taxonID	type	description	language	source
039D8245D41F783FFF62FC768ADDFD21.taxon	type_taxon	Type species. Tympanoterpes colombiae Distant 1892: 60 (Colombia).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D41F783FFF62FC768ADDFD21.taxon	description	Description. Body size variable (19 – 36 mm body length). Head wider than mesonotum, eyes protruding beyond anterior pronotum, vertex at area of ocelli as long as or a little shorter than front, postclypeus globose, centrally sulcate, rounded or transverse anteriorly, rostrum generally reaching the hind coxae or hind trochanters, extending to abdominal sternites in the occasional specimen. Pronotum shorter than mesonotum, trapezoidal with anterior margin narrower than lateral margins of pronotal collar, lateral angles of pronotal collar expanded, metanotum extends laterally beyond wing groove, cruciform elevation smoothly arched posteriorly. Metasternum transverse central plate moderately elevated, not angularly produced anteriorly. Fore wings and hind wings generally hyaline, with eight and six apical cells, respectively, radial and radiomedial crossveins generally parallel, obliquely oriented to radius posterior and median veins respectively and without infuscation. Basal cell longer than broad with variable markings. Basal area of fore wing clavus and hind wing costal cell may be infuscated. Fore femora with oblique primary spine, upright secondary spine, and a small tertiary spine, tarsi three-segmented. Male operculum triangular not covering the tympanal cavity completely laterally or posteromedially, nor medially in some species, almost or reaching to sternite II, opercula well separated along midline, meracanthus tapering to a point, reaching to or passing posterior margin of operculum, female operculum and meracantus generally of similar shape to male for each species, opercula smaller extending medially only to lateral base of meracanthus. Abdomen about as long as distance between apex of head and posterior of cruciform elevation. Abdominal segments with sides expanding laterally to tergite 4 where the abdomen begins narrowing posteriorly to the genitalia, epipleurites folded toward dorsal surface producing a channel on the lateral margins of the abdomen, posterior sternites translucent in males of some species. Female sternite VII with sinuate or straight posterior margin and medial notch. Timbal cover incomplete, with rounded anterior apex, concealing only the lateral aspect of the timbal, timbal cover with concave anteromedial margin that can curve toward apex or produce approximate parallel dorsal and ventral margins to anterior of extension above the lateral timbal before reaching the rounded apex, ventral margin is parallel to long body axis, timbals extend below wing bases. Pygofer distal shoulder rounded, dorsal beak absent, pygofer basal lobe indistinct, pygofer upper lobe large and generally well separated from pygofer, paddle-like often twisting towards apex, median uncus lobe extending posteriorly, lateral uncus lobes curve under median uncus lobes forming an arch when viewed from the side and restraining aedeagus, and male aedeagus a simple tubular structure with a membranous region near the terminus. Female abdominal segment 9 with dorsal beak well defined and sinuate posterior margin, ovipositor sheath extends beyond dorsal beak. Measurements (MM). Length of body: 19 – 36; length of fore wing: 27 – 46; width of fore wing: 7.7 – 13.1; length of head: 1.7 – 5.7; width of head including eyes: 7.5 – 13.6; width of pronotum including suprahumeral plates: 7.6 – 13.6; width of mesonotum: 6.5 – 11.3.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D41F783FFF62FC768ADDFD21.taxon	diagnosis	Diagnosis. Within the Fidicinini, Ariasa can be distinguished from the three genera (Fidicina Amyot & Audinet-Serville, 1843, Fidicinoides Boulard & Martinelli, 1996, and Berglana Boulard & Martinelli, 1996) of the Fidicinina by the presence of three-part tarsi. Hemisciera Amyot & Audinet-Serville, 1843 is simply distinguished from all other Fidicinini by the bright green basal area of the fore wings and bright red on the basal area of the hind wings. The adpressed pronotal margins, non-parallel radial and radiomedial crossveins, relatively small eyes, small anterolaterally extended timbal covers, and the sternite separating the male opercula distinguish the members of Beameria Davis, 1934. Majeorona Distant, 1905, has a head width wider than the mesonotum, with the prominent eyes stylate and extended anteriorly. The genera Proarna Stål, 1864, Prasinosoma Torres, 1963, Tympanoterpes Stål, 1861, all possess a head that is about as wide as the mesonotum with the eyes barely or not extending beyond the anterior angle of the pronotum, radial crossvein vertically oriented to the radius anterior 2 and radius posterior veins with a radiomedial crossvein that is obliquely oriented, large semicircular male opercula, and triangular or small, curved timbal covers. The single species of Elassoneura Torres, 1964 shares most of the characters in these genera but the head is wider than the mesonotum and there is significant infuscation along the majority of the fore wing venation. The single species of the genus Pompanonia Boulard, 1982 can be distinguished by a head that is wider than the mesonotum, the radial crossvein vertically oriented to the radius anterior 2 and radius posterior veins with a radiomedial crossvein that is obliquely oriented, semicircular male opercula, triangular timbal covers, and the very wide base of the abdomen with conspicuous timbals. Guyalna Boulard & Martinelli, 1996 can be distinguished by the head being as wide or slightly wider than the mesonotum, eyes that only protrude slightly beyond the anterior angles of the pronotum, parallel sides of the abdomen, and the median uncus lobes are bent forming a horse-shoe shape when viewed from the posterior. Dorisiana Metcalf, 1952 can be distinguished by its head being as wide or wider than the mesonotum with the eyes not extending beyond the lateral edges of the pronotum, triangular timbal covers, ventral margin of the timbal cover angled dorsally not parallel to the long body axis, and the median uncus lobes are bent. Pacarina Distant, 1905 can be distinguished by the non-parallel radial and radiomedial crossveins, the vertex at the ocelli being much longer than the front, and the triangular timbal covers that cover the majority of the timbal. Similary, the genus Ollanta Distant, 1905 can be distinguished by the lobate lateral angles of the pronotal collar, the non-parallel radial and radiomedial crossveins, and triangular timbal covers concealing the majority of the timbal. Cracenpsaltria n. gen. lacks the posteriorly extending median uncus lobes of Ariasa instead possessing separate, long hooked lateral uncal lobes, the pygofer distal shoulder is angled in the new genus, lacks the laterally expanding abdominal segments, and the length of the abdomen is greater than the distance between the apex of the head and posterior cruciform elevation.