taxonID	type	description	language	source
039C6B74B71BFFAABAEB05B1FB16FE84.taxon	materials_examined	TYPE: Peru. Amazonas: Luya, Camporredondo, El Cedro, 6 ° 14 ’ 07 ” S, 78 ° 20 ’ 01 ” W, 1550 m, 18 June 1999, J. Campos [de la Cruz] 6049 (holotype, USM- 164664!; isotype, MO- 6147261! [barcode MO- 2080242]). Figures 1, 2.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71BFFAABAEB05B1FB16FE84.taxon	diagnosis	Diagnosis: — Similar to P. andicola Esquerre (2019: 280) but the outer corona differs in the new species by being (2.0 –) 2.9 – 3.7 (– 4.4) mm long and having a single purplish band, instead of 4 – 5 mm long and with three purplish bands. Furthermore, the leaves of P. luyensis are abaxially densely pubescent as opposed to only slightly pubescent in P. andicola.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71BFFAABAEB05B1FB16FE84.taxon	description	Vine, densely pubescent throughout. Stems striate. Stipules (2.9 –) 3.3 – 4.0 (– 5.0) × 0.3 – 0.4 (– 0.5) mm, falcate, sparsely pubescent abaxially, petioles (8.4 –) 11.1 – 17.3 (– 21.8) mm long, glandless; laminas (5.1 –) 5.6 – 7.5 (– 8.8) × (2.1 –) 2.3 – 3.6 (– 4.7) cm, entire, narrowly obovate to obovate, not variegated, round at the base, densely pubescent abaxially, tomentose trichomes 0.2 – 0.3 mm long, glabrous adaxially, 3 - lobed, the lateral lobes slightly obtuse, the central lobe slightly obtuse to acute; the angle between the lateral lobes veins (14 –) 16 – 18 (– 22) °, with 4 – 8 ocellate yellowish glands (0.9 –) 1.1 – 1.7 (– 2.5) mm diam. in two lines between primary veins, with two of the glands located at the very base, and rarely and only on the largest leaves two additional glands outside the base of the two primary lateral veins, each primary vein ending in a distinct mucro. Peduncles (9.6 –) 17.0 – 18.0 (– 25.2) mm long, in pairs, bracts (1.2 –) 1.8 – 2.5 (– 3.5) × 0.3 – 0.4 mm, linear-triangular, entire or 2 - cleft, scattered along the peduncle; floral stipe (1.4 –) 2.2 – 2.7 (– 3.4) mm long to (2.4 –) 2.8 – 3.9 (– 4.1) mm long in fruit. Flowers (32.2 –) 33.4 – 33.6 (– 36.5) mm diam., white, born suberect or facing above the horizontal plane; sepals (13.2 –) 14.4 – 15.4 (– 17.0) × (2.8 –) 4.0 – 4.5 (– 5.5) mm, oblong-triangular, densely minutely pubescent outside, obtuse to rounded apex, occasionally with pinkish base; petals (5.8 –) 8.2 – 8.7 (– 10.2) × (1.4 –) 2.1 – 2.4 (– 3.1) mm, ca. ½ as long as sepals, narrowly oblong to oblong-ovate, rounded apex, white; coronal filaments in 2 series, filaments of outer row (2.0 –) 2.9 – 3.7 (– 4.4) mm long, filiform, distally slightly dilated, white or cream, with one subapical purple or reddish purple band, rarely with pink base; filaments of the inner row (2.1 –) 2.6 – 2.8 (– 2.9) mm long, capillary, erect, green, fimbriate; operculum (1.5 –) 1.9 – 2.3 (– 3.1) mm long, green with purplish apex; nectar ring 0.42 – 0.48 mm high; androgynophore (4.5 –) 6.0 – 7.5 (– 9.4) mm long, pale green to whitish; filaments (4.3 –) 4.9 – 5.5 (– 6.0) mm long; anthers (2.8 –) 3.8 – 4.3 (– 4.9) × (1.2 –) 1.8 – 2.0 (– 2.5) mm; ovary (1.6 –) 2.4 – 3.1 (– 4.9) × (1.2 –) 2.2 – 2.5 (– 3.7) mm, ovoid to globose, densely pubescent; styles (3.1 –) 4.7 – 5.0 (– 6.2) × 0.3 – 0.4 (– 0.5) mm, flushed purple; stigma (0.4 –) 0.9 – 1.1 (– 1.3) mm diam., capitate. Fruit (6.9 –) 10.1 – 13.6 (– 18.0) × (6.9 –) 10.6 – 12.3 (– 16.6) mm, subglobose; aril unknown; seeds 2.7 – 3.2 (– 3.5) × (2.1 –) 2.2 – 2.4 (– 2.5) mm, transversely sulcate with 8 sulci, rugulose.