taxonID	type	description	language	source
039E355D6420FFAD8F9CF94CFB48FBAF.taxon	distribution	Distribution. Species amply distributed from the Sierra Madre Occidental in the Mexican states of Sonora and Chihuaha to the north of Nicaragua, following the Pacific mountain chains in Central America (Fig. 5, black circles). C. multipustulata is distributed in the Mexican Transition and Caribbean Mesoamerican domains.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6420FFAD8F9CF94CFB48FBAF.taxon	materials_examined	Material examined (192 specimens). GUATEMALA NHM: (1) one specimen: Guatim, multipustulata Stål, Type Stål Coll: Deyrolle, Baly Coll., multipustulata Stål Mexico, Guatemala [underneath: Type Stål Col. Deyrolle]. OUMNH: (1) one specimen: V. de Atitlan 25 – 3000 ft. Champion, B. C. A. duplicates pres. 1909 by F. D. Godman Cat. No. 84, Calligrapha multipustulata Stål J. Gómez- Zurita det. 2012. HONDURAS NMNH: (1) one specimen: 11964, 30 - 12 - 77, Muchilena, Hond., J. V. Mankins Collector, Calligrapha diversa Det. 1977, J. V. Mankins, Calligrapha multipustulata Stål J. Gómez-Zurita det. 2011. MEXICO EGRC: (1) one specimen: Mexico, Jalisco, 5.5 mi NE Tecolotlán, 13 vii 1982, A. J. Gilbert, det. Daccordi ’ 86; (2) one specimen: Mexico, Cuernavaca, Dr. A. Fenyes; (3) two specimens: Mexico, Chiapas, El Aguacero, 14 vi 1988, D. B. & A. M. Thomas, Calligrapha multipustulata Stål J. Gómez-Zurita det. 2011; (4) ten specimens 1: Mexico, Morelos, Cañón de Lobos, 19 km E Cuernavaca, 1220 – 1375 m, 3 vii 1992, C. Bellamy; (5) one specimen: Mexico, Puebla, 5 km NW Petlacingo 1400 m, 7 vii 1992, C. Bellamy, Calligrapha multipustulata Stål J. Gómez-	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6420FFAD8F9CF94CFB48FBAF.taxon	description	Variation. Some specimens seem to be less melanised than the type in several respects, not matching the most usual configuration of markings for the species, including a largely separated humeral spot and lunule (seen in the Guatemalan specimen at NHM, and also in NMB specimens at G. Frey collection), which are most usually confluent at least for 2 / 3 of the length of the humeral spot; the humeral markings nearly always reach the basal margin of elytra, even if narrowly in some specimens. In one specimen from Guadalajara, Mexico (MZC), the basal part of the humeral spot is missing, but not so the apical part, and the humeral lunule is also split in two halves, so that the humeral marking appears as three free spots at obtuse angles. The spot enclosed by the humeral lunule can have its two constituent longitudinal spots free, with only apices confluent forming a U shape, with only basal ends free appearing as an emargination, or closed at both ends but not in the middle, enclosing a small pale marking. At least in two specimens at MfN and two at ZSM the midlateral spot is very reduced, almost obsolete, and those at NMB have this spot reduced and only narrowly touching the margin of elytron; in the specimens from Sinaloa (NMNH), slightly different from typical C. multipustulata [I originally determined some of them as C. geographica], this feature is large and confluent with elytral margin, but it is narrowed before reaching it. A few specimens have a slightly violaceous tint to dark parts instead of the typical greenish bronze.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6427FFB38F9CFF06FAE0FC51.taxon	materials_examined	Holotype: Oaxaca / Sallé / Type / Typus [red] (NRM). The holotype lacks seven apical antennomeres of right antenna and the claws of the right mesotarsus appear broken. Habitus (Fig. 3 a). Length: 9.00 mm, width: 5.86 mm. Body elongated oval, moderately convex. Head, base of mandibles, pronotum, scutellum, ventral surfaces and legs black with bronzy metallic gloss. Labrum, basal antennomeres ventrally, apex of clypeus, apex of mandibles, knees, claws, epipleura and elytral markings dark rufous. Apical antennomeres and dorsum of basal antennomeres darkened. Abdominal segments with rufous tint. Ground of elytra creamy yellow. Head sparsely punctured, smooth on antennal calli; supraocular sulci impressed, long, from antennal calli reaching well behind the eye; clypeal suture broadly U-shaped. Antennae slender but short, reaching humeri; first antennomere long, thick, posteriorly curved; antennomeres 2 – 6 elongated, thin, relatively smooth and nearly glabrous, second less than half as long as first, third nearly as long as first; antennomeres 3 – 6 shortening progressively, but sixth longer than second, and segments lengthening gradually beyond sixth antennomere, latest longer than first; antennomeres 7 – 11 thicker, and 8 – 11 rugose and pubescent; eighth antennomere 0.65 x as wide as long. Labrum relatively short, emarginate anteriorly. Mandibles strong, largely protruding beyond labrum; external border feebly concave. Last segment of maxillary palpi large, with sides divergent, truncated obliquely at apex. Pronotum transverse (W / L = 1.77), margined laterally and apically; surface microreticulate; rather densely, strongly punctured on disc, punctures larger, confluent towards sides. Hypomeral suture strongly impressed, not reaching base of pronotum basaly and gradually diverging from pronotal border towards apex; hypomera shagreened, unpunctured. Prosternal process truncated at apex; longitudinally carinated. Metanepisterna strongly punctured. Metaventrite with few relatively strong basal punctures, homogeneously sparse fine punctures elsewhere. Scutellum short (W / L = 0.96), flat and smooth, microreticulate. Elytral punctures strong around and within markings; pale surface almost unpunctured except for premarginal dense row of finer punctures; scutellar row present. Elytral margin dark; numerous dark markings on elytra, most prominent being: (i) sutural stripe entire, width nearly constant from base, surrounding scutellum, to sudden preapical narrowing; (ii) subsutural stripe entire, confluent with sutural stripe from scutellar area nearly to elytral apex; (iii) basal arm of arcuate band slightly oblique, largely confluent with subsutural stripe; apical arm of arcuate band slightly divergent from suture as irregular spot connected to subsutural stripe; (iv) humeral spot large, irregularly delimited apically owing to confluence with additional minor markings; (v) humeral lunule large, confluent with humeral spot at basal half; apical half obliquely produced towards suture, surpassing posteriorly basal end of arcuate band; (vi) spot enclosed by humeral lunule large, occupying most of space between lunule and subsutural stripe, ovoid with loosely defined basal margin and enclosing a small pale spot; (vii) spot of apical declivity large, irregular, connected to subsutural stripe by dark suffusions; (viii) apical spot present and irregularly delimited; (ix) midlateral spot elliptical and elongated (covering eight premarginal line punctures), free from elytral dark margin; (x) most of free area of elytra occupied by many marks and spots of heterogeneous sizes and shapes; pale area among spots usually as wide as diameter of punctures surrounding markings. Tibiae externally canaliculated from 1 / 2 to 1 / 3 of their length, with some longitudinal ridges within. Abdominal ventrites finely punctured. Penis figured in Figs 4 c, 4 f.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6427FFB38F9CFF06FAE0FC51.taxon	distribution	Distribution. This species is restricted to the Pacific slopes of the Sierra Madre del Sur in the states of Guerrero and Oaxaca in Mexico (Fig. 5, white circles). This species is only found in the Caribbean Mesoamerican domain.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6427FFB38F9CFF06FAE0FC51.taxon	materials_examined	Material examined (21 specimens). MEXICO EGRC: (1) one specimen: Mexico, Oaxaca, 5 mi S Candelaria Loxicha, 18 – 19. vii. 1974, Clark, R. R. Murray, J. S. Ashe, Joseph C. Schaffner coll. MCZ: (1) one specimen: Panistlahuca, Mexico, Salle Coll., 1 st Jacoby Coll. MfN: (1) one specimen: Mexico, v. Studt, Calligrapha consputa det. Daccordi ' 80. NHM: (1) one specimen: named by Stål, Baly Coll., consputa Stål Mexico; (2) two specimens: Oax., Baly Coll.; (3) one specimen: Oax., 143, consputa Stål, 67 - 56; (4) one specimen: Mex., consputa Stål, 67 - 56; (5) one specimen: Panistlahuca, Mexico, Sallé Coll., Calligrapha consputa Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (6) one specimen: Panistlahuca, 640, Calligrapha consputa Stål apud Sallé, Sp. figured, Godman-Salvin Coll., Biol. Centr. - Amer. NMB: (1) one specimen: Mexico, Calligrapha consputa Stal J. Bechyne det. 1954 2. NRM: (1) one specimen: Oaxaca, Sallé. TAMUIC: (1) two specimens: [X 0536352 and X 0538391], Mexico, Oaxaca, 5 mi S Candelaria Loxicha, 18 – 19. vii. 1974, Clark, R. R. Murray, J. S. Ashe, Joseph C. Schaffner coll.; (2) two specimens: [X 0536353 and X 0547093], Mexico, Guerrero, Hwy 95 on Hwy 200, 2.8 km W Veintidos, James E. Wappes; (3) one specimen: X 0548166, Mexico, Oaxaca, 4.7 mi S San Gabriel Mixtepec, 16. vii. 1985, Robert W. Jones & Joseph C. Schaffner. UNKNOWN SOURCE MCZ: (1) one specimen: Calligrapha signata Ol., diversa Dj., Hagewisch, Calligrapha consputa Stål J. Gómez-Zurita det. 2010; (2) one specimen: Calligrapha consputa Stål J. Gómez-Zurita det. 2011; (3) one specimen: 172, CM, ex. Chev. coll.?, Calligrapha consputa Stål J. Gómez-Zurita det. 2011.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6427FFB38F9CFF06FAE0FC51.taxon	description	Variation. The pale minute spot within the spot enclosed by humeral lunule can be absent, but it is always indicated by the punctures associated with it. The midlateral spot can be narrow, as a dark line covering a line of punctures of the premarginal row, or more rarely, in particularly dark specimens, can emit dark suffusions laterally confluent with dark margin of elytra. Pale surface of elytra can be greatly reduced by spot confluence.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6438FFB18F9CFB80FC7FFC1A.taxon	distribution	Distribution. This subspecies of C. geographica occupies the northern parts of the species range in the southern states of Chiapas, Tabasco and Veracruz in Mexico (Fig. 5, white squares). This is an endemic subspecies of the Caribbean Mesoamerican region.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6438FFB18F9CFB80FC7FFC1A.taxon	materials_examined	Material examined (40 specimens). MEXICO EGRC: (1) one specimen: Mexico, Chiapas, E Tuxtla Gtz, El Chorreadero, 13 / 06 / 1989, P. K. Lago & S. Testa, Calligrapha consputa Stal det. Daccordi ’ 86, Calligrapha geographica Stål J. Gómez-Zurita det. 2011. MCZ: (1) one specimen: Calligrapha geographica Stal, Mex., Jacoby 2 nd Coll.; (2) one specimen: Cosamaloapam, Mexico, Salle Coll., 1 st Jacoby Coll.; (3) one specimen: 96, Mexico, M. Chevrolat, Jacoby 2 nd Coll.; (4) one specimen: Jalapa, Vera Cruz, Höge, Jacoby 2 nd Coll.; (5) one specimen: Chrysomela geographica Mex.; (6) one specimen: Mex. Chrysomela pardalina; (7) one specimen: Mex.; (8) one specimen: Mex., Calligrapha geographica Stal J. Gómez-Zurita det. 2010. MfN: (1) two specimens: Cosamaloapam 6, geographica St. [one specimen without data]. NHM: (1) one specimen: geographica Chevt. Mexico, Baly Coll.; (2) one specimen: E. Coll. Laferté, Mexico, 67 - 56; (3) one specimen: Ent. Club. 44 - 12, [...] Newman, Mexico; (4) two specimens: Mexico, Z. Walker, 6 35 13; (5) one specimen: ex Mus Murray, Mexico, Fry Coll. 1905.100; (6) one specimen: [female], geographica Chv t, Mexico, Alvarado Sallé var., 67 - 56; (7) one specimen: Mex., Baly Coll.; (8) one specimen: Playa Vicente, Mexico, Sallé Coll., Calligrapha geographica Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (9) one specimen: Cosamaloapan, Mexico, Sallé Coll., 641, Calligrapha geographica Stål apud Sallé, Sp. figured, Godman-Salvin Coll., Biol. Centr. - Amer. NMB: (1) one specimen: Mexico, Calligrapha geographica Stål, J. Bechyné det. 1957; (2) one specimen: Mexiko, Calligrapha geographica Stål J. Gómez-Zurita det. 2010. NMCZ: (1) one specimen: [female], geographica Chev., Mex., Coll. Achard Mus. Pragense; (2) one specimen: Mexique, geographica Stål, Coll. Achard Mus. Pragense; (3) one specimen: Tacotalpa, Coll. Achard Mus. Pragense. NRM: (1) one specimen: Mexic, Chevrol, Schh.; (2) one specimen: Mexico, Tarnier. OUMNH: (1) one specimen: F. Wallan Mexico, Ent. Club 44 - 12, From the Brit. Mus. duplicates Pres. 1899, Calligrapha geographica Stål J. Gómez-Zurita det. 2011. UNKNOWN SOURCE MfN: (1) two specimens: consputa [...] [one without data]. NHM: (1) one specimen: geographica, Baly Coll.; (2) one specimen: Mex., Baly Coll.; (3) two specimens: E. Coll. Laferté, geographica Chevrolat, 67 - 56; (4) one specimen: [male], 67 - 56. NMCZ: (1) one specimen: Calligrapha consputa Stål J. Achard det. in BCA, Coll. Achard Mus. Pragense.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6438FFB18F9CFB80FC7FFC1A.taxon	description	Variation. Species rather constant in general features and colouration, displaying some variation on confluence of elytral markings. Subsutural stripe can be slightly more divergent basally than in type or almost entirely confluent. The midlateral spot can be absent. The apex of the humeral spot can be free or narrowly joined to basalmost additional spot on disc. Apex of humeral lunule can be sometimes very narrowly confluent with base of the arcuate band, and more rarely with the apex of the spot enclosed by humeral lunule. The strongest variation is observed in size and number and confluence of additional spots on disc, since they can be broken in smaller spots or confluent among them or with the predominant features, like the innermost spot sometimes confluent very narrowly with the external concavity of arcuate band, as in the type.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D643AFFB78F9CFC45FCDCFF4F.taxon	distribution	Distribution. This subspecies of C. geographica occupies the southern parts of the species range in the southern states of Chiapas and Tabasco in Mexico (Fig. 5, black squares). Subspecies endemic of the Caribbean Mesoamerican domain.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D643AFFB78F9CFC45FCDCFF4F.taxon	materials_examined	Material examined (seven specimens). MEXICO EGRC: (1) one specimen: Mexico, Chiapas, Cuxtepeques area, 15 / 06 / 1987, D. B. Thomas & A. Mendoza, Calligrapha famularis Stål J. Gómez-Zurita det. 2011. NHM: (1) one specimen: Teapa, Tabasco, Feb., H. H. S., Calligr. multipustulata var., Godman-Salvin Coll., Biol. Centr. - Amer.; (2) one specimen: Teapa, Tabasco, Jan., H. H. S., Godman-Salvin Coll., Biol. Centr. - Amer.; (3) one specimen: Teapa, Pilate, Baly Coll. NRM: (1) one specimen: Mexico, Tarnier.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D643AFFB78F9CFC45FCDCFF4F.taxon	description	Variation. Apex of arcuate band subsquare; markings showing a lesser degree of confluence (e. g., spot of apical declivity and apical spot free from subsutural stripe).	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D643CFFB58F9CFEB6FBA3F9EE.taxon	distribution	Distribution. This species is known from a single locality in Panama, Bugaba (Fig. 7, white circle). This narrow endemism can be considered part of the Caribbean Mesoamerican region. Material examined (21 specimens). PANAMA NMCZ: (1) one specimen: Panama, Coll. Achard Mus. Pragense.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D643CFFB58F9CFEB6FBA3F9EE.taxon	description	Variation. The large posterior roundish lobe of the arcuate band can be apical instead of preapical, or subtriangular, directed outwards and backwards. The apical spot can be present, minute, or its position can be indicated by a golden small marking (as on the right elytron of type). Some of the small additional markings can be missing, and the spot of apical declivity sometimes is free from the subsutural stripe.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D643EFFB88F9CF956FDA1FB8A.taxon	materials_examined	Holotype: Mexique / 68 [small pink square] / ancoralis Stål / Type Stål Coll: Deyrolle / Baly Coll. / ancoralis Stål Mexico [handwritten underneath: Type by Stål Col Deyrolle] (NHM). Specimen lacks right mesotarsus. Habitus (Fig. 6 f). Length: 9.46 mm, width: 6.06 mm. Body elongated oval, moderately convex. Head, pronotum, scutellum, legs, ventral surfaces, elytral markings, fine margin of elytra and epipleura very dark reddish, almost black; antennae, labrum and mouth parts paler; mandibles darker, almost black at apex; ground of elytra dark yellow. Head large, deeply inserted in pronotum; rather strongly and evenly punctured on frons, clypeus and near eyes, unpunctured on antennal calli; supraocular sulci deep, straight, surpassing eye margin above; frontal suture joining broadly V-shaped frontoclypleal suture; clypeal margin feebly concave. Antennae relatively short, reaching humeri; first antennomere long, thick but narrower basally, posteriorly curved; antennomeres 2 – 5 elongated, thin, relatively smooth and nearly glabrous; second antennomere half as long as first, third slightly shorter than first; antennomeres 3 – 5 shortening progressively, sixth longer than second, and 5 – 7 subequal; antennomeres 7 – 11 thicker, widening from base to apex, rugose and pubescent, lengthening gradually beyond seventh antennomere; eleventh about as long as first, tappering towards blunt apex and slightly emarginate dorsally; eighth antennomere 0.8 x as wide as long. Labrum small, moderately emarginate apically. Mandibles strong, with straight sides before pronounced preapical curvature. Last maxillary palpomere subsquare, parallel-sided, obliquely truncated at apex; preceding palpomere slightly broader at apex than width of last segment, much narrower basally, outer edge regularly curved. Pronotum transverse (W / L = 1.93), wider at base than apically; surface very finely microreticulate, almost smooth, with moderately strong and evenly distributed punctures, disappearing narrowly at sides and on longitudinal disc line; sides and apical border finely margined. Hypomeral suture deep and entire, only interrupted near base of pronotum, and slightly and gradually diverging from pronotal border anteriorly; hypomera finely microreticulate, unpunctured, concave at base, convex anteriorly. Prosternum finely microreticulate, densely punctured except on two lateral transversal calli, each puncture with fine whitish seta; prosternal process narrow between procoxae, expanding apically to twice median width; apex feebly curved, almost straight. Mesanepisterna finely microreticulate, unpunctured. Metepisterna coarsely punctured. Metaventrite relatively strongly punctured on anterior half, gradually becoming almost unpunctured posteriorly. Scutellum narrower at base than long (W / L = 0.83), sides regularly curved towards a round pointed apex. Ground of elytra finely, rather densely punctured with dark minute spots, forming regular premarginal line laterally, slightly confused in humeral area; punctures somewhat stronger around predominant markings, arranged as: (i) sutural stripe entire from base of elytra, surrounding scutellum, to gradual preapical narrowing, almost disappearing at apex; (ii) subsutural stripe almost entirely confluent with sutural stripe, except briefly basally, starting at level with apex of scutellum; slightly enlarged preapically; (iii) arcuate band anchor-shaped [giving name to the species], entire, gently curved and connected laterally to subsutural stripe by long dark area for most of its length, except briefly basally and less so behind; apical end prolonged as broadly curved hook-shape, bent forward and confluent in posterior border with some small discal markings (two on the left elytron and three on the right elytron of type); (iv) humeral spot large, elongated and laterally confluent with (v) humeral lunule, both joining basal elytral margin at base; apical end of humeral lunule obliquely directed towards suture, ending near and slightly beyond basal tip of arcuate band; (vi) spot enclosed by humeral lunule large, broadly emarginate anteriorly and laterally confluent with humeral lunule, appearing as bifurcation of humeral marking; (vii) midlateral spot large, subrectangular, laterally confluent with dark margin of elytron; (viii) spot of apical declivity large, roundish, weakly connected to apical enlargement of subsutural stripe by areas of dark suffusion; (ix) apical spot small, roundish, free; (x) seven additional markings on disc, free or connected to prolongation of arcuate band; three forming diagonal row behind and parallel to apical arm of humeral lunule; two other aligned spots present laterally in apical declivity of elytron. Femora shiny, sparsely and finely punctured, with fine short whitish hairs; transversal section of tibiae roundish, slightly flattened externally with feebly marked longitudinal furrow for half their length. Abdominal ventrites delicately, rather densely punctured medially and at sides; first ventrite with several short longitudinal furrows at coxal margin. Penis figured in Figs 9 a, 9 d.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D643EFFB88F9CF956FDA1FB8A.taxon	distribution	Distribution. This species is widely distributed in the northern part of Mesoamerica, along both Sierra Madre chains and reaching the Transvolcanic Belt, from southern Arizona in the USA to the state of Veracruz in Mexico (Fig. 8, black circles). C. ancoralis is also present in the southern tip of Baja California in Mexico and it has been reported from the Lee County in Texas, although the latter is an extremely doubtful record. This species is one of the fews that has colonized the California-Rocky Mountain domain to the north, otherwise is mostly present in the Mexican Transition and Caribbean Mesoamerican zones.