identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039F8D3EFFBEFFE2969647DFFAEDF9E6.text	039F8D3EFFBEFFE2969647DFFAEDF9E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon	<div><p>Nothocyphon, new genus</p><p>Type species: Nothocyphon lindensis (Blackburn, 1892), new combination</p><p>Diagnosis. Small (BL 1.5−3.5mm) oval to elongate-oval relatively flat marsh beetles. The transverse head rests between the slightly projecting blunt angles of the pronotum and is in dorsal view concealed by the front margin of pronotum. Eyes projecting. Mandibles short, with one small tooth or without tooth. Terminal segment of maxillary palpus bottle-shaped, of labial palpus bean-shaped, inserted at an angle on the penultimate segment. Antennae filiform, base not modified. No antennal sulcus. Pronotum with wide base, sides curving down, rounded, converging towards front corners. Unmodified ambulatory legs with paired keels on tibiae. Scutellum an equilateral triangle. Elytra unmodified. Dorsal side of body irregularly punctate, finely pubescent.</p><p>Underside. The subgenal crest meets the gular suture and then the border of head capsule next to the mouthparts, there is no buttonhole configuration. Prosternal process slender, teardrop-shaped. Receiving mesoventral groove elongate, U- or V-shaped, anteriorly open. Mesoventral process parallel, the finely incised end meets the short metaventral process. Discrimen visible over most of metaventrite. Abdominal segments 3−7 exposed, unmodified, except female N. lanceolatus n. sp.</p><p>Male: T8 well developed, S8 large, Y-shaped. T9 with straight apodemes, plate membranous. S9 soft, base rarely visible, caudally setose, mostly bilobed. Tegmen with parameres, penis with parameroids; shapes variable but no essential modifications.</p><p>Female: Ovipositor long, unmodified. Prehensor present. Bursella without major sclerites or ornaments.</p><p>Description. The body contour at rest in dorsal view is oval to elongate oval, with the wide head concealed under the slightly projecting pronotal front margin (e.g., Figs. 62, 165−168). In side view the body is relatively flat to slightly domed (Figs. 172, 173). Head across eyes wider than long from front of frontoclypeus to edge of occiput (Fig. 1). Eyes bulging. The frontoclypeus projects little between the antennae, the small labrum is rectangular, with rounded corners. The mandibles are largely concealed, the space between their condyli corresponds to about 40% of HCW. A single or no tooth on the mandibles; if present, the right tooth tends to be larger (Figs. 1, 2, 5). Mola without spinous area. The terminal segment of the maxillary palpus is bottle-shaped, narrowed in its distal half (Fig. 6). The bean-shaped terminal labial palpus segment inserts subterminally on the penultimate segment (Figs. 3, 7). The angle between the two segments seems to increase with the degree of dehydration, for example during preparation for mounting. Antenna (Figs. 4, 25): scape large, unmodified. Pedicel thinner and shorter, first flagellar segment slender, length not much different from pedicel.</p><p>The head sits between the front angles of the pronotum which are blunt but marked. Pronotum at base little narrower than the elytra (Figs. 165−171). It is much longer medially than along the downcurved sides. The rear edge is sinuous, the rear angles are obtuse, rounded. In side view, the pronotum is little arched along the median line, more strongly across its anterior part. The scutellum is an equilateral triangle between the elytral bases. Humeri distinct, the elytra are gently arched, without carinae or sutural stripe. Unmodified ambulatory legs, tibiae on outside with paired carinae. Fully winged beetles.</p><p>The dorsal body surface is covered with fine semi-erect pubescence and is irregularly punctate. Punctures on head, pronotum and scutellum are granular, raised, those on elytra are normal.</p><p>Most species are of uniform brown colour (e.g., Fig. 171) but several have patterned pronota and elytra (e.g., Figs. 62, 63, 165−167) which is often not very apparent. Distinctness of patterns seems to vary with the degree of maturity of the specimen. Further, pigment on the elytra is often difficult to distinguish from dark shades caused by the folded wings which are seen by partial transparency. Pigment patterns alone do not permit species identifications.</p><p>Lower face. The front end of the subgenal ridge joins a branch from the gular suture. Together they enter the head capsule border near the mandibular insertion (Fig. 1). No buttonhole configuration. A very shallow ridge obliquely crosses the area between eye and mouthparts. The pronotal process is narrow, the tip resembling a teardrop bears fine hairs. The U- or bluntly V-shaped receiving groove of the mesoventrite is longer than it is wide in front, bordered by shallow ridges only along the sides, open in front. Mesoventral process parallel, its end finely incised, meeting the short metaventral process. Discrimen visible over most of metaventrite. Elytral epipleura narrow, smooth. Abdominal sternites 3−7 exposed, unmodified, except the usual row of microscopic piliferous granules along front edge of anterior sternites.</p><p>Males. T8: Apodemes straight, about as long as the plate, basally connected by a thin curved or angular sclerite along the antecosta (e.g., Figs. 21, 34). Caudal edge with marginal setae and a microtrichial pecten (Zwick 2013c). Distal half of plate covered with microtrichia; a band in front of the caudal edge remains usually bare. The area covered with microtrichia often has a tongue-shaped forward extension between which and the apodemes is often a paler, window-like area (e.g., Fig. 143). S8 is distinct, Y-shaped (e.g., Figs. 35, 57, 70). T9: Apodemes about as long as those of T8, antecosta between them weakly sclerotized. Plate soft, colourless and bare (e.g., Figs. 22, 58). Thin sclerite rods resembling continuations of the apodemes may occur along the sides of the membranous plate. S9: The setose mostly bilobed caudal portion is often all that is seen, the plate is soft, usually there is no well defined base (e.g., Figs. 17, 66, 71). Penis with trigonium and parameroids, tegmen and parameres specifically distinct.</p><p>Females. Known from only a few species. Ovipositor (e.g., Figs. 15, 39) slender, bacula very long, with subterminal recurrent branchlet. Gonocoxae long and slender, caudally very narrow, gonostyli small, tubular, with apical sensilla. The apodemes of T8 are shorter than the bacula but much longer than the sclerotized lateral rods of S8 which may anteriorly be connected in various ways. T8 with continuous fringe of delicate microtrichia along caudal edge. S8 is similar but caudal microtrichial fringe interrupted. Prehensor specific (e.g., Figs. 14, 16, 28, 31, 40), bursella with small pores (Libonatti &amp; Ruta 2013), no large sclerites on surface.</p><p>Note. In order to be able to name the new species assignment to a genus is needed. The beetles redescribed or described in the present study share a generalized body structure and lack apomorphic character expressions seen in other genera. Some support for Nothocyphon can be derived from a recent molecular study of Australian marsh beetles (Cooper et al. 2014).</p><p>Cooper et al. (2014) recognized an Austrocyphon -group, and three clades within it. One clade included the Austrocyphon species which grouped closely (Cooper et al. 2014, their fig. 4).</p><p>The sister clade to Austrocyphon contained still undescribed species which were called Contacyphon spp., mostly of the lindensis- group. C. H. S. Watts has now identified several, from a draft of the present manuscript (Watts, personal communication). Specimens of N. lanceoloatus n. sp. form one cluster grouping with N. vandiemeni n. sp., N. multidentatus n. sp., and N. triangulum n. sp. Together they are sister group of N. ypsilon n. sp. This pattern generally agrees with the taxonomy proposed here.</p><p>However, Nothocyphon frater and its siblings formed the sister clade to the other two (Cooper et al. 2014), as though it also deserved genus rank. Presently, I cannot take this step because I find nothing in body structure to justify generic separation of the frater -group from the rest.</p><p>The numerous species of Nothocyphon can reliably be distinguished by the male genitalia which, however, do not fundamentally differ from the Microcara - type (Nyholm 1972). Similarities in the genitalia permit the provisional distinction of some groups but differences between groups are not striking, elevation to genus group rank is not appropriate. Numerous species stand by themselves, their closest relatives are unknown. Females are in most cases unknown, female genital character sets are only fragmentarily available.</p><p>Keys to males are provided. For females, users are referred to the figures of the few known ones. In view of the overall similarity between the included species, the following descriptions are essentially brief diagnoses based on terminalia. Where possible, male segments 8 and 9 are illustrated, but often without repetitive descriptive text.</p></div>	https://treatment.plazi.org/id/039F8D3EFFBEFFE2969647DFFAEDF9E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFB8FFE3969643F9FCABF9B0.text	039F8D3EFFB8FFE3969643F9FCABF9B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon lindensis	<div><p>The lindensis -group</p><p>Members of this group share a ventral armature between the penis and the bilobed S9, either a large hollow spinelike cone or an armature composed of patches of slender spines. On the everted genitalia, these structures stand perpendicular to the penis, tip (or tips, respectively) pointing cephalad.</p><p>In the known females the front ends of the rods of S8 are connected by a wide flat ring (e.g., Fig. 15). The complex prehensor is composed of large sclerotized areas, often including pocket- or sleeve-like structures, and on the opposite side of the oviduct of a sclerite loop armed with spines (e.g., Figs. 15, 16). There may also be teeth and spines.</p><p>The larvae of N. lindensis and its close relatives differ from similar small taxa by a long narrow extension of the last tergite and form a group of their own (Watts 2014).</p></div>	https://treatment.plazi.org/id/039F8D3EFFB8FFE3969643F9FCABF9B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFB8FFE0969641CFFA1AFEE8.text	039F8D3EFFB8FFE0969641CFFA1AFEE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon lindensis	<div><p>Key to males of the lindensis -group</p><p>1 A large hollow cone-like spine between penis and S9 (e.g., Figs. 12, 19). Yellow or light brown beetles with a brown spot close to the suture near the top of elytra (Fig. 165), rarely more extended maculae (Fig. 166)............................... 2</p><p>1' Groups of slender spines between penis and S9 (Figs. 26, 29). Brown beetles...................................... 5</p><p>2 Parameres enveloping the penis, one or two huge curved spines along their medioventral edge (Figs. 10, 11)............. 3</p><p>2' Parameres slender, lying alongside penis, not surrounding it.................................................... 4</p><p>3 Apex of trigonium entire, with spines in irregular dorsoventral rows (Figs. 8−10)................ N. lindensis (Blackburn)</p><p>3' Trigonium forked, the hollow spine visible between the short branches (Figs. 12, 13).................... N. ypsilon n. sp.</p><p>4 Trigonium truncate, short, with curved lateral horns (Fig. 18)....................................... N. taurus n. sp.</p><p>4' Trigonium deeply excised, each lobe medially with many spinules (Fig. 24)......................... N. banksiae n. sp.</p><p>5 Notch in trigonium deeper than wide. Each lobe of S9 with a rounded notch. A single triangular patch of spines, parameres with strong teeth (Fig. 26).............................................................. N. donnabuangi n. sp.</p><p>5' Notch of trigonium shallower, wider than deep. Caudal lobes of S9 not notched. Paired spinule patches, parameres with fine teeth (Fig. 29)........................................................................... N. imitator n. sp.</p></div>	https://treatment.plazi.org/id/039F8D3EFFB8FFE0969641CFFA1AFEE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFBBFFEF96964670FAC5FB2B.text	039F8D3EFFBBFFEF96964670FAC5FB2B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon lindensis (Blackburn 1892) Blackburn 1892	<div><p>Nothocyphon lindensis (Blackburn, 1892), new combination</p><p>(Figs. 1−3, 8−10, 14, 165)</p><p>Helodes ( Cyphon ?) lindensis Blackburn, 1892: 522 . Helodes lindensis— Klausnitzer, 1981: 118, figs. 1–6.</p><p>Type material examined: To stabilize the use of the name a lectotype is here designated: 1♂: Helodes lindensis, Blackb. Co-type \ Port Lincoln Blackburn \ 433 [red, on specimen card itself; also a red line across base of card] \ Lectotype design. P.Zwick 2014 \ Nothocyphon lindensis (Blackburn) ♂, det. P. Zwick 2014 (SAMA). Paralectotypes here designated: 1♀: Cyphon lindensis Bl. S. Australia \ Paralectotype design. P.Zwick 2014 \ Nothocyphon lindensis (Blackburn) ♀, det. P. Zwick 2014 (SAMA). 1♂, paralectotype here designated from Klausnitzer's (1981) illustrations (ICZN, Article 74.4); the actual specimen in the Natural History Museum, London, was not examined during the present study.</p><p>Additional material examined: SA: 1♀: lindensis, Blackb. \ Quorn, S.A. Blackb's Coll. \ C 192 [circular label] \ Quorn [red, on specimen card itself] (SAMA; the specimen was received as a syntype of H. lindensis, as one of 7 specimens originally mounted on 4 cards on the same pin. Since Quorn is not mentioned in the original description the specimens are not eligible as syntypes of H. lindensis . The other six specimens are actually Austrocyphon robustus Zwick). 5♂: Mt. Lofty Rgs, S.H.Curnow; 1♂: Adelaide, J.G.O.Tepper; 1♂, 1♀: Orroroo [32.44S 138.36E] S. Australia; 2♀: Orroroo S. Australia \ [illegible] seem to be unusually longe [sic!] lindensis Bl.; 1♂, 5♀: Parachilna Flinders Range Hale; 1♂: Pt. Lincoln, S.A. Lea; 10♀: Adelaide C.Watts 28-9-58; 1♀: 6k S.Willunga S.A. 9.12.96 C.Watts; 1♂, 1♀: SW Willow Springs Stn 27/4/12 CHS.Watts under bark nr creek \ Watts det. lindensis (all SAMA). ♀ torso: Mt. Lofty Rgs. A.H.Elston \ A.H.Elston collection Cyphon \ 1757 Id. by A.M.Lea \ K249511; 1♂: Quorn S. Australia A.H.Elston \ A.H.Elston collection \ Cyphon 1171 fenestratus Blckb. \ K249506; 1♂: Quorn S. Australia A.H.Elston \ A.H.Elston collection \ Cyphon fenestratus Bl. Id. by A.M.Lea \ K249508 (all AMS). 1♂: 31.47S, 138.21E Nooltana Creek 13km NW by N of Hawker, S.A. 16. Sept. 1978 M.S.Upton; 1♂: 31.45S 138.21E Nooltana Ck 13km NW by N of Hawker, SA, 16 Sep. 1978 DCF.Rentz; 2♂, 2♀: Angorichina Hostel 7km E of Parachilna 23.Oct.1978 E.B.Britton (all ANIC). 2♂, 2♀: Mt Remarkable S. Austr. Oct. 1925 F.E.Wilson \ F.E.Wilson Coll. (MV). ─ VIC: 1♂: Lilydale distr. Victoria J.E.Dixon \ Cyphon lindensis Blackb. Id. by J.Armstrong; 1♂, 1♀: Lilydale dist. Victoria J.E.Dixon \ Cyphon lindensis Blackb. Id. by HJCarter; 1♀: Western District Victoria Nat. Mus. Victoria C.French's Coll. 6.1.0 8 Cyphon lindensis Blackb. Id. By HJCarter (all MV). ─ ACT: 1♀: 35.16S 149.06E, Black Mtn ACT 600m Dec. 1986 Weir, Lawrence, Dressler \ flight interception—window trap; 1♀: 36.16S 149.06E ACT Black Mtn light trap 4.Mar.1991 W.Dressler (all ANIC). ─ NSW: 1♂: 34.44S, 150.32E, Kangaroo Valley, NSW, N.Slope 22.1.71, S.Misko &amp; K.Pullen; 1♀: Acacia Flat NSW J. Armstrong \ in copula; 1♂: Acacia Plateau N.S. Wales H.Davidson; 1♂: 1km W Barrengarry [34.72°S, 150.52E] 27.Sept.1986 C.Reid on Ac. melanoxylon flwr; 5♀: S.E. Border NSW-Qld H.Davidson (all ANIC). 1♂, 1♀: AUSTRALIA, N.S.W. Mt. Kaputar, Bullawa Creek 29.I.1985 G.Hangay; 1♀: AUSTRALIA, N.S.W., Bigga 24– 25.XI.2000 S 34°05' E 149°09' leg. A.Podlussány; 2♀: AUSTRALIA, N.S.W. Casilis, „Kuuloo Station“ 710m \ 31°50'S 150°08'E 24–25.X.2000 leg. A.Podlussány; 1♀: AUSTRALIA, NSW Dorrigo N.P., rain forest 770m, 3– 6.I.2006 \ leg. G.Hangay, I.Rozner, R. de Keyzer and A.Podlussány (all HNHM). 1♀: Australia, NSW O'Sullivan's Gap 22 September 1991 Vince Lorimer \ in leaf litter by creek \ VWHL-108 32°20'S, 152°15'E \ K249526; 1♂: Sydney N.S. Wales X.31 Dr. K.K.Spence \ K.K.Spence collection \ K249515 (all AMS). 1♂: 2.5km W Forest Reefs NSW 4 April 1993 C.Reid coll. on eucalypts (ANIC). 3♂, 7♀: Cyphon N.S. Wales 8788 Forest Reefs N.S.W. Lea; 3♂, 9♀: Cyphon N.S. Wales 8788 Sydney Lea (all SAMA). 1♂: TI 10C 19.08.2013 Binni Creek Site C, 33.69937S, 148.77011E Lachlan wetlands, J.Miller (PZ). ─ NT: 2♂, 1♀: Stanley Chasm NT 9\10\68 C.Watts (SAMA). ─ QLD: 1♀: 24 mi. SW. by W. of Kingaroy, Q., 26.44S 151.31E, 21.xi.1968, Britton &amp; Misko; 1♂: same, but at light; 1♀: Australia, QLD, Eungella 27 November 1988 Richard Bejsak \ Attracted to black light (all ANIC). 