taxonID	type	description	language	source
039A9827FF9AFFF0CBD7895DFDCEEC85.taxon	materials_examined	Material examined One female, cl 3.4 mm, URB, Doline No. 19 (Cave), Minami Daito Island, coll. K. Kinjo, 18 March 2000; 1 specimen, cl 3.5 mm, ZRC 2004.0562, Cave Gushiken-Dou, Minami Daito Island, Ryukyu Islands, coll. T. Naruse et al., 3 November 2002; 1 male, cl 3.9 mm, ZRC 2004.0563, Cave Mizuana, Minami Daito Island, Ryukyu Islands, coll. T. Naruse et al., 3 November 2002. Diagnosis Rostrum sharp, unarmed, reaching to middle of second segment of antennular peduncle. Antennal spine fused with inferior orbital angle. Pterygostomian margin with a spine. Telson normally with two pairs of dorsal spines and its distal margin rounded, with no posteromedian projection. Eyes with cornea strongly reduced, with a small spot of pigment. Posterior end of caridean lobe of first maxilliped truncated, with no long curved setae. Exopods present on all pereiopods, epipods on first four pereiopods. First pereiopod with carpus excavated deeply anteriorly, twice as long as high, slightly longer than merus but shorter than chela; finger as long as palm. Second pereiopod with carpus excavated anteriorly, four times as long as high, longer than merus and chela, finger longer than palm. Third pereiopod with propodus more than 2.5 times as long as dactylus, dactylus ending in two claws, flexor margin with three spines. Fifth pereiopod with propodus more than twice as long as dactylus. Endopod of male first pleopod with an appendix interna at its anterior margin. Appendix interna of male second pereiopod longer than appendix masculina. Uropodal diaeresis with one long spine. Habitat Anchialine caves. Remarks	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF9AFFF0CBD7895DFDCEEC85.taxon	distribution	Distribution Disjunct distribution in Indo-West Pacific.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF9DFFFDCBFB89B2FE8AEB8F.taxon	description	(Figures 1 – 3)	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF9DFFFDCBFB89B2FE8AEB8F.taxon	materials_examined	Material examined One male, cl 5.4 mm, 1 female, cl 7.3 mm, ZRC 2004.0564, upper stream of Tima River, about 4 km from river mouth, Okinawa Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng, T. Naruse and S. Islam, 11 June 2000; 1 male, cl 5.6 mm, 1 female, cl 7.5 mm, 2 ovigerous females, cl 6.6 – 6.8 mm, URB, Henan River, Okinawa Island, Ryukyu Islands, 17 April 1999; 12 females, cl 3.5 – 6.8 mm, 3 males, cl 3.8 – 4.6 mm, Yakukachi River, Amamiohshima, Ryukyu Islands, 29 July 1992; 5 males, cl 5.0 – 5.6 mm, 2 females, cl 5.3 – 7.1 mm, 5 juveniles, ZRC 2004.0565, fast flowing water, upper part of Omija River, Iriomote Island, Ryukyu Islands, coll. Y. Cai and T. Naruse, 15 June 2000; 3 females, cl 7.2 – 7.9 mm, ZRC 2004.0566, Okuma River, Okinawa Island, Ryukyu Islands, 31 May 1998. Comparative material examined	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF9DFFFDCBFB89B2FE8AEB8F.taxon	description	Description Rostrum (Figures 1 A and 3 A) reaching to or slightly beyond end of scaphocerite, dorsal margin nearly horizontal, or slightly sigmoid, armed with 16 – 25 (mode 17 – 24) teeth throughout dorsal margin, two or three of them situated on carapace posterior to orbital margin, armed ventrally with one to six (mode one to three) teeth. Antennal spine slightly lower than inferior orbital angle; pterygostomian margin subrectangular. Sixth abdominal somite 0.65 times length of carapace, 1.6 times as long as fifth somite, subequal to length of telson. Telson (Figure 1 B, C) 3.2 times as long as wide, not terminating in a posteromedian projection, with two pairs of dorsal spinules and one pair of dorsolateral spinules; dorsal pairs situated near edges; distal end broadly rounded, with about 11 setae, lateral pair slightly longer than intermediates. Preanal (Figure 1 J) carina with a spine. Eyes well developed, anterior end reaching to 0.8 times length of basal segment of antennular peduncle. Antennular peduncle 0.70 times as long as carapace; basal segment of antennular peduncle distinctly longer than sum of second and third segment lengths, anterolateral angle reaching 0.30 times length of the second segment, second segment distinctly longer than third segment. Stylocerite reaching to 0.7 times length of second segment of antennular peduncle. Scaphocerite (Figure 1 D) 3.0 times as long as wide. Incisor process of mandible (Figure 1 E) ending in irregular teeth, molar process truncated. Lower lacinia of maxillula (Figure 1 F) broadly rounded, upper lacinia elongate, with a number of distinct teeth on inner margin, palp slender. Upper endites of maxilla (Figure 1 G) subdivided, palp short, scaphognathite tapering posteriorly with numerous long, curved setae at posterior end. Palp of first maxilliped (Figure 1 H) ending in a fingerlike projection. Second maxilliped (Figure 1 I) typical. Third maxilliped (Figure 2 A) reaching to end of antennular peduncle, with ultimate segment slightly shorter than penultimate segment. Epipods on first four pereiopods. Exopod on all pereiopods. First pereiopod (Figures 2 B and 3 B) reaching to distal end of eyes; merus 3.2 times as long as broad, as long as carpus; carpus excavated anteriorly, shorter than chela, 1.6 times as long as high; chela 1.6 times as long as broad; fingers shorter than palm. Second pereiopod (Figures 2 C and 3 C) reaching to end of second segment of antennular peduncle; merus shorter than carpus, 5.0 times as long as broad; carpus longer than chela, 5.0 times as long as high; chela 3.0 times as long as broad; fingers slightly longer than palm. Third pereiopod (Figures 2 D, E and 3 D, E) reaching to end of scaphocerite, propodus 11 times as long as broad, 4.2 times as long as dactylus; dactylus 2.1 times as long as wide (spines included), terminating in one claw, with five to seven accessory spines on flexor margin. Fifth pereiopod (Figures 2 F, G and 3 F, G) reaching slightly beyond end of second segment of antennular peduncle, propodus 11 times as long as broad, 4.3 times as long as dactylus, dactylus 2.7 times as long as wide (spinules included), terminating in two claws, with 61 spinules on flexor margin. Endopod of male first pleopod (Figure 3 H) elliptical, 2.2 times as long as wide, one quarter length of exopod; appendix interna elongated, with most of its length reaching beyond distal end of endopod. Appendix masculina of male second pleopod (Figure 3 I) very small, appendix interna much longer than appendix masculina. Uropodal diaeresis (Figure 1 K) with 1 movable spinule. Eggs 0.35 – 0.45 × 0.24 – 0.30 mm in diameter. Habitat Lower reaches of rivers and streams which discharge to the sea. Remarks De Haan (1849) described Paratya compressa under the name of Ephyra, from Japan with no specific locality being mentioned. Nishino (1981) showed the distributions of two forms of P. compressa, one with a large egg, in central Honshu and the other, with a small egg, in southern Japan. Specimens reported by Kemp (1917) as Paratya compressa are most probably an undescribed species. According to Kemp (1917, p 296), his material was from Komatsu Lake near the eastern shore of Lake Biwa and from the Ogura and Yodo ponds near Kyoto, Honshu. ‘‘ The dactylar spines (of third pereiopod) vary in number from 19 to 22, very rarely 18. In the fifth pereiopods (text figs. Ie, f) the propodus is also from 2.1 to 2.5 times as long as dactylus. ’’ This description, however, is very different from the type material of P. compressa. Dr. C. Fransen, curator of the Leiden Museum, checked the syntypes on the request of the first author, as they are all too fragile to be posted. Result of his examination revealed that of the eight syntypes of Paratya compressa (RMNH D 1007), ‘‘ only two specimens have a third pereiopod attached. In one specimen the ratio propodus / dactylus (of third pereiopod) is 4.7, the number of spines on the carpus of the dactylus (of third pereiopod) seven. In the other (smaller) specimen, the ratio propodus / dactylus is 4.0 and the number of spines on the carpus of the dactylus five’ ’ (C. Fransen, pers. commun.). A review study on the basis of specimens from the whole distribution range is necessary to clarify their identities. Kemp (1917) proposed a subspecies Paratya compressa improvisa (Figure 4) for the specimens from Lake Haruna, Honshu Island, Japan. It differs from the nominal subspecies mainly on the basis of the rostral teeth arrangement. It has no teeth on the carapace while those of P. compressa have two to three. But as shown in Figures 1 – 4, the propodus and dactylus of the male third pereiopods are very different in the two subspecies and very easy to separate from each other, we hereby recognize both as distinct species. Distribution Southern Japan.