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71BFFAABAEB05B1FB16FE84.taxon	etymology	Etymology. The species is named after the province of Luya (Department of Amazonas), where it was found.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71BFFAABAEB05B1FB16FE84.taxon	conservation	Conservation status. The EOO is estimated as 129 Km 2 and AOO as 12 km 2. The species is known from four locations. None of these locations are protected, and deforestation represents a clear threat to the species. A decline in AOO, extent and quality of habitat, number of locations and number of individuals is therefore expected, and P. luyensis is assigned a conservation status of “ Endangered ” [EN B 1 a + B 2 a].	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71BFFAABAEB05B1FB16FE84.taxon	biology_ecology	Phenology. The species has been collected from May to June in flowers and nearly mature fruit, and photographed in late October in bud, flowers, and immature fruit.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71BFFAABAEB05B1FB16FE84.taxon	discussion	Discussion. Passiflora luyensis is similar to P. andicola and P. pascoensis L. Escobar (1989: 880). The easiest character to distinguish these three species is the flower diameter, which is smallest (32.2 – 36.5 mm) in P. luyensis, 40 – 43 mm in P. andicola, and largest (70 – 80 mm) in P. pascoensis. Passiflora luyensis can also be distinguished by the number of the purple bands in its outer row of filaments, which have one subapical purple or reddish-purple band, rarely with pink base, while P. andicola has two or three purple bands and P. pascoensis has three to six purple bands. Paratypes: PERU. Amazonas: Luya. Camporredondo, Fundo Cedro, 2450 – 2550 m, 26 May 1989, C. Diaz et al. 3549 (MO- 4355746, MYF, QCNE, USM- 127287). Cajamarca: Cutervo, outskirts of Socota, weedy roadside, 06 ° 19 ’ S, 078 ° 41 ’ W, 1800 m, 10 Feb. 1988, Gentry et al. 61472 (MO- 6147775, USM- 121749).	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71EFFABBAEB012BFD31FABC.taxon	materials_examined	TYPE: Peru. Pasco: Oxapampa: road from Oxapampa to Yaupi, Torrebamba, hedges between fields, 10 ° 44 ’ S, 75 ° 28 ’ W, 1830 m, 6 Apr. 2001, M. Weigend et al. 5504 (Ref. Nr. 2001 / 504. 9) (holotype, NY!; isotypes, HUT- 41618 [image!]; M- 0142684! [barcode], MO- 5731213! [barcode MO- 1272544], MO- 6147706! [barcode MO- 1183418]). Figure 3.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71EFFABBAEB012BFD31FABC.taxon	diagnosis	Diagnosis: — Similar to P. pilpintu T. Boza but the lamina differs by the wider angle between the lateral lobes veins (22 – 28 ° vs. 10 – 17 °). Furthermore, P. hexadenia has extrafloral laminar nectaries outside (exmedial) the primary lateral veins at the leaf base in addition to between the primary veins, instead of having nectaries only between the primary veins. Moreover, the corona is white and diagnostically unbanded in P. hexadenia, while in P. pilpintu it is purple banded.