khulo, Georgia. B: Kubaniulus gracilis Lohmander, 1936, male from near Gelenjik, Krasnodar Prov. C: Kubaniulus lativelatus sp. nov., male paratype from Pasture Abago, Krasnodar Prov. Scale bar: 1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC3472CFF6D5E2FFDF9FA56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826229/files/figure.png	https://doi.org/10.5281/zenodo.3826229	FIGURE 10. Kubaniulus lativelatus sp. nov., male paratype from near Khosta, Krasnodar Prov. A–C: anterior, middle and posterior parts of body, respectively, lateral view. D: head, ventral view. E: posterior part of body, ventral view. F: midbody ring, frontal view; G: leg 2, frontal view. H & I: leg-pair 1, caudolateral and frontoventral views, respectively. J: segment 7 and leg-pair 7, lateral view. Abbreviations: ep, epiproct; hp, hypoproct; lg, lamella lingualis of gnathochilarium; mz, metazona; pr, promere; pz, prozona; sg, stipes of gnathoсhilarium; sl, solenomere; ve, velum. Scale bars: 0.2 mm (A), 0.1 mm (B–F) or 0.05 mm (G–J).	FIGURE 10. Kubaniulus lativelatus sp. nov., male paratype from near Khosta, Krasnodar Prov. A–C: anterior, middle and posterior parts of body, respectively, lateral view. D: head, ventral view. E: posterior part of body, ventral view. F: midbody ring, frontal view; G: leg 2, frontal view. H & I: leg-pair 1, caudolateral and frontoventral views, respectively. J: segment 7 and leg-pair 7, lateral view. Abbreviations: ep, epiproct; hp, hypoproct; lg, lamella lingualis of gnathochilarium; mz, metazona; pr, promere; pz, prozona; sg, stipes of gnathoсhilarium; sl, solenomere; ve, velum. Scale bars: 0.2 mm (A), 0.1 mm (B–F) or 0.05 mm (G–J).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC3472CFF6D5E2FFDF9FA56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826231/files/figure.png	https://doi.org/10.5281/zenodo.3826231	FIGURE 11. Kubaniulus lativelatus sp. nov., female from near Khosta, Krasnodar Prov. A: head, ventral view B: leg-pair 2 and gonopores, ventral view. Abbreviations: lg, lamella lingualis of gnathochilarium; sg, stipes of gnathochilarium. Scale bars: 0.1 mm.	FIGURE 11. Kubaniulus lativelatus sp. nov., female from near Khosta, Krasnodar Prov. A: head, ventral view B: leg-pair 2 and gonopores, ventral view. Abbreviations: lg, lamella lingualis of gnathochilarium; sg, stipes of gnathochilarium. Scale bars: 0.1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC3472CFF6D5E2FFDF9FA56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826233/files/figure.png	https://doi.org/10.5281/zenodo.3826233	FIGURE 12. Kubaniulus lativelatus sp. nov., male paratype from near Khosta, Krasnodar Prov. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2 and penes, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	FIGURE 12. Kubaniulus lativelatus sp. nov., male paratype from near Khosta, Krasnodar Prov. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2 and penes, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC3472CFF6D5E2FFDF9FA56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826235/files/figure.png	https://doi.org/10.5281/zenodo.3826235	FIGURE 13. Kubaniulus lativelatus sp. nov., male paratype near Khosta, Krasnodar Prov. A & B: both gonopods, ventral and ventrolateral views, respectively. C: left gonopod, mesal view. D: right gonopod, lateral view (solenomere is broken). Abbreviations: ms, mesomere; op, opisthomere; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.05 mm.	FIGURE 13. Kubaniulus lativelatus sp. nov., male paratype near Khosta, Krasnodar Prov. A & B: both gonopods, ventral and ventrolateral views, respectively. C: left gonopod, mesal view. D: right gonopod, lateral view (solenomere is broken). Abbreviations: ms, mesomere; op, opisthomere; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.05 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC3472CFF6D5E2FFDF9FA56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826237/files/figure.png	https://doi.org/10.5281/zenodo.3826237	FIGURE 14. Kubaniulus lativelatus sp. nov., male paratype from near Khosta, Krasnodar Prov. A: promere of gonopod, caudal view. B: gonopod, mesal view. Abbreviations: op, opisthomere; pr, promere; ms, mesomere; sl, solenomere; ve, velum. Scale bar: 0.2 mm.	