1♀: Australia: QLD., 8.3.1997 Lamington N.P., O'Reilly's, seapage near Elabama Falls 28°17'S, 153°10'E, P.Zwick (NHMW). S.E.Q., Boggomoss, via Taroom (all QMSB): 25°27'Sx 150°02'E: 1♀ No.8, 12 Nov 1996 G.B.Monteith at light 0 40 \ QM Reg.No. T37689; 1♂: 30, 12 Nov 96 – Jan.1997 Cook &amp; Monteith Baited Flight Intercept 0 35 QM Reg.No. T37688; 25°31'Sx 150°03'E: 1♀, 13 Nov 96 — Jan 1997 Cook &amp; Monteith Baited Flight Intercept 0 63 \ QM Reg.No. T37692; 25°26'Sx 150°08'E, No. 30: 1♂: 14 Nov. 1996 G.B.Monteith At Light 0 62 \ QM Reg.No. T37691; 1♂: 14 Nov. 1996 – Jan. 1997 Cook &amp; Monteith Baited Flight Intercept 0 57 \ QM Reg.No. T37690.</p><p>Habitus. BL 2.3−3.6mm, BL/BW ~1.6. Body elongate oval. Surface punctures are very fine everywhere, finest on head. The pilosity is yellowish. Most specimens are yellowish brown, with a small brown spot at the highest point of the elytra, next to the suture, a little behind midlength (Fig. 165). Sometimes, specimens are darker with brown head and pronotum and brown maculae on the elytra reaching obliquely forward to the elytral margin, and back along the side. All marks are diffusely delimited. Appendages yellowish to light brown, antenna distally infuscate.</p><p>Male (Figs. 8−10). Segments 8 and 9, as described for the genus. On a transparent membrane between S9 and the penis stands a slender backward pointing spine-like hollow cone which points basad on the everted penis (Figs. 8, 10). Pala large, front rounded. The penis narrows caudally, a continuous sclerite extends from the sides of the pala to the apices of the parameroids. A slight medial fold or crest suggests there is a delicate cover over the entire pala. The basally wide bulbous trigonium narrows caudally. Its apex is expanded in dorsoventral direction and carries on each side a row of strong teeth. In precisely dorsoventral view the two rows are separate, in oblique or in side view the apex resembles a rooster with a crest. The trigonium is much shorter than the rod-like parameroids. Their expanded apex is beset with sensillar pores, some colourless membrane attaches to the outside. The tegmen is a large ventrally split colourless sleeve surrounding the penis. On each side it supports one very large spine (Figs. 8, 10), homology with parameres uncertain. At high magnifications the spine appears longitudinally twisted.</p><p>Female. Front ends of rods of S8 connected by a sclerotized ring (compare Fig. 15, N. ypsilon, which is similar). T8 and ovipositor unmodified. Prehensor (Fig. 14) composed of a loop-shaped narrow sclerite armed with some spinules and anterolaterally from it two bands of large tooth-like scales on one side of the oviduct, and on the opposite side an anteriorly bilobed sclerite which bears two elongate rough areas with irregular polygonal structure (Fig. 14 a). Medially a sclerotized nipple projects into the lumen of the duct, standing vertical on its wall (Fig. 14 b).</p><p>Notes. Klausnitzer (1981) realized that H. lindensis does not belong in genus Elodes but provisionally accepted the original combination. He mentioned faint elytral ribs which I have not seen. The species is externally indistinguishable from N. ypsilon n. sp. Blackburn, Lea, and other students did not study the genitalia and failed to distinguish the taxa whose ranges overlap.</p><p>N. lindensis inhabits an exceptionally wide range on the mainland but is not known from Tasmania. Larvae of N. lindensis live in “still water in dams or side pools of small creeks or in trapped leaf masses” (Watts 2014).</p><p>Nyholm (1969) documented how during copula of European Contacyphon spp. the prehensor (Latin: the grasper) is extruded into the male gonoduct and seizes the spermatophore. Among material from coll. Armstrong a pair taken in copula was glued on one card, but only the female remains. Its prehensor protrudes between the components of the ovipositor in a way similar to what Nyholm described. It may function in a similar way.</p></div>	https://treatment.plazi.org/id/039F8D3EFFBBFFEF96964670FAC5FB2B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFB4FFEC96964236FB14F9E6.text	039F8D3EFFB4FFEC96964236FB14F9E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon ypsilon	<div><p>Nothocyphon ypsilon, n. sp.</p><p>(Figs. 11−13, 15, 16)</p><p>Type material: holotype ♂, 1♂ paratype: 35.16S 149.06E, Black Mtn ACT 600m Dec. 1986 Weir, Lawrence, Dressler \ flight interception—window trap (ANIC).</p><p>Additional paratypes: ACT: 9♂: 35.16S 149.06E, Black Mtn ACT 600m Dec. 1987 M.E.Irwin ex Malaise trap; 1♂: Black Mtn, light trap, ACT, 29.1. 68 M.S.Upton; 1♂: 35.14S 149.11 ACT Mt. Majura Oct. C.Reid beating dead Eucalypt leaves.— NSW: 1♂: 30.30S 152.23E NSW Thungutti Camp New England N.P. 16–18.Nov.1990 T.A.Weir at light; 1♂: 31.52D 152.22 E, Wingham Scrub N.S.W. 3.i. 70 in UV light trap, Brittain, Holloway, Misko; 1♂: 34.43S, 150.11E, Surf Beach, NSW (Bateman's Bay) light trap, 2.iii.72, S.Misko; 1♂: Bruxner Park, Coffs Harbour, N.S.W. ii. 65 M.S.Upton; 1♂: Minnamurra Falls NSW 34.38S, 156.43E, 26.ii.1969, Brittain &amp; Misko; 1♂: Upper Allyn River NSW, 14.Feb.1968, D.H.Colless (all ANIC). 1♂, 1♀: nr Yowaka River NSW, 59 05208N 557 42731E J &amp; R Miller 15.6.69; 1♂: NSW Brindale Creek campground. At light Border Rngs. Nt. Pk. 22/19/09 [sic!] CHS.Watts \ DNA VOUCHER CW Sc #241; 1♀: NSW 5 km E Dorrigo At Light 20/10/09 CHS.Watts \ DNA VOUCHER CW Sc #248 (all SAMA). 1♂: Williams R., Salisbury NSW 29.v.63 G. Monteith \ UQIC Reg. #88679 (QMSB). 1♀: Australia, NSW O'Sullivan's Gap 22 September 1991 Vince Lorimer m\ in leaf litter by creek VWHL-108 \ 32°20'S, 152°15'E \ K249526 (AMS).— SA: 2♂, 3♀: Parachilna Flinders Range Hale; 1♀: Melrose S.A. Oct. A.M.Lea; 1♀: Mt. Lofty, S.A. J.G.O.Tepper (SAMA). 2 ♂, Aldgate S.A. Oct. 1921 Wilson \ Cyphon lindensis Blackb. Id. by J.Armstrong (MV, T-21993, T-21994).— VIC: 1♂: Eltham 24.6.25 \ Cyphon lindensis Blackb. Id. by J.Armstrong (T-21995); 1♀: 7 km W Neerim, Vic., Tarago River 1.Mar.1972. Neboiss (T- 21998) (all MV). 1♂: 37.44S 145.43E VIC Warburton 2.2 km NE on Acheron Way 320m 27.Feb,1993 931 A.Newton &amp; M.Thayer \\ Euc.regnans forest w.Acacia dealbata pyr.fogging fungusy logs (ANIC).− TAS: 1♂: 42.34S 147.50E 4km W Oxford TAS 27.Jan.1983 J.C.Cardale coll. at light (ANIC). 1♀: Clarendon, Griffith collection Id. By A.M.Lea Macrohelodes sp. Tas. 469; 2♂: Maitland Tas. 471 Griffith collection id. By A.M.Lea; 1♂: Launceston Tas. Lea; 1♂: Hobart Tas. Lea; 1♂: Hobart Tas. Lea \ Cyphon 8788 Tasmania; 3♂: R. Isis Tas Dec. 0 1 Griffith collection Id. by A.M.Lea (all SAMA). 2♂: Tasmania \ Cyphon lindensis Blackb. Id. by J.Armstrong (MV; T-21996, T-21997)— QLD: 1♂: Ewan Rd 4 mi W of Paluma, N.Q., at light, 9.1.68, J.G.Brooks; 1♂: Wallaville, Qld. T.L.Bancroft (all ANIC). 1♀: QLD Ellis Beach CHSWatts 12\10\12 \ beating riparian vegetation \ 749 \Watts det. lindensis (SAMA). 1♀: Brisbane QLD. Oct. 61 Light Trap \ UQIC Reg. #88663; 1♀: N.E.Qld. 19°30'Sx 146°57'E, Mt. Elliott summit 1150m, 12 May 1991, D.Cox, Pyrethrum, tree &amp; logs (QM Reg.No. T227415); 1♀: Bluewater range, 50 km WNW Townsville, N.E.Qld, 6–9.Dec. 1986, 700m, Monteith, Thompson &amp; Hamlet Flight intercept Trap (QM Reg.No. T227416); 1♀: Bulburin S.F., 600m, 9km E Many Peaks, C.Qld. 17 Sept. 1989, G.B.Monteith, Pyrethrum, rainforest (QM Reg.No. T227417); 1♂: C.Qld., 21°52'Sx 147°18'E Upper Hall Ck, via Camilla, 4.Dec.96 – 6.Apr. 1997, G.Monteith and E.Mulder, Intercept, RF at Creek (QM Reg.No. T227413); 1♂: MEQ: 21°36'Sx 148°58'E, Blue Mt. 0.6 km SE, 930m, 4.Oct, 1999, G.B.Monteith, rainforest, Pyrethrum trees &amp; logs, 7806 (QM Reg.No. T227414) (all QMSB). 1♀: Mt. Spec N.Q. 1/67 G.B. \ J.G. Brooks, Bequest (ANIC).</p><p>Additional material: 1♂: Blue Mts., Blackburn [NSW; body missing, genitalia slide only] (SAMA). 1♀: 34.24S 150.50E Mt. Keira Scout Camp NSW ca. 320m 4/ 5 Mar.1981 Lawrence &amp; Calder \ under bark rotten logs (ANIC).</p><p>Habitus. BL 2.5−3.5mm, BL/BW ~1.6. Similar to N. lindensis, most specimens yellow with a round brown spot near the light sutural interval at half length of elytron. In some, there is also a brown stripe along the suture extending forward to the scutellum and sometimes further, to just inside of the humerus.</p><p>Male (Figs. 11−13): Segments 8 and 9 similar to N. lindensis . Penis more slender than in N. lindensis, pala rounded. The parameroids are a little longer than the trigonium, curving around its tip. The sclerotized trigonium apex is shaped like a wide Y. The truncate end of each branch is rough and bears ventrally some strong teeth (Fig. 12, seen by transparency). The long hollow cone between penis and S9 is beset with fine microtrichia and projects ventrally from under the trigonium. The tegmen is funnel-shaped, basally wide, ventrally on each side with two (occasionally one) huge falcate claws, no separate parameres. The caudo-lateral portions of the tegmen are soft and unpigmented, with small spinules and asperities.</p><p>Female. Tip of S7 with denser pilosity than rest, segment 8 resembles N. lindensis . Ovipositor (Fig. 15) unmodified. Prehensor (Fig. 16) similar to N. lindensis but the central sclerite with the sclerotized nipple possesses two rough deep pockets, the front end of the plate is parabolic. On the opposite side of the duct lies a narrow loopshaped sclerite and in front of it a wide sclerotized area whose anterolateral ends carry the spinule bands which are composed of larger teeth but less dense than in N. lindensis .</p><p>Notes. Nothocyphon ypsilon and N. lindensis are close relatives that are sympatric in much of their large mainland range. However, of the two only N. ypsilon occurs in Tasmania which permitted the definite association of sexes.</p><p>Etymology. The name alludes to the resemblance of the penis tip to the Greek letter Y.</p></div>	https://treatment.plazi.org/id/039F8D3EFFB4FFEC96964236FB14F9E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFB7FFED9696417DFD22FC49.text	039F8D3EFFB7FFED9696417DFD22FC49.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon taurus	<div><p>Nothocyphon taurus, n. sp.</p><p>(Figs. 17−20, 166, 173)</p><p>Type material: 1♂ holotype, 1 ♀ paratype: Paluma, N.Q. 10.i.68, J.G.Brooks (ANIC).</p><p>Additional paratypes: 1♂: Mt.Lewis 800m QLD 26.Dec.1986 H&amp;A.Howden flight intercept.trap; 6♀: Mt.Spec N.Q. 1/67 G.B. \ J.G. Brooks, Bequest; 1♂, 1♀: Mt.Spec N.Q. 1/69 G.B. \ J.G. Brooks, Bequest [originally on one card]; 1♀: Cardstone N.Q. 16.xii.65 at light K.Hyde (all ANIC). 1♀: Australia: N.Qld, 7 km WNW Paluma, 16.xii.1988 – 13.i.1989 Storey &amp; Dickinson MDP 1 Malaise trap (QDPC).</p><p>Additional material examined: 1 ♀: Paluma, N.Q. 10.i.68, J.G.Brooks (no prehensor; ANIC).</p><p>Habitus. BL 2.4−2.8mm, BL/BW ~1.5. Broadly oval with brown spot on yellow to light ochre elytron, fairly convex (Fig. 173). One dark specimen has a partly brown pronotum and a brown macula behind the scutellum next to the suture but the sutural interval remains pale (Fig. 166).</p><p>Male (Figs. 17−19). A slender hollow spine between S9 and penis. Penis slender, the pala is an elongate caudally narrowing oval. Trigonium inserted near 3/5th of penis length, wide, short, caudally truncate, the caudal edges extended into large caudo-laterally curved horns, each with a few coarse spines at tip. Parameroids much longer than trigonium, with a long paramedian sclerite strip ending in a sinuous tip. Laterally, an extension of the sclerite resembling a fish hook supports delicate membranes. Apical area of parameroids with sensory pores.</p><p>Tegmen lying over penis base, sides downcurved and continued caudad as slender parameres which are longer than the trigonium but shorter than the parameroids. Caudo-laterally each paramere with several sharp hooklets.</p><p>Female (Fig. 20). Abdominal sternites unmodified except last sternite indistinctly truncate. The ovipositor and the segment 8 resemble N. lindensis . Prehensor: The wrinkly internally spiny oviduct widens to a parallel-sided pocket which is weakly sclerotized and has a scaly inner surface. Caudally the pocket narrows, the tip seems to be extensible. Inside the pocket lie two anteriorly diverging sclerite bands with several strong teeth. Laterally from their truncate caudal ends are groups of hair-like slender spines. Opposite the sclerite bands lies an arched line from which originates a pair of thread-like sclerites with sharp teeth.</p><p>Notes. Represents the lindensis -group in the tropics. Association of sexes is by agreement in habitus and pigmentation, in combination with common occurrence. The spine-like cone beneath the penis is very similar to N. lindensis, N. ypsilon, and N. banksiae n. sp. while the prehensor resembles those of N. donnabuangi n. sp. and N. imitator n. sp., see below.</p><p>Etymology. The scientific name of buffalos (a noun in apposition) was chosen because in ventral view the trigonium resembles the horned front of a bull.</p></div>	https://treatment.plazi.org/id/039F8D3EFFB7FFED9696417DFD22FC49	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFB6FFED969644D4FB6DF928.text	039F8D3EFFB6FFED969644D4FB6DF928.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon banksiae	<div><p>Nothocyphon banksiae, n. sp.</p><p>(Figs. 21−24)</p><p>Type material: 1♂ holotype: NSW New England NP Banksia Point 22/11/06 CHS.Watts / 597 (SAMA).</p><p>Habitus. Oval, stout, BL 2.6mm, BL/BW ~1.55. Light brown, shining. Centre of head, a narrow strip along front margin of elytra including humeri and scutellum yellowish brown. Sutural interval lighter than rest of elytra. The granular punctures on the head are very fine and dense, those on pronotum little larger. The normal punctures on the elytra again larger but still relatively fine, over 2 diameters apart. The pilosity is light brown to yellowish. Antenna short and stout, flagellar segments only about 1.5 times longer than wide at apex.</p><p>Male (Figs. 21−24). The base of S9 is not visible, caudally S9 is indistinctly bilobed, slightly scooped out, the apical setae are unusually small. In the centre rises a sharp caudally pointing hollow tooth. Penis with long anteriorly rounded pala that narrows gradually caudad. The transverse bridge in the distal third of penis supports a U-shaped trigonium. Each of its two lobes is caudally rounded and caudo-medially densely beset with a pelt of spines that are directed medio-caudad. The parameroids are narrow transparent unsclerotized fingers a little longer than the trigonium (left one in Fig. 24 twisted, an artifact). The tegmen is a thread-like sclerite that widens continuously on the sides and supports the parameres. Each paramere is a slender strut with several spines, the subapical spine is largest.</p><p>Female. Unknown.</p><p>Note. The hollow spine on S9 (Fig. 19) shows that N. banksiae belongs in the N. lindensis species group. The shape of the trigonium and parameres differ much from all other species. The only ♂ was taken together with N. auritus n. sp.</p><p>Etymology. Named after the type locality which is itself named after the plant genus, Banksia .</p></div>	https://treatment.plazi.org/id/039F8D3EFFB6FFED969644D4FB6DF928	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFB6FFE896964037FF6CFA9F.text	039F8D3EFFB6FFE896964037FF6CFA9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon donnabuangi	<div><p>Nothocyphon donnabuangi, n. sp.</p><p>(Figs. 25−28, 169)</p><p>Type material: ♂ holotype, 1♂ paratype: 37.43S 145.41E VIC Mt Donna Buang 1200 m N of Warburton 811 26.Jan–11.Feb. 1987 A.Newton &amp; M.Thayer \ wet scler. Noth. cunn. Pyrethrin fogging old fungusy logs (ANIC).</p><p>FIGURES 25–31. Nothocyphon lindensis -group, N. donnabuangi (25−28) and N. imitator (29−31). 25, antenna; 26, 29, cleared extracted male terminalia, parts shown as trasparent; 27, 30, contour of female sternite 7; 28, 31, prehensor. 26, 29 and 28, 31 to the same scales, respectively; 27, 30 not to scale. Abbreviation: nS9, notch in caudal edge of S9.</p><p>Additional paratypes: 8 ♂: 37.43S 145.41E VIC Mt Donna Buang 1200 m N of Warburton 811 26.Jan– 11.Feb. 1987 A.Newton &amp; M.Thayer \ wet scler. Noth. cunn. FMHD #87 216 flight intercept (window) trap; 6 ♂: 37.43S 145.41E VIC Mt Donna Buang 1200 m N of Warburton 811 26.Jan–11.Feb. 1987 A.Newton &amp; M.Thayer \ wet scler. Noth. cunn. FMHD #87 219 flight intercept (window) trap (ANIC, 2 in PZ).</p><p>Additional material: 1♀ (presumed): 37.43S 145.41E VIC Mt Donna Buang 1200 m N of Warburton 811 26.Jan–11.Feb. 1987 A.Newton &amp; M.Thayer \ wet scler. Noth. cunn. FMHD #87 216 flight intercept (window) trap (ANIC).</p><p>Habitus. Elongate oval, elytra almost parallel over much of their length. Body flat. BL 2.4−2.8mm, BL/BW ~1.7. Light to dark brown, visibility of pronotal and elytral patterning decreases with degree of overall darkening. Head darkest, labrum and frons between antennae light, rufous. Pronotum with dark band across posterior half. Elytra with a narrow dark loop starting as a stripe behind the scutellum, next to the sutural space. Near 1/3 of elytral length the narrow stripe curves outward and forward without reaching the side margin of the elytron (Fig. 169).</p><p>Granular punctures coarser on pronotum than on head but both fine, the impressed punctures on the elytra are much larger.