taxonID	type	description	language	source
039D87A4AC052013FF337A4FDF528D73.taxon	description	Figures 1 – 4, 12 http: // zoobank. org / NomenclaturalActs / 8 B 307 A 58 - A 49 F- 487 D- 8486 - 893086 B 96 AF 4	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC052013FF337A4FDF528D73.taxon	materials_examined	Type material. Holotype: male (HNU) China, Hunan Province, Xinning County, Langshan National Geological Park, rock cave of Yunv, 110 ° 44.012 ' E, 26 ° 16.543 ' N, 311 m, 25 November 2014, collected from cracks between stones, Haiqiang Yin, Cheng Wang, Bing Zhou, Jiahui Gan, Yuhui Gong leg. Paratypes: 42 females, 14 males, same data as holotype.	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC052013FF337A4FDF528D73.taxon	etymology	Etymology. The specific name is a combination of the Latin prefix “ bi- ” (two, twice, double) and the Latin “ ocellatus ” (having ocular spots), referring to the two white eyes which are strongly reduced in size.	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC052013FF337A4FDF528D73.taxon	diagnosis	Diagnosis. This species resembles Leptonetela quinquespinata (Chen & Zhu, 2008) in similar proportions of each part of the male palp; slightly wide, flattened conductor; similar shape of the atrium (Figs 1 B – D, 2 C, 3 A, B, 4 A, C in this present paper; figs 44 B – D, 45 C, 46 A, B, 47 B, D in Wang & Li 2011). The new species can be distinguished by eyes reduced to two white spots; median apophysis completely covered by the conductor, neither present in L. quinquespinata (Figs 1 A, B, 2 A, 4 C in this present paper; figs 44 A, B, 45 A, 47 B, D in Wang & Li 2011); chelicerae with ten promarginal and seven small retromarginal teeth in new species (Fig. 4 D), but nine promarginal and five small retromarginal teeth in L. quinquespinata (fig. 47 C in Wang & Li 2011). Males of the new species resemble L. anshun Lin & Li, 2010, L. grandispina Lin & Li, 2010, L. liping Lin & Li, 2010, L. maxillacostata Lin & Li, 2010, L. tetracantha Lin & Li, 2010, L. zhai Wang & Li, 2011, L. yaoi Wang & Li, 2011, L. thracia Gasparo, 2005, L. mita Wang & Li, 2011, and L. microdonta (Xu & Song, 1983) in pedipalpal tarsus with a transverse groove, slightly contracted mesially, with no distinct wrinkles (figs 4 A, B, 19 A, B, 31 A, B, 33 A, B, 49 A, B in Lin & Li 2010; figs 70 A, B, 67 A, B, 59 A, B, 30 A, B, 34 A, B in Wang & Li 2011). It can be distinguished from all the former species by the presence of two white eyespots (Figs 1 A, 2 A); membranous and slightly wide flattened conductor; embolus distally twisted at one third; median apophysis hidden by the conductor (Figs 1 B, 4 C). Females resemble L. parlonga Wang & Li, 2011 by the shape of atria and spermathecae; but can be distinguished by eyes reduced to two eyespots in the new species (Figs 2 A, 3 C), normal six eyes in L. parlonga (fig. 37 A in Wang & Li 2011); anterior margin of the atrium decorated with dense short and slender hairs in L. parlonga (fig. 37 C in Wang & Li 2011), absent in the new species (Fig. 2 C).	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC052013FF337A4FDF528D73.taxon	description	Description. Male. Total length 2.45 (Fig. 1 A). Carapace 1.13 long, 0.94 wide. Opisthosoma 1.23 long, 0.99 wide. Prosoma yellowish, with several setae near the anterior margin of carapace and a short seta situated before ALE (anterior lateral eyes) (Fig. 3 C). ALE strongly reduced in size, only two white eyespots present; PME completely absent; PLE barely visible, with slight vestiges left. Eye sizes: ALE 0.03, the vestige of PLE less than 0.01 long. Thoracic median groove short, brown, needle-shaped. Cervical grooves and radial furrows light brown, indistinct. Clypeus 0.22 high. Chelicerae light brown, with ten promarginal and seven small retromarginal teeth, both rows of teeth gradually becoming smaller and denser from the base to the distal end of fang furrow (Fig. 4 D). Endites reddish brown. Labium yellowish brown and plump, fused to sternum. Sternum pale yellow, peltate. Legs yellowish. Leg measurements: I 12.33 (3.51, 0.40, 3.56, 3.07, 1.79); II 9.72 (2.99, 0.31, 2.75, 2.45, 1.22); III 8.57 (2.32, 0.35, 2.36, 2.27, 1.27); IV 10.74 (3.17, 0.25, 3.00, 2.75, 1.57). Leg formula: I – IV – II – III. Opisthosoma pale brown, ovoid, lacking distinct patterns (Fig. 1 A). Male pedipalpus as illustrated in Figs 1 B – D, 3 A, B, 4 C. Femur without any spines. Patella distally provided with a dorsal spine. Tibia with three trichobothria dorsally, five long spines prolaterally and eight spines (five spines in a longitudinal row along the tibia and the other three in a transversal row along distal margin of the tibia) retrolaterally. Tarsus sunken and contracted at middle position, attaching to an earlobe-shaped process retrolaterally. One distal short spine, three retrolateral and three prolateral long spines present on the distal half of the tarsus (Figs 1 C, D, 3 A, B). Pedipalpal bulb oval in shape, smooth. Conductor membranous, slight wide, flattened. Embolus with distal one third twisted towards prolateral side and the distal end slightly broader, not sharp. Median apophysis completely covered by the conductor and its vague outline can be seen through the conductor (Figs 1 B, 4 C). Prolateral lobe medium sized, triangular (Fig. 3 A). Female. Similar to male in general features, somatic coloration and body size, but legs shorter. Total length 2.43 (Figs 2 A, B). Carapace 0.98 long, 0.90 wide. Opisthosoma 1.42 long, 1.13 wide. Eye sizes: ALE 0.03, the vestige of PLE 0.02 long. Clypeus 0.22 high. Leg measurements: I 9.43 (2.55, 0.36, 2.83, 2.23, 1.46); II 8.19 (2.31, 0.39, 2.31, 1.92, 1.26); III 7.88 (1.89, 0.32, 1.92, 1.67, 1.08); IV 8.49 (2.51, 0.38, 2.45, 1.98, 1.17). Leg formula: I – IV – II – III. Genital area densely covered with long hairs. Atrium much wider than long. Internal genitalia consist of paired spermathecae and sperm ducts. Spermathecae highly twisted, distal end more strongly sclerotized than proximally, and the distal ends separated slightly far from each other (Figs 2 B, C, 4 A, B).	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC052013FF337A4FDF528D73.taxon	distribution	Distribution. Known only from the type locality (Fig. 12).	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC02201EFF337A19DF528ACF.taxon	description	Figures 5 – 8, 12 http: // zoobank. org / NomenclaturalActs / 9 D 53 BD 4 D-BCDF- 4 DA 2 - 9369 - 6 AADA 3 C 89151	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC02201EFF337A19DF528ACF.taxon	materials_examined	Type material. Holotype: male (HNU) China, Hunan Province, Shaoyang County, Hebo Town, Chengbei Village, Jigong Cave, 111 ° 17.460 ' E, 26 ° 45.438 ' N, 576 m, 23 November, 2011, Xiang Xu, Jinlong Wan, Yi Zhao, Shihong Peng leg. Paratypes: 14 females, 15 males, same data as holotype.	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC02201EFF337A19DF528ACF.taxon	etymology	Etymology. The specific name is an adjective in apposition and derived from the Latin words “ lata ” (broad) and “ apicalis ” (apical), in reference to the presence of broad and peak-shaped median apophysis.	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC02201EFF337A19DF528ACF.taxon	diagnosis	Diagnosis. Males resemble those of Leptonetela hexacantha Lin & Li, 2010, L. jinsha Lin & Li, 2010, L. reticulopecta Lin & Li, 2010, L. kanellisi (Deeleman-Reinhold, 1971), and L. tianxingensis Wang & Li, 2011 by having depressed and strongly contracted structure in the middle of the pedipalpal tarsus (Figs 5 C, D, 7 A, B in the present paper; figs 24 A, B, 26 A, B, 45 A, B in Lin & Li 2010; figs 18 A, B, 63 A, B in Wang & Li 2011); short distance between apical tarsus and pedipalpal bulb (Figs 5 C, D, 7 A, B in the present paper; figs 24 A, B, 26 A, B, 45 A, B in Lin & Li 2010; figs 18 A, B, 63 A, B in Wang & Li 2011), the new species is distinguished from these species by presence of three prolateral, nine retrolateral tibial spines on male pedipalpus; broad and peak-shaped median apophysis; short, wide, translucent conductor on male bulb (Figs 5 C, D, 7 A, B). The