FIGURE 14. Kubaniulus lativelatus sp. nov., male paratype from near Khosta, Krasnodar Prov. A: promere of gonopod, caudal view. B: gonopod, mesal view. Abbreviations: op, opisthomere; pr, promere; ms, mesomere; sl, solenomere; ve, velum. Scale bar: 0.2 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFC3472CFF6D5E2FFDF9FA56.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826299/files/figure.png	https://doi.org/10.5281/zenodo.3826299	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFDC4733FF6D5FA2FB39FA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826239/files/figure.png	https://doi.org/10.5281/zenodo.3826239	FIGURE 15. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Leptoiulus hastatus Lohmander, 1932, male from near Shnokh, Armenia. B: Leptoiulus tanymorphus (Attems, 1901), male from Ajikent, Azerbaijan. C: Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. D: Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. Scale bar: 1 mm.	FIGURE 15. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Leptoiulus hastatus Lohmander, 1932, male from near Shnokh, Armenia. B: Leptoiulus tanymorphus (Attems, 1901), male from Ajikent, Azerbaijan. C: Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. D: Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. Scale bar: 1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFDC4733FF6D5FA2FB39FA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826241/files/figure.png	https://doi.org/10.5281/zenodo.3826241	FIGURE 16. Leptoiulus hastatus Lohmander, 1932, male from near Lichk, Armenia. A: anterior part of body, lateral view. B: middle part of body, lateral view. C: posterior part of body, dorsolateral view. D: anterior part of body, ventral view. E: midbody ring, frontal view. F: head, ventral view. Abbreviations: ep, epiproct; hp, hypoproct; lg, lamella lingualis of gnathochilarium; ls, labral seta; mz, metazona; pz, prozona; sg, stipes of gnathoсhilarium. Scale bars: 0.2 mm (A–C, E, F) or 0.1 (D).	FIGURE 16. Leptoiulus hastatus Lohmander, 1932, male from near Lichk, Armenia. A: anterior part of body, lateral view. B: middle part of body, lateral view. C: posterior part of body, dorsolateral view. D: anterior part of body, ventral view. E: midbody ring, frontal view. F: head, ventral view. Abbreviations: ep, epiproct; hp, hypoproct; lg, lamella lingualis of gnathochilarium; ls, labral seta; mz, metazona; pz, prozona; sg, stipes of gnathoсhilarium. Scale bars: 0.2 mm (A–C, E, F) or 0.1 (D).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFDC4733FF6D5FA2FB39FA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826243/files/figure.png	https://doi.org/10.5281/zenodo.3826243	FIGURE 17. Leptoiulus hastatus Lohmander, 1932. A-C: male from Toðanalý, Azerbaijan. D-F: Morph B, male from Ceyrankeçməz, Qobustan, Azerbaijan. A: leg pair 1, anterolateral view. B: leg pair 1, anterior view. C: gnathochilarium, ventral view. Abbreviations: lg, lamella lingualis of gnathochilarium; sg, stipes of gnathoсhilarium. Scale bars: 0.1 mm (A–C, E, F) or 0.05 mm (D).	FIGURE 17. Leptoiulus hastatus Lohmander, 1932. A-C: male from Toðanalý, Azerbaijan. D-F: Morph B, male from Ceyrankeçməz, Qobustan, Azerbaijan. A: leg pair 1, anterolateral view. B: leg pair 1, anterior view. C: gnathochilarium, ventral view. Abbreviations: lg, lamella lingualis of gnathochilarium; sg, stipes of gnathoсhilarium. Scale bars: 0.1 mm (A–C, E, F) or 0.05 mm (D).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFDC4733FF6D5FA2FB39FA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826245/files/figure.png	https://doi.org/10.5281/zenodo.3826245	FIGURE 18. Leptoiulus hastatus Lohmander, 1932, male from Aganos, Azerbaijan. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	FIGURE 18. Leptoiulus hastatus Lohmander, 1932, male from Aganos, Azerbaijan. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFDC4733FF6D5FA2FB39FA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826249/files/figure.png	https://doi.org/10.5281/zenodo.3826249	FIGURE 19. Leptoiulus hastatus Lohmander, 1932, male from near Lichk, Armenia. A: segment 7, ventral view. B-C: gonopod, frontoventral and mesal views, respectively. Abbreviations: fl, flagellum; lp, lamella of promere; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.5 mm (A), 0.2 mm (B, C).	