identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039E87C6FFF9E463D8BFD380FB0FF90B.text	039E87C6FFF9E463D8BFD380FB0FF90B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nossidium Erichson 1845	<div><p>Nossidium Erichson 1845</p><p>Darby (2015b) revised the generic description, discussed the relationships with other closely related genera, and described Nossidium katyae from Bolivia, the only known representative of the genus in S. America. The only mention of the genus in Central America is by Dybas (1976) who recorded seeing a species possibly related to Nossidium collected at 7,600 feet on the Volcan Chiriqui in Panama on the Costa Rica border. Three species have been described from North America, N. americanum and N. amoenum by Motschulsky 1868 (as Anisarthria) and N. posthumum by Matthews 1874. Examples of these species in BMNH have been examined to confirm possible synonymy with the new Costa Rican species but have proved to be quite different particularly in their more parallel, less rounded form, which is similar to that of N. pilosellum, the type species of the genus.</p></div>	https://treatment.plazi.org/id/039E87C6FFF9E463D8BFD380FB0FF90B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Darby, Michael	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFF9E461D8BFD153FAB8FE51.text	039E87C6FFF9E461D8BFD153FAB8FE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nossidium issyae	<div><p>Nossidium issyae sp. n.</p><p>(Figs. 3 A–H)</p><p>Habitus Fig. 3 Aa/b. Length 0.96 mm. Body yellowish brown covered with grey/yellow overlapping pubescence, legs and antennae dusky yellow. Antennomeres 3–8 length 0.4 mm; antennomeres 4–8 elongate, 10,11 globular, with sensillae. Width across eyes 0.38 mm. Mentum broadest at anterior margin tapering towards base, two longest setae situated medially on lateral margins, Fig. 3F. Pronotum length 0.33 mm, width 0.63 mm, posterior margin sinuate before the angles. Elytra length 0.56 mm, width 0.63 mm, widest just behind humeri. Wings of usual Nossidium type, Fig. 3H. Mesoventral collar reaching onto the humeri with a raised anterior border and a vertically oriented medial ridge, Fig. 3G; mesoventral keel wide between the mesocoxae, Fig. 3D. Metaventrum short, pubescence denser on disc, posterior intercoxal process comprised of two short, sharp points, Fig. 3E.</p><p>Male: not known.</p><p>Female: spermatheca globular, Fig. 3C.</p><p>Etymology. Named after Miss Isobel 'Issy' Millett, a close friend of the author’s family.</p><p>Diagnosis. The new species may be separated from N. schuelkei sp. n. and N. katyae by the form of the mesoventral keel which does not extend below the mesocoxae. In this respect it is similar to N. pilosellum, and the North American species N. americanum Motschulsky and N. amoenum Motschulsky but these are less rounded and more parallel sided species.</p><p>Type data. Holotype: ♀, Costa Rica, Alajuela, Peñas Blancas, 820m, 19 May 1989, J. Ashe, R. Brooks, R. Leschen; Snow Entomological Museum , Costa Rican Expedition,#279; ex. with ants under bark (MM). Fragments of a second specimen from the same locality, ex. Xylaria, 17 May 1989, mounted as a slide, are also in MM.</p></div>	https://treatment.plazi.org/id/039E87C6FFF9E461D8BFD153FAB8FE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Darby, Michael	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFBE466D8BFD0FCFD03FCC5.text	039E87C6FFFBE466D8BFD0FCFD03FCC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nossidium schuelkei	<div><p>Nossidium schuelkei sp. n.</p><p>(Figs. 4 A –F, 5 A–E)</p><p>Length 0.84 mm. Body dark brown covered with grey/yellow overlapping pubescence, legs and antennae dusky yellow, lateral and posterior borders with a sharply defined yellow margin in transmitted light. Antennomeres 3–8 length 0.33 mm, 4–8 elongate, 10,11 globular, Fig. 4D. Width across eyes 0.31 mm. Mentum broadest at anterior margin tapering towards base, two longest setae situated medially on lateral margins, Fig. 5C. Pronotum length 0.24 mm, width 0.50 mm, posterior margin sinuate before the angles. Elytra length 0.56 mm, width 0.53 mm, widest just behind humeri, wings of usual Nossidium type, Fig. 4E. Mesoventral collar reaching onto the humeri with a raised anterior border and a vertically oriented medial ridge, Fig. 5B; mesoventral keel wide, reaching below the mesocoxae in dorsal view and with a flattened cap rounded anteriorly and with a raised border, closely fitting posteriorly with the metaventrum so as to appear an anterior metaventral process, Fig. 5A. Metaventrum with line of metepisternal suture reaching mesocoxae but not directed posteriad at lateral margins; lateral points on the posterior intercoxal process broadened and flattened towards the base with convex lateral margins, Fig. 5 F. Pygidium with +/– 15 teeth Fig. 5 G.