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D41F783FFF62FC768ADDFD21.taxon	distribution	Distribution. The genus is found over much of South America including references to Argentina, Bolivia, Brazil, Colombia, French Guiana, Guyana, Paraguay, Surinam, Uruguay, and Venezuela (Metcalf 1963; Sanborn 2011 a; 2011 b; 2013; 2016 a; Sanborn & Heath 2014). It is doubtful that all of the six undetermined species attributed to the genus by Pogue (1996) in Peru are species of Ariasa but confirmed records from Peru are provided here.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D41F783FFF62FC768ADDFD21.taxon	discussion	Key to the species of Ariasa	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D41D7834FF62FF6D8BA0FEA3.taxon	description	(Fig. 1)	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D41D7834FF62FF6D8BA0FEA3.taxon	materials_examined	Type material. Holotype. “ Bonda / Colombia / Acc. No. 1999 // May ” male (CMNH). Paratypes. Same data as holotype, three females (CMNH); one male and two females (AFSC); “ Bonda / Colombia / Acc. No. 1999 // June ” one male and two females (CMNH).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D41D7834FF62FF6D8BA0FEA3.taxon	etymology	Etymology. The species is named for the white (L. alba white) spots (L. maculosa spotted) on the lateral mesothorax.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D41D7834FF62FF6D8BA0FEA3.taxon	description	Description. Ground color castaneous marked with black and tawny. Head. Head wider than mesonotum, medial area black surrounding ocelli extending anterolaterally along posterior epicranial suture almost to lateral supra-antennal plate, black anterior, medial and posterior to eye, anterior and medial black areas not connected. Tawny along posterior epicranial suture, between posterior cranial depression and posterior eye extending anteriorly to lateral ocelli, ovoid mark on anterior vertex, and lateral frontoclypeal suture, tawny regions of anterior vertex and frontoclypeal suture expanded in some paratypes. Supra-antennal plate black with green posteromedial margin with postclypeus and anterolateral margin. Covered with short golden pile dorsally, longer silvery pile posterior to eye. Ocelli rosaceous, eyes castaneous. Ventral head ground color, transverse black mark surrounding eye extending medially toward postclypeus, tawny posteriorly from postclypeus to eye and along margin with lorum. Lorum tawny along margins. Thick white pubescence and long white pile on lorum and gena. Postclypeus centrally sulcate, sulcus tawny on dorsum and apex, black in middle, and castaneous on posterior ventral region, tawny region green on apex and expanding to medial triangle along frontoclypeal suture in some paratypes. Postclypeus with eleven transverse grooves, short golden pile on dorsum, thick white pubescence and long silvery pile on lateral postclypeus. Anteclypeus castaneous with black laterally and posterior margin covered with thick white pubescence and long silvery pile. Mentum tawny, labium proximal half tawny striped with castaneous, darkening to piceous tip reaching to hind trochanters with sparse short and long silvery pile. Scape castaneous with tawny medial side, pedicel castaneous with black tip, antennal flagella black. Thorax. Dorsal thorax castaneous. Pronotum with black mark on either side of midline expanding anteriorly and posteriorly into triangular forms continuing to encircle scutes in ambient fissure, black within lateral fissure, black mark in paramedian fissure not reaching midline extending posteriorly onto disc from middle of fissure, tawny anterior margin. Pronotal collar green with castaneous posterior margin, black mark on margin of pronotal collar lateral angle. Pronotum covered with short golden pile, longer pile laterally. Mesonotum castaneous, lighter along parapsidal suture, cruciform elevation light castaneous with lateral green regions between arms, wing groove green laterally and posteriorly, lateral and submedian sigillae black, lighter mark extending anteriorly from terminus of anterior arm of cruciform elevation to posteromedial lateral sigillae, mark extends to parapsidal suture in some paratypes. White pubescent spot on anterolateral margin of mesonotum. Metanotum castaneous with green posterior margin. Long, dense golden pile on mesonotum, more dense on anterior margin, laterally and posteriorly and between arms of cruciform elevation, in wing groove, and on posterior metanotum. Ventral thoracic segments with thick white pubescence and long white pile, tawny with castaneous basisternum 2 and 3. Wings. Fore wings and hind wings hyaline with eight and six apical cells respectively. Venation castaneous, costa tawny to node, becoming black distally, proximal half of median vein, arculus, proximal cubitus anterior to distal curve, cubitus posterior + anal vein 1, and anal vein 2 + 3 green, black mark between radius & subcostal vein, posterior extension from anal vein 2 + 3 castaneous with black posterior margin. Anterior three quarters and proximal margin of basal cell clouded. Pterostigma to three quarters the length of apical cell 1. Basal membrane of fore wing white at base becoming dark gray distally. Venation of hind wing cataneous except green cubitus anterior, cubitus posterior, and anal veins 1 and 2. Anal cell 3 and anal cell 2 along anal vein 3 dark gray, anal cell 1 and 2 along anal vein 2 white as is anterior cubital cell 2 along proximal seven-eighths of cubitus posterior vein. Small infuscation on proximal cubital cells 1 and 2. Legs. Legs light castaneous, with greenish on distal coxa, femora and proximal tibiae, trochanters striped with castaneous. Fore femora with proximal spine oblique, secondary spine largest and upright and small tertiary spine, all castaneous and connected by a castaneous fascia. Fore tibiae proximally, middle tibiae proximal half and all but distal end of hind tibiae green. Tarsi castaneous, tibial spurs and comb castaneous with green base. Operculum. Male operculum triangular with smoothly rounded posterolateral margin, reaching to anterior of sternite II, tawny with light castaneous base, covered with white pubescence. Medial margin extending to just mediad of meracanthus not covering tympanal cavity medially. Meracathus pointed, tawny with a castaneous base not reaching to posterior