FIGURE 19. Leptoiulus hastatus Lohmander, 1932, male from near Lichk, Armenia. A: segment 7, ventral view. B-C: gonopod, frontoventral and mesal views, respectively. Abbreviations: fl, flagellum; lp, lamella of promere; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.5 mm (A), 0.2 mm (B, C).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFDC4733FF6D5FA2FB39FA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826251/files/figure.png	https://doi.org/10.5281/zenodo.3826251	FIGURE 20. Leptoiulus hastatus Lohmander, 1932, male from Toðanalý, Azerbaijan. A: gonopods, mesal view. B: gonopods, mesodistal view. C: gonopods, posterodistal view. D: promere of gonopod, posterior view. E: gonopods without promere, mesal view. F: gonopods without promere, lateral view. Abbreviations: fl, flagellum; lp, lamella of promere; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: A: 0.2 mm, rest: 0.1 mm.	FIGURE 20. Leptoiulus hastatus Lohmander, 1932, male from Toðanalý, Azerbaijan. A: gonopods, mesal view. B: gonopods, mesodistal view. C: gonopods, posterodistal view. D: promere of gonopod, posterior view. E: gonopods without promere, mesal view. F: gonopods without promere, lateral view. Abbreviations: fl, flagellum; lp, lamella of promere; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: A: 0.2 mm, rest: 0.1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFDC4733FF6D5FA2FB39FA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826253/files/figure.png	https://doi.org/10.5281/zenodo.3826253	FIGURE 21. Leptoiulus hastatus Lohmander, 1932, aberrant male from Ceyrankeçməz, Qobustan, Azerbaijan. A: gonopods, lateral view. B: gonopods, mesal view. C: promere of gonopod, posterior view. Female from same place as male. D: vulva, posterior view, E: vulva, lateral view. F: vulva, mesal view. Abbreviations: bu, bursa; lp, lamella of promere; ms, mesomere; om, operculum; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.1 mm.	FIGURE 21. Leptoiulus hastatus Lohmander, 1932, aberrant male from Ceyrankeçməz, Qobustan, Azerbaijan. A: gonopods, lateral view. B: gonopods, mesal view. C: promere of gonopod, posterior view. Female from same place as male. D: vulva, posterior view, E: vulva, lateral view. F: vulva, mesal view. Abbreviations: bu, bursa; lp, lamella of promere; ms, mesomere; om, operculum; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFDC4733FF6D5FA2FB39FA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826255/files/figure.png	https://doi.org/10.5281/zenodo.3826255	FIGURE 22. A-B: Leptoiulus hastatus Lohmander, 1932, male holotype, after Lohmander, 1932 (A-B). A: gonopod, mesal view. B: opisthomere of gonopod, lateral view. C-F: male holotype of L. disparatus Lohmander, 1936, after Lohmander, 1936. C: promere of gonopod, caudal view. D: gonopod, mesal view. E: end of opisthomere, frontolateral view. F: end of opisthomere, caudal view. Abbreviations: fl, flagellum; lp, lamella of promere; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Drawn not to scale.	FIGURE 22. A-B: Leptoiulus hastatus Lohmander, 1932, male holotype, after Lohmander, 1932 (A-B). A: gonopod, mesal view. B: opisthomere of gonopod, lateral view. C-F: male holotype of L. disparatus Lohmander, 1936, after Lohmander, 1936. C: promere of gonopod, caudal view. D: gonopod, mesal view. E: end of opisthomere, frontolateral view. F: end of opisthomere, caudal view. Abbreviations: fl, flagellum; lp, lamella of promere; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Drawn not to scale.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFDC4733FF6D5FA2FB39FA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826257/files/figure.png	https://doi.org/10.5281/zenodo.3826257	FIGURE 23. Leptoiulus hastatus Lohmander, 1932. A-E: end of gonopod opisthomere, mesal views. A: male from Vardzia, Georgia. B: male from Tba Bazaleti Lake. C: male from Vardzia, Georgia. D: male from Vardzia, Georgia. E: male from Toðanalý, Azerbaijan. F–G: vulvae, posterior view. F: female from Sevanavank Monastery. Armenia. G: female from Tba Bazaleti Lake Abbreviations: bu, bursa; om, operculum; ph, phylacum; sl, solenomere; ve, velum. Scale bars: 0.02 mm (A–D), 0.1 mm (F–G) or not to scale (E).	FIGURE 23. Leptoiulus hastatus Lohmander, 1932. A-E: end of gonopod opisthomere, mesal views. A: male from Vardzia, Georgia. B: male from Tba Bazaleti Lake. C: male from Vardzia, Georgia. D: male from Vardzia, Georgia. E: male from Toðanalý, Azerbaijan. F–G: vulvae, posterior view. F: female from Sevanavank Monastery. Armenia. G: female from Tba Bazaleti Lake Abbreviations: bu, bursa; om, operculum; ph, phylacum; sl, solenomere; ve, velum. Scale bars: 0.02 mm (A–D), 0.1 mm (F–G) or not to scale (E).