identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039B87FDCC75C73B9D99DF1CFA71FC29.text	039B87FDCC75C73B9D99DF1CFA71FC29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera Chaowasku 2012	<div><p>Hubera Chaowasku, gen. nov. Type: ⎯ Hubera cerasoides (Roxburgh 1795: 30) Chaowasku.</p> <p>Shrubs, treelets or medium-sized trees, evergreen or rarely deciduous (so far observed only in H. cerasoides). Young twigs glabrous or hairy. Leaves petiolate; elliptic(-oblong), oblong or seldom obovate, base equal or rarely unequal, attenuate, wedge-shaped, obtuse or rarely (sub)cordate, apex usually obtuse, acute(- acuminate) or (caudate-)acuminate, rarely rounded or emarginate; upper surface of midrib (slightly) sunken, flat, or (both edges) slightly raised [with one or two groove(s) in the middle]; domatia of aggregated tuft of hairs present in some species, at the axils where secondary veins meet primary veins; tertiary venation reticulate. Inflorescences usually on branches and axillary, one-flowered or seldom two-flowered, rarely cauliflorous and then&gt; 3-flowered; pedicels often articulated above the base; bracts (and bracteoles) present or sometimes not seen (absent?). Flowers bisexual; sepals free or rarely basally connate, triangular to ovate; petals of the outer and inner whorls equal or slightly subequal, ovate, elliptic(-ovate) or (elliptic-)tongueshaped, erect or spreading at anthesis; in dried condition often marked by yellow powdery material. Stamens generally ≥ 40 per flower, apex of anther connective flat-topped, covering the thecae. Carpels generally ≥ 10 per flower, cylindrical or occasionally flask-shaped, peripheral ones sometimes slightly incurved; stigmas ± globose, (ellipsoid-)obovoid, ellipsoid(-ovoid) or ellipsoid-cylindrical, generally ± hairy; ovaries hairy, 1 ovule per ovary, sub-basal. Torus (nearly) flat or (slightly) raised with flat-topped (or with rarely rounded) apex, generally ± hairy, often ± enlarged in fruit. Monocarps per fruit few to many, red in vivo, stipitate or rarely (sub)sessile; seed-containing portion (sub)globose, ellipsoid(-cylindrical), cylindrical or rarely ± obovoid, apex sometimes (slightly) apiculate. Seed per monocarp 1, (sub)globose, ellipsoid, cylindrical or rarely ± obovoid; seed coat smooth; raphe flat or slightly raised; endosperm ruminations spiniform(-flattened peg).</p> <p>Etymology: ⎯Named in honour of Prof. Herbert Huber (1931-2005), who was the first to distinguish the three clades discussed in the present paper: Hubera (clade A), Monoon, and Polyalthia s.s. as informal groups of Polyalthia sensu lato based solely on morphology (Huber 1985).</p> <p>Distribution: ⎯Twenty-seven species are formally transferred here (see below); they are distributed from East Africa and Madagascar through southern and southeastern Asia to Malesia and the southwestern Pacific. It is anticipated that when the species of Hubera are thoroughly revised, an additional 10–20 species will be added.</p> <p>Given the morphological criteria mentioned above, the following species are transferred to the new genus:</p></div> 	https://treatment.plazi.org/id/039B87FDCC75C73B9D99DF1CFA71FC29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC76C73B9D99DA80FE19FB01.text	039B87FDCC76C73B9D99DA80FE19FB01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera amoena (A. C. Sm.) Chaowasku 2012	<div><p>1. Hubera amoena (A.C.Sm.) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia amoena Smith (1950: 159).</p> <p>Distribution: ⎯ Fiji.</p> <p>Notes: ⎯Study of the type specimen of this species (see Table 1) was the basis for this transfer; no DNA was available for this species.</p></div> 	https://treatment.plazi.org/id/039B87FDCC76C73B9D99DA80FE19FB01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC76C73B9D99DDA8FE19FA5E.text	039B87FDCC76C73B9D99DDA8FE19FA5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera capillata (A. C. Sm.) Chaowasku 2012	<div><p>2. Hubera capillata (A.C.Sm.) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia capillata Smith (1950: 158).</p> <p>Distribution: ⎯ Fiji.</p> <p>Notes: ⎯Study of the type specimen of this species (see Table 1) was the basis for this transfer; no DNA was available for this species.</p></div> 	https://treatment.plazi.org/id/039B87FDCC76C73B9D99DDA8FE19FA5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC76C73B9D99DF50FD24F8BB.text	039B87FDCC76C73B9D99DF50FD24F8BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera ceramensis (Boerl.) Chaowasku 2012	<div><p>3. Hubera ceramensis (Boerl.) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia ceramensis Boerlage (1899: 106).</p> <p>Distribution: ⎯Ceram.</p> <p>Notes: ⎯Study of a drawing and description of this species in Boerlage (1899) was the basis for this transfer; no DNA was available for this species.</p></div> 	https://treatment.plazi.org/id/039B87FDCC76C73B9D99DF50FD24F8BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC76C73B9D99DE6DFB8EF84F.text	039B87FDCC76C73B9D99DE6DFB8EF84F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera cerasoides (Roxburgh 1795) Chaowasku. 2012	<div><p>4. Hubera cerasoides (Roxb.) Chaowasku, comb. nov.</p> <p>Basionym: Uvaria cerasoides Roxburgh (1795: 30). Homotypic synonyms: Guatteria cerasoides (Roxb.) Dunal (1817: 127). Unona cerasoides (Roxb.) Baillon (1868: 272). Polyalthia cerasoides (Roxb.) Beddome (1869: t. 1).</p> <p>Distribution: ⎯Mainland Asia except southern Thailand and Peninsular Malaysia.</p></div> 	https://treatment.plazi.org/id/039B87FDCC76C73B9D99DE6DFB8EF84F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC77C73A9D99D9D4FE06FE86.text	039B87FDCC77C73A9D99D9D4FE06FE86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera decora (Diels) Chaowasku 2012	<div><p>5. Hubera decora (Diels) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia decora Diels (1925: 342).</p> <p>Distribution: ⎯ Madagascar.</p></div> 	https://treatment.plazi.org/id/039B87FDCC77C73A9D99D9D4FE06FE86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC77C73A9D99D831FB7EFDDE.text	039B87FDCC77C73A9D99D831FB7EFDDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera forbesii (F. Muell. ex Diels) Chaowasku 2012	<div><p>6. Hubera forbesii (F.Muell. ex Diels) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia forbesii Mueller ex Diels (1912b: 131).</p> <p>Distribution: ⎯New Guinea (to Kei Islands?).</p> <p>Notes: ⎯Study of herbarium specimens including the type (see Table 1) and pollen morphology of this species (Table 3) was the basis for this transfer; no DNA was available for this species.</p></div> 	https://treatment.plazi.org/id/039B87FDCC77C73A9D99D831FB7EFDDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC77C73A9D99DBD9FE19FC2F.text	039B87FDCC77C73A9D99DBD9FE19FC2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera gracilis (Burck) Chaowasku 2012	<div><p>7. Hubera gracilis (Burck) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia gracilis Burck (1911: 430)</p> <p>Distribution: ⎯New Guinea.</p> <p>Notes: ⎯Study of the type specimen of this species (see Table 1) was the basis for this transfer; no DNA was available for this species.</p></div> 	https://treatment.plazi.org/id/039B87FDCC77C73A9D99DBD9FE19FC2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC77C73A9D99DA8BFE06FBCE.text	039B87FDCC77C73A9D99DA8BFE06FBCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera henrici (Diels) Chaowasku 2012	<div><p>8. Hubera henrici (Diels) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia henrici Diels (1925: 342).</p> <p>Distribution: ⎯ Madagascar.</p></div> 	https://treatment.plazi.org/id/039B87FDCC77C73A9D99DA8BFE06FBCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC77C73A9D99DDE9FE19FA1F.text	039B87FDCC77C73A9D99DDE9FE19FA1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera hirta (Miq.) Chaowasku 2012	<div><p>9. Hubera hirta (Miq.) Chaowasku, comb. nov.</p> <p>Basionym: Monoon hirtum Miquel (1865: 16). Homotypic synonym: Polyalthia hirta (Miq.) Mueller (1877: 95).