</p><p>Male (Fig. 26). T8, S8, and T9 as for the group. S9 deeply and widely divided, each lobe notched and itself divided into two sclerotized pigmented hairy lobes (nS 9 in Fig. 26). A membrane between S9 and penis bears a large triangular field of sharp spinules, tips directed cephalad. Tegmen widely divided into elongate parameres that are shorter than the penis. End of paramere with strip of forward-pointing irregular teeth, the caudal ones largest. Penis slender, pala narrow, its front end truncate. Trigonium apex forked, notch parabolic, deeper than wide, arms of fork gently upcurved, with irregularly serrate outer margin. There is a transverse crest at the base of the fork. The flat parameroids are little longer than the penis tip which they embrace from the outside.</p><p>Female (presumed; Figs. 27, 28). Last sternite ending in a slender triangular tip. Apices of rods of S8 separate, enlarged, spatulate. The ovipositor resembles other species in the group, with slender gonocoxites which are about 4 times longer than the tubular style with 2 sensory pegs. The anterior portion of oviduct containing the prehensor is covered with many fine spinules. The prehensor consists of a tongue-shaped sclerite that branches anteriorly. The arms diverge strongly. Along the edge of the structure stand about 11 very coarse teeth. In front of these sclerites a delicate sclerite band delimits a long parallel area leading to the bursal area covered with only minute pores. They are little discs surrounded by a corona of minute radial plicae, together resembling little suns.</p><p>Notes. The window trap samples on Mt. Donna Buang contained males of the related species N. horridus, N. imitator, and N. donnabuangi . The syntopic simultaneous occurrence corroborates their status as distinct biospecies. The samples contained only 2 females which by pigmentation and surface structure are tentatively assigned to N. donnabuangi and N. imitator, respectively.</p><p>Etymology. Named after the type locality, name formed as though Mount Donna Buang were a name in male gender.</p></div>	https://treatment.plazi.org/id/039F8D3EFFB6FFE896964037FF6CFA9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFB3FFE99696429FFE38FD6D.text	039F8D3EFFB3FFE99696429FFE38FD6D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon imitator	<div><p>Nothocyphon imitator, n. sp.</p><p>(Figs. 29−31, 170)</p><p>Type material: ♂ holotype, 1♂ paratype: 37.43S 145.41E VIC Mt Donna Buang 1200 m N of Warburton 811 26.Jan–11.Feb. 1987 A.Newton &amp; M.Thayer \ wet scler. Noth. cunn. FMHD #87 216 flight intercept (window) trap (ANIC).</p><p>Additional paratypes: VIC: 1♂: 337.43S 145.37E VIC Mt Ben Cairn 960m, NW of Warburton 30.Jan.– 11.Feb.1987 A.Newton &amp; M.Thayer \ wet scler.for. FMHD 87 250 Flight interception trap; 5♂: 37.43S 145.41E VIC Mt Donna Buang 1200 m N of Warburton 811 26.Jan–11.Feb. 1987 A.Newton &amp; M.Thayer \ wet scler. Noth. cunn. FMHD #87 216 flight intercept (window) trap.— NSW: 1♂: South Ramshead 1850 Kosciusko Ntl Park, NSW, March 1982 Ken Green pitfalls 1♂: South Ramshead 2000 Kosciusko Ntl Park, NSW, April 1982 Ken Green pitfalls (all ANIC).</p><p>Additional material examined: 1♀ (presumed), same data as holotype (ANIC).</p><p>Habitus. BL 2.6−3.0mm, BL/BW ~1.8. A sibling of N. donnabuangi n. sp., a little bit darker (which, however, may be related to specimen age), the granular punctation of prothorax slightly finer than in N. donnabuangi, and the dark brown band on the elytra a bit wider (Fig. 170). However, definite identification requires study of the terminalia. These are similar, only distinctive details are described.</p><p>Male (Fig. 29). Apical penis fork wider than in N. donnabuangi, wider than deep, the lateral lobes are flat, irregular teeth on the outside occur only near the tip. The parameres are largely membranous with a lateral row of large coarse teeth on a sclerotized fold which in slide mounts appears as elongate sclerite with shallowly serrate outer margin. S9 of same general shape as in N. donnabuangi but the hairy caudal lobes are entire, not excised. The armature ventrally from the penis consists of two separate patches of needle-like spinules. They are caudally connected to a triangular field with a median strip of teeth.</p><p>Female (presumed, see under N. donnabuangi; Figs. 30−31). Sternite 7 is shorter than in N. donnabuangi, it ends in a blunt triangular tip. The tooth-bearing sclerites of the prehensor are not bent but only gently curved. Each bears only about 9 strong teeth.</p><p>Note. Several males preserved dry on cards have everted genitalia. They are easily recognized by the entire lobes of S9 and two separate spinule patches between them, as well as the widely separated, only apically spinose branches of the penis tip. The presumed female has somewhat finer pronotal punctures than the female assigned to N. donnabuangi .</p><p>Etymology. The name is a noun in apposition highlighting the great similarity, also of the genitalia, almost imitating N. donnabuangi .</p></div>	https://treatment.plazi.org/id/039F8D3EFFB3FFE99696429FFE38FD6D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFB2FFE99696420EFA1AF8C8.text	039F8D3EFFB2FFE99696420EFA1AF8C8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon lanceolatus	<div><p>Key to males of the lanceolatus -group (the fragmentarily known sp. A not in key)</p><p>1 Trigonium flat, only fine denticles along edges (e.g., Figs. 33, 51)............................................... 2</p><p>1' Trigonium with raised ridges bearing large spines (e.g., Figs. 44, 47)............................................. 6</p><p>2 Caudal tips of the parameres of the two body sides separate from each other, not connected (e.g., Figs. 37, 51)............ 3</p><p>2' Caudal tips of left and right parameres connected by a transverse spine-bearing sclerite (Fig. 56)....... N. denticulatus n. sp.</p><p>3 Each paramere is a single continuous sclerite armed with teeth, each ends in a bare sclerotized knob or process (e.g., Figs. 32, 36)................................................................................. N. lanceolatus n. sp.</p><p>3’ Each paramere subdivided into a large membranous lobe and a toothed ventral sclerite (e.g., Figs. 48, 51)............... 4</p><p>4 Trigonium with long parallel apex, a large eversible band of spines on the membranous part of paramere (Figs. 50, 51); elytra with dark curved band in anterior half (Fig. 167)............................................... N. taeniatus n. sp.</p><p>4' Apex of trigonium different, parameres without eversible spinule band, elytra brown................................ 5</p><p>5 Trigonium oval, regular serrations along outer edge of both parts of the parameres (Fig. 48)............ N. biserratus n. sp.</p><p>5' Trigonium tongue-shaped, the parameral sclerite with very irregularly shaped large teeth, near base and distally on otherwise membranous lobe sclerotized processes with numerous spines along edge or at apex (Fig. 49)........... N. nungatta n. sp.</p><p>6 Trigonium narrow, the paramedian dentiferous ridges start some distance from tip (Figs. 43, 44)....... N. serratipenis n. sp.</p><p>6' Trigonium wider, dentiferous edges beginning at tip, rising gradually, end declining abruptly (Fig. 47). N. multidentatus n. sp.</p></div>	https://treatment.plazi.org/id/039F8D3EFFB2FFE99696420EFA1AF8C8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFB2FFF49696404FFB0EFC4A.text	039F8D3EFFB2FFF49696404FFB0EFC4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon lanceolatus	<div><p>Nothocyphon lanceolatus, n. sp.</p><p>(Figs. 32−40)</p><p>Type material: 1♂ holotype: 42.08S 146.13E, TAS, Tas.World.Her.Area, Lake St.Clair Site 1 HEC Road, 18 Feb.1997, sweep, P.Greenslade (ANIC). 1♂ p aratype, 1 presumed ♀ taken together with the holotype (ANIC).</p><p>Additional ♂ paratypes and presumed (non-type) females (TAS): 1♂: Cyphon fenestratus Blackb. Id. by HJCarter/ 5840 TAS [plus illegible text; red ink on card] (SAMA). 1♂: 42.08S 146.13E, TAS, Tas.World.Her.Area, Lake St.Clair Site 2, Lake St. Clair Road, 18 Feb.1997, sweep, P.Greenslade; 2♂: L. St. Clair Tasmania J. Armstrong; 10♂, 1♀: Cradle Mtn Tasmania J. Armstrong. 1♀: 41.50S 148.03E TAS Pelion Hut 3 km S Mt.Oakleigh, 860m 15–10 Feb. 1990 I. D. Naumann, rainforest; 1♂ same, but: 6 Feb. 1990, sweeping closed forest edge I. Naumann; 1♂ same, but: 13 Feb.–7 Mar.1991, I.Naumann, M.Horak, Malaise #1 closed forest; 3♀: 41.50S 148.03E TAS Pelion Gap, 2 km ENE Mt. Ossa, 1200m, 13 Feb.–17 Mar. 1991, I.Naumann, M.Horak, Malaise trap \ Eucalyptus coccifera woodland with alpine heath mats; 1♂: TAS, Cradle Mtn. Dove R. 1000m, 30.1.1989, D. Bickel alpine yellowpane (all ANIC). 4♂: Cradle Mtn. Tasmania, Carter &amp; Lea (SAMA, 1 in PZ). 1♂: N.W. Tasmania Cradle Valley 3.000 feet 12.1.1923 /Brit. Mus. 1925-120; 2♂: Strahan Tasmania J. Armstrong/ Paratype / [unpublished manuscript name] Armst. Id. J. Armstrong/ Brit. Mus. 1961 61 (all NHML). 1♂: 41.16S 145.17E Hellyer Gorge TAS 17 Jan.1983 I.D.Naumann &amp; J.C.Cardale coll.; 1♂: 42.01S 145.32E TAS 8km N Queenstown 22 Nov.1988 P.B.McQuillan &amp; E.S.Nielsen; 2♂: AUSTRALIA, TAS, Strahan, 5km W 42°09'S, 145°16'E 6 February 1992 Tom Gush \ on flowering Leptospermum sp. Tom Gush Collection 3367; 8♂: Strahan Tasmania J. Armstrong (all ANIC). 1♂: SW TAS, Lower Gordon R., 42.56S 145.50E, 42.54S 145.54E, Howard, Hill M.E.C. 128L7090 Feb. 1977 moss /12; 1♂: 42.10S 146.10E, 7km SWbyW Derwent Bridge, 16 Jan–2 Feb. 1989 I. D.Naumann &amp; J.C.Cardale Malaise \ ethanol (all ANIC). 2♂: TAS 18K SW Derwent Bridge 29/11/00 C.H.S.Watts; 4♂: TAS Franklin beaches 4K N Derwent bridge 30/1/06 CHS.Watts on Leptospermum flws (all on same card, penes projecting); 1♂, 1♀: TAS 15k N Tullah 41°35'48S 145°40'43E 30/10/02 CHS.Watts; 1♂: Huon R. Tas Lea \ Cyphon 8802 Tasmania; 15♂: Summit of Mt. Wellington Tas. (Lea) \ Cyphon Tasmania 8801; 1♂: Mt. Wellington, Tas. 12.12.0 1 Griffith \ Griffith collection Id. by A.M.Lea \ Cyphon nr. adelaidae Bl. S.A. 7946 (all SAMA). 2♂: Summit of Mt. Wellington, Tas. (Lea) (QMSB; QM Reg.No. T227409, T227410)). 1♂: Summit of Mt. Wellington TAS Lea; 1♂: Mt. Wellington TAS Lea; 1♂: 42.54S 147.13E TAS, Mt. Wellington summit 1240m, 5 Feb.1992 C.Reid under stones or on flowers; 2♂: Hobart Tasmania J.J.Walker \ G.C.Champion Coll. B.M. 1927- 409 (all ANIC). 4♂: Hobart Tasmania J.J.Walker \ G.C.Champion coll. B.M. 1927-409; 1♂: Hobart 91-88 \ 2463 (all NHML). 1♂: Australia Tasm. Hartz Mts NP., 8–10.ii.1980, nr. Leptospermum A.Newton, M.Thayer; 1♂: 42.43S 147.50E TAS Sandpit For. Reserve S of Orford 200m 915 17.Jan–2.Feb. 1993 A.Newton &amp; M.Thayer \ Euc.globulus w/rainf understory FMHD #93-45 ex window trap (all ANIC).</p><p>Additional material examined: TAS: 2♂, 2♀: TAS 18K SW Derwent Bridge 29/11/00; 8♂: TAS Narcissus Bay Lake St. Clair 30/1/06 on Lepto &amp; Baeckea flws (all CHS.Watts, SAMA). 10♂, 1 presumed ♀: Gordon R. Tasmania J.Armstrong; 1♂: 42.41S 146.37E TAS Mt.Field NP Lake Fenton by dam 1000m 6.Feb. 1992 C.Reid Nothofagus gunni; 1 damaged ♂: 42.54S 147.13E TAS, Mt. Wellington summit 1240m, 5 Feb.1992 C.Reid under stones or on flowers (all ANIC).— VIC: 7♂: Grampians Vic. E.T.Smith (ANIC; originally mounted on 3 cards).</p><p>Habitus. Elongate oval, elytral margins almost parallel. BL 2.2−3.2 mm, BL/BW ~1.7. Brown to almost black, in relatively pale specimens the sutural interval is vaguely reddish. The appendages are yellowish to brown. The antennal base is unmodified, distal flagellar segments are just twice as long as distally wide. Head and scutellum with fine granular punctures, the granular punctures on the pronotum are larger. Elytra with still larger normal punctures. The pilosity is short, semi-erect, yellowish.</p><p>Male (Figs. 32−37). Segments 8 and 9 as for the genus. Penis slender, pala very narrow, longitudinally folded, forming a slender anterior tip. Trigonium and parameroids originate near the caudal third of penis. Trigonium flat, sides and apex with small teeth. Shape varies from triangular to broadly tongue-shaped and truncate; see Notes section below. The parameroids are gently curved flat rods that narrow caudally and are longer than the trigonium. The inner edge is straight, the outer one convex. The tegmen is a long thin sclerite loop caudally supporting the band-like parameres whose caudal portion widens, surrounding the penis ventrolaterally. Parameres with variable armature (see notes), the caudal teeth are always largest. A blunt spur-like sclerite supports the ventrolateral end of the paramere (e.g., Figs. 32, 33, 37).</p><p>Female (presumed). S7 with two paramedian oval patches of finely porous cuticle from where minute converging canals lead inward. Near the medial edge of each modified cuticular area stands a group of erect hairs that differ from the normal fine tomentum (Fig. 38). T8 and ovipositor unmodified, rods of S8 connected by a wide ring in front (Fig. 39).</p><p>Prehensor (Fig. 40) located in an abruptly widening section of the oviduct. Two parallel bands of slender spines lead towards the wide section. Its projecting caudo-lateral angles are two apparently extensible pockets with concentric folds. Between them lies a trapezoidal sclerite with two paramedian ridges armed with strong anteriorly directed teeth. Anteriorly, on the opposite side of the duct a toothed sclerite ring is located. It ends in two slender strips extending forward towards the balloon-shaped bursella, surface with minute sun-like cuticular ornaments.</p><p>Notes. The male genitalia vary. In most specimens the trigonium is bluntly rounded but the tip may also be narrow and pointed, or spade-shaped (Figs. 33, 36, 37). The armature of the parameres varies also. A continuous band of coarse teeth may extend from the large caudal hooks to the anterior end of the paramere (Fig. 33). In other specimens the armature may be altogether finer (Fig. 37). In still others, a few fine teeth only may be present between the large distal hooks and the basal patch of teeth (Fig. 36). In groups of specimens taken together the various morphs coexist, variation seems to be individual. However, ten males taken together at the Gordon River are all small (2.2−2.6mm; Fig. 37), have relatively short genitalia, a sharply pointed trigonium, more strongly bulging parameroids and weak armature of the parameres. The female in that sample has a spermatophore in the bursella. Another male taken at the Lower Gordon R. is normal and resembles Fig. 33. The presumed females agree in habitus and were taken together with male N. lanceolatus .</p><p>J. Armstrong attached type labels to specimens of N. lanceolatus but assigned them to two different taxa none of which he described. At the same time he confused N. lanceolatus with other taxa, e.g., N. frater and another presently undescribed species.</p><p>N. lanceolatus is widespread in Tasmania. Its occurrence in the Grampians should be confirmed by contemporary collections.</p><p>Etymology. The name is a Latin adjective describing the lanceolate shape of the pala.</p></div>	https://treatment.plazi.org/id/039F8D3EFFB2FFF49696404FFB0EFC4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFAFFFF4969644D6FAFDF8A5.text	039F8D3EFFAFFFF4969644D6FAFDF8A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon serratipenis	<div><p>Nothocyphon serratipenis, n. sp.</p><p>(Figs. 41−45)</p><p>Type material: 1♂ holotype, 2 ♂ paratypes: Tasmania \ K 24-144 \? Cyphon var. pictus Blck. ? Id. by JHCarter \ Cyphon very near but prob. not pictus Bl. Id. by A.M.Lea \ K249509 (AMS).</p><p>Additional paratypes: 1♂: Hobart TAS. Lea; 2♂♂: 42.41S 146.43 TAS, National Park, low bushes entrance to NP, 150m, Feb. 1992, C. Reid; 1♂ paratype, 1 probable ♀: 1090 [red ink] \ Tasmania \ Cyphon doctus Lea? var. cf. pictus Blkb. (originally together on the same card which has a green band across the base, perhaps taken in copula; female genitalia missing) (all ANIC). 2♂: TAS, South Esk R. Evandale, 1.3.67 A.Neboiss (MV, T-22023, T-22024).</p><p>Habitus. BL 2.9−3.0 mm. BL/BW ~1.7. Elongate-suboval. Head and pronotum yellowish brown, pronotum with brown marks. Elytra light to dark brown, with variably distinct pattern (Fig. 45). On either side of the scutellum the front edge of the elytra is pale. A dark band extends over the anterior 2/3 of the elytral side margin. Directly behind the humerus is a short curved spot. Near midlength a dark band extends obliquely across the elytra, with a short backward spur along the light sutural interval.</p><p>Male (Figs. 41−44). Segments 8 and 9 as standard in the group. Tegmen a narrow band supporting the elongate parameres which have no armature except a blunt curved caudoventral spur.</p><p>Penis slender, the hairpin-like pala occupies 2/3 of its length. The widest point is at the origin of the slender parameroids. They are longer than the trigonium which they embrace. The trigonium is carrot-shaped with roughened apex. Two prominent ventral keels rise cephalad beginning at some distance from apex and side edge, each with a row of large sharp teeth (Figs. 43, 44).</p><p>Female (presumed). The only specimen is very dark but the elytral pattern can still be recognized. The female abdominal sternites are unmodified, damaged, genitalia missing.