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFDC4733FF6D5FA2FB39FA1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826299/files/figure.png	https://doi.org/10.5281/zenodo.3826299	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFD74738FF6D58CFFC2CFB75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826239/files/figure.png	https://doi.org/10.5281/zenodo.3826239	FIGURE 15. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Leptoiulus hastatus Lohmander, 1932, male from near Shnokh, Armenia. B: Leptoiulus tanymorphus (Attems, 1901), male from Ajikent, Azerbaijan. C: Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. D: Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. Scale bar: 1 mm.	FIGURE 15. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Leptoiulus hastatus Lohmander, 1932, male from near Shnokh, Armenia. B: Leptoiulus tanymorphus (Attems, 1901), male from Ajikent, Azerbaijan. C: Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. D: Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. Scale bar: 1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFD74738FF6D58CFFC2CFB75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826263/files/figure.png	https://doi.org/10.5281/zenodo.3826263	FIGURE 25. Leptoiulus tanymorphus, male from Zərgəran, Ýsmayýllý, Azerbaijan. A: leg pair 1, anteriolateral view. B: leg pair 1, anterior view. C: gnathochilarium, lateral view. Abbreviations: lg, lamella lingualis of gnathochilarium; sg, stipes of gnathoсhilarium. Scale bars: A, B: 0.05 mm, C: 0.1 mm.	FIGURE 25. Leptoiulus tanymorphus, male from Zərgəran, Ýsmayýllý, Azerbaijan. A: leg pair 1, anteriolateral view. B: leg pair 1, anterior view. C: gnathochilarium, lateral view. Abbreviations: lg, lamella lingualis of gnathochilarium; sg, stipes of gnathoсhilarium. Scale bars: A, B: 0.05 mm, C: 0.1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFD74738FF6D58CFFC2CFB75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826265/files/figure.png	https://doi.org/10.5281/zenodo.3826265	FIGURE 26. Leptoiulus tanymorphus (Attems, 1901), male from Ismailly, Azerbaijan. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	FIGURE 26. Leptoiulus tanymorphus (Attems, 1901), male from Ismailly, Azerbaijan. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFD74738FF6D58CFFC2CFB75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826267/files/figure.png	https://doi.org/10.5281/zenodo.3826267	FIGURE 27. Leptoiulus tanymorphus (Attems, 1901), male from near Kuba, Azerbaijan. A: tip of leg 2, caudal view. B: legpair 1, caudal view. C & D: gonopods, ventral and ventrolateral views, respectively. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.01 mm (A), 0.05 mm (B), 0.1 mm (C, D).	FIGURE 27. Leptoiulus tanymorphus (Attems, 1901), male from near Kuba, Azerbaijan. A: tip of leg 2, caudal view. B: legpair 1, caudal view. C & D: gonopods, ventral and ventrolateral views, respectively. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.01 mm (A), 0.05 mm (B), 0.1 mm (C, D).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFD74738FF6D58CFFC2CFB75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826269/files/figure.png	https://doi.org/10.5281/zenodo.3826269	FIGURE 28. Leptoiulus tanymorphus (Attems, 1901), male from Khosrov Nature Reserve, Armenia. A: head, ventral view. B: leg-pair 1, caudal view. C & D: gonopods, ventral and frontoventral views, respectively. E: opisthomere of gonopod, frontal view. Abbreviations: fl, flagellum; lg, lamella lingualis of gnathochilarium; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sg, stipes of gnathochilarium; sl, solenomere; ve, velum. Scale bars: 0.1 mm (A), 0.05 mm (B-D), 0.02 mm (E).	FIGURE 28. Leptoiulus tanymorphus (Attems, 1901), male from Khosrov Nature Reserve, Armenia. A: head, ventral view. B: leg-pair 1, caudal view. C & D: gonopods, ventral and frontoventral views, respectively. E: opisthomere of gonopod, frontal view. Abbreviations: fl, flagellum; lg, lamella lingualis of gnathochilarium; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sg, stipes of gnathochilarium; sl, solenomere; ve, velum. Scale bars: 0.1 mm (A), 0.05 mm (B-D), 0.02 mm (E).