margin of operculum. Female operculum with rounded posterolateral margin, terminating medially at middle meracanthus reaching to anterior of sternite II, tawny with a light castaneous base, covered with thick white pubescence. Female meracanthus almost as long as operculum, tawny with a castaneous base. Opercula with long silvery pile radiating from edge. Abdomen. Abdomen castaneous covered with long golden pile, especially dense laterally. Tergites with medial transverse black mark on anterior tergites, additional transverse black marks posterior to timbal cavity on tergite 2 and on lateral tergites 5 and 6, membrane between tergites tawny. Timbal cover incomplete exposing timbal dorsally, anterior margin of tergite 2 curled posteriorly with golden pile, angled at dorsal base and slightly arching to rounded anterior apex, ventral margin straight and parallel to long body axis. Female abdominal segment 9 with castaneous dorsolateral surfaces and stigma, tawny ventrally and posteriorly, with dense, long, golden pile. Posterior margin of abdominal segment 9 sinuate, black on dorsomedial curve and posterior of dorsal beak. Sternites and epipleurites tawny except castaneous sternite I and longitudinal castaneous mark along posterior midline of sternite VIII in males, male sternites II – VII translucent. Female sternite VII with sinuate posterior margin and medial notch, light castaneous spot laterally in some paratypes. Sternites and epipleurites with white pubescence, very dense laterally, and long silvery pile. Genitalia. Male pygofer castaneous, darker along midline, laterally, and on posterior margin, tawny between pygofer basal lobes, with rounded distal shoulder, dorsal beak absent, with long silvery pile posteriorly. Pygofer basal lobe indistinct, pygofer upper lobes tawny, elongated, separated from pygofer, flattening and twisting to curved apex, short golden pile radiating medially. Anal styles castaneous, anal tube tawny. Median uncus lobes castaneous, flattend laterally with rounded apex and upturned at the posterior terminus, lateral uncus lobe ventral side straight as triangular extension is folded under posterior extension of median uncus lobes, basal lateral uncus lobes rounded and separated, long golden pile radiating dorsally and laterally. Aedeagus tubular, dark castaneous. Female gonapophyses VIII and IX castaneous extending beyond dorsal beak with radiating golden pile. Gonocoxite IX tawny or light castaneous. Anal styles castaneous. Measurements (MM). N = 3 males or 7 females, mean (range). Length of body: male 31.00 (30.0 – 32.0), female 28.77 (27.1 – 30.8); length of fore wing: male 40.03 (38.9 – 40.8), female 39.70 (37.2 – 43.6); width of fore wing: male 12.30 (11.9 – 12.8), female 11.89 (11.5 – 12.8); length of head: male 4.83 (4.8 – 4.9), female 4.82 (4.6 – 5.1); width of head including eyes: male 11.57 (11.2 – 12.0), female 11.86 (11.1 – 12.8); width of pronotum including suprahumeral plates: male 11.87 (11.5 – 12.3), female 11.77 (10.9 – 12.9); width of mesonotum: male 9.87 (9.7 – 10.1), female 9.93 (9.4 – 10.7).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D41D7834FF62FF6D8BA0FEA3.taxon	diagnosis	Diagnosis. The lateral red abdomen promptly distinguishes A. nigrorufa from the new species. The lateral ochraceous areas of the abdomen, ochraceous pronotal collar and costal margin distinguish A. arechavaletae from the new species. The white basal area of the hind wings and unicolorous dorsal abdominal segments are unique to A. albiplica. The smaller body size (19 mm vs greater than 30 mm in males) simply distinguishes A. diupsilon and A. urens from the new species. The green and brown body, the white pubescent spots lateral to the genitalia, and smaller body size (about 28 mm) simply distinguish A. alboapicata from the new species. The smaller body size (about 28 mm), black basal cell of the fore wing, and primarily black mesothorax distinguish A. nigrovittata from the new species. Both A. egregia and A. maryannae differ from the new species in that they possess a variegated pattern of silvery and black pile on the dorsal abdomen that form an arch and have black in the basal cell of the fore wing. A black arch formed from pile is found on the dorsal abdomen of A. bilaqueata and A. bartletti n. sp. along with black marking in the basal cell of the fore wing. Ariasa russelli also has a black arch formed from pile on the dorsal abdomen along with primarily black abdominal tergites and black in the basal cell all of which distinguish it from the new species. The new species is most similar in general appearance to A. colombiae with which it was confused in the past (Sanborn 2010). However, A. colombiae can be distinguished by its smaller body size (25 – 28 mm vs greater than 30 mm for males), the pronotal collar is tawny rather than green, lack of obvious white spots on the anterolateral mesonotum, the basal cell of the fore wing has a dark tawny spot, lateral silvery pile on the abdomen rather than the golden pile of the new species, and the uncus is longer and more robust in A. colombiae than in the new species.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D41D7834FF62FF6D8BA0FEA3.taxon	distribution	Distribution. The species is known only from the type series collected in northern Colombia.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187834FF62FEDF8BF3FD05.taxon	materials_examined	Type locality. Brazil.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187834FF62FEDF8BF3FD05.taxon	discussion	Remarks. The white basal areas of the hind wings and unicolorous black abdomen are unique to this species within the genus.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187834FF62FEDF8BF3FD05.taxon	distribution	Distribution. Specimens have been reported previously only from Brazil (Metcalf 1963).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187834FF62FCBE8EEAFB75.taxon	materials_examined	Type locality. Argentina.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187834FF62FCBE8EEAFB75.taxon	discussion	Remarks. The white spots on the lateral regions of tergite 8 are unique to this species in the genus. The species is known currently only from Argentina (Metcalf 1963; Duffels & van der Laan 1985; Sanborn 2013) having been collected in most of the north-central region of the country (Sanborn & Heath 2014). The species has been associated with the Chaco and Espinal floristic provinces in Argentina (Sanborn, et al. 2011 a; Sanborn & Heath 2014).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187834FF62FCBE8EEAFB75.taxon	distribution	Distribution. Specimens have been reported previously only from Argentina (Metcalf 1963; Sanborn 2013; Sanborn & Heath 2014).