new species resembles L. quinquespinata (Chen & Zhu, 2008) in being eyeless, having membranous embolus, slightly twisted prolaterally (Figs 5 A, B, 8 B in this present paper; figs 44 A, B, 47 D in Wang & Li 2011); it can be distinguished by nine strong spines located in retrolateral side of pedipalpal tibia (Figs 5 D, 7 A in this present paper; figs 44 B, 46 C in Wang & Li 2011); nine small retromarginal teeth on the chelicerae in the new species (Fig. 8 D in this present paper; fig. 47 C in Wang & Li 2011), while six strong spines retrolaterally-directed on the pedipalpal tibia; five small retromarginal teeth on the chelicerae in L. quinquespinata (Figs 5 D, 7 A in this present paper; figs 44 D, 46 B in Wang & Li 2011). Females resemble those of L. quinquespinata in being eyeless; having highly twisted sperm, but can be distinguished by the distal ends of sperm ducts closer placed to each other in the new species (Figs 6 C, 8 B), slightly further away from each other in L. quinquespinata (figs 45 C, 47 B in Wang & Li 2011).	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC02201EFF337A19DF528ACF.taxon	description	Description. Male. Total length 2.69 (Fig. 5 A). Carapace 1.26 long, 1.12 wide. Opisthosoma 1.37 long, 1.03 wide. Prosoma reddish brown, with several setae near the anterior margin of carapace. Ocular area with a pair of setae, eyes absolutely absent. Median groove short, cervical grooves and radial furrows light brown. Clypeus 0.24 high. Chelicerae brown, with nine promarginal and eight small retromarginal teeth, promarginal row of teeth gradually becoming smaller and denser from the base to distal end of fang furrow (Fig. 8 D). Endites brown. Labium brown and plump, fused with sternum. Sternum and legs yellowish. Leg measurements: I 12.72 (3.53, 0.45, 3.98, 2.97, 1.79); II 10.03 (2.91, 0.41, 3.09, 2.12, 1.50); III 8.66 (2.52, 0.42, 2.41, 2.10, 1.21); IV 10.34 (3.12, 0.36, 2.98, 2.53, 1.35). Leg formula: I – IV – II – III. Opisthosoma pale brown, ovoid, lacking distinctive patterns. Male pedipalpus (Figs 5 B – D, 7 A, B, 8 C): femur covered with long and thin hairs; tibia with three trichobothria dorsally; three slender spines prolaterally and nine strong spines retrolaterally (six spines along the tibia form a longitudinal row and the other three spines along distal margin of the tibia form a transversal row) exist on the pedipalpal tibia (Figs 7 A, B). Tarsus rugose and contracted mesially, attaching to an earlobe-shaped process retrolaterally, with long spines distally (Figs 5 C, D, 7 A, B). Pedipalpal bulb nearly round; embolus membranous, slightly twisted towards the prolateral side; conductor translucent, broad, flat; median apophysis broad, peakshaped (Figs 5 B, 8 C). Prolateral lobe cuspate (Fig. 7 B). Female. Similar to male in coloration of opisthosoma and general features, but larger body size, shorter legs, and prosoma yellowish. Total length 3.06 (Figs 6 A, B). Carapace 1.23 long, 1.15 wide. Opisthosoma 1.70 long, 1.24 wide. Clypeus 0.24 high. Leg measurements: I 10.94 (3.06, 0.40, 3.33, 2.42, 1.73); II 9.46 (2.62, 0.41, 2.88, 2.13, 1.42); III 7.93 (2.38, 0.33, 2.21, 1.86, 1.15); IV 10.26 (2.95, 0.36, 2.95, 2.48, 1.52). Leg formula: I – IV – II – III. Genital area densely covered with long hairs (Figs 6 B, 8 A). Internal genitalia with a pair of spermathecae and sperm ducts: spermathecae dark brown, sclerotized and highly twisted, with the distal ends very close to each other; sperm ducts pale brown, less sclerotized (Figs 6 C, 8 B). The atrium broad, nearly triangular, slightly procurved at anterior median margin (Fig. 8 B).	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC02201EFF337A19DF528ACF.taxon	distribution	Distribution. Known only from the type locality (Fig. 12).	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC0F2005FF337DB5DF5289B4.taxon	description	Figures 9 – 12 http: // zoobank. org / NomenclaturalActs / 59 BAAE 5 B- 7 E 35 - 44 E 8 - 9576 - 5 F 8 F 9 CEBED 3 D	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC0F2005FF337DB5DF5289B4.taxon	materials_examined	Type material. Holotype: male (HNU) China, Hunan Province, Yuanling County, Xiazhai Village, Wuyuan cave, 110 ° 35.518 ’ E, 28 ° 57.537 ’ N, 315 m, 1 November, 2006, Xiang Xu and Guo Tang leg.	