</p><p>Male: not known.</p><p>Female: spermatheca Fig. 4 F.</p><p>Etymology. Named after the collector Michael Schülke of the Museum für Naturkunde, Berlin.</p><p>Diagnosis. The new species is similar to N. katyae but may be distinguished by its less shining, broader and more rounded form with less posteriorly tapering elytra; by the form of the mesoventral keel which is more rounded anteriorly and with a raised border (Figs. 5A /B, katyae Fig. 5 F) and by the form of the lateral points on the posterior intercoxal process of the metaventrum which are broader basally and much less sharply pointed (Fig. 5D, katyae Fig. 5 G) For differences of N. schuelkei and N. katyae with N. pilosellum, see Darby (2015).</p><p>Type data. Holotype: ♀, Costa Rica, Osa Peninsula, Road, Bahia Drake to Rincon de Osa, forest and plantations, car net devart, 8.41–42 N 83.29 –40W, 0–400m, 9.x.2012, 1500–1700 hrs, Schülke &amp; Grünberg (INBio) . Paratype: ♀, same data as holotype (INBio) .</p></div>	https://treatment.plazi.org/id/039E87C6FFFBE466D8BFD0FCFD03FCC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Darby, Michael	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFCE467D8BFD196FB7DFE75.text	039E87C6FFFCE467D8BFD196FB7DFE75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptenidium Erichson 1845	<div><p>Ptenidium Erichson 1845</p><p>Blackwelder (1944–57) lists 8 species of Ptenidium occurring in Central America: concinnum from St. Vincent and Grenada, foveatum from Mexico and Guatemala, fuscipenne from Panama, nitens from Nicaragua, and ignobile, impunctatum, obesum and strangulatum from Guatemala . With the exception of fuscipenne described by Motschulsky, all these specimens were collected by G. Champion and described by A. Matthews (1872). The types of all the Matthews species are in BMNH and were examined and found to be different species from the Costa Rican specimens. The only other species of Ptenidium known from Central and South America are those from Peru described by Darby 2016, who erected a new subgenus, Peruvium, to accomodate several species bearing patches of posteriorly directed setae at the hind angles of the pronotum and globular spermathecae.</p></div>	https://treatment.plazi.org/id/039E87C6FFFCE467D8BFD196FB7DFE75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Darby, Michael	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFDE467D8BFD11DFE85F852.text	039E87C6FFFDE467D8BFD11DFE85F852.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bambara invisibilis Nietner 1856	<div><p>Bambara invisibilis Nietner 1856</p><p>8 examples of this very common, worldwide species. Costa Rica, Osa Peninsula, Road, Bahia Drake to Rincon de Osa, forest and plantations, car net devart, 8.41–42 N 83.29 –40W, 0–400m, 9.x.2012, 1500–1700 hrs, Schülke &amp; Grünberg (INBio).</p></div>	https://treatment.plazi.org/id/039E87C6FFFDE467D8BFD11DFE85F852	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Darby, Michael	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFDE467D8BFD57EFC2AF9DA.text	039E87C6FFFDE467D8BFD57EFC2AF9DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptenidium gruenbergae	<div><p>Ptenidium gruenbergae sp.n.