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187834FF62FAAE8EB3F92D.taxon	materials_examined	Type locality. Carmelo and Montevideo, Uruguay.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187834FF62FAAE8EB3F92D.taxon	discussion	Remarks. Within the genus, the species is unique in possessing an ochraceous pronotal collar, lateral abdominal tergites and costal margin. The species has been associated with the Chaco and Espinal floristic provinces in Argentina (Sanborn, et al. 2011 a; Sanborn & Heath 2014).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187834FF62FAAE8EB3F92D.taxon	distribution	Distribution. Specimens have been reported previously from Argentina and Uruguay (Metcalf 1963; Sanborn 2013; Sanborn & Heath 2014).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187838FF62F9568B01FD72.taxon	description	(Fig. 2)	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187838FF62F9568B01FD72.taxon	materials_examined	Type material. Holotype. “ PERU Madre de Dios / nr Puerto Maldonado / Posadas Amazonas / S 12 48.115 W 69 18.019 / 609 ft; (30 – IX) – (3 – X) – 2004 / C R Bartlett ” female (USNM). Paratypes. Same data as holotype, two females (UDCC); one male and one female (AFSC); “ PERU Madre de Dios / nr Puerto Maldonado / Posadas Amazonas / lodge @ Rio Tambopata / S 12 48.115 W 69 18.019 / 609 ft; (7 – 10) – X – 2004 / C R Bartlett ” one female (UDCC) (this specimen is missing the left wings and the right wings are detached but it was captured with a net and has all its pile and pubescence giving a better representation of the species than those specimens captured in light traps so it is included in the type series). A female was chosen as the holotype based on the apparent preference for females to be attracted to light traps as has been seen in some other cicadas (Young & Sanborn 2015) and thus will be the most likely sex to need determination.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187838FF62F9568B01FD72.taxon	etymology	Etymology. The species is named for Charles Bartlett who collected the type series and has provided numerous specimens to the author over the years.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187838FF62F9568B01FD72.taxon	description	Description. Ground color castaneous marked with black and tawny. One of the paratype female specimens is tinged with green and the male paratype is darker with more expansive black markings. These two specimens appear to have been captured with a net rather than in the light traps so there is no discoloration due to the preservative fluid and are covered with more dense pile. Head. Head wider than mesonotum, longitudinal black mark on either side of medial epicranial suture extending from posterior of head, widening to enclose the lateral ocellus, and extending medially to the frontoclypeal suture, mark angles anterolaterally to anterior vertex lateral to supra-antennal plate, continuing on ventral head laterally to eye where the mark splits and encircles the eye. Vertex lighter lateral to lateral ocelli and on anterior margin. Supra-antennal plate tawny with black spot on anteromedial margin. Covered with short silvery pile dorsally, longer silvery pile posterior to eye. Ocelli rosaceous, eyes castaneous. Ventral head ground color with transverse black mark between antenna and eye. Lorum black except castaneous spot on anteromedial region and lateral posterior margin. Thick white pubescence and long white pile on lorum and gena. Postclypeus centrally sulcate, sulcus with elongated tawny spot on dorsum and apex, black on ventral sulcus and within transverse grooves, transverse ridges castaneous. Postclypeus with thirteen transverse grooves, short golden pile on dorsum, thick white pubescence and long silvery pile on lateral postclypeus. Anteclypeus black with elongated castaneous mark on anterior three fourths of midline covered with thick white pubescence and long silvery pile. Mentum tawny with small lateral black spot near apex, labium proximal half tawny striped with castaneous, darkening to piceous tip reaching to hind trochanters with sparse short silvery pile. Antennae black except for castaneous annulus on proximal scape. Thorax. Dorsal thorax castaneous. Pronotum with black mark on either side of midline angled laterally towards anterior extending from ambient fissure to medial pramedian fissure. Elongate transverse black spot on either side of anterior midline. Black within anterior two thirds of lateral fissure, black mark in paramedian fissure continuing in ambient fissure to encircle disc. Pronotal collar ground color. Pronotum covered with short silvery pile, sparse short black pile on disc between medial ends of paramedian and lateral fissures. Mesonotum castaneous, lighter along parapsidal suture, along midline and connecting to cruciform elevation, wing groove black with tawny lateral margin, triangular black marks on anteromedial lateral sigilla and medial submedian sigilla, posterior submedial sigilla black, medial lanceolate mark, scutal depression and spot anterior to anterior arm of cruciform elevation near terminus black. Marks in submedial sigilla connected by incomplete fascia. Mark in lateral sigillae extends laterally along anterior margin in male paratype. Mark anterior to anterior arm of cruciform elevation extends across anterior arm in some paratypes. Male paratype with white pubescent spot on anterolateral margin of mesothorax. Lateral and submedial sigillae dark castaneous in some paratypes. Metanotum black with castaneous posterior margin. Long, dense silvery pile on mesonotum laterally and posteriorly and between arms of cruciform elevation, in wing groove, and on posterior metanotum, shorter silvery pile on dorsal surface. Ventral thoracic segments castaneous except black katepisternum 2, basisternum 2, meron 2, and episternum 2 with long and short white pile. Wings. Fore wings and hind wings hyaline with eight and six apical cells respectively. Venation castaneous, costa tawny to node, becoming black distally, proximal third of median vein, arculus, proximal third of cubitus anterior, cubitus posterior + anal vein 1, and anal vein 2 + 3 castaneous. Basal cell black with castaneous mark at base, basal half of clavus black. Pterostigma to one third the length of apical cell 1. Basal membrane of fore wing black, greenish in male paratype. Venation of hind wing tawny except green median vein, distal half of cubitus posterior, and middle of anal vein 2, ambient vein, distal radius posterior, and median veins 1 and 2 black. Black mark at base of cubital cells 1 and 2. Anal cell 3 and anal cell 2 along anal vein 3 dark gray, anal cell 1 and 2 along anal vein 2 dark gray. Base