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D643EFFB88F9CF956FDA1FB8A.taxon	materials_examined	Material examined (201 specimens). MEXICO EGRC: (1) one specimen: Mexico, Nayarit, 3 km WNW Jala, 3800 ft, 24. vii. 93, C. Bellamy, tropical thorn forest, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (2) one specimen: Mexico, Baja California Sur, vic. Todos Santos, 30. viii. 1994, J. Huether, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (3) nine specimens: Mexico, Tamaulipas, Bocatoma, 7 km SSE Gomez Farias, 19 – 23. v. 1979, E. G. Riley, Calligrapha ancoralis Stål det. Daccordi ’ 79 [one with: Calligrapha ramulifera Stal det. Daccordi ’ 81, Calligrapha ancoralis Stål J. Gómez- Zurita det. 2011]; (4) four specimens: Mexico, Tamaulipas, Bocatoma, 6 mi S Gomez Farias, 12 – 23. v. 1979, Marlin E. Rice Coll.; (5) three specimens: Mexico, Tamaulipas, Bocatoma w. s., 7 km SSE Gomez Farias, 1. vi. 1982, R. Turnbow; (6) one specimen: Mexico, Tamaulipas, Bocatoma w. s., 7 km SSE Gomez Farias, 15. x. 1985, R. Turnbow; (7) one specimen: Mexico, Tamaulipas, Bocatoma area, 7 km SSE Gomez Farias, 1 – 4. vi. 1982, J. E. Wappes; (8) one specimen: Mexico, Veracruz, Puente Nacional, 25. x. 1979, J. E. Wappes, Calligrapha ancoralis Stål det. Daccordi ’ 81; (9) one specimen: Mexico, Sinaloa, 30 mi N Mazatlan, 18. viii. 1964, H. R. Burke & J. Apperson; (10) one specimen: Mexico, Sinaloa 10 mi NW Mazatlan, 15. viii. 1965, H. Burke & J. Meyer; (11) one specimen: Mexico, Tamaulipas, 5 mi N Llera, 4. vi. 1965, H. R. Burke, J. Meyer & J. C. Schaffner; (12) one specimen: Mexico, Nuevo León, Linares, 1. vii. 1973, taken at light, Mastro & Schaffner; (13) four specimens: Mexico, Nuevo León, 2400 ft, 16 mi W Linares, Hwy 58, 11. x. 1982, C. W. and L. O’Brien & G. Wibmer; (14) one specimen: Mexico, Nuevo León, 20 mi W Linares, 3250 ft, Hwy 58, 21. vii. 1982, C. and L. O’Brien & G. Wibmer; (15) two specimens: Mexico, Nuevo León, 15 mi W Linares, 1800 ft, Hwy 58, 21. vii. 1982, C. and L. O’Brien & G. Wibmer; (16) one specimen: Mexico, Tamaulipas, Grutas de Quintero, 8 mi S Ciudad Mante, 24. viii. 1965, Reiber & Hollingsworth; (17) one specimen: Mexico, Tamaulipas, 8 km E Campo, 16. x. 1985, R. Turnbow; (18) one specimen: Mexico, Veracruz, Puente Nacional, 17. viii. 1960, H. Howden, Calligrapha stillatipennis det. Daccordi ’ 86, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011. FSCA: (1) one specimen: Mexico, Sinaloa, 14 – 15 mi N Los Mochis, 16. vii. 1984, B. K. Dozier, Calligrapha ancoralis St. Det. J. Watts 1993; (2) one specimen: Mexico, Tamaulipas, Galeana, 31. v. 1982, R. Turnbow, Calligrapha ancoralis St. Det. E. G. Riley ’ 82; (3) one specimen: Mexico, San Luis de Potosí, El Salto Falls, 1. vii. 1968, Debra Weems; (4) one specimen: Mexico, Baja California Sur, Sa. Victoria trail, La Burrera-La Laguna 550 – 730 m, 13. x. 1978, Dozier & Wescott; (5) four specimens: Mexico, Sonora, 13 mi SE Los Alamos, 30. x. 1972, K. Stephan; (6) one specimen: Mexico, Veracruz, Puente Nacional, 25. x. 1979, J. E. Wappes, Calligrapha ancoralis Stal Det. E. G. Riley ’ 82; (7) four specimens: Mexico, Tamaulipas, Bocatoma Area, 7 km SSE Gomez Farias, 1 – 4. vi. 1982, J. E. Wappes; (8) one specimen: Mexico, Tamaulipas, 9 mi W of 85, on Ocampo rd, 16. x. 1985, R. Morris II, Calligrapha ancoralis (Stal) det. I. S. Askevold 1990; (9) one specimen: Mexico, San Luis de Potosi, 12 km E Ocampo, 16. x. 1985, R. Morris II; (10) two specimens: Mexico, Sonora, 25 km W Santa Ana Viejo, 31. vii. 1985, W. Opitz, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (11) two specimens: Mexico, Tamaulipas, Cd. Victoria, 1. x. 1985, H. Dozier, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (12) one specimen: Mexico, San Luis Potosi, El Salto Falls, 1. vii. 1968, P. Weems, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (13) one specimen: Sinaloa, Puente de Quemalo, 30. viii. 1962, W. Rossember, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011. IBE-JGZ: (1) one specimen: IBE-JGZ-C 328, Mexico, Tamaulipas, Ciudad Victoria, spring 2011, Alexey G. Moseyko leg. MCZ: (1) one specimen: Ventanas, Durango, Höge, Calligrapha ancoralis; (2) eight specimens: Ventanas, Durango, Höge, Jacoby 2 nd Coll. [one with: Calligrapha ancoralis Stål J. Gómez-Zurita det. 2010]; (3) one specimen: Jalapa, Mexico, Hoege, 1 st Jacoby Coll.; (4) one specimen: Mexico, Sallé Coll., 1 st Jacoby Coll.; (5) one specimen: Ventanas, Mex. 2000 ft, Forres, Jacoby 2 nd Coll.; (6) one specimen: Guadaljiro [Guadalajara], Mexico; (7) two specimens: Guadalajara, Mexico, Dr. Buller, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2010; (8) one specimen: Sierra San Lazaro, Horn Coll., C. Calligr. ancoralis. MfN: (1) one specimen: Ventanas; (2) one specimen: Canel., Canelas Becker; (3) one specimen: Ventanas, Durango, Höge, ancoralis Stål; (4) one specimen: 585, San Andres de la Sierra, Calligrapha ancoralis det. Daccordi ' 80; (5) one specimen: Ven., suffriani Jac., Caligrapha ancoralis Stål 1860, J. Gómez-Zurita det. 2009. NHM: (1) one specimen: E. Coll. Thomson, Mexiq., 182; (2) two specimens: Ventanas, Mex., 2000 ft, Forrer, Godman-Salvin Coll., Biol. Centr. - Amer.; (3) seven specimens: Ventanas, Durango, Höge, Godman-Salvin Coll., Biol. Centr. - Amer.; (4) one specimen: Jalapa, Mexico, Hoege, Godman-Salvin Coll., Biol. Centr. - Amer.; (5) one specimen: Mexico, Sallé Coll., 646, Calligrapha ancoralis Stål, Godman-Salvin Coll., Biol. Centr. - Amer.; (6) one specimen: Ex Coll. J. Sturm, Mexico, C. imperialis Mihi, Godman-Salvin Coll., Biol. Centr. - Amer.; (7) one specimen: Mex., Baly Coll. NMB: (1) one specimen: Ventanas, Durango, Höge, Calligrapha ancoralis Stal J. Bechyné det. 1951; (2) one specimen: Mexico, 1888, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2010; (3) two specimens: Mazatlan, Sin., Mex., x. 1958, leg. G. F r ey. NMNH: (1) one specimen: on orchid foliage, Mex., Feb. 18 ’ 39, Laredo Tex. # 15715, Calligrapha notatipennis (Stål) HSB 1939, return to the national collection, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (2) one specimen: Venodio [El Venadillo], Sinaloa, Mex., Kusche, vii. 31. ’ 18, F. Monrós Collection 1959, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (3) one specimen: 32 km E of Guerrero, San Luis Potosi, Mex., v. 29.1948, dry scrub area, F. Werner & W. Nutting, F. Monrós Collection 1959; (4) one specimen: 47 km S of Ciudad Victoria, Tamaulipas, Mex. (Rt. 1 km 659) 900 ft, vii. 5.1948, semi arid plateau and jungle valley, F. Werner leg., F. Monrós Collection 1959; (5) two specimens: Mexico, Chiapas, 1 - III- 65, Heinrich et al. on Bromeliads, Bro. 88450, 65 - 9816, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2003 [one with: Calligrapha sp. det. R. White]; (6) one specimen: Ventanas, Durango, Höge, Calligrapha ancoralis; (7) two specimens: Mex., Tamaulipas, Bocotoma area, 7 km SSE Gomez Farias, June 1 – 4, 1982, J. E. Wappes [one with: Calligrapha anchoralis Stal det. JE Wappes 2002]; (8) one specimen: Mexico, Mazatlan, Sin., Mex., Kusche 15. ix. 1917, F. Monrós Collection 1959, Calligrapha ancoralis Stål F. Monrós det. 1954; (9) one specimen: 20 mi W Linares, N. León, Mex. xi- 8 - 46, E. C. Van Dyke collector, Van Dyke Collection, Calligrapha ancoralis Stal F. Monrós det. 1953; (10) one specimen: Miraflores, L. Cal., vii. 8.38, Michelbacher & Ross collectors; (11) one specimen: Alamos, Sonora, viii. 2.1940, R. P. Allou Collector; (12) one specimen: Mexico, 14. xii. 62, Randolph with Tillandsia, Laredo 63695, 63 - 33 Calligrapha sp. prob ancoralis Stal, Det. G. B. Vogt; (13) two specimens: Mex., S. L. P., 1.8 mi N El Naranjo 1000 ’, x. 4 & 13.65, George E. Ball, D. R. Whitehead collectors; (14) sixteen specimens: Mexico, Veracruz, 22.0 mi E Jalapa, Rte. 140, 1000 ’ iii. 9.66, in bromeliads, George E. Ball & D. R. Whitehead collectors [one with: Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011]; (15) two specimens: Mexico, Veracruz, 4.0 mi SW Puente Nacional 600 ’ iii. 8.1966, in bromeliads, George E. Ball & D. R. Whitehead collectors; (16) four specimens: Venodio [Venadillo], Sin., Mex., Kusche, B. P. Clark donor [one with: 7. vii. ’ 18; one with: 27. vi. ’ 18; one with: 31. vii. ’ 18; one with: 5 – 6. vii. ’ 18]; (17) one specimen: Mazatlan, Sin. Mex., Kusche, 15 – 17. ix. ’ 18, B. P. Clark donor, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (18) one specimen: Cordoba VC, Mex., Mch. 11 ’ 08, Fredk Knab Collector; (19) one specimen: Mexico, Veracruz, at Brownsville, vi. 3.66, Heinrich, Burgess on Tillandsia ionantha (bromeliad) 66 - 20832, Calligrapha sp. d. RE White; (20) one specimen: Mexico, Sonora, Alamos, 15. vii. 1963, Paul J. Spangler; (21) one specimen: Laredo Tex. ex. S. L. Potosi Mex. iii- 19 - 56, Watt, Tillandsia plants 56 - 3677; (22) one specimen: Tampico, Mex. 7.12, E. A. Schwarz Collector; (23) nine specimens: Mexico, at Miami, 6. xi. 1968, w. Tillandsia plants 68 - 27129 [one with: Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011]; (24) one specimen: Mexico, Veracruz, 16. xi. 65, Brown et al. on Tillandsia, Bro. 88417, 65 - 11596, Calligrapha sp. Dt. R. White; (25) one specimen: Mexico, Veracruz, Ver. at Laredo 9. iv. 1969, D. L. Dillehay on Bromeliad plants 69 - 5807, Calligrapha sp. d. RE White; (26) one specimen: Mexico, Veracruz, 30. i. 67, Gonzalez et al., # Brownsville 90502 Lot #: 67 - 4685, host: Tillandsia ionantha, Calligrapha sp. Det. R. E. White; (27) one specimen: Monterrey, Mex., Lar. Tex. 60648, 3 - 15 - ’ 60, 8555, E. Tillandsia plants, Calligrapha sp. DMW; (28) one specimen: Mexico, San Luis Potosi at Brownsville, v. 16.66, Burgess, Tillandsia ionanth. (Brom.) 66.21549, Calligrapha sp. d. R. E. White; (29) one specimen: Laredo, Tex. ex Huichinrayan, S. L. P., Mexico, iii. 19.56, Watt, Tillandsia plant 56 - 3678, Calligrapha sp. DMW 56; (30) three specimens: Laredo, Tex., ex. S. L. Potosi, Mex., iii. 19.56, Watt, Tillandsia plants 56 - 3677 [one with: Calligrapha sp. DMW 56]; (31) one specimen: Tampico, Mex. 7 / 12, E. A. Schwarz collector, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (32) one specimen: Ventanas, Durango, Höge, 386, gift of F. C. Bowditch, gift ex MCZ dupl. series, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (33) one specimen: Mexico at Miami, 6. xi. 1968, B. K. Dozier, w. Tillandsia plants 68 - 27129, Calligrapha sp. nr. ramulifera Stal d. R. E. White, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (34) one specimen: Vera Cruz, Mex., Lar. Tex. 59422, Jan. 13, ’ 59, 1685, bromeliad plant, Zygogramma sp. DMW, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (35) three specimens: Mexico, 26. iii. 1970, A. S. Millis, Bromeliad Lot 70 - 6302 [one with: Zygogramma sp. det. RE White, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011]. TAMUIC: (1) two specimens: X 0533714 and X 0535353, Mexico, Tamaulipas, 97 km E Ciudad Victoria, Hwy 70, vii. 3.1986, Robert W. Jones, Peter Kovarik, Joseph C. Schaffner leg., Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (2) one specimen: X 0535667, Mexico, Veracruz, La Cumbre, 29. viii. 1962, William F. Chamberlain, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (3) one specimen: X 0536715, Mexico, Sonora, 53 km SW Moctezuma, 23. vii. 1980, S. McCleve & P. Jump; (4) four specimens: X 0538805, X 0540387, X 0543221 and X 0548738, Mexico, Tamaulipas, 5 mi N Llera, 4. vi. 1965, Horace R. Burke, J. R. Meyer & Joseph C. Schaffner; (5) two specimens: X 0538979 and X 0546003, Mexico, Tamaulipas, 5 mi SSE Gomez Farias, 19. vii. 1970, at light, Joseph C. Schaffner, Ellwood R. Hart, Robert R. Murray & Robert G. Phelps; (6) one specimen: X 0539658, Mexico, Tamaulipas, 3 mi W Ciudad Victoria, behind Army Camp, 6. x. 1985, Robert W. Jones, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (7) one specimen: X 0540728, Mexico, Nuevo León, 9 mi S Monterrey, 11. viii. 1972, E. E. Grissell; (8) two specimens: X 0541548 and X 0550793, Mexico, Tamaulipas, 5.7 mi N La Encarnación, 11. vii. 1973, Gaumer & Clark; (9) one specimen: X 0542299, Mexico, Tamaulipas, 21 mi N Ciudad Victoria, 19. vii. 1970, Robert G. Phelps, Robert R. Murray, Hart & Schaffner; (10) one specimen: X 0542904, Mexico, San Luis de Potosí, 8.7 mi S Santa María del Río, 5. vii. 1974, Clark, Murray, Ashe & Schaffner leg.; (11) one specimen: X 0544229, Mexico, Tampico, 27. viii. 1962, William F. Chamberlain, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (12) one specimen: X 0544629, Mexico, Aguascalientes, 6 mi E Calvillo, 11. vii. 1983, P. W. Kovarik, Harrison, Joseph C. Schaffner, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011; (13) one specimen: X 0547099, Mexico, Nuevo León, 23.6 mi SW Linares, 3. vii. 1974, Wayne Clark, Robert R. Murray, James S. Ashe & Joseph C. Schaffner; (14) one specimen: X 0547429, Mexico, Sonora, 19 km NW Nacori Chico, 5. viii. 1982, S. McCleve, G. E. Ball & K. E. Ball; (15) three specimens: X 0548127, X 0549273 and X 0550400, Mexico, San Luís de Potosí, Tamazunchale, 22. vii. 1970, Robert R. Murray, Robert G. Phelps, Ellwood R. Hart & Joseph C. Schaffner; (16) two specimens: X 0548748 and X 0550035, Mexico, Tamaulipas, 2 mi SE Gomez Farias, 20. vii. 1970, Robert R. Murray, Robert G. Phelps, Hart & Joseph C. Schaffner; (17) one specimen: X 0549943, Mexico, Tamaulipas, 7 km SSE Gomez Farias, Bocatoma w. s., 20. vii. 1982, R. Turnbow; (18) one specimen: X 0552047, Mexico, Durango, El Salto, 7. viii. 1967, Horace R. Burke & J. Hafernik; (19) one specimen: X 0552320, Mexico, San Luis de Potosí, 2 mi SE Pedro Montoya, 25. vii. 1970, Robert R. Murray, Robert G. Phelps, Hart & Joseph C. Schaffner. USA FSCA: (1) one specimen: USA, Arizona, Santa Rita Mts., Madera Cyn., 21. vii. 1968; (2) one specimen: USA, AZ, Santa Cruz, Pena Blanca Pajarito, 25. vii. 1962, R. H. Arnett, Jr. & E. Van Kassell, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011. NMNH: (1) one specimen: Nogales, Ariz. ix. 25.1959, D. E. Noel, Calligrapha sp. det ’ 60 D. M. Weisman; (2) one specimen: U. S. A. Texas, Lee Co., Fedor, F. Monrós Collection 1959, Calligrapha wickhami Bowditch, F. Monrós det. 1954, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011. UNKNOWN SOURCE MfN: (1) one specimen: ancoralis St.; (2) two specimens: Calligrapha ancoralis Stål 1860, J. Gómez-Zurita det. 2009.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D643EFFB88F9CF956FDA1FB8A.taxon	description	Variation. Dark parts of body can have a blackish or even greenish or bluish more or less metallic sheen, while pale areas of elytra can be orange. Of the additional markings on disc of elytra, 3 – 4 are always free: in the 3 - spot row behind humeral lunule, the spots at ends are free, while the middle one can be confluent with the apex of the curved prolongation of arcuate band (as seen on the right elytron of the holotype); the two additional markings at the apical declivity of elytra are nearly always free, although the internal spot can be sometimes confluent with other markings (as seen on the right elytron of the holotype), or they can be narrowly connected with each other or missing altogether (as respectively seen in two specimen from Ventanas, Mexico; MfN). In two specimens held at MCZ, the apex of the humeral marking is connected to the base of the arcuate band. The anchor-like marking can be more or less broken at apical arms, the apical spot can be minute or missing, and the spot on apical declivity can be largely confluent with sutural stripe.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6433FF838F9CFBF5FA8CFC1A.taxon	distribution	Distribution. This species is restricted to the Pacific domains of southern Mesoamerica, from the state of Chiapas in Mexico to the provinces of Alajuela and Cartago in Costa Rica (Fig. 10, white circles). C. ramulifera is only present in the Caribbean Mesoamerican domain.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6433FF838F9CFBF5FA8CFC1A.taxon	materials_examined	Material examined (123 specimens). COSTA RICA FSCA: (1) three specimens: Costa Rica, Punt. Prv., 6 km S Sta. Elena, 6 – 7. vi. 1983, J. E. Wappes; (2) one specimen: Costa Rica, Puntarenas, Monteverde, 26. ii. 1987, E. Giesbert, Calligrapha ramulifera Stål J. Gómez- Zurita det. 2011. MCZ: (1) one specimen: Collection Schild-Burgdorf, Costa Rica, Piedras Negras 10, F. A. Eddy Collection, Calligrapha ramulifera Stål J. Gómez-Zurita det. 2010; (2) one specimen: Irazu, 6 – 7000 ft, H. Rogers, 1 st Jacoby Coll. MfN: (1) one specimen: 29785, ramulifera Stål, Costarica, Wagner; (2) one specimen: Collection Schild-Burgdorf, Costa Rica, Piedras Negras; (3) one specimen: Collection Schild-Burgdorf, Costa Rica, Atenas. NHM: (1) one specimen: Irazu, 6 – 7000 ft, H. Rogers, Godman-Salvin Coll., Biol. Centr. - Amer.; (2) one specimen: Pozo Azul, Costa Rica, C. E. Tottenham BM 1969 - 77. NMB: (1) one specimen: Turrialba, Costarica, Polyspila sylvia St., Calligrapha ramulifera Stål J. Gómez-Zurita det. 2010. NMCZ: (1) two specimens: Turrialba, Costa Rica, Coll. Achard Mus. Pragense. NMNH: (1) one specimen: Costa Rica, Ala. Pr., 6 – 8 km W Atenas, June 1 – 2, 1980, J. E. Wappes, J. E. Wappes Collection, Calligrapha ramulifera Stål det. M. Daccordi 1986. ZSM: (1) one specimen: San Mateo, Costa Rica, Calligrapha ramulifera Stål, Staatssammlung München 1975 Erwerb Coll. Machatschke. EL SALVADOR FSCA: (1) one specimen: El Salvador, Landaverde, S. & L. Steinhauser, Calligrapha ramulifera Stal Det. J. Watts 1993; (2) one specimen: El Sal., San Isidro, Co. Verde 1000 m, 2. vi. 1972, S. & L. Steinhauser, 2563, Calligrapha ramulifera Stål J. Gómez-Zurita det. 2011; (3) one specimen: El Sal., Los Chorros 700 m, 4. vi. 1972, S. & L. Steinhauser, Calligrapha ramulifera Stål J. Gómez-Zurita det. 2011. MfN: (1) three specimens: Servicios Técnicos Cafetalería, El Salvador C. A. [one with: 14 - E- 42]. NMNH: (1) one specimen: nº 631 – 39, 17. vii. 1955, Rosario, Cuzcatlan, Col. M. S. V.; (2) one specimen: San Salvador, 25. ii. 20, E. S. C. A., K. A. Salman collector, Calligrapha ramulifera Stål EAC’ 27. GUATEMALA EGRC: (1) one specimen: Guatemala, Guat. City, Las Hamacas Tr. Tk., 5 – 8. viii. 1979, Thomas & Case, E. Riley Colin., C. serpentina var. Daccordi 1981. MCZ: (1) two specimens: Zapote, Guatemala, C. Champion, 1 st Jacoby Coll. MfN: (1) one specimen: Guatemala, Calligrapha ramulifera Stål det. Daccordi ' 79; (2) two specimens: Guatemala. NHM: (1) one specimen: Guatemala C. A., 14, Biol. Cent. - Am. 1905 - 17, Calligrapha ramulifera Stål, Godman-Salvin Coll., Biol. Centr. - Amer.; (2) three specimens: Zapote, Guatemala, G. C. Champion, Godman-Salvin Coll., Biol. Centr. - Amer.; (3) one specimen: Cerro Zunil, 4000 ft, Champion, Godman- Salvin Coll., Biol. Centr. - Amer.; (4) one specimen: San Isidro, 1600 ft., Champion, Godman-Salvin Coll., Biol. Centr. - Amer.; (5) one specimen: Guatemala, named by Stål, Sp. figured, Baly Coll., ramulifera Stål, Guatemala, Mexico [underneath: named by Stål, Col: Deyrolle]; (6) one specimen: Guatemª 55.71. NMB: (1) one specimen: Guatemala, G. Bermoully, Calligrapha ramulifera det. Daccordi ‘ 78. NMNH: (1) six specimens: Mauricio, Guatemala, July 88, Dr. G. W. Bock collector, Collection F. Knab [one with: Calligrapha ramulifera Stål F. Monrós det. 1953; one with Calligrapha ramulifera Stål]; (2) five specimens: Guatemala, San Jose, May ’ 43, D. G. Hall; (3) two specimens: Guatemala, S. Sebastian, Retalhueleu, 1925, L. Thiel, F. Monrós Collection 1959; (4) two specimens: Guat., ’ 25, L. Thiel, S Sebastian, Retalhuleu; (5) one specimen: Guatemala, C. A. Champion, 14, Wickham Collection 1933, Calligrapha ramulifera; (6) one specimen: Los Patos Riv., 14 mi up, vi. 18 – 23.1926, Suchetepequez Province, Guatemala, Pres. by J. R. Slevin Collector, Calligrapha ramulifera Stål J. Gómez-Zurita det. 2011. TAMUIC: (1) one specimen: X 0543052, Guatemala, Escuintla, 25. v. 1923, E. G. Smyth, Calligrapha ramulifera Stål J. Gómez-Zurita det. 2011. ZSM: (1) ten specimens: Guatemala, Suchitepequez, Patului, A. Poll [one with: 1.7.83, Calligrapha ramulifera Stål J. Gómez-Zurita det. 2011; five with: 1.7.83; two with: 10.7.83; two with: 10.8.83]. HONDURAS EGRC: (1) one specimen: Honduras, Copan, 16 km NW Sta. Rosa de Copan, 8. x. 1993, R. Turnbow, Calligrapha aeneopicta St. det. Daccordi ’ 86, Calligrapha ramulifera Stål J. Gómez-Zurita det. 2011; (2) two specimens: Honduras, Com., 5 km NW Taulabé, Río Jaitique, 29. vii. 1977, O’Brien & Marshall. FSCA: (1) one specimen: Honduras, Copan, 16 km NW Sta. Rosa de Copan, 8. x. 1993, R. Turnbow, Calligrapha aeneopicta Stal det. E. G. Riley ’ 93, Calligrapha ramulifera Stål J. Gómez-Zurita det. 2011; (2) two specimens, Honduras, Copan, 16 km NW Santa Rosa de Copan, 8. x. 1993, F. W. Skillman, Calligrapha ramulifera Stål J. Gómez-Zurita det. 2011. NMNH: (1) one specimen: Dépt. Copon, May 1919, Honduras, S. F. Blanc Coll., Calligrapha ramulifera Stål J. Gómez-Zurita det. 2011. MEXICO MCZ: (1) two specimens: Tapachula, Chiapas, Höge, Jacoby 2 nd Coll. MfN: (1) one specimen: Tapachula 16. NHM: (1) two specimens: Tapachula, Chiapas, Höge, Godman-Salvin Coll., Biol. Centr. - Amer.; (2) one specimen: Mex., Baly Coll.; (3) one specimen: Chiapas, Mexico, Escuintla, VI- 1925, 69, F. R. Mason Coll., Pres. by Imp. Bur. Ent. Brit. Mus. 1926 - 83; (4) one specimen: E. Coll. Thomson, Mexiq.?. NMCZ: (1) one specimen: Guanajuato, E. Dugés, Coll. Achard Mus. Pragense 3. NMNH: (1) one specimen: banana debris, Mex., 8 - 8 - ’ 08, Galv. 5077. NICARAGUA IBE-JGZ: (1) one specimen: IBE-JGZ-C 107, Nicaragua, Rivas, Sierra Serena UTM 16 P- 657865 - 1242291, 21 September 2009, J. - M. Maes leg.; (2) five specimens: [IBE-JGZ-C 231, IBE-JGZ-C 249, IBE-JGZ-C 250, IBE-JGZ- C 251, IBE-JGZ-C 252], Nicaragua, Rivas, Finca Guadalupe, 10 – 12 / 07 / 2010, J. - M. Maes leg.; (3) four specimens: [IBE-JGZ-C 234, IBE-JGZ-C 253, IBE-JGZ-C 261, IBE-JGZ-C 262], Nicaragua, Estelí, P. T. P. Miraflor- Moropotente, Finca Las Flores 13.192570 N 86.331465 W 865 m, Abril 2010, J. - M. Maes leg.; (4) two specimens: [IBE-JGZ-C 255, IBE-JGZ-C 256], Nicaragua, Rivas, Cárdenas, Finca Sierra Serena, 14 / 07 / 2010, J. - M. Maes leg.; (5) one specimen: IBE-JGZ-C 265, Nicaragua, Estelí, P. T. P. Miraflor-Moropotente, Finca Bartolo UTM 16 P- 0573112 - 1459212 993 m, Junio 2010, A. del Socorro & B. Nimia leg.; (6) one specimen: IBE-JGZ- 2166, Nicaragua, Estelí, Reserva Natural Miraflor, Isidrillo [51], Agosto 2010, J. - M. Maes leg.; (7) one specimen: IBE- JGZ- 2172, Nicaragua, Estelí, Reserva Natural Miraflor, Miraflor [209], Septiembre 2010, J. - M. Maes leg. MCZ: (1) one specimen: Nicaragua, Calligrapha ramulifera Stål J. Gómez-Zurita det. 2010; (2) one specimen: Nicaragua; (3) one specimen: Chontales, Nicaragua, T. Belt, 1 st Jacoby Coll., Calligrapha ramulifera Stål J. Gómez-Zurita det. 2010. NMB: (1) one specimen: Nicaragua, Managua Low, Calligrapha ramulifera Stål J. Bechyné det. 1951. NMNH: (1) two specimens: Nicaragua, A. D. Harvey coll. 1923 [one with: Calligrapha ramulifera Stål F. Monrós det. 1953]; (2) four specimens: Managua Cy, Nicaragua, A. D. Harvey Coll. 1923; (3) one specimen: Ameya, Nicaragua, A. D. Harvey coll. 1923; (4) one specimen: Nicaragua, 11. ii. 1938, F. Monrós Collection 1959, Calligrapha ramulifera Stål F. Monrós det. 1954; (5) one specimen: Managua, Nicaragua [illegible] 1925; (6) one specimen: Managua, Nic., v. 28.1956, Mabry, 56 - 9599, Calligrapha sp. DMW 56; (7) one 3. The State of Guanajuato in Central Mexico is very far from the general area of distribution of C. ramulifera, therefore this record most likely represents either a labeling mistake or a reference to a locality in Chiapas, in the Pacific slopes of the Sierra Madre de Chiapas. specimen: Nica., Chinandega, Volcan Casita, 1. vii. 95, Maes / Hernández, duplicated retained from the Museo Entomologico S. E. A. Collection, Leon, Nicaragua, Calligrapha ramulifera Stål J. Gómez-Zurita det. 2011. UNKNOWN SOURCE MfN: (4) one specimen: picta Chev. Mex. NHM: (1) one specimen: Baly Coll.; (2) one specimen: E. Coll. Laferté, 67 - 56. NRM: (1) one specimen: Guatimala, [illegible]. OUMNH: (1) two specimens: [no data] [one with: Calligrapha ramulifera Stål J. Gómez-Zurita det. 2011].	