taxonID	type	description	language	source
039D87A30729FF9EFF11F9D53DD6FDD0.taxon	description	section Calusparassus subsection Quadripartitae (Steenis) Steenis (1953 a: 337). Type: — Nothofagus fusca (Hook. f.) Oerst. [≡ Fuscospora fusca (Hook. f.) Heenan & Smissen]. Steenis (1952 b) provided the superfluous name of sect. Planae for the previously published Nothofagus section Sempervirentes (Steenis 1952 a) in an erratum. Steenis (1953 a) gives the type of Nothofagus section Calusparassus subsection Quadripartitae as N. betuloides, but clearly cites Steenis (1952 a) for the basionym where N. fusca is given as the type.	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A30729FF9EFF11F9D53DD6FDD0.taxon	materials_examined	Type: — Nothofagus gunnii (Hook. f.) Oerst. [≡ Fuscospora gunnii (Hook. f.) Heenan & Smissen].	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A30729FF9EFF11F9D53DD6FDD0.taxon	description	Trees up to 30 m high. Leaves distichous, revolute when evergreen or plicate when deciduous, entire or teeth with 1 – 2 serrations, fimbrial veins incomplete or absent; with or without large globular glandular trichomes on cuticle; with solitary unicellular trichome type A with a very small, unthickened base and with a large, unicellular, thin-walled trichome emerging; without solitary unicellular trichome type C with a large base equal to or greater than the diameter of the trichome and with a large, unicellular, thin-walled trichome emerging; with or without conical unicellular trichomes with a heavily thickened foot cell. Stomata randomly oriented; without thickened T-pieces of cuticle at the poles separating the two guard cells; without giant stomata over the major veins; stomata size within the areoles variable. Upper epidermal cells over veins more elongate than areolar cells, with granular cell walls. Stipules not peltate. Dichasia with 1 central dimerous flower and 2 lateral trimerous flowers, or 1 or 0 dimerous flower and 1 trimerous flower. Male floral meristem round. Staminate flowers with narrowly campanulate, often bell-shaped perianth; lobes 4, rarely 3 or 5, prominent, symmetric and regular; stamens <20, centripetal development; anthers 2.5 – 4.5 mm long, slightly curved, distal connective protrusion weakly developed, filament connective free, without epidermal papillae. Pollen peritreme, mesocolpia straight to convex; colpi short, spatulate or parallel-sided, with U-shaped ends, margins inwardly conspicuously heavily thickened, aperture 4 – 11 µm long, polar to equatorial lengths ratio ≤ 0.3. Cupule valves 2 – 4; outer surface of valves with simple trichomes; lamellae membranous, glabrous; fruits 0 – 1 dimerous, 1 – 2 trimerous, or 4 – 7. Six species in New Zealand, southern South America, and Tasmania.	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A3072FFF9DFF11FDD13AC7F91F.taxon	description	Govaerts & Frodin (1999) placed N. truncata in synonymy of Nothofagus fusca var. colensoi (Hooker 1853: 229) Cheeseman (1906: 641). This treatment is incomprehensible and contradicts the available ecological, morphological and genetic evidence on the distinctiveness of F. fusca and F. truncata and is therefore not accepted here or elsewhere (e. g., Allan 1961, Wardle 1984, Mark & Lee 1985, Haase 1990, 1992, Dawson & Lucas 2011).	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A3072FFF9DFF11FDD13AC7F91F.taxon	description	Fuscospora × eugenananus (Gilland.) Heenan & Smissen, comb. nov. ≡ Nothofagus × eugenananus Gillanders (2008: 56). Type: — AUSTRALIA. Woodbank gardens, Longley, Tasmania, May 2007, K. D. Gillanders s. n. (holotype HO 544640). Hybrid parentage: — Fuscospora fusca (Hook. f.) Heenan & Smissen × F. alessandri (Espinosa) Heenan & Smissen (Gillanders 2008).	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A3072FFF9DFF11FDD13AC7F91F.taxon	description	Seven species in New Zealand, southern South America, and Australia.	