</p> <p>Distribution: ⎯New Guinea.</p> <p>Notes: ⎯Study of the type specimen of this species (see Table 1) was the basis for this transfer; no DNA was available for this species.</p></div> 	https://treatment.plazi.org/id/039B87FDCC77C73A9D99DDE9FE19FA1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC77C73A9D99DC9BFAA6F942.text	039B87FDCC77C73A9D99DC9BFAA6F942.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera humblotii (Drake ex Cavaco & Keraudren) Chaowasku 2012	<div><p>10. Hubera humblotii (Drake ex Cavaco &amp; Keraudren) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia humblotii Drake ex Cavaco &amp; Keraudren (1957: 77).</p> <p>Distribution: ⎯ Comoro Islands.</p> <p>Notes: ⎯This species was transferred to Hubera based on the fact that related species [e.g. H. henrici, H. perrieri (Cavaco &amp; Keraudren) Chaowasku] with similar morphology (presence of domatia on the lower leaf surface, Schatz &amp; Le Thomas 1990) belong to Hubera (Fig. 1); no DNA was available for this species.</p> </div>	https://treatment.plazi.org/id/039B87FDCC77C73A9D99DC9BFAA6F942	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC77C73A9D99DE6DFCB4F84F.text	039B87FDCC77C73A9D99DE6DFCB4F84F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera jenkinsii (Hook. f. & Thomson) Chaowasku 2012	<div><p>11. Hubera jenkinsii (Hook.f. &amp; Thomson) Chaowasku, comb. nov.</p> <p>Basionym: Guatteria jenkinsii Hooker &amp; Thomson (1855: 141). Homotypic synonym: Polyalthia jenkinsii (Hook.f. &amp; Thomson) Hooker &amp; Thomson (1872: 64).</p> <p>Distribution: ⎯Southeastern Asia (west of Wallace’s Line).</p></div> 	https://treatment.plazi.org/id/039B87FDCC77C73A9D99DE6DFCB4F84F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC68C7259D99D9D4FCC5FE03.text	039B87FDCC68C7259D99D9D4FCC5FE03.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera keraudreniae (Le Thomas & G. E. Schatz) Chaowasku 2012	<div><p>12. Hubera keraudreniae (Le Thomas &amp; G.E.Schatz) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia keraudreniae Le Thomas &amp; Schatz in Schatz &amp; Le Thomas (1990: 124).</p> <p>Distribution: ⎯ Madagascar.</p> <p>Notes: ⎯This species was transferred to Hubera based on the fact that a related species (H. decora) with similar morphology (absence of domatia on the lower leaf surface, Schatz &amp; Le Thomas 1990), belongs to Hubera (Fig. 1); no DNA was available for this species.</p> </div>	https://treatment.plazi.org/id/039B87FDCC68C7259D99D9D4FCC5FE03	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC68C7259D99D8AEFCAAFD22.text	039B87FDCC68C7259D99D8AEFCAAFD22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera korinti (Chaowasku 1864) Chaowasku 2012	<div><p>13. Hubera korinti (Dunal) Chaowasku, comb. nov.</p> <p>Basionym: Guatteria korinti Dunal (1817: 133). Homotypic synonym: Polyalthia korinti (Dunal) Thwaites (1864: 398).</p> <p>Distribution: ⎯Southern/southwestern India to Sri Lanka.</p></div> 	https://treatment.plazi.org/id/039B87FDCC68C7259D99D8AEFCAAFD22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC68C7259D99DB8DFE19FC73.text	039B87FDCC68C7259D99DB8DFE19FC73.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera leptopoda (Diels) Chaowasku 2012	<div><p>14. Hubera leptopoda (Diels) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia leptopoda Diels (1915: 179).</p> <p>Distribution: ⎯New Guinea.</p> <p>Notes: ⎯Study of the type specimen of this species (see Table 1) was the basis for this transfer; no DNA was available for this species.</p></div> 	https://treatment.plazi.org/id/039B87FDCC68C7259D99DB8DFE19FC73	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC68C7259D99DABEFD17FB41.text	039B87FDCC68C7259D99DABEFD17FB41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera loriformis (Gillespie) Chaowasku 2012	<div><p>15. Hubera loriformis (Gillespie) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia loriformis Gillespie (1931: 4).