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF90FFF8CB578EBBFCF2ECD7.taxon	materials_examined	Material examined Holotype: male, cl 2.4 mm, URB- 497, well near Kaneshi, Okinawa Island, Ryukyu Islands, coll S. Shokita, November 1971. Allotype: 1 female, cl 2.8 mm, URB- 498, well near Kaneshi, Okinawa Island, Ryukyu Islands, coll S. Shokita, November 1971. Paratypes: 2 males, cl 2.0 – 2.4 mm, URB- 499, well near Kaneshi, Okinawa Island, Ryukyu Islands, coll S. Shokita, November 1971. Others: 1 specimen, URB, Shiokawa, Okinawa Island, Ryukyu Islands, coll T. Nagai, March 1997; 4 female cl 3.0 – 3.6 mm (RUB-ZC- 78 - 81), 1 male cl 2.6 mm (RUB-ZC- 82), Hatoma Island, Yaeyama Group, Ryukyu Islands, coll. M. Toda, 18 October 2002. Diagnosis Rostrum short, not reaching beyond end of eyestalk; unarmed. Antennal spine fused fully with inferior orbital angle. Pterygostomian margin broadly rounded. Telson not terminating in a projection, with only two pairs of dorsal spinules on distal two-thirds of telson; with two pairs of distal spines, lateral pair of distal spines distinctly longer than intermediates. Antennular peduncle 0.55 times as long as carapace, stylocerite not reaching to end of basal segment of antennular peduncle. Epipods on first three pereiopods. Merus of first pereiopod as long as ischium; carpus of first pereiopod deeply excavated, subequal to chela in length, 3.3 times as long as high, chela 2.3 times as long as broad, fingers distinctly longer than palm. Merus of second pereiopod as long as ischium; carpus of second pereiopod excavated anteriorly, very slender, seven times as long as high, more than twice longer than chela; chela 2.5 times as long as broad, fingers 1.6 times as long as palm. Propodus of third pereiopod three times as long as dactylus, dactylus 2.5 times as long as wide, with eight spines on flexor margin. Propodus of fifth pereiopods three times as long as dactylus, dactylus 3.5 times as long as broad, with 18 – 19 spinules on flexor margin. Endopod of male first pleopod subrectangular, no appendix interna, slightly shorter than exopod. Appendix masculina of male second pleopod slender. Uropodal diaeresis with four movable spinules. Habitat Anchialine caves. Remarks	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF90FFF8CB578EBBFCF2ECD7.taxon	distribution	Distribution Known from the Ryukyu Islands, Japan, Palau, and Guam.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF95FFF9CB9B8861FEA6EEC1.taxon	materials_examined	Material examined Neotype of Caridina brevirostris: 1 male, cl 6.8 mm, NSMT, River Aritsu, Kume-Jima Island, Ryukyu Islands, coll. A. Kawakami, 8 December 1995. Others: 2 males, cl 6.2 – 7.1 mm, URB, Omija River, Iriomote Island, Ryukyu Islands, coll. T. Naruse, 1 August 1999. Habitat Lower reaches of rivers and streams which discharge to the sea. Remarks Cai et al. (2006) recently designated neotypes for C. acuminata Stimpson 1860 from the Ogasawara (Bonin) Islands and Caridina brevirostris Stimpson 1860 from Kume Island, Ryukyus, and synonymized both species with Atyoida pilipes. Atyoida pilipes has been reported from the Philippines and eastern Lesser Sunda Islands at about 120 east longitude eastward through the Pacific high island (Chace 1983). In Japan, it has been reported from Kume Island, Ishigaki Island (Shokita 1979) Iriomote Island of the Ryukyu Islands (Shokita 1997). Distribution West Pacific.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF94FFF9CBE78A76FEA6EB6D.taxon	materials_examined	Material examined Seven males, cl 10 – 15 mm, 4 females, cl 15 – 18 mm, 2 ovigerous females, cl 14 – 15 mm, ZRC 2004.0567, 124 ° 15.269 E 24 ° 30.119 N, Gaburumata River, Ishigaki Island, Ryukyu Islands, pH 7.3, coll. Y. Cai and T. Naruse, 13 June 2000; 1 female, cl 7.1 mm, ZRC, 124 ° 14.809 E 24 ° 22.769 N, Gaburumata River, Ishigaki Island, Ryukyu Islands, pH 7.3, coll. Y. Cai and T. Naruse, 13 June 2000; 1 male, cl 12 mm, 1 female, cl 15 mm, 1 ovigerous female, cl 16 mm, ZRC 2004.0568, 124 ° 14.529 E 24 ° 26.059 N, stream to Tsuru River, Ishigaki Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 17 June 2000; 1 ovigerous female, cl 14.8 mm, Aira River, Iriomote Island, Ryukyu Islands, 8 August 1969; 1 female, cl 7.1 mm, ZRC 2004.0569, Inoda River, Ishigaki Island, coll. N. K. Ng et al., 13 June 2000. Habitat Lower reaches of rivers and streams which discharge to the sea. Remarks Compared to the material of Chace (1983), the number of ventral rostral teeth for this species is more variable in our specimens. It ranges from one to seven. Atyopsis spinipes occurs from Philippines and eastern Lesser Sunda Islands (ca. 120 ° 009 E) northwards to Taiwan and as far as Tokuno-shima in the Ryukyus, and eastwards as far as Samoa (Chace 1983). Distribution West Pacific.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF94FFFACBC78E9DFED5EADB.taxon	materials_examined	Material examined Syntypes: 3 females, cl 7.0 – 8.4 mm, MNHN-Na 930, no date. Neotype of Caridina exilirostris: ovigerous female, cl 7.4 mm, NSMT, Okuma River, Okinawa Island, Ryukyu Islands, coll. P. K. L. Ng et al., 31 May 1998. Others: 3 females, cl 3.0 – 3.2 mm, 4 males, cl 4.4 – 4.7 mm, upstream of Nakama river, Iriomote Island, Ryukyu Islands, coll. M. Tomokuni, 3 November 1985; 1 female, cl 7.7 mm, upper of Kawauchi-gawa River, Amami-oshima Island, Ryukyu Islands, coll. M. Takeda, 15 July 1988; 1 female, cl 54 mm, URB 1230, Sumiyoh-gawa River, Amami-ohshima Island, Ryukyu Islands, coll. M. Takeda, 17 July 1988; 1 male, cl 4.8 mm, 1 ovigerous female, cl 6.9 mm, URB 1112, Shino-kawa River, Amami-oshima Island, Ryukyu Islands, coll. M. Takeda, 17 July 1988; 4 females, cl 5.4 – 6.8 mm, URB 1182, Kawauchi-gawa River, Amami-ohshima Island, Ryukyu Islands, coll. M. Takeda, 15 July 1988; 3 males, 4.9 – 5.9 mm, 4 females, cl 6.5 – 7.5 mm, 2 ovigerous females, cl 6.1 – 6.5 mm, URB 1218, Kawauchi-gawa River, Amami-ohshima Island, Ryukyu Islands, coll. Takeda, 18 July 1988; 1 male, cl 4.2 mm, 2 females, cl 4.8 – 5.2 mm, 2 ovigerous females, cl 5.2 – 5.6 mm, URB 1135, Kanyu-gawa River, Amami-ohshima Island, Ryukyu Islands, coll. M. Takeda, 16 July 1988. Habitat Rivers and streams which discharge to the sea. Remarks Cai et al. (2006) recently designated a neotype for C. exilirostris Stimpson, 1860 from Okinawa Island, Ryukyu Islands, and synonymized it with Caridina typus. This is the most common species found from the Ryukyu Islands. Distribution Indo-West Pacific.