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A3072CFF9CFF11F89B3C8AFC4B.taxon	description	F. M. Vásquez (in sched., 20 September 1995) annotated SGO 63396 as lectotype. We are unaware of any other published lectotypification and as we concur with this assessment we formally designate this specimen as lectotype.	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A3072DFF9CFF11FC673C99F9DE.taxon	description	& Smissen (Grant & Clement 2004: 448).	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A3072DFF82FF11F9F13AF8FD7E.taxon	description	nom. inval. Conserved type (designated by Lanjouw et al. 1961: 251): — Nothofagus antarctica (G. Forst.) Oerst. In proposing the conservation of Nothofagus, Steenis (1953 b) did not nominate a type, but the proposal was nonetheless recommended by the Committee for Spermatophyta with the provision that this deficiency was remedied (Rickett 1958). When the conservation of the name was published, N. antarctica was given as the type (Lanjouw et al. 1961, Appendix 3). This was an unfortunate choice as Blume (1851) did not include N. antarctica among the species he placed in Nothofagus. Nothofagus subgenus Molischia and Nothofagus section Calucechinus subsection Antarcticae each include the type of Nothofagus (N. antarctica) and are therefore invalid (McNeill et al. 2012, ICN Art. 22.2).	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A3072DFF82FF11F9F13AF8FD7E.taxon	description	for deciduous species (section Eufagus). Fagus section Eufagus is treated here as invalid (McNeil et al. 2012, ICN Art. 21.3).	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A3072DFF82FF11F9F13AF8FD7E.taxon	description	41.5). Autonyms: — Nothofagus Blume subgenus Nothofagus, in Philipson & Philipson (1988: 33); Nothofagus Blume subgenus Nothofagus section Nothofagus, in Philipson & Philipson (1988: 33). Trees up to 45 m high. Leaves distichous, planar when evergreen or plicate when deciduous, teeth with 1 – 2 or 2 - or-more serrations, fimbrial veins incomplete or absent; with large globular glandular trichomes on cuticle; without solitary unicellular trichome type A with a very small, unthickened base and with a large, unicellular, thin-walled trichome emerging; without solitary unicellular trichome type C with a large base equal to or greater than the diameter of the trichome and with a large, unicellular, thin-walled trichome emerging; with or without conical unicellular trichomes with a heavily thickened foot cell. Stomata parallel to long axis of leaf; with thickened T-pieces of cuticle at the poles separating the two guard cells; without giant stomata over the major veins; stomata size within the areoles variable or more-or-less even. Upper epidermal cells over veins more elongate than areolar cells or not distinguishable from areolar cells, with or without granular cell walls. Stipules peltate. Dichasia with 1 central dimerous flower and 2 lateral trimerous flowers, or 1 trimerous flower. Male floral meristem round. Staminate flowers with narrowly campanulate, often bell-shaped perianth; lobes 4, rarely 3 or 5, prominent, symmetric and regular; stamens <20, centripetal development; anthers 2.5 – 4.0 mm long, slightly curved, distal connective protrusion weakly developed, filament connective free, without epidermal papillae. Pollen peritreme, mesocolpia straight to convex; colpi short, spatulate or parallel-sided, with U-shaped ends, margins conspicuously thickened, annulate, aperture 4 – 11 µm long, polar to equatorial lengths ratio ≤ 0.3. Cupule valves (2 –) 4 (– 8), or 2 asymmetrical; outer surface of valves with simple trichomes; lamellae membranous, usually glabrous, or sometimes glandular; fruits 1 dimerous or 1 – 2 trimerous. Five species in southern South America.	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A30733FF87FF11FD383F33F9DE.taxon	description	C. Muñoz (in sched., February 1960) annotated SGO 63009 as type specimen. We are unaware of any other published lectotypification and as we concur with this assessment we formally designate this specimen as lectotype.	