of cubital cell and medial anal cell 3 whitish green. Legs. Legs light castaneous, with darker regions on lateral coxae and black on medial coxae, darker spot on medial proximal trochanter and majority of distal trochanter, and femora striped with castaneous. Fore femora with proximal spine oblique, secondary spine largest and upright and small tertiary spine, all castaneous and connected by a black fascia. Fore tibiae proximally tawny then black, middle and hind tibiae castaneous with distal end black. Tarsi black, tibial spurs and comb dark castaneous. Operculum. Female operculum with angled posterolateral margin, terminating medially at middle meracanthus reaching to posterior of sternite I, tawny with a black base and castaneous spot lateral to meracanthus, covered with white pubescence and short white pile radiating from margin. Black cover basal half in one paratype. Female meracanthus as long as operculum, tawny with a black spot on base. Male operculum triangular with smoothly rounded posterolateral margin, reaching to anterior of sternite II, tawny with black base, covered with thick white pubescence and silvery pile radiating from margin. Medial margin extending to medial meracanthus not covering tympanal cavity medially. Meracathus pointed, tawny with a black spot on base not reaching beyond opening of tympanal cavity and covered with thick white pubescence. Abdomen. Abdomen castaneous with dark transverse mark on anterolateral tergite 2 and dorsolateral tergite 3, covered with long silver and black pile, especially dense laterally. Pile more dense in male and female partypes captured with net. Black pile more dense on anterolateral tergites 2 – 4, medial tergites 3, 7 and 8, and dorsolateral tergite 4 – 8 forming a black arch on the dorsal surface. Paratype male tergites with transverse black and castaneous anterior. Female abdominal segment 9 with castaneous dorsolateral surfaces and stigma, tawny ventrally and posteriorly, with dense, long, golden pile. Posterior margin of abdominal segment 9 sinuate, black fascia on dorsolateral surface not reaching posterior margin, a small black spot on either side of base of dark castaneous dorsal beak, connected across midline in one paratype. Timbal cover incomplete exposing timbal dorsally, anterior margin of tergite 2 curved posteriorly with silvery pile, angled at dorsal base and slightly arching to rounded anterior apex, ventral margin straight and parallel to long body axis. Sternites and epipleurites tawny except black anteromedial sternite I and black marks on anterolateral epipleurites, larger on epipleurites 4 – 6. Male paratype also has dark castaneous mark along anteromedial margin across sternite II and across the middle of sternite VII and some female paratypes also have a mark across sternite II and the posterior margin of sternite VII. Female sternite VII with sinuate posterior margin and medial notch, lateral margin arching posteriorly. Male sternites II – VII translucent. Sternites and epipleurites with long silvery pile, more dense laterally and posteriorly, shorter pile radiating from posterior margin. White pubescence, dense laterally, in paratype male and female collected by net. Genitalia. Female gonapophyses VIII and IX black extending beyond dorsal beak with long radiating golden pile. White pubescence on gonapophyses in some paratypes. Gonocoxite IX tawny. Anal styles castaneous. Male pygofer dark castaneous at base, black in middle and tawny posterior margin, dark tawny between pygofer basal lobes, with rounded distal shoulder, dorsal beak absent, with long silvery pile laterally. Pygofer basal lobe indistinct, pygofer upper lobes castaneous, elongated, separated from pygofer, flattening and twisting to curved apex, short golden pile radiating medially at base. Anal styles black, anal tube tawny. Median uncus lobes black with tawny midline, flattend laterally with rounded apex and slightly upturned at posterior terminus, lateral uncus lobe black, tawny laterally, ventral side with tawny triangular extension, basal lateral uncus lobes rounded meeting medially, long golden pile radiating dorsally and laterally. Aedeagus tubular, dark castaneous. Measurements (MM). N = 1 male or 5 females, mean (range). Length of body: male 34.6, female 33.60 (32.7 – 35.0); length of fore wing: male 46.9, female 47.40 (45.9 – 48.5); width of fore wing: male 13.3, female 13.30 (13.0 – 13.6); length of head: male 5.6, female 5.86 (5.5 – 6.2); width of head including eyes: male 13.1, female 13.84 (13.4 – 14.1); width of pronotum including suprahumeral plates: male 13.1, female 13.70 (13.1 – 14.0); width of mesonotum: male 11.3, female 11.80 (11.4 – 12.4).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187838FF62F9568B01FD72.taxon	diagnosis	Diagnosis. The lateral red abdomen promptly distinguishes A. nigrorufa from the new species. Similarly, the lateral ochraceous areas of the abdomen, ochraceous pronotal collar and costal margin distinguish A. arechavaletae from the new species. The white basal area of the hind wings and unicolorous dorsal abdominal segments are unique to A. albiplica. The smaller body size (19 mm vs greater than 32.5 mm) simply distinguishes A. diupsilon and A. urens from the new species. The green and brown body, the white pubescent spots lateral to the genitalia, and smaller body size (about 28 mm) simply distinguish A. alboapicata from the new species. The smaller body size (about 28 mm), black basal cell of the fore wing, and primarily black mesothorax distinguish A. nigrovittata from the new species. The new species can be distinguished from both A. colombiae and A. albimaculosa n. sp. by the more expansive black markings on the mesothorax, the black markings on the adominal tergites crossing the midline and not forming an arch on the dorsal surface, and the basal cell for the fore wing is not black in these species. The transverse band on the head is complete between the epicranial suture and eye, the basal cell of the forewing is black in the distal region only, the pronotal collar is marked with green and a rounded uncal extension is folded under the uncus in A. egregia. The abdominal tergites are mostly black rather than being mainly tawny or castaneous, the pronotal collar is marked with green rather than being castaneous, the basal cell of the fore wing is marked with green proximally rather than being completely black, and the uncus terminus is flattened and bulbous in A. maryannae rather than being rounded as in the new species. The abdominal tergites are mostly black, including completely black tergites 1 and 2, rather than being mainly tawny or castaneous, abdominal sternites are heavily marked with black rather than being tawny, the basal cell of the fore wing is marked with tawny proximally rather than being completely black, and the uncus terminus is upturned at the terminus in A. russelli rather than being straight as in the new species. The most similar species in general appearance to the new species is A. bilaqueata. The new species has more dense pile, particularly on the lateral mesonotum and on the abdomen, the triangular extension in the middle of the ventral uncus is about one fifth the length of the uncus and upright seen as a triangle in lateral view, and female abdominal segment 9 dorsal surface is straight with a slightly upturned dorsal beak while the triangular extension in the middle of the ventral uncus is about one third the length of the uncus and folded mediad so it is not seen in a lateral view and female abdominal segment 9 is smoothly arched dorsally in A. bilaqueata.