identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039FCF4AFFB4FF842AB26E8DFBAFF9ED.text	039FCF4AFFB4FF842AB26E8DFBAFF9ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus Weise 1901	<div><p>Genus Horniolus Weise</p><p>Horniolus Weise, 1901: 442 . Type species: Horniolus dispar Weise, 1901, by monotypy.</p><p>Diagnosis. Form elongate, oblong to broadly oval, dorsum moderately to strongly convex, densely pubescent, elytral pubescence often comprising a mixture of dark and paler hairs. Head with expanded clypeus, eyes with a distinct but narrow ocular canthus.Antennae with 11 antennomeres. Scutellar shield large, triangular and prominent. Prosternal carinae inverted Y-shaped with a short stem. Abdominal postcoxal line complete, deeply semicircular, or apically obliterated and apparently incomplete (Chen et al. 2016). Tarsal formula 4-4-4. Female genitalia with a complex, coiled spermatheca, coxites elongate triangular.</p><p>Note. Many species of this genus have a similar external appearance with the ground colour black to dark pitchy brown with yellowish-orange elytral maculae, often in the form of four orange yellow elytral spots of various sizes / shapes. This genus is generally poorly represented in institutional collections and due to its general resemblance to other Scymnini, it is likely to be difficult to spot in mixed collections.</p><p>Biology/ Hosts. Members of Horniolus mostly prey on mealybugs, such as Planococcus lilacinus (Cockerell) and Dysmicoccus brevipes (Cockerell) ( Hemiptera: Pseudococcidae), infesting coffee, tea and pineapple. These beetles also feed on spiralling whitefly, Aleurodicus dispersus Russell ( Hemiptera: Aleyrodidae) (Irulandi et al. 2001; Ramani et al. 2002; He et al. 2013; Poorani 2015; Chen et al. 2016; label data).</p></div>	https://treatment.plazi.org/id/039FCF4AFFB4FF842AB26E8DFBAFF9ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFB4FF872AB26B5BFE68FEA2.text	039FCF4AFFB4FF872AB26B5BFE68FEA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus dispar Weise, R. G. Booth 1901	<div><p>Horniolus dispar Weise</p><p>(Fig. 1a–d)</p><p>Horniolus dispar Weise, 1901: 443 (lectotype male; DEI).– Korschefsky 1931: 110; Poorani 2002: 349.</p><p>Diagnosis. This species (Fig. 1a, b) has reddish-brown head and pronotum and the elytra are dark brown to black, with each elytron having three bright yellow-orange maculae as follows (Fig. 1a): a small basal spot adjacent to the humeral callus; a large, transverse spot, roughly trapezoidal in shape and slightly oblique with a deep median constriction, reaching a little beyond midline; and a smaller, transverse oval macula near apical margin, narrowed and curved outward; the smallest basal spot fused with the large anterior macula in one example (Fig. 1b). Male genitalia as illustrated (Figs 1c, d) (Image credit: Keita Matsumoto, Copyright BMNH). (Based on the material studied at BMNH).</p><p>Material examined. Ceylon, G. Lewis, 1910-320 / Dikoya, 3800-4200 ft, 21.I.-7.II.22 (2: BMNH); one male specimen with same data, genitalia dissected and in glass vial pinned with the specimen, bearing the label “ Horniolus dispar Weise, R. G. Booth det. 1985” (BMNH) .</p><p>Distribution. Sri Lanka.</p><p>Note. Bielawski (1957) described H. dispar as Scymnus (Pullus) sp. from Sri Lanka and later corrected this error (Bielawski 1961).</p></div>	https://treatment.plazi.org/id/039FCF4AFFB4FF872AB26B5BFE68FEA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFB6FF862AB26DB5FB80FD5A.text	039FCF4AFFB6FF862AB26DB5FB80FD5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus guimeti (Mulsant)	<div><p>Horniolus guimeti (Mulsant)</p><p>(Fig. 1e)</p><p>Scymnus guimeti Mulsant, 1850: 979 (lectotype female; OUMNH).– Weise 1879: 145; Korschefsky 1931: 143.</p><p>Horniolus guimeti: Booth &amp; Pope 1989: 354 (lectotype designation).– Poorani 2002: 349; Chen et al. 2016: 119.</p><p>Distribution. Malaysia (Sarawak and Sabah); Bangladesh.