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A30733FF87FF11FD383F33F9DE.taxon	description	indicate a type (McNeill et al. 2012, ICN Art. 40.1).	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A30733FF87FF11FD383F33F9DE.taxon	description	name because it lacked a full and direct reference to the basionym (McNeill et al. 2012, ICN Art. 41.5).	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A30733FF87FF11FD383F33F9DE.taxon	description	time because it lacked a full and direct reference to the basionym (McNeill et al. 2012, ICN Art. 41.5).	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A30733FF87FF11FD383F33F9DE.taxon	description	= Trisyngyne subgenus Trisyngyne section Trisyngyne series Balansae Baumann-Bodenheim (1992: 110 – 111), nom. inval. This name is invalid as Baumann-Bodenheim (1992) included three species in this taxon but did not indicate a type (McNeill et al. 2012, ICN Art. 40.1).	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
039D87A30733FF87FF11FD383F33F9DE.taxon	materials_examined	Type: — Trisyngyne baumanniae Baum. - Bod. Baumann-Bodenheim (1992) did not explicitly indicate a type for this taxon but included only T. baumanniae which is acceptable as an indication of the type (McNeill et al. 2012, ICN Art. 40.3). Trees up to 40 m high, or sometime shrubs. Leaves distichous or spiralled, conduplicate, evergreen, entire or teeth with 1 – 2 serrations, fimbrial veins incomplete or absent; with large globular glandular trichomes on cuticle; without solitary unicellular trichome type A with a very small, unthickened base and with a large, unicellular, thin-walled trichome emerging; without solitary unicellular trichome type C with a large base equal to or greater than the diameter of the trichome and with a large, unicellular, thin-walled trichome emerging; with or without conical unicellular trichomes with a heavily thickened foot cell. Stomata randomly oriented; without thickened T-pieces of cuticle at the poles separating the two guard cells; with giant stomata over the major veins; stomata size within the areoles more-or-less even. Upper epidermal cells over veins thinner than areolar cells or not distinguishable from areolar cells, without granular cell walls. Stipules peltate. Dichasia 1 - or 3 - dimerous. Male floral meristem round. Staminate flowers with a tubular strongly connate perianth; lobes 4, rarely 2, 3 or 5, reduced, symmetric and regular; stamens <20, centripetal development; anthers 2.0 – 8.0 mm long, straight with only the apex slightly curved, distal connective protrusion pronounced and strongly developed, filament connective connate, with epidermal cells consisting of longitudinal ridges of striated papillae, often with papillose ornamentation on the distal connective protrusion and weakly ornamented cells on the lateral, dorsal and ventral sides. Pollen goniotreme, mesocolpia straight to concave; colpi parallel-sided with U-shaped ends, margins inwardly thickened and / or inflexed; aperture 4 – 11 µm long, polar to equatorial lengths ratio> 0.3. Cupule valves 2 or absent; outer surface of valves glabrous; lamellae membranous or woody, glabrous, entire; fruits 1 or 3 dimerous. Twenty-five species from Papua New Guinea, Indonesian Papua and New Caledonia. Notes: — Baumann-Bodenheim (1992) proposed the transfer of a number of names in Nothofagus to Trisyngyne. However, he did not provide a full and direct reference to the basionym and place of publication and therefore his combinations are not validly published (McNeill et al. 2012, ICN Art. 41.5). It is therefore necessary to make new combinations in Trisyngyne for names published in Nothofagus, and this action is taken below.	en	Heenan, Peter B., Smissen, Rob D. (2013): Revised circumscription of Nothofagus and recognition of the segregate genera Fuscospora, Lophozonia, and Trisyngyne (Nothofagaceae). Phytotaxa 146 (1): 1-31, DOI: 10.11646/phytotaxa.146.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.146.1.1