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4187838FF62F9568B01FD72.taxon	distribution	Distribution. The species is known only from the type series collected in Peru.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4147838FF62FCAF8B46F93C.taxon	materials_examined	Type locality. Chapada, Brazil.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4147838FF62FCAF8B46F93C.taxon	discussion	Remarks. A species that shows morphological similarities to A. bartletti n. sp., A. egregia, A. maryannae, and A. russelli. These species may form a complex within the genus. The species is associated with the Chaco floristic province based on the collection localities of specimens studied from Argentina (Sanborn & Heath 2014). The species extends into other floristic provinces as the Chaco does not cover the range of reported localities for the species.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4147838FF62FCAF8B46F93C.taxon	distribution	Distribution. Specimens have been reported previously from Brazil (Metcalf 1963; Sanborn 2013), Venezuela (Sanborn 2007), and Argentina (Sanborn & Heath 2014). The distribution of the species is expanded here to include Bolivia and French Guiana.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4147838FF62FCAF8B46F93C.taxon	materials_examined	Material examined for new record. “ BOLIVIA: Santa Cruz, 4 km / SSE Buena Vista, Flora / & Fauna Hotel, XI – 2003 / R. F. Morris & J. E. Wappes ” one male (UGCA); “ BOLIVIA: Santa Cruz, 3.7. km / SSE Buena Vista, Hotel Flora / & Fauna 430 m; 5 – 15 – XI – 2001 / coll. M. C. Thomas & B. Dozier / tropical transition forest, BLT ” two females (AFSC); “ Bolivia, Dpto. Pando / Provincia Federico Román / Nueva Esperaza / 10 ° 03 ’ 29.76 ” S / 65 ° 20 ’ 26.54 ” W / Altitud 110 m / 17 / July / 2007 ” one male (AFSC); “ FRENCH GUIANA / Saül Point-de-Vue 13 – X – 15 / Automatic Light Trap, Blue light, Soc. Ent. Antilles-Guyane leg. ” two females (AFSC); “ FRENCH GUIANA / Saül Point-de-Vue 01 – IX – 15 / Automatic Light Trap, Blue light, Soc. Ent. Antilles-Guyane leg. ” four males and four females (AFSC); “ FRENCH GUIANA / Saül Point-de-Vue 15 – IX – 15 / Automatic Light Trap, Blue light, Soc. Ent. Antilles-Guyane leg. ” eleven males and eight females (AFSC).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4147839FF62F9658DE8FEA2.taxon	materials_examined	Type locality. Manaure and Bogota, Colombia and Venezuela.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4147839FF62F9658DE8FEA2.taxon	discussion	Remarks. The species is most similar morphologically to A. albimaculosa n. sp. The species has been associated with the Chaco floristic province in Argentina (Sanborn, et al. 2011 a; Sanborn & Heath 2014) but is associated with other plant communities across the range of the species as the Chaco does not extend to the type locality.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4147839FF62F9658DE8FEA2.taxon	distribution	Distribution. Specimens have been reported previously from Colombia, Venezuela (Metcalf 1963; Sanborn 2013), and Argentina (Sanborn & Heath 2014).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62FEDF8A20FD05.taxon	materials_examined	Type locality. British Guiana.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62FEDF8A20FD05.taxon	discussion	Remarks. One of the two small species in the genus along with A. urens. These species may form a complex within the genus as they share general morphological characteristics as well.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62FEDF8A20FD05.taxon	distribution	Distribution. Specimens have been reported previously from British Guiana and Guiana (Metcalf 1963).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62FCBE8B23FBE1.taxon	materials_examined	Type locality. Rio de Janiero, Brazil.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62FCBE8B23FBE1.taxon	discussion	Remarks. A species recently transferred to the genus (Sanborn 2016 a) that shows morphological similarities to A. bartletti n. sp., A. bilaqueata, A. maryannae, and A. russelli. These species may form a complex within the genus.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62FCBE8B23FBE1.taxon	distribution	Distribution. Specimens have been reported previously from Brazil (Uhler 1903).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62FB028A35FA7E.taxon	materials_examined	Type locality. Piste d'Apatou pk 25, Commune de Saint-Laurent, French Guiana.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62FB028A35FA7E.taxon	discussion	Remarks. A recently described species that shows morphological similarities to A. bartletti n. sp., A. bilaqueata, A. egregia, and A. russelli. These species may form a complex within the genus.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62FB028A35FA7E.taxon	distribution	Distribution. The species is known only from the type series collected in French Guiana (Sanborn 2016 a).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62F9A48E05F8E1.taxon	materials_examined	Type locality. Venezuela.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62F9A48E05F8E1.taxon	discussion	Remarks. The species is unique in the genus with its red lateral abdominal tergites.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4157839FF62F9A48E05F8E1.taxon	distribution	Distribution. Specimens have been reported previously from Colombia and Venezuela (Metcalf 1963; Sanborn 2013).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D412783EFF62FF278CFFFDAC.taxon	materials_examined	Type locality. Central Brazil, Chapada (Brazil), and Bolivia.