taxonID	type	description	language	source
039E87B4FFE3FFD82D91FA1AFAA2FCB2.taxon	diagnosis	Diagnosis. Head yellowish-brown with all the areas before the antennae dark brown except the clypeus; small brown spots present in the vertex and densely covered with long bristles. Forewing and hind wing oval with outer margin slightly pointed; forewing with sagittal stripes and sparse brown spots present along the Cu and 3 m-cu. Male: anteroventral edge of ectoproct developed into stout arm, ended flat with small teeth covered the margin; terminal lobe of parabaculum separated from four fifth and supraapophyseal plate laterally splayed outwards.	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE3FFD82D91FA1AFAA2FCB2.taxon	description	Measurements. Forewing length 8.3 – 9.5 mm, width 3.1 – 4.0 mm. Hind wing length 7.5 – 8.2 mm, width 2.6 – 3.2 mm. Body length 5.2 – 6.0 mm. Description. Head. Yellowish-brown. All the areas before the antennae dark brown except the clypeus, small brown spots present in the vertex and densely covered with long bristles. Antennae amber, brown spot present in the ventral scape, with more than fifty segments. Eyes black with a metallic luster. Thorax. Yellowish-brown, with brown longitudinal stripes along both sides. Legs yellowish-brown, with a brown spot present at the apical tibiae. Wings (Fig. 1). Forewing oval, yellowish-brown, hyaline with sagittal stripes and sparse brown spots present along the Cu vein and 3 m-cu. Veins brown with transparent intervals, except the crossveins of gradate series all brown. Anterior radial trace bearing 3 ORB’s; ORB 3 forked into 4 branches, with 2 r – rs. M simple with 2 branches. CuA forked into 6 braches after the inner gradate series and CuP simple. Four gradate series, inner gradate series 3 crossveins; middle 7; outer 7 and cubitus 3. Hind wing oval, pale yellowish-brown, hyaline and immaculate. Veins brown and transparent, with Rs and M slightly lighter and Cu slightly darker. Rs with 4 branches, with 2 r – rs. M forked into 2 branches. CuA 5 branches and CuP simple. Two gradate series, inner gradate series 2 crossveins; and outer 7. Male terminalia (Fig. 4). 8 th tergite trapezoidal, posteroventral edge declining and not include the spiraculae. 9 th tergite rectangle from lateral view. Ectoproct developed with anteroventral edge developed into stout arm, bend inward but not intersected, ended flat with small teeth covered the margin. Mediuncus of gonarcus (Fig. 5 – 6) broadened basally, tapering distally and bend downwards; intrahemigonarcus ovoid and extrahemigonarcus subtriangular from dorsal view with paramediuncus triangular. Terminal lobe of parabaculum (Fig. 7 – 8) separated from four-fifths, bend inwards and tapering distally; supraapophyseal plate laterally splayed. Hypandrium internum (Fig. 9) approximate trapezoidal from ventral view with both sides slightly curled. Female terminalia (Fig. 10 – 11). Lateral of 9 th tergite extended sidewards. gonapophyses laterales developed, hind margin obviously exceed the hind margin of ectoproct, approximate elongated from ventral view. Ectoproct ovoid. Subgenitale elongated into stripe, slightly expanded at both ends and contracted in the middle, gonapophyses posteriors consist of two lamella respectively, subtriangular and semi-circular from ventral view.	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE3FFD82D91FA1AFAA2FCB2.taxon	distribution	Distribution. China (Sichuan)	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE3FFD82D91FA1AFAA2FCB2.taxon	materials_examined	Material examined. CHINA: Holotype, 1 ♂, Sichuan province, Dujiangyan city, Qingchengshan town, Qingcheng Mountains. 1. v. 1978, Fasheng Li (CAU). Paratypes, 2 ♂ 1 ♀, Sichuan province, Chengdu city, Dayi county. 3. v. 1978, Fasheng Li (CAU).	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE3FFD82D91FA1AFAA2FCB2.taxon	etymology	Etymology. The specific name refers to the special parabaculum, with the terminal lobe deeply forked, which separated from four-fifths.	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE3FFD82D91FA1AFAA2FCB2.taxon	discussion	Remarks. This new species is different from other species by the special parabaculum with the terminal lobe separated from four-fifths. Although it is similar to the W. helanensis in forewing pattern, it can be easily distinguished by the color of head and the shape of anteroventral edge of ectoproct. This species is also similar to W. transsylvanicus and W. subnebulosus, it can be distinguished from W. subnebulosus by the conspicuous cubitus gradate series in forewing while in W. subnebulosus the cubitus gradate series is inconspicuous. As for W. transsylvanicus, the 8 th tergite approximate trapezoid from lateral view, the hind margin of 9 th sternite curved upwards and the anteroventral edge of ectoproct ended with obvious brush-like protrusion while in W. dissectus the posteroventral margin of 8 th tergite extended downward obviously, the 9 th sternite trapezoid from lateral view and the anteroventral edge of ectoproct developed into stout arm, ended flat with small teeth covered the margin.	