</p><p>(Figs. 6 A–J)</p><p>Length 0.80 mm. Dark, shining, appearing polished, with very short, barely visible, scattered pubescence. Antennomeres 3–8 length 0.33 mm, Fig. 6B. Head with longer pubescence and a group of much longer setae medially on the anterior margin, the central stouter ones appearing to form a bifid process, Fig. 6E. Width across eyes 0.32 mm. Mentum with rounded lateral margins, Fig. 6G. Pronotum length 0.24 mm, width 0.45 mm, with a small group of setae on the posterior margin at the corners, Fig. 6H; without any fovea on the anterior and posterior margins. Scutellum with a depression at the anterior corners and a short median keel with rounded margins, Fig. 6J. Elytra length 0.54 mm, width 0.47 mm. Proventral process parallel sided, rounded posteriorly and with a shallow median depression, Fig. 6F. Mesoventral collar and keel, Fig. 6I.</p><p>Male: aedeagus Figs. 6 Ca/b.</p><p>Female: spermatheca Fig. 6D.</p><p>Etymology. Named after the collector Brigitte Grünberg (Berlin).</p><p>Diagnosis. The only other described species in this subgenus, which is typified by the patches of setae on the pronotal posterior angles and the form of the genitalia, are from Peru. P. gruenbergae can be separated from those by the group of setae on the front margin of the head.</p><p>Type data. Holotype: ♂, Costa Rica, Osa Peninsula, Road, Bahia Drake to Rincon de Osa, forest and plantations, car net devart, 8.41–42 N 83.29 –40W, 0–400m, 9.x.2012, 1500–1700 hrs, Schülke &amp; Grünberg (INBio) . Paratypes: 6 examples, same data as holotype (INBio, BMNH) .</p></div>	https://treatment.plazi.org/id/039E87C6FFFDE467D8BFD57EFC2AF9DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Darby, Michael	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFDE467D8BFD438FE0BFD37.text	039E87C6FFFDE467D8BFD438FE0BFD37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ptenidium nitidum Heer 1841	<div><p>Ptenidium nitidum Heer 1841</p><p>A single specimen of this common and widespread species. Costa Rica, Osa Peninsula, Road, Bahia Drake to Rincon de Osa, forest and plantations, car net devart, 8.41–42 N 83.29 –40W, 0–400m, 9.x.2012, 1500–1700 hrs, Schülke &amp; Grünberg (INBio).</p></div>	https://treatment.plazi.org/id/039E87C6FFFDE467D8BFD438FE0BFD37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Darby, Michael	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFEE465D8BFD19CFCB6FF28.text	039E87C6FFFEE465D8BFD19CFCB6FF28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Petrotrichis rotundata Darby 2015	<div><p>Petrotrichis rotundata Darby 2015</p><p>A single example of this species. The only other specimens known to date are from Bolivia. Costa Rica, Osa Peninsula, Road, Bahia Drake to Rincon de Osa, forest and plantations, car net devart, 8.41–42 N 83.29 –40W, 0– 400m, 9.x.2012, 1500–1700 hrs, Schülke &amp; Grünberg (INBio).</p></div>	https://treatment.plazi.org/id/039E87C6FFFEE465D8BFD19CFCB6FF28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Darby, Michael	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFF0E46AD8BFD68DFB4AFA68.text	039E87C6FFF0E46AD8BFD68DFB4AFA68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Seminis	<div><p>Seminis gen. n.</p><p>(Figs. 7 A–K)</p><p>From the Latin noun semen meaning a plant seed, gender neuter. Type species Seminis factiosum sp. n.</p><p>Description. Small, rounded, pubescent species easily mistaken for a plant seed. Colour dark brown. Antennomeres 11 segmented, antennomeres 1 and 2 larger than 3–8; 4–8 elongate (5 &amp; 7 longer than remainder), 9–11 forming a loosely jointed club, Fig. 7B. Eyes present. Mentum +/– square. Pronotum without obvious reticulation, slightly wider than elytra, hind angles smoothly rounded in lateral view. Scutellum triangular without any distinctive features. Elytra widest at apex of scutellum, with a short narrow epimeron at the humerus, Fig. 7I. Proventrum narrow, coxae almost contiguous and almost reaching anterior margin, coxal cavities open behind. Mesoventrum, clearly reticulate, collar strongly curved posteriorly at humeri, lateral margins very narrow, hind margins directed anteriorly from procoxal margin, keel reaching beneath mesocoxae in life, ventral surface narrowly eliptical, pubescent, flattened, Figs.7E, F. Metascutellum with two marginal spines Fig. 7K. Metaventrum clearly reticulate, posterior with two sharp points at corners, Fig. 7F. Abdomen: reticulation of ventrites distinct, cells bearing a single posteriorly directed seta alternating with others fringed with microtrichiae, Fig. 7H; tergites II–V with wing folding spicule patches, palisade fringe on tergite VII absent; pygidium composed of tergites IX–X fused, with small apical and two lateral points on posterior margin, anterior margin with row of setae. Fig. 7J; glands as described by Hall (2003) absent. Wings of usual ptiliid type.