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71EFFABBAEB012BFD31FABC.taxon	description	Vine, sparsely puberulous. Stems striate. Stipules 2.9 – 3.2 × 0.3 – 0.4 mm, narrowly triangular, more or less falcate; petioles 11.5 – 17.5 mm long, glandless; laminas 4.8 – 8.3 × 3.2 – 5.3 cm, entire, obovate, round at the base, glabrous abaxially and adaxially, (2 –) 3 - lobed, the lateral and central lobe emarginate; the angle between the lateral lobes veins 22 – 28 °, with 12 – 15 ocellate glands 0.8 – 1.3 mm diam., 6 glands located at the base of the lamina, four in two lines between the primary veins and two outside of the primary veins, each primary vein ending in a distinct mucro. Peduncles 13.3 – 16.1 mm long, slender, in pairs; bracts 1.8 – 2.1 × 0.2 – 0.3 mm, scattered along the peduncle, floral stipe 1.1 – 1.8 mm long to 2.1 – 7.7 mm long in fruit. Flowers 3.5 – 3.6 cm diam., greenish white or pale yellow-green; sepals 15.7 – 16.8 × 3.7 – 3.9 mm, narrowly oblong triangular, glabrous outside, acute apex, occasionally pink near base; petals 4.6 – 6.1 × 1.5 – 1.7 mm, narrowly obovate; obtuse apex, ca. 1 / 3 as long as sepals, white; coronal filaments in 2 series, filaments of outer row 7.4 – 8.7 mm long, filiform, pure white, very rarely with pink base; filaments of the inner row 1.5 – 2 mm long, operculum 1.5 – 1.7 mm long, membranous, plicate; nectar ring 1.6 mm high; limen annular, membranaceous, erect; androgynophore 5.0 – 5.5 mm long, green; filaments 3.3 – 3.7 mm long; anthers 2.8 – 3.6 × 1.2 – 1.5 mm; ovary 1.7 – 3.0 × 1.6 – 2.4 mm, ellipsoid, sparsely hirsute; styles 3.6 – 4.3 × 0.2 – 0.3 mm; stigma 0.8 – 1.1 mm diam. capitate. Immature fruit 6.0 – 7.7 × 6.2 – 6.5 mm, globose, sparsely hirsute. Seeds unknown.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71EFFABBAEB012BFD31FABC.taxon	etymology	Etymology. The name refers to the six glands crowded at the base of the leaf blade, which looks like six eyes.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71EFFABBAEB012BFD31FABC.taxon	conservation	Conservation status. The EOO could not be estimated because the species is only known from the type; the AOO is 4 km 2. The type locality is 21 km from the Yanachaga Chemillen National Park, in an area that is highly degraded and under logging pressure. Passiflora hexadenia is provisionally assigned a conservation status of “ Critically Endangered ” [CR B 2 a].	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71EFFABBAEB012BFD31FABC.taxon	biology_ecology	Phenology. Passiflora hexadenia was collected in flower and immature to nearly mature fruit in April.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71EFFABBAEB012BFD31FABC.taxon	discussion	Discussion. Passiflora hexadenia resembles P. pascoensis by having 2 – 3 - lobed leaves, however it differs from P. pascoensis by having the lateral and central lobe emarginate (vs. acute) and the four ocellate glands located in two lines between the primary veins along the lamina (vs. at the apex). Passiflora hexadenia is characterized by having smaller flowers than P. pascoensis (3.5 – 3.6 vs. 7.0 – 8.0 cm in diam.). Acha et al. (2021) reported a DNA analysis under their sample 565 as “ P. hexadenia, ” but the identification was corrected later to P. pilpintu (see next species).	