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6433FF838F9CFBF5FA8CFC1A.taxon	description	Variation. This species is rather constant in appearance and the observed variability in appearance does not generate identification problems. The sutural stripe can be suddenly narrowed at apex (as seen in the paralectotypes). The basal ends of the subsutural stripe can converge sometimes with the sutural stripe, and in one specimen from Pozo Azul (Costa Rica), the ends of the arcuate band are also completely fused with the subsutural stripe, appearing as lateral lumps. Humeral markings are generally broadly fused, but in some specimens the apical ends of the humeral spot and lunule are free and slightly divergent (as seen in PLT 1, among others). The spot enclosed by the humeral lunule can be very small or maybe absent altogether (as seen on one elytron of a single specimen). The midlateral spot can be subrectangular or ellipsoidal and in the latter case, the connection to dark margin of elytron can be narrow and through dark suffusion. The spot of apical declivity can be very small and free. Spots on disc can appear indicated only by the pale or golden surrounding area, and instead of ten spots as in the lectotype, there can be arrangements of eight (2 - 2 - 2 - 2) or nine (3 - 2 - 2 - 2) spots. In some specimens, the sculpture of the epipleura differs from the type showing a leathery and shiny appearance. Two specimens from Turrialba (MNCP) and two from ca. Santa Elena, Costa Rica (FSCA) are more slender, longer specimens than the type.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6408FF818F9CFC45FB3AF8D7.taxon	materials_examined	Holotype: 1351 / San Antonio, Chiapas, v. Jurckhein /? argus (MfN). Paratypes: (1) one specimen: Cerro Zunil, 4000 ft., Champion / 96425 (MfN); (2) two specimens: Guatemala [one with: geographica Stål] (MfN); (3) one specimen: Distrikt Socomusco, Hotzen S. V. (MfN); (4) one specimen: Sinanja, Vera Paz, Champion / Godman-Salvin Coll., Biol. Centr. - Amer. (NHM); (5) one specimen: Purula, Vera Paz, Champion / Godman-Salvin Coll., Biol. Centr. - Amer. (NHM). Habitus (Fig. 6 c). Length: 8.84 mm; width: 5.61 mm. Body elongated oval, moderately convex. Head, disc of labrum, mandibles basally, first antennomere dorsally, pronotum, scutellum, all ventral surfaces, coxae, trochanters apically, femora, tibiae, and first tarsal segment shiny black with slight bronzy metallic reflection; narrow apex of clypeus, margins of labrum, apex of mandibles, mouth parts, apical antennomeres, tarsi, dark elytral markings, and epipleura very dark testaceous; basal segments of antennae ventrally, base of trochanters, knees, and tarsal articulations bright reddish brown; background of elytra dark yellow with dark markings margined golden creamy yellow. Head broad, deeply inserted in pronotum; surface very finely microreticulate, relatively densely and homogenously covered with moderately strong punctures; supraocular sulci deep, slightly bisinuous, from antennal calli well beyond upper eye margin; few sparse setae near eyes, particularly on antennal calli; frontal suture very fine, obsolete behind, apically joining strongly bisinuated clypeal suture; clypeus long, slightly depressed at sides, conspicuously sloping ventrally at apex, sculptured as frons, with few sparse fine setae laterally and longer, larger and medially convergent setae at apical angles. Antennae slender, surpassing humeri; first antennomere long, thick, nearly cylindrical at apical 2 / 3; antennomeres 2 – 6 elongated, relatively thin, smooth, with very sparse setae; second antennomere half as long as first, third subequal to first; antennomeres 3 – 6 shortening progressively, sixth slightly longer than second; segments lengthening gradually beyond sixth antennomere; antennomeres 7 – 11 thicker, widening from base to apex, 8 – 11 rugose and pubescent, eleventh about as long as first, tappering towards blunt apex and slightly emarginate dorsally; eighth antennomere 0.68 x as wide as long. Labrum with regularly curved sides and emarginate at apex; with very long lateral and apical straight or slightly bent setae, convergent medially. Mandibles large, very strong, well surpassing labrum by 2 / 3 their length; sides slightly concave basally before broad apical curvature; laterally and frontolaterally very strongly punctured, with long recumbent translucent setae. Maxillary palpi long; apical palpomere subtrapezoidal, broader than long, with slightly curved longer external margin and slightly compressed dorsoventrally and weakly concave at apex; previous palpomere slightly broader apically than base of last segment, long, regularly curved externally to narrow base; first palpomere long, feebly curved, gradually thickening from base. Pronotum transverse (W / L = 1.87), broader anteriorly before middle; sides straight basally, slightly divergent up to midlength, and gently curving towards strongly protruding apical angles; apical border slightly bisinuated, slightly convex medially; basal border convex; apically finely margined, laterally and at angles with broader, slightly explanate margin; surface very finely microreticulate, with irregularly scattered small and medium-sized punctures, becoming large, deep and sometimes convergent at sides; primary punctures intermingled with very fine secondary punctures; round deep convergent punctures at basal margin except on median lobe; setae at apical angles long, fine, bent dorsolaterally backwards; setae at basal angles long, fine, bent dorsolaterally forwards. Hypomeral suture impressed, slightly wavy, regularly slightly diverging from pronotal border, more weakly impressed apically; hypomera very finely microreticulate, shiny, unpunctured, convex medially, transversally depressed basally. Prosternum relatively narrow, feebly concave, finely margined anteriorly; surface finely microreticulate except at glossy process; prosternal process narrow, convex, feebly longitudinally raised medially, setose, apically depressed, gradually sloping towards straight apex; sides of process strongly rugosely punctured, with scattered long setae; apex of process punctured, setose, feebly raised at sides. Mesanepisterna finely microreticulate. Metepisterna moderately and sparsely punctured; apically densely punctured with elongated convergent punctures. Metaventrite finely leathery at sides, smooth and glossy medially, strongly closely punctured near mesocoxae, with some strong punctures near lateral border; finely punctured, with very fine hairs laterally on disc; deeply transversally furrowed at posterior border. Scutellum small, longer than wide at base (W / L = 0.90) with regularly curved sides towards round blunt apex; finely microsculptured, unpunctured. Elytra long, with regularly broadly curved shoulders, weakly curved laterally towards regular elongated apical curvature; surface very finely microsculptured, smooth, shiny, with dense, very fine dot-like punctures in pale areas, almost imperceptible apically on elytra, and stronger, deeper punctures around and within dark markings; very tight, regularly impressed premarginal line of punctures, only becoming slightly confused at subhumeral area; scutellar row present, double basally. Markings: (i) sutural stripe complete, continuous, broadly surrounding scutellum basally, gradually narrowing posteriorly, barely reaching apex of elytra; (ii) subsutural stripe broad, completely confluent laterally with sutural stripe, starting basally shortly before apex of scutellum, gradually narrowing posteriorly, slightly widening right before apex and irregularly disappearing abruptly slightly before apex; (iii) arcuate band long, set medially on disc, completely confluent laterally with subsutural stripe, weakly concave externally, with slightly divergent but entirely laterally confluent basal end; apical end strongly widened as slightly elongated rounded, weakly divergent shape; (iv) humeral spot short, elongated oval on shoulder, completely confluent laterally with (v) humeral lunule, gently curved apically towards suture, ending near and only slightly beyond basal end of arcuate band; basally very narrowly connected to dark basal margin of elytron at humeral internal declivity; (vi) spot enclosed by humeral lunule large, round, narrowly separated from internal concavity of lunule; (vii) midlateral spot large, subrectangular, completely confluent externally with narrow dark margin of elytron; (viii) spot of apical declivity large, semicircular, completely fused laterally with subsutural stripe; (ix) apical spot medium-sized, round, slightly transversally elongated, free; (x) nine additional roundish, small to relatively large spots on disc; four equidistantly separated and aligned longitudinally parallel to elytral margin on lateral declivity, the basalmost at level with basal end of arcuate band; four more as an internal series, parallel to former series with each spot set slightly forwards compared to nearest spot in external row, except basalmost spot; one round spot on external basal concavity of arcuate band, at level with midlateral spot. Legs long, slender. Femora fusiform, with surface delicately microsculptured, shiny, with relatively dense fine punctures bearing short fine setae. Tibiae slender, faintly curved ventrally; feebly carinated longitudinally at ventral margin, neatly furrowed externally at apical half, furrow widening towards emargination for tarsal insertion; sparsely punctured with longitudinally elongated punctures, becoming rugose at apex, with golden reddish dense pubescence. Abdominal ventrites shiny, smooth, with very fine sparse punctures bearing a fine translucent hair; punctures denser, stronger near metacoxae. Penis figured in Figs 9 c, 9 f.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6408FF818F9CFC45FB3AF8D7.taxon	distribution	Distribution. Species endemic of the Sierra Madre de Chiapas (Mexico) and chain extensions in Guatemala (Fig. 8, black squares). C. catarinae sp. nov. is endemic from the Caribbean Mesoamerican domain. Material examined (13 specimens). GUATEMALA NMNH: (1) two specimens: Guatemala, Suchitepequez, Zapotitlan, 1525 meters, Finca Las Nubes, 16 June ’ 73, Ginter Ekis, Calligrapha catarinae i. l. J. Gómez-Zurita det. 2011; (2) one specimen: Cerro Zunil, 4000 ft, Champion, Calligrapha catarinae i. l. J. Gómez-Zurita det. 2011. OUMNH: (1) one specimen: Cerro Zunil, 4000 ft., Champion, B. C. A. duplicates pres. 1909 by F. D. Godman Cat. No. 84, Calligrapha catarinae i. l. J. Gómez- Zurita det. 2011; (2) one specimen: Cerro Zunil, 4 – 5000 ft., Champion, B. C. A. duplicates pres. 1909 by F. D. Godman Cat. No. 84, Calligrapha catarinae i. l. J. Gómez-Zurita det. 2011. ZSM: (1) Cerro Zunil, 4 – 5000 ft, Champion, Polyspila argus Stal, Staatssammlung München 1975 Erwerb Coll. Machatschke, Calligrapha catarinae i. l. J. Gómez-Zurita det. 2011. MEXICO NMNH: (1) one specimen: Chiapas, Mex., L. Holzen, ’ 19, Pacific Slope Cordilleras 800 – 1000 m, Calligrapha catarinae i. l. J. Gómez-Zurita det. 2011. ZSM: (1) one specimen: Distrikt Socomusco, Hotzen S. W., Staatssammlung München 1975 Erwerb Coll. Machatschke.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6408FF818F9CFC45FB3AF8D7.taxon	description	Variation. In one female specimen from Guatemala, the basal connection of humeral lunule with base of elytra is broader, more conspicuous. The spot enclosed by the humeral lunule can be narrowly connected to inner concavity of lunule by areas of dark suffusion. In most specimens there are ten additional markings on elytra, and not nine like in the holotype, with the internal row of spots including five and not four spots, the three basalmost small and round, with a trend of third and larger fourth to fuse together (a trend apparently completed in the holotype). The single additional spot on concavity of arcuate band can be very narrowly connected to the latter by dark suffusions.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6408FF818F9CFC45FB3AF8D7.taxon	diagnosis	Diagnosis. The new species is very similar and possibly most closely related to the Panamanian C. fulvitarsis, sharing a similar shape, colouration and disposition of spots, including a basal blunt end of the arcuate band and a roundish spot enclosed by the humeral lunule, generally with small basal emargination in the other species in this group. However, these species can be easily recognized by the larger size of C. catarinae sp. nov., larger and neater spots on elytra, larger midlateral spot, confluent with margin of elytron, and legs completely black. This species is also very similar to C. argus Stål, sharing the lack of subhumeral spot and the presence of a large midlateral spot, confluent with margin, as well as a characteristic bitonality to pale areas of elytra, with a paler rim around dark markings. However, C. catarinae sp. nov. is consistently darker, blackish, and shows other differences, including a more compact humeral marking, a blunt basal end of the subsutural stripe, a sudden preapical disappearance of the subsutural stripe, an elongated spot enclosed by humeral lunule, the semicircular shapes of the spot of apical declivity and generally of the apex of the arcuate band, and it is also generally larger and more elongated. The size of the specimens and the general disposition of markings is also reminiscent of C. geographica Stål, among which it can be found sometimes in collections, but a closer examination reveals many differences, most notably the type of midlateral spot, narrow and free in C. geographica Stål, large and confluent with margin in C. catarinae sp. nov. The pattern of markings may also induce the mistake to place the specimens with C. diversa Stål, but the absence of midlateral spot in the latter makes them easily distinguishable. Finally, there could be some confusion with C. suboculata Stål, particularly since this species can lack one of the two spots enclosed by the humeral lunule, but the shape of most markings on elytra separate them.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6408FF818F9CFC45FB3AF8D7.taxon	etymology	Derivatio nominis. These large and striking Calligrapha are amongst the most beautiful representatives of the genus, which compelled me to name this species with love after my beautiful daughter, Catarina.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D640BFF858F9CFF06FB2FFD1F.taxon	materials_examined	Holotype: Mexico / HOLOTYPE Calligrapha anabelae sp. nov. J. Gómez-Zurita [red] (MfN). Paratypes: MfN: (1) one specimen: Mexico / PARATYPE Calligrapha anabelae sp. nov. J. Gómez-Zurita; (2) one specimen: PARATYPE Calligrapha anabelae sp. nov. J. Gómez-Zurita. NHM: (3) one specimen: 23842 / Truqui, Mexico / Fry Coll. 1905.100 / PARATYPE Calligrapha anabelae sp. nov. J. Gómez-Zurita; (4) one specimen: Baly Coll. / PARATYPE Calligrapha anabelae sp. nov. J. Gómez-Zurita; (5) one specimen: Mexique / Baly Coll. / PARATYPE Calligrapha anabelae sp. nov. J. Gómez-Zurita. Habitus (Fig. 6 e). Length: 9.78 mm, width: 6.22 mm. Body elongated oval, moderately convex. Head, mandibles, dorsum of first antennomere and pronotum very dark reddish brown, with dull greenish metallic reflection; mouth pieces, antennae, labrum, scutellum, dark markings of elytra, epipleurae, ventral surfaces and legs dark reddish brown, with slight greenish metallic reflection on scutellum, elytral markings and legs; background of elytra custard yellow. Head large, wide, deeply inserted in pronotum; surface finely microreticulate, densely, rather uniformly punctured; punctures somewhat larger and denser near and behind eyes; almost unpunctured near frontal suture and antennal calli; punctures bearing short translucent seta and one much longer, semierect seta near upper eye margin; deep, slightly curved furrow above eye; frontal suture very fine in all length, joining at apex broadly V-shaped clypeal suture. Clypeus finely microreticulate, with smaller, shallower punctures than head; anterior border laterally with 4 – 6 long pale yellowish setae, convergent medially, almost as long as labrum. Eyes dorsoventrally elongated. Genae very short. Labrum transverse, relatively short (ratio L: W = 0.4), emarginate anteriorly, with 6 – 7 long yellowish translucent setae at sides, thicker than clypeal setae, convergent medially. Antennae short, conspicuously thickened beyond sixth antennomere; antennomeres 3 – 6 slender, gradually shortening and thickening; antennomeres fifth and sixth subequal to second; five last antennomeres darkened, rugose, densely pubescent; eighth antennomere 0.87 x as wide as long. Mandibles large, robust, protruding by 1.5 x length of labrum; external border markedly concave; surface strongly, densely punctured, with large punctures bearing long yellowish translucent seta; very dense and very fine secondary punctation. Last maxillary palpomeres gradually widening towards apex, with external margin weakly convex, internal margin weakly concave and shorter, and apical margin oblique, straight; apex of previous palpomere slightly broader than base of last segment, curved and gradually narrowing towards base, more accentuated at external edge; first palpomere club-shaped, obliquely cut at apex. Pronotum transverse (W / L = 1.81), sides narrowly margined, feebly and regularly convex from base to strongly produced acute and rounded anterior angles; posterior angle obtuse; anterior border concave, with broader margin at sides, straight or very slightly convex medially, very finely edge; posterior border regularly convex, slightly sinuous at sides, unmargined; surface alutaceous, very finely microreticulate, with dense, small, shallow punctures on disc, gradually becoming larger and deeper, sometimes confluent towards sides, with premarginal area and anterior angles nearly devoid of large punctures, and with longitudinal row of small punctures very close to margin; sides of disc with two round depressions in front of middle; row of elongated punctures along basal margin at sides. Hypomeral suture very deep, slightly wavy, continuous from near pronotal basal angle to base of anterior angle, gradually but weakly diverging from pronotal border; hypomera alutaceous, unpunctured, weakly concave basally with few transverse wrinkles. Prosternum flat at sides and process, convex, only weakly raised at middle; anterior border feebly concave, finely margined; surface finely microreticulate, nearly unpunctured anteriorly, with large, deep punctures near procoxae and at process, bearing fine translucent setae; process narrow between coxae, as narrow as width of eighth antennomere, gradually widening to apex to nearly 1.9 x width between coxae; process unmargined laterally, weakly convex at apex, rugosely punctured and feebly carinated longitudinally at sides and medially. Mesanepisterna very finely microreticulate, not as alutaceous as hypomera; with sparse small, shallow punctures. Metepisterna more finely alutaceous than mesanepisterna, with shallow elongated punctures, deeper and larger along sides, rugosely punctate only at apex. Metaventrite finely leathery, sparsely, regularly impressed with fine small punctures, hairy medially with very fine translucent setae. Scutellum long (W / L = 0.82), acute, with sides feebly curved towards blunt apex; surface finely microreticulate, unpunctured. Elytra long, convexly raised above pronotum; humeri round, sides weakly convex and regularly curved at apical third towards broadly rounded apex; margined from humeral angle to apical sutural angle and at suture on apical declivity; surface very finely alutaceous, sparsely and irregularly punctured with shallow, dot-like punctures, deeper at sides and around dark markings at disc; scutellar row of punctures present, long, starting close to base of elytra, with some 15 – 17 tight, sometimes connected punctures, confused, double basally; premarginal row of very tight punctures present, regularly arranged except slightly confused ends; margin punctured all along. Epipleura very finely alutaceous, unpunctured, feebly sloping ventrally internally, slightly visible from side. Markings: (i) sutural stripe narrow, continuous from base of elytra surrounding scutellum to sutural angle, gradually narrowing to disappear; (ii) subsutural stripe narrow, starting slightly before apex of scutellum, gradually narrowing towards apex of elytra, disappearing before reaching it, slightly widening at apical declivity; entirely confluent with sutural stripe except basally for a relatively long distance (9 – 10 punctures); (iii) arcuate band with narrow blunt basal end, weakly convex externally and enlarged apically as roundish large spot; laterally confluent for most of its length, except for short distance at both ends, with subsutural stripe by darkened area between both features; (iv) humeral spot and lunule entirely confluent laterally forming large humeral marking; humeral spot reaching basally very close to narrow dark basal margin of elytra, nearly connected to it by dark suffusions; apical third of humeral lunule bent at obtuse angle towards suture, ending behind and very close to basal end of arcuate band; (v) spot enclosed by humeral lunule large, nearly U-shaped, with inner space also darkened, appearing as slightly basally emarginate large roundish spot; largely confluent externally and apically with humeral lunule at internal angle; (vi) midlateral spot large, subtrapezoidal, slightly inclined backwards, externally confluent with elytral margin, and internally convering about seven punctures of premarginal line of punctures; (vii) spot of apical declivity longitudinally elongated, confluent by most of its length, except briefly anteriorly with enlargement of subsutural stripe; (viii) apical spot smaller than spot of apical declivity, roundish, free; (ix) nine additional spots on disc; eight roundish, subequal, one larger and of irregular shape (possibly two spots combined); four spots parallel to margin at more or less regular intervals, second at level with midlateral spot, third (largest spot) at level with apical enlargement of arcuate band; one spot on external concavity of arcuate band; longitudinal and slightly curved towards suture series of three spots more or less surrounding previous spot, second in this series at level with midlateral spot; last spot medially in space delimited by apex of arcuate band, base of spot of apical declivity and last spot of external marginal series. Femora spindle-shaped, with surface almost glossy, sparsely punctured, with punctures deeper at apex, each puncture with very fine translucent seta; tibiae slender, gradually widening towards apex, weakly longitudinally carinated ventrally, furrowed on apical half dorsally, with furrow progressively widening towards tarsal insertion; surface almost glossy, with sparse punctures, denser at angles and apically, pubescent ventrally and densely at apex with thick golden setae. Tarsi elongated, slender, entirely and densely pubescent underneath, except at basal narrow half of second segment; onychium with small apical tooth underneath; claws weakly divaricate. Abdominal ventrites alutaceous, very weakly and sparsely punctured, finely pubescent medially and at sides of three apical segments. Penis figured in Figs 9 b, 9 e.