</p><p>Note. Horniolus guimeti (Mulsant) (Fig. 1e, BMNH) was originally described by Mulsant (1850) as Scymnus guimeti based on Hope’s material. Booth &amp; Pope (1989) mentioned that “this species appears not to have been correctly recognized since its original description”, transferred it to Horniolus and designated a lectotype for it with illustrations of the male and female genitalia. As observed by Poorani (2002, 2015), it is a doubtful record for India and the specimens identified as H. guimeti by R. D. Pope in the collections of the erstwhile Commonwealth Institute of Biological Control, Bangalore, were found to be Scymnus (Pullus) latemaculatus Motschulsky, 1858, a common species with a superficially similar elytral pattern consisting of four yellow spots. Poorani (2002) and Chen et al. (2016) included Bangladesh in its distribution range and it remains an unverified record.</p></div>	https://treatment.plazi.org/id/039FCF4AFFB6FF862AB26DB5FB80FD5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFB6FF862AB26971FD92FA62.text	039FCF4AFFB6FF862AB26971FD92FA62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus hisamatsui Miyatake 1976	<div><p>Horniolus hisamatsui Miyatake</p><p>Horniolus hisamatsui Miyatake, 1976: 29 (holotype female; EUMJ).</p><p>Horniolus hisamatsui: Canepari 2003: 262 .– Chen et al. 2016: 114; Szawaryn &amp; Větrovec 2020: 576.</p><p>Distribution. India (Szawaryn &amp; Větrovec 2020), Nepal (Canepari 2003), and China (Guangdong, Hainan, Guangxi, Tibet) (Ren et al. 2009; Chen et al. 2016).</p><p>Note. Miyatake (1976) described it from Hong Kong. Chen et al. (2016) redescribed it with illustrations of specimens from China. Recently Szawaryn &amp; Větrovec (2020) recorded it from northeastern India (Assam) and illustrated the habitus and male genitalia.</p></div>	https://treatment.plazi.org/id/039FCF4AFFB6FF862AB26971FD92FA62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFB6FF862AB26FD2FC2AFBC6.text	039FCF4AFFB6FF862AB26FD2FC2AFBC6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus nigripes Miyatake	<div><p>Horniolus nigripes Miyatake</p><p>Horniolus nigripes Miyatake, 1976: 33 (holotype male; BPBM).– Poorani 2002: 350.</p><p>Distribution. India: Tamil Nadu.</p><p>Note. Miyatake (1976) described it based on a single male specimen collected from Valparai (Western Ghats / Nilgiris range) and illustrated the habitus and the male genitalia. It appears to be a very rare species and any representatives could not be found in any recent collections from the same area.</p></div>	https://treatment.plazi.org/id/039FCF4AFFB6FF862AB26FD2FC2AFBC6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFB6FF802AB268DAFE08FDF8.text	039FCF4AFFB6FF802AB268DAFE08FDF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus sororius Poorani 2015	<div><p>Horniolus sororius Poorani</p><p>(Fig. 2)</p><p>Horniolus sororius Poorani, 2015: 7 (holotype male; NBAIR).</p><p>Diagnosis. Length: 2.20–2.65 mm; width: 1.65–1.80 mm. Form (Fig. 2a, b) elongate, oblong oval, dorsum moderately convex and densely pubescent. Head and pronotum dark testaceous brown, elytra dark pitchy brown to black, each elytron with a pair of bright orange-yellow spots, anterior spot transverse, subrectangular with a wavy margin, posterior spot transverse oval and smaller, not reaching sutural or lateral margins, both spots occasionally larger, almost touching lateral margins of elytra. Ventral side dark reddish brown, except legs lighter, yellowish brown. Abdominal postcoxal line (Fig. 2c) complete and semicircular, area enclosed by postcoxal line sparsely punctate with few punctures along anterior margins. Male genitalia (Fig. 2e–g) and spermatheca (Fig. 2d) as illustrated.</p><p>Distribution. India: Karnataka; Kerala; Tamil Nadu.</p><p>Prey/associated habitat. Predatory on Aleurodicus dispersus Russell ( Hemiptera: Aleyrodidae); Ferrisia virgata (Cockerell) ( Hemiptera: Pseudococcidae); associated with mealybugs infesting coffee, garden plants, hedgerows and avenue trees such as Spathodea campanulata and Bauhinea purpurea; on Eleusine coracana (label data).</p><p>This species was recorded by Ramani et al. (2002) (as Horniolus sp.) as a predator of A. dispersus . Irulandi et al. (2001) recorded this species as H. vietnamicus Miyatake, 1976 and studied its biology on coffee mealybug, Planococcus lilacinus . Najitha (2016) and Vidya &amp; Bhaskar (2017) recorded it as a predator of pepper root mealybug ( Formicococcus polysperes Williams) from Kerala, South India.