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFD74738FF6D58CFFC2CFB75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826271/files/figure.png	https://doi.org/10.5281/zenodo.3826271	FIGURE 29. Leptoiulus tanymorphus, male from Zərgəran, Ýsmayýllý, Azerbaijan. A: gonopods without promere, lateral view. B: gonopods, mesal view. C: promere of gonopod, posterior view. Female from same place as male. D: vulva, posterior view, E: vulva, lateral view. F: vulva, mesal view. Abbreviations: bu, bursa; fl, flagellum; om, operculum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.05 mm (A, D, E) or 0.1 mm (B, C, F).	FIGURE 29. Leptoiulus tanymorphus, male from Zərgəran, Ýsmayýllý, Azerbaijan. A: gonopods without promere, lateral view. B: gonopods, mesal view. C: promere of gonopod, posterior view. Female from same place as male. D: vulva, posterior view, E: vulva, lateral view. F: vulva, mesal view. Abbreviations: bu, bursa; fl, flagellum; om, operculum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.05 mm (A, D, E) or 0.1 mm (B, C, F).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFD74738FF6D58CFFC2CFB75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826273/files/figure.png	https://doi.org/10.5281/zenodo.3826273	FIGURE 30. Leptoiulus tanymorphus (Attems, 1901) (A, male, after Lohmander (1936), B and C, male from near Shamshadyn, Armenia). A: gonopods, mesal view. B: promere of gonopod, caudal view. C: gonopod, mesal view. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bar 0.2 mm (B) or drawn not to scale (A, C).	FIGURE 30. Leptoiulus tanymorphus (Attems, 1901) (A, male, after Lohmander (1936), B and C, male from near Shamshadyn, Armenia). A: gonopods, mesal view. B: promere of gonopod, caudal view. C: gonopod, mesal view. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bar 0.2 mm (B) or drawn not to scale (A, C).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFD74738FF6D58CFFC2CFB75.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826299/files/figure.png	https://doi.org/10.5281/zenodo.3826299	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEE4701FF6D5BDBFD3CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826239/files/figure.png	https://doi.org/10.5281/zenodo.3826239	FIGURE 15. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Leptoiulus hastatus Lohmander, 1932, male from near Shnokh, Armenia. B: Leptoiulus tanymorphus (Attems, 1901), male from Ajikent, Azerbaijan. C: Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. D: Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. Scale bar: 1 mm.	FIGURE 15. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Leptoiulus hastatus Lohmander, 1932, male from near Shnokh, Armenia. B: Leptoiulus tanymorphus (Attems, 1901), male from Ajikent, Azerbaijan. C: Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. D: Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. Scale bar: 1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEE4701FF6D5BDBFD3CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5917994/files/figure.png	https://doi.org/10.5281/zenodo.5917994	FIGURE 31. Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. A-C: anterior, middle and posterior parts of body, respectively, lateral views. D-F: anterior, middle and posterior parts of body, respectively, ventral views. G-I: same, respectively, dorsal views. J: midbody ring, frontal view. K: ozopore, lateral view. L hypoproct, ventral view. M: head, ventral view. N: tip of leg 2, ventrocaudal view. Abbreviations: ep, epiproct; hp, hypoproct; lg, lamella lingualis of gnathochilarium; mz, metazona; pz, prozona; sg, stipes of gnathoсhilarium. Scale bars: 0.2 mm (A–E), 0.1 mm (F, I, M) or 0.05 mm (K, L, N).	FIGURE 31. Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. A-C: anterior, middle and posterior parts of body, respectively, lateral views. D-F: anterior, middle and posterior parts of body, respectively, ventral views. G-I: same, respectively, dorsal views. J: midbody ring, frontal view. K: ozopore, lateral view. L hypoproct, ventral view. M: head, ventral view. N: tip of leg 2, ventrocaudal view. Abbreviations: ep, epiproct; hp, hypoproct; lg, lamella lingualis of gnathochilarium; mz, metazona; pz, prozona; sg, stipes of gnathoсhilarium. Scale bars: 0.2 mm (A–E), 0.1 mm (F, I, M) or 0.05 mm (K, L, N).