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71FFFA9BAEB0563FB36FA7D.taxon	materials_examined	TYPE: Peru. Pasco: Oxapampa: Distr. Chontabamba. Carretera la suiza nueva, bosque intervenido en borde la carretera, 10 ° 40 ’ 44 ” S, 75 ° 26 ’ 34 ” W, 2360 m, 8 Aug. 2008, D. Rodríguez M., Antonio Peña, & R. Rivera 100 (holotype, MO- 6253990! [barcode MO 1183704] and MO- 6253991! [barcode MO 1183703]; HOXA 031903 [image!], HUT, USM). Figures 4, 5.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71FFFA9BAEB0563FB36FA7D.taxon	diagnosis	Diagnosis: — The flower and narrow leaves of this new species are very similar to those of P. quadriflora Killip (1927: 424), as is the subangular stem, which often dries flattened and scabrous, but the leaves differ by having the two primary lateral veins spreading 10 ° – 17 °, reaching the laminar margins, and forming mucros and a lobed leaf (vs. lateral veins arching inwards distally toward the central vein and not reaching the margin in P. quadriflora).	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71FFFA9BAEB0563FB36FA7D.taxon	description	Vine 1 – 2 m long, glabrescent. Stems striate, subangulate, drying slightly scabrous. Stipules 1.9 – 2.7 × 0.3 – 0.4 mm, narrowly triangular, more or less falcate, glabrous; petioles 5.2 – 12.5 mm long; laminas 4.6 – 5.7 × 2.0 – 2.3 cm, entire, variegated as juveniles, variegation often retained at maturity, narrowly oblong-obovate (obovate), rounded at the base, often reddish purple abaxially, glabrous or abaxially very sparsely glabrescent and with a few lax trichomes on the primary veins, 3 - lobed, the lobes ca. equal or the center slightly longer, or slightly shorter or obscure and the leaf thus very shallowly bilobed, the lateral and central lobes obtuse to acute, variegated along veins only; the angle between the lateral lobes veins 10 ° – 17 °, with (7 –) 10 – 14 ocellate yellow glands 0.5 – 0.8 mm diam. in lines between primary veins only, each primary vein ending in a distinct mucro. Peduncles 9.4 – 21.3 mm long, in pairs, bracts 1.4 – 4.6 × 0.1 – 0.2 mm, linear, scattered along the peduncle; floral stipe ca. 3.5 mm long to 3.4 – 4.2 mm long in fruit. Flowers 3.5 – 6.3 cm diam., greenish white or pale yellow-green; sepals 20.9 – 24.8 × 4.4 – 4.6 mm, narrowly ovate-oblong, glabrous, acute apex, pale green to white; petals 10.5 – 8.9 × 0.3 – 0.5 mm, narrowly oblong-ovate; about ½ as long as the sepals or slightly less, round apex, white; coronal filaments in 2 series, filaments of outer row 5.9 – 10.0 mm long, white with 3 – 4 purple bands; filaments of the inner row 2.4 – 3.2 mm long; operculum 3.1 – 3.4 mm long, membranous, plicate; nectar ring present; limen annular, membranaceous, erect; androgynophore 10.5 – 12.2 mm long, pale green; stamens with filaments 4.2 – 5.6 mm long; anthers 4.3 – 4.4 × 1.7 – 2.0 mm; ovary 2.1 – 2.8 × 1.3 – 1.4 mm, ovoid, glabrous to minutely puberulous; styles 3.9 – 4.3 × 0.5 – 0.6 mm, purplish; stigma 1.2 – 1.7 mm diam., capitate. Fruit 11.2 – 13.9 × 9.1 – 12.7 mm, globose to widely ellipsoid, glabrous to sparsely pubescent, aril unknown, seeds 3.3 – 3.7 × 2.3 – 2.9 mm, transversely ridged with rugulose or verrucose ridges with 7 – 9 sulci.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71FFFA9BAEB0563FB36FA7D.taxon	etymology	Etymology. This new species is named for the shape of the leaf, which resembles the shape of Lepidopteran wing scales. The indeclinable term “ pilpintu ” comes from the Quechua language meaning butterfly and is allowed under International Code of Nomenclature [ICN] Art. 23.2, (Turland et al. 2018).	