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE4FFDB2D91FC55FDCDF8FD.taxon	diagnosis	Diagnosis. This species is characterized by the yellowish-brown head with all the areas before the antennae dark brown; brown longitudinal stripe present in the middle of vertex. Forewing hyaline with no obvious spots and hind wing immaculate, with 1 r-rs absent, only 2 r-rs present. Male: anteroventral edge of ectoproct developed upwards as a strong arm; basal apophyseal lamella of parabaculum laterally splayed from dorsal view and supraapophyseal plate laterally splayed. Femal: subgenitale consist of a pair of rhombic lamella, which separate mutually.	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE4FFDB2D91FC55FDCDF8FD.taxon	description	Measurements. Forewing length 7.2 – 7.6 mm, width 3.1 – 3.3 mm. Hind wing length 6.1 – 6.3 mm, width 2.7 – 2.9 mm. Body length 5.7 – 7.2 mm. Description. Head. Yellowish-brown. All the areas before the antennae dark brown, a subtriangular brown spot present near the hind margin of antennal socket, brown longitudinal stripe present in the middle of vertex. Antennae amber, scape and pedicel brown, obviously darker than flagellum, with more than sixty-five segments. Eyes black with a metallic luster. Thorax. Yellowish-brown, with brown longitudinal stripes along the sides of pronotum and mesonotum, scutum of metanotum obviously brown. Legs yellowish-brown with no spots and last tarsi darker than others. Wings (Fig. 2). Forewing ovoid, yellowish-brown, hyaline with no obvious spots. Veins brown with transparent intervals, crossveins of gradate series all brown. Anterior radial trace bearing 3 ORB’s; ORB 3 forked into 3 – 4 branches, with 2 r – rs. M simple with 2 branches. CuA forked into 5 braches after the inner gradate series and CuP simple. Four gradate series, inner gradate series 3 crossveins; middle 6; outer 6 – 8 and cubitus 3. Hind wing ovoid with outer margin slightly pointed, pale yellowish-brown, hyaline and immaculate. Veins pale yellowish-brown. Rs with 5 branches, 1 r-rs absent, only 2 r-rs present. M forked into 2 branches. CuA 5 branches and CuP simple. Two gradate series, inner gradate series 2 crossveins; and outer 7. Male terminalia (Fig. 12). 8 th tergite rectangle, posteroventral edge declining and not include the spiraculae. 9 th tergite partly fused with ectoproct, dorsal of hind margin slightly depressed forward. Ectoproct developed, extending backward; anteroventral edge developed upwards as a strong arm, with a blunt end, densely covered with small spines. Mediuncus of gonarcus (Fig. 13 – 14) strong, tapering distally; paramediuncus strong, curved inward from dorsal view, with apex slightly pointed. Parabaculum (Fig. 15 – 16) special, terminal lobe separated from half, tapering gradually and sparsely covered with small spines; base of apophyseal lamella laterally splayed from dorsal view; supraapophyseal plate laterally splayed, more than half length of the parabaculum, with a small upward hook in the end. Hypandrium internum (Fig. 17) approximate triangle from ventral view. Female terminalia (Fig. 18 – 19). 9 th tergite approximate " L " shaped from lateral view, lateral extended backwards. gonapophyses laterales developed, hind margin obviously exceed the hind margin of ectoproct. Ectoproct ovoid, middle of the hind margin slightly depressed. Subgenitale consist of a pair of rhombic lamella, which separate mutually.	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE4FFDB2D91FC55FDCDF8FD.taxon	distribution	Distribution. Europe, Japan, Transbaikalian Russia and China (Hubei, Inner Mongolia)	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE4FFDB2D91FC55FDCDF8FD.taxon	materials_examined	Material examined. CHINA: 3 ♂, Hubei province, Yichang city, Xingshan county, Longmen. 28. ix. 1994, Chikun Yang (CAU). 1 ♀, Inner Mongolia Autonomous Region, Alxa Left Banner, Helan Mountains. 31. vii. 2010. Yanlin Tian (CAU).	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE4FFDB2D91FC55FDCDF8FD.taxon	discussion	Remarks. This species was found in United Kingdom by Withycombe with two male specimens and Nakahara found female specimens from Japan in 1956. This is the first time for the species reported in China. This species seems related to W. mortoni based on the forewing; however, it can be distinguished from W. mortoni by the absence of the prominent dorsal tuft of hairs at the apex of abdomen and the more elongate and slender ectoproct with the turner inwards and upwards.	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE6FFD72D91F8C6FCFCFC54.taxon	diagnosis	Diagnosis. This species is characterized by the yellowish-brown head with only the frons before the antennae obviously dark brown. Brown longitudinal stripes along the sides present in the thorax and brown longitudinal thin stripe present in the middle of pronotum and mesonotum. Legs yellowish-brown, with brown spot present at the basal and apical tibiae. Male: posteroventral edge of 8 th tergite declining and not include the spiraculae; terminal lobe of parabaculum separated from one fifth, expanded distally and rolled up, a transparent small hook present in the end. Female: ectoproct ovoid, posteroventral edge slightly pointed, middle of hind margin with an obvious projection.	