</p><p>Seasonal occurrence. Collected in June, September, December (label data).</p><p>Note. This species has a superficial similarity with Scymnus (Pullus) latemaculatus Motschulsky, 1858, one of the most commonly collected species of this region. Horniolus sororius can be differentiated from S. latemaculatus by the comparatively more elongate body outline, presence of a mixture of dark and paler setae on the elytra, prosternal carinae, and the genitalia.</p></div>	https://treatment.plazi.org/id/039FCF4AFFB6FF802AB268DAFE08FDF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFB0FF802AB26FACFBC9F8C1.text	039FCF4AFFB0FF802AB26FACFBC9F8C1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus darjeelingensis Poorani 2022	<div><p>Horniolus darjeelingensis sp. n.</p><p>(Fig. 3)</p><p>Diagnosis. This species can be differentiated from the other known species of Horniolus by its much larger size (2.80 mm long), more robust, broadly oval outline and the colour pattern with reddish brown head and pronotum and yellowish brown, medially infuscate elytra with a pair of basal maculae and a pair of median black spots and the male genitalia. It superficially resembles H. sonduongensis Hoàng, 1979 (as illustrated by Szawaryn &amp; Větrovec 2020) in terms of elytral pattern but the latter appears to be a much darker species with different male genitalia. The male genitalia in H. darjeelingensis sp. n. also somewhat resemble those of H. vietnamicus but the penis guide in the former has distinct lamellate expansions (absent in H. vietnamicus) and is apically broader and less acutely curved in lateral view (apically narrowed and acutely pointed in H. vietnamicus).</p><p>Description. Holotype male: Length: 2.80 mm; width: 1.81 mm; PL: 0.97 mm; PW: 1.29 mm; EL: 1.84 mm; EW: 1.81 mm. Form (Fig. 3a) broad oval, dorsum convex with dense yellowish white pubescence, eyes with erect silvery white hairs. Head and pronotum reddish brown with dark brown lateral margins, scutellar shield medially reddish brown with dark brown borders, elytra yellow, sutural and adjoining areas reddish brown, basal margin of elytra with a black median fascia with an emarginate lower margin, not reaching scutellar shield / lateral or humeral area; middle of each elytron with a black transverse spot (outer half narrow, inner half distinctly broader); margins of elytra narrowly black to dark brown; ventral side yellowish-testaceous, antenna and legs yellow. Head nearly 5x as wide as an eye, frons broad, interocular distance slightly more than thrice as wide as eye. Pronotum densely punctate on disc, punctures separated by 1–3 diameters, very shallowly impressed, interspaces between punctures shiny, punctures distinctly larger and more closely spaced and denser on lateral and anterolateral areas. Elytral punctures densely punctate, shallowly impressed, separated by 2–4 diameters, interspaces shiny. Abdominal postcoxal lines (Fig. 3b, c) complete, deeply semicircular, enclosed area with punctures progressively larger towards posterior. Male genitalia as illustrated (Figs. 3d–g), penis guide in ventral view (Fig. 3d) parallel-sided for up to two-thirds of its length, abruptly narrowed and triangular in the apical third, shorter than parameres, parameres with dense, elongate lateral and apical setae (Fig. 3e); penis (Fig. 3f) with a well-developed capsule having a short outer process and a distinctly longer inner process, apically modified as illustrated (Fig. 3g).</p><p>Female: Not studied.</p><p>Material examined. Holotype male: “ India W. Bengal, Darjeeling dist., Singla 300m 17.x.78, Besuchet Löbl” (NBAIR).</p><p>Distribution. India: West Bengal (Darjeeling).</p><p>Etymology. The specific epithet is in reference to its type locality (Darjeeling).</p></div>	https://treatment.plazi.org/id/039FCF4AFFB0FF802AB26FACFBC9F8C1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFB0FF8D2AB26A69FAE1FD18.text	039FCF4AFFB0FF8D2AB26A69FAE1FD18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus wiolettae Poorani 2022	<div><p>Horniolus wiolettae sp. n.