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71FFFA9BAEB0563FB36FA7D.taxon	conservation	Conservation Status. The species is known from four neighboring localities, none of which are protected. Logging activities and deforestation represent a potential threat to the species. The AOO is estimated at 8 km 2, a decline in its extent and number of individuals may therefore be expected. Passiflora pilpintu is assigned a conservation status of “ Endangered ” [EN B 2 a]. Phenology. Flowering material of P. pilpintu has been collected in August – September and fruits have been found in January and August – September.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71FFFA9BAEB0563FB36FA7D.taxon	discussion	Discussion. The taxonomic placement of this species in section Decaloba was investigated in detail by Acha et al. (2021). Samples from the specimens Valenzuela 13876 and Vásquez 28889 (as samples “ sp. nov. 388 ” and “ P. hexadenia 565 ”, respectively) were genotyped and they emerged together into the southern Andean “ South American Clade 7 ” of 20 – 25 species of this section, including P. indecora Kunth (1817: 134) and P. quadriflora.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71FFFA9BAEB0563FB36FA7D.taxon	materials_examined	Paratypes: PERU. Pasco: Oxapampa: Dist. Huancabamba, zona de amortiguamiento del Parque Nacional Yanachaga Chemillén, Sector Torre Bamba, camino a Oso Playa, 10 ° 20 ’ 09 ” S, 075 ° 34 ’ 58 ” W, 2444 m, 10 sept. 2010, E. Briceño et al. 229 (HOXA 044866); Sector Oso Playa, camino hacia el campamento, 10 ° 18 ’ 52 ” S, 075 ° 34 ’ 31 ’’ W, 2478 m, 22 Oct. 2009, L. Valenzuela et al. 13876 (MO- 6748874 [barcode MO- 2080699] SRA acc. no. — SRS 7821085); Parque Nacional Yanachaga-Chemillén, Sector Chacos, 10 ° 37 ’ S, 075 ° 17 ’ W, 2471 m, 24 Jan. 2006, R. Vásquez et al. 28889 (HOXA, MO- 6147476 [barcode MO- 1183262] SRA acc. no. — SRS 7820982, USM).	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71DFFA7BAEB0622FD17FD34.taxon	materials_examined	TYPE: Peru. Amazonas: Distr. Chachapoyas, remnants of forest a few km past Molinopampa, 06 ° 12 ’ S, 76 ° 40 ’ W, 2450 m, 13 Apr. 2001, H. Van der Werff, R. Vásquez, B. Gray, & R. Rojas 17007 A (holotype, MO- 04838344! barcode MO- 1103150). Figure 6.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71DFFA7BAEB0622FD17FD34.taxon	diagnosis	Diagnosis: — Differs from all other species of Passiflora in Peru by the combination of bilobed leaves densely hirsutepubescent abaxially, with the angle between the veins of the lateral lobes of the leaf 28 – 48 ° and laminar nectary glands present, bracts at least 1 mm wide, and the outer corona 11 – 12.3 mm long.