</p><p>Note. The male genitalia readily separate N. serratipenis from its relatives, especially N. multidentatus n. sp., see account of that species.</p><p>Etymology. The name is an adjective referring to the saw-like crests on the penis apex (Latin serra, a saw).</p></div>	https://treatment.plazi.org/id/039F8D3EFFAFFFF4969644D6FAFDF8A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFAEFFF596964365FD84F871.text	039F8D3EFFAEFFF596964365FD84F871.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon multidentatus	<div><p>Nothocyphon multidentatus, n. sp.</p><p>(Figs. 46, 47)</p><p>Type material: 1♂ holotype: Launceston Tas. Lea \ Cyphon 8805 Tasmania (SAMA).</p><p>Paratypes: 1♂ on roof of tent with light inside Rav. de Casoars [34.6565S, 136.5773E], K.I. [Kangaroo Island], 24.Oct 1951 G.F.Gross; 1♂: Kangaroo I. S.A. \ prob. Cyphon near adelaidae Bl.; 1♂: SA 13k W Meadows [35.1790S, 138.7629E] 9/12/96 C.H.S.Watts (all SAMA).</p><p>Habitus. Size 3.0− 3.2mm, BL/BW ~1.65. The elytra are dark, with some still darker fascia that seems to resemble N. taeniatus (Fig. 167).</p><p>Male (Figs. 46, 47). Generally similar to N. serratipenis, trigonium and parameres differ. Trigonium wider, the toothed ridges are connected to the rough field at the penis apex and run along the sides rising high in front and then declining abruptly. The ridges carry numerous slender hooks, the longest ones near the highest point. The parameroids are curved around the trigonium and partly concealed by it in ventral view. Tegmen with thin basal sclerite and complexly folded ventrolateral paramere lobes that seem to wrap around the pala. There is no armature, also no spur. Dorsally, some transparent thin sclerite extends caudad from the basal sclerite of the tegmen, branching at some distance from it. The branching is strengthened by a sclerotized U-shaped bracket (arrow in Fig. 47).</p><p>Female. Unknown.</p><p>Note. The few specimens were taken in South Australia and northern Tasmania, the presently known range is disjunct. At first glance the genitalia are similar to N. serratipenis from which it can be distinguished by the different position of the toothed ridges on the penis, the lack of any armature on the parameres and the presence of the U-shaped bracket in N. multidentatus .</p><p>Etymology. The multitude of slender teeth on the trigonium is reflected by the name (the Latin adjective, multidentatus, with many teeth).</p></div>	https://treatment.plazi.org/id/039F8D3EFFAEFFF596964365FD84F871	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFA9FFF29696420FFEE2F81A.text	039F8D3EFFA9FFF29696420FFEE2F81A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon biserratus	<div><p>Nothocyphon biserratus, n. sp.</p><p>(Fig. 48)</p><p>Type material: 1♂ holotype: Hazelbrook [~ 33.23°S, 150.43°'E] N.S.W. Mrs L.Smith (ANIC); 1 ♂ paratype: Sydney Lea (SAMA).</p><p>Habitus. Elongate oval, BL 2.1−2.4mm, BL/BW ~1.55. Entirely light brown. Surface sculpture and pilosity as in the congeners. Antennae short and stout, flagellar segments only about 1.5 times longer than wide at apex.</p><p>Male. Segments 8 and 9 typical of the group. Penis long and slender, trigonium apically wider than in the congeners, a little scooped, the raised margins with faint scaly marks. Parameroids similar to the related species. Some transparent sclerotized projection stands perpendicular to the penis axis immediately in front of the origin of parameroids and trigonium. It was somehow connected to the parameres, details not recognized. The anterior part of the pala is narrow, folded. The tegmen is a thin arched sclerite supporting the parameres. Each is a large membranous folded structure with a slender relatively short sclerite dorsally and a wider one with a long tongueshaped apex on the ventral side. Both sclerites bear numerous sharp teeth along the outer edge.</p><p>Female. Unknown.</p><p>Note. The peculiar ventral process of the penis appears to be homologous to less obvious structures in N. taeniatus . There is not enough material to study details. Presently N. biserratus marks the northern border of the group range.</p><p>Etymology. The adjectival name biserratus (Latin, with two saws) describes the two long rows of regular teeth on each paramere.</p></div>	https://treatment.plazi.org/id/039F8D3EFFA9FFF29696420FFEE2F81A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFABFFF096964795FC13FC21.text	039F8D3EFFABFFF096964795FC13FC21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon nungatta	<div><p>Nothocyphon nungatta, n. sp.</p><p>(Fig. 49)</p><p>Type material: 1 ♂ holotype, 1 possibly conspecific ♀ (no type status) originally mounted on one card: NSW nr Nungatta Creek [~ 37.19S 149.43E] 58 88083N 557 20893E [sic!] 13.2.0 0 \ J&amp;R Miller (SAMA).</p><p>Habitus. Elongate oval, BL 2.8mm, BL/BW ~1.7. Granular punctures on head fine, a little larger on pronotum, spaced, integument shining. Normal elytral punctures fine and dense, although distinctly larger than on pronotum. Dorsal face dark brown, head darkest. Antennal flagellum infuscate, flagellar segments barely 2x as long as wide at apex. Pilosity dark, semi-erect.</p><p>Male. Segments 8 and 9 typical of the group, unmodified, lobes of S9 visible under the penis. Pala longer than half total length of penis, the lateral sclerites lean against each other but diverge again slightly anteriorly. The parameroids are simple flat lobes a little longer than the trigonium. Laterally they are intimately connected with surrounding membranes. The trigonium narrows caudally, it is tongue-shaped, the apex is bluntly rounded and armed with few small denticles. The tegmen (torn on one side) is a narrow sclerite band laterally supporting large complex parameres. The curved rod-like base divides into a medial lobe with huge teeth separated by deep rounded notches; the apex of this portion seems to taper dorsally from the penis. Near the forking point the basal rod supports a rounded unarmed medial process and a curved lateral one with an apical row of stout teeth (mp in Fig. 49). The outer paramere lobe is membranous except for a small distal area supporting a medially directed short process with a crown of irregular sharp teeth (Fig. 49, lp).</p><p>Female (presumed). Resembles the male. Abdominal sternites unmodified, the rods of S8 anteriorly separate. The ovipositor is of the standard build, bursellar sclerite or prehensor not found.</p><p>Note. Structural details of the parameres remain unknown, the complex folding of the two partly overlapping lobes was not disentangled.</p><p>Etymology. Named after the type locality; a noun in apposition.</p></div>	https://treatment.plazi.org/id/039F8D3EFFABFFF096964795FC13FC21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFABFFF09696433CFB27F87C.text	039F8D3EFFABFFF09696433CFB27F87C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon taeniatus	<div><p>Nothocyphon taeniatus, n. sp.</p><p>(Figs. 50, 51, 167)</p><p>Type material: 1 ♂ holotype, 2 ♂ paratypes: Mt. Gingera [35.58S, 148.78E] NSW 18.ii.1969 light trap S.Misko (ANIC).</p><p>Habitus. Elongate oval, BL 2.5−2.7mm, BL/BW ~1.55. Yellowish brown, pronotum with darker centre. Each elytron with an arched brown band which is narrow behind the scutellum and widens laterally, meeting the elytral margin a little before midlength (Fig. 167). Punctures on fore body fine, granular, the normal punctures on the elytra a little larger, denser, integument shiny. The semi-erect pilosity is yellowish. Appendages light brown, antennae moderately slender, flagellar segments about twice longer than apically wide.</p><p>Male (Figs. 50, 51). T8 as for the group, S8 not observed. T9 with long straight apodemes which are relatively strong but the plate is completely membranous. Base of S9 not recognized, caudal portion with some setae, narrowly and deeply divided, each lobe shallowly excised between prominent lateral corners, edge irregular.</p><p>Penis slender, with narrow spatulate crested base. Trigonium with long almost parallel-sided apical portion, apex with asperities. Parameroids little longer than trigonium, tips triangular, apex blunt, sides connected to membranes. In front of the trigonium base appears an oval window (w) laterally delimited by raised crests (c) and some projecting sclerotized structures (p). The tegmen is a narrow arch, sides directed backward. Immediately after the bend each side divides into two large lobes. The medio-ventral lobe (mv) widens caudally and bears on its external edge huge widely spaced teeth; the unarmed apex tapers caudally. The dorsolateral lobe (dl) bears a long row of strong teeth along its outer edge before it tapers apically. Between the two lobes a band armed with many sharp hooks curves sideward and forward.</p><p>Female. Unknown.</p><p>Note. A close relative of N. nungatta and N. biserratus; the present species stands out by its elytral pattern. The three related species differ mainly in the complex parameres.</p><p>Etymology. The name (Latin taeniatus, banded) alludes to the dark band on the elytron.</p></div>	https://treatment.plazi.org/id/039F8D3EFFABFFF09696433CFB27F87C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFAAFFFE96964795FBCEFEAF.text	039F8D3EFFAAFFFE96964795FBCEFEAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon	<div><p>Nothocyphon sp. A</p><p>(Fig. 52)</p><p>Material examined: 1♂: 35.51S 146.15E VIC Baw Baw Alpine Res. 1.2 km NE Neulynes Mill, 1145m, 28.Jan.– 10.Feb. 1987 A.Newton &amp; M.Thayer \ wet scler.for. FMHD 87 236 Flight interception trap (ANIC).</p><p>Note. The 2.6 mm long brown beetle is severely damaged and not formally named but is mentioned to document the diversity of the group.</p><p>Segment 8 missing. Of segment 9 remain one apodeme of the tergite (not shown) and the unusual, strongly sclerotized angular distal lobes of S9. Of the penis only the slender pala is undamaged. The parameroids seem to resemble the related species, shape of trigonium unknown. The tegmen is strong and supports parameres composed of three large components. The median one has a large medially projecting knee-like sclerite with a few teeth. Caudally it is tongue-shaped, with a few basolateral teeth and a large subapical one. The medial lobe bears a single series of spines along its edge, the basolateral lobe several, in irregular arrangement.</p></div>	https://treatment.plazi.org/id/039F8D3EFFAAFFFE96964795FBCEFEAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFA5FFFE969646B5FB91F9CD.text	039F8D3EFFA5FFFE969646B5FB91F9CD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon denticulatus	<div><p>Nothocyphon denticulatus n. sp.</p><p>(Figs. 53−56, 168)</p><p>Type material: 1♂ holotype, 1♂ paratype: South Ramshead NSW Kosciusko NP 1850m subalpine grid Jan. 1983 pitfalls Ken Green coll (ANIC).</p><p>Habitus. BL 2.9−3.1mm, BL/BW 1.7. Body elongate oval, elytra almost parallel over most of their length. Antennae slender, base unmodified, flagellar segments a little more than twice as long as apically wide. Head and pronotum with dense granular punctation, a little coarser on pronotum. The elytra have fine dense normal punctures, the semi-erect pilosity is yellowish.</p><p>The specimens are differently pigmented. The head is dark brown, the frontoclypeus rufous. The pronotum is brown to dark brown with diffusely lighter margins. The elytra of the smaller holotype are dark brown, the humeral area a little lighter, sutural interval and scutellum rufous. The larger paratype (Fig. 168) has yellowish brown elytra with a dark brown spot posterolaterally from the scutellum and a longitudinal band. The wide band starts in the second third of elytral length. It runs parallel to the side edge, being separated from it by a pale space similar to its own width. The band follows the caudal curvature of the elytron so that the ends of the bands are opposed but remain separated by the light sutural interval. The appendages are brownish, distal flagellar segments infuscate.</p><p>Male (Figs. 53−56). T9 with unconnected delicate apodemes which are slightly longer than those of T8. S9 has a weakly sclerotized rounded base, the plate is deeply divided into two tongue-shaped caudally setose lobes. The tegmen is a narrow arched sclerite. The elongate parameres are armed with fang-like teeth. Their caudal ends are connected by a transverse weak sclerite covered with minute denticles and a few large teeth along its rear edge. A vaguely sclerotized median area extends cephalad; probably the entire area between it and the basal tegminal sclerite is covered by a transparent membrane. Penis with slender pala whose anterior band-like sclerites lie flat, side by side, with a shallow median ridge. Trigonium and parameroids occupy the caudal third. The sides of the long trigonium are almost parallel, faintly concave. Trigonium apex shallowly excised, caudolaterally with a few denticles. The finger-shaped parameroids are longer than the trigonium, their rounded apices curve gently mediad.</p><p>Female. Unknown.</p><p>Note. The wide, plate-like marginal sclerites of the pala are typical of the group. Those in N. denticulatus lie flat, which seems to be an ancestral trait suggesting that the other species in which they are folded with opposed faces are a monophylum and together represent the sister-group of N. denticulatus .</p><p>Etymology. The species name refers to the spinous transverse caudal edge of the complex formed by tegmen and parameres. It is a Latin adjective ( denticulatus —provided with small teeth).</p></div>	https://treatment.plazi.org/id/039F8D3EFFA5FFFE969646B5FB91F9CD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFA4FFFF96964526FE62F958.text	039F8D3EFFA4FFFF96964526FE62F958.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon vandiemeni	<div><p>Nothocyphon vandiemeni, n. sp.</p><p>(Figs. 57−61)</p><p>Type material: 1♂ holotype, 3♂ paratypes: [TAS] 17 km W Maydena 28/1/00, C.H.S.Watts (SAMA).</p><p>Additional paratypes: 2♂: Cradle Mtn. Tasmania Carter &amp; Lea (SAMA). 1♂: 41.50S 148.03E TAS Pelion Gap, 2 km ENE Mt. Ossa, 1200m, 30 Nov.1990 – 8 Jan.1991, E.Nielsen, E.Edwards malaise #6; 1♀: 41.50S 148.03E TAS Pelion Hut 3 km S Mt.Oakleigh, 860m, 5-10 Feb.1990, I.D.Naumann, rainforest; 2♂: Cradle Mtn Tasmania J. Armstrong; 1♂: Strahan Tasmania J.Armstrong [as paratype of an undescribed new species]; 5♂, 2♀: 42.10S 146.07E 4 km SSE of Mt. Rufus 800m TAS 26–28 Jan 1980 Lawrence &amp; Weir (all ANIC).</p><p>Additional material examined: 2♂ taken together with the holotype can be identified with difficulty because of an unsuitable medium used during previous mounting (SAMA).</p><p>Habitus. BL 2.2−2.7mm. BL/BW ~1.7. Elongate, elytra almost parallel. Dark brown to black, front corners and a narrow seam along sides of pronotum reddish, tibiae and tarsi also reddish, antennae infuscate. Distal flagellar segments cylindrical, a little more than twice as long as wide. Punctures on head granular, rough, on pronotum finer and sparse, elytra with dense and much larger normal punctures.</p><p>Male. Segment 8 as described for the genus, unmodified, apodemes of T9 with basal transverse connection, S9 caudally pilose, shallowly bilobed, base indistinct (Figs. 57, 58). Penis slender, pala elongate, parallel-sided, front truncate, trigonium long and conical, ventrally beset with sharp hooklets, parameroids a little longer, tongueshaped, wide near midlength but tapering caudally, medial edge straight, outer edge angular. Tegmen a thin sclerite arch connecting the parameres which are flat triangular plates laterally armed with some sharp teeth. Near the mediobasal corner of the triangle inserts a slender conical flap, with slender teeth, mainly along the outside. At rest, the flaps are directed caudad (Figs. 59, 60), but point forward from the base of the fully everted genitalia.</p><p>Female. The exposed sternites are unmodified. The rods of S8 are parallel and in front connected by a short transverse bar. T8 and ovipositor unmodified. The prehensor (Fig. 61) consists of a wide bare caudally truncate sclerite loop which narrows anteriorly where it reaches the gonoduct. It bears a single row of large bursellar pores (Libonatti &amp; Ruta 2013), the bursella itself is bare. Opposite the loop lie two elongate plates which narrow caudally. Anterolaterally their surface has large, forward pointing scales.</p><p>Note. The beetles are very dark. The externally indistinguishable sexes were taken together at two sites. However, in the same area occur additional similar species, e.g., N. lanceolatus which may also be almost black but is distinguished by its unicolourous pronotum and (in direct comparison) slightly stouter antennae.</p><p>Etymology. Named after Anton van Diemen, name patron of van Diemens Land, the former name of Tasmania where the species is endemic.</p></div>	https://treatment.plazi.org/id/039F8D3EFFA4FFFF96964526FE62F958	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFA4FFFA969641D8FEE9FBB6.text	039F8D3EFFA4FFFA969641D8FEE9FBB6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon auritus	<div><p>Nothocyphon auritus, n. sp.</p><p>(Figs. 62−68)</p><p>Type material: 1♂ holotype, 11♂ paratypes: NSW New England NP Banksia Point 22/11/06 CHS.Watts (SAMA).