taxonID	type	description	language	source
039E87C6FFF9E461D8BFD153FAB8FE51.taxon	description	Habitus Fig. 3 Aa / b. Length 0.96 mm. Body yellowish brown covered with grey / yellow overlapping pubescence, legs and antennae dusky yellow. Antennomeres 3 – 8 length 0.4 mm; antennomeres 4 – 8 elongate, 10,11 globular, with sensillae. Width across eyes 0.38 mm. Mentum broadest at anterior margin tapering towards base, two longest setae situated medially on lateral margins, Fig. 3 F. Pronotum length 0.33 mm, width 0.63 mm, posterior margin sinuate before the angles. Elytra length 0.56 mm, width 0.63 mm, widest just behind humeri. Wings of usual Nossidium type, Fig. 3 H. Mesoventral collar reaching onto the humeri with a raised anterior border and a vertically oriented medial ridge, Fig. 3 G; mesoventral keel wide between the mesocoxae, Fig. 3 D. Metaventrum short, pubescence denser on disc, posterior intercoxal process comprised of two short, sharp points, Fig. 3 E. Male: not known. Female: spermatheca globular, Fig. 3 C.	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFF9E461D8BFD153FAB8FE51.taxon	etymology	Etymology. Named after Miss Isobel ' Issy' Millett, a close friend of the author’s family.	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFF9E461D8BFD153FAB8FE51.taxon	diagnosis	Diagnosis. The new species may be separated from N. schuelkei sp. n. and N. katyae by the form of the mesoventral keel which does not extend below the mesocoxae. In this respect it is similar to N. pilosellum, and the North American species N. americanum Motschulsky and N. amoenum Motschulsky but these are less rounded and more parallel sided species.	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFF9E461D8BFD153FAB8FE51.taxon	materials_examined	Type data. Holotype: ♀, Costa Rica, Alajuela, Peñas Blancas, 820 m, 19 May 1989, J. Ashe, R. Brooks, R. Leschen; Snow Entomological Museum, Costa Rican Expedition, # 279; ex. with ants under bark (MM). Fragments of a second specimen from the same locality, ex. Xylaria, 17 May 1989, mounted as a slide, are also in MM.	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFBE466D8BFD0FCFD03FCC5.taxon	description	Length 0.84 mm. Body dark brown covered with grey / yellow overlapping pubescence, legs and antennae dusky yellow, lateral and posterior borders with a sharply defined yellow margin in transmitted light. Antennomeres 3 – 8 length 0.33 mm, 4 – 8 elongate, 10,11 globular, Fig. 4 D. Width across eyes 0.31 mm. Mentum broadest at anterior margin tapering towards base, two longest setae situated medially on lateral margins, Fig. 5 C. Pronotum length 0.24 mm, width 0.50 mm, posterior margin sinuate before the angles. Elytra length 0.56 mm, width 0.53 mm, widest just behind humeri, wings of usual Nossidium type, Fig. 4 E. Mesoventral collar reaching onto the humeri with a raised anterior border and a vertically oriented medial ridge, Fig. 5 B; mesoventral keel wide, reaching below the mesocoxae in dorsal view and with a flattened cap rounded anteriorly and with a raised border, closely fitting posteriorly with the metaventrum so as to appear an anterior metaventral process, Fig. 5 A. Metaventrum with line of metepisternal suture reaching mesocoxae but not directed posteriad at lateral margins; lateral points on the posterior intercoxal process broadened and flattened towards the base with convex lateral margins, Fig. 5 F. Pygidium with + / – 15 teeth Fig. 5 G. Male: not known. Female: spermatheca Fig. 4 F.	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFBE466D8BFD0FCFD03FCC5.taxon	etymology	Etymology. Named after the collector Michael Schülke of the Museum für Naturkunde, Berlin.	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFBE466D8BFD0FCFD03FCC5.taxon	diagnosis	Diagnosis. The new species is similar to N. katyae but may be distinguished by its less shining, broader and more rounded form with less posteriorly tapering elytra; by the form of the mesoventral keel which is more rounded anteriorly and with a raised border (Figs. 5 A / B, katyae Fig. 5 F) and by the form of the lateral points on the posterior intercoxal process of the metaventrum which are broader basally and much less sharply pointed (Fig. 5 D, katyae Fig. 5 G) For differences of N. schuelkei and N. katyae with N. pilosellum, see Darby (2015).	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFBE466D8BFD0FCFD03FCC5.taxon	materials_examined	Type data. Holotype: ♀, Costa Rica, Osa Peninsula, Road, Bahia Drake to Rincon de Osa, forest and plantations, car net devart, 8.41 – 42 N 83.29 – 40 W, 0 – 400 m, 9. x. 2012, 1500 – 1700 hrs, Schülke & Grünberg (INBio). Paratype: ♀, same data as holotype (INBio).	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFDE467D8BFD57EFC2AF9DA.taxon	description	Length 0.80 mm. Dark, shining, appearing polished, with very short, barely visible, scattered pubescence. Antennomeres 3 – 8 length 0.33 mm, Fig. 6 B. Head with longer pubescence and a group of much longer setae medially on the anterior margin, the central stouter ones appearing to form a bifid process, Fig. 6 E. Width across eyes 0.32 mm. Mentum with rounded lateral margins, Fig. 6 G. Pronotum length 0.24 mm, width 0.45 mm, with a small group of setae on the posterior margin at the corners, Fig. 6 H; without any fovea on the anterior and posterior margins. Scutellum with a depression at the anterior corners and a short median keel with rounded margins, Fig. 6 J. Elytra length 0.54 mm, width 0.47 mm. Proventral process parallel sided, rounded posteriorly and with a shallow median depression, Fig. 6 F. Mesoventral collar and keel, Fig. 6 I. Male: aedeagus Figs. 6 Ca / b. Female: spermatheca Fig. 6 D.	