</p> <p>Distribution: ⎯ Fiji.</p> <p>Notes: ⎯Study of herbarium specimens including the type of this species (see Table 1) was the basis for this transfer; no DNA was available for this species.</p></div> 	https://treatment.plazi.org/id/039B87FDCC68C7259D99DABEFD17FB41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC68C7259D99DC68FDD6F99E.text	039B87FDCC68C7259D99DC68FDD6F99E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera mossambicensis (Vollesen) Chaowasku 2012	<div><p>16. Hubera mossambicensis (Vollesen) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia mossambicensis Vollesen (1980b: 403).</p> <p>Distribution: ⎯East Africa.</p> <p>Notes: ⎯Study of herbarium specimen of this species (see Table 1) was the basis for this transfer; no DNA was available for this species.</p></div> 	https://treatment.plazi.org/id/039B87FDCC68C7259D99DC68FDD6F99E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC68C7259D99DF19FCC5F8C5.text	039B87FDCC68C7259D99DF19FCC5F8C5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera multistamina (G. E. Schatz & Le Thomas) Chaowasku 2012	<div><p>17. Hubera multistamina (G.E.Schatz &amp; Le Thomas) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia multistamina Schatz &amp; Le Thomas (1990: 126).</p> <p>Distribution: ⎯ Madagascar.</p> <p>Notes: ⎯This species was transferred to Hubera based on the fact that a related species (H. decora) with similar morphology (absence of domatia on the lower leaf surface, Schatz &amp; Le Thomas 1990), belongs to Hubera (Fig. 1); no DNA was available for this species.</p> </div>	https://treatment.plazi.org/id/039B87FDCC68C7259D99DF19FCC5F8C5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC69C7249D99D9D4FB98FEDF.text	039B87FDCC69C7249D99D9D4FB98FEDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera nitidissima (Chaowasku 1863) Chaowasku 2012	<div><p>18. Hubera nitidissima (Dunal) Chaowasku, comb. nov.</p> <p>Basionym: Unona nitidissima Dunal (1817: 109). Homotypic synonym: Polyalthia nitidissima (Dunal) Bentham (1863: 51).</p> <p>Distribution: ⎯New Guinea and northern/northeastern Australia to New Caledonia.</p></div> 	https://treatment.plazi.org/id/039B87FDCC69C7249D99D9D4FB98FEDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC69C7249D99D8DBFE06FDFE.text	039B87FDCC69C7249D99D8DBFE06FDFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera pendula (Capuron ex G. E. Schatz & Le Thomas) Chaowasku 2012	<div><p>19. Hubera pendula (Capuron ex G.E.Schatz &amp; Le Thomas) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia pendula Capuron ex Schatz &amp; Le Thomas (1990: 128).</p> <p>Distribution: ⎯ Madagascar.</p></div> 	https://treatment.plazi.org/id/039B87FDCC69C7249D99D8DBFE06FDFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC69C7249D99DB39FE06FC91.text	039B87FDCC69C7249D99DB39FE06FC91.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera perrieri (Cavaco & Keraudren) Chaowasku 2012	<div><p>20. Hubera perrieri (Cavaco &amp; Keraudren) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia perrieri Cavaco &amp; Keraudren (1957: 75).</p> <p>Distribution: ⎯ Madagascar.</p></div> 	https://treatment.plazi.org/id/039B87FDCC69C7249D99DB39FE06FC91	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC69C7249D99DA18FE69FB27.text	039B87FDCC69C7249D99DA18FE69FB27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera rumphii (Blume ex Hensch.) Chaowasku 2012	<div><p>21. Hubera rumphii (Blume ex Hensch.) Chaowasku, comb. nov.</p> <p>Basionym: Guatteria rumphii Blume ex Henschel (1833: 153). Homotypic synonym: Polyalthia rumphii (Blume ex Hensch.) Merrill (1923: 162).</p> <p>Distribution: ⎯Southeastern Asia (west of Wallace’s Line) to Ambon (and New Guinea?).</p> <p>Notes: ⎯This and the other two morphologically similar species, H. ceramensis and H. jenkinsii, constitute a species complex (Turner 2011). To resolve this complex, detailed revisionary and phylogenetic studies are required.