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D640BFF858F9CFF06FB2FFD1F.taxon	distribution	Distribution. This species is endemic from central Mexico, in the general area around the Transvolcanic Belt, from the states of Jalisco to Guerrero and Morelos (Fig. 8, white squares). C. anabelae sp. nov. is thus present in the Mexican Transition and Caribbean Mesoamerican domains.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D640BFF858F9CFF06FB2FFD1F.taxon	materials_examined	Material examined (23 specimens). MEXICO EGRC: (1) one specimen: Mexico, Queretaro, Queretaro, 10. viii. 1980, Z. A. Wibmer, C. multipustulata Stal det. Daccordi ’ 86, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011; (2) one specimen: Mexico, Jalisco, 5.5 mi NE Tecolotlan, 13. vii. 1982, A. J. Gilbert, Calligrapha multipustulata St. det. Daccordi ’ 86, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011. FSCA: (1) one specimen: Mexico, Jalisco, Est. Biol. Chamela, 10 – 20. vii. 1985, E. Giesbert, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011; (2) one specimen: Mexico, 8 km N Netapa, 12. vii. 1971, S. R. & L. M. Stenhauser, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011. MCZ: (1) one specimen: Morelos, Tepoztlan, 14 June 1993, leg. D. Furth, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2010; (2) one specimen: Tuxpan, Jalisco, Mex., 9.6.1903, J. F. McClendon, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2010. NMB: (1) one specimen: Mexique, Coll. Chapuis, det. Calligrapha suboculata Stal, Calligrapha argus Stal J. Bechyné det. 1954. NMNH: (1) one specimen: Mexcala, Gro. Mex. vi. 29.51, F. Monrós Collection 1959, Calligrapha anabelae i. l. J. Gómez-Zurita det.; (2) one specimen: Colima Vulcano Mex., L. Conrad, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011; (3) four specimens: 19 mi SW Colima, Colima, Mexico, 26. vii. 63, Ackerman & Whitehead, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011. TAMUIC: (1) two specimens: [X 0532852 and X 0585222], Mexico, Jalisco, 8 mi S Autlan, 8. vii. 1984, Lynn Carroll, Joseph C. Schaffner & Timothy P. Friedlander, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011; (2) four specimens: [X 0533713, X 0534763, X 0535248 and X 0535313], Mexico, Jalisco, 11 mi N Autlan, 6. vii. 1984, Carroll, Schaffner & Friedlander, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011; (3) five specimens: [X 0533782, X 0534181, X 0534286, X 0535329 and X 0544282], Mexico, Jalisco, 16 km N Autlan, 7. vii. 1984, Lynn Carroll, Joseph C. Schaffner & Timothy P. Friedlander, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011. ZSM: (1) three specimens: Vulkan Colima, coll. Joh. Laue, 25.7.1918, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011; (2) two specimens: Vulkan Colima, coll. Joh. Laue, 20.6.1918, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011; (3) three specimens: Vulkan Colima, coll. Joh. Laue 1918 [two with: Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011]; (4) four specimens: Vulcan Colima, Mexico, 1918, coll. Joh. Laue [three with: Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011]; (5) one specimen: Mexico, Guadalajara, Juni 1982, leg. Fittkau, Calligrapha anabelae i. l. J. Gómez-Zurita det. 2011.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D640BFF858F9CFF06FB2FFD1F.taxon	description	Variation. Sides of the apical segment of maxillary palpi are subparallel in females. Sides of pronotum can be almost straight, convergent before apical fourth, regularly curving towards produced anterior angles. Punctation of pronotum can be denser than in type and abdominal segments can be uniformly pubescent. The humeral marking on elytra can be clearly and relatively broadly connected to base of elytra; the spot enclosed by the humeral lunule can be narrowly connected to apical oblique area of lunule; two median discal spots at level with midlateral spot can be fused; and the midlateral spot can be narrowly confluent with elytral margin. But the most remarkable variation is found on the trend of this species to lose most of its elytral markings, including basal ends of subsutural stripe and arcuate band, humeral marking and spot enclosed by lunule, most of additional spots on disc, except for spot at concavity of arcuate band and apical spot (Fig. 11). But, even if the spots are missing, their position is still indicated by puncture arrangements, particularly at basal half of elytra, or the corresponding areas are slightly darkened. One specimen from Tuxpan (MCZ) has all elytral markings present, but precisely the midlateral spot is very reduced, appearing as a spot just below premarginal row of punctures; this specimen is otherwise identical to type. This trait is very unique among Calligrapha, whereby specimens with incomplete elytral patterns look like the intermediate stages in the darkening process that recently emerged, teneral adults experiment to acquire their adult appearance. However, all these specimens had hardened teguments, not different from these seen in the fully pigmented specimens.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D640BFF858F9CFF06FB2FFD1F.taxon	diagnosis	Diagnosis. At first sight C. anabelae sp. nov. looks very similar to Northeastern North American Calligrapha species in the C. scalaris group, such as C. tiliae Brown and C. virginea Brown, but they can be recognized at once by the conspicuous hypomeral suture, absent in all North American species, and by the dark epipleura in the Mexican species. The most similar species, and possibly the closest relative is C. argus, but they can be readily distinguished by the different colouration to darker areas, reddish in the latter and blackish in the new species, as well as by a generally more compact humeral marking, among other minor details. From the allopatric relative C. catarinae sp. nov., they can be recognized easily from the basal ends of subsutural stripe completely fused with the sutural stripe in this species while these ends are divergent in C. anabelae sp. nov., but also by stronger elytral punctation in the latter. Finally, the new species is recognizable from other blackish species with similar marking disposition, such as C. simillima or C. diversa, by the presence of a conspicuous midlateral spot, absent in these species.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D640BFF858F9CFF06FB2FFD1F.taxon	etymology	Derivatio nominis. This species belongs into the same group of neat forms such as C. argus Stål and C. catarinae sp. nov. This new species is dedicated with love to my partner for fifteen years, Anabela R. P. Cardoso, entomologist enthusiastic of tiger beetles, who has always supported and encouraged my work.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D640EFF8E8F9CFD46FDBEFD8D.taxon	distribution	Distribution. This is one of the species in the group with a wider distribution, ranging from the southern half of Mexico (particularly in the Caribbean domain) to Venezuela, through Guatemala, Belize, Honduras, El Salvador, Nicaragua, Costa Rica, Panama and Colombia (Fig. 10). It is one of the three species in this evolutionary lineage that possibly reached the Caribbean NW South American region from its main distribution in the Caribbean Mesoamerican domain in recent times (Montelongo & Gómez-Zurita, 2014).	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D640EFF8E8F9CFD46FDBEFD8D.taxon	materials_examined	Material examined (350 specimens). BELIZE NMB: (1) one specimen: Br. Honduras, Punta Gorda, 1915, Col. R. Vitalis, Calligrapha argus Stål J. Bechyné det. 1954. NMNH: (1) one specimen: Belize, Toledo dist., Blue Creek Village, 18 June 1981, W. E. Steiner, Earthwatch Belize Expedition 1981, D. H. Messersmith, W. E. Steiner et al., Calligrapha argus Stål J. Gómez-Zurita det. 2011. COLOMBIA MCZ: (1) one specimen: Sevilla, Mgd. Colombia, v. 6.28, Darlington, Calligrapha argus Stål J. Gómez-Zurita det. 2010. NHM: (1) one specimen: Calligrapha pacta Dej. Columbia, [illegible], Chrysomela argus Stål, Baly Coll.; (2) one specimen: E. Coll. Laferté, 426, Calligrapha pacta Dej. Colombia, argus Stål Stål, 67 - 56; (3) one specimen: E. Coll. Laferté, 479, Granada, 67 - 56. NRM: (1) one specimen: Columbia, Stål. COSTA RICA EGRC: (1) one specimen: Costa Rica, Monte Verde, Cordillera de Tilarán, 10. iii. 1991, M. E. Rice, Calligrapha argus Stål J. Gómez-Zurita det. 2011. FSCA: (1) one specimen: Costa Rica, Guanacaste, Finca Taboga Res. Farm, 9. vii. 1966, G. R. Buckingham; (2) two specimens: Costa Rica, Ala. Pr., 8 km S S. Ramon, 31. v. 1980, J. E. Wappes, Calligrapha argus St. Det. E. G. Riley ’ 80; (3) one specimen: Costa Rica, Guanacaste, La Pacífica, nr. Cañas, 8. vi. 1983, J. E. Wappes; (4) two specimens: Costa Rica, Pun., 13 mi NW Esparza, 250 ’, 19. vi. 1974, C. W. & L. B. O’Brien & G. B. Marshall, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (5) one specimen: Costa Rica, Puntarenas, Monteverde, 23 – 27. v. 1987, E. Giesbert, Calligrapha argus Stål J. Gómez- Zurita det. 2011; (6) four specimens: Costa Rica, Puntarenas, 4 km NE San Luis de Guacimal, 22. ii. 1987, E. Giesbert, Calligrapha argus Stål J. Gómez-Zurita det. 2011. IBE-JGZ: (1) one specimen: IBE-JGZ- 0077, Costa Rica, Guanacaste, Area Conservación Guanacaste, Sector Santa Elena, Potrero Grande, 27 / 05 / 2000, A. Solís leg. MfN: (1) one specimen: Costa Rica, Piedras Negras, Collection Schild-Burgdorf, Calligrapha notatipennis Stål. NHM: (1) one specimen: Costa Rica, 92 - 18, C. suboculata Stål; (2) one specimen: Irazu, 6 – 7000 ft, H. Rogers, Godman-Salvin Coll., Biol. Centr. - Amer.; (3) one specimen: Surface of fallen tree, Dry tropical forest, Costa Rica, Volcán Rincón de la Vieja, 10 º 47 ' N 85 º 19 ' W 2700 ', 18. iii. 1982, R. J. Kirby & S. A. Speight BM 1982 - 260. NMB: (1) one specimen: Turrialba, Costa Rica, C. argus ab. famularis St. J. Bechyné det. 1951; (2) one specimen: Bebedero, Costa Rica, Reimoser; (3) one specimen: Turrialba, Costa Rica. NMCZ: (1) two specimens: Costa Rica, Coll. Achard Mus. Pragense [one with: Calligrapha argus St.]. NMNH: (1) one specimen: Costa Rica, F. Nevermann, 17. iii. 30, Turrucares 600 m, Pacifikseite, unter loser Rinde vorsteckt, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (2) one specimen: Costa Rica, F. Nevermann, Guanacaste, Mai 1932, P. Assmann leg.; (3) one specimen: Costa Rica, F. Nevermann, 17. i. 30, Sta Maria (Tilerán) 800 m, W. S. Thomas leg.; (4) one specimen: Costa Rica, Guanacaste, Hacienda Palo Verde, 6 – 15 July 1976, J. C. Solomon coll., 122, Calligrapha argus Stål J. Gómez- Zurita det. 2011; (5) one specimen: Costa Rica, Cent. Am. ix. 1.1906, F. Monrós Collection 1959; (6) two specimens: Finca La Pacífica, 7 km NW Cañas, Guanacaste Prov., Costa Rica, DHJ, Jun, 17 – 24, 1969 [one with: Calligrapha argus Stål J. Gómez-Zurita det. 2011]; (7) one specimen: Costa Rica, Rio Corobici, Las Canas, 15 June 1967, Flint & Ortiz; (8) one specimen: Costa Rica, Guanacaste Prov., Lomas Barbudel Res., 13 July 1989, leg. David G. Furth, Calligrapha argus Stål J. Gómez-Zurita det. 2011. TAMUIC: (1) one specimen: Costa Rica, Guanacaste, 29 km WSW Cañas, Sta. OTS Paloverde, 10 º 21 ’ N 85 º 21 ’ W, 30 June 1976, H. A. Hespenheide, Calligrapha argus Stål J. Gómez-Zurita det. 2011. EL SALVADOR EGRC: (1) one specimen: El Salvador, Ahuachapan, Apaneca, 4500 ft, 7 – 12. ix. 2002, D. Marqua, Calligrapha argus Stål J. Gómez-Zurita det. 2011. FSCA: (1) one specimen: El Salvador, Tamanique 1000 m, 5. xii. 1971, S. & L. Steinhauser, Calligrapha argus Stål J. Gómez-Zurita det. 2011. NMNH: (1) one specimen: El Salvador, Vol. Conchagua, 27 – 29 May 1958, O. L. Cartwright, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (2) two specimens: No. 444 - 2806, 5. vii. 54, Metapan, M. S. V.; (3) two specimens: No. 444 - 7813, 15. vi. 54, La Unión, M. S. V. GUATEMALA EGRC: (1) one specimen: Guatemala, Guat. City, Las Hamacas Tr. Pk., 5 – 8. viii. 1979, Thomas & Case, Calligrapha argus St. det. Daccordi ’ 81. FSCA: (1) one specimen: Guatemala, Baja Veracruz, 6 – 9 km E Purulhá 5000 ’, 15 – 24. iv. 1990, E. Giesbert, Calligrapha argus Stål J. Gómez-Zurita det. 2011. MCZ: (1) three specimens: Chacoj, Vera Paz, Champion, Ex Godman and Salvin [one with: Calligrapha argus Stål]; (2) one specimen: Capetillo, Guatemala, C. Champion, 1 st Jacoby Coll.; (3) one specimen: Chacoj, Vera Paz, Champion, 1 st Jacoby Coll.; (4) one specimen: Panzos, Vera Paz, Conradt, Jacoby 2 nd; (5) one specimen: Teleman, Vera Paz, Champion, Jacoby 2 nd Coll.; (6) one specimen: Chacoj, Vera Paz, Champion, Jacoby 2 nd Coll. MfN: (1) two specimens: Chacoj, Vera Paz, Champion, 96424 [one with: Calligrapha argus St.]; (2) one specimen: Guatemala, [illegible]. MTJM: (1) one specimen: Guatemala, Suchitepequez Dept., Finca Los Tarrales, ca. 8 km N of Patulul, 14 º 32.37 ’ N 91 º 08.17 ’ W, 760 m, 4 June 2005, R. S. Zack, Calligrapha argus Stål J. Gómez-Zurita det. 2009. NHM: (1) one specimen: El Reposo, 800 ft., Champion, Godman-Salvin Coll., Biol. Centr. - Amer.; (2) four specimens: Chacoj, Vera Paz, Champion, Godman-Salvin Coll., Biol. Centr. - Amer.; (3) two specimens: S. Geronimo, Guatemala, Champion, Godman-Salvin Coll., Biol. Centr. - Amer.; (4) one specimen: Cerro Zunil, 4000 ft., Champion, Godman- Salvin Coll., Biol. Centr. - Amer.; (5) one specimen: Capetillo, Guatemala, Rodríguez, Biol. Centr. Amer. Collection; (6) one specimen: Checoj, Vera Paz, Champion, Biol. Centr. Amer. Collection; (7) one specimen: Cerro Zunil, 4 – 5000 ft., Champion, Godman-Salvin Coll., Biol. Centr. - Amer. NMNH: (1) one specimen: Guatemala, Chicacao, 13. vii. 1949, T. H. Farr, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (2) two specimens: Chacoj, Vera Paz, Champion [one with: Calligrapha argus Stål]; (3) one specimen: Guatemala, San Pedro, 11823, 9 - 13 - ’ 60, 22083, in Orchidaceae, Calligrapha sp. DMW. NRM: (1) two specimens: Chacoj, Vera Paz, Champion; (2) one specimen: Cerro Zunil, 4 – 5000 ft. Champion, Calligrapha argus Stål; (3) one specimen: Cerro Zunil, 4 – 5000 ft. Champion; (4) one specimen: Capetillo, Guatemala, G. C. Champion. OUMNH: (1) nine specimens: Chacoj, Vera Paz, Champion, B. C. A. duplicates pres. 1909 by F. D. Godman Cat. No. 84; (2) one specimen: Panajachel, 5000 ft, Champion, B. C. A. duplicates pres. 1909 by F. D. Godman Cat. No. 84, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (3) one specimen: Panzos, Vera Paz, Conradt, B. C. A. duplicates pres. 1909 by F. D. Godman Cat. No. 84, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (4) one specimen: Capetillo, Guatemala, G. C. Champion, B. C. A. duplicates pres. 1909 by F. D. Godman Cat. No. 84, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (5) six specimens: Cerro Zunil, [four with: 4 – 5000 ft; two with: 4000 ft], Champion, B. C. A. duplicates pres. 1909 by F. D. Godman Cat. No. 84, Calligrapha argus Stål J. Gómez-Zurita det. 2011. ZSM: (1) sixteen specimens: Guatemala, Suchitepequez, Patului, [one with: 15.8.83; three with: 1.7.83; two with: 10.8.83; three with: 16.8.83; seven with: 10.7.83], A. Poll, Calligrapha argus Stål J. Gómez-Zurita det. 2011. HONDURAS EGRC: (1) one specimen: Honduras, Cortes Hill, behind San Pedro Sula, 17. ix. 1984, C. W. O’Brien. FSCA: (1) two specimens: Honduras, Intibuca, 18 km W La Esperanza, 3. xii. 1995, F. W. Skillman; (2) one specimen: Honduras, Choluteca, Cerro Guanacuare, 4. vi. 1993, F. W. Skillman, Jr.; (3) one specimen: Honduras, Francisco Morazon Zamorano, 4. x. 1993, R. Turnbow, Calligrapha argus Stal Det. E. G. Riley ’ 93; (4) one specimen: Honduras, Francisco, Morazon, 25.5 km SSW Talanga, 3. vi. 1993, M. C. Thomas, Calligrapha argus Stål J. Gómez- Zurita det. 2011; (5) four specimens: Honduras, El Paraíso, Yuscarán, R. Turnbow, Calligrapha argus St. Det. E. G. Riley ’ 03 [three with: 14. vii. 2001; one with: 21. vii. 2001]; (6) one specimen: Honduras, El Paraíso, Yuscarán 840 m, 4 – 8. viii. 1992, L. Stange & C. Porter, degradated wet forest, Calligrapha argus Stål J. Gómez-Zurita det. 2011. MCZ: (1) one specimen: Sn. P. Sula, Hond. NHM: (1) one specimen: Honduras, 45 - 123; (2) two specimens: Hond. [illegible], Baly Coll. NMNH: (1) one specimen: Hond., El P. vic Yuscaran, 18 May 1995, J. E. Wappes, Calligrapha argus (Stål) det. J. E. Wappes 2002. TAMUIC: (1) one specimen: X 0534460, Honduras, Comayagua, 30 mi E Tegucigalpa, 31. vii. 1982, Robert W. Jones, Calligrapha argus Stål J. Gómez-Zurita det. 2011. MEXICO EGRC: (1) three specimens: Mexico, Tamaulipas, Bocatoma, 7 km SSE Gomez Farias, 27 – 28. v. 1979, E. G. Riley, on Guazuma ulmifolia (Sterculiaceae); (2) two specimens: Mexico, Tamaulipas, Bocatoma, 7 km SSE Gomez Farias, [one with: 25 – 30. iii. 1978; one with: 19 – 23. v. 1979], E. G. Riley; (3) four specimens: Mexico, Tamaulipas, Bocatoma, 6 mi S Gomez Farias, 25 – 30. iii. 1978 [one with: F. Breitenbach; one with: Marlin E. Rice Coll.]; (4) six specimens: Mexico, San Luis de Potosí, El Salto, at the falls, 26. v. 1979 [three with: Marlin E. Rice Coll.; three with: E. G. Riley]; (5) one specimen: Mexico, Oaxaca, 37 mi NW Tehuantepec, 1500 ft, 12. viii. 1974, O’Brien & Marshall; (6) one specimen: Mexico, Chiapas, 16 km W Ocozocautla, Aguacera, 16. vi, D. B. Thomas; (7) one specimen: Mexico, Chiapas, Oaxaca border, Hwy 90, 28. vii. 1988, D. B. and A. M. Thomas; (8) one specimen: Mexico, Chiapas, El Aguacero, 25. vi. 1989, P. Lago, J. Burne & D. Thomas; (9) three specimens: Mexico, Chiapas, El Aguacero, D. B. & A. M. Thomas [one with: 14. vi. 1988; two with: 29. vii. 1988, Calligrapha argus Stål J. Gómez-Zurita det. 2011]; (10) one specimen: Mexico, Chiapas, Simojovel, 20. x. 1988, Thomas, Calligrapha argus Stål J. Gómez-Zurita det. 2011. FSCA: (1) one specimen: Mexico, Chiapas, 16 km W Ocozocoautla, 20. vi. 1987, E. Giesbert, Calligrapha argus St. Det. J. Watts 1993; (2) one specimen: Mexico, San Luis de Potosí, El Salto Falls, 3. vii. 1990, J. K. Adams; (3) one specimen: Mexico, San Luis de Potosi, El Salto Falls, 29. vi. 1968, H. V. Weems, Jr.; (4) one specimen: Mexico, Tamaulipas, 1 – 2 mi E N Morelos, 2. vi. 1982, J. E. Wappes; (5) sixteen specimens: Mexico, Veracruz, 16.5 mi S Catemaco, Hwy 180, 17 – 25. vi. 1985, Askevold & Heffern [one with: Calligrapha argus Stal det. M. Daccordi 1986]; (6) two specimens: Mexico, Veracruz, 5.2 mi S Catemaco, Hwy 180, 17. vi. 1985, Askevold & Heffern [one with: Calligrapha argus Stal det. I. Askevold 1986]; (7) two specimens: Mexico, Veracruz, 1.5 mi NE Tatahuicapan, 25. vi. 1985, Askevold & Heffern; (8) two specimens: Mexico, Veracruz, vic. of El Salto de Eyipantla, 15 km S San Andres Tuxtla, 15 – 28. vi. 1985, Askevold & Heffern; (9) three specimens: Mexico, Veracruz, Lake Catemaco, 7. vii. 1965, G. H. Nelson, sweeping roadside vegetation, Calligrapha argus St. det. A. J. Gilbert ’ 87; (10) one specimen: Mexico, Veracruz, Hwy 145, 6 mi S Tinaja, 25. vii. 1972, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (11) one specimen: Mexico, San Luis de Potosí, 7.4 km E Tancuayalab, 24. vii. 1988, R. Turbow, Calligrapha argus St. det. E. G. Riley ’ 93; (12) one specimen: Mexico, San Luis de Potosí, El Salto, 19. vi. 1973, H. W. Weems, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (13) one specimen: Mexico, Chiapas, Hwy 190, Chiapas de Corsa, 3. viii. 2001, W. Opitz, Calligrapha argus Stål J. Gómez- Zurita det. 2011; (14) one specimen: Mexico, Tamaulipas, Mun. de Aldana, Rancho Nuevo, Barra Coma, 11 – 28. vi. 1979, D. F. Gicca, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (15) one specimen: Mexico, Chiapas, 9 km N Arriaga, 15. x. 1988, E. Giesbert, Calligrapha argus Stål J. Gómez-Zurita det. 2011. MCZ: (1) one specimen: Mex.; (2) one