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE6FFD72D91F8C6FCFCFC54.taxon	description	Measurements. Forewing length 7.4 – 8.4 mm, width 3.0 – 4.2 mm. Hind wing length 6.6 – 7.3 mm, width 2.2 – 4.0 mm. Body length 4.8 – 6.4 mm. Description. Head. Yellowish-brown. Frons obviously dark brown, oval brown spot present in the middle of vertex. Antennae amber, ventral of scape brown, with more than fifty segments. Eyes big and black with a metallic luster. Thorax. Yellowish-brown, with obvious brown longitudinal stripes along the sides and brown longitudinal thin stripe present in the middle of pronotum and mesonotum. Legs yellowish-brown, with brown spot present at the basal and apical tibiae. Wings (Fig. 3). Forewing oval with outer margin slightly pointed, yellowish-brown, hyaline with uneven grey sagittal stripes and sparse pale brown spots in the margin. Veins yellowish-brown with transparent intervals. Anterior radial trace bearing 3 ORB’s; ORB 3 forked into 2 branches, with 2 r – rs. M simple with 2 branches. CuA forked into 6 braches after the inner gradate series and CuP simple. Four gradate series, inner gradate series 3 crossveins; middle 5; outer 6 and cubitus 3. Hind wing oval with outer margin slightly pointed, pale yellowishbrown, hyaline and immaculate. Veins pale brown and transparent, with Cu slightly darker. Rs with 4 branches, with 2 r – rs. M forked into 2 branches. CuA 4 branches and CuP simple with 2 branches. Two gradate series, inner gradate series 2 crossveins; and outer 7. Male terminalia (Fig. 20). 8 th tergite trapezoidal, posteroventral edge declining and not include the spiraculae. 9 th tergite rectangle from lateral view. Ectoproct subtriangular with anteroventral edge developed into slender arm, bend inwards and intersected, ended into a hook. Mediuncus of gonarcus (Fig. 21 – 22) broadened basally, tapering distally and slightly bend downwards; paramediuncus small, curved upwards as a hook. Terminal lobe of parabaculum (Fig. 23 – 24) separated from one fifth, expanded distally and rolled up, densely covered with small spines in the surface, ending in a small transparent hook; supraapophyseal plate wide, subtriangular from dorsal view. Hypandrium internum (Fig. 25) approximate triangle from ventral view, split in the central base. Female terminalia (Fig. 26 – 27). Lateral of 9 th tergite extended sidewards. gonapophyses laterales developed, hind margin obviously exceed the hind margin of ectoproct, approximate long oval from ventral view. Ectoproct ovoid, posteroventral slightly pointed, middle of hind margin with an obvious projection. Subgenitale broadened basally, tapering distally with apex flat; gonapophyses posteriors distally pointed and exceed the apex of subgenitale.	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE6FFD72D91F8C6FCFCFC54.taxon	distribution	Distribution. China (Inner Mongolia, Qinghai, Gansu)	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE6FFD72D91F8C6FCFCFC54.taxon	materials_examined	Material examined. CHINA: 1 ♂, Inner Mongolia Autonomous Region, Alxa Left Banner, Helan Mountains. 29. vii. 2010. Yanlin Tian (Holotype) (CAU); 1 ♂, Inner Mongolia Autonomous Region, Alxa Left Banner, Helan Mountains. 6. viii. 2010. Yanlin Tian (CAU); 1 ♂, Inner Mongolia Autonomous Region, Alxa Left Banner, Helan Mountains. 9. viii. 2010. Yanlin Tian (CAU); 1 ♀, Gansu province, Tianshui city, Qingshui county, Dalong forest. 4. viii. 2013, Xinyu Luo (CAU); 1 ♀, Qinghai province, Tibetan Autonomous Prefecture of Haibei, Menyuan Hui Autonomous County, Xianmi National Forest Park. 9. viii. 2013, Xinyu Luo (CAU).	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
039E87B4FFE6FFD72D91F8C6FCFCFC54.taxon	discussion	Remarks. This species was described by Tian & Liu in 2011 only by the male specimens; in this paper we describe the female specimen and the new distributions in China for the first time. W. helanensis is similar to W. sufuensis, but obviously different in many characteristics, there is no brown stripe along the middle gradate series in forewing in W. helanensis, but they present in W. sufuensis; in W. helanensis the mediucus of gonarcus is slender and paramediuncus curved upwards as a hook while in W. sufuensis the mediucus is short and paramediuncus extend backward; the terminal lobe of parabaculum in W. helanensis expanded distally and rolled up, with a transparent small hook present in the end while in W. sufuensis the distal of terminal lobe extended in to a upward hook. W. helanensis is also similar to W. baikalensis, while considering the color of head, it can be easily distinguished, all the areas before antennae in head are dark brown in W. baikalensis, while in W. helanensis only the frons before the antennae are obviously dark brown.	en	Zhao, Yang, Tian, Yanlin, Liu, Zhiqi (2017): New data on brown lacewing genus Wesmaelius Krüger, 1922 from China (Neuroptera, Hemerobiidae), with a key to Chinese species. Zootaxa 4273 (1): 19-30, DOI: 10.11646/zootaxa.4273.1.2