</p> </div>	https://treatment.plazi.org/id/039B87FDCC69C7249D99DA18FE69FB27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC69C7249D98DD93FC70FA4A.text	039B87FDCC69C7249D98DD93FC70FA4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera sambiranensis (Capuron ex Le Thomas & G. E. Schatz) Chaowasku 2012	<div><p>22. Hubera sambiranensis (Capuron ex Le Thomas &amp; G.E.Schatz) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia sambiranensis Capuron ex Le Thomas &amp; Schatz in Schatz &amp; Le Thomas (1990: 116).</p> <p>Distribution: ⎯ Madagascar.</p> <p>Notes: ⎯This species was transferred to Hubera based on the fact that related species (e.g. H. henrici, H. perrieri) with similar morphology (presence of domatia on the lower leaf surface, Schatz &amp; Le Thomas 1990), belong to Hubera (Fig. 1); no DNA was available for this species.</p> </div>	https://treatment.plazi.org/id/039B87FDCC69C7249D98DD93FC70FA4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC69C7249D99DF65FE0EF94D.text	039B87FDCC69C7249D99DF65FE0EF94D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera stuhlmannii (Engl.) Chaowasku 2012	<div><p>23. Hubera stuhlmannii (Engl.) Chaowasku, comb. nov.</p> <p>Basionym: Unona stuhlmannii Engler (1895: 179). Homotypic synonym: Polyalthia stuhlmannii (Engl.) Verdcourt (1969: 94).</p> <p>Distribution: ⎯East Africa.</p></div> 	https://treatment.plazi.org/id/039B87FDCC69C7249D99DF65FE0EF94D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC69C7249D99DE64FE0EF86F.text	039B87FDCC69C7249D99DE64FE0EF86F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera tanganyikensis (Vollesen) Chaowasku 2012	<div><p>24. Hubera tanganyikensis (Vollesen) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia tanganyikensis Vollesen (1980a: 56).</p> <p>Distribution: ⎯East Africa.</p></div> 	https://treatment.plazi.org/id/039B87FDCC69C7249D99DE64FE0EF86F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC6AC7279D99D9D4FE19FE35.text	039B87FDCC6AC7279D99D9D4FE19FE35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera trichoneura (Diels) Chaowasku 2012	<div><p>25. Hubera trichoneura (Diels) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia trichoneura Diels (1912a: 871).</p> <p>Distribution: ⎯New Guinea.</p> <p>Notes: ⎯Study of the type specimen of this species (see Table 1) was the basis for this transfer; no DNA was available for this species.</p></div> 	https://treatment.plazi.org/id/039B87FDCC6AC7279D99D9D4FE19FE35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC6AC7279D99D8FCFD2AFD5B.text	039B87FDCC6AC7279D99D8FCFD2AFD5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera verdcourtii (Vollesen) Chaowasku 2012	<div><p>26. Hubera verdcourtii (Vollesen) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia verdcourtii Vollesen (1980a: 56).</p> <p>Distribution: ⎯East Africa.</p> <p>Notes: ⎯This species was transferred to Hubera based on the fact that a related species (H. stuhlmannii) with similar morphology (presence of domatia on the lower leaf surface, Vollesen 1980a), belongs to Hubera (Fig. 1); no DNA was available for this species.</p> </div>	https://treatment.plazi.org/id/039B87FDCC6AC7279D99D8FCFD2AFD5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
039B87FDCC6AC7219D99DA56FE17FDB7.text	039B87FDCC6AC7219D99DA56FE17FDB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hubera vitiensis (Chaowasku 1865) Chaowasku 2012	<div><p>27. Hubera vitiensis (Seem.) Chaowasku, comb. nov.</p> <p>Basionym: Polyalthia vitiensis Seemann (1865: 4).</p> <p>Distribution: ⎯ Fiji.</p> <p>Notes: ⎯Study of herbarium specimens of this species (see Table 1) was the basis for this transfer; no DNA was available for this species.