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71DFFA7BAEB0622FD17FD34.taxon	description	Vine, densely to sparsely puberulent throughout. Stems striate. Stipules (3.1 –) 4.0 – 4.5 (– 6.1) × (0.2 –) 0.4 – 0.5 mm, linear to linear triangular, more or less falcate, very sparsely pubescent abaxially; petioles (13.4 –) 14.6 – 16.7 (– 18.5) mm long, glandless; laminas (3.8 –) 6.1 – 7.1 (– 8.6) × (4.0 –) 4.4 – 5.3 (– 6.0) cm, entire, very broadly obovate to broadly obovate, cordate at the base, densely hirsute abaxially, trichomes 0.2 – 0.3 mm long, sparsely hirsute adaxially, trichomes 0.2 mm long; 3 - lobed, the lateral lobes acuminate to acute, central lobe cusp-like; the angle between the lateral lobes veins (28 –) 33 – 38 (– 48) º, with 4 – 10 ocellate glands 0.6 – 0.9 (– 1.1) mm diam. in lines between the primary veins only, 4 glands located at the base between primary veins, each primary vein ending in a distinct mucro. Peduncles (8.5 –) 14.7 – 21.8 (– 30.2) mm long, in pairs, bracts (3.6 –) 4.4 – 5.3 (– 6.6) × (1.0 –) 1.1 – 1.2 mm, linear, scattered along the peduncle; floral stipe (1.9 –) 2.1 – 2.4 (– 2.9) mm long to 7.5 mm long in fruit. Flowers 4.7 – 4.8 cm diam., greenish-white or pale yellow-green; sepals (11.4 –) 16.3 – 18.1 (– 21.8) × (3.8 –) 5.1 – 5.7 (– 7.0) mm, hirsute outside, obtuse apex; petals 10.1 – 11.2 × 3.7 – 3.8 mm, linear to narrowly elliptic, acute apex, white, base purple; coronal filaments in 2 series; filaments of outer row 11.0 – 12.3 mm long, filiform, truncate, color unknown; filaments of the inner row (2.0 –) 2.4 – 2.7 (– 3.5) mm long, capillary, slightly clavate at apex; operculum (2.3 –) 2.9 – 3.2 (– 3.8) mm long, membranous, plicate, crenate; androgynophore 7.3 – 8.1 (– 8.8) mm long; filaments (5.4 –) 5.9 – 6.4 (– 6.6) mm long; anthers (5.1 –) 5.3 – 5.5 (– 5.6) × (1.4 –) 1.6 – 1.8 (– 2.3) mm; ovary 3.6 – 5.1 (– 6.4) × 2.3 – 3.7 mm, ovoid, densely hirsute; styles (4.9 –) 5.7 – 7.0 (– 8.4) × (0.2 –) 0.4 – 0.5 (– 0.8) mm; stigma (1.7 –) 2.1 – 2.4 (– 2.7) mm diam., capitate. Fruit (immature) ca. 22.0 × 13.4 mm, ellipsoid, densely hirsute, mature size unknown but color at maturity “ dark blue ” (Weigend 98 / 412); aril unknown, seeds 3.0 – 3.6 × 2.4 – 2.6 mm, transversely sulcate with smooth ridges with 7 – 9 sulci, acute at the base.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71DFFA7BAEB0622FD17FD34.taxon	etymology	Etymology. This new species is named after the Chachapoyas district in northern Peru, where it grows.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71DFFA7BAEB0622FD17FD34.taxon	conservation	Conservation status. The species is only known from two populations in Chachapoyas (Amazonas). The AOO is estimated as 4 km 2. Based on this restricted distribution and habitat, P. chachapoyensis is assigned a conservation status of “ Endangered ” [EN B 2 a].	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71DFFA7BAEB0622FD17FD34.taxon	biology_ecology	Phenology. Flowers observed in April and June, fruits have been found in June.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B71DFFA7BAEB0622FD17FD34.taxon	discussion	Discussion. Passiflora chachapoyensis is one of the few passifloras in Peru with bilobed leaves that are densely hirsute beneath and can be distinguished by the characters described in the diagnosis. Passiflora cisnana Harms (1894: 5), though not closely related (Boza et al. 2018), is similar in leaf shape, both having pubescent broadly obovate leaves with cordate base, but it can be distinguished from the new species by the absence of laminar nectaries and floral bracts. Paratypes: PERU. Amazonas: Prov. Mendoza, road Chachapoyas to Mendoza, 40 – 42 km E of Chachapoyas, 2400 – 2500 m, 5 Jun. 1988, M. Weigend et al. 98 / 412 (F- 2211490, M- 0007958 [barcode], USM- 172620); M. Weigend et al. 98 / 441 (F- 2209984, M- 0007957 [barcode]).	