specimen: Mex., A. P. Morse Coll., Calligrapha argus Stål J. Gómez-Zurita det. 2010; (3) one specimen: Tuxtla, San Andrés, Mexico, Sallé Coll., 1 st Jacoby Coll.; (4) one specimen: Yucatan, contigua Chev. [illegible], 1 st Jacoby Coll.; (5) two specimens: Tapachula, Chiapas, Höge, Jacoby 2 nd Coll. MfN: (1) one specimen: 29787, argus Stål, Mexico, Ehrenb.; (2) two specimens: Atoyac; (3) one specimen: Tapachula; (4) one specimen: Sierra de Zungolica [Zongolica], 583, scalaris Lec.; (5) one specimen: Tupataro, 21; (6) two specimens: Mexico, Flohr. NHM: (1) one specimen: Temax, N. Yucatan, Gaumer; (2) one specimen: Tepachula, Chiapas, Höge, Godman-Salvin Coll., Biol. Centr. - Amer.; (3) one specimen: Juquila, Mexico, Sallé Coll., Calligrapha multipunctata Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (4) one specimen: Veracruz, Mexico, Sallé Coll., 643, Sp. figured, Godman-Salvin Coll., Biol. Centr. - Amer.; (5) one specimen: Minas Viejas, Mexico, Dr. Palmer, Godman-Salvin Coll., Biol. Centr. - Amer.; (6) one specimen: Almolonga, Mexico, Hoege, Godman- Salvin Coll., Biol. Centr. - Amer.; (7) one specimen: C. Campeche, Mexico, Sallé Coll., Calligrapha argus Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer. [one with: 644]; (8) one specimen: Calligrapha, Mexico, Pascoe Coll. 9360; (9) one specimen: Chiapas, Mexico, Sallé Coll., Calligrapha famularis Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (10) one specimen: Tuxtla, Mexico, Sallé Coll., Calligrapha famularis Stål apud Sallé, Biol. Centr. Amer. Collection; (11) one specimen: Etla, Mexico, Sallé Coll., Calligrapha famularis Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (12) one specimen: Atoyac, Vera Cruz, May, H. H. S., Calligrapha argus, Godman-Salvin Coll., Biol. Centr. - Amer.; (13) one specimen: Tapachula, Chiapas, Höge, Godman-Salvin Coll., Biol. Centr. - Amer. NMCZ: (1) one specimen: Mexique, famularis Stål, Coll. Achard Mus. Pragense, Calligrapha argus Stål J. Gómez-Zurita det. 2009; (2) three specimens: Mexique, Coll. Achard Mus. Pragense. NMNH: (1) one specimen: Mexico, at Miami, 6. xi. 1968, H. L. Rubin, with Tillandsia sp. plants 68 - 27132, Calligrapha sp. nr. labyrinthica Stal d. R. E. White, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (2) one specimen: Chilpancingo, Guerrero, Mex. ii. 11.46, Laredo Tex. on orchid plant 46 - 1591, Calligrapha sp. perhaps notatipennis Stal HSB’ 46, 46 - 1591; (3) one specimen: Sierra de Durango, gift F. C. Bowditch, Calligrapha argus Stål J. Gómez- Zurita det. 2011; (4) one specimen: Calligrapha sp. Det. S. W. Lingafelter 1997, Ex Mexico on Chamaedorea, III- 17 - 97, Calligrapha argus Stål J. Gómez-Zurita det. 2003; (5) one specimen: Temax, N. Yucatan, Gaumer, Calligrapha multipustulata Stål; (6) one specimen: Tampico Mex., Lar. Tex. 60649, Mar. 13 - 60, 8556, bromeliad, Calligrapha sp. prob. argus Stål DMW; (7) one specimen: Mexico Dist. Federal 31. i. 1949, Laredo, F. Monrós Collection 1959, Calligrapha argus Stål F. Monrós det. 1954; (8) one specimen: Mexique, gift of F. C. Bowditch; (9) one specimen: Chiapas, 19 km N Arriaga, June 23, 1987, J. E. Wappes; (10) one specimen: Tamazunchale, S. L. Potosí, Mex., vi. 19.41, col. & pres. by Henry S. Dybas, F. Monrós Collection 1959, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (11) one specimen: Chiapas, Mex., L. Hotzen ’ 19, Pacific Slope Cordilleras 800 – 1000 m, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (12) five specimens: Mexico, Chiapas, 33.7 mi N Huixtla 6000 ’, ii. 26 – 27.1966, in bromeliads, George E. Ball & D. R. Whitehead collectors; (13) one specimen: Mexico, Veracruz, 22.0 mi E Jalapa, Rte. 140, 1000 ’, iii. 9.66, in bromeliads, George E. Ball & D. R. Whitehead collectors; (14) one specimen: Mexico, Ver., Cordoba, vii. 28.1964, A. B. Lau, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (15) one specimen: Veracruz, Mexico, Sallé Coll.; (16) two specimens: Mexico, Vera Cruz, 30. i. 67, Gonzalez et al. # Brownsville 90583, Lot. # 67 - 4686, Tillandsia sp., Calligrapha sp. Det. R. E. White; (17) one specimen: Mexico, Vera Cruz, 30. i. 67, Gonzalez et al. # Brownsville 90587, Lot. # 67 - 4690, Bromeliads sp.; (18) two specimens: Atoyac, vii. 14.41, Vera Cruz, Mex., Col. by H. S. Dybas, F. Monrós Collection 1959 [one with: Calligrapha argus Stål J. Gómez-Zurita det. 2011; (19) one specimen: Mexico, Campeche, Edizna, 150 ’, iv. 22.1966 in bromeliads, George E. Ball & D. R. Whitehead collectors; (20) two specimens: Mexico, at Laredo 30. xi. 67, Warren & Bauman on bromeliad plnts [one with: Calligrapha sp. nr. diversa Stal d. R. White]; (21) one specimen: Mexico, Laredo 29. iii. 1947, F. Monrós Collection 1959; (22) one specimen: Laredo, Tex., ex. S. L. Potosi, Mex., iii. 19.56, Watt, Tillandsia plant 56 - 3676, Calligrapha sp. DMW 56; (23) one specimen: Mexico, Huichihuayan, S. L. P., iii. 27.58, Fouts colr., on Tillandsia plant, Laredo Tex. 58517, 58 - 6331, Calligrapha sp. det. D. M. Weisman, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (24) one specimen: Mexico, SLP at Brownsville, 11.16.66, Heinrich & Burgess w. bromeliads 66 - 30323, Calligrapha sp. d. R. White; (25) one specimen: Tamazanchule, S. L. P., Mex., ix. 23.46, E. S. Ross collector, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (26) one specimen: Texas / Mexico, 4 I 1972, D. Johnston, San Antonio 2914, Florist whse., Lot. 72 - 2882, Calligrapha sp. det. R. White, Calligrapha argus Stål J. Gómez-Zurita det. 2009. TAMUIC: (1) two specimens: X 0533764 and X 0547082, Mexico, Chiapas, [Aguaceros], 40 km W Tuxtla Gutiérrez, 21. vi. 1987, at light, William F. Chamberlain, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (2) one specimen: X 0534994, Mexico, Chiapas, [Aguaceros], 40 km W Tuxtla Gutiérrez, 20. vi. 1987, William F. Chamberlain, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (3) four specimens: [X 0533940, X 0538585, X 0540150 and X 0544532], Mexico, Veracruz, Panuco, 27. viii. 1962, William F. Chamberlain, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (4) one specimen: X 0537162, Mexico, Veracruz, San Andrés Tuxtla, 15. vii. 1964, Merril H. Sweet; (5) two specimens: [X 0539034 and X 0541214], Mexico, Veracruz, Los Tuxtlas Range, Laguna Encantada, 18. viii. 1960, D. C. Robinson; (6) two specimens: [X 0542378 and X 0546254], Mexico, Veracruz, Catemaco, 22. viii. 1967, Horace R. Burke & J. Hafernik; (7) one specimen: X 0547338, Mexico, Tabasco, 20 km W Cardenas, Campo Exp. CSAT, 22. vii. 1980, Joseph C. Schaffner, Janis Weaver & Timothy P. Friedlander; (8) one specimen: X 0552475, Mexico, Chiapas, 9.5 mi NW Pichucalco, 3. viii. 1980, Joseph C. Schaffner, Jannis Weaver & Timothy P. Friedlander. NICARAGUA IBE-JGZ: (1) two specimens: [IBE-JGZ-C 105, IBE-JGZ-C 106], Nicaragua, Rivas, Finca Sierra Serena UTM 16 P- 657865 - 1242291, 21 September 2009, J. - M. Maes leg.; (2) one specimen: IBE-JGZ-C 232, Nicaragua, Rivas, Finca Guadalupe, 10 – 12 / 07 / 2010, J. - M. Maes leg.; (3) one specimen: IBE-JGZ-C 233, Nicaragua, Estelí, P. T. P. Miraflor-Moropotente, Finca Las Flores, 13.192742 N 86.331123 W 870 m, Abril 2010, J. - M. Maes leg.; (4) two specimens: [IBE-JGZ-C 240, IBE-JGZ-C 254], Nicaragua, Rivas, Finca Isla Vista, 6 – 7 / 07 / 2010, J. - M. Maes leg.; (5) four specimens: [IBE-JGZ-C 257, IBE-JGZ-C 258, IBE-JGZ-C 259, IBE-JGZ-C 260], Nicaragua, Rivas, Cárdenas, Finca Sierra Serena, 14 / 07 / 2010, J. - M. Maes leg.; (6) one specimen: IBE-JGZ- 2181, Nicaragua, Estelí, P. T. P. Miraflor-Moropotente, Finca Bartolo UTM 16 P- 0573112 - 1459212 993 m, Junio 2010, A. del Socorro & B. Nimia leg.; (7) one specimen: IBE-JGZ- 2183, Nicaragua, Montelimar, Reserva Natura, parcela 1, 8 / 10 / 2010, J. - M. Maes leg. MCZ: (1) one specimen: Chontales, Nicaragua, 1 st Jacoby Coll. NHM: (1) one specimen: Chontales, Nicaragua, T. Belt, Godman-Salvin Coll., Biol. Centr. - Amer.; (2) one specimen: Janson, Nicarag., Chont. s, Fry Coll. 1905.100. NMNH: (1) one specimen: La Calera, Nic., 15 / viii / 55, trampa luminica, C. Espinoza R. Bradley 57 - 70, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (2) one specimen: Nicaragua, Pachomil, vi. 71, J. Maldonado C.; (3) one specimen: Consequina Slope, Nicaragua, vii. 7.32, M. Willows Jr. collector, Templeton Crocker Exp. 1932, Calligrapha argus Stål J. Gómez-Zurita det. 2011. PANAMA EGRC: (1) one specimen: Panama, Canal Zone, La Pita Signal Station rd., 8. vi. 1976, E. G. Riley; (2) three specimens: Panama, Canal Zone, Fort Kobbe, [one with: 28. vi. 1970; one with: 6. vi. 1976; one with: 15. vi. 1976], E. G. Riley; (3) two specimens: Panama, Canal Zone, Diablo Heights, [one with: 30. iv. 1971; one with: 15. vi. 1970], E. G. Riley; (4) one specimen: Panama, Canal Zone, Gamboa, 11. vi. 1976, E. G. Riley; (5) one specimen: Panama, Canal Zone, Achiote rd, 10 km SW Gatun, 12. vi. 1976, E. G. Riley; (6) one specimen: Panama, C. Z., Cardenas Village, 22. v. 1980, E. G. Riley & D. LeDoux; (7) two specimens: Panama, C. Z., K- 1 rd., near Fort Kobbe, 19. v. 1980, E. G. Riley & D. LeDoux; (8) one specimen: Panama, C. Z., Gamboa, 18. v. 1980, E. G. Riley & D. LeDoux. FSCA: (1) one specimen: Panama, Summit Gardens, Canal Zone, 25. v. 1977, R. Davis, Jr.; (2) one specimen: Panama, Colon, vic. Ft. Sherman, 24. ii. 1999, J. E. Wappes, Calligrapha argus Stal Det. E. G. Riley ’ 01; (3) one specimen: Panama, Panama, 10 km N El Llano, 15. ii. 1987, E. Giesbert, Calligrapha argus Stål J. Gómez- Zurita det. 2011; (4) two specimens: Panama, Canal Zone, Ft. Gulick, H. J. Harlan, E. J. Gerberg Collection, Calligrapha argus Stål J. Gómez-Zurita det. 2011 [one with: June 1979; one with: 17. v. 1979]. IBE-JGZ: (1) three specimens: Panama, Isla Colón, ii / 95, lgt. D. Buzga. MCZ: (1) four specimens: Panama, Panama [one with: Cal. argus St.]; (2) one specimen: Panama, Bella Vista, 2. vii. 1924, N. Banks, Calligrapha argus Stål J. Gómez-Zurita det. 2010; (3) one specimen: V. de Chiriqui, 25 – 4000 ft, Champion, Jacoby 2 nd Coll.; (4) four specimens: Panama, Panama. NHM: (1) one specimen: V. de Chiriqui, 25 – 4000 ft, Champion, Godman-Salvin Coll., Biol. Centr. - Amer.; (2) one specimen: San Lorenzo, Panama, Champion, Godman-Salvin Coll., Biol. Centr. - Amer. NMNH: (1) one specimen: Panama, Chiriqui Prv., 4 km E Boqueti, 16 May 1996, Wappes, Huether & Morris; (2) one specimen: April 23, 1911, Panama Pan., E. A. Schwarz collector; (3) one specimen: Calebra, C. Z., Pan., Collection F. Knab; (4) one specimen: Canal Zone 100 m, 5.0 mi NW Gamboa 9 º 10 ’ 00 ’’ N 79 º 45 ’ 00 ’’ W, sample 5 – 7, 6 Apr 1976, Montgomery & Lubin coll., canopy fogging experiment in Luehea seemanni, Pyrethrin fog; (5) one specimen: Paraiso CZ, Pan, May 9, 11, E. A. Shwarz, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (6) one specimen: Pan., Panama Prov., Cerro Campana 17. v. 1999, Morris / Wappes, Calligrapha argus Stål J. Gómez-Zurita det. 2011. TAMUIC: (1) one specimen: X 0535600, Panamá, Chiriquí, vic. Hornito, 14 – 18. v. 1996, James E. Wappes, Huether & Morris, Calligrapha argus Stål J. Gómez-Zurita det. 2011; (2) one specimen: X 0536682, Panama, Cerro Campana, 7. i. 1994, James E. Wappes, Calligrapha argus Stål J. Gómez-Zurita det. 2011. VENEZUELA NMCZ: (1) one specimen: Caracas, Sallé, simillima var. Stal Caracas, famularis Stal Mex., Coll. Achard Mus. Pragense, Calligrapha argus Stål J. Gómez-Zurita det. 2009; (2) two specimens: Vénézuéla, L. Laglaize, Déc. 1896, Coll. Achard Mus. Pragense [one with: Calligrapha argus Stål J. Gómez-Zurita det. 2009]. NMNH: (1) two specimens: Venezuela, Trujillo, La Puerta 1700 mts., 1. iv. 1947, H. E. Box. NRM: (1) one specimen: Venezuela, Tarnier, … Deg. UNKNOWN SOURCE MfN: (1) five specimens: [no data]. NHM: (1) one specimen: [illegible], Baly Coll.; (2) one specimen: C., 160, Baly Coll.; (3) two specimens: E. Coll. Laferté, 67 - 56; (4) one specimen: 67 - 56; (5) one specimen: E. Coll. Thomson, 67 - 56; (6) one specimen: Baly Coll. NMCZ: (1) one specimen: Coll. Achard Mus. Pragense; (2) one specimen: Calligrapha famularis St., Coll. Achard Mus. Pragense, Calligrapha argus Stål J. Gómez-Zurita det. 2009. NMNH: (1) one specimen: Amer. Mer., gift of F. C. Bowditch; (2) one specimen: Norway, 17. ix. 1971, L. Beikman, aircraft Lot 72 - 1433, Calligrapha sp. d. R. White. NRM: (1) one specimen: 342, Calligrapha argus.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D640EFF8E8F9CFD46FDBEFD8D.taxon	description	Variation. My interpretation of C. argus is perhaps more consistent with considering this taxon a species complex, given that it exhibits a certain degree of variation that would allow further subdividing it into more taxa. Nevertheless, in general most specimens conform to the type in several characters, including divergent basal ends of subsutural stripe, free humeral marking, free and round spot enclosed by humeral lunule, divergent basal end and roundish divergent apical end of arcuate band, well-delimited and orderly arranged additional spots on disc, and brick-reddish tint to dark body parts. However, some specimens tentatively placed within this group with large midlateral spot, laterally confluent with margin of elytron, deviate in one or several characters from the type and are reminiscent of C. multipustulata. These include a narrow connection of humeral marking with dark basal margin of elytron; an irregular spot enclosed by humeral lunule narrowly connected to it; an humeral lunule with its apical half separated from the rest; a broken, irregular apex of arcuate band; certain variation in the number of additional spots due to confluence / fragmentation (although they are generally ten), the innermost spot usually confluent with external concavity of arcuate band; and / or the pigmentation can be much darker, blackish bronzy, and the contrast between pale background and paler area surrounding markings can be exacerbated. In my opinion, and judging from the illustration in Biologia Centrali Americana for C. famularis, Jacoby (1882) could have interpreted this taxon as some of these atypical forms of C. argus, in particular the large darker forms similar to C. multipustulata typical from Nicaragua. This interpretation would have grounded his proposition of C. famularis as an aberration of C. argus. The specimens from Venezuela held at MNPC have somewhat larger markings, with the spot enclosed by humeral lunule large, ovoid, elongated anteriad, and the spot on the concavity of arcuate band large, slightly oblique and narrowly connected to the latter marking, and they are also reminiscent of the configuration observed in C. famularis.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6405FF928F9CFDFDFA78FA7F.taxon	distribution	Distribution (Fig. 7). The species, which is not rare in collections, was until now considered endemic from Costa Rica, most frequent in the northern half of the country. However, in the course of this study I have found specimens collected in coastal localities along the Bahia of Coronada in the Pacific south of Costa Rica, and one specimen seemingly collected in a Panamian locality, nearby the Costarican border, expanding its known range to the south. In any case, it is a Caribbean Mesoamerican endemic species. The Colombian and Mexican records at NMNH most likely represent mislabelled specimens.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6405FF928F9CFDFDFA78FA7F.taxon	materials_examined	Material examined (291 specimens). COLOMBIA NMNH: (1) one specimen: Colombie, gift of F. C. Bowditch, gift ex MCZ Dupl. Series. COSTA RICA EGRC: (1) one specimen: Costa Rica, Punt. Pr., Monteverde area, 4 – 6. vi. 1980, J. E. Wappes, Calligrapha elegantula Jac. Daccordi ’ 81; (2) twenty-six specimens: Costa Rica, Puntarenas, Monteverde, 26. v – 3. vi. 1984, E. Riley, D. Rieder & D. LeDoux; (3) three specimens: Costa Rica, Puntarenas Prov., Monteverde 1300 – 1600 m, 6. i. 1974, C. Barfield, taken from vegetation; (4) two specimens: Costa Rica, Guan., La Pacífica nr. Cañas, 22 – 26. v. 1984, E. Riley, D. Rieder & D. LeDoux. FSCA: (1) one specimen: Costa Rica, Puntarenas, Monteverde area, 4 – 6. vi. 1980, J. E. Wappes, Calligrapha elegantula Jac. Det. E. G. Riley ’ 82; (2) two specimens: Costa Rica, Puntarenas, Monteverde, 3. vi. 1984, E. Riley, D. Rider & D. LeDoux; (3) one specimen: Costa Rica, Ala. Province, 6 – 8 km W Atenas, 1 – 2. vi. 1980, J. E. Wappes, Calligrapha elegantula Jac., Det. E. G. Riley ’ 86; (4) one specimen: Costa Rica, Puntarenas, San Luis (Monteverde) 3900 ’, 12 – 13. v. 1996, E. Giesbert; (5) one specimen: Costa Rica, San Jose, Monteverde, 21. vi. 1976, at blacklight, J. K. Balciunas & P. P. Habeck; (6) one specimen: Costa Rica, Puntarenas, Monteverde, 27. ii. 1987, E. Giesbert; (7) two specimens: Costa Rica, Puntarenas, 4 km NE San Luis de Guacimal, 22. ii. 1987, E. Giesbert. IBE-JGZ: (1) three specimens: [IBE-JGZ- 0073, IBE-JGZ- 0074, IBE-JGZ- 0075] Costa Rica, Zarcero, Alfaro Ruíz, 4 / vi / 2000, A. Solís leg., Calligrapha elegantula Jacoby, 1877 J. Gómez- Zurita det. 2003. MCZ: (1) one specimen 4: Cache, Costa Rica, H. Rogers, 1 st Jacoby Coll., Type 17438 [red label], 4. Specimen labelled as type prior to this study. Jan. Jul. 2004. MCZ Image Database; (2) one specimen: 473, Costa Rica, 473 Calligrapha elegantula Jac. [male; female] […]; (3) one specimen: C. Rica, elegantula; (4) one specimen: Costa Rica; (5) one specimen: Cartago; (6) one specimen: Costa Rica, Jacoby 2 nd Coll.; (7) two specimens: Costa Rica, Van Patten [one with: C. elegantula Jac.]; (8) one specimen: C. Rica, Calligrapha elegantula Jac.; (9) one specimen: Collection Schild-Burgdorf, Costa Rica, Zarzero, 7, F. A. Eddy Collection; (10) one specimen: Collection Schild-Burgdorf, Costa Rica, San Jose; (11) five specimens: Hamburg Farm, March, Costa Rica, C. P. Dodge; (12) seven specimens: C. Rica [one with: Cal. elegantula Jac.]; (13) three specimens: Cache, Costa Rica, H. Rogers, 1 st Jacoby Coll.; (14) one specimen: Irazu 6 – 7000 ft, H. Rogers, 1 st Jacoby Coll.; (15) four specimens: Costa Rica, Jacoby 2 nd Coll.; (16) one specimen: Bebedero, Costa Rica (Underwood, 94); (17) one specimen: San José, Costa Rica (Underwood); (18) seven specimens: C. Rica; (19) one specimen: Escazu, Costa Rica, Calligrapha elegantula Jacoby J. Gómez-Zurita det. 2010; (20) one specimen: Cariblanco, Costa Rica, Calligrapha elegantula Jacoby J. Gómez-Zurita det. 2010. MfN: (1) four specimens: Costa Rica, Van Patten, 96374 [one with: C. elegantula Jac.]; (2) one specimen: Irazu, 6 – 7000 ft., H. Rogers, Calligrapha elegantula Jac.; (3) one specimen: R. Susio, Costa Rica, H. Rogers; (4) three specimens: Carthago, S. José, Costa Rica, C. Wercklé G.; (5) five specimens: 39600, Costa Rica, Fratz. [four specimens without data]; (6) three specimens: Collection Schild-Burgdorf, Costa Rica, San Jose [one with: elegantula Jac.]; (7) two specimens: Collection Schild-Burgdorf, Costa Rica, Zarzero. MTJM: (1) two specimens: Costa Rica, Monteverde, 10 º 29 ’ N 84 º 50 ’ W, 1400 m, R. D. Akre, Calligrapha elegantula Jacoby J. Gómez-Zurita det. 2009 [one with: Feb. 5, 1963; one with: Feb. 6, 1963]. NHM: (1) twelve specimens: Costa Rica, Sharp Coll. 1905 - 313; (2) four specimens: Costa Rica, Rogers, Sharp Coll. 1905 - 313; (3) two specimens: Costa Rica, 92 - 48 [one with: Calligrapha elegantula Jac.]; (4) two specimens: Costa Rica, Van Patten, Godman-Salvin Coll., Biol. Centr. - Amer. [one with: Calligrapha elegantula]; (5) seven specimens: R. Susio, Costa Rica, H. Rogers, Godman- Salvin Coll., Biol. Centr. - Amer. [one with: C. elegantula Jac.]; (6) three specimens: Irazu, 6 – 7000 ft, H. Rogers, Godman-Salvin Coll., Biol. Centr. - Amer. NMB: (1) one specimen: Collection Schild-Burgdorf, Costa Rica, Atenas, Calligrapha elegantula J. Bechyné det. 1951; (2) one specimen: Collection Schild-Burgdorf, Costa Rica, Zarcero; (3) two specimens: Carpintera, Costa Rica, Reimoser. NMCZ: (1) one specimen: Costa Rica, 475, Coll. Achard Mus. Pragense; (2) six specimens: Costa Rica, Coll. Achard Mus. Pragense. NMNH: (1) two specimens: Quepos, Puntarenas C. R., 19 - 5 - 55, on Sida sp. [one with: Calligrapha elegantula Jacoby J. Gómez-Zurita det. 2011]; (2) one specimen: La Carpentera, Costa Rica, F. Monrós Collection 1959, Calligrapha elegantula Jac., F. Monrós det. 1954; (3) two specimens: San Jose, Costa Rica, F. Monrós Collection 1959; (4) two specimens: C. Rica, F. Monrós Collection 1959; (5) three specimens: Costa Rica, 30 - iii- 1948, F. Monrós Collection 1959; (6) twelve specimens: Costa Rica, El Alto de Ochomogo, 28. vi. 934, C. H. Ballon s / maiz escobilla, F. Monrós Collection 1959; (7) one specimen: Costa Rica, San Ramón de Tres Ríos, ix. 1991, J. M. Maes, Calligrapha elegantula Jacoby 1991 Det. C. L. Staines; (8) one specimen: Irazu, Costa Rica, Coll. Schild & Burgdorf; (9) four specimens: Zarzero, Costa Rica, Coll. Schild & Burgdorf; (10) one specimen: Costa Rica, Van Patten; (11) two specimens: San Jose, C. R., J. F. Tristan coll. [one with: Apr. 28]; (12) three specimens: C. Rica, F. Monrós Collection 1959; (13) one specimen: La Palma, C. R., 1500 m alt., C. H. Lankester collector, Calligrapha elegantula Jac. F. Monrós det. 1953; (14) one specimen: San Jose, C. R., 1160 m, M. Valerio coll., Calligrapha elegantula Jac. F. Monrós det. 1953; (15) eight specimens: El Alto de Ochomogo, C. R., 6.28.34, C. H. Ballou, on corn escobilla, C. R. No 1368 [one with: Calligrapha elegantula Jac. HSB]; (16) twelve specimens (within a capsule): El Alto de Ochomago, C. R., 6 - 28 - 34, C. H. Ballou, on corn + escobilla, C. R. No. 1368, Calligrapha elegantula Jacoby J. Gómez-Zurita det. 2003; (17) one specimen: Irazu, 6 – 7000 ft, H. Rogers; (18) one specimen: Costa Rica, V. Patten, Calligramma elegantula Jac.; (19) one specimen: La Carpentera, Costa Rica, April 1924, W. M. Mann; (20) one specimen: San Jose, C. R., 1160 m, M. Valerio coll.; (21) one specimen: Costa Rica, J. F. Tristan coll.; (22) one specimen: 11990, Turrialba C. R., 27 - 2 - 71, J. V. Mankins collector, Calligrapha elegantula Det. 1977, J. V. Mankins; (23) two specimens: Costa Rica, Heredia, Finca La Selva, 21 – 30 July 1976, J. C. Solomon Coll.; (24) six specimens: Costa Rica, F. Nevermann, iii. 33, Guayabillos SW Abhang Irazu 2200 m [one with: iii. 33]; (25) one specimen: Costa Rica, F. Nevermann, 25. viii. 28, San Jose 1000 – 1200 m, an grasnarbe; (26) three specimens: Costa Rica; (27) three specimens: Costa Rica, Oct.; (28) two specimens: Coronado, Costa Rica, July 1923, T. Assman, Nevermann Collection 1940; (29) two specimens: San Jose, C. R., March 1921, Nevermann Collection 1940; (30) one specimen: Costa Rica, Monteverde, November 1981, N. J. Besansky; (31) ten specimens: Costa Rica, Puntarenas, Monteverde area, 6 – 14 June 1973, 1400 – 1700 m, Erwin & Hevel Central American Expedition 1973; (32) two specimens: Costa Rica, Punt. Prov., Monteverde 1430 mts., 23 February 1978, E. R. Snyder Hodges; (33) four specimens: Costa Rica, Heredia, 1800 m, Monte de la Cruz, 1 July 2001, C. L. Staines; (34) one specimen: on bromeliad plants, San Jose, Costa Rica, iv. 5.61, A. S. Mills Coll, Miami 16264, 10 t 61 - 11378, Calligrapha sp. det. D. M. Weisman; (35) one specimen: Irazu, 6 – 7000 ft, H. Rogers; (36) one specimen: Costa Rica, San Jose 10 km N, 9 Aug 1972; (37) one specimen: Costa Rica, Volcan Irazu, 15 May 1951, O. L. Cartwright; (38) one specimen: NW Costa Rica, Oct. 22 – 31, 1992, Coll. R. Pienkowski; (39) one specimen: Costa Rica, San Jose 1100 – 1200 m, August 1980, N. L. H. Krauss; (40) one specimen: Costa Rica, Vista de Mar, xi. 21.1965, N. L. H. Krauss; (41) two specimens: Costa Rica, Heredia, Hwy 32 at San Jose border, 8 January 1995, C. L. & S. L. Staines. NRM: (1) one specimen: Costa Rica, Van Patten, Calligr. elegantula Jac. OUMNH: (1) one specimen: Cache, Costa Rica, H. Rogers, B. C. A. duplicates, presented 1909 by F. D. Godman, Cat. No. 84, Calligrapha diversa Stål, Calligrapha elegantula Jacoby J. Gómez-Zurita det. 2011. TAMUIC: (1) two specimens: [X 0537769 and X 0540719], Costa Rica, Alajuela, 10 km N San Pedro, 26. vii. 1990, William F. Chamberlain; (2) one specimen: X 0540763, Costa Rica, Cartago, Juan Viñas, 5. viii. 1990, William F. Chamberlain; (3) one specimen: X 0540842, Costa Rica, Puntarenas, Monteverde, 2. viii. 1990, G. M. Chamberlain; (4) one specimen: X 0541102, Costa Rica, Puntarenas, Monteverde, 20 – 24. vi. 1986, W. Hanson & G. Bohart; (5) two specimens: [X 0541496 and X 0547150], Costa Rica, San José, San José, 19 – 20. xii. 1987, F. D. Parker; (6) two specimens: [X 0547858 and X 0549945], Costa Rica, San José, San José, 27 – 29. xii. 1987, F. D. Parker; (7) five specimens: [X 0542347, X 0546293, X 0546589, X 0547506 and X 0550458], Costa Rica, Puntarenas, Monte Verde, 6. i. 1974, taken from vegetation, Carl Barfield; (8) one specimen: Costa Rica, San José, Escazu, 5 – 6. vii. 1988, F. D. Parker. MEXICO NMNH: (1) one specimen: Mexico, R. F. Pearsall, Brooklyn Museum Colln 1929. PANAMA FSCA: (1) one specimen: Panama, Chiriquí, Hartmann’s Finca, 4 – 6. vii. 1997, J. Huether. UNKNOWN SOURCE MCZ: (1) one specimen: elegantula; (2) one specimen: F. A. Eddy Collection; (3) one specimen: 38, Jacoby 2 nd Coll. MfN: (1) one specimen: [no data]. OUMNH: (1) one specimen: E. Coll. Laferté, bor, Calligrapha persimilis Reiche, Amer. bor., From the Brit. Mus. duplicates Pres. 1899, Calligrapha elegantula Jacoby J. Gómez- Zurita det. 2011.