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF97FFE4CBFB8E4FFC32ECB1.taxon	materials_examined	Material examined None. Habitat Lower reaches of rivers and streams which discharge to the sea. Remarks This species was recently reported by Shokita (2003). The single specimen on which Shokita’s (2003) record was based could not be located in the present study. Distribution Indo-West Pacific.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF97FFE4CBFB8E4FFC32ECB1.taxon	materials_examined	Material examined Neotype: 1 male, cl 4.9 mm, NSMT, upper stream of Tima River, about 4 km from river mouth, Okinawa Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng, T. Naruse and S. Islam, 11 June 2000. Others: 3 males, cl 3.2 – 3.8 mm, 1 female, cl 4.3 mm, 6 ovigerous females, cl 4.2 – 5.3 mm, ZRC, 128 ° 02.489 E 26 ° 33.489 N, freshwater stream draining to a small patch of mangrove, Oura River, Okinawa Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng, T. Naruse and S. Islam, 11 June 2000; 13 ovigerous females, cl 4.6 – 5.6 mm, 4 females, cl 3.6 – 4.6 mm, 10 males, cl 3.1 – 3.8 mm, ZRC 2004.0519, 128 ° 04.609 E 26 ° 33.429 N, upstream of Tima River, about 1 – 2 km from river mouth, Okinawa Island, Ryukyu Islands, pH 7.4, coll. Y. Cai, N. K. Ng, T. Naruse and S. Islam, 11 June 2000; 46 males, cl 3.6 – 4.6 mm, 12 females, cl 3.6 – 4.9 mm, 1 ovigerous female, cl 4.7 mm, ZRC 2004.0520, 128 ° 05.529 E 26 ° 33.639 N, upstream of Tima River, about 3 km from the river mouth, Okinawa Island, Ryukyu Islands, pH 7.4, coll. N. K. Ng and S. Islam, 11 June 2000; 6 males, cl 3.8 – 5.1 mm, 12 females, cl 4.7 – 6.5 mm, 4 ovigerous females, cl 5.8 – 6.5 mm, ZRC 2004.0521, upper stream of Tima River, about 4 km from river mouth, Okinawa Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng, T. Naruse and S. Islam, 11 June 2000; 20 males, cl 3.5 – 4.4 mm, 1 female, cl 5.2 mm, 25 ovigerous females, cl 5.7 – 6.1 mm, ZRC 2004.0522, 128 ° 07.049 E 26 ° 36.619 N, Arume River, Okinawa Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng, T. Naruse and S. Islam, 11 June 2000; 8 males, cl 2.9 – 4.0 mm, 10 females, cl 5.0 – 5.6 mm, 15 ovigerous females, cl 5.0 – 5.6 mm, ZRC 2004.0523, 124 ° 15.269 E 24 ° 30.119 N, Gaburumata River, Ishigaki Island, Ryukyu Islands, pH 7.3, coll. Y. Cai and T. Naruse, 13 June 2000; 2 males, cl 3.6 – 4.1 mm, 1 female, cl 6.8 mm, 1 ovigerous female, cl 6.1 mm, ZRC 2004.0524, 124 ° 09.809 E 24 ° 24.429 N, fast flowing stream, with pH 7.6 at one of the tributary of Nagura River, below reservoir, Ishigaki Island, Ryukyu Islands, coll. Y. Cai and T. Naruse, 13 June 2000; 2 males, cl 3.9 – 4.0 mm, 1 female, cl 5.6 mm, 1 ovigerous female, cl 5.3 mm, ZRC 2004.0525, 124 ° 09.869 E 24 ° 25.009 N, fast flowing tributary of Nagura River, below a reservoir, Ishigaki Island, Ryukyu Islands, pH 7.2, coll. Y. Cai and T. Naruse, 13 June 2000; 1 female, cl 6.1 mm, ZRC 2004.0526, 123 ° 51.849 E 24 ° 23.659 N, fast flowing water, about 200 m from sea, Omija River, Iriomote Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 15 June 2000; 1 male, cl 4.0 mm, ZRC 2004.0527, fast flowing water, upper part of Omija River, Iriomote Island, Ryukyu Islands, coll. Y. Cai and T. Naruse, 15 June 2000; 6 males, cl 3.6 – 3.7 mm, 8 females, cl 4.8 – 5.5 mm, 3 ovigerous females, cl 5.0 – 5.3 mm, ZRC 2004.0528, 124 ° 33.769 E 24 ° 17.819 N, pH 7.7, Kara Stream, Ishigaki Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 17 June 2000; 1 male, cl 4.3 mm, ZRC 2004.0529, Taiho River, Okinawa Island, Ryukyu Islands, coll. P. K. L Ng et al., 31 May 1998; 2 females, cl 5.0 – 5.6 mm, 2 ovigerous females, cl 5.3 – 6.1 mm, ZRC 2004.0530, Okuma River, Okinawa Island, Ryukyu Islands, coll. P. K. L. Ng et al., 31 May 1998. Habitat Lower reaches of rivers and streams which discharge to the sea. Remarks Cai et al. (2006) recently designated a neotype for C. grandirostris Stimpson, 1860 and redescribed it in detail. Distribution Caridina grandirostris has thus far only been reported from Japan.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF89FFE5CBFB8998FF7BED90.taxon	materials_examined	Material examined Neotype: ovigerous female, cl 7.7 mm, NSMT (NIES-Iz- 1), 138 ° 56.439 E 34 ° 41.839 N, Inouzawa River, freshwater, lotic, EC 134, Shimoda City, Izu Peninsula, coll. K. Satake, 10 Aug 2004. Others: one male, cl 4.4 mm, 1 female, cl 6.3 mm, 2 ovigerous females, cl 5.8 – 5.9 mm, ZRC, Okuma River, Okinawa Island, Ryukyu Islands, 31 June 1998; 14 males, cl 3.5 – 3.9 mm, 1 female, cl 4.0 mm, 7 ovigerous females, cl 4.5 – 5.8 mm, ZRC 2004.0530, 128 ° 02.489 E 26 ° 33.489 N, freshwater stream draining to a small patch of mangrove, Oura River, Okinawa Island, Ryukyu Islands, coll. Y. Cai et al., 11 June 2000; 5 males, cl 3.0 – 4.0 mm, 20 females, cl 4.5 – 5.2 mm, 19 January 1975, Yona River, Okinawa Island, Ryukyu Islands; 1 male, cl 4.8 mm, Aritsu River, Kume-Jima, Ryukyu Islands, coll. A. Kawakami, 8 December 1995; 1 male, cl 4.3 mm, 5 ovigerous females, cl 5.5 – 6.2 mm, ZRC 2004.0531, 128 ° 07.049 E 26 ° 36.619 N, Arume River, Okinawa Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng, T. Naruse and S. Islam, 11 June 2000; 1 male, cl 3.8 mm, 1 ovigerous female, cl 6.5 mm, ZRC, 123 ° 52.809 E 24 ° 16.649 N, shallow freshwater stream of Aira River, Iriomote Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 14 June 2000; 9 males, cl 3.0 – 4.2 mm, 12 ovigerous females, cl 4.7 – 6.5 mm, ZRC 2004.0532, 123 ° 52.749 E 24 ° 16.609 N, shallow freshwater stream, downstream of Aira River, Iriomote Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 14 June 2000; 7 males, cl 3.1 – 3.5 mm, 7 ovigerous females, cl 4.4 – 4.9 mm, ZRC 2004.0533, 123 ° 51.299 E 24 ° 18.399 N, downstream from the headwater of Nakama River, Iriomote Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 15 June 2000; 6 males, cl 4.2 – 4.7 mm, 8 females, cl 5.8 – 6.4 mm, 15 ovigerous females, cl 5.8 – 6.3 mm, ZRC 2004.0534, fast flowing water, upper part of Omija River, Iriomote Island, Ryukyu Islands, coll. Y. Cai and T. Naruse, 15 June 2000; 3 males, cl 2.4 – 3.9 mm, 8 females, cl 4.6 – 6.4 mm, 1 ovigerous female, cl 5.7 mm, ZRC 2004.0535, upper Hiji fall, Hiji River, Okinawa Island, Ryukyu Islands, 4 November 1987; 3 females, cl 5.0 – 5.9 mm, URB 1750, no data; 1 ovigerous female, cl 5.7 mm, URB 1221, Kawauchi-gawa River, Amami-Ohshima Island, Ryukyu Islands, coll. M. Takeda, 15 July 1988; 1 female, cl 3.0 mm, Kanyu-gawa River, Amami-ohshima, Ryukyu Islands, coll M. Takeda, 16 July 1988; 4 males, cl 4.4 – 5.0 mm, 6 ovigerous females, cl 6.6 – 6.8 mm, URB 1111, Kawauchi-gawa River, Amami-oshima, Ryukyu Islands, coll M. Takeda, 18 July 1988; 1 ovigerous female, cl 6.2 mm, URB 1219, Kawauchi-gawa River, Amami-ohshima, Ryukyu Islands, 18 July 1988. Habitat Lower reaches of rivers and streams which discharge to the sea. Remarks Cai et al. (2006) recently designated a neotype for C. leucosticta Stimpson, 1860 and redescribed it in detail. Distribution Caridina leucostica has only been reported with certainty from Japan thus far (Cai et al. 2006).	