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC0F2005FF337DB5DF5289B4.taxon	etymology	Etymology. The species is named after one of the collectors, Mr. Guo Tang, a deceased colleague at Hunan Normal University, China; name is in genitive case.	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC0F2005FF337DB5DF5289B4.taxon	diagnosis	Diagnosis. Leptonetela tangi sp. nov. resembles L. gigachela (Lin & Li, 2010) by strong chelicerae; male pedipalp tibia subequal to tarsus in length (Figs 9 C, D, 10 A, B, 11 B, C in the present paper; figs 1 A, B, 2 D, E in Lin & Li, 2010); it can be distinguished by presence of median apophysis in new species, absent in L. gigachela; mid pedipalpal tarsus ventral apophysis absent in L. tangi sp. nov.; present in L. gigachela (Figs 9 B, 10 A, B, 11 A); Leptonetela tangi sp. nov. also resembles L. chakou Wang & Li, 2017, L. kangsa Wang & Li, 2017, L. langdong Wang & Li, 2017, L. qiangdao Wang & Li, 2017, and L. shanji Wang & Li, 2017 by having a strong second spine close to proximal male palp tip (Figs 9 C, D, 10 A, B in the present paper; figs 4 C, D, 20 C, D, 22 C, D, 54 C, D, 58 C, D in Wang, Xu & Li, 2017); it can be distinguished by median apophysis with thick base and sharp tip in new species (Fig. 9 B in the present paper; figs 4 B, 20 B, 22 B, 54 B, 54 B in Wang, Xu & Li, 2017).	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC0F2005FF337DB5DF5289B4.taxon	description	Description. Male. Total length 1.91 (Fig. 9 A). Carapace 0.97 long, 0.88 wide. Opisthosoma 0.93 long, 0.77 wide. Prosoma yellowish brown and pear-shaped, with two setae at anterolateral area of PME and many short setae in the anterior margin of clypeus. Thoracic median groove dark-brown, needle-shaped. Cervical groove and radial furrows light brown. Clypeus 0.17 high, slightly sloped anteriorly. Six eyes, ALE and PLE connected to each other by the black bases, PME separated from ALE and PLE (Figs 9 A, 11 D). Eye sizes: ALE 0.10, PLE 0.09, PME 0.06. ALE-ALE 0.01, ALE-PME 0.14, PLE-PLE 0.13, PLE-PME 0.05, AER 0.19, PER 0.24. Chelicerae yellowish, long and strong, fang furrow with nine promarginal (the first tooth sturdy) and six retromarginal teeth (the first tooth especially sturdy, even stronger than the first promarginal tooth) (Figs 11 B, C). Endites pale yellow. Labium yellowish and plump, fused to sternum. Sternum yellowish, peltate. Legs yellowish. Leg measurements: I 7.25 (2.01, 0.35, 2.12, 1.68, 1.09); II 5.22 (1.53, 0.30, 1.28, 1.20, 0.91); III 4.41 (1.27, 0.29, 1.16, 0.93, 0.76); IV 6.25 (1.82, 0.29, 1.74, 1.43, 0.97). Leg formula: I – IV – II – III. Opisthosoma deep brown, ovoid, covered with short hairs. Male pedipalpus (Figs 9 B – D, 10 A, B, 11 A): tibia and tarsus subequal in length; femur slightly longer than tibia and tarsus, with four short ventral spines; patella with one dorsal spine distally; tibia with seven long spines ventrally (the second one near the basal end especially strong), but without any trichobothria; tarsus with a long seta ventrally, a strong apical seta, three prolateral, and three retrolateral setae at the distal half, and attaching to an earlobe-shaped process medially (Figs 9 C, D, 10 A, B); bulb ovoid, with a smooth surface; embolus and conductor thin and translucent; median apophysis distinct, strongly cuticularized, with a thick base and sharp end (Figs 9 B, 11 A). Female. Unknown.	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
039D87A4AC0F2005FF337DB5DF5289B4.taxon	distribution	Distribution. Known only from the type locality (Fig. 12).	en	He, Ailan, Liu, Jinxin, Xu, Xiang, Yin, Haiqiang, Peng, Xianjin (2019): Description of three new species of spider genus Leptonetela Kratochvíl, 1978 from caves of Hunan Province, China (Araneae, Leptonetidae). Zootaxa 4554 (2): 584-600, DOI: 10.11646/zootaxa.4554.2.10