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B713FFA5BAEB02DBFE51F9A9.taxon	materials_examined	TYPE: PERU. Cajamarca: Prov. San Ignacio, mountain slope above San José de Lourdes, 1300 – 1600 m, [05 ° 06 ' 16 " S, 078 ° 53 ' 50 " W], 9 Jun. 1998, M. Weigend, Th. Franke, J. Skrabal, & M. A. Gonzales B. 98 / 488 (holotype, USM- 175434!; isotypes, F- 2211422!, M- 0007959! [barcode], MO 5297020! [barcode]). Figures 7, 8.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B713FFA5BAEB02DBFE51F9A9.taxon	diagnosis	Diagnosis: — The flowers of this new species are similar to those of P. pilpintu T. Boza, also with purple banded corona, but the sepals are shorter (15.3 – 16.1 vs 20.9 – 24.5 mm), there are 3 (4) coronal rows [vs 2], and the androgynophore is shorter (7.2 – 7.7 vs 10.5 – 12.5 mm long). Furthermore, P. coronapapillata has 2 - lobed leaves with wide angle between the lateral lobes veins (60 ° –) 70 ° – 80 ° (– 90 °), instead of 3 - lobed leaves with narrow angle (10 ° – 17 °).	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B713FFA5BAEB02DBFE51F9A9.taxon	description	Vine, very sparsely minutely puberulous to mostly glabrescent throughout. Stems striate. Stipules (2.5 –) 3.0 – 4.7 × (0.5 –) 1.2 – 1.4 mm, linear, more or less falcate, glabrous; petioles 12.7 – 13.7 mm long, glandless; laminas (1.5 –) 3.5 – 4.8 × (2.5 –) 6.6 – 7.5 cm, entire, depressed ovate, rounded at the base, glabrous throughout, 2 - lobed, the lateral lobes acute to rounded; the angle between the lateral lobes veins (60 ° –) 70 ° – 80 ° (– 90 °), with (7 –) 8 – 12 ocellate glands, glands 0.8 – 1.0 mm diam., only in lines between the primary veins, each primary vein ending in a distinct mucro. Peduncles 23.6 – 43.0 mm long, in pairs, bracts 2.7 – 3.0 × 1.0 – 1.1 mm, narrowly elliptic tapering at both ends, scattered along the apical 1 / 3 of the peduncle; floral stipe 1.5 – 2.3 mm to 4.0 mm in fruit. Flowers 3.4 – 3.7 cm diam., white to greenish white; sepals 15.3 – 16.1 × 4.4 – 4.9 mm, narrowly ovate, acute to rounded apex, green flushed with red outside, offwhite inside; petals 8.5 – 10.1 × 2.3 – 3.9 mm, narrowly ovate; about 2 / 3 as long as the sepals, obtuse apex, snow white; coronal filaments in 3 (– 4) series, filaments of outer row 7.9 – 9.0 mm long, linear, apical 1 / 3 acicular and adaxially with a zone of papillae, base white, then three dark violet or purplish bands, then several paler blue or purplish bands towards the apex, filaments of the 2 (– 3) inner rows similar and difficult to distinguish, 2.7 – 3.1 mm long, capillary, obconic-papillate apex, yellowish green with purple bands; operculum ca. 2.1 – 2.6 mm long, membranous, plicate; androgynophore 7.2 – 7.7 mm long, green; filaments 3.7 – 4.6 mm long; anthers 3.8 – 4.4 × 1.8 – 2.1 mm; ovary 2.7 – 2.8 × 1.7 – 2.1 mm, ovoid to ellipsoid, sparsely puberulous, the trichomes in longitudinal rows; styles 4.6 – 5.8 mm, purplish red; stigma 1.4 – 1.5 mm diam. capitate. Fruit 14 – 18 mm × 14 – 16 mm (including 1 mm long fruit stipe), widely ellipsoid, color unknown; seeds 3.5 – 4.0 × 2.5 – 2.8 mm, transversely sulcate with rugulose ridges, (5 –) 6 – 7 sulci.