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF88FFE6CBC288FBFDD7ED4E.taxon	materials_examined	Material examined Neotype: ovigerous female, cl 8.8 mm, NMST, from a stream in Tenno-ura, Chichi Jima Island, Ogasawara Islands, coll. K., Satake, 4 May 2000. Lectotype of Caridina japonica De Man, 1892 b, ZMA De 102876, Japan, coll. J. Anderson, 1881, 4 males, cl 7.8 – 8.2 mm, paralectotype of Caridina japonica De Man, 1892 b, data same as lectotype; 1 male, cl 8.8 mm, paralectotype, MNHN-Na 731, Kagar, Hayagana, Japan. Others: 1 female, cl 9.9 mm, NMST 1745, no data; 1 male, cl 7.5 mm, NMST 1121, upper part of Kawauchigawa River, Amami-oshima Island, Ryukyu Islands, coll M. Takeda, 15 July 1988; 1 ovigerous female, cl 7.9 mm, NMST 1754, Shino-Kawa River, Amami-oshima Island, Ryukyu Islands, coll. M. Takeda, 17 July 1988; 1 male, cl 8.3 mm, 2 ovigerous females, cl 7.3 – 9.8 mm, ZRC 2004.0537, Omija River, Iriomote Island, Ryukyu Islands, coll. T. Naruse, 1 August 1999; 1 ovigerous female, cl 8.2 mm, NMST, Amami-ohshima Island, Ryukyu Islands, 29 July 1992; 1 male, cl 16 mm, 2 ovigerous females, cl 10 – 12 mm, ZRC, 128 ° 15.319 E 26 ° 39.949 N, Arakawa River, Okinawa Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng, T. Naruse and S. Islam, 11 June 2000; 1 female, cl 4.9 mm, ZRC, fast flowing water, upper part of Omija River, Iriomote Island, Ryukyu Islands, coll. Y. Cai and T. Naruse, 15 June 2000; 1 female, cl 9.5 mm, 2 ovigerous females, cl 6.2 – 9.8 mm, ZRC. 2004.0539, Taiho River, Okinawa Island, Ryukyu Islands, coll. P. K. L. Ng et al., 31 May 1998; 1 female, cl 4.9 mm, ZRC 2004.0538, fast flowing water, upper part of Omija River, Iriomote Island, Ryukyu Islands, coll. Y. Cai and T. Naruse, 15 June 2000. Habitat Lower reaches of rivers and streams which discharge to the sea. Remarks Cai et al. (2006) recently designated a neotype for C. multidentata Stimpson, 1860 from Ogasawara (Bonin) Islands and synonymized C. japonica De Man, 1892 with it. Distribution	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF8BFFE7CBF488DDFCE0EC26.taxon	materials_examined	Material examined One ovigerous female, cl 4.2 mm, ZRC 2004.0570, 128 ° 07.049 E 26 ° 36.619 N, Arume River, Okinawa Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng, T. Naruse and S. Islam, 11 June 2000; 1 male, cl 3.2 mm, 2 ovigerous females, cl 4.9 – 5.0 mm, ZRC 2001.0571, 123 ° 52.809 E 24 ° 16.649 N, shallow freshwater stream of Aira River, Iriomote Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 14 June 2000; 1 male, cl 40 mm, 5 ovigerous females, cl 4.8 – 5.3 mm, ZRC 2004.0572, 123 ° 52.749 E 24 ° 16.609 N, shallow freshwater stream, downstream of Aira River, Iriomote Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 14 June 2000; 1 female, cl 4.4 mm, 3 ovigerous females, cl 4.4 – 4.5 mm, ZRC 2004.0573, 123 ° 51.849 E 24 ° 23.659 N, fast flowing water, about 200 meters from sea, Omija River, Iriomote Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 15 June 2000; 1 male, cl 3.4 mm, ZRC 2004.0574, 123 ° 51.849 E 24 ° 23.659 N, fast flowing water, about 200 m from sea, Omija River, Iriomote Island, Ryukyu Islands, coll. Y. Cai and T. Naruse, 16 June 2000. Habitat Lower reaches of rivers and streams which discharge to the sea. Remarks Cai and Shokita (2006) recently discussed the taxonomy of Caridina serratirostris and C. celebensis, and pointed out that ‘‘ In Ryukyus, specimens with short rostrum are mostly of C. celebensis, while those with long rostrum, almost all having arthrobranch on the first pereiopod, are of C. serratirostris. ’’ Distribution Japan, Philippines, Fiji, Malaysia, Indonesia and Madagascar.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF8AFFE0CBFA8932FD83EF9E.taxon	materials_examined	Material examined Two ovigerous females, cl 3.9 – 4.1 mm, ZRC 2004.0575, upper stream of Tima River, about 4 km from river mouth, Okinawa Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng, T. Naruse and S. Islam, 11 June 2000; 1 ovigerous female, cl 4.5 mm, Yakugachi River, Amami-ohshima, Ryukyu Islands, 30 July 1992; 2 ovigerous females, cl 4.8 – 5.3 mm, Tabaru River, Yonaguni Island, coll. S. Shokita, 31 March 2000; 26 ovigerous females, cl 3.2 – 4.1 mm, 7 females, cl 2.1 – 3.4 mm, 2 males, cl 2.4 – 2.5 mm, ZRC 2004.0576, 128 ° 04.609 E 26 ° 33.429 N upstream of Tima River, about 1 – 2 km from river mouth, Okinawa Island, Ryukyu Islands, pH 7.4, coll. Y. Cai, N. K. Ng, T. Naruse and S. Islam, 11 June 2000; 1 ovigerous female, cl 4.7 mm, ZRC, Okuma River, Okinawa Island, Ryukyu Islands, 31 May 1998; 2 females, cl 4.1 – 4.2 mm, ZRC, Tima River, Okinawa Island, Ryukyu Islands, 25 April 1998; 1 female, cl 2.9 mm, ZRC 2004.0577, upper Hiji fall, Hiji River, Okinawa Island, Ryukyu Islands, 4 November 1987. Habitat Lower reaches of rivers and streams which discharge to the sea. Remarks Caridina celebensis was recently reviewed by Cai and Shokita (2006). It is characterised by the absence of an arthrobranch on the base of the first pereiopod. Caridina celebensis is distributed from Sulawesi, Indonesia to Honshu, Japan. Distribution Japan, China, Philippines, Sulawesi.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF8DFFE3CBE98A8BFEE1E885.taxon	description	(Figures 5 and 6 A – E)	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF8DFFE3CBE98A8BFEE1E885.taxon	materials_examined	Material examined One male, cl 4.9 mm, URB, Aritsu River, Kume-jima Island, Ryukyu Islands, coll. A. Kawakami, 8 December 1995; 1 male, cl 3.5 mm, 124 ° 14.809 E 24 ° 22.769 N, Gaburumata River, Ishigaki Island, Ryukyu Islands, coll. Y. Cai and T. Naruse, 13 June 2000; 1 male, cl 4.3 mm, ZRC 2004.0578, 123 ° 52.809 E 24 ° 16.649 N, shallow freshwater stream of Aira River, Iriomote Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 14 June 2000. 1 male, cl 3.8 mm, URB, Omija River, Iriomote Island, Ryukyu Islands, 1 August, 1998, coll. T. Naruse; 1 male, cl 8.1 mm, 1 ovigerous female, cl 7.8, RUB, a small stream near Haemida beach, Iriomote, Ryukyu Islands, coll T. Naruse, 28 July 1999. Description Rostrum straight (Figure 5 A), reaching to base of second segment of antennular peduncle, dorsal margin nearly horizontal, elevated slightly above dorsal margin of carapace, rostral formula 0 + 9 – 17 / 1 – 6, dorsal teeth evenly spaced, all on rostrum considerably anterior to orbital margin. Antennal spine fused with inferior orbital angle; pterygostomian margin rounded. Sixth abdominal somite 0.40 times length of carapace, 1.3 times as long as fifth somite, shorter than telson. Telson (Figure 5 C) 2.5 times as long as wide, terminating in a projection, with five pairs of dorsal spinules and one pair of dorsolateral spinules; distal end with two pairs of spines and three to four pairs of plumose setae, lateral pair of spines distinctly longer than sublateral pair, intermediate pairs of setae distinctly longer than later spines, distal margin broadly rounded. Preanal carina high, without spine (Figure 5 B). Eyes well developed, anterior end reaching to 0.7 times length of basal segment of antennular peduncle. Antennular peduncle 0.52 times as long as carapace; basal segment of antennular peduncle longer than sum of second and third segment lengths, anterolateral angle reaching 0.20 times length of the second segment, second segment distinctly longer than third segment. Stylocerite (Figure 5 D) reaching 0.8 length of basal segment of antennular peduncle. Scaphocerite 2.6 times as long as wide. Incisor process of mandible ending in irregular teeth, molar process truncated. Lower lacinia of maxillula broadly rounded, subtriangular, upper lacinia elongate, with a number of distinct teeth on inner margin, palp slender. Upper endites of maxilla subdivided, palp short, scaphognathite tapering posteriorly with numerous long, curved setae at posterior end. Palp of first maxilliped ending in a finger-like projection. Second maxilliped typical, arthrobranch well developed. Third maxilliped reaching to end of antennular peduncle, with ultimate segment shorter than penultimate segment. Epipods well developed on first four pereiopods. First pereiopod (Figure 5 E) reaching to distal end of basal segment of antennular peduncle; merus 2.1 times as long as broad, as long as, or slightly longer than carpus; carpus excavated anteriorly, shorter than chela, 1.3 times as long as high; chela 2.4 