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D412783EFF62FF278CFFFDAC.taxon	discussion	Remarks. The large proportion of black on the head and mesothorax along with the transverse black fascia on the anterior pronotal collar are unique to the species within the genus. The species has been associated with the Chaco floristic province in Argentina (Sanborn, et al. 2011 a; Sanborn & Heath 2014) being collected over much of the north-central region of the country (Sanborn & Heath 2014).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D412783EFF62FF278CFFFDAC.taxon	distribution	Distribution. Specimens have been reported previously from Argentina, Bolivia, Brazil (Metcalf 1963; Sanborn 2013; Sanborn & Heath 2014), and Paraguay (Sanborn 2011 b).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D412783EFF62FDD58A35FC29.taxon	materials_examined	Type locality. Mont Saint Marcel de la Haute-Camopi, N 2 ° 23 ' 03.00 " O 53 ° 00 ' 37.00 ", altitude 635 m, Commune de Camopi, French Guiana.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D412783EFF62FDD58A35FC29.taxon	discussion	Remarks. A recently described species that shows morphological similarities to A. bartletti n. sp., A. bilaqueata, A. egregia, and A. maryannae. These species may form a complex within the genus.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D412783EFF62FDD58A35FC29.taxon	distribution	Distribution. The species is known only from the type series collected in French Guiana (Sanborn 2016 a).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D412783EFF62FC5A8A20FAEB.taxon	materials_examined	Type locality. British Guiana.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D412783EFF62FC5A8A20FAEB.taxon	discussion	Remarks. One of the two small species in the genus along with A. diupsilon. These species may form a complex within the genus as they share general morphological characteristics and are of similar size.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D412783EFF62FC5A8A20FAEB.taxon	distribution	Distribution. Specimens have been reported previously from British Guiana and Guiana (Metcalf 1963).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D413783DFF62FD558EA5FA59.taxon	description	(Fig. 3)	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D413783DFF62FD558EA5FA59.taxon	type_taxon	Type species. Cicada marginata Olivier 1790: 755 (Surinam).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D413783DFF62FD558EA5FA59.taxon	etymology	Etymology. The genus name is a combination of cracens (L. slender, graceful) and psaltria (L. musician) in reference to the slender appearance of the genus in comparison to other members of the Fidicinini. The genus is feminine.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D413783DFF62FD558EA5FA59.taxon	description	Description. Medium sized species (body length 26 – 35 mm). Head wider than mesonotum, eyes protruding beyond anterior pronotum, vertex at area of ocelli about the same length as front, postclypeus globose, centrally sulcate, rounded anteriorly, rostrum reaching the hind trochanters. Pronotum shorter than mesonotum, trapezoidal with anterior margin narrower than lateral margins of pronotal collar, lateral angles of pronotal collar expanded, lateral part of pronotal collar smoothly curved, metanotum extends laterally beyond wing groove, cruciform elevation smoothly arched posteriorly. Fore wings and hind wings hyaline, with eight and six apical cells, respectively, radial and radiomedial crossveins almost parallel, obliquely oriented to radius posterior and median veins respectively and without infuscation. Basal cell longer than broad, infuscated in anterior half. Basal area of hind wing costal cell infuscated. Fore femora with oblique primary spine that may be adpressed against femur, upright secondary spine, and very small tertiary spine, tarsi three-segmented. Male operculum triangular not covering the tympanal cavity completely posterolaterally, reaching to sternite II, opercula well separated along midline, meracanthus tapering to a point, almost reaching posterior margin of operculum. Female operculum and meracantus of similar shape to male, opercula smaller extending medially only to lateral base of meracanthus which extends beyond posterior margin of operculum. Abdomen longer than the distance between apex of head and posterior of cruciform elevation. Abdominal segments with parallel sides to tergite 4 where the abdomen begins narrowing posteriorly to the genitalia, epipleurites folded toward dorsal surface producing a channel on the lateral margins of the abdomen. Female sternite VII with sinuate posterior margin and medial notch. Timbal cover incomplete, with rounded anterior apex, concealing only the lateral aspect of the timbal, timbal cover with concave anteromedial margin smoothly curving towards rounded apex above lateral timbal, ventral margin slightly curved ventrad, timbals extend below wing bases. Pygofer distal shoulder angled, dorsal beak absent, pygofer basal lobe indistinct, pygofer upper lobe large, pointed, adpressed to pygofer, median uncus lobe indistinct, lateral uncus lobes meeting along midline posterior to median uncus and restraining aedeagus, elongated and recurved, widening in distal third before angling to a pointed terminus, aedeagus tubular. Female abdominal segment 9 with dorsal beak well defined and sinuate posterior margin, ovipositor sheath barely extends beyond dorsal beak. Measurements (MM). Length of body: 24.5 – 34; length of fore wing: 32 – 42; width of fore wing: 10 – 13; length of head: 3.7 – 5; width of head including eyes: 9.5 – 11; width of pronotum including suprahumeral plates: 10 – 17; width of mesonotum: 8 – 10.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D413783DFF62FD558EA5FA59.taxon	diagnosis	Diagnosis. The only species in this genus was classified last within the Ariasa but it has significantly different genitalia and a thinner appearance than the species of Ariasa. The contrasting coloration pattern of the type species is also different from Ariasa species. This becomes the 14 th genus in the subtribe Guyalnina. Within the Fidicinini, Cracenpsaltria n. gen. can be distinguished from Fidicina Amyot & Audinet-Serville, 1843, Fidicinoides Boulard & Martinelli, 1996, and Berglana Boulard & Martinelli, 1996 (the Fidicinina) by the presence of three-part tarsi. The bright green basal area of the fore wings and bright red on the basal area of the hind wings of Hemisciera Amyot & Audinet-Serville, 1843 simply distinguish it from all other Fidicinini. Beameria Davis, 1934 can be distinguished by its adpressed pronotal margins, non-parallel radial and radiomedial crossveins, relatively small eyes, small anterolaterally extended timbal covers, and the sternite separating the male opercula. Majeorona Distant, 1905 differs from the new genus by its head width being wider than the mesonotum, with the prominent eyes stylate and