</p><p>(Fig. 4)</p><p>Diagnosis. Horniolus wiolettae sp. n. resembles H. nigripes Miyatake and the male genitalia are also superficially similar. However, the head and pronotum in H. nigripes are black and the male genitalia also differ as follows: the penis guide is as long as parameres and distinctly narrower in the apical one-third and the penis is more strongly curved in Horniolus wiolettae sp. n. while in H. nigripes, the penis guide is shorter than parameres, elongate lanceolate and not narrowed in the apical one-third, and distinctly narrower in lateral view. It also resembles H. sororius and H. hisamatsui in having four-spotted elytra but the latter two species differ from it in having distinctly more transverse, narrower and smaller elytral maculae and different male genitalia. Horniolus dispar differs from the newly described species in having a much darker brown head and pronotum and anterior pair of elytral maculae differently shaped and more deeply constricted in the middle.</p><p>Description. Male: Length: 1.42–1.68 mm; width: 1.01–1.19 mm. TL/TW: 1.41; PL/PW: 0.49–0.53; EL/EW: 1.00–1.03. Form (Fig. 4a) elongate oval, dorsum moderately convex, densely pubescent, head and pronotum with yellowish hairs, lateral sides of pronotum with a mixure of dark brown and paler hairs, elytra with bright yellowish hairs on yellow maculae, rest of elytra with dark brown pubescence. Head and pronotum dark reddish brown; scutellar shield black; elytra black, each elytron with a pair of yellowish maculae, anterior one large, broad with a wavy margin, just below basal margin, not reaching suture, almost touching lateral margin, posterior spot smaller, elongate, oblique oval. Head about 3.5x as wide as an eye, interocular distance about 1.6x as wide as an eye; Pronotum densely punctate, punctures separated by 1–2 diameters. Elytra densely punctate, punctures shallowly impressed, separated by 1–3 diameters, interspaces smooth. Abdominal postcoxal lines (Fig.4b) complete, broadly and deeply semicircular. Male genitalia (Figs. 4c–f) with penis guide in ventral view (Fig. 4c) basally broadest, its sides gradually narrowed towards middle, abruptly narrowed posteriorly into a triangular apex; penis guide in lateral view (Fig. 4d) basally curved, apically strongly narrowed to an acute tip; parameres (Fig. 4c, d) with inner and apical margins densely setose, subequal in length to penis guide; penis (Fig. 4e) with a distinct capsule, strongly curved, apex as illustrated on Fig. 4f.</p><p>Female: Not studied.</p><p>Material examined. Holotype male; “ INDIA, TN, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=77.18&amp;materialsCitation.latitude=8.26" title="Search Plazi for locations around (long 77.18/lat 8.26)">Kanyakumari</a>, Pechiparai, 18.vi.15, N08.26 o E077.18 o SM Team” (NBAIR) ; Paratype: One male, with the same data as holotype (NBAIR) .</p><p>Distribution. India: Tamil Nadu.</p><p>Etymology. This species is named in honour of Dr. Wioletta Tomaszewska, Polish Academy of Sciences, Warsaw, for her significant contributions to Coccinellidae systematics.</p><p>Note. A single specimen identified as Horniolus nr. nigripes (BMNH, Fig. 1f) corresponds to this species. The specimen bears two identification labels, one as “ Horniolus ? dispar Weise, det. R. B. Madge 1985” and the other as “ Horniolus nigripes Miyatake, det. R.G. Booth 1985” [“ INDIA: Chrys S. Kerala / on Lagerstroemia sp., CIE A16725 / Nymph on leaf, Lagerstroemia sp., Vazhani, 16.8.84, K.M. Das C.M., P.P., KFRI / genitalia in glass vial / Horniolus ? dispar Weise, det. R. B. Madge 1985 / Horniolus ? nigripes Miyatake, det. R. G. Booth 1985” (BMNH)].</p></div>	https://treatment.plazi.org/id/039FCF4AFFB0FF8D2AB26A69FAE1FD18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFBDFF8D2AB26894FDB0FA30.text	039FCF4AFFBDFF8D2AB26894FDB0FA30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus darjeelingensis Poorani 2022	<div><p>1. Horniolus darjeelingensis sp. n.</p><p>Distribution: India (West Bengal).</p></div>	https://treatment.plazi.org/id/039FCF4AFFBDFF8D2AB26894FDB0FA30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFBDFF8D2AB268DEFE25F9FD.text	039FCF4AFFBDFF8D2AB268DEFE25F9FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus dispar Weise, R. G. Booth 1901	<div><p>2. Horniolus dispar Weise, 1901</p><p>Distribution: Sri Lanka.