times as long as broad; fingers subequal to length of palm. Second pereiopod (Figure 5 F) reaching to end of second segment of antennular peduncle; merus slightly shorter than carpus, 4.3 times as long as broad; carpus slightly longer than chela, 4.3 times as long as high; chela 2.9 times as long as broad; fingers 1.6 times as long as palm. Third pereiopod (Figure 5 G, H) reaching to end of scaphocerite, propodus 8.3 times as long as broad, 3.7 times as long as dactylus; dactylus 2.2 times as long as wide (spines included), terminating in one stout claw, with five accessory spines on flexor margin. Fifth pereiopod (Figure 5 I, J) reaching to end of antennular peduncle, propodus 9.2 times as long as broad, 3.3 times as long as dactylus, dactylus 3.1 times as long as wide (spinules included), terminating in one elongated claw, with 28 – 42 spinules on flexor margin. Endopod of male first pleopod sub-triangular, reaching to one-thirds length of exopod, appendix interna reaching beyond distal end of endopod by most of its length. Appendix masculina of male second pleopod reaching to two-thirds length of exopod. Uropodal diaeresis (Figure 5 K) with 16 – 18 movable spinules. Eggs 0.40 – 0.42 × 0.20 – 0.25 mm in diameter. Habitat Rivers and streams which discharge to the sea. Remarks Chace (1997) recently redescribed this species on the basis of specimens from Philippines. He (Chace 1997, p 13) also commented that it ‘‘ … is possible that C. laoagensis will eventually fall into synonymy with the variable C. weberi from Indonesia, … ’’ According to an on-going revision by one of the authors (YC), most of the subspecies of C. weberi, are in fact, distinct species. Among all the allied species, C. laoagensis is most similar to C. weberi De Man, 1908 a by the form of the rostrum, which is straight, pointed and crested at the base of the rostrum; the spines and setae on the distal margin of the telson (the longer setae arranged between two pairs of strong spines, of which, the sublateral pair shorter than the lateral pair vs. the longer setae arranged between one pair of strong spines). Caridina laoagesis appears to be rare in Ryukyu Islands. It has previously been reported from Tabaru River of Yonaguni Island, Nagura and Miyara Rivers of Ishigaki Island, and Shigema, Yona and Manna Rivers of Okinawa Island by Shokita (1979, p 2003) as Caridina weberi. Distribution Philippines, Japan	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF81FFEFCBC18BCFFE7BEFD0.taxon	description	(Figures 6 F – J, 7 and 8)	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF81FFEFCBC18BCFFE7BEFD0.taxon	materials_examined	Material examined Holotype: female, cl 6.1 mm (dry), URB 495; Izaga Cave, Miyako Island, Ryukyu Islands, Japan, coll. S. Shokita, 19 September 1964; paratype: 1 female, cl 5.4 mm (dry), URB 496, data same as holotype. Others: 1 male cl 4.8 mm, 7 females, cl 3.9 – 8.8 mm, URB Murikawa, Yamato-ga, Miyako Island, Ryukyu Islands, Japan, 23 October 1982. Diagnosis Rostrum straight or slightly upturned, reaching to end of second segment of antennular peduncle, or to end of scaphocerite. Rostrum with teeth throughout its dorsal margin, rostral formula 9 – 14 + 15 – 24 / 9 – 23. Antennal spine prominent, situated lower than suborbital angle; pterygostomian margin rounded; eyes strongly reduced. Preanal carina with spine. Telson with five pairs of dorsal spines, small posteromedial projection, lateral pair of spines longer than or as long as intermediate pairs of setae. Antennular peduncle 0.5 – 0.6 times as long as carapace. Stylocerite reaching to end of basal segment of antennular peduncle, or slightly beyond it. Scaphocerite 3.1 times as long as wide. First pereiopod with ischium much shorter than merus, carpus 1.0 – 1.5 times as long as chela, 4.2 times as long as high, chela 2.0 – 3.0 times as long as broad, finger distinctly longer than palm. Second pereiopod with ischium much shorter than merus, carpus 1.5 – 2.0 times as long as chela, 8 – 12 times as long as high, chela 2.4 – 4.0 times as long as broad, fingers 1.5 – 2.0 times as long as palm. Third pereiopod with propodus not enlarged, its length 4.0 – 5.0 times as long as dactylus; dactylus ending in two claws, with two to four spines on flexor margin. Fifth pereiopod with propodus 5.0 – 6.5 times as long as dactylus, dactylus ending in one claw, with 13 – 17 spinules on flexor margin. Endopod of male first pleopod sub-triangular, no appendix interna. Uropodal diaeresis with 11 – 13 spinules. Habitat Subterranean water. Remarks The forms of the rostrum, the first two pereiopods and telson are much more variable than in the original description by Fujino and Shokita (1975). Holthuis (1978) described Caridina troglodytes, on the basis of four incomplete specimens from Danmin Cave near Konogusgus, New Ireland. Caridina troglodytes is very similar to C. rubella, most of its characters fall within the range of variation of C. rubella, except for the smaller number of ventral rostral teeth (2 – 8 vs. 9 – 23) and the fewer spinules on the dactylus of the fifth pereiopods (6 vs. 13 – 15). As the ventral rostral teeth are quite variable in both species and the supposed fifth pereiopod is unattached, and thus could be the fourth pereiopod instead, the identity of Holthuis’ species need more specimens for it to be ascertained. Shokita (1979) reported it from two more locations in southern Japan, one from subterranean water of Shiokawa River, Okinawa Island, Ryukyus, and the other from Suirendo Cave, Okinoerabu Island, Ryukyu Islands. Cai and Anker (2004) reported the occurrence of Caridina rubella from Palawan Island, the Philippines. Distribution Ryukyu Islands, Philippines.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF82FFE9CBD28ABBFDA4ED26.taxon	description	(Figure 9)	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF82FFE9CBD28ABBFDA4ED26.taxon	materials_examined	Material examined Paratype of Caridina sakishimensis: 1 female, cl 7.2 mm, URB, Arakawabanna River, Yonaguni Island, Ryukyu Islands, coll. S. Shokita, 27 March 1972. Others: 2 females, cl 3.9 – 4.0 mm, ZRC 2004.0581, fast flowing water, upper part of Omija River, Iriomote Island, Ryukyu Islands, coll. Y. Cai and T. Naruse, 15 June 2000; 1 ovigerous female, cl 6.2 mm, ZRC 2004.0582, coll. A. Kawakami, Aritsu River, Kume-jima, Ryukyu Islands, 8 December 1995; 1 male, cl 3.8 mm, URB, Omija River, Iriomote, Ryukyu Islands, coll. T. Naruse, 1 August 1999; 1 ovigerous female, cl 7.8 mm, 1 male, cl 8.1 mm, URB, a stream flowing to Haemida beach, Iriomote Island, Ryukyu Islands, coll. T. Naruse, July 1999. Diagnosis Rostrum straight, reaching near to middle of second segment of antennular peduncle. Rostral formula 1 – 6 / 3 – 6. Suborbital angle acute, fused with antennal spines; pterygostomian margin rectangular. Preanal carina with no spine. Telson with small posteromedial projection, lateral pair of spines shorter than intermediate pairs. Antennular peduncle 0.6 times as long as carapace. Stylocerite reaching 0.8 times length of basal segment of antennular peduncle. Scaphocerite 2.9 times as long as wide. First pereiopod with carpus 1.1 – 1.3 times as long as high, chela 2.0 times as long as broad, finger distinctly shorter than palm. Second pereiopod with carpus 4.5 times as long as high, chela 2.6 times as long as broad, fingers 1.5 times as long as palm. Third pereiopod with propodus 4.0 – 4.8 times as long as dactylus; dactylus ending in two claws, with two to four spines on flexor margin. Fifth pereiopod with propodus 4.5 – 5.0 times as long as dactylus, dactylus ending in one claw, with 29 spinules on flexor margin. Endopod of male first pleopod sub-triangular, with appendix interna. Uropodal diaeresis with 21 – 22 spinules. Habitat Mountain streams or rivers. Remarks Tiwari and Pillai (1971) described a new species, Caridina prashadi, from the southern Andaman Islands. Fujino and Shokita (1975) did not compare it with C. prashadi when they described C. sakishimensis from Southern Ryukyus. It was subsequently recorded from Kume Island of the northern Ryukyus and Okinoerabu Island of the central Ryukyus (Shokita 1979; Suzuki and Sato 1994). Examination of one fresh collection from the Andaman Islands (2 females, cl 4.0 – 4.1 mm, 1 ovigerous female, ZRC, Mount Harriet National Park, south Andaman Islands, India, coll. Indranei Das, 20 August 1997) and re-examination of the types in the Ryukyu University, and the examination of fresh material from Iriomote Island indicated that C. sakishimensis is, in fact, identical with C. prashadi. The record of C. rapaensis by Shokita (2003) was based on these specimens and should thus be reassigned here. Distribution Andaman Islands, Ryukyu Islands.