</p><p>Additional paratypes: 4♂, 1♀: 30.30S, 152.24 E. New Engand Ntl Pk, NSW, 2–15.Oct.1985, I.Naumann, J.Cardale, Malaise trap (ethanol) (ANIC). 2♂: Australia NSW, Tom's Cabin Point Lookout, New England NP 30°29'54''S, 153°23'52.6''E, 14 November 2006, coll. D.Britton (AMS, K249619 &amp; K249620).</p><p>Habitus. Oval, BL 2.4−2.8mm, BL/BW ~1.6. Antennae short, the subterminal segments only about twice as long as wide at apex. Head dark brown, rest of dorsal face variably patterned, probably depending on specimen age. Fig. 62 shows an intermediate pattern. Centre of pronotum dark, narrow rufous seams along all margins or only the sides contrastingly yellow. Pale specimens (Fig. 63) with yellowish elytra are infuscate only between scutellum and humerus, and laterally from the humerus along the elytral margin. There is also a dark (sometimes almost black) streak parallel to the suture near the tip of the elytra but the sutural interval itself remains yellow. Dark specimens have dark brown elytra with slightly paler humerus and a paler area around the suture in the declivous caudal third but not reaching the elytral tip. This caudal area and the scutellum are always yellowish.</p><p>Male. Segments 8 and 9 as described for the genus (Figs. 64−66). Penis (Fig. 67) long and slender, pala long, anteriorly oval, caudally waisted, then again widened, the straight sclerites of the parameroids diverge caudally a little. The trigonium is much shorter than the parameroids, a slender triangle with some teeth along midline. From its tip two curved setae seem to arise which are supporting a wide membranous flange on either side of the trigonium tip. Laterally the flange is attached to the parameroid. The sclerotized rod of the parameroid is caudally bent mediad, the straight obtuse tips of the parameroids converge. Externally, a wide membrane along the said sclerite is basally indistinguishable from the membranous part of the paramere. Tegmen a simple arched sclerite supporting the straight parameres with medially curved membranous tips embracing the parameroids from the outside. The membranous part is only basally supported by a slender caudally tapering sclerite. Near the base the sclerite bears ventrally an ear-like flap with teeth along the caudal edge. The flap is provided with muscle fibres and apparently movable.</p><p>Female. Sternites unmodified. T8 and ovipositor unmodified. Rods of S8 connected by a short transverse bar. The gonoduct widens to a membranous pear-shaped pouch with a weakly pigmented sclerite arch in front, with several teeth at each end. On one side of the pouch is a finely reticulate longitudinal strip (Fig. 68) leading to the bursellar duct. The opposite side of the pouch carries many fine teeth and sharp asperities.</p><p>Note. Several males of Austrocyphon ovensensis (Blackburn) and Austrocyphon wattsi Zwick were at Banksia Point taken together with the present species. The Austrocyphon spp. are more slender than N. auritus, uniformly dark brown, also the scutellum. Their antennae are more slender, with subterminal segments almost 3x longer than wide at the apex. Similar habitus and the pale scutellum and subterminal mark along the elytral suture served to associate the sexes of N. auritus .</p><p>Some specimens preserved in ethanol have everted male genitalia which are now stored in glycerine, in microvials on the specimen pin (SAMA).</p><p>The intact genitalia resemble a whitish membranous conical envelope around the sclerotized central structures. The envelope is formed by the parameres and parameroids which are difficult to separate in preparations. When successful, a few fine teeth appear on the outside of the parameroid which mediate this firm connection. The earlike flaps stand perpendicular to the body axis and are directed ventrad, directly behind S7. The same is seen in a male preserved dry and mounted on a card (AMS).</p><p>Etymology: The name is an adjective. It refers to the ear-shaped mobile appendages, Lat. auritus, eared, provided with ears.</p></div>	https://treatment.plazi.org/id/039F8D3EFFA4FFFA969641D8FEE9FBB6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFA1FFF8969643CDFA06FD3A.text	039F8D3EFFA1FFF8969643CDFA06FD3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon wattsi	<div><p>Nothocyphon wattsi, n. sp.</p><p>(Figs. 69−76)</p><p>Type material: 1♂ holotype: [VIC] Melbourne, pond in Botanic Gardens, ~1957 C.H.S.Watts (SAMA); 3♂, 6♀ paratypes with the same data (SAMA). Additional paratypes: 3♂, 3♀: VIC Sassafras D.A.Duckhouse 06/60 C.H.S.Watts; 4♂: VIC Sassafras Dandenongs D.A.Duckhouse. 08/62 C.H.S.Watts (all SAMA). 1♂, 1♀: Olinda Victoria F.E.Wilson 30.11.29 \ in cop.; 1♂: 37.43S 145.42E VIC Cement Creek 625m N of Warburton 814 27.Jan– 11.Feb.1987 A.Newton &amp; M.Thayer \ Euc. regnans Notho.cunn. FMHD #87-232 flight interception (Window) trap (ANIC).</p><p>Additional material examined: 6♂, 4♀: Melbourne, pond in Botanic Gardens, ~1957 C.H.S.Watts; 1♀: VIC Sassafras D.A.Duckhouse 06/60 C.H.S.Watts; 1♀: VIC Sassafras Dandenongs D.A.Duckhouse. 08/62 C.H.S.Watts (all SAMA).</p><p>Habitus. BL 2.5−2.9mm, BL/BL ~1.6. A stout oval beetle, its pigmentation varies from brown with reddish seams of pronotum, reddish scutellum and sutural interval to entirely almost black. Antennae short, base unmodified, flagellar segments just twice as long as wide, terminal segment more elongate, almost cylindrical.</p><p>Male (Figs. 69−75). Segments 8 and 9 as for the genus. The penis is slender, subparallel but has an exceptionally wide front part of pala which is in situ curved up towards the tegmen. The parameroids insert near the caudal third of the penis, their sclerotized tongue-shaped apices curve mediad. On the outside, the parameroids are membranous and intimately connected to the closely appressed caudal lobe of the paramere. The trigonium is in dorsal view a triangle with apical teeth. In oblique view it looks rather massive, with many apical teeth. In ventral view, the trigonium is partly concealed by two slender sclerotized tubes originating from a common median base on the penis just basally from the trigonium. Each tube is covered with numerous sharp teeth (Fig. 74).</p><p>Membranes tightly connect the outer face of the parameroids with the parameres which originate from a strong basal tegmen across the pala. Near the base of each paramere articulates a triangular sclerite with a subterminal median patch of slender spines pointing caudad. Ventrally the paramere bears a sausage-shaped spiny swelling which is directed cephalad and forms a spiny bulb under the triangular appendage. The membranous distal part of the paramere is frazzled, with numerous superfine colourless filaments.</p><p>A hyaline tube armed with very fine spinules and some longitudinally striate weak sclerotization resembles an endophallus but is actually part of the vas deferens (Fig. 75). It is located in front of the penis, at a considerable distance, but was sometimes extracted with it during dissection.</p><p>Female. T8 and ovipositor unmodified. The rods of S8 are apically widened and connected by a transverse bracket which is wider than the space between the two rods further caudally (not shown). The prehensor (Fig. 76) is equipped with a trapezoidal sclerite from which rises a slender tube, extending forward into the lumen. Opposite this structure and a little further in front is the sinuous thickened front edge of a sclerite loop whose thinning sides extend cephalad and are accompanied by dense spinule bands. The bursellar pores are minute discs with radial plicae.</p><p>Note. The articulated triangle on the paramere and the connection between parameres and the membranous outer face of the parameroids are characteristic of the auritus -group. N. auritus shares inwardly directed tips of the parameroids. By the shape of the pala N. scutiger is most similar.</p><p>Etymology. Dedicated to Chris Watts, Adelaide, successful student of Australian marsh beetles who provided most of the present beetles. Some of them he collected himself and recognized that they are „close to lindensis “.</p></div>	https://treatment.plazi.org/id/039F8D3EFFA1FFF8969643CDFA06FD3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFA3FFF896964446FE95F8A5.text	039F8D3EFFA3FFF896964446FE95F8A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon scutiger	<div><p>Nothocyphon scutiger, n. sp.</p><p>(Figs. 77−81)</p><p>Type material: 1♂ holotype: 38.42S 143.34E VIC Bin Rd. 5.8km N of Cape Horn, Otway NP 450m 24 Jan.1987 A.Newton &amp; M.Thayer \ wet sclerophyll forest, at UV light at night (ANIC).</p><p>Habitus. Elongate oval, BL ~ 2.7mm, BL/BW ~1.7 (body broken between pronotum and elytra). Head brown, pronotum with transverse brown central mark, sides and front edge yellow, elytra and scutellum light brown. Punctures on elytra normal, rather fine, although larger than granular punctures on forebody. Pilosity semi-erect, light brown. Antennae slender, flagellar segments about twice as long as apically wide. Right mandible with small tooth, left one sinuous along medial edge.</p><p>Male (Figs. 77−81). Segments 8 and 9 as for the genus, except pilosity of T8 unusually short and fine, barely longer and stronger than the microtrichia that cover much of the plate. Penis of unusual shape, the very long anteriorly wide pala narrows in almost straight line. The base of the parameroids is in dorsal view (Fig. 79) concealed by a shield that is along its sides covered by fine backward-directed scales and spines and ends in two sinuous horns between which the tip of the trigonium is seen. The shield arises from the trigonium which in ventral view (Fig. 80) is a small slender triangle. The parameroids diverge from the base but are bent mediad near midlength. Their terminal portion is sinuous and partly membranous. The tegminal complex (Fig. 81) is very wide, the tegmen is a thin sclerite arch. Caudally, the long rod-like parameres take a hairpin bend, the apical portion with a series of moderate spines is turned back by 180°. Between the two parts of the paramere are remains of strong longitudinal muscles. On the outside of each paramere articulates near the base a slender sclerite with a small subterminal tooth.</p><p>Two sclerites resembling arched bands (arrow in Fig. 81) lie between the parameres. Together, the bands form a ring open at both sides. Both ends of one of the bands are connected by muscles remaining after incomplete clearing in KOH.</p><p>Female. Unknown.</p><p>Note. The slender sclerite articulating basolaterally on the paramere shows N. scutiger is a member of the auritus -group. The shield-like structure of the penis is exceptional. The homology of the arched bands between the parameres is doubtful.</p><p>Etymology. The shape of the penis suggested the name: Latin scutiger, somebody carrying a shield; a noun in apposition.</p></div>	https://treatment.plazi.org/id/039F8D3EFFA3FFF896964446FE95F8A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFFA2FFF996964177FB90F83F.text	039F8D3EFFA2FFF996964177FB90F83F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon amphora	<div><p>The amphora -group</p><p>T9 strongly reduced, in preparations often barely perceptible. The anteriorly truncate penis resembles an amphora, with handles formed by the abruptly outcurving parameroids (e.g., Figs. 83, 85: pb). The trigonium is basally wide, then restricted to a long bottleneck, the tip mostly again wider. The flat band-like parameroids are shorter than the apparently rigid trigonium which is not visibly articulated with the pala. There is a bracket-like transverse sclerite of uncertain homology, possibly the tegmen. The parameres are independent from it, separate, each with a basal rod and a caudal plate of variable form, armed with spines. Females (known only of N. armstrongi): S7 with two areas with micropores and fine canals (Fig. 89). The apodemes of S8 are anteriorly connected by a ring (not shown). Prehensor with two large strongly spinose sclerites (Fig. 90).</p><p>The few species live in the southeast of Australia and were rarely collected.</p></div>	https://treatment.plazi.org/id/039F8D3EFFA2FFF996964177FB90F83F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF9DFFC696964225FBC3F8B5.text	039F8D3EFF9DFFC696964225FBC3F8B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon amphora	<div><p>Nothocyphon amphora, n. sp.</p><p>(Figs. 83, 91)</p><p>Type material: 1 ♂ holotype: Pierces Ck Forest ACT 20 Jan.1989 I.McArthur IPS trap (ANIC). Paratypes: 1♂: 38.39S 143.42E VIC Haines Junc. 525m 1.9km W. on Turtons Track 809 25.Jan.–8.Feb.1987 A.Newton &amp; M.Thayer \ wet scler. forest FMHD #87-213 flight interc. (window) trap [body in poor shape]; 1♂: 38.43S 145.35E VIC Otway NP 390m Binn Rd 4.3 km N of Cape Horn 808 25.Jan.–8.Feb. 1987 A.Newton &amp; M.Thayer \ wet scler. forest FMHD #87/210 flight interc. (window) trap [fore body missing] (all ANIC).</p><p>Habitus. BL 2.7−2.8mm, BL/BW ~1.7. Very similar to N. isolaeregis in all respects.</p><p>Male. Similar to N. isolaeregis, trigonium a little stouter. The parameres are longer than the parameroids and very slender. Their medial edge is nearly straight, smooth, the outer edge bears one to a few angular projections near the apex (Fig. 83).</p><p>Female. Unknown.</p><p>Notes. The holotype from near Canberra has a few teeth on the paramere, the specimens from the Otway ranges only one tooth (Fig. 91). The present gap in the range may be due to the scarcity of specimens. If correct, N. amphora probably coexists with N. alces n. sp.</p><p>Etymology. The name is a noun in apposition describing the shape of the penis.</p></div>	https://treatment.plazi.org/id/039F8D3EFF9DFFC696964225FBC3F8B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF9DFFC696964513FC8CFB1F.text	039F8D3EFF9DFFC696964513FC8CFB1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon isolaeregis	<div><p>Nothocyphon isolaeregis, n. sp.</p><p>(Figs. 82, 86, 91)</p><p>Type material: 1 ♂ holotype, 1 ♂ paratype: Helodes (Cyphon) ovensensis Blkb. King Island 10553 \ King I. TAS. Lea (SAMA). Additional paratypes: 5♂, King Island, TAS. Lea \ Helodes ovensensis, Blackb. (SAMA). 1♂ (genitalia naturally everted): King Isl. coll [illegible] 12.06 \ Helodes (Cyphon) ovensensis Bl. King Island [on reverse side:] A. M. Lea coll 12 0 6 \ Cyphon fenestratus Id. by J. Armstrong \ det fenestratus (MV, T-21999).</p><p>Habitus. BL 2.6−2.8mm, BL/BW ~1.7, slender. Light brown, head darkest, humeral area a little paler than rest. One specimen has a faint curved macula on the elytron like N. donnabuangi has. The head punctures are fine, dense, and rough, the integument appears scaly. Pronotal punctures little larger, granular, spaced, integument shiny. Elytral punctures normal, much larger. Semi-erect short yellowish pilosity.</p><p>Male (Figs. 82, 86). Pala truncate and parallel-sided. The parameroids are of equal width over their entire length. They curve around the trigonium, their apex is scooped out, spoon-like. The flat trigonium is longer than the parameroids, it resembles a bottle: base about three times wider than bottleneck, apex again wider, truncate or indistinctly concave, with a few rough scales along edge.</p><p>The separate flat parameres have a long basal rod and a short, caudally toothed plate. Left and right paramere of a given specimen often differ in details (Fig. 91).</p><p>Female. Unknown.</p><p>Etymology. Endemic to King Island. The name is a noun in apposition, an agglutination of two Latin nouns in the genitive case meaning “from the island of the king“.</p></div>	https://treatment.plazi.org/id/039F8D3EFF9DFFC696964513FC8CFB1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF9FFFC49696458CFC6BFBF8.text	039F8D3EFF9FFFC49696458CFC6BFBF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon alces	<div><p>Nothocyphon alces, n. sp.</p><p>(Figs. 85, 91)</p><p>Type material: 1♂ holotype: 37.44S 145.43E VIC Warburton 2.2 km NE on Acheron Way 320m 27.Feb,1993 931 A.Newton &amp; M.Thayer \ Euc.regnans forest w. Acacia dealbata pyr. fogging old fungusy logs (ANIC).</p><p>Habitus. BL 2.4mm. BL/BW ~1.7. Very similar to N. isolaeregis and N. amphora .</p><p>Male. Similar to N. isolaeregis, but the pala is anteriorly narrowed and the trigonium is apically not widened. The parameres are short and wide, medial portion with some large teeth, outer edge drawn out into a large triangular tooth with some small teeth on its base.</p><p>Female. Unknown.</p><p>Etymology. The strong teeth on the plate of the paramere make the ensemble resemble the antlers of a moose. The scientific name of that huge deer, Alces, is used as a noun in apposition.</p></div>	https://treatment.plazi.org/id/039F8D3EFF9FFFC49696458CFC6BFBF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF9FFFC496964795FDF1FDF1.text	039F8D3EFF9FFFC496964795FDF1FDF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon pacificus	<div><p>Nothocyphon pacificus, n. sp.