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFDE467D8BFD57EFC2AF9DA.taxon	etymology	Etymology. Named after the collector Brigitte Grünberg (Berlin).	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFDE467D8BFD57EFC2AF9DA.taxon	diagnosis	Diagnosis. The only other described species in this subgenus, which is typified by the patches of setae on the pronotal posterior angles and the form of the genitalia, are from Peru. P. gruenbergae can be separated from those by the group of setae on the front margin of the head.	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFFDE467D8BFD57EFC2AF9DA.taxon	materials_examined	Type data. Holotype: ♂, Costa Rica, Osa Peninsula, Road, Bahia Drake to Rincon de Osa, forest and plantations, car net devart, 8.41 – 42 N 83.29 – 40 W, 0 – 400 m, 9. x. 2012, 1500 – 1700 hrs, Schülke & Grünberg (INBio). Paratypes: 6 examples, same data as holotype (INBio, BMNH).	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFF0E46AD8BFD68DFB4AFA68.taxon	description	From the Latin noun semen meaning a plant seed, gender neuter. Type species Seminis factiosum sp. n. Description. Small, rounded, pubescent species easily mistaken for a plant seed. Colour dark brown. Antennomeres 11 segmented, antennomeres 1 and 2 larger than 3 – 8; 4 – 8 elongate (5 & 7 longer than remainder), 9 – 11 forming a loosely jointed club, Fig. 7 B. Eyes present. Mentum + / – square. Pronotum without obvious reticulation, slightly wider than elytra, hind angles smoothly rounded in lateral view. Scutellum triangular without any distinctive features. Elytra widest at apex of scutellum, with a short narrow epimeron at the humerus, Fig. 7 I. Proventrum narrow, coxae almost contiguous and almost reaching anterior margin, coxal cavities open behind. Mesoventrum, clearly reticulate, collar strongly curved posteriorly at humeri, lateral margins very narrow, hind margins directed anteriorly from procoxal margin, keel reaching beneath mesocoxae in life, ventral surface narrowly eliptical, pubescent, flattened, Figs. 7 E, F. Metascutellum with two marginal spines Fig. 7 K. Metaventrum clearly reticulate, posterior with two sharp points at corners, Fig. 7 F. Abdomen: reticulation of ventrites distinct, cells bearing a single posteriorly directed seta alternating with others fringed with microtrichiae, Fig. 7 H; tergites II – V with wing folding spicule patches, palisade fringe on tergite VII absent; pygidium composed of tergites IX – X fused, with small apical and two lateral points on posterior margin, anterior margin with row of setae. Fig. 7 J; glands as described by Hall (2003) absent. Wings of usual ptiliid type. Separation from related species. The distinctive spatulate form of the mesoventral keel is similar to that of Nossidium katyae, species of the genus Kuschelidium Johnson and, in a reduced form, to those of Limulopteryx Hall. Of these Seminis is only likely to be confused, if the ventral characters are examined, with Kuschelidium. From that genus it may be separated by the absence of antennal grooves on the proventrum, the shorter elytra, the absence of short carinae running posteriad from the mesocoxal cavities, and the widely separated metacoxae. A similar spatulate mesoventral keel is also present in members of the subfamily Cephaloplectinae but in that case it is a feature of the proventrum and not of the mesoventrum.	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFF0E46BD8BFD3F3FF1BFF05.taxon	description	Habitus, Fig. 7 A. Length 0.89 mm. Colour dark brown. Pubescence, antennae and legs dusky yellow. Antennomeres 3 – 11 length 0.34 mm, Fig. 7 B. Mentum and submentum chaetotaxy Fig. 7 G. Width across eyes 0.38 mm. Pronotum 0.60 mm wide, 0.43 mm long. Elytra 0.41 mm long, 0.57 mm wide, detached elytron Fig. 7 I. Mesoventrum with strong downwardly projecting keel between mesocoxae in dorsal view, Figs. 7 E, F. Metascutellum Fig. 7 K. Metaventrum clearly reticulate, posterior margin between metacoxa + / – 1 / 4 the width of the sclerite, straight with two sharp points at corners, Fig. 7 F. Abdominal reticulation with alternate cells of single setae and microtrichiae, Fig. 7 H. Pygidium, Fig. 7 J. Wings of usual Ptiliid type. Male: aedeagus, Figs. Ca / b. Ventrite VI with semicircular incision, ventrite VII with anteriorly projected apodeme, Fig. 7 D. Female: unknown.	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFF0E46BD8BFD3F3FF1BFF05.taxon	etymology	Etymology. From the Latin adjective factiosus meaning intriquing.	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
039E87C6FFF0E46BD8BFD3F3FF1BFF05.taxon	materials_examined	Type data. Holotype: ♂, Costa Rica, Osa Peninsula, Road, Bahia Drake to Rincon de Osa, forest and plantations, car net devart, 8.41 – 42 N 83.29 – 40 W, 0 – 400 m, 9. x. 2012, 1500 – 1700 hrs, Schülke & Grünberg (INBio).	en	Darby, Michael (2016): New Species and Records of Costa Rican Featherwing Beetles (Coleoptera: Ptiliidae). Zootaxa 4184 (1): 41-51, DOI: 10.11646/zootaxa.4184.1.2