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF84FFD5CBFA8832FC84EF27.taxon	description	(Figures 10 and 11)	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FF84FFD5CBFA8832FC84EF27.taxon	materials_examined	Material examined One male, cl 2.9 mm (USNM- 264045, holotype of Caridina blancoi), near mouth of Tayabas River, Luzon Island, Philippines, 25 February 1909; 1 male, cl 2.2 mm (SHU- 79 - 310 - 1, holotype of Caridina hainanensis), 1 female, cl 4.0 mm (SHU- 79 - 310 - 2, paratype of C. hainanensis), Wenchang County, Hainan Island, China, coll. X. Liang and S. Yan, 14 April 1979; 11 males, cl 2.6 – 3.3 mm, 6 females, cl 3.5 – 3.8 mm, 16 ovigerous females, cl 3.4 – 4.0 mm, URB, mangrove of Nakama River, Iriomote Island, Ryukyu Islands, coll. S. Shokita, 3 – 5 June 1985; 30 specimens, ZRC 2004.0579, tide-pool in mouth of Shira River, Iriomote Island, Ryukyu Islands, 5 October 1982. Diagnosis Rostrum (Figure 10 A, G) straight or sloping ventrally anteriorly, reaching near middle of second segment of antennular peduncle, to end of third segment. Rostral formula: 3 – 4 + 9 – 16 / 1 – 7 / 1 – 6 (3 – 4), no apical teeth. Suborbital angle acute, distinctly separated from antennal spines; pterygostomian margin rounded. Preanal carina (Figure 10 B) with a spine. Telson (Figure 10 C, D) with three to four pairs of dorsal spines and small posteromedial projection, lateral pair of spines longer than intermedial pairs. Antennular peduncle long, subequal to carapace length. Stylocerite reaching 0.8 times length of basal segment of antennular peduncle. Scaphocerite (Figure 10 F) 4.0 times as long as wide. First pereiopod (Figure 11 A) with carpus 2.3 times as long as high, chela 2.5 times as long as broad, finger longer than palm. Second pereiopod (Figure 11 B) with carpus 6.8 times as long as high, chela 4.1 times as long as broad, fingers 2.0 times as long as palm. Third pereiopod (Figure 11 C, D) with propodus 3.2 times as long dactylus, propodus without enlargement. Dactylus ending in two claws, with one to four spines on flexor margin. Fifth pereiopod (Figure 11 E, F) with propodus 2.5 times as long as dactylus; dactylus ending in one claw, with 57 – 76 spinules on flexor margin. Endopod of male first pleopod with no appendix interna. Uropodal diaeresis with 13 – 19 spinules. Egg size: 0.38 – 0.48 × 0.25 – 0.30 mm. Habitat Lower reaches of rivers or mountain streams which discharge to the sea. Commonly found in mangrove creeks. Remarks Liang and Yan (1983) described Caridina hainanensis on the basis of specimens from Hainan Island. Their species has the same typical characteristics as C. propinqua, namely the form of the rostrum and the telson, the form of various pereiopods, especially the third, which has a very small number of spines (one to three in C. hainanensis vs. one to four in C. propinqua) on the flexor margin of the dactylus, the identical form of the endopod of male first pleopod. Re-examination of the types indicated that C. hainanensis should be regarded as a junior synonym of C. propinqua. Chace (1997) proposed a new species, based on a single specimen. He commented that ‘‘ (t) he proposal of a new species, based on a single specimen in a genus that is noteworthy for its variable species, may be questionable, but it seems desirable to call attention to a taxon that apparently differs from all others known in a combination of characters; the form and dentition of the rostrum and telson; the prominence of the suborbital angle; and the form of the chelae and carpi of two anterior pereiopods and of the dactyls of the third and fourth pereiopods. ’’ Re-examination of the holotype confirms that it is identical with C. propinqua. Egg size of this species is quite variable. It is 0.39 – 0.45 × 0.24 – 0.27 mm in specimens from Hainan (Liang and Yan 1983), 0.54 × 0.36 mm in populations from Malaysia (Johnson 1961) and 0.38 – 0.48 × 0.25 – 0.30 mm in the present specimens from the Ryukyus. Kemp (1918) reported C. propinqua from southern Thailand. The identity of his species is doubtful as it had an egg size of 0.64 – 0.70 × 0.39 – 0.44 mm, which is much larger than in specimens from elsewhere. This record has recently been reported by Shokita (2003). Previously, Caridina propinqua has been reported from Sri Lanka (De Silva 1982), India (De Man 1908 b; Kemp 1915), Malaysia (Johnson 1961; Ng and Choy 1990), southern China (Liang and Yan 1983) and the Philippines (Chace 1997). Distribution Sri Lanka, India, Malay Peninsula, Philippines, and China.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FFB8FFD7CBE98A2BFC15ECA5.taxon	description	(Figure 12) Material examined Holotype: female, cl 6.5 mm, USNM- 123426, Ingue Cave at Yomitan Village, Okinawa Island, Ryukyu Islands, Japan, coll. D. W. Rhodes, 3 March 1967. Description Rostrum (Figure 12 A) short, straight, reaching near end of second segment of antennular peduncle; dorsal margin with 12 teeth, three of them on carapace, ventral margin with five very small teeth; inferior orbital angle of carapace fused with antennal spine, sharp and long; pterygostomial angle rectangular. Sixth abdominal somite 0.50 times as long as carapace, 1.6 times as long as fifth somite, slightly shorter than telson. Telson terminating in median projection; five pairs of dorsal spinules, one pair of dorsolateral spines near distal end, four pairs of spines on distal margin, lateral pair shortest, sublateral pair longest; preanal carina high, with a small spine. Scaphocerite 2.8 times as long as wide. Eyestalk short, eye with an undeveloped cornea. Antennular peduncle stout, 0.5 times as long as carapace; stylocerite reaching slightly beyond end of basal segment of antennular peduncle. Third maxilliped reaching to end of third segment of antennular peduncle, with ultimate segment shorter than penultimate segment. Epipods on first four pereiopods. First pereiopod (Figure 12 C) stout, reaching to middle of second segment of antennular peduncle, ischium much shorter than merus, merus 2.0 times as long as broad, as long as carpus; carpus excavated anteriorly, shorter than chela, 1.2 times as long as high; chela 1.9 times as long as broad; fingers 0.6 times as long as palm. Second pereiopod (Figure 12 D) reaching end of second segment of antennular peduncle, ischium much shorter than merus, merus slightly longer than carpus, 5.4 times as long as broad; carpus 1.3 times as long as chela, 5.0 times as long as high; chela 2.4 times as long as broad; fingers 1.3 times as long as palm. Third pereiopod (Figure 12 E, F) reaching end of third segment of antennular peduncle, propodus normal shape, distinctly shorter than