</p></div>	https://treatment.plazi.org/id/039FCF4AFFBDFF8D2AB268DEFE25F9FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFBDFF8D2AB26B63FDAAF9BA.text	039FCF4AFFBDFF8D2AB26B63FDAAF9BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus guimeti (Mulsant 1850)	<div><p>3. Horniolus guimeti (Mulsant, 1850)</p><p>Distribution: Malaysia, Bangladesh.</p></div>	https://treatment.plazi.org/id/039FCF4AFFBDFF8D2AB26B63FDAAF9BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFBDFF8D2AB26BA4FC74F900.text	039FCF4AFFBDFF8D2AB26BA4FC74F900.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus hisamatsui Miyatake 1976	<div><p>4. Horniolus hisamatsui Miyatake, 1976</p><p>Distribution: India (Assam, West Bengal), Nepal, Hong Kong.</p></div>	https://treatment.plazi.org/id/039FCF4AFFBDFF8D2AB26BA4FC74F900	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFBDFF8D2AB26E0BFC3BFAE7.text	039FCF4AFFBDFF8D2AB26E0BFC3BFAE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus nepalensis (Canepari 1997) Poorani 2022	<div><p>Horniolus nepalensis (Canepari, 1997) comb. n.</p><p>(Fig. 5)</p><p>Platynaspis nepalensis Canepari, 1997: 20, 45 (Holotype female; MHNG).</p><p>Platynaspis nepalensis Canepari, 1997, described from Nepal, is transferred to Horniolus . Canepari’s illustrations and the images of the holotype female (obtained through the courtesy of Dr. Giulio Cuccodoro, MHNG) (Fig. 5) clearly establish it as a species of Horniolus . The dorsal habitus (Fig. 5a) of P. nepalensis is reminiscent of some of the known species of Horniolus such as H. hisamatsui and H. sonduongensis (as illustrated by Szawaryn &amp; Větrovec 2020) in having a discal black spot on each elytron. The abdominal postcoxal lines (Fig. 5c) are deeply semicircular and complete and typical of Horniolus . The female genitalia (Fig. 5d) (illustrated in Canepari 1997: p. 40, Figs 123–125) with intricately coiled spermatheca and the elongate triangular coxites are characteristic of Horniolus and not found in any of the Platynaspis species of this region or elsewhere, all of which have broad and transverse coxites. Only Crypticolus, a genus of Platynaspini from Madagascar, has elongate triangular coxites with a complex, distally coiled spermatheca (Ślipiński &amp; Tomaszewska 2002).</p></div>	https://treatment.plazi.org/id/039FCF4AFFBDFF8D2AB26E0BFC3BFAE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFBDFF8D2AB26BEEFE5EF8CD.text	039FCF4AFFBDFF8D2AB26BEEFE5EF8CD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus nepalensis (Canepari 1997) Poorani 2022	<div><p>5. Horniolus nepalensis (Canepari, 1997) comb. n.</p><p>Distribution: Nepal.</p></div>	https://treatment.plazi.org/id/039FCF4AFFBDFF8D2AB26BEEFE5EF8CD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFBDFF8D2AB26A33FDCBF88B.text	039FCF4AFFBDFF8D2AB26A33FDCBF88B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus nigripes Miyatake 1976	<div><p>6. Horniolus nigripes Miyatake, 1976</p><p>Distribution: India (Tamil Nadu).</p></div>	https://treatment.plazi.org/id/039FCF4AFFBDFF8D2AB26A33FDCBF88B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFBDFF8D2AB26A75FD72F850.text	039FCF4AFFBDFF8D2AB26A75FD72F850.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus sororius Poorani 2015	<div><p>7. Horniolus sororius Poorani, 2015</p><p>Distribution: India (Karnataka, Kerala).</p></div>	https://treatment.plazi.org/id/039FCF4AFFBDFF8D2AB26A75FD72F850	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
039FCF4AFFBDFF8D2AB26ABEFDCBF81E.text	039FCF4AFFBDFF8D2AB26ABEFDCBF81E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Horniolus wiolettae Poorani 2022	<div><p>8. Horniolus wiolettae sp. n.</p><p>Distribution: India (Tamil Nadu).</p></div>	https://treatment.plazi.org/id/039FCF4AFFBDFF8D2AB26ABEFDCBF81E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Poorani, J.	Poorani, J. (2022): A review of Horniolus Weise (Coleoptera: Coccinellidae) from the Indian region, including descriptions of two new species. Zootaxa 5104 (4): 545-556, DOI: 10.11646/zootaxa.5104.4.4