</p><p>Separation from related species. The distinctive spatulate form of the mesoventral keel is similar to that of Nossidium katyae, species of the genus Kuschelidium Johnson and, in a reduced form, to those of Limulopteryx Hall. Of these Seminis is only likely to be confused, if the ventral characters are examined, with Kuschelidium . From that genus it may be separated by the absence of antennal grooves on the proventrum, the shorter elytra, the absence of short carinae running posteriad from the mesocoxal cavities, and the widely separated metacoxae. A similar spatulate mesoventral keel is also present in members of the subfamily Cephaloplectinae but in that case it is a feature of the proventrum and not of the mesoventrum.</p><p>Kuschelidium was placed by Johnson in the Ptinellini, and by Sawada and Hirowatari (2002) close to Mikado (in the Nanosellini) although their decision was based on Japanese specimens which differ markedly from K.maori Johnson, the type species of the genus. On ventral characters in particular Seminis looks closest to Acrotrichis and its place in the subfamily Acrotrichinae appears to be confirmed by the presence of the lateral spines on the metascutellum noted by Hall to be absent in all other ptiliid subgroups (Hall 2003, 2005). The symmetrical form of the aedeagus, the incised sternite VI, the form of sternite VII with a short apodeme and the fused tergites IX and X forming the pygidium support that, but it should also be noted that the palisade fringe on tergite VII present in Acrotrichis is absent in Seminis .</p><p>Species of Limulopteryx and the Cephaloplectinae are known to be termitophilous, and the possibility exists that species of Seminis (and Nossidium katyae) may also be associated with termites or ants.</p></div>	https://treatment.plazi.org/id/039E87C6FFF0E46AD8BFD68DFB4AFA68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Darby, Michael	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFF0E46BD8BFD3F3FF1BFF05.text	039E87C6FFF0E46BD8BFD3F3FF1BFF05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Seminis factiosum	<div><p>Seminis factiosum sp. n.</p><p>(Figs 7A–K)</p><p>Habitus, Fig. 7A. Length 0.89 mm. Colour dark brown. Pubescence, antennae and legs dusky yellow. Antennomeres 3–11 length 0.34 mm, Fig. 7B. Mentum and submentum chaetotaxy Fig. 7G. Width across eyes 0.38 mm. Pronotum 0.60 mm wide, 0.43 mm long. Elytra 0.41 mm long, 0.57 mm wide, detached elytron Fig. 7I. Mesoventrum with strong downwardly projecting keel between mesocoxae in dorsal view, Figs. 7E,F. Metascutellum Fig. 7K. Metaventrum clearly reticulate, posterior margin between metacoxa +/– 1/4 the width of the sclerite, straight with two sharp points at corners, Fig. 7F. Abdominal reticulation with alternate cells of single setae and microtrichiae, Fig. 7H. Pygidium, Fig. 7J. Wings of usual Ptiliid type.</p><p>Male: aedeagus, Figs. Ca/b. Ventrite VI with semicircular incision, ventrite VII with anteriorly projected apodeme, Fig. 7D.</p><p>Female: unknown.</p><p>Etymology. From the Latin adjective factiosus meaning intriquing.</p><p>Type data. Holotype: ♂, Costa Rica, Osa Peninsula, Road, Bahia Drake to Rincon de Osa, forest and plantations, car net devart, 8.41–42 N 83.29 –40W, 0–400m, 9.x.2012, 1500–1700 hrs, Schülke &amp; Grünberg (INBio).</p></div>	https://treatment.plazi.org/id/039E87C6FFF0E46BD8BFD3F3FF1BFF05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Darby, Michael	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFF1E46BD8BFD756FAECFE57.text	039E87C6FFF1E46BD8BFD756FAECFE57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acrotrichis Motschulsky 1848	<div><p>Acrotrichis Motschulsky, 1848</p><p>More than 400 specimens including several new species which will be dealt with in a separate paper.</p></div>	https://treatment.plazi.org/id/039E87C6FFF1E46BD8BFD756FAECFE57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Darby, Michael	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