merus, 9.0 times as long as broad, 4.5 times as long as dactylus; dactylus ending in two strong claws; 2.6 times as long as wide (spines included), with five accessory spines on flexor margin. Fifth pereiopod (Figure 12 G, H) reaching to end of basal segment of antennular peduncle, propodus distinctly longer than merus, 11 times as long as broad, 4.7 times as long as dactylus; dactylus 2.6 times as long as wide, ending in two large claws, with 26 spinules on its flexor margin. Uropodal diaeresis with 17 spinules. Habitat The specimen was collected from subterranean water. Etymology The new species is named after the type locality, Okinawa Island, Ryukyus, used as a noun in apposition. Remarks With respect to the rostral formula, and the long stylocerite, Caridina okinawa, new species, is very different from all the epigean Caridina species from the Ryukyus. It is however, most similar to C. cantonensis Yu, 1938 (cf. Cai and Ng 1999) from southern China. It could be distinguished by the shorter stylocerite (reaches to the end of basal segment of antennular peduncle vs. distinctly beyond); the less developed eyes (vs. well developed in C. cantonensis); the shorter finger of the first pereiopod (0.6 times as long as palm vs. as long as palm). Distribution Known only from the type locality, Okinawa Island, Ryukyu Islands.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FFBAFFD3CBC989B2FCBCEC8F.taxon	description	(Figures 13 and 14)	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FFBAFFD3CBC989B2FCBCEC8F.taxon	materials_examined	Material examined Holotype: Ovigerous female, cl 4.8 mm, eggs 0.25 × 0.45 mm, NMST, 123 ° 51.849 E 24 ° 23.659 N, fast flowing water, about 200 m from sea, Omija River, Iriomote Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 15 June 2000. Paratypes: One female, cl 4.8 mm, RUB, 1 ovigerous female, cl 4.8 mm, ZRC 2004.0583, eggs 0.50 × 0.30 mm, small river in Ohno, Ishigaki Island, Ryukyu Islands, coll. T. Naruse, 23 March 2001. Comparative material examined One ovigerous female, cl 6.1 mm, NMMBA, Dongqing river, Lanyu Island, Taiwan, coll. C. C. Han, 19 April 2002; 1 damaged female, cl 5.2 mm, ZRC, Sungai Ifis, Halmahera, Indonesia, coll. D. Robb, September 1994; 1 ovigerous female, cl 4.7 mm, ZRC, Philippines, coll M. Kottelat, 1993; 1 female, cl 5.5 mm, SMF 947 a, Mariveles Island, Philippines, coll. Semper, 1876. Description Rostrum (Figures 13 A and 14 A) short, straight, with a pointed end, reaching middle of second segment of antennular peduncle; dorsal margin with 12 – 15 large teeth, two to three of them on carapace, ventral margin with three to five very small teeth; inferior orbital angle of carapace fused with antennal spine, sharp, long; pterygostomial angle broadly rounded. Sixth abdominal somite 0.41 times as long as carapace, 1.2 times as long as fifth somite, slightly shorter than telson. Telson (Figure 13 B, C) 2.8 times as long as wide, terminating in median projection; five to seven pairs of dorsal spinules, one pair of dorsolateral spines near distal end, one pair of spines and three pairs of subequal spiniform setae on distal margin; lateral spine slightly shorter than intermediate pairs of setae. Preanal carina high, without spine. Eyes (Figures 13 A and 14 A) well developed, reaching to 0.8 times length of basal segment of antennular peduncle. Antennular peduncle stout, 0.6 times as long as carapace; basal segment of antennular peduncle longer than sum of second and third segment lengths of antennular peduncle, anterolateral angle reaching 0.2 times length of second segment; second segment slightly longer than third. Stylocerite reaching 0.9 times length of basal segment of antennular peduncle. Scaphocerite (Figure 14 B) 2.8 times as long as wide. Incisor process of mandible (Figure 13 D) ending in irregular teeth, molar process truncated. Lower lacinia of maxillula (Figure 13 E) broadly rounded, subtriangular, upper lacinia elongate, with a number of distinct teeth on inner margin, palp slender. Upper endites of maxilla (Figure 13 F) subdivided, palp short, scaphognathite tapering posteriorly with numerous long, curved setae at posterior end. Palp of first maxilliped (Figure 13 G) terminating in broad triangular end. Second maxilliped (Figure 13 H) typical, arthrobranch well developed. Third maxilliped (Figure 13 I) reaching to end of antennular peduncle, with ultimate segment shorter than penultimate segment. Epipods on first four pereiopods. First pereiopod (Figure 14 C) stout, reaching to end of basal segment of antennular peduncle, merus 2.0 times as long as broad, slightly longer than carpus; carpus strongly excavated anteriorly, shorter than chela, 1.2 times as long as high; chela 2.0 times as long as broad; fingers as long as palm. Second pereiopod (Figure 14 D) reaching end of second segment of antennular peduncle, merus slightly longer than carpus, 4.4 times as long as broad; carpus slightly longer than chela, 3.8 times as long as high; chela 2.5 times as long as broad; fingers 1.7 times as long as palm. Third pereiopod (Figure 14 E, F) reaching end of third segment of antennular peduncle, propodus distinctly shorter than merus, 8.5 times as long as broad, 4.3 times as long as dactylus; dactylus ending in two strong claws; 2.5 times as long as wide (spines included), with three accessory spines on flexor margin. Fifth pereiopod (Figure 14 G, H) reaching to end of basal segment of antennular peduncle, propodus distinctly longer than merus, 11 times as long as broad, 4.7 times as long as dactylus; dactylus 2.4 times as long as wide, ending in two large claws, with 26 spinules on flexor margin. Uropodal diaeresis (Figure 14 J) with 19 – 20 spinules. Eggs 0.25 – 0.30 × 0.45 – 0.50 mm in diameter. Habitat The holotype specimen was caught from a fast flowing stream, hiding under vegetation at the edge of the stream. The collection site is about 200 m from the sea. Etymology The name is derived from Latin, macro, large, and dentata, toothed, alluding to the large teeth on the dorsal margin of the rostrum. Remarks With regard to the form of the rostrum, the large number of uropodal spinules and the form of various pereiopods, Caridina macrodentata should be referred to the C. weberi species group. In the group, it is most similar to C. weberi papuana, which, however, can be separated from C. weberi papuana by its longer rostrum, the larger rostral teeth and the biunguiculate dactylus of the fifth pereiopod. The new species is also close to C. okinawa, new species. The rostral formula, the form of the preanal carina, and pereiopods are shared only by these two species in the Ryukyus. Caridina okinawa was found from subterranean water in Okinawa, central Ryukyus, while C. macrodentata was caught from epigean waters in Iriomote, southern Ryukyus. Caridina macrodentata differs from C. okinawa by its well developed eyes, larger teeth on the dorsal margin of the rostrum, the relatively broad scaphocerite (2.5 time as long as wide vs. 2.8 times); the longer fingers of the first pereiopod (as long as palm vs. 0.6 times), the longer fingers of the second pereiopod (1.7 times as long as palm vs. 1.3 times) and the shorter stylocerite (does not reach the end of basal segment of antennular peduncle vs. reaching). Cai and Ng (2001) doubtfully referred a damaged specimen from Halmahera, Indonesia, to Caridina pareparensis De Man, 1892 a. According to the available characters, especially the rostrum and the first pereiopod, it most probably belongs to the present new species, and is formally referred to it here. Distribution Japan, Taiwan, Philippines and Indonesia (Halmahera).	