</p><p>(Figs. 84, 91)</p><p>Type material: 1♂ holotype: NSW New England NP Thungutti Camp 22/11/01 CHS.Watts (SAMA); 1♂ paratype: NSW nr Nungatta Creek 58 88083N 557 20893E 13.2.0 0 \ J&amp;R Miller (SAMA). Habitus. BL 2.4−2.6 mm, BL/BW ~1.7. Very similar to N. isolaeregis and N. amphora . Male. Penis relatively slender, pala waisted, narrow bottleneck, apex strongly widened and shallowly excised. The parameres are longer than the parameroids and slender, inner face with a strong curved tooth (which differs in details of shape between the specimens), outer face long and slender, externally serrate (Fig. 84). Female. Unknown.</p><p>Etymology. The adjectival Latin (peaceful) name refers to the relative proximity of the known localities to the Pacific Ocean, but the sites are not coastal.</p></div>	https://treatment.plazi.org/id/039F8D3EFF9FFFC496964795FDF1FDF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF9FFFC596964378FF44F87A.text	039F8D3EFF9FFFC596964378FF44F87A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon armstrongi	<div><p>Nothocyphon armstrongi, n. sp.</p><p>(Figs. 87−91)</p><p>Type material: holotype 1 ♂: Orange [33.2842S, 149.1024E] N.S.W. J. Armstrong (ANIC). Paratypes: 19 ♂, 1♀ with the same data (all ANIC). 1♂: Cyphon fenestratus Bl. N.S. Wales \ Mt. Victoria [33.5907S, 150.2550E] N.S.W. Lea (SAMA).</p><p>Habitus. Very similar to other members of the group, contour regularly elongate oval, BL 2.5−2.7mm, BL/BW ~1.6. Body and appendages ochre to light brown, occiput and middle of pronotum darker, humeral area and sutural interval a bit lighter than rest, pilosity pale. The granular punctures on the head are very fine and dense, those on the pronotum a little coarser but wide-spaced, the integument is shiny. The normal punctures on the elytra are much larger, moderately dense, integument shining. Antennae slender, flagellar segments more than twice as long as apically wide.</p><p>Male (Figs. 87, 88). Penis and parameres slenderer than in other members of the group, trigonium with longer and narrower caudal portion, apex at best insignificantly widened. Parameroids much shorter than trigonium. Parameres in the form of two separate ventrolateral sclerites with slender basal rod. The plate-like part ends in a long straight dorso-lateral process, the blunt medial corner bears on its upper side a short oblique tooth.</p><p>Female. Last exposed sternite (S7; Fig. 89) with sparse pilosity, except along edges. Two oblique oval paramedian shallow depressions have fine pores and canals. A small group of distinct pores is seen near the mediocaudal end of each depression. T8 and ovipositor unmodified, rods of S8 anteriorly connected by a sclerite ring.</p><p>The prehensor (Fig. 90) is located in a wide anterior section of the oviduct and includes a pair of elongate sclerites which are medially bare but on the outside beset with large curved hooks. Around these spiny sclerites the integument is brownish, with a variety of scale- or spine-like minute acanthae. Further in front, on the opposite face of the duct, lies a thin oval sclerite ring with some small teeth along the sides. The surface of the large bursella is densely covered with minute circular areas surrounded by fine radial plicae.</p><p>Note. Occurs further inland than the other species in the group; there is no habitat information for any of them.</p><p>Etymology. Named in memory of the collector, the late J. Armstrong of Nyngan, NSW. His interest in Scirtidae is documented by a single publication (Armstrong 1953) and by numerous “ Cyphon -like” specimens he collected, identified, or even designated as types of new species—but none of these was ever described. Unfortunately, J. Armstrong relied solely on habitus characters. Accordingly, many specimens were misidentified, the intended type series sometimes included several different species. Armstrong's labels never provide details of collection, rarely a collecting date. He mounted many specimens in groups on pieces of card. For example, the present specimens [plus 5♂ of Austrocyphon pictus (Blackburn)] had been glued onto 3 cards, together on a single pin.</p></div>	https://treatment.plazi.org/id/039F8D3EFF9FFFC596964378FF44F87A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF98FFC3969645FDFCB5F9BD.text	039F8D3EFF98FFC3969645FDFCB5F9BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon patruelis	<div><p>Nothocyphon patruelis, n. sp.</p><p>(Figs. 98, 99)</p><p>Type material: holotype 1♂: Cradle Mtn. Tasmania Carter &amp; Lea \ on King Pine [handwritten] (SAMA). Paratypes: 7♂: Cradle Mtn. Tasmania Carter &amp; Lea; 1♂: TAS Cradle Valley Cradle Mountain-Lake St Clair N Pk 19/1/00 C.H.S.Watts (SAMA, 1 in PZ). 5♂: Cradle Mtn. Tasmania J.Armstrong; 1♂: Henty R. Tasmania J.Armstrong; 1♂: Strahan Tasmania J.Armstrong [labelled as paratype of a new species which J.Armstrong never published] (all ANIC).</p><p>Habitus. Very similar to N. frater but more slender, BL 2.2–2.6 mm, BL/BW ~1.9.</p><p>Male. Differs from N. frater by a well developed S8 with many setae. The penis (Fig. 98) is more slender, the pala resembles an elongate rectangle. It is longer than the parallel parameroids which diverge slightly and then narrow to slender tips located in continuation of the straight median edge of the parameroids. The median slit divides the parameroids forward up to the transverse bridge where the very small sclerotized triangular trigonium stands. In side view it lies mostly flat on the parameroids but in one specimen the trigonium stood at right angle to the penis axis, it is apparently movable. Tegmen with a triangular bare sclerotized area between the membranous lobe and the spinule group which is composed of numerous very fine spinules (Fig. 99).</p><p>Female. Unknown.</p><p>Notes. N. patruelis seems to be the most primitive of the three siblings. Its trigonium remains next to the transverse bridge at the caudal end of the pala. In N. frater and N. amita the trigonium shifts back onto the parameroids which probably excludes all movement. S8 is well sclerotized and large.</p><p>Nothocyphon patruelis and N. frater both occur in Tasmania. Despite their similarity they are probably different biospecies. N. patruelis is known from the area between Cradle Mt., Strahan, and tributaries to Lake Margaret while N. frater is widespread on the island. Both species were taken at Strahan.</p><p>Etymology. The name alludes to the close relation between this species, N. frater and N. amita: Latin patruelis designates a cousin from father's side; a noun in apposition.</p></div>	https://treatment.plazi.org/id/039F8D3EFF98FFC3969645FDFCB5F9BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF98FFC1969641BDFE8EF817.text	039F8D3EFF98FFC1969641BDFE8EF817.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon frater (Blackburn 1892) Blackburn 1892	<div><p>Nothocyphon frater (Blackburn, 1892), new combination</p><p>(Figs. 4−7, 92–97, 171, 172)</p><p>Helodes ( Cyphon ?) frater Blackburn, 1892: 520 .</p><p>Type material: Redescribed by Klausnitzer (1981), the actual specimen in the Natural History Museum, London, was not examined for this study. The penis of the type is illustrated in Klausnitzer (1981: fig. 16). In the interest of a stable nomenclature, this specimen is here designated as Lectotype (ICZN, Article 74.4). The label is “Blackburn coll., 1910/236, Type ” (Klausnitzer 1981). For the probable type locality see the Notes section below.</p><p>FIGURES 92–100. Nothocyphon frater- group, males. N. frater: 92, T8; 93, S8; 94, T9; 95, specimen from Dilgry R., penis, tegmen and S9, superimposed; 96, penis and tegmen, superimposed, specimen from Little Yarra R.; 97, armature of paramere.— N. patruelis: 98, penis and tegmen, superimposed; 99, armature of paramere.—100, N. amita, penis and tegmen, superimposed. 92−96, 98, 100, and 97, 99 to same scales, respectively.</p><p>Other material examined: VIC: 1♂: Dandongadale, 11.1.55, Neboiss \ Neboiss det.? ovensensis; 1♂: Tanjil Bren, Vic. 22.3.1958 A. N. (all MV). 1♂: Nat. Mus. Victoria G.French's coll. 5.11.08; 2♂: VIC 10K E Warburton 14/1/87 C.H.S.Watts; 7♂: VIC 5km NW Marysville 2/12/98 C.H.S.Watts; 1♂: VIC Panmure Mt Emu Creek 30/5/ 0 5 C.H.S.Watts; 1♂: VIC Fisherman's rest 6/11797 C.Watts (all SAMA). 8♂: VIC, Jamieson R., 29.12.1972; 2♂: VIC, Ovens R. 2 mls downstream from Harrietville, 23.11.1972; 2♂, 2 probable ♀: VIC, Little Yarra R. nr. Yarra Junction, 7.10.1972; 1♂: Delatite R. 7.7 mls downstream from Mirrimbah, 24.11.1972; 1♂: VIC, Dandenong ranges, Monbulk, 29.11.1972; 1♂: VIC, Royston R., 28.12.1972; 1♂: VIC, Mt. Buffalo, Eurobin Falls, 24.11.1972; 1♂: VIC, Pretty Valley, Mt. McKay, 7.1.1971; 2♂: VIC, Steavenson R. near Buxton, 28.12.1972 (all leg. P.Zwick, PZ).— NSW: 1♂: NSW, 3km N Lansdown, nr Taree G.Williams 13.Sept.1991 \ ex Acradenia euodiifolia blossoms; 1♂: Dorrigo N.S. Wales; 1♂: 31.53S 151.32E Barrington Tops Dilgry R. NSW 26.Nov.1985 C.Reid on Tree Ferns &amp; Nothofagus; 1♂: Adaminaby-Kiandra Rd., NSW, 27.1.66, E.Britton; 1♂: 36.25S, 148.22E, NSW, Kosciusko NP, Betts Ck NE 833 1740m 14. Feb.1987, A.Newton &amp; M.Thayer \ Alpine meadow FMHD #87-267 Sphagnum at stream edge and in bog (all ANIC). 1♂: National Park—Lea \ 8809 Cyphon N.S. Wales; 1♂: NSW 15 K W Jindabyne Little Thredbo River 6/1/06 on Leptospermum flowers C.H.S.Watts; 1♂: NSW Dead Horse Gap 4.5 km W Thredbo 6/1/06 C.H.S.Watts; 8♂: NSW Thredbo River 6 K N Jindabyne 6/1/06 on Leptospermum flws C.H.S.Watts (all SAMA). 1♂: NSW, Geehi R. on Alpine Way, 24.11.2005, P.Zwick (PZ).— TAS: 1♂: Helodes (Cyphon) ovensensis Blkb. King Island; 1♂: 41.36S 145.41E, TAS, Murchison Hwy/Que R. Mine Rd. 680m, 907, 11–27 Jan.1993 A.Newton &amp; M.Thayer \ Noth. cunn. rainforest FMHD #93-22 window trap (all ANIC). 1♂: King Island; 4♂: Arthur Plains S.W. Tas 3.2.1965 Neboiss; 2♂: Lake Dobson, Tas. 20. Feb. 1967 A. Neboiss (all MV). 1♂: West Tamar \ 20/10/94 [1894! on reverse side]; 1♂: Hobart Tas. Lea; 1♂: Helodes (Cyphon) ovensensis Blkb. King Island 10553 \ King I. Tas. Lea; 1♂: King Island, Tas., Lea; 1♂: Strahan, Tas. Lea &amp; Carter; 4♂: TAS Dove Lake Cradle Mtn—Lake St. Clair N. Pk. 19/1/00 C.H.S.Watts; 3♂: TAS Black River 5km NW Mawbanna 27/11/00 C.H.S.Watts (all SAMA). 1♂: TAS, Ellendale, 4.12.1972; 1♂: Australia, TAS, Wedge R., 5.12.1972; 1♂: TAS, Guide R., 2.12.1972; 1♂: TAS, Sassafras Ck nr Mole Ck, 6.12.1972; 6♂: TAS, Clarence R., 4.12.1972 (all leg. P.Zwick; PZ).</p><p>Habitus. Elongate, anterior two thirds of elytra almost parallel, body in side view rather flat, not domed (Figs. 171, 172). BL 2.1–2.4 mm, BL/BW ~1.7. Mature specimens are uniformly dark brown, also legs and antennae. Antennal segments 2 and 3 of subequal length, flagellar segments slightly conical, little more than twice as long as apically wide, terminal segment little longer (Fig. 4).</p><p>Male. Apodemes of T8 shorter than the truncate plate. S8 weak, pale sclerites in Y-like arrangement. T9 represented by straight apodemes continued backward along the segment's side edge, eventually curving slightly mediad. S9: two large lobes beset with long setae. S9 is laterally quite firmly attached to the sides of T9 and lacks a sclerotized base (Figs. 92–94). Penis (Figs. 95, 96) with narrow oval pala and much wider plate-like parameroids separated by a fine median slit, never divergent. The widest point is near the base. The parameroids are almost parallel, in the caudal fifth narrowed to blunt tips. A stripe along the slit is more strongly sclerotized than the rest and connected to a faint curved line delimiting a slightly more sclerotized apical area with few sensory pores. The short bifid sharply pointed trigonium stands at the start of the slit, basally from the widest point of the parameroids. The ring-shaped tegmen supports a simple ear-shaped membranous lobe and bears a few (ca 5–7) short forward pointing spines on the arched sclerite caudally from the membranous appendage (Fig. 97).</p><p>Female (presumed, not figured). T8 and ovipositor unmodified. S8 with narrow front end, the slender rods converge and lie side by side in the anterior section but do not meet. Prehensor and surface of bursella similar to N. amita, but sclerites more elongate and weakly sclerotized, spinules seem to be finer and sparser.</p><p>Notes. Blackburn apparently collected Austrocyphon ovensensis (Blackburn) and N. frater together and described them in succession, saying about H. frater “Victoria; also by sweeping on the banks of the Ovens”. Why he called the species frater (Latin, brother) is unknown. Klausnitzer's drawing (1981: fig. 15) misses the tegminal armature.</p><p>There are no definite females of N. frater but some taken together with males at Little Yarra River are most probably conspecific because they closely resemble the female of N. amita; see below.</p><p>Nothocyphon frater is common along the banks of fast flowing streams. N. patruelis and N. amita resemble it much but are regarded as separate biospecies in view of consistently different details of genitalia. Also, N. frater and N. patruelis coexist in Tasmania. In contrast, according to present knowledge, N. frater and N. amita are allopatric.</p></div>	https://treatment.plazi.org/id/039F8D3EFF98FFC1969641BDFE8EF817	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF95FFCF96964795FEDDFD4E.text	039F8D3EFF95FFCF96964795FEDDFD4E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon amita	<div><p>Nothocyphon amita, n. sp.</p><p>(Figs. 100–103)</p><p>Type material: holotype 1♂: [QLD] Cairns dist. P.E.Dodd (SAMA). Paratypes: 2 ♂, 2♀: QLD Zillie Falls 6 km E Millaa Millaa CHS.Watts 21/8/07 (reared, mounted from ETOH) (all SAMA). 1♂: QLD 13km NW Kuranda 6 Dec 1982 J.Doyen (ANIC). 1♂: Mt Lewis Road, via Julatten, NE Qld, 12 Oct. 1980, G.B.Monteith, Rainf. (QM Reg.No. T227411); 1♂: Mossman Bluff Summit 10km W Mossman N.Qld., 18 Dec 1988, 1300m, Monteith &amp; Thompson, Pyrethrum \ Trees &amp; rocks (QM Reg.No. T227412) (QMSB). 2♂: Australia: N.Qld, 40 km up Windsor T'land Rd 12.xi.1990 R.Storey &amp; S.DeFaveri and K. Halfpapp. 1♂: Australia: N.Qld, 40 km up Windsor T'land Rd 12.xi.1990 R.Storey &amp; S.DeFaveri and K. Halfpapp; 1♂: Tinaroo Dam N.Qld 10.xi.1982 L.J.Morgan (all QDPC).</p><p>Additional material examined: Several larvae, pupae, and exuviae: QLD Zillie Falls 6 km E Millaa Millaa CHS.Watts 21/8/07, in ethanol (SAMA).</p><p>Habitus. Smaller (BL 1.8–2.4mm) and mostly stouter than N. frater, with distinctly rounded sides of elytra (BL/BW ~1.5) but the smallest specimen (from Mossman Bluff Summit) is about 1.8 times longer than wide and has nearly parallel elytra. Brown, margins of pronotum and humeri a bit paler than rest.</p><p>Male (Fig. 100). Differs from N. frater and N. patruelis by the stouter and shorter penis, pala and parameroids are of similar length. The pala is wide at the base, much narrower caudally. The penis contour widens much where the parameroids originate whose sclerotized medial edges lie parallel to each other. Outer edges curved, first convex, then slightly concave, distal half convex, tips together forming an ovoid apex. In the widest area the outer edge is very delicate, with small irregular notches and lobes. Trigonium bifid, longer than wide and inserted well behind the transverse bridge. The median slit between the parameroids reaches to this point but seems to continue forward as a faint line. S8 Y-shaped, delicate.</p><p>Female (Figs. 101–103). Resembles male in habitus. T8 and ovipositor unmodified. The front of S8 is slender, tongue-shaped, the lateral rods converge in front until they meet. However, a fine separation line is visible. Shortly before the front end the rods separate again, their slightly widened ends lie in close proximity. The prehensor consists of two longitudinal sclerites in a bell-shaped widening of the gonoduct. Caudally each sclerite is covered with sharp triangular spinules, tips directed backward. The narrow front end is close to the bursella which is covered with approximately circular depressions of variable size, each with minute peripheral plicae.</p><p>Note. N. amita varies in body shape and slightly in trigonium shape. In the male from Mossman Bluff summit the trigonium appears conical, the tips are suppressed. In the paratypes from Zillie Falls the trigonium is a littler shorter than in Fig. 100 which shows the holotype; other males are similar. The specimens from Zillie Falls had been reared, conspecificity of sexes is certain and the distinctive females are assigned type status.</p><p>The same sample includes larval and pupal exuviae. The larvae agree with the description of N. frater (Watts 2014) . The pupa of N. amita has 4 long pronotal horns, like most scirtid pupae. Several pupal exuviae projected from the dorsal cleft of the larval exuviae in which they were anchored by a pair of small hook-organs (Zwick &amp; Zwick 2008), each with about 20 slender hooklets.</p><p>In N. frater, N. patruelis and N. amita, shape of pala and parameroids as well as point of insertion of the trigonium form North-South morphoclines. In the absence of intermediate morphs the allopatric N. frater and N. amita are believed to be separate biospecies. The genetic distance between the two (Cooper et al. 2014) is important.</p><p>Etymology. Latin amita designates father's sister, a noun in apposition alluding to the close relationship between the taxa.