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B713FFA5BAEB02DBFE51F9A9.taxon	etymology	Etymology. The name refers to the conspicuous obconic-papillate apices of the flower’s inner rows of filaments. The outer coronal filaments also have a region with conspicuous papillae, a feature that is rare in P. subg. Decaloba.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B713FFA5BAEB02DBFE51F9A9.taxon	conservation	Conservation status. Passiflora coronapapillata is known from five collections in one general location. Based on its very restricted distribution and habitat, this species is assigned a conservation status of “ Critically Endangered ” [CR B 2 a].	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B713FFA5BAEB02DBFE51F9A9.taxon	biology_ecology	Phenology. Flowering and fruiting known in May – June and in October.	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
039C6B74B713FFA5BAEB02DBFE51F9A9.taxon	discussion	Discussion. The flowers of this species have a complex condensed inner corona of 2 (– 3) series that are inconspicuously ranked and may appear as one series to the unaided eye. The taxonomic placement of this species was studied by Acha et al. (2021). A sample from the paratype Campos 3901 (as “ P. coronapapillata 421 ”) was genotyped and the phylogenetic analysis surprisingly nested the species inside their “ Central American Clade 2 ” near the West Indian P. penduliflora Bertero ex DC. [de Candolle 1828: 326], a dubious placement that would require an exceptional case of long-distance dispersal to the Peruvian Andes and convergent evolution of a several-banded corona. The authors did not discuss this anomalous result, which may have been due to experimental or other error, as the sequence obtained was short compared to the other samples (Acha et al. 2021). Passiflora coronapapillata shares with P. punctata similar leaves with ocellate glands but differs by laminas with (7 –) 8 – 12 ocellate glands arranged along lines between the primary veins [vs. (12 –) 20 – 35 ocellate glands between and often outside primary veins in P. punctata]. Passiflora coronapapillata is also characterized by having corona filaments in 3 (– 4) series, with the outer row filaments acicular in the apical 1 / 3 and adaxially with a zone of papillae, while P. punctata has corona filaments in 2 series, with the outer row filaments falcate, dilated near the middle, boomerang-shaped, and with a triangular apex. Paratypes: PERU. Cajamarca: Prov. San Ignacio, Distr. San José de Lourdes, localidad Laurel, relicto de bosque primario (bosque intervenido), 05 ° 01 ' 00 " S, 078 ° 57 ' 00 " W, 1500 – 1600 m, 17 May 1997 (imm. fr.), Campos & Vargas 3901 (MO 5297020 [barcode MO- 1273943] SRA acc. no. - SRS 7820969, USM 171579); Distr. San José de Lourdes, Cerro Parcos, near Villa Rica town, 05 ° 03 ’ 51 ’’ S, 078 ° 53 ’ 29 ’’ W, 1780 m, 2 Oct. 2023 (imm. fr.), Esquerre 232 (PRG); 05 ° 04 ’ 05 ’’ S, 078 ° 53 ’ 24 ’’ W, 1830 m, 2 Oct. 2023 (buds), Esquerre 233 (PRG); 05 ° 04 ’ 04 ’’ S, 078 ° 53 ’ 8 ’’ W. 1790 m, 2 Oct. 2023, Esquerre 234 (PRG).	en	Espinoza, Tatiana Erika Boza, Macdougal, John M., Jørgensen, Peter Møller (2025): Five New Species of Passiflora (subgenus Decaloba, Passifloraceae) from Peru. Phytotaxa 716 (2): 72-86, DOI: 10.11646/phytotaxa.716.2.1, URL: https://doi.org/10.11646/phytotaxa.716.2.1