</p> <p>Several species (e.g. H. henrici, H. korinti, H. nitidissima, H. perrieri, H. stuhlmannii, H. vitiensis) exhibit domatia on the lower leaf surface at the axils where the secondary veins meet the primary veins (Figs. 5A–D; see comparisons of lower leaf surface without domatia in Figs. 5E–H). They are characterized by a tuft of aggregated hairs visible to the naked eye. In Annonaceae, this kind of domatium has been reported to occur in Mitrephora Hooker &amp; Thomson (1855: 112) (Weerasooriya &amp; Saunders 2010), which is also a member of the Miliuseae, and Annona Linnaeus (1753: 536) [including Rollinia Saint-Hilaire (1824: 28)] (Van den Bos et al. 1989), a member of the subfamily Annonoideae (Chatrou et al. 2012).</p> <p>The genus Miliusa was recovered as sister to Hubera. So far, no morphological synapomorphy linking these genera has been observed. They only share some characters considered as symplesiomorphies, such as reticulate tertiary leaf venation and pollen with verrucate to rugulate ornamentation (cerebroid sensu Mols et al. 2004b) and germination zone(s) characterized by enlargements/reductions of the intine sublayers (Chaowasku et al. 2008).</p> <p>Among genera of Malmeoideae, Hubera exhibits the widest distribution, ranging from East Africa and Madagascar across southern and southeastern Asia through Malesia and the southwestern Pacific. It is the only genus of Miliuseae that occurs in Madagascar and East Africa. Phylogenetic analysis of Hubera (Fig. 1) shows some clear biogeographic patterns. The Afro-Madagascan species are grouped together in a strongly supported clade (clade A2), as do the species occurring in the Austro-Papuasian area, which are clustered, with strong support, in clade A1. The biogeographic scenario explaining this distribution will be the focus of another study.</p> <p>Schatz &amp; Le Thomas (1990) revised Polyalthia species occurring in Madagascar and distinguished five informal groups (groups A–E) based on macromorphological and pollen characters. Species of groups B and C possess monosulcate pollen and were found to form a strongly supported clade (now transferred to Fenerivia) recovered outside Miliuseae (Saunders et al. 2011), which is congruent with the phylogenetic results. Genera outside Miliuseae exhibit monosulcate pollen, whereas genera belonging to Miliuseae possess cryptoaperturate/disulculate pollen (Chaowasku et al. 2012). Species of groups A, D, and E have cryptoaperturate pollen, and their membership in Hubera, which is a member of the Miliuseae, was thus not unexpected.</p> <p>It is generally difficult to distinguish Hubera from Fenerivia using only macromorphology. Both genera share some similar morphological features, e.g. axillary inflorescences, uniovulate carpels, and spiniform(- flattened peg) endosperm ruminations (Schatz &amp; Le Thomas 1990). However, Fenerivia possesses a more pronounced seed raphe that is rib-like (Saunders et al. 2011), whereas that of Hubera is flat to slightly raised (Fig. 4a). In addition, Fenerivia exhibits a pronounced (± thickened) receptacle rim (vestigial calyx flange sensu Saunders et al. 2011). This feature is considered one of the diagnostic characters of Fenerivia. It is absent (or rarely slightly observed) in Hubera. Nevertheless, the presence of domatia on the lower leaf surface should be a primary character in distinguishing Madagascan Hubera from Fenerivia because the latter does not possess this character, whereas Hubera species formerly known as Polyalthia group A of Schatz &amp; Le Thomas (1990) do. Domatia on the lower leaf surface can also be used to quickly distinguish certain Afro- Asian species of Hubera from Polyalthia s.s. and other genera formerly known as Polyalthia. Another consequence of this study is elimination of Polyalthia s.s. from the floras of Africa and Madagascar; thus, it is strictly a genus of Australasia.</p> </div>	https://treatment.plazi.org/id/039B87FDCC6AC7219D99DA56FE17FDB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Chaowasku, T;Johnson, DM;van der Ham, RWJM;Chatrou, LW	Chaowasku, T, Johnson, DM, van der Ham, RWJM, Chatrou, LW (2012): Characterization of Hubera (Annonaceae), a new genus segregated from Polyalthia and allied to Miliusa. Phytotaxa 69: 33-56, DOI: 10.11646/phytotaxa.69.1.6