extended anteriorly. The genera Proarna Stål, 1864, Prasinosoma Torres, 1963, Tympanoterpes Stål, 1861, can be distinguished by their head that is about as wide as the mesonotum with the eyes barely or not extending beyond the anterior angle of the pronotum, radial crossvein vertically oriented to the radius anterior 2 and radius posterior veins with a radiomedial crossvein that is obliquely oriented, large semicircular male opercula, and triangular or small, curved timbal covers. The single species of Elassoneura Torres, 1964 shares most of the characters in the three previous genera but the head is wider than the mesonotum and there is significant infuscation along the majority of the fore wing venation. The single species of the genus Pompanonia Boulard, 1982 can be distinguished by possessing a head that is wider than the mesonotum, the radial crossvein vertically oriented to the radius anterior 2 and radius posterior veins with a radiomedial crossvein that is obliquely oriented, semicircular male opercula, triangular timbal covers, and the very wide base of the abdomen with conspicuous timbals. Members of Guyalna Boulard & Martinelli, 1996 can be distinguished from the new genus by a head being as wide or slightly wider than the mesonotum, eyes that only protrude slightly beyond the anterior angles of the pronotum, parallel sides of the abdomen, and the median uncus lobes are bent forming a horse-shoe shape when viewed from the posterior. Species of Dorisiana Metcalf, 1952 can be distinguished by a head being as wide or wider than the mesonotum with the eyes not extending beyond the lateral edges of the pronotum, triangular timbal covers, ventral margin of the timbal cover angled dorsally not parallel to the long body axis, and the median uncus lobes are bent. Species of Pacarina Distant, 1905 can be distinguished by the nonparallel radial and radiomedial crossveins, the vertex at the ocelli being much longer than the front, and the triangular timbal covers that cover the majority of the timbal. Similary, the members of the genus Ollanta Distant, 1905 can be distinguished by the lobate lateral angles of the pronotal collar, the non-parallel radial and radiomedial crossveins, and triangular timbal covers that conceal the majority of the timbal. The new genus shares many characteristics with Ariasa but the genitalia are significantly different lacking the posteriorly extending median uncus lobes and possessing separate, long recurved lateral uncus lobes, the pygofer distal shoulder is angled not curved, the abdominal segments do not expand laterally, and the length of the abdomen is greater than the distance between the apex of the head and posterior cruciform elevation.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D413783DFF62FD558EA5FA59.taxon	distribution	Distribution. Specimens of the type species have been reported from Brazil, Surinam, South America (Metcalf 1963; Sanborn 2013) and French Guiana (Sanborn 2011 a). The genus is also recorded for the first time below from Bolivia and Peru.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4117822FF62F98A8AD7FB83.taxon	materials_examined	Type locality. Surinam.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4117822FF62F98A8AD7FB83.taxon	discussion	Remarks. Boulard (2001) describes in detail the identity, history and synonymy of the species and came to the conclusion that Cicada marginata Olivier, 1790 was a valid species and was assigned to Ariasa. However, after reviewing Sulzer (1761) it is clear that Kirkaldy (1909) was correct in his interpretation of the validity of C. marginata Olivier because it does not matter that Latreille changed the criteria of the genus Cicada L. when Kirkaldy provided the nomen novum as Boulard (2001) suggested. Although Sulzer (1761) used a colloquial name in the text (p. 95 of the first section of the book) to describe his C. marginata, the name marginata is clearly identified as being associated with Cicada L. from Systema Naturae in the second section of the book containing the figure legends (p. 25) and both sections refer to the same figure (Fig. 66, Tab. X) so that the description and the name correspond to the same species. The name C. marginata Sulzer, 1761 meets the criteria of publication, the criteria of availability, the use of the Latin alphabet (in a font distinct from the remaining text and matching the font of the genus Cicada), the name is used as a binomen, there is a description of the species, and the name was valid when proposed meaning the name is available under the Code of Zoological Nomenclature (ICZN 1999). Therefore, C. marginata Sulzer has priority over C. marginata Olivier and Kirkaldy was correct in assigning a nomen novum for Olivier’s species as the name Cicada marginata was preoccupied when proposed by Olivier. As a result, Cracenpsaltria brasiliorum (Kirkaldy, 1909) rev. stat., n. comb. is the valid name for the taxon with the erection of the new genus here.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4117822FF62F98A8AD7FB83.taxon	distribution	Distribution. Specimens have been reported from previously from Brazil, Surinam, South America (Metcalf 1963; Sanborn 2013), and French Guiana (Sanborn 2011 a). The range of the species is extended to include Boliva and Peru with the new records reported here.	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
039D8245D4117822FF62F98A8AD7FB83.taxon	materials_examined	Material examined for new record. “ Rurrenabaque / Beni River // 450 mt. / Bolivia, 1956 // LEPena col. ” (two males FSCA; one male AFSC); “ BOLIVIA: Santa Cruz / Potrerillos del Guenda Res. / Elev. 1322 ft. / 17 ° 40.262 ’ S 63 ° 27.445 ’ W / 6 – 12 January 2005 / J. V. McHugh lab. exped. ” (one female UGCA); “ Prov. del Sara / Bol. Steinbach // Carn. Mus. / Acc. 5604 ” (one female CMNH); “ Prov. del Sara / Bolivia, 450 m. / J. Steinbach // Carn. Mus. / Acc. 6443 ” (four females CMNH, two females AFSC); “ Ituralde Prov., Dpto La Paz, San Miguel, 14 ° 30.602 ' S 67 ° 29.555 ' W, 30 – IX – 2007 ” (a specimen UCMS); “ PERU Madre de Dios / nr Puerto Maldonado / Posada Amazonas / lodge @ Rio Tambopata / S 12 48.115 W 69 18.019 / 609 ft; (30 – IX) – (3 – X) – 2004 / C R Bartlett ” (two males UDCC); and “ PERU Madre de Dios / nr Puerto Maldonado / Posada Amazonas / lodge @ Rio Tambopata / S 12 48.115 W 69 18.019 / 609 ft; (7 – 10) – X – 2004 / C R Bartlett ” (one male UDCC, one male AFSC).	en	Sanborn, Allen F. (2016): Generic redescription, two new species, and a key to the species of the cicada genus Ariasa Distant, 1905 with the description of a new genus (Hemiptera: Cicadidae: Cicadinae: Fidicinini). Zootaxa 4137 (4): 501-519, DOI: 10.11646/zootaxa.4137.4.3