</p></div>	https://treatment.plazi.org/id/039F8D3EFF95FFCF96964795FEDDFD4E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF94FFCF969645D5FC25FA28.text	039F8D3EFF94FFCF969645D5FC25FA28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon soror	<div><p>Nothocyphon soror, n. sp.</p><p>(Figs. 104–108)</p><p>Type material: 1♂ holotype: 42.10S 146.07E 4 km SSE of Mt. Rufus 800m TAS 26–28 Jan 1980 Lawrence &amp; Weir (ANIC). Paratypes: 1♂: Strahan Tasmania J. Armstrong (ANIC). NSW: 1♂: Wentworth Fs. (SAMA). 1♂: Sydney \ K.K.S. \ Cyphon fenestratus Blckb. Sydney \ K249505 (AMS).</p><p>Habitus. Elongate oval, BL 2.4 mm, BL/BW ~1.6. Colour of upper side varies (probably with specimen age) from uniformly yellowish (the holotype) to head and centre of pronotum dark brown, remainder brown (specimen from Strahan).</p><p>Male. T8 unmodified, S8 not observed, of T9 only the slender apodemes are sclerotized (Figs. 104, 105). S9: base not noticed, caudally deeply and widely divided into two lobes with numerous straight setae (Fig. 106).</p><p>Tegmen and parameres form a strong sclerotized ring (Figs. 106, 108). The narrow anterior part lies over the penis, the wider caudal portion with spreading transparent wings and heavy dark knobs that meet medially lies beneath the penis. Penis (Figs. 106, 107) with long narrow pala which is anteriorly a little expanded. Penis width more than doubles where trigonium and parameroids originate. The trigonium is a large spade-shaped plate with gently serrate sides and shallowly excised caudal edge. The straight slender flat parameroids lie dorsally from the trigonium, only their bare externally rounded tips project.</p><p>Female. Unknown.</p><p>Notes and etymology. The flat penis, the ring-shaped tegmen and parameres and the position of the parameroids dorsally (instead of laterally) from the trigonium suggest a close relationship with N. frater . The name soror (Latin: sister) was chosen to underline this. The habitat of the rare species is unknown. It shares the wide range from TAS to NSW with its stream-inhabiting relative, N. frater .</p></div>	https://treatment.plazi.org/id/039F8D3EFF94FFCF969645D5FC25FA28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF94FFCD96964134FBE7FCF2.text	039F8D3EFF94FFCD96964134FBE7FCF2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon platyphallus	<div><p>Nothocyphon platyphallus, n. sp.</p><p>(Figs. 109–111)</p><p>Type material: 1♂ holotype: Mt. Wellington, Tas. 12.12.0 1 Griffith \ Griffith collection Id. by A.M.Lea \ Cyphon nr. adelaidae Bl. S.A. \ 2946 (SAMA). Paratypes: 1♂: Huon R. Tas Lea \ Cyphon 8802 Tasmania; 1♂: C.E.Cole Tyenna Tas. 6.– 7.12.17 /16 (both SAMA). 1♂: 41.11S, 148.00E, TAS, Mt. Michael 740m, 27.Nov.1989, sweeping Tube 202 M. Mitchell; 1♂: 41.20N, 147.50E, TAS, Mt. Victoria 900m, 25.Nov.1989, pyrethrin knockdown tree 1, Tube 237 R.Roy coll.; 1♂: 42.42S 147.52E TAS Big Sassy Creek 12 May 1989 Tube 137 pyrethrin knockdown Atherospermum J.Diggle rainforest \ ANIC Coleoptera Voucher No. 88-0316; 1♂: Frankford Tasmania (all ANIC).</p><p>Habitus. BL 2.9–3.4 mm. BL/BW ~1.7. Elongate oval, elytra about 1.8 times wider than the pronotum. Head, pronotum and scutellum with granular, fine punctation, surface shining. Elytral punctures simple, larger than on pronotum.</p><p>Colour varies from light brown to dark brown with lighter shoulders and a narrow reddish stripe along suture. Front corners and stripes along sides of pronotum lighter than disk. The specimen from Mt. Michael is the palest, with pale humeri and a long caudally narrowing pale parasutural stripe plus a pale fascia along the outer edge in the caudal half of elytra.</p><p>The prosternal process resembles a tennis racket, narrow base, wide oval plate. Despite this, the U-shaped long receiving mesoventral groove is narrow. It appears like the continuation of the narrow space between the paramedian crests on the declivous front of the segment.</p><p>Male. S8 is Y-shaped, weak, observed only by transparency of T8 (Fig. 109). Only apodemes of T9 sclerotized, S9 represented by two widely separate caudally pilose lobes (Fig. 110).</p><p>Penis elongate, with trapezoidal base (Fig. 111). Trigonium flat, leaf-shaped, with small serrations along the truncate tip. Where the trigonium is widest are two paramedian crests with a few forward pointing teeth. The narrow parameroids surround the trigonium apex but their apices do not meet. Tegmen a narrow sclerite forming a caudally open ring. Its separate widened ends represent the parameres, anterolaterally with a few strong teeth.</p><p>Female. Unknown.</p><p>Notes. This Tasmanian endemic shares the short ring-shaped tegmen with widened externally hooked ends as parameres with the other members of the group. However, N. platyphallu s is larger and differs by possessing a mandibular tooth, and by a wider end of the prosternal process. Two specimens were taken near rivers and streams, development may take place in flowing water.</p><p>Etymology. The Greek name describing the flat trigonium is a noun in apposition.</p></div>	https://treatment.plazi.org/id/039F8D3EFF94FFCD96964134FBE7FCF2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF96FFCD9696448EFBBBFBF9.text	039F8D3EFF96FFCD9696448EFBBBFBF9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon triangulum	<div><p>The triangulum -group</p><p>In the two Tasmanian species the penis is sub-parallel, only the front of the pala may be wide. The triangular trigonium is a little shorter than the simple club- or tongue-shaped parameroids and has only small spinules along its margins. Tegmen and parameres are short, the latter are armed with strong teeth.</p></div>	https://treatment.plazi.org/id/039F8D3EFF96FFCD9696448EFBBBFBF9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF96FFCD9696437EFA1AFAD8.text	039F8D3EFF96FFCD9696437EFA1AFAD8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon triangulum	<div><p>Key to males of the triangulum -group</p><p>1 Trigonium very wide, triangular, with straight sides. The blunt basal corners conceal the base of the parameroids (Fig. 112)....................................................................................... N. triangulum n. sp.</p><p>1' Base of parameroids exposed, trigonium narrow, without basolateral corners, sides a little convex (Fig. 114). N. radula n. sp.</p></div>	https://treatment.plazi.org/id/039F8D3EFF96FFCD9696437EFA1AFAD8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF96FFCA96964255FE14FABA.text	039F8D3EFF96FFCA96964255FE14FABA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon triangulum	<div><p>Nothocyphon triangulum, n. sp.</p><p>(Figs. 112, 113)</p><p>Type material: 1 ♂ holotype: Mt. Wellington Tas. Lea (SAMA). Paratypes: 1♂ with the same data as the holotype; 8♂: Cyphon Tasmania 8801 Summit of Mt. Wellington Tas. (Lea) (SAMA). 1♂: 42.41S 146.43 TAS, National park, low bushes entrance to NP, Feb. 1992, C.Reid (naturally everted genitalia; ANIC). 1♂: 42.40S 146.46E TAS, Mt. Field NP, Robert Tarn 1220m, 7 Feb 1992 C.Reid on flowers; 1♂: 42.40S 146.41E TAS 2.5km W Mt. Field NP c. 600m, 6. Feb.1992 C.Reid on Nothofagus cunninghamii; 1♂: Mt. Wellington TAS Lea Hobart Tasmania J.J.Walker (all ANIC); 1♂: Hobart Tasmania J.J.Walker \ G.C.Champion coll. B.M. 1927-409 (NHML).</p><p>Habitus. BL 1.6–1.8mm, BL/BW ~1.7. Elongate oval, dorsal side entirely brown, head darkest, the sutural interval may have a reddish tinge. The semi-erect pilosity is brownish. Head with fine, pronotum with slightly coarser dense granular punctation, normal dense punctures on elytra little larger.</p><p>Male. Segments 8 and 9 as for the genus, the pilose lobes of S9 large. The slender penis is waisted between the wide truncate front edge of pala and the transverse bridge where parameroids and trigonium originate. The trigonium is flat, triangular. The widest point is near the base where blunt lateral angles project over the base of parameroids. The convergent sides are straight and meet at a blunt angle of about 40°. Sides of triangle with small teeth. The finger-shaped slightly curved parameroids are a little longer than the trigonium, their apices almost meet behind the trigonium tip. The tegmen is a narrow sclerite bracket supporting the short parameres. Each is an elongate somewhat twisted sclerite with several large hooks along the sides.</p><p>Female. Unknown.</p><p>Note. Endemic to Tasmania. The holotype has partly extruded genitalia. The large lobes of S9 stand in vertical position laterally from the penis, like shields. The penis base is straight, the parameroids and the trigonium curve gently into opposite directions. Parameres not visible.</p><p>Etymology. The specific name describes the shape of the trigonium and is a Latin noun in apposition, triangulum, a triangle.</p></div>	https://treatment.plazi.org/id/039F8D3EFF96FFCA96964255FE14FABA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF91FFCB969642C1FA0CFED6.text	039F8D3EFF91FFCB969642C1FA0CFED6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon radula	<div><p>Nothocyphon radula, n. sp.</p><p>(Fig. 114)</p><p>Type material: 1♂ holotype: Australia, TAS, nr. Humbug Hill nr Binalong Bay 13.Febr. 1980, Euc. Casuarina woodland, A.Newton, M.Thayer \ pyrethrin fogging Eucalyptus bark (ANIC). Paratypes: 1♂: Australia Tasm. Hastings St. Res., nr Newdegate Cave 130m, 9.ii.1980, A.Newton, M.Thayer/ trapped in floating debris forest stream; 1♂: TAS, Fortescue Bay, Tasman Pen. wet scler. 7–9 II 1989 D.Bickel Malaise; 1♂: 43.07S 146.47E Edwards Rd TAS Hartz Mts 4.Feb.1983 I.D.Naumann J.C.Cardale coll. (all ANIC)</p><p>Habitus. BL 2.4–2.6mm, BL/BW ~1.85. The sides of the slender body are parallel over most of their length. The beetle is light to dark brown, rear of head and centre of pronotum darkest, appendages yellowish. Similar to N. triangulum .</p><p>Male. Segments 8 and 9 as for the genus. Penis slender and parallel, front of the pala little wider. Trigonium and parameroids occupy the caudal third of penis length. The trigonium is elongate, tongue-shaped, the gently curved sides and the blunt tip with small scale-like teeth. The slender parameroids are a little longer than the trigonium, narrowed to a rounded apex inclined mediad. The tegmen is a thin sclerite band that widens abruptly laterally and supports the wide obliquely triangular parameres provided with many coarse teeth.</p><p>Female. Unknown.</p><p>Note. A close relative of N. triangulum, both are endemic to Tasmania. They are easily distinguished by trigonium and parameres.</p><p>Etymology. The name radula (Latin, a rasp) is a noun in apposition describing the structure of the paramere.</p></div>	https://treatment.plazi.org/id/039F8D3EFF91FFCB969642C1FA0CFED6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF90FFCB969645F8FDA5FABC.text	039F8D3EFF90FFCB969645F8FDA5FABC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon thylacinus	<div><p>Nothocyphon thylacinus, n. sp.</p><p>(Figs. 115–120)</p><p>Type material: 1♂ holotype: Cradle Mtn Tasmania J. Armstrong (ANIC).</p><p>Habitus. An elongate-oval Nothocyphon . BL 2.7mm, BL/BW 1.7. Dark brown, punctation granular and fine on head and pronotum, spaced on pronotum, normal and dense on elytra. Flagellar segments twice as long as wide at apex.</p><p>Male. Segments 8 and 9 as in the generic description except that T8 (damaged) appears fairly narrow (Figs. 115–117). The tegmen is a narrow transverse arch supporting slender, blade-like parameres which carry some large teeth near midlength and are serrate along the edge (Fig. 120). The penis (Figs. 118, 119) has an anteriorly wide pala (damaged) which narrows to where the trigonium is inserted. The trigonium is a small sharply pointed triangle. Ventrally it is armed with sharp teeth standing in two irregular oblique rows. Some teeth are unusually large and project laterally across the parameroids. The gently arched parameroids are narrow and extend far beyond the trigonium. They lack spines, to their outside attach membranes (torn in the only specimen).</p><p>Female. Unknown.</p><p>Etymology. The large teeth on the trigonium reminded me of the fangs of vertebrate carnivores. Named after the largest marsupial predator, the Marsupial Wolf, Thylacinus, which survived the arrival of Man in Australia longer in Tasmania than elsewhere.</p></div>	https://treatment.plazi.org/id/039F8D3EFF90FFCB969645F8FDA5FABC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF90FFC8969642BCFEE6F838.text	039F8D3EFF90FFC8969642BCFEE6F838.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon sarcophilus	<div><p>Nothocyphon sarcophilus, n. sp.</p><p>(Figs. 121–123)</p><p>Type material: 1♂ holotype: SW TAS, Lower Gordon R., 42.56S 145.50E, 42.54S 145.54E, Howard, Hill \ H.E.C. Survey 12R 600–1000 Feb. 1978 beating \ 10 (ANIC).</p><p>Habitus. BL 2.4 mm, BL/BW ~1.7, elongate oval, similar to N. thylacinus .</p><p>Male. Segments 8 (Figs. 121, 122) and 9 (Fig. 123) as described for the genus. Penis (Fig. 123) parallel-sided, with rounded pala. The short trigonium inserts near the basal third. The contour of the trigonium resembles an inverted heart, its sides are thickened, the bulges bear huge teeth along the medial edge. The straight parameroid is much longer than the trigonium, the long bare apex bears externally a toothed sclerotized lobe pointing cephalad. In dorsoventral view the tips of the parameroid and of the paramere overlap. The paramere is short, slender and heavily toothed along the outside and distally. The tegmen is a thin sclerite arch connecting the parameres.</p><p>Female. Unknown.</p><p>Etymology. The trigonium reminded me of a strongly toothed lower jaw of a predator. This suggested the name: Sarcophilus, the Tasmanian Devil, the largest extant carnivore in Tasmania.</p><p>FIGURES 115–123. Nothocyphon thylacinus -group, males. N. thylacinus: 115, T8; 116, S8; 117, T9; 118, penis, dorsal; 119, trigonium, ventral; 120, tegmen and parameres.— N. sarcophilus: 121, T8, with details of grouped microtrichia on plate; 122, S8; 123, segment 9, penis and tegmen with parameres, superimposed, ventral view. 115–117, 118–120, 121–123 to the same scales, respectively.</p></div>	https://treatment.plazi.org/id/039F8D3EFF90FFC8969642BCFEE6F838	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF92FFD69696466CFE94FCB5.text	039F8D3EFF92FFD69696466CFE94FCB5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon armatus	<div><p>Nothocyphon armatus, n. sp.</p><p>(Figs. 124–127)</p><p>Type material: 1 ♂ h olotype, 1 ♀ paratype: Mt. Edith Forest Road, 1 km off Danbulla Road [QLD] 6.May 1967 D.H. Colless (ANIC); 1 ♀ paratype: Mt. Edith 4– 7 M. off Danbulla rd. N.QLD 27.Apr. 1967 D.H. Colless (ANIC). 1 ♂ paratype: Baldy Mtn road SW of Atherton N.Qld, 1000m, 10 Oct 1980, G. Monteith, Pyrethrum, rainforest (QMSB; QM Reg.No. T227418).</p><p>Habitus. BL 2.1–2.2mm, BL/BW 1.5. Oval, rather convex. Dorsal side yellowish-orange to brown, elytra with diffuse pattern of variable intensity. A narrow strip along the base of the elytra on either side of the dark scutellum is pale. The strip surrounds the humerus on the outside, along the edge of elytron. Sutural interval also pale but interrupted by brown at midlength from where a brownish arch may extend forward to near the humerus. Appendages yellowish. Lower face and mouthparts as described for the genus. Flagellar segments a little conical, barely twice as long as wide at apex, last segment oval.</p><p>Male (Figs. 124–126). Segments 8 and 9 as described for the genus. Penis elongate, pala broadly rounded, almost truncate. The penis widens just caudally from midlength at the bases of trigonium and parameroids. The parameroids are flat, with blunt ends curving around the shorter trigonium. The trigonium is a simple cone supporting a crown of several strong teeth. The most basal teeth are the largest, sickle-shaped, and curve outward and ventrad. The tegmen is a transverse bar which supports short rod-like parameres ending in several very large straight teeth.</p><p>Female (Fig. 127). T8 and ovipositor unmodified. Rods of S8 very long, in front connected by a wide semicircular sclerite (not shown). The prehensor is elongate, anteriorly with a sclerite loop and two divergent scaly bands. Caudally from this lies a pair of narrow sleeves ending in dark, scaly pockets, plus a smaller pocket laterally from each. Caudo-laterally are two finely scaly bands.</p><p>Note. Habitus, pigmentation and female genitalia resemble N. lindensis . However, the male genitalia differ much. For example, they lack the median hollow spine or spinule patches ventrally from the penis which N. lindensis and relatives have. The parameres are very different.</p><p>Etymology. The Latin adjective armatus, armed, alludes to the large spines on the trigonium and the parameres.</p></div>	https://treatment.plazi.org/id/039F8D3EFF92FFD69696466CFE94FCB5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF8DFFD6969644D6FBFFF95D.text	039F8D3EFF8DFFD6969644D6FBFFF95D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon brevihamatus	<div><p>Nothocyphon brevihamatus, n. sp.</p><p>(Figs. 128–131)</p><p>Type material: holotype ♂: New England Nat. Park, NSW, rainforest below Wright's L'kout, 11.x.66 E.Britton (ANIC).