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6405FF928F9CFDFDFA78FA7F.taxon	description	Variation. Species very constant in appearance, showing slight differences in elytral maculation, for instance different confluence patterns among additional spots on disc, these with apex of arcuate band, or with apical spot or the spot on apical declivity; the basalmost spot can also be narrowly confluent with humeral marking externally. The basal end of the subsutural stripe is commonly confluent with sutural stripe, but sometimes it is slightly separated from it for a short distance. I have seen one specimen with the spot enclosed by humeral lunule confluent with the latter and another with the basal end of the arcuate band. The spot of apical declivity appeared very small in one specimen. Finally, in a single specimen at NMCZ, the apex of arcuate band presents two consecutive lobes.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6419FF988F9CF9A6FB7BFE4A.taxon	distribution	Distribution. This species is, together with C. argus, the one with the largest geographic range in this group, from southern Arizona (where it has been confused with C. sylvia), and one of the few reaching South America, to Venezuela (Fig. 14, black circles). For this work, I have seen specimens from Colombia, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, USA and Venezuela. Considering the information compiled for this study, in my opinion Crotch's (1873) record of C. sylvia from Arizona must have corresponded to C. diversa. Unfortunately, other authors perpetuated that interpretation until today, even though some early accounts, like that of Linell (1896), were critical about the presence of this species in the United States. In the current interpretation for the species as proposed here, this would be the Calligrapha species in this group with the largest geographic range, from the California-Rocky Mountain to the Caribbean Northwest South American domain. However, it is worth noting that the species seems to be absent from Panama, and the records from Costa Rica are scarce and without locality data, thus representing the only known case of a Calligrapha species with a disjunct distribution. This special distribution, coupled with a certain morphological diversity (see below) may suggest that C. diversa represents a species complex with several species involved, but I find it premature to split it in additional taxa.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6419FF988F9CF9A6FB7BFE4A.taxon	materials_examined	Material examined (578 specimens). CANADA MCZ: (1) one specimen: Calligrapha spiraea Say Canada. COLOMBIA MCZ: (1) one specimen: Cali, Colombia, ix. x. 1.94, W. Rosenberg, Calligrapha diversa Stål J. Gómez-Zurita det. 2010. NHM: (1) three specimens: E. Coll. Laferté, 67 - 56 [one with: Colombia]; (2) one specimen: Colom. COSTA RICA NHM: (1) four specimens: Costa Rica, V. Patten, Godman-Salvin Coll., Biol. Centr. - Amer.; (2) one specimen: Costa Rica, 201, 67 - 56, diversa Stål. EL SALVADOR NMHN: (1) one specimen: No. 444.2813, 5. vii. 54, Metapan, Col. M. S. V., Calligrapha diversa Stål J. Gómez- Zurita det. 2011. GUATEMALA FSCA: (1) one specimen: Guatemala, Baja Veracruz, rd to Santa Rosa, 6. x. 2007, J. & M. Huether, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (2) one specimen: Guatemala, Guatemala, Guatemala, 7. vii. 1984, R. Pérez, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (3) one specimen: Guatemala, Antigua, 25 – 28. iv. 1999, P. J. Landolt, Calligrapha diversa Stål J. Gómez-Zurita det. 2011. MCZ: (1) one specimen: Capetillo, Guatemala, G. C. Champion; (2) one specimen: Gua.; (3) seven specimens: Guatem.; (4) fourteen specimens: Guatemala; (5) one specimen: Duenas, Guatemala, C. Champion, 1 st Jacoby Coll.; (6) one specimen: Capetillo, Guatemala, C. Champion, 1 st Jacoby Coll.; (7) one specimen: Mexico, 1 st Jacoby Coll.; (8) one specimen: Coban, Vera Paz, Conradt, Jacoby 2 nd Coll.; (9) five specimens: Gua. MfN: (1) one specimen: Duenas, Guatemala, G. C. Champion, 96385, Callig. diversa Stål; (2) one specimen: Capetillo, Guatemala, G. C. Champion, 96387; (3) one specimen: Duenas, Guatemala, G. C. Champion, 96387. NHM: (1) one specimen: Guatemala, Sallé Coll., Calligrapha diversa Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (2) four specimens: Duenas, Guatemala, G. C. Champion; (3) one specimen: Guatemala, Sallé Coll., 673, Godman-Salvin Coll., Biol. Centr. - Amer.; (4) one specimen: Zapote, Guatemala, G. C. Champion, Godman-Salvin Coll., Biol. Centr. - Amer.; (5) one specimen: Capetillo, Guatemala, G. C. Champion, Godman-Salvin Coll., Biol. Centr. - Amer.; (6) one specimen: S. Geronimo, Guatemala, Champion, Godman-Salvin Coll., Biol. Centr. - Amer.; (7) one specimen: Purula, Vera Paz, Champion, Godman-Salvin Coll., Biol. Centr. - Amer.; (8) four specimens: Guat.; (9) one specimen: Guatemala; (10) two specimens: Guatem a 55.71 [one with: 67 - 56]. NMB: (1) one specimen: Guatemala, Escuintla, 1879. viii, Calligrapha diversa Stål J. Bechyné det. 1954. NMCZ: (1) one specimen: Las Nubes, Guatemala, Coll. Achard Mus. Pragense; (2) three specimens: San Marcos, Guatemala, Coll. Achard Mus. Pragense; (3) one specimen: Canalitos, Guatemala, Coll. Achard Mus. Pragense. NMNH: (1) five specimens: Senahu, Alta V. Paz, Guatemala, Paul Haase collector [one with: Calligrapha diversa Stål J. Gómez-Zurita det. 2011]; (2) four specimens: Acatenango, Guatemala, May 1948, H. T. Dalmat Collector [one with: Calligrapha diversa Stål J. Gómez-Zurita det. 2011]; (3) two specimens: Yepocapa, Guatemala, Sept. 1949, H. T. Dalmat collector; (4) two specimens: Coban, Guat., Alta Vera Paz, May 19, 1920, J. M. Aldrich Collector [one with: Calligrapha 20 - maculata Stål, Calligrapha diversa Stål J. Gómez-Zurita det. 2011]; (5) one specimen: Purula, Vera Paz, Champion, Calligrapha diversa Stål; (6) two specimens: Capetillo, Guatemala, G. C. Champion [one with: Calligrapha diversa Stål J. Gómez-Zurita det. 2011]; (7) one specimen: Guatemala, Guatemala City, ix. 1959, NLH Krauss; (8) one specimen: Guatemala, Collection F. Knab; (9) one specimen: Guatemala, Coll. Van Patten, gift ex MCZ dupl. series; (10) one specimen: Guatemala, 17. iii. 65, Lindsay, Gillagry on orchid plants, San Ped 15579, 65 - 5911, Calligrapha sp. nr. geographica Det. G. B. Vogt, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (11) one specimen: San Jose Pinula, Guat., 26. viii. 52, coll. corn, R. H. Painter Coll., Altitude 6500 ft, Calligrapha diversa Stål J. Gómez-Zurita det. 2011. ZSM: (1) three specimens: Guatemala, Alta Verapaz, San Cristobal, [two with: 18.5.79; one with: 4.6.79], Gaigl, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (2) two specimens: Guatemala, Quezaltenango, 6.8.83, A. Poll. HONDURAS FSCA: (1) one specimen: Honduras, F. Morazan, B. Rincon, 15. x. 1993, F. W. Skillman, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (2) one specimen: Honduras, Francisco Morazan, Zamorano, 2. x. 1993, R. Turnbow, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (3) one specimen: Honduras, El Paraíso, Montserrat [Cerro Moncerrato] 5750 ’, 3. x. 1993, F. W. Skillman, Calligrapha diversa Stål J. Gómez-Zurita det. 2011. NMNH: (1) one specimen: 11811, Tola [Tela], Hond. 27 - 10 - 77, J. V. Mankins Collector, Calligrapha diversa Stål J. Gómez- Zurita det. 2011. TAMUIC: (1) one specimen: X 0547342, Honduras, Santa Bárbara, E slope Santa Bárbara, 11. vii. 1968, J. R. Meyer. MEXICO EGCR: (1) one specimen: Mexico, Jalisco, 9 mi SW Autlan, 4300 ft, Hwy 80, 11. viii. 1982, C. and L. O’Brien & G. Wibmer, Calligrapha multipustulata St. det. Daccordi ’ 86, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (2) two specimens: Mexico, Mich., 5 mi NW Tuxpan 6200 ft, 7. viii. 1982, C. W. and L. O’Brien & G. Wibmer; (3) one specimen: Mexico, Ags., Pabellón, 24. vi. 1956, R. P. Allen; (4) one specimen: Mexico, Tamaulipas, along rd to Rancho de Cielo, W Gomez Farias, 2. i. 1981, E. G. Riley; (5) one specimen: Mexico, Tamaulipas, Bocatoma, 7 km SSE Gomez Farias, 19 – 23. v. 1979, E. G. Riley; (6) one specimen: Mexico, Durango, Durango 6500 ft, 31. vii. 1964, J. F. McAlpine; (7) one specimen: Mexico, Durango, 21 mi E Durango, 24. vi. 1964, H. F. Howden; (8) one specimen: Mexico, Durango, 6 mi S Durango, 14. vii. 1964, H. F. Howden; (9) one specimen: Mexico, Mich., 4 mi SE Tuxpan, 6500 ft, Hwy 15, 16. viii. 1982, C. W. and L. O’Brien & G. Wibmer; (10) one specimen: Mexico, Mich., 5 mi NE Patzcuaro 7200 ft, 15. viii. 1982, C. W. and L. O’Brien & G. Wibmer; (11) one specimen: Mexico, Tamaulipas, 11 mi SW Cd. Victoria, 4000 ft, Hwy 101, 22. vii. 1982, C. W. and L. O’Brien & G. Wibmer; (12) one specimen: Mexico, Jalisco, 10 mi NE Tecolotlan, Hwy 80, 5400 ft, 11. viii. 1982, C. W. and L. O’Brien & G. Wibmer; (13) one specimen: Mexico, Nuevo León, 20 mi W Linares, 3250 ft, Hwy 58, 21. vii. 1982, C. W. and L. O’Brien & G. Wibmer; (14) one specimen: Mexico, Mich., 7 mi E Quiroga, 8100 ft, Hwy 15, 15. viii. 1982, C. W. and L. O’Brien & G. Wibmer; (15) one specimen: Mexico, Nuevo León, Cyn. Santa Rosa, 10 km W Linares, 20. v. 1992, D. Thomas, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (16) one specimen: Mexico, Chiapas, Ruinas Chinkultic, 18. x. 1988, J. E. Wappes, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (17) one specimen: Mexico, Chiapas, Ocozocuatla, 23. x. 1988, D. B. & A. M. Thomas, Calligrapha diversa Stål J. Gómez-Zurita det. 2011. FSCA: (1) two specimens: Mexico, Guerrero, 2 km S Ahuacatitlán 5500 ’, 20. ix. 1989, J. E. Wappes; (2) one specimen: Mexico, Colima, W rd to El Terrero, 3 – 5000 ’, 3 – 5. x. 1992, J. E. Wappes; (3) one specimen: Mexico, Guerrero, 10 km S Ixcateopan, 4000 ’, 18 – 20. ix. 1989, E. Giesbert, Calligrapha diversa Stal Det. J. Watts 1993; (4) one specimen: Mexico, Morelos, 6 km W Yautepec, 17 – 18. x. 1984, E. Giesbert; (5) one specimen: Mexico, Oaxaca, Oaxaca, 8 – 16. viii. 1956, D. Muller, coll. at light; (6) one specimen: Mexico, Guerrero, 10.6 km S Ixcateopan, 5400 ’, 18. ix. 1989, R. Turnbow, Calligrapha diversa Stal Det. E. G. Riley ’ 93; (7) two specimens: Mexico, Nayarit, Volcan Ceboruco, 4 – 11 km S Jala, 8. x. 1992, R. Turnbow; (8) one specimen: Mexico, Jalisco, 10.8 mi S Talpa de Allende 4900 ’, 9. viii. 1967, Ball, T. L. Erwin & N. E. Leech, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (9) one specimen: Mexico, Oaxaca, rte. 175, 25.2 mi S Suchixtepec, 8 – 9. vii. 1972, P. A. Meyer & G. E. Ball, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (10) four specimens: Mexico, Chiapas, 8 mi E Rizo de Oro, Hwy 190, 22. vi. 1985, Askevold & Heffern, Calligrapha multipustulata Stal det. Daccordi 1986, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (11) two specimens: Mexico, Jalisco, 1 mi E Acatlan, 14. vi. 1987, B. K. Dozier; (12) one specimen: Mexico, Colima, 4500 ’ El Terrero rd (vic. El Sauz), 30. ix – 4. x. 1991, E. Giesbert; (13) ten specimens: Mexico, Zacatec., Zacateca City, 24. viii. 1985, W. Opitz, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (14) three specimens: Mexico, Chiapas, Hwy 190, Amatenango del Valle, 1. viii. 2001, W. Opitz, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (15) four specimens: Mexico, Guanajuato, San Miguel de Allende, 7 – 11. viii. 1988, G. B. Edwards, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (16) two specimens: Mexico, Jalisco, 54 mi S Autlan de Navarro, 31. viii. 1971, J. M. Cicero, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (17) one specimen: Mexico, Oaxaca, La Pluma Hidalgo 1200 m, 21. vii. 1993, P. J. Landolt, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (18) one specimen: Mexico, Chiapas, Comitlan, 31. vii. 2001, W. Opitz, Calligrapha diversa Stål J. Gómez-Zurita det. 2011. MCZ: (1) two specimens: Amula, Guerrero 6000 ft, Aug. H. H. Smith [one with: Calligrapha diversa Stål]; (2) one specimen: Cuernavaca, Mexico, Calligrapha diversa Stål; (3) three specimens: Tuxpan, Jalisco, Mex. 7 / 9 / 1903, J. F. McClendon; (4) four specimens: Guadalajara, Jalisco, Mex. 6 / 26 / 1903, J. F. McClendon; (5) ten specimens: Tepehuanes, Dgo., Mex., Wickham [one with: Calligrapha diversa Stål J. Gómez-Zurita det. 2010]; (6) two specimens: Tuxpan, Jalisco, Mex. 9 / 6 / 1903, J. F. McClendon; (7) four specimens: Guadalajara, Mex. 8 / 8 / 03 [one with: Calligrapha diversa Stål J. Gómez-Zurita det. 2010]; (8) one specimen: Mexico, Mendoza [?], E. Palmer; (9) one specimen: Guadalupe, Hidalgo, vii. 10.32, Calligrapha diversa Stål J. Gómez-Zurita det. 2010; (10) one specimen: Jacala, Hidalgo 4500 ft, 5. vii. 1989, Mexico, Ralph Haag; (11) one specimen: Bowditch Collection, Mex.; (12) one specimen: Guerr. Mex. 2.26.0 1, F. A. Eddy Collection; (13) one specimen: Chiapas, Mex.; (14) five specimens: Chiapas, Mex., Coll. Van Patten, Peab. Acad.; (15) one specimen: Mexico, Guerrero, 37 km W Iguala, 17 June 1993, leg. D. Furth, Calligrapha diversa Stål J. Gómez-Zurita det. 2010; (16) one specimen: Mexico, Oaxaca, Monte Alban, 23 Aug. 1991, leg. D. Furth; (17) one specimen: Mexico, Sallé Coll., 1 st Jacoby Coll.; (18) one specimen: Oaxaca, Mexico, Hoege, 1 st Jacoby Coll.; (19) two specimens: Cuernavaca, Morelos, June, H. H. S., Jacoby 2 nd Coll.; (20) two specimens: Amula, Guerrero, 6000 ft, Sept., H. H. Smith, Jacoby 2 nd Coll.; (21) one specimen: Chilpancingo, Guerrero, 4600 ft, June, H. H. Smith, Jacoby 2 nd Coll.; (22) one specimen: Xucumanatlan, Guerrero, 7000 ft, July, H. H. Smith, Jacoby 2 nd Coll.; (23) one specimen: Tuxpan, Jalisco, Mex. 7 / 9.1903, J. F. McClendon; (24) one specimen: Guadalajara, Jalisco, Mex., 6 / 26.1903, J. F. McClendon; (25) one specimen: Cuernavaca, Mexico; (26) one specimen: Durango, Mex.; (27) one specimen: Chrysomela signata Klug Mexico; (28) one specimen: Mex., ex. Chev. coll.?; (29) one specimen: Pte. de Ixtla, Mor., Mex., Wickham, Calligrapha diversa Stål J. Gómez-Zurita det. 2010. MfN: (1) two specimens: 29789, diversa Dej., signata Kl., Mexico Deppe [one without data]; (2) one specimen: Guadalajara; (3) two specimens: Mexico, Hoege S., Kotze G.; (4) one specimen: diversa Stål, Mexico; (5) one specimen: V. de Mexico; (6) three specimens: Guanajuato 21 [two without data]; (7) one specimen: Maravatio 13; (8) one specimen: Juquila 15; (9) one specimen: Mex.; (10) one specimen: Canelas; (11) one specimen: Tula, Hidalgo, Höge; (12) one specimen: Mexico, Ob [...]; (13) twenty-one specimens: Mexico, Durango City [one with: diversa Stål]; (14) six specimens: Mexico; (15) one specimen: Mexico, Colima; (16) one specimen: La Borrega, Sª Durº; (17) nine specimens: Durango City; (18) one specimen: Pilar, Durango; (19) one specimen: Mexico, Puebla; (20) one specimen: Mexico, Guadalajara; (21) two specimens: Teopisca; (22) one specimen: Comilan; (23) two specimens: Mexico-Hidalgo, Tula, 31.8.1993, Erber leg. [one with: Calligrapha geographica Stål, D. Erber det. 2000]; (24) one specimen: Mexico-Hidalgo, 20 km westl. Actopan, 1.9.1993, Erber leg., Calligrapha geographica Stal, D. Erber det. 2000; (25) one specimen: diversa Dej. signata Klug, Mexico; (26) one specimen: argus Chevr., formularis Stål Mexico; (27) one specimen: 2, Durango City, C. argus. MTJM: (1) one specimen: Mexico, Mich., Ixtlan, 9. vi. 1950, D. Enkerlin, Calligrapha diversa Stål J. Gómez-Zurita det. 2009; (2) one specimen: Mexico, Chapingo, 7. vii. 1950, C. F. Dowling, Calligrapha diversa Stål J. Gómez-Zurita det. 2009. NHM: (1) one specimen: Named by Stål, Baly Coll., diversa Stål Mexico; (2) one specimen: Calligrapha diversa Dej., signata Klug, Mexico, D r Klug, Baly Coll.; (3) six specimens: Truqui, Mexico, Fry Coll. 1905.100 [one with: 23843]; (4) one specimen: Cuernavaca, Mexico, Sallé Coll., Calligrapha diversa Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (5) one specimen: Etla, Mexico, Sallé Coll., Calligrapha diversa Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (6) one specimen: Juquila, Mexico, Sallé Coll., Calligrapha diversa Stål apud Sallé, Sp. figured, Godman-Salvin Coll., Biol. Centr. - Amer.; (7) one specimen: Oaxaca, Mexico, Sallé Coll., 638, Calligrapha diversa Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (8) one specimen: Ex Coll. J. Sturm, Mexico, C. diversa Dej., Mexico, Sallé Coll., Calligrapha diversa Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (9) two specimens: Xucumanatlan, Guerrero 7000 ft July, H. H. Smith, Godman-Salvin Coll., Biol. Centr. - Amer.; (10) one specimen: Atlisco, Puebla, Höge, Godman-Salvin Coll., Biol. Centr. - Amer.; (11) two specimens: Amula, Guerrero, 6000 ft, Aug., H. H. Smith, Godman-Salvin Coll., Biol. Centr. - Amer.; (12) one specimen: Cuernavaca, Morelos, June, H. H. Smith, Godman-Salvin Coll., Biol. Centr. - Amer.; (13) one specimen: Tupataro, Guanajuato, Höge, Godman-Salvin Coll., Biol. Centr. - Amer.; (14) one specimen: Tula, Hidalgo, Höge, Godman-Salvin Coll., Biol. Centr. - Amer.; (15) one specimen: Chilpancingo, Guerrero, 4600 ft, July, H. H. Smith; (16) one specimen: Mexico City, May. 88, H. H. S., Godman-Salvin Coll., Biol. Centr. - Amer.; (17) one specimen: Cerro de Plumas, Mexico, Hoege, Godman-Salvin Coll., Biol. Centr. - Amer.; (18) one specimen: Morelia, Michoacan, Höge, Godman-Salvin Coll., Biol. Centr. - Amer.; (19) two specimens: Tepetlapa, Guerrero, 3000 ft, Oct., H. H. Smith, Godman-Salvin Coll., Biol. Centr. - Amer.; (20) fifteen specimens: Temascaltepec 1931 [two with: 1831], Mex. D. F., G. B. Hinton Collector, Hinton Coll. B. M. 1937 - 173; (21) two specimens: Mex.; (22) one specimen: Oax.; (23) one specimen: foveicollis Dej. Mexico Sallé; (24) one specimen: Guadalajara, Crawford, 47; (25) one specimen: Mex. M. Coffiss, 58 / 60; (26) one specimen: Cuernavaca VI- 1934 Mex, H. E. Hinton Collector, Hinton Coll. B. M. 1939 - 583; (27) one specimen: Real de Arriba, Temescaltepec, Mex. 6 – 7000 ft v – vi 1933, B. M. 1959 - 100, H. E. Hinton, R. L. Usinger Collectors. NMB: (1) two specimens: Irapuato, Guanajuato; (2) three specimens: Mexico, Erwerb 1955, Coll. Brancsik; (3) one specimen: Durango, Mexico, Erwerb 1955, Coll. Brancsik; (4) one specimen: Durango, Erwerb 1955, Coll. Brancsik; (5) three specimens: Durango, Mexico; (6) three specimens: Mexico [one with: Calligrapha diversa Stål J. Bechyné det. 1951]; (7) two specimens: Mexico 600 m Umg. Monterrey, G. Frey, 27.12.1966, Calligrapha diversa Stål J. Gómez-Zurita det. 2010; (8) four specimens: Mexiko [two with: Calligrapha diversa Stål J. Gómez-Zurita det. 2010]. NMCZ: (1) three specimens: Mexico, Mus. Pragense Col. Kambersky [one with: Calligrapha diversa St.; one with: Calligr. signata]; (2) one specimen: diversa Stål, signata Kl., Spinola Mexico,? diversa Stål Mex., Coll. Achard Mus. Pragense; (3) one specimen: Mexique, Mexico, Calligrapha diversa Stål J. Achard det. in BCA; (4) two specimens: Mexique, Coll. Achard Mus. Pragense; (5) six specimens: Sierra de Durango, Coll. Achard Mus. Pragense [one with: diversa Stål]; (6) one specimen: Mex., Coll. Achard Mus. Pragense; (7) two specimens: Mexique, Guanajuato, Coll. Achard Mus. Pragense. NMNH: (1) one specimen: Mexico, Tamps vic. Gom Farias 1000 ’, 8 May 1994, J. E. Wappes, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (2) one specimen: Mexico, Chiapas, 45 km SE Comitan, Chincultic Ruinas, June 16, 1987, J. E. Wappes, Calligrapha spp. Det. E. G. Riley ’ 89, Calligrapha diversa Stål J. Gómez-Zurita det. 2003; (3) one specimen: Mexico, Chiapas, Ruinas Chinkultic, X- 18 - 1988, J. E. Wappes, Calligrapha diversa Stål J. Gómez-Zurita det. 2003; (4) one specimen: with mustard greens from Mex., 1 July 42, El Paso Tx. # 34245, C. sp. nr. diversa (Stal) HSB 42, Calligrapha diversa Stål J. Gómez-Zurita det. 2009; (5) one specimen: Mexico, Mex., June 9, 1922, E. G. Smyth on malva; (6) thirteen specimens: S. Jacinto, D. F., 8 - 1923 Mex., E. G. Smyth, Chittenden No. 12523 [one with: Calligrapha diversa Stål; one with: Calligrapha diversa Stål J. Gómez-Zurita det. 2011]; (7) sixteen specimens: Colima Vulcano Mex., L. Conrad [one with: Calligrapha diversa Stål; one with: Calligrapha diversa Stål J. Gómez-Zurita det. 2011]; (8) one specimen: Chapultepec, Mex., 6. vii. 01, R. H. Hay collector; (9) one specimen: Mexico; (10) one specimen: State of Colima, Mex., L. Conradt coll.; (11) three specimens: Cuernavaca, Mex., 7 - 7 - 00, El. 5000 ft, O. C. Dean collector [one with: Calligrapha diversa Stål J. Gómez-Zurita det. 2011]; (12) four specimens: Mexico, Morelos, Cuernavaca, NLH Krauss [one with: vii- 45, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; one with: vi- 45; one with: viii- 15 - 44; one with: xi- 9 - 44]; (13) three specimens: Mexico C., Mex., 11 Jly 0 4, F. Kraemer Collector [one with: Calligrapha diversa Stål J. Gómez-Zurita det. 2011]; (14) two specimens: Mexico, D. F., Teotihuacan ruins, 7 September 1972, G. F. & S. Hevel; (15) three specimens: Mexico, Zacatecas, Loreto, 1034 elev. 22 º 16 ’ N 101 º 58 ’ W, 4 July 1995, lg. D. Furth & G. Chavarria; (16) one specimen: Mexico DF, Mex., R. Müller nº 211; (17) one specimen: Mexico, Michoacan, 8.1 mi E Villamar, 5500 ’, Rte 15, vii. 31.1966, George E. Ball & D. R. Whitehead collectors; (18) one specimen: Mexico, Chiapas, 3.1 mi N Pueblo Nuevo, Rte 195, 5800 ’, iv. 27.66, George E. Ball & D. R. Whitehead collectors; (19) two specimens: Jalisco, Mex., June ’ 04, from Dr. G. W. Bock, Collection F. Knab; (20) one specimen: Xucumanatlan, Guerrero 7000 ft. July, H. H. Smith; (21) one specimen: Mexico, Morelos, Hacienda Cocoyotla, nr. Cuatlan del Rio, vii. 31.44, NLH Krauss; (22) one specimen: Cuernavaca, Morelos, Mex., Wickham, Wickham collection 1933; (23) one specimen: Mexico; (24) two specimens: Mexico, Sonora, 3 km E Yecora, Puente La Ventana 28 º 22.640 N 108 º 53.842 W, 6 July 2006, 1510 m, leg. David G. Furth; (25) two specimens: Wickham, Tacuba D. F. Mex., ix. 2; (26) one specimen: Mexico, Imuris, Son. at Nogales 281, viii- 29 - 1969, J. Backe-Wiig, with Zea mays fruit 69 - 20143, Calligrapha sp. Det. R. E. White; (27) one specimen: Colima, Col. Mex., Conradt, 636; (28) one specimen: Ocotlan, Mex. Tabasco, L. H. Weld, Van Dyke Collection, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (29) one specimen: Jesus Maria, Nayarit, Mexico, vii. 55, Col. Melkin, F. Monrós Collection 1959; (30) one specimen: Chapultepec, Mex., 6. vii. 01, R. H. Hay Collector; (31) one specimen: Morelia, Michoacan, Höge, gift of F. C. Bowditch, gift ex MCZ dupl. series; (32) one specimen: Mexico City, Höge, 383, gift of F. C. Bowditch, gift ex MCZ dupl. series; (33) one specimen: Savula, Jalisco, Mexico, Nov. 23, WM Mann collector; (34) one specimen: Dist. Fedrl. Mex. 7 + 8, 10, 132; (35) one specimen: Oaxaca Mex., May 13, 1938, R. Greenfield; (36) one specimen: Mexico, Calligrapha diversa Stål Dugès; (37) one specimen: Mexico, Durango, 8. x. 65, J. W. Green with Rupa graveolus leaf + stem, El Paso 65277, 65 - 24826, Calligrapha near scalaris Det. R. White; (38) one specimen: vic. Compostela, Nayarit, Mexico, viii. 1934, Coll. of J. R. Fisher; (39) one specimen: Mexico, Progreso, Valle Hidalgo, July 27, 1963, Alfred B. Lau, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (40) one specimen: Mexico, Hidalgo, 16. xi. 1970, R. L. Hodgdon, on cacti, Lot. 71 - 267, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (41) one specimen: on orchid plants, Morelia, Mich., Mexico, xi. 18.58, Heinrich colr., Brownsville Tex., 80228, 10 t 59 - 1270, Zygogramma sp. det. D. M. Weisman, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (42) one specimen: Mexico, Chihuahua, San Rafael-Cuiteco Rd. 17 º 21.111 N 107 º 56.597 W, 8 August 2007, 1870 m, leg. David G. Furth, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (43) one specimen: Mexico, Mexico, Aldama, 20 July 1995, leg. D. Furth & G. Chavarria, Calligrapha diversa Stål J. Gómez-Zurita det. 2011. NRM: (1) two specimens: Mexico, Stål; (2) one specimen: Soledad, Guerrero, 5500 ft. July. H. H. Smith, Calligrapha diversa St. OSAC: (1) one specimen: Mexico, Mich., Santiago, W of Zamora on St. 110, 14. vi. 1967, on Sida carpinifolia, Calligrapha diversa Stål J. Gómez-Zurita det. 2009. OUMNH: (1) three specimens: W. Mexico, Coffin, Named 1899 by M. Jacoby Calligrapha diversa Stål; (2) two specimens: Mex., Named 1899 by M. Jacoby Calligrapha diversa Stål; (3) one specimen: Mex., Named 1898 by M. Jacoby Calligrapha diversa St.; (4) one specimen: W. Mexico Coffin, Named 1898 by M. Jacoby Calligrapha diversa Stål, 204; (5) one specimen: 199, Calligrapha signata Kl., W. Mexico, Coffin, 199. TAMUIC: (1) one specimen: X 0397020, Mexico, Nuevo León, 15 mi W Linares, 1 – 2. vii. 1973, Víctor C. Mastro & Joseph C. Schaffner, at light; (2) two specimens: [X 0534070 and X 0549245], Mexico, Chiapas, Tuxtla Gutierrez, Chorreadero Canyon, 20. vi. 1987, William F. Chamberlain, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (3) one specimen: X 0534310, Mexico, Aguascalientes, 14 mi W Aguascalientes, 10. vii. 1983, Peter Kovarik, Harrison & Joseph C. Schaffner, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (4) one specimen: X 0534616, Mexico, Jalisco, 11 mi N Autlan, 6. vii. 1984, Carroll, Schaffner & Friedlander, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (5) one specimen: X 0535093, Mexico, Guanajuato, 10 mi S San Luis de la Paz, 2. ix. 1962, G. M. Chamberlain, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (6) one specimen: X 0536749, Mexico, Guerrero, 2.1 mi NW Cacahuamilpa, 11. viii. 1978, Plitt & Joseph C. Schaffner; (7) one specimen: X 0538313, Mexico, San Luis de Potosí, 13.4 mi W El Naranjo, 12. vii. 1973, Gaumer & Clark; (8) one specimen: X 0538531, Mexico, Morelos, 5.1 mi E Cuernavaca, 29. vi. 1973, Robert R. Murray & Maria E. Murray; (9) one specimen: X 0541424, Mexico, Queretaro, 7 mi N Queretaro, 13. ix. 1968, Veryl V. Board; (10) one specimen: X 0550339, Mexico, Chiapas, 17 km E La Trinitaria, L. Monte Bella road, 14. viii. 1967, J. Hafernik & Horace R. Burke; (11) one specimen: X 0550617, Mexico, San Luis de Potosí, 10 mi E Ciudad del Maíz, 23. viii. 1964, Horace R. Burke & J. Apperson; (12) one specimen: X 0550735, Mexico, Tamaulipas, 7 km SSE Gomez Farias, Bocatoma, 13. vii. 1982, Robert H. Turnbow; (13) one specimen: X 0551166, Mexico, Nuevo León, Linares, 1. vii. 1973, Víctor C. Mastro & Joseph C. Schaffner, at light. UWIRC: (1) one specimen: Chapingo, Mex., ix- 52, F. González, Calligrapha diversa Stål J. Gómez- Zurita det. 2010. ZSM: (1) fourteen specimens: Vulkan Colima, coll. Joh Laue 1918 [one with: Calligrapha diversa Stål det. Dr. J. Bechyné 1949]; (2) two specimens: Vulkan Colima, coll. Joh Laue, 1.8.1918; (3) four specimens: Vulkan Colima, Mexiko 1918, coll. Joh. Laue, Esperanza 1000 m [one with: Calligrapha diversa Stål Gómez-Zurita det. 2011]; (4) one specimen: Vulkan Colima, Coll. Joh. Laue 20.6.1918; (5) one specimen: Vulkan Colima, Mexiko, 20.6.1918, coll. Joh. Laue; (6) one specimen: Vulkan Colima, coll. Joh. Laue, 25.7.1918; (7) one specimen:?. F. vii. 11, Vulkan Colima, Mexiko, 1918, coll. Joh. Laue; (8) one specimen: Calligrapha diversa Stål Mexique; (9) one specimen: Cuernavaca, Morelos, June, H. H. S., 96386, Calligrapha diversa Stål, Staatssammlung München 1975 Erwerb Coll. Machatschke; (10) one specimen: Mexic, Staatssammlung München 1975 Erwerb Coll. Machatschke; (11) one specimen: Mexico, Taxco 1800 m, 1.6.81, leg. Gfeller; (12) one specimen: Mexico, Guadalajara, Juni 1982, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (13) one specimen: Mexico, Tapalpa, 25 - 5 - 82, leg. Fittkau, Calligrapha diversa Stål J. Gómez-Zurita det. 2011. NICARAGUA IBE-JGZ: (1) one specimen: IBE-JGZ-C 245, Nicaragua, Estelí, P. T. P. Miraflor-Moropotente [49], Julio 2010, J. - M. Maes leg.; (2) one specimen: IBE-JGZ-C 269, Nicaragua, Estelí, P. T. P. Miraflor-Moropotente, Los Cerritos [121], Julio 2010, J. - M. Maes leg.; (3) one specimen: IBE-JGZ- 2168, Nicaragua, Estelí, Reserva Natural Miraflor, El Tunal [177], Agosto 2010, J. - M. Maes leg.; (4) one specimen: IBE-JGZ- 2176, Nicaragua, Estelí, Reserva Natural Miraflor, Miraflor [15], Septiembre 2010, J. - M. Maes leg.; (5) one specimen: IBE-JGZ- 2193, Nicaragua, Estelí, Reserva Natural Miraflor 13 º 09 ' 04 '' N 86 º 19 ' 35 '' W 978 m (no. Herb. 6369), 9 / 09 / 2011, J. - M. Maes leg. NHM: (1) one specimen: Chontales, Nicaragua, T. Belt, Godman-Salvin Coll., Biol. Centr. - Amer. NMNH: (1) one specimen: Nicaragua, Matagalpa, 10 km NW Matagalpa, Selva Negra, 10 – 18. iv. 2005, 1280 m, 12 º 59 ’ N 85 º 54 ’ W, W. Opitz, beating forest edge vegetation; (2) one specimen: Nicaragua, Santa Maria de Ostuma, xi. 1959, N. L. H. Krauss, Calligrapha diversa Stål J. Gómez-Zurita det. 2011. USA FSCA: (1) one specimen: USA, Arizona, Cruz Co., Patagonia, 28. ix. 1968; (2) one specimen: USA, Arizona, Cochise Co., Pt. Huachuca, 17. x. 1968, Dwight Schuh, det. J. Watts ’ 93; (3) four specimens: USA, Arizona, 1 mi N Nogales, Mariposa Canyon, R. L. Jacques [one with: 11. viii. 1971; two with: 13. viii. 1971; one with: 24. viii. 1971]; (4) one specimen: USA, AZ, Huachuca Mts., Copper Canyon W of Montezuma pass, 30. vii. 1987, G. H. Nelson, on Quercus arizonica, Calligrapha diversa Stål J. Gómez-Zurita det. 2011. NMNH: (1) one species: Garden Canon, Huachuca Mts., Ariz., W. H. Mann, WMMann Collection 1954, 6808, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (2) one specimen: Patagonia, Ariz., VII- 9 - 30, J. O. Martin Collector, Calligrapha diversa Stål J. Gómez- Zurita det. 2009; (3) one specimen: Patagonia, Ariz., VIII- 21 - 40, Coll’d by F. W. Nunenmacher, Chicago N. H. Mus. (F. W. Nunenmacher Collection), F. Monrós Collection 1959, Calligrapha diversa Stål J. Gómez-Zurita det. 2009; (4) two specimens: Patagonia, Ariz., E. P. Van Duzee Collector, Calligrapha diversa Stål J. Gómez-Zurita det. 2009 [one with: VIII- 1 ’ 24; one with: VIII- 2 ‘ 24]; (5) one specimen: Patagonia, Sta. Cruz Co., Ariz., Oct 3, 1971, Coll. D. Richman, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (6) one specimen: USA, Arizona, Santa Cruz Co., Sycamore Cyn., Alascoa Mts., 2 August 1991, Lee Guidry, Calligrapha diversa Stål J. Gómez-Zurita det. 2011. VENEZUELA MCZ: (1) one specimen: Venezuela, Calligrapha diversa Stål J. Gómez-Zurita det. 2010. NHM: (1) one specimen: simillima Stål, Chry simillima Stål Caracas, 67 - 56; (2) one specimen, Venezuela [illegible], 67 - 56, Chrysomela diver [...] Stål, Type from C. Stål. NMB: (1) one specimen: Venezuela, Calligrapha simillima Stal J. Bechyné det. 1952; (2) one specimen: Umg. Caracas, Venezuela, i. 54, G. u. Helga Frey, Calligrapha diversa Stål J. Gómez-Zurita det. 2010. NMCZ: (1) one specimen: Caracas, Chrysomela simillima Stål, diversa Dej. Caracas, Calligrapha diversa Dej. Caracas, Coll. Achard Mus. Pragense, Calligrapha diversa Stål J. Gómez-Zurita det. 2009; (2) three specimens: Caracas, Coll. Achard Mus. Pragense; (3) one specimen: Venezuela, Coll. Achard Mus. Pragense. NMNH: (1) one specimen: on Sida sp., Jul 12 ’ 39, G. Vivas B., El Valle D. F. Venez., 11 – 34, C. H. Ballou, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (2) one specimen: on Sida rhombifolia, XI- 30 ’ 38, Ident. 39 10879, Florida, Venez. 1134, S. & C. H. Ballou, Calligrapha diversa Stål J. Gómez-Zurita det. 2009; (3) one specimen: from G. V. Berthier, Caracas, Venez. 1935 # 5, Calligrapha diversa Stål F. Monrós det. 1953; (4) one specimen: Venezuela, Lara, Parque Nac. Yacambu, 6 – 8. iv. 1981, A. S. Menke & L. Hollenbey collectors, Calligrapha diversa Stål J. Gómez-Zurita det. 2011; (5) one specimen: Guaparo, Venezuela, Dec. 20, ’ 38, C. H. Ballou, 1134, on grass. ZSM: (1) one specimen: Caracas, Berg Avila, P. Cor. Vogl., Calligrapha simillima Stål det. Dr. J. Bechyné 1949; (2) one specimen: Venezuela, Hacienda San Eborras, 15.1.28, bei El Negrito, 700 m, südl. d. Marakaibo See's bei San Kasemiro, G. Klaebisch. UNKNOWN SOURCE MfN: (1) one specimen: diversa Stål; (2) nine specimens: [no data]; (3) one specimen: Calligr. sylvia Stål; (4) one specimen: 2. NHM: (1) one specimen: 1267; (2) one specimen: 55 / 24; (3) one specimen: 55 / 24, 284. NMCZ: (1) four specimens: Coll. Achard Mus. Pragense; (2) one specimen: 12 / 955; (3) one specimen: 126, Calligrapha diversa; (4) one specimen: [illegible], simillima Stål, Coll. Achard Mus. Pragense, Calligrapha diversa Stål J. Gómez-Zurita det. 2009. NMNH: (1) two specimens: Gift ex MCZ dupl. series. OUMNH: (1) two specimens: [no data]; (2) two specimens: 15.202, Miers Coll. Presented 1880 by J. W. Miers, Named [one with: 1897; one with: 1899] by M. Jacoby Calligrapha diversa Stål.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6419FF988F9CF9A6FB7BFE4A.taxon	description	Variation. Despite its large geographic range, this species is rather constant in appearance across its distribution, although some features can vary. For instance, the base of the subsutural stripe can be completely fused with the sutural stripe, and actually reaching quite close to the base of elytra basally. This characteristic, which appears in some specimens of C. diversa from Venezuela, but also present in some from Guatemala held at NMCZ or from Chiapas and Guatemala at MCZ, is a defining trait of C. simillima, and in fact these specimens had been ranked with this taxon. These Mesoamerican specimens also have slightly larger markings, filling more elytral space, and their spot enclosed by humeral lunule is larger and more elongated, thus more centred in the available space, contrasting with the backwards position observed in typical C. diversa, and they also show a more reddish tint to their pale elytral areas. Yet, the shape of the specimens is slender, and not as convex as observed in C. simillima, and the base of their arcuate band is divergent from subsutural stripe, again unlike in C. simillima, with a nicely curved apex. Venezuelan specimens are, in turn, just like typical C. diversa but with the mentioned particularity to their subsutural stripes. More rarely, the base of the arcuate band can be confluent with the subsutural stripe and in a single specimens from Teopisca (at MfN), the arcuate band is completely detached from the subsutural stripe, very much as it occurs in C. elegantula, although the remaining characters are clearly these of C. diversa. The apex of the arcuate band can show a variable shape, as a divergent rounded spot to a transversal ovoid spot, and very rarely broken or with irregular profile, as an apical hump. The apex of the humeral lunule generally reaches beyond the basal end of the arcuate band. The spot enclosed by the humeral lunule ranges from small and round (or with a small basal emargination) to large, longitudinally elongated, and it can appear, even if rarely, narrowly confluent with the apical inner concavity of humeral lunule. The spot of apical declivity can be narrowly to broadly confluent laterally with subsutural stripe, and the spot described as “ minute ” in the type can be as big as the others, resulting in a pattern of eight additional spots, roughly 3 - 2 - 2 - 1; these additional discal spots can be very regularly arranged, as in the type, or more often with slightly irregular shape and size, showing variable degrees of confluence among them (and in one specimen the innermost spot is confluent with external concavity of arcuate band). Finally, the black parts of body can have a greenish or bluish metallic sheen.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6413FF9E8F9CFD87FC44FDA5.taxon	materials_examined	Holotype: Type Stål Coll: Deyrolle / Baly Coll. / simillima Stål Venezuela, Caraccas [underneath: Type Stål Col: Deyrolle] (NHM). The specimen lacks six and two apical antennal segments of left and right antennae, respectively. Habitus (Fig. 12 e). Length: 9.79 mm, width: 5.89 mm. Body elongated oval, convex. Head, labrum, mandibles, pronotum and margin of scutellum very dark reddish brown to blackish, with slight bronzy metallic reflection; scutellum, elytral markings, fine margin of elytra, epipleura, ventral surfaces and legs with slightly paler reddish shade and very faint metallic reflection; sides of labrum, antennae and mouth pieces brownish orange; pale areas of elytra creamy yellow. Head large, deeply inserted in pronotum, very finely microreticulate, densely and rather strongly punctured on clypeus, epistome and behind eyes, with scattered punctures on frons; slightly depressed at junction of frontal suture (more impressed anteriorly) with broadly V-shaped clypeal suture; supraocular furrow deeply impressed, running a short distance behind upper eye margin. Antennae relatively short, reaching humeri; first antennomere relatively long and thick, straight at posterior border, convex at anterior border, narrow basally; antennomeres 2 – 5 elongated, thin, relatively smooth and almost glabrous; second antennomere shortest, about half as long as first, third as long as first; antennomeres 3 – 5 shortening progressively, sixth clearly longer than second; segments progressively longer beyond fifth antennomere; seventh and following antennomeres thicker, widening from base to apex; beyond eighth rugose and pubescent; eighth antennomere 0.71 x as wide as long. Labrum short, small, feebly emarginate anteriorly. Mandibles large, robust, surface strongly punctured, with punctures bearing pale yellow long setae; margins subparallel, weakly concave basally. Last maxillary palpomere rather elongated, enlarged apically, weakly convex, sides parallel at apical 1 / 3, curved to narrower basal end; previous palpomere slightly broader apically than base of last segment and considerably narrower basally, with regularly curved exernal margin and shorter, strongly bent interior margin. Pronotum transverse (W / L = 1.83), with sides subparallel in basal half, regularly and gradually converging to moderately protruding anterior angles apically, finely margined; anterior border slightly produced forward medially, convex, finely margined; basal border regularly convex, unmargined; surface microreticulate, with rather densely irregularly scattered moderately strong and deep punctures, becoming stronger and confluent at sides; strong elongated punctures on basal border laterally, confluent with margin. Hypomeral suture deeply impressed, basally curved to form angular margin of hypomera, slightly gradually diverging from lateral border of pronotum towards anterior angle; hypomera very finely microreticulate, unpunctured, with few transversal wrinkles basally. Prosternum relatively long, concave and margined anteriorly, very finely microreticulate, with few moderately strong punctures bearing fine whitish setae; process convex, margined and punctured at sides, gradually slightly widening towards weakly convex apex, slightly surpassing procoxae posteriorly. Mesanepisterna microreticulate, with sparse, moderately strong punctures. Metepisterna microreticulate, with strong, dense, sometimes confluent punctation. Metaventrite finely microsculptured, with dense moderately strong punctures at anterior angles, much finer and sparser on posterior half; transversally furrowed and deeply punctured at metacoxal border. Scutellum small, short (W / L = 0.97), subtriangular with blunt pointed apex; finely microsculptured, unpunctured. Elytra smooth, very finely sparsely punctured with dot-like punctures with dark background; premarginal line of dense, slightly stronger punctures from humerus to nearly sutural apical angle; larger and deeper punctures surrounding and within dark markings; scutellar row present. Markings as follows: (i) sutural stripe broadly surrounding scutellum basally, narrowing gradually preapically before reaching apex of elytra as thin stripe; (ii) subsutural stripe broad, entirely confluent with sutural stripe from near base of elytra; (iii) arcuate band nearly straight, very faintly concave externally and completely confluent laterally with subsutural stripe; apical end enlarged as slightly divergent big rounded spot; stripe emiting dark suffusions laterally at apical half; (iv) humeral spot and humeral lunule completely fused as a large shoulder spot, elongated, with apical 1 / 3 slightly narrower and bent towards suture, ending slightly behind and near base of arcuate band; separated basally from basal margin of elytra; (v) spot enclosed by humeral lunule large, roundish and laterally confluent externally with concavity of humeral lunule; (vi) spot of apical declivity large, broadly elliptic, laterally confluent with subsutural stripe; (vii) apical spot roundish, slightly transverse; (viii) eight additional spots on disc; six forming triangle with base parallel to elytral margin and vertex at and narrowly confluent with weak concavity of arcuate band, two spots in the middle round and smaller, and three spots at marginal line increasingly larger towards apex (last spot large, irregularly shaped, broken in smaller spots); two additional roundish medium sized spots slightly obliquely arranged on apical declivity of elytra. Femora smooth, sparsely punctured, with fine whitish setae. Tibiae slender, subcylindrical, weakly carinated at interior angles, with apex slightly rugose covered by long dense pale yellowish setae; gradually expanded furrow from midlength towards apical emargination of tibiae. First abdominal ventrite strongly margined anteriorly, with dense, strong elongated and deep punctures confluent with margin, strong punctures at sides, and scattered, finer punctures elsewhere; other ventrites with denser smaller punctation basally and at sides; sides slightly raised internally to lateral fovea. Penis figured in Figs 13 a, 13 d.