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FFBEFFD8CB6289BBFD9DEFF2.taxon	description	(Figures 15 – 17)	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FFBEFFD8CB6289BBFD9DEFF2.taxon	materials_examined	Material examined Twenty six males, cl 3.3 – 4.4 mm, 8 females, cl 3.0 – 5.2 mm, ZRC 2004.0584, 124 ° 12.749 E 24 ° 25.199 N, freshwater stream connected to Sokobaru Dam Reserve, Ishigaki Island, Ryukyu Islands, coll. Y. Cai, N. K. Ng and T. Naruse, 13 June 2000; 1 ovigerous female, cl 4.9 mm, ZRC 2004.0585, 124 ° 09.869 E 24 ° 25.009 N, fast flowing tributary of Nagura River, below a reservoir, Ishigaki Island, Ryukyu Islands, pH 7.2, coll. Y. Cai and T. Naruse, 13 June 2000; 1 male, cl 4.0 mm, ZRC, 123 ° 51.219 E 24 ° 18.249 N, freshwater, very small, shallow stream, headwater of Nakama River, Iriomote Island, Ryukyu Islands, pH 7.4, coll. Y. Cai, N. K. Ng and T. Naruse, 15 June 2000; 9 males, cl 3.5 – 5.0 mm, 8 females, cl 4.0 – 5.2 mm, ZRC 2004.0586, 124 ° 12.649 E 24 ° 25.989 N, river below Sokobaru Dam Reserve, Ishigaki Island, Ryukyu Islands, pH 7.5, coll. Y. Cai and T. Naruse, 13 June 2000. Description Rostrum (Figures 15 A and 17 A) narrow, straight, reaching to middle of second segment of antennular peduncle, or slightly beyond end of this segment, rostral formula: 1 – 2 + 1 + 10 (mode 5 – 7) / 0 – 4 (mode 1 – 2). Antennal spine fused with inferior orbital angle; pterygostomian margin with a spine. Sixth abdominal somite 0.49 times length of carapace, 1.4 times as long as fifth somite, as long as telson. Telson (Figure 16 A, B) 3.0 times as long as wide, terminating in a projection, with five to six pairs of dorsal spinules and one pair of dorsolateral spinules; distal end with about four to five pairs of spines, lateral pair as long as or slightly longer than intermediate pairs, sublateral pair shortest. Preanal carina (Figure 20 K) without spine. Eyes well developed, anterior end reaching to 0.8 times length of basal segment of antennular peduncle. Antennular peduncle 0.57 – 0.63 times as long as carapace; basal segment of antennular peduncle longer than sum of second and third segment lengths, anterolateral angle reaching 0.25 times length of the second segment, second segment distinctly longer than third segment. Stylocerite reaching to 0.85 – 0.9 length of basal segment of antennular peduncle. Scaphocerite (Figure 15 B) 2.8 times as long as wide. Incisor process of mandible (Figure 15 C) ending in irregular teeth, molar process truncated. Lower lacinia of maxillula (Figure 15 D) broadly rounded, upper lacinia elongate, with a number of distinct teeth on inner margin, palp slender. Upper endites of maxilla (Figure 15 E) subdivided, palp short, scaphognathite tapering posteriorly with numerous long, curved setae at posterior end. End of palp of first maxilliped (Figure 15 F) truncate. Podobranch of second maxilliped (Figure 15 G) well developed. Third maxilliped (Figure 15 H) reaching to end of scaphocerite, with ultimate segment as long as or slightly shorter than penultimate segment. Epipods present on first four pereiopods. First pereiopod (Figures 16 C and 17 B) reaching to distal end of basal segment of antennular peduncle; merus 2.6 – 2.7 times as long as broad, slightly longer than carpus; carpus excavated anteriorly, shorter than chela, 1.6 – 1.8 times as long as high; chela 2.1 – 2.3 times as long as broad; fingers 1.1 – 1.4 times as long as palm. Second pereiopod (Figures 16 D and 17 C) reaching to end of antennular peduncle; merus slightly shorter than carpus, 5.1 – 5.3 times as long as broad; carpus 1.15 – 1.25 times as long as chela, 5.0 – 5.1 times as long as high; chela 2.6 – 2.9 times as long as broad; fingers 1.4 – 1.7 times as long as palm. Third pereiopod (Figures 16 E, F and 17 D, E) reaching to end of scaphocerite, propodus incurved strongly in male but straight in female, 7.4 – 8.4 times as long as broad, 3.4 – 4.2 times as long as dactylus; dactylus 2.0 times as long as wide in male and 2.6 times in female (spines included), terminating in two claws, with three to four accessory spines on flexor margin. Fifth pereiopod (Figures 16 G, H and 17 F, G) reaching beyond end of basal segment of antennular peduncle, propodus 8.6 – 10 times as long as broad, 3.1 – 3.3 times as long as dactylus, dactylus 3.0 – 3.3 times as long as wide (spinules included), terminating in one claw, with 25 – 46 spinules on flexor margin. Endopod of male first pleopod (Figure 16 I) inflated at distal three-quarters into palmshape, with numerous tiny spinules on distal portion of dorsal surface; appendix interna at base of endopod, small, elongate, or reduced, 1.5 times as long as wide, 0.7 times length of exopod. Appendix masculina of male second pleopod (Figure 16 J) cylindrical, reaching to half length of endopod, armed with numerous stout spines on surface, with appendix interna reaching to distal quarter of appendix masculina or near its distal end. Uropodal diaeresis (Figure 16 K) with 19 – 20 movable spinules. Eggs 1.1 × 0.7 mm in diameter. Habitat Mountain streams. Remarks Fujino and Shokita (1975) described it as a subspecies of Caridina denticulata (De Haan, 1849), on the basis of specimens from Ishigaki Island. Shokita (1979) raised it to a full species, Neocaridina ishigakiensis. In his revision of the genus Neocaridina, Cai (1996) treated it as a subspecies of N. palmata (Shen, 1948), as the sexual appendages are very similar to the nominal subspecies. Apart from the differences in the rostrum, the appendix interna of the male first pleopod is also shorter than that of N. palmata. While N. ishigakiensis is morphologically closer to N. palmata than N. denticulata (cf. Cai 1996), we nevertheless follow Shokita (1979) in regarding it as a distinct species as recent investigations have shown that it is restricted to Ishigaki Island, and there are no intermediate forms, and no range overlap in the distributions of N. ishigakiensis and N. palmata. Liang (2004) recently reported the occurrence of Neocaridina denticulata ishigakiensis from Wenling, Zhejiang Province. His drawing (Liang 2004, fig. 42) shows that sexual appendages of his specimens are much more like those of Neocaridina sinensis than N. ishigakiensis. Distribution Several localities on Ishigaki Island.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
039A9827FFB5FFD8CB7F8966FD5AEBCE.taxon	materials_examined	Material examined Five males, cl 3.3 – 4.8 mm, 1 female, cl 5.0 mm, 3 ovigerous females, cl 4.1 – 5.0 mm, ZRC 2004.0587, headwater of Nakama River, Iriomote Island, Ryukyu Islands, Cai et al., 16 June 2000; 19 males, cl 3.4 – 5.1 mm, 7 females, cl 4.0 – 5.2 mm, 4 ovigerous females, ZRC 2004.0588, slightly downstream of the head water area of Nakama River, Ishigaki Island, Ryukyu Islands, Cai et al., 16 June 2000; 2 males, cl 4.4 – 4.8 mm, 1 female, cl 5.8 mm, ZRC 2004.0589, upper stream of Nakama River, Ishigaki Island, Ryukyu Islands, Cai et al., 2 males, cl 4.1 – 4.2 mm, 1 ovigerous female, cl 4.5 mm, ZRC, Iriomote Island, Ryukyu Islands, Cai et al, 15 June 2000; 27 males, cl 3.2 – 4.3 mm, ZRC 2004.0590, headwater of Nakama river, Ishigaki Island, Ryukyu Islands, coll. Y. Cai and T. Naruse, 2000. Habitat Headwater of Nakama River, Iriomote Island. Remarks This species has been described in detail, with comparisons being made against several congeners (see Naruse et al. 2006). Distribution Known only from the type locality, Iriomote Island.	en	Cai, Y. (2006): Atyid shrimps (Crustacea: Decapoda: Caridea) of the Ryukyu Islands, southern Japan, with descriptions of two new species. Journal of Natural History 40 (38 - 40): 2123-2172, DOI: 10.1080/00222930601084049, URL: http://dx.doi.org/10.1080/00222930601084049