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEE4701FF6D5BDBFD3CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826281/files/figure.png	https://doi.org/10.5281/zenodo.3826281	FIGURE 32. Leptoiulus meskhii sp. nov., male from Hoveri, Lerik, Azerbaijan. A: leg pair 1, anterolateral view. B: leg pair 1, anterior view. C: gnathochilarium, ventral view. Abbreviations: lg, lamella lingualis of gnathochilarium; sg, stipes of gnathoсhilarium. Scale bars: A: 0.05 mm (A) or 0,1 mm (B, C).	FIGURE 32. Leptoiulus meskhii sp. nov., male from Hoveri, Lerik, Azerbaijan. A: leg pair 1, anterolateral view. B: leg pair 1, anterior view. C: gnathochilarium, ventral view. Abbreviations: lg, lamella lingualis of gnathochilarium; sg, stipes of gnathoсhilarium. Scale bars: A: 0.05 mm (A) or 0,1 mm (B, C).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEE4701FF6D5BDBFD3CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826283/files/figure.png	https://doi.org/10.5281/zenodo.3826283	FIGURE 33. Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	FIGURE 33. Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEE4701FF6D5BDBFD3CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826285/files/figure.png	https://doi.org/10.5281/zenodo.3826285	FIGURE 34. Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. A: leg-pair 1, frontoventral view. B & C: gonopods, ventrolateral and ventrocaudal views, respectively. D: opisthomere of gonopod, mesal view. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.05 mm (A), 0.1 mm (B, C) or 0.02 mm (D).	FIGURE 34. Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. A: leg-pair 1, frontoventral view. B & C: gonopods, ventrolateral and ventrocaudal views, respectively. D: opisthomere of gonopod, mesal view. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.05 mm (A), 0.1 mm (B, C) or 0.02 mm (D).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEE4701FF6D5BDBFD3CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826287/files/figure.png	https://doi.org/10.5281/zenodo.3826287	FIGURE 35. Leptoiulus meskhii sp. nov., male from Hoveri, Lerik, Azerbaijan. A: gonopods, lateral view. B: gonopods, mesal view. C: promere of gonopod, posterior view. Female from same place as male. D: vulva, posterior view, E: vulva, lateral view. F: vulva, mesal view. Abbreviations: bu, bursa; om, operculum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.05 mm (A, C, D, E) or 0.1 mm (B, F).	FIGURE 35. Leptoiulus meskhii sp. nov., male from Hoveri, Lerik, Azerbaijan. A: gonopods, lateral view. B: gonopods, mesal view. C: promere of gonopod, posterior view. Female from same place as male. D: vulva, posterior view, E: vulva, lateral view. F: vulva, mesal view. Abbreviations: bu, bursa; om, operculum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.05 mm (A, C, D, E) or 0.1 mm (B, F).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEE4701FF6D5BDBFD3CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826289/files/figure.png	https://doi.org/10.5281/zenodo.3826289	FIGURE 36. Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. A: promere of gonopod, caudal view. B: gonopod, meal view. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bar: 0.2 mm.	FIGURE 36. Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. A: promere of gonopod, caudal view. B: gonopod, meal view. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bar: 0.2 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEE4701FF6D5BDBFD3CFF2A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826299/files/figure.png	https://doi.org/10.5281/zenodo.3826299	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEA4705FF6D5884FDC1FBF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826239/files/figure.png	https://doi.org/10.5281/zenodo.3826239	FIGURE 15. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Leptoiulus hastatus Lohmander, 1932, male from near Shnokh, Armenia. B: Leptoiulus tanymorphus (Attems, 1901), male from Ajikent, Azerbaijan. C: Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. D: Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. Scale bar: 1 mm.	