</p><p>Habitus. Elongate oval, BL 2.3mm, BL/BW ~1.7. Light brown, legs yellowish brown, flagellar segments 2–8 infuscate. The dorsal punctation is fine, with blond semi-erect pilosity. The antennae are slender, the distal flagellar segments are cylindrical, about twice as long as wide, tip of last segment blunt.</p><p>Male. Segments 8 and 9 as described for the genus (Figs. 128, 129). Tegmen and parameres together form a thin rectangular frame (Fig. 130). The straight parameres are long. The thin sclerites of the tegmen widen abruptly and form a triangular lobe caudally from which the long straight parameres narrow continuously. They end as gently inwardly curved rods. Penis (Fig. 131) with oval pala supporting long thin inwardly curved parameroids with asperous tip. On their medial side they are excised in the area occupied by the short trigonium. Its basal arms are connected by a transverse bridge from which originate two parallel rods with rectangularly downcurved sharp tip.</p><p>Female. Unknown.</p><p>Note. The unusual shape of the genitalia, especially the profoundly divided trigonium reminded me of Petrocyphon species. However, the animal's body structure agrees with genus Nothocyphon . No similar species is known.</p><p>Etymology. Latin brevihamatus translates as short hooked; the name is an adjective.</p></div>	https://treatment.plazi.org/id/039F8D3EFF8DFFD6969644D6FBFFF95D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF8DFFD4969641E8FBEEFD07.text	039F8D3EFF8DFFD4969641E8FBEEFD07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon crux	<div><p>Nothocyphon crux, n. sp.</p><p>(Figs. 132–136)</p><p>Type material: holotype ♂: 30.30S 152.23E NSW Thungutti Camp New England N.P. 16–18.Nov.1990 A.Calder (ANIC).</p><p>Habitus. BL 3.4 mm, BL/BW ~1.8. Elongate, parallel, brown, elytron with 3 weak costae. The punctures on head, pronotum and scutellum are very fine and indistinctly granular. The normal elytral punctures are larger but still fine and dense. The yellowish-brown pilosity is semi-erect. Overall body shape, mouthparts (mandibles not examined), head sutures and ventral face of thorax as described for the genus. The antennal base is unmodified, the flagellar segments are slender, indistinctly conical, about 2.5x as long as wide.</p><p>FIGURES 128–136. Nothocyphon spp., males. N. brevihamatus: 128, T8 and S8; 129, T9 and S9, superimposed; 130, tegmen and parameres; 131, penis, ventral view. 128, 129 and 130, 131 to the same scales, respectively.— N. crux: 132, S8; 133, T9; 134, S9; 135, penis; 136, tegmen (broken) and parameres. 132−134 and 135, 136 to the same scales, respectively.</p><p>Male. Segment 8 (Figs. 132–136; T8 broken, not shown) unmodified. The thin apodemes of T9 are connected by a forward-bent fine sclerite and continue for a long distance before they taper away on the membranous plate. From their diffuse ends extend two arched lightly pigmented areas to the anterior connecting sclerite. S9 is broad, composed of two short lobes separated by a narrow membranous strip over their entire length, each half with a vaguely sclerotized outwardly curved short apodeme and apical setae.</p><p>The penis (Fig. 135) is elongate, its strongly sclerotized components are arranged in cross-shape. From an almost circular soft pala with a median line of weakness originate two strong sub-parallel rods. In the distal fourth they are briefly curved outward and then converge as the edges of a short cordiform trigonium with very fine denticles around its tip. A strong outward directed thumb projects near the base of the trigonium. The parameroids are short approximately triangular platelets which are little longer than the trigonium. The tegmen (Fig. 136) is a thin lace supporting the parameres which are elongate short sclerites, each with 2 or 3 large hooks that are directed obliquely outward and forward.</p><p>Female. Unknown.</p><p>Note. N. crux has unique genitalia, no similar species is known. The genital preparation was unfortunately damaged and several parts are broken. The figures are partly based on photographs taken before the accident.</p><p>Etymology. The Latin noun, crux, a cross, describes the overall shape of the penis.</p></div>	https://treatment.plazi.org/id/039F8D3EFF8DFFD4969641E8FBEEFD07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF8FFFD49696441DFE5EF8ED.text	039F8D3EFF8FFFD49696441DFE5EF8ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon esau	<div><p>Nothocyphon esau, n. sp.</p><p>(Figs. 137–142)</p><p>Type material: NSW. 1♂, holotype: Kosciusko Nt.Pk Batt's Creek 1750m SW pitfall traps 1983–1986 Ken Green No. 39 (ANIC). Paratypes: 1♂ with the same data as the holotype; 1♂: Kosciusko Nat.Pk Batts Creek NSW 1700m 1983–1986 sweep sample Ken Green No.41 (both ANIC).</p><p>Habitus. BL 2.8–3.1mm, BL/BW ~1.75. Body structure as described for the genus. An elongate-oval flat beetle, sides of elytra almost parallel. The light brown pilosity is erect, the animal appears quite hairy. There is a colour gradient from the dark brown head to the yellowish-brown apical portion of elytra. The punctation on head and scutellum is dense, fine and rough, that of the pronotum is granular and rather coarse. The normal elytral punctures are large near the base but grow finer and denser caudally. The antenna is slender, the flagellar segments are cylindrical and a little more than twice as long as wide.</p><p>Male. Segment 8 as for the genus (Figs. 137, 138). T9 (Fig. 139) is bare, the strong apodemes are connected by delicate struts. The plate is soft and colourless except the caudolateral angles which are slightly pigmented. Of S9, only the pilose caudal lobes were observed (not shown).</p><p>Penis (Fig. 140) with deeply excised pala. Its lateral sclerites are straight convergent strong rods. They continue caudad and form the parameroids. Each ends in a flat, tongue-shaped sclerotized tip and supports a subterminal recurved outer lobe which is armed with fine spinules and teeth on the outside. The area between the strut and the medial lobe is transparent, window-like. The basal arms of the trigonium are attached to the lateral rods. The trigonium is very short, X-shaped, caudally with irregular teeth (Figs. 140–142). The tegmen is a thin long sclerite band which turns into strong longitudinal rods. Caudally, each rod ends in a short thick thumb-like sclerite resting beneath the hyaline window of the parameroid. Caudo-medially, each paramere bears a membranous bare lobe (Fig. 140, pel).</p><p>Female. Unknown.</p><p>Note. An exceptional species. There is no visible limit between the lateral sclerites of the pala and the parameroids, and the trigonium is exceptionally stout and short. Details of its shape and the arrangement of its caudal spines vary slightly (Figs. 140–142). The shape of the parameres is peculiar. No similar species is known.</p><p>Etymology. The name was taken from the Old Testament: Esau (Hebrew: hairy) was the very hairy brother of Jacob. A noun in apposition.</p></div>	https://treatment.plazi.org/id/039F8D3EFF8FFFD49696441DFE5EF8ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF89FFD296964795FB15FB9C.text	039F8D3EFF89FFD296964795FB15FB9C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon horridus	<div><p>Nothocyphon horridus, n. sp.</p><p>(Figs. 143–148)</p><p>Type material: 1♂, holotype, 3♂, paratypes: 37.43S 145.41E VIC Mt Donna Buang 1200m N of Warburton 811 26.Jan–11.Feb. 1987 A.Newton &amp; M.Thayer \ wet scler. Noth. cunn. FMHD #87 216 flight intercept (window) trap (ANIC).</p><p>Habitus. Oval, BL 2.5–2.6mm, BL/BW 1.6. Brown, head and centre of pronotum dark brown, margins diffusely reddish. Elytra with narrow dark brown sutural seam from the scutellum to near elytron midlength where the band curves outward and forms an anteriorly open semicircle ending at the elytral margin. The granular punctures on the forebody are fine, dense on the head, widely spaced on the pronotum. The dense normal punctures on the elytra are larger but comparatively fine. The semi-erect pilosity is light brown. The short antenna reaches the basal fifth to sixth of the elytron. The base is unmodified, the distal segments are about twice as long as wide.</p><p>Male. Middle of last sternite projecting a little, contour angular. Segments 8 and 9 as for the genus (Figs. 143– 146), except the Y-shaped S8 unusually large. Base of S9 not visible. Tegmen and parameres (Fig. 147) exceptionally strongly sclerotized, of unusual shape. Two separate basally slightly scooped curved sclerites converge, meet, and then widen and diverge again. The short apical portion of each bears 3 massive teeth between which rise fine hair-like structures. At the narrow meeting point each paramere has a medial extension connected to its counterpart by a membrane with some anteriorly directed paramedian spine-like sclerite. At this point they seem to be connected to membranes around the penis, details are unknown.</p><p>Penis (Fig. 148) long, slender, with parallel sides. The front of the pala is anteriorly shallowly excised and almost twice as wide as the caudal portion. The parameroids and the trigonium originate in the distal third. The trigonium is strongly sclerotized, from a transverse bracket-like base originates a long slender gently downcurved tip. The narrow apically slightly spatulate parameroids are straight and longer than the trigonium.</p><p>Female. Unknown.</p><p>Note. The species was taken together with N. donnabuangi and N. imitator whom it resembles in the brown elytral band. It differs by the very fine widely spaced pronotal granula, and of course by the exceptional genitalia.</p><p>Etymology. Latin horridus, awful, an allusion to the powerful claw-shaped parameres.</p></div>	https://treatment.plazi.org/id/039F8D3EFF89FFD296964795FB15FB9C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF89FFD3969643A8FCD7F869.text	039F8D3EFF89FFD3969643A8FCD7F869.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon longiphallus	<div><p>Nothocyphon longiphallus, n. sp.</p><p>(Figs. 149–155)</p><p>Type material: 1♂ holotype: “Lam.[ington] Nat. Pk., QLD. 28 Jan–3 Feb., 1963 G. Monteith” (QMSB, UQIC Reg. #88655).</p><p>Habitus. BL 2.6mm, BL/BW ~1.5. Broadly oval, flat. Dark brown, legs a little lighter. Structure as described for the genus. Both mandibles with a tooth, the one on the right side larger. Terminal segment of maxillary palpus missing. Flagellomeres barely twice as long as wide, widened towards apex, segments 10 and 11 missing.</p><p>Male. Segments 8 and 9 as for the genus, except S9 small, with two rounded pilose lobes. Basal portion narrow, with unpaired small apodeme (Figs. 149–152). Penis and tegmen thin and unusually long, curving ventrad. Penis with narrow elongate scoop-shaped pala (Fig. 154; very base missing). It supports an extremely short trigonium resembling an open mussel (Fig. 155), and rod-shaped, gently curved parameroids which are several times longer than the pala. Tegmen consisting of two long blade-like sclerites (Fig. 153). Base damaged, some connection between them may be missing. At about 1/3 of total length, each sclerite is on its medial face armed with a flat curved toothed appendage. Parameres with fine sensory pores, especially near the end.</p><p>Female. Unknown.</p><p>Note. The type is in poor shape. It had been pinned with a minuten pin and was slightly mildewed; it fell to pieces during study. The fragments are mounted on a card and 2 plastic slides on the same pin. Nothocyphon brevihamatus from the New England area in New South Wales has similarly long parameroids but in view of marked differences in other parts it is doubtful if it can be regarded as a close relative of N. longiphallus .</p><p>Etymology. The name describes the extraordinary length of the phallic complex.</p><p>FIGURES 149–155. Nothocyphon longiphallus, male. 149, T8; 150, S8; 151, T9; 152, S9; 153, tegmen and parameres; 154, penis; 155, detail of the trigonium. All to the same scale, except 155.</p></div>	https://treatment.plazi.org/id/039F8D3EFF89FFD3969643A8FCD7F869	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF8BFFD096964795FE98FBC5.text	039F8D3EFF8BFFD096964795FE98FBC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon naso	<div><p>Nothocyphon naso, n. sp.</p><p>(Figs. 156–159)</p><p>Type material: 1 ♂, holotype, Mossman Bluff Track 5–10km NW Mossman, N.Qld., 17–31 Dec 1988, Monteith, Thompson &amp; ANZSES, Site 9, 1260m, flt. intercept (QM Reg.No. T227420); 1 ♂, paratype: Mossman Bluff 10km NW Mossman, N.Qld., 17–18 Dec 1988, 1100–1300m, Monteith &amp; Thompson (QM Reg.No. T227419) (QMSB).</p><p>Habitus. BL 2.4mm, BL/BW ~1.6, elongate-oval, fairly convex. The pattern resembles N. lindensis, the dorsal side is yellow with a brown parasutural spot. Antennae short, flagellar segments slightly conical, not even twice as long as wide at apex. The punctation of the forebody is very fine, on the pronotum essentially only the spaced setal insertions are visible. Punctures on elytra normal, fine, separated by twice their own diameter, the yellow pilosity rather apparent.</p><p>Male. Segments 8 and 9 as for the genus (Figs. 156, 157). The penis is slender, the front end of the pala is rounded (Fig. 158). The trigonium inserts just beyond midlength. It is carrot-shaped with wide base narrowing in almost straight line, apex pointed. Raised cones stand in an irregular row along the side, the basal cones are largest, tooth-like and reach over the edge, growing finer towards the tip. The parameroids are closely appressed to the trigonium, straight, with obtuse tip and large outwardly directed triangular process near midlength. Immediately basally from trigonium and parameroids occur two crescent-shaped pale sclerites that are laterally connected to a slender forward-pointing strut. The parameres are connected by a strong arched bar-like tegmen (Fig. 159). Each paramere is a strong rod which widens abruptly a short distance from the base and is divided into 2−3 very large rigid spines and an elongate swelling beset with numerous teeth along its medial face. The largest teeth occur near the base of the swelling.</p><p>Female. Unknown.</p><p>Note. Resembles N. lindensis, N. ypsilon as well as N. armatus in habitus, but not in the genitalia.</p><p>Etymology. The name (the Latin noun naso, the nose) alludes to the nose-like triangular process on the parameroid.</p></div>	https://treatment.plazi.org/id/039F8D3EFF8BFFD096964795FE98FBC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
039F8D3EFF8BFFD196964360FE91F83A.text	039F8D3EFF8BFFD196964360FE91F83A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nothocyphon plicatus	<div><p>Nothocyphon plicatus, n. sp.</p><p>(Figs. 160–164)</p><p>Type material: 1♂, holotype: South Ramshead NSW Kosciusko NP 1850m subalpine grid Jan. 1983 pitfalls Ken Green coll. (ANIC); 1♂, paratype: Kosciusko Nat. Pk Smiggins Bog NSW 1600m 1983–1986 pitfall traps Ken Green No.63 (ANIC).</p><p>Habitus. BL 2.7−2.8mm, BL/BW 1.7. Brown, pronotal disc darkest, head anteriorly reddish, scutellum and sutural interval with reddish tinge and lighter than remainder of elytra.</p><p>Male. Segments 8 and 9 (Figs. 160−162) as for the genus, S8 with large densely pilose lateral remnants of the plate. The penis is stout, the wide pala occupies half of the length (Figs. 163, 164). Remains of muscles fill much of the pala, its side sclerites are strong, the front edge is truncate or bluntly rounded. The penis widens much at the origin of trigonium and parameroids. The trigonium is a slender triangle with rounded basal edges, shallowly concave sides of the long distal portion, and obtuse slightly asperous tip. The angularly projecting base of the parameroids carries a slender caudal lobe whose externally serrate outer half is folded mediad and concealed at rest. There is no tegmen, the parameres are separate sinuous rods with blunt ends that lean closely against penis and parameroids (Fig. 164).</p><p>Female. Unknown.</p><p>Note. Separate parameres not connected by a tegmen occur also in other Australian marsh beetles (e.g., several Petrocyphon spp.). Therefore, the similarity in this detail between N. plicatus and the Nothocyphon amphora -group is assumed to have arisen independently.</p><p>Etymology. The longitudinally folded parameroids suggested the name, the Latin adjective plicatus, folded.</p><p>FIGURES 156–164. Nothocyphon spp., males. N. naso: 156, T8, S8, T9 (from top); 157, S9; 158, penis; 159, tegmen and parameres.— N. plicatus: 160, T8, interrupted line delimits area covered with microtrichia, stippling indicates window-like pale area; 161, S8; 162, T9 and S9 superimposed, the folded condition is an artifact. 163, 164, penis in dorsoventral view: 163, partly torn, one parameroid unfolded; 164, intact, with the rod-like parameres; 160–172 and 163, 164 to the same scales, respectively.</p></div>	https://treatment.plazi.org/id/039F8D3EFF8BFFD196964360FE91F83A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Zwick, Peter	Zwick, Peter (2015): Australian Marsh Beetles (Coleoptera: Scirtidae). 7. Genus Nothocyphon, new genus. Zootaxa 3981 (3): 301-359, DOI: 10.11646/zootaxa.3981.3.1