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6413FF9E8F9CFD87FC44FDA5.taxon	distribution	Distribution (Fig. 15, white circles). This species is scarcely present in entomological collections. Most of the specimens I found are from Mexico, but a couple of records can be attributed with certain confidence to Guatemala and Honduras. Records from the USA derive from intercepted fruit from Central America in Texas, or specimens of dubious identification. Considering that the type is labelled as originating from Venezuela, the highly scattered distribution for this species would be highly unusual for the genus. Alternatively, it could be that the species is not frequent, simply rare in entomological collections, or admitedly affected by a problematic taxonomy.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6413FF9E8F9CFD87FC44FDA5.taxon	materials_examined	Material examined (16 specimens). GUATEMALA EGRC: (1) one specimen: Guatemala, Guatemala City, Las Hamacas T. P., 23. vii. 1979, D. B. Thomas & E. P. Case. HONDURAS NMNH: (1) one specimen: on banana debris from Honduras, Apr. 29, 1935, Charleston, # 5530, Gay, Calligrapha simillima Stål J. Gómez-Zurita det. 2011. MEXICO EGRC: (1) one specimen: Mexico, Chiapas, Chorreadero Cyn., 5 mi E Chiapa de Corzo, 15. vi. 1986, D. Thomas, Calligrapha multipustulata Stal det. Daccordi ’ 90, Calligrapha simillima Stål J. Gómez-Zurita det. 2011; (2) one specimen: Mexico, Hidalgo, 4 mi SW Chapulhucan, 25. v. 1979, E. G. Riley, Calligrapha suffriani St. det. Daccordi ’ 81, Calligrapha simillima Stål J. Gómez-Zurita det. 2011; (3) one specimen: Mexico, Yucatán, Chichén- Itzá, 10. viii. 1990, D. A. Rider, Calligrapha simillima Stål J. Gómez-Zurita det. 2011. NMB: (1) three specimens: Chitzen Itza, Yucatan, Mex., G. Frey, x. 58, Calligrapha simillima Stål J. Gómez-Zurita det. 2010. ZSM: (1) one specimen: Mexico, Guadalajara, Juni 1982, leg. Fittkau. NICARAGUA IBE-JGZ: (1) one specimen: IBE-JGZ-C 230, Nicaragua, Matagalpa, Selva Negra, Mayo 2010, J. - M. Maes leg. USA MCZ: (1) one specimen: Tex., F. A. Eddy Collection 5. NHM: (1) one specimen: Baly Coll., Calligrapha geographica? Ch., Texas, Pilate. UNKNOWN SOURCE NMB: (1) one specimen: Encantado, M., 8. x. 50, Racenis l., R: 331. NMNH: (1) one specimen: banana trash from Hond. or Pan., Mar. 2 ’ 36, No. 16262, [male], Calligrapha simillima Stål J. Gómez-Zurita det. 2009; (2) one specimen: F. Monrós Collection 1959, Calligrapha geographica Stal F. Monrós det. 1957, Calligrapha simillima Stål J. Gómez-Zurita det. 2011.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6413FF9E8F9CFD87FC44FDA5.taxon	description	Variation. The additional markings on disc can show different confluence patterns as compared with type, e. g. the interior vertex of triangle of spots not confluent with arcuate band.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6415FFE38F9CF93FFE1CFB89.taxon	distribution	Distribution. The species has been reported from North America, but these US records belong instead to the closely related C. diversa (see above). Instead, C. sylvia is only found in the Atlantic domain of Northern Mexico, east from the Sierra Madre Oriental, but it may reach further south, always on the eastern slopes of the Central American mountain chains, based on the single record of a specimen from Guatemala (Fig. 14). The species is at present considered endemic of the Caribbean Mesoamerican domain.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6415FFE38F9CF93FFE1CFB89.taxon	materials_examined	Material examined (49 specimens). GUATEMALA MCZ: (1) one specimen: Sinanja, Vera Paz, Champion, 1 st Jacoby Coll., Calligrapha nr. sylvia J. Gómez- Zurita det. 2010. MEXICO EGRC: (1) one specimen: Mexico, Hidalgo, 40 mi SW Jacala, 3000 ’ Hwy 85, 24. vii. 1982, C. and L. O’Brien & G. Wibmer, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (2) one specimen: Mexico, Hidalgo, Minera Outland (= Otongo) 3400 ’, 31. vii. 1982, C. W. O’Brien & G. Wibmer, at night, Calligrapha diversa St. det. Daccordi ’ 90, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (3) one specimen: Mexico, Tamaulipas, Bocatoma, 7 km SSE Gomez Farias, 25 – 30. iii. 1978, E. G. Riley, Calligrapha ramulifera Stal det. Daccordi ’ 81, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (4) one specimen: Mexico, Tamaulipas, Bocatoma, 6 mi S Gomez Farias, 19 – 23. v. 1979, Marlin E. Rice Coll., Calligrapha ramulifera Stal det. Daccordi ’ 81, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (5) one specimen: Mexico, Tamaulipas, 6 – 9 km N Rio Guayalejo, 14. x. 1985, R. Turnbow, Calligrapha ramulifera Stal det. Daccordi ’ 81, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (6) one specimen: Mexico, Nuevo Leon, Santa Rosa Canyon, 28 km W Linares, 1. vi. 1987, R. Turnbow, Calligrapha ramulifera Stal det. Daccordi ’ 81, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011. FSCA: (1) one specimen: Mexico, Tamaulipas, Bocatoma w. s., 7 km SSE Gomez Farias, 15. x. 1985, R. Turnbow, Calligrapha ramulifera Stal det. E. G. Riley ’ 88, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (2) one specimen: Mexico, Tamaulipas, 1 – 3 mi E Ocampo, 26. x. 1979, J. E. Wappes, Calligrapha ramulifera Stal det E. G. Riley ’ 82, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (3) one specimen: Mexico, Veracruz, Catemaco-Tuxtlas Biol. Sta. Rd, 15 – 17. vi. 1985, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (4) two specimens: Mexico, Nuevo León, 20 km W Montemorelos, camino a Royones, 21. vi. 1981, B. Miller & L. Stange, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (5) seven specimens: Mexico, San Luis de Potosí, El Salto Falls, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011 [two with: 3. vii. 1968, H. V. Weems, Jr.; one with: 1. vii. 1968, H. V. Weems, Jr.; two with: 1. vii. 1968, D. Weems; two with: 1. vii. 1968, P. Weems]; (6) one specimen: Mexico, Nuevo León, Horsetail Falls, 25. vi. 1965, N. Chernoff. NHM: (1) two specimens: Mex., Baly Coll.; (2) one specimen: Chrysom. consimilis? Chevr., Mexico, Baly Coll.; (3) one specimen: Cosamaloapam, Mexico, Sallé Coll., Calligr. sylvia Stål, Godman-Salvin Coll., Biol. Centr. - Amer. NMB: (1) one specimen: Temascal, Oaxaca, Mexico, x. 1962, leg. Epping. NMCZ: (1) one specimen: Mexique, Coll. Achard Mus. Pragense. NMNH: (1) two specimens: Tuxtepec, Oax., Mex., J. Camelo G., Apr. 1934, No. 969 [one with: Calligrapha sylvia Stål J. Gómez-Zurita det. 2011]; (2) one specimen: Mexico, Oaxaca, Temascal, Oct. 1963, D. H. Janzen, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (3) one specimen: orchids from Tamazunchale, S. L. P. Mex., Laredo Tex. no. 45689, 5 Jan 48,? suboculata Stal HSB 48, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (4) one specimen: Mexico, S. L. P., El Naranjo, vi. 29.1965, Paul J. Spangler, El Salto, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (5) one specimen: Mex. S. L. P., 24.7 mi E Landa de Matamoros Qro. 5000 ’ xi. 18 – 19.65, George E. Ball & D. R. Whitehead collectors, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (6) one specimen: Mexico, Tamazunchale, 8. iii. 65, Sullins & Heinrich with orchid, Bro. 88492, 65 - 9817, Calligrapha sp. Det. R. White, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (7) one specimen: Mexico 22. xii. 66, Coll. L. J. Wheadon, # Brownsville 90526, Lot # 67 - 6715, Calligrapha sp. Det. R. E. White, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (8) one specimen: orchid, Maiz, S. L. P., Mex., Lar. 93693, v. 6.49, 47 - 6851; (9) one specimen: Mexico, San Luis, at Brownsville, 16. iii. 64, Burgess, Heinrich, Dieball on bromeliads 67 - 10164, Calligrapha sp. nr. ramulifera Stal d R. White, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (10) one specimen: Mexico, Nuevo León, Linares, Cañón de Santa Rosa, 19. vii. 952, Coll. G. Halffter, F. Monrós Collection 1959, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (11) one specimen: Tuxtepec, Oax., Mex., J. Camelo G., Nov. 1982, No. 328, Calligrapha ancoralis Stål J. Gómez-Zurita det. 2011. TAMUIC: (1) one specimen: X 0534183, Mexico, Veracruz, Tepetzintla, 28. viii. 1962, William F. Chamberlain, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (2) one specimen: X 0535608, Mexico, Tamaulipas, 9.5 km E San Carlos, Sierra Chiquita de San Carlos, 16. vii. 1988, L. A. Ruedas, at light, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011; (3) one specimen: X 0535717, Mexico, Veracruz, Panuco, 27. viii. 1962, William F. Chamberlain, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011. USA TAMUIC: (1) one specimen 6: X 0535279, USA, Utah, Garfield Co., 13 mi W Long Valley Junction, Dixie National Forest, 16. vii. 1967, Horace R. Burke, Calligrapha sylvia Stål J. Gómez-Zurita det. 2011. UNKNOWN SOURCE MCZ: (1) one specimen: 1 st Jacoby Coll., Calligrapha sylvia Stål J. Gómez-Zurita det. 2010. NHM: (1) one specimen: Ent. Club 44 - 12, Z. Wal [...], N. America; (2) one specimen: sylvia Stål, 67 - 56. NMCZ: (1) one specimen: Coll. Achard Mus. Pragense.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6415FFE38F9CF93FFE1CFB89.taxon	description	Variation. Head and pronotum can be very dark, almost black, with slight metallic green tinge, or alternatively dark red and ventrally pale brown with very feeble metallic shine, as seen in the MfN paralectotype. Punctation on the head can be strong, dense, also on frons and clypeus and mesanepisterna can be punctured close to posterior border. The last maxillary palpomere of some specimens is subtrapezoidal. The spot enclosed by humeral lunule can be missing, or it can be emarginate anteriorly and postero-laterally narrowly confluent with humeral lunule, as seen in the MfN paralectotype. Finally, the small additional markings may also be reduced or absent, or more or less numerous than in the type.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6468FFE08F9CFBF7FA73F8D7.taxon	distribution	Distribution. The species is a Mexican endemic in the Atlantic plains east from the Sierra Madre Oriental, from the states of Nuevo León and Tamaulipas to Oaxaca and Veracruz (Fig. 15, black circles). From a biogeographic perspective, the species is characteristic of the Mexican Transition Zone and the Caribbean Mesoamerican domain.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6468FFE08F9CFBF7FA73F8D7.taxon	materials_examined	Material examined (210 specimens). MEXICO EGRC: (1) one specimen: Mexico, San Luis de Potosí, Tamazunchale, 24 – 25. v. 1979, coll. E. Riley, Calligrapha aeneopicta Stål J. Gómez-Zurita det. 2011; (2) one specimen: Mexico, Hidalgo, 40 mi NW Jacala, 3000 ’, Hwy 85, 24. vii. 1982, C. and L. O’Brien & G. Wibmer; (3) one specimen: Mexico, Hidalgo, 33 mi NE Jacala, 3700 ’, 27. v. 1974, C. W. and L. O’Brien & Marshall; (4) one specimen: Mexico, San Luis de Potosí, Xilitla, 1800 ft, 24. vii. 1954, J. G. Chillcott; (5) one specimen: Mexico, San Luis de Potosí, Hwy 85, 8 mi N Tamazunchale, 700 ft, 24. vii. 1982, C. W. and L. O’Brien & G. Wibmer; (6) one specimen: Mexico, Hidalgo, 4 km N Chapulhuacan 2800 ft, 8. viii. 1967, H. R. Burke & J. Hafernik; (7) one specimen: Mexico, San Luis de Potosí, 11.1 km NE Xilitla, 1500 ft, 3. vi. 1987, R. Turnbow; (8) one specimen: Mexico, Veracruz, 1 mi E Rinconada, 25. x. 1979, J. E. Wappes; (9) one specimen: Mexico, Tamaulipas, Canyon, 5 mi W Cd. Victoria, 3. vi. 1984, D. Thomas, B. Ratcliffe, D. Caldwell, T. Taylor & A. Reifschneider, Calligrapha aeneopicta Stål J. Gómez-Zurita det. 2011. FSCA: (1) one specimen: Mexico, San Luis de Potosí, 1 km S Tamazunchale, 3 – 5. vi. 1987, R. Turnbow, Calligrapha aeneopicta Stål det. E. G. Riley ’ 88; (2) one specimen. Mexico, Veracruz, 3 km SW Puente Nac., 6. vii. 1992, M. C. Thomas, Calligrapha aeneopicta Stål det. J. Watts 1993; (3) one specimen: Mexico, Veracruz, Rinconada, SE of Jalapa, 13. viii. 1963, F. A. Lee; (4) two specimens: Mexico, Veracruz, Rte. 140, SE Jalapa, 1080 m, trop. evergreen trees, in bromeliads, 18. xii. 1978, G. E. & K. E. Ball [one with: Calligrapha aeneopicta Stål det. I. S. Askevold 1990]. MCZ: (1) four specimens: Huachinago, Mexico [one with: Calligrapha aeneopicta Stål]; (2) one specimen: Paso de Telaya, Jicaltepec, V. Cruz, Mexico, 4 / 9,96; (3) four specimens: Jicaltepec, V. Cruz, Mex. 3 - 14 - 96; (4) five specimens: Sn. Rafael, Jicaltepec, V. Cruz, Mex. [one with: March 13, 96; one with: 3 / 9 / 96; three with: March 12, 96]; (5) one specimen: Mexico (Reitter), 225; (6) one specimen: Jalapa, Mexico, Hoege, Callig. aeneopicta; (7) two specimens: Orizaba, Mex., Mann.; (8) two specimens: Mex. [one with: Chrysomela aeneopicta Stål]; (9) one specimen: Jalapa, Mex., O. W. Banalt, 6.18 - 23.98 c, O. W. B. Mex 7.15.1898, F. A. Eddy Collection; (10) four specimens: Jalapa, Mexico, Hoege, 1 st Jacoby Coll.; (11) one specimen: Jalapa, Vera Cruz, Höge, Jacoby 2 nd Coll.; (12) one specimen: Atoyac, Vera Cruz, May, H. H. S., Jacoby 2 nd Coll.; (13) five specimens: Orizaba, H. H. S. & F. D. G., Jacoby 2 nd Coll.; (14) three specimens: San Rafael, Jicaltepec, V. Cruz, Mex. 3 / 12, 96; (15) one specimen: Huachinango, Mexico; (16) one specimen: Calligrapha picra Chev., Mexico, Mex.; (17) one specimen: Mexico, Calligrapha aeneopicta Stål J. Gómez-Zurita det. 2010; (18) one specimen: Mex. ex. Chev. coll.? Calligrapha aeneopicta Stål J. Gómez-Zurita det. 2010. MfN: (1) eight specimens: 29784, aeneopicta Stål, picta Chevr. Dej. amoena, N. A., Mexico, Deppe [seven specimens without data]; (2) two specimens: Jalapa, Mexico, Hoege, 96398 [one with: Callig. aeneopicta]; (3) three specimens: Mexico, Hoege S., Kotze G.; (4) one specimen: aeneopicta Stahl, Mexico; (5) one specimen: var. Mx. H.; (6) one specimen: sp. ign. Mex; (7) one specimen: Manantial, Veracruz; (8) one specimen: Mexico, Jalapa, aeneopicta Stål; (9) one specimen: Mexico, Oaxaca; (10) one specimen: Mexico; (11) one specimen: 792, Jalapa; (12) one specimen: Mexico, Puebla; (13) four specimens: Manantial [one with: 792]. NHM: (1) one specimen: Named by Stål, Baly Coll, aeneopicta Stål Mexico, named by Stål, Col: Deyrolle; (2) three specimens: Mexico, 56 / 55 [two with: 56 / 143]; (3) one specimen: Mexico; (4) one specimen: Mex., Baly Coll.; (5) one specimen: Truqui, Mexico, Fry Coll. 1905.100; (6) one specimen: Mexico, Fry Coll. 1905.100, Calligrapha aeneopicta Stål Mexico; (7) three specimens: Mexico, Fry Coll. 1905.100 [one with: 23839]; (8) one specimen: Orizaba, H. H. S. & F. D. G. Dec. 1857, Godman-Salvin Coll., Biol. Centr. - Amer.; (9) one specimen: Cordova, Mexico, Sallé Coll., Calligrapha aeneopicta Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer. [one with: Sp. figured]; (10) two specimens: Toxpam, Mexico, Sallé Coll., 636, Calligrapha aeneopicta Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (11) one specimen: Orizaba, Mexico, Sallé Coll., Calligrapha aeneopicta Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (12) five specimens: Jalapa, Mexico, Hoege, Godman-Salvin Coll., Biol. Centr. - Amer.; (13) one specimen: Ex Coll. J. Sturm, Mexico, Guyan. Calligrapha suturalis Mihi, Mexico Sallé Coll., Calligrapha aeneopicta Stål apud Sallé, Godman-Salvin Coll., Biol. Centr. - Amer.; (14) one specimen: 150, Vera Cruz, Mexico, Koebele, R. C. L. Perkins, B. M. 1942 - 95; (15) one specimen: Mexico, Pascoe Coll. 93 - 60, Calligrapha aeneopicta. NMB: (1) one specimen: Orizaba, Mex., Dr. A. Fenyes, Erwerb 1955, Coll. Brancsik; (2) one specimen: Mexico 1888, Calligrapha aeneopicta Stål J. Bechyné det. 1951; (3) four specimens: Mexiko. NMCZ: (1) one specimen: Calligrapha aeneopicta Stål Mexique, Coll. Achard Mus. Pragense; (2) two specimens: Mexique, Coll. Achard Mus. Pragense [one with: Calligrapha aeneopicta]; (3) one specimen: icterica Chevr. Mexiq., Coll. Achard Mus. Pragense; (4) two specimens: Mex., Coll. Achard Mus. Pragense; (5) four specimens: Mexique, Mexico, Coll. Achard Mus. Pragense; (6) one specimen: aeneopicta Mexiq., Coll. Achard Mus. Pragense; (7) one specimen: Mexico, Coll. Achard Mus. Pragense; (8) three specimens: Mexico, Mus. Pragense Col. Kambersky [one with: Call. aeneopicta, Calligrapha aeneopicta St. J. Achard det.]. NMNH: (1) two specimens: Mexico, F. Monrós Collection 1959 [one with: Calligrapha aeneopicta Stål F. Monrós det. 1954; (2) two specimens: Mexico, Veracruz, 17. vi. 1897, R. Hañover, F. Monrós Collection 1959; (3) six specimens: San Rafael, Jicaltepec, Vera Cruz, Mex., 25 June, Townsend [one with: Calligrapha aeneopicta Stål F. Monrós det. 1953]; (4) one specimen: Nogales V. C., Mex. 16.6.0 5, Fredk Knab collector; (5) one specimen: Cordoba, V. C., Mex., Mch. 1.07, Fredk Knab collector; (6) one specimen: Cordoba, Mex., Dec. 20. ’ 07, Fredk Knab Collector; (7) one specimen: Mexico DF, J. R. Inda Collector; (8) one specimen: on leaf banana from Mex., 17.12.1934, New Orleans # 10610, Calligrapha aeneopicta Stål F. Monrós det. 1953; (9) one specimen: Jalapa, Mexico, Hoege; (10) two specimens: Almo, iii; (11) one specimen: Orizaba, V. C., Mex., Jan. 9 – 16, ’ 92, H. Osborn Collector; (12) one specimen: Mex., Coll. M. L. Linell; (13) one specimen: Ogumba, Mexico, alt. 8.088 ft, Collection Wm Schaus; (14) one specimen: Mexico, Veracruz, Orizaba, 1 June 1981, W. E. Steiner, Calligrapha aeneopicta (Stål) Det. C. L. Staines 2004; (15) one specimen: Mexico, Ver., Huatusco, vii. 25.1964, A. B. Lau; (16) one specimen: Ojo de Agua, Sabinas, Hidalgo, Nuevo León, Mexico, Alt. 1000 ft., June 14, 1940, Hoogstraal and Knight, F. Monrós Collection 1959; (17) one specimen: Dos Amates, Veracruz, Mexico, P. Hubbell, 5.1975, Calligrapha aeneopicta Stål J. Gómez-Zurita det. 2011; (18) one specimen: Jalapa, Mex., Wickham Collection 1933; (19) two specimens: Jalapa, mex., vi. 1, Smith, Wickham Collection 1933 [one with: Calligrapha aeneopicta Stål J. Gómez-Zurita 2011]; (20) one specimen: Mexico, Veracruz, 11.9 mi E Jalapa, 2600 ’, Rte. 140, iv. 9.1966, in bromeliads, George E. Ball & D. R. Whitehead collectors, Calligrapha aeneopicta Stål J. Gómez- Zurita det. 2011; (21) one specimen: Mexico, Veracruz, 22.0 mi E Jalapa, Rte. 140, 1000 ’, iii. 9.66, in bromeliads, George E. Ball & D. R. Whitehead collectors; (22) one specimen: Mexico, Veracruz, 4.7 mi E Jalapa, 3600 ’, Rte. 140, iv. 9.1966, in bromeliads, George E. Ball & D. R. Whitehead collectors; (23) one specimen: Mexico, Veracruz, 6.2 mi NW Jalapa, 5600 ’, Rte. 140, iv. 8.1966, in bromeliads, George E. Ball & D. R. Whitehead collectors. NRM: (1) one specimen: Mexico, Tarnier; (2) two specimens: Mexico, Sallé; (3) one specimen: Mexico, Stål; (4) one specimen: Mexico, aeneopicta Stål. TAMUIC: (1) one specimen: X 0532849, Mexico, Veracruz, Paraje Nuevo, vi. 1942, Calligrapha aeneopicta Stål J. Gómez-Zurita det. 2011; (2) two specimens: [X 0538610 and X 0548062], Mexico, Hidalgo, 4 km N Chapulhuacan, 8. viii. 1967, Horace R. Burke & J. Hafernik; (3) two specimens: [X 0547547 and X 0549309], Mexico, Veracruz, 4 mi E Guadalupe, 31. viii. 1962, William F. Chamberlain, Calligrapha aeneopicta Stål J. Gómez-Zurita det. 2011. ZSM: (1) one specimen: Mexico, Hoege S. & Kotze G., Calligrapha aeneopicta Stål, Staatssammlung München 1975 Erwerb Coll. Machatschke; (2) one specimen: Mexico, Oaxaca, Calligrapha aeneopicta Stål, Staatssammlung München 1975 Erwerb Coll. Machatschke. UNKNOWN SOURCE MCZ: (1) two specimens: F. C. Bowditch Collection [one with: 924]; (2) one specimen: Calligrapha aeneopicta Stål J. Gómez-Zurita det. 2010; (3) one specimen: 1 st Jacoby Coll., Calligrapha aeneopicta Stål J. Gómez-Zurita det. 2010. MfN: (1) one specimen: Calligrapha aeneopicta Stål; (2) nine specimens: [no data]. NHM: (1) one specimen: 61, Baly Coll.; (2) two specimens: Baly Coll.; (3) three specimens: 67 / 56; (4) one specimen: C. picta Duf., Baly Coll.; (5) two specimens: 4852 [one with: C. picta Duf.]; (6) one specimen: picta, 67 - 56; (7) one specimen: E. Coll. Laferté, 67 - 56; (8) one specimen: E. Coll. Thomson 67 - 56; (9) one specimen: E. Coll. Thomson, 67 - 56. NMCZ: (1) five specimens: Coll. Achard Mus. Pragense; (2) one specimen: 125; (3) one specimen: N. Granada, Coll. Nickerl Mus. Pragense. NMNH: (1) one specimen: Gift Ex MCZ Dupl. Series; (2) one specimen: Gift of F. C. Bowditch, Calligrapha aeneopicta Stål F. Monrós det. 1953. OUMNH: (1) one specimen: Named 1898 by M. Jacoby, Calligrapha aeneopicta Stål; (2) one specimen: Named 1899 by M. Jacoby, Calligrapha aeneopicta Stål; (3) one specimen: 45, Calligrapha picta. ZSM: (1) one specimen: Coll. Haag, Polyspila aeneopicta Stal, Staatssammlung München 1975 Erwerb Coll. Machatschke.	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
039E355D6468FFE08F9CFBF7FA73F8D7.taxon	description	Variation. This species is quite constant in its shape and elytral simplified patterns, but interestingly, some individuals display minute spots where other species show the typical Calligrapha markings. For instance, some specimens present a very small spot enclosed by humeral lunule near the apex of the latter marking, or more rarely they can show faint traces of the discal spot on external concavity of arcuate band. However, very often, they do not have any trace of spots on the elytra, or have two little markings instead of one as in the type. Thus, the small spot of the lateral declivity of elytra is often lacking and very rarely there may be additional minute spots or traces of them (ground colour becoming paler) on disc. Rarely, some specimens may lack spot on apical declivity of elytra or have it minute and detached from subsutural stripe, but it can be also relatively large and broadly fused laterally to subsutural stripe, small and also fused or, more rarely, free (in three of the specimens I have seen, it is free on one elytron and fused on the other; and in a couple of specimens it is missing altogether in just one elytron).	en	Gómez-Zurita, Jesús (2015): Systematic revision of the genus Calligrapha Chevrolat (Coleoptera: Chrysomelidae, Chrysomelinae) in Central America: The group of Calligrapha argus Stål. Zootaxa 3922 (1): 1-71, DOI: 10.11646/zootaxa.3922.1.1