FIGURE 15. Habitus pictures of some species of Leptoiulini from the Caucasus, lateral views. A: Leptoiulus hastatus Lohmander, 1932, male from near Shnokh, Armenia. B: Leptoiulus tanymorphus (Attems, 1901), male from Ajikent, Azerbaijan. C: Leptoiulus meskhii sp. nov., male paratype from Galabyn, Azerbaijan. D: Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. Scale bar: 1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEA4705FF6D5884FDC1FBF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826291/files/figure.png	https://doi.org/10.5281/zenodo.3826291	FIGURE 37. Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. A-C: anterior, middle and posterior parts of body, respectively, lateral views. D: anterior part of body, frontolateral view. E: tip of leg 2, frontal view. F: head, ventral view. G: midbody ring, frontal view. Abbreviations: ep, epiproct; lg, lamella lingualis of gnathochilarium; mz, metazona; pz, prozona; sg, stipes of gnathoсhilarium. Scale bars: 0.1 mm (A–D, G), 0.02 (E) or 0.2 (F).	FIGURE 37. Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. A-C: anterior, middle and posterior parts of body, respectively, lateral views. D: anterior part of body, frontolateral view. E: tip of leg 2, frontal view. F: head, ventral view. G: midbody ring, frontal view. Abbreviations: ep, epiproct; lg, lamella lingualis of gnathochilarium; mz, metazona; pz, prozona; sg, stipes of gnathoсhilarium. Scale bars: 0.1 mm (A–D, G), 0.02 (E) or 0.2 (F).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEA4705FF6D5884FDC1FBF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826293/files/figure.png	https://doi.org/10.5281/zenodo.3826293	FIGURE 38. Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	FIGURE 38. Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. A: antenna, lateral view. B: gnathochilarium, ventral view. C: leg 1, caudal view. D: leg 2, caudal view. E: ventral edge of segment 7, lateral view. Scale bar: 0.2 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEA4705FF6D5884FDC1FBF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826295/files/figure.png	https://doi.org/10.5281/zenodo.3826295	FIGURE 39. Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. A: segment 7, frontoventral view. B: leg-pair 2, caudal view. C & D: gonopod, ventral and mesal views, respectively. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.1 mm.	FIGURE 39. Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. A: segment 7, frontoventral view. B: leg-pair 2, caudal view. C & D: gonopod, ventral and mesal views, respectively. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bars: 0.1 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEA4705FF6D5884FDC1FBF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826297/files/figure.png	https://doi.org/10.5281/zenodo.3826297	FIGURE 40. Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. A: promere of gonopod with flagellum, caudal view B: gonopod, mesal view. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bar: 0.2 mm.	FIGURE 40. Leptoiulus gonopodialis sp. nov., male paratype from Lagodekhi Nature Reserve, Georgia. A: promere of gonopod with flagellum, caudal view B: gonopod, mesal view. Abbreviations: fl, flagellum; ms, mesomere; op, opisthomere; ph, phylacum; pr, promere; sl, solenomere; ve, velum. Scale bar: 0.2 mm.	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
039087E0FFEA4705FF6D5884FDC1FBF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3826299/files/figure.png	https://doi.org/10.5281/zenodo.3826299	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	FIGURE 41. Female vulvae of Leptoiulini from the Caucasus, caudolateral views. A: Chaetoleptophyllum flexum Golovatch, 1979 (female from Algeti Nature Reserve, Georgia). B: Kubaniulus gracilis Lohmander, 1936 (female from Pasture Abago, Krasnodar Prov.). C: Kubaniulus lativelatus sp. nov. (paratype from near Gelenjik, Krasnodar Prov). D: Leptoiulus hastatus Lohmander, 1936 (female from Totev,Armenia). E: Leptoiulus tanymorphus (Attems, 1901) (paratype, after Lohmander, 1936). F: Leptoiulus meskhii sp. nov. (paratype from Galabyn, Azerbaijan). G: Leptoiulus gonopodialis sp. nov. (paratype from Lagodekhi Nature Reserve, Georgia). Abbreviations: a, ampulla/ -ae; bu, bursa; em, emargination of median field; mf, median field; om, operculum; rs, receptaculum seminis. Scale bar: 0.2 mm (A–E, G), drawn not to scale (H, F).	2020-05-14	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier		Zenodo	biologists	Evsyukov, Aleksandr;Golovatch, Sergei;Reip, Hans;Vandenspiegel, Didier			
