taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
039B9101BC572A47165CFB6834F5F90F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990443/files/figure.png	https://doi.org/10.5281/zenodo.3990443	Figure 1. Gromphas jardim sp. nov. (A–D) Holotype. (A) Dorsal view. (B) Lateral view. (C) Aedeagus. From left to right, dorsal, lateral and ventral views. (D) Labels. (E) Female paratype from Bolivia. (F) Dorsal view of head (holotype). (G) Frontal view of cephalic projection (male paratype).	Figure 1. Gromphas jardim sp. nov. (A–D) Holotype. (A) Dorsal view. (B) Lateral view. (C) Aedeagus. From left to right, dorsal, lateral and ventral views. (D) Labels. (E) Female paratype from Bolivia. (F) Dorsal view of head (holotype). (G) Frontal view of cephalic projection (male paratype).	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC572A47165CFB6834F5F90F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990445/files/figure.png	https://doi.org/10.5281/zenodo.3990445	Figure 2. Protibia of Gromphas jardim sp. nov. (A, B) Male protibia: (A) Ventral view, (B) Dorsal view. Red arrow indicates apical tubercle modified in a tapered spur. (C) Female protibia. Note that the apical tuft of setae is much denser and longer in male than in female.	Figure 2. Protibia of Gromphas jardim sp. nov. (A, B) Male protibia: (A) Ventral view, (B) Dorsal view. Red arrow indicates apical tubercle modified in a tapered spur. (C) Female protibia. Note that the apical tuft of setae is much denser and longer in male than in female.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC572A47165CFB6834F5F90F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990447/files/figure.png	https://doi.org/10.5281/zenodo.3990447	Figure 3. Lateral view of pronotum. (A) Gromphas jardim sp. nov. (B) Gromphas amazonica. Note that the granulation extends much more posteriorly in G. jardim, reaching the pronotal posterior margin, than in G. amazonica, wherein the granulation is absent or rudimentary in posterolateral region after lateral fossa.	Figure 3. Lateral view of pronotum. (A) Gromphas jardim sp. nov. (B) Gromphas amazonica. Note that the granulation extends much more posteriorly in G. jardim, reaching the pronotal posterior margin, than in G. amazonica, wherein the granulation is absent or rudimentary in posterolateral region after lateral fossa.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC572A47165CFB6834F5F90F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990449/files/figure.png	https://doi.org/10.5281/zenodo.3990449	Figure 4. Male paratype of Gromphas jardim sp. nov. and its labels (braces ‘{’ joining two sides of a same label). We believe that the French zoologist and explorer Alcide d’Orbigny was the collector of this specimen in 1832 (see the text).	Figure 4. Male paratype of Gromphas jardim sp. nov. and its labels (braces ‘{’ joining two sides of a same label). We believe that the French zoologist and explorer Alcide d’Orbigny was the collector of this specimen in 1832 (see the text).	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC572A47165CFB6834F5F90F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990451/files/figure.png	https://doi.org/10.5281/zenodo.3990451	Figure 5. Updated distribution of the six species of Gromphas.	Figure 5. Updated distribution of the six species of Gromphas.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC572A47165CFB6834F5F90F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990455/files/figure.png	https://doi.org/10.5281/zenodo.3990455	Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).	Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC572A47165CFB6834F5F90F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990459/files/figure.png	https://doi.org/10.5281/zenodo.3990459	Figure 7. Metepisterna. (A) Gromphas jardim sp. nov.: gromphas is unique among the Phanaeini by not showing any trace of a metepisternal tab (MTab) covering the epipleural margin, a synapomorphy of this genus. (B) Oruscatus davus: Both species of Oruscatus have a small but evident MTab (indicated by red arrow). (C) Bolbites onitoides: The MTab of Bolbites is strongly developed and the dorsal margin of metepisternum is highly curved. This is the same form seen in the rest of Phanaeina and is a synapomorphy of this taxon. (D) Dichotomius fissus (Harold, 1867): MTab in a non-Phanaeini. This structure is present also in a series of other Coprini, as well as in at least some Oniticellini and Onitini.	Figure 7. Metepisterna. (A) Gromphas jardim sp. nov.: gromphas is unique among the Phanaeini by not showing any trace of a metepisternal tab (MTab) covering the epipleural margin, a synapomorphy of this genus. (B) Oruscatus davus: Both species of Oruscatus have a small but evident MTab (indicated by red arrow). (C) Bolbites onitoides: The MTab of Bolbites is strongly developed and the dorsal margin of metepisternum is highly curved. This is the same form seen in the rest of Phanaeina and is a synapomorphy of this taxon. (D) Dichotomius fissus (Harold, 1867): MTab in a non-Phanaeini. This structure is present also in a series of other Coprini, as well as in at least some Oniticellini and Onitini.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC5F2A4616B3FF6A348BFB36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990451/files/figure.png	https://doi.org/10.5281/zenodo.3990451	Figure 5. Updated distribution of the six species of Gromphas.	Figure 5. Updated distribution of the six species of Gromphas.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC5F2A4616B3FF6A348BFB36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990455/files/figure.png	https://doi.org/10.5281/zenodo.3990455	Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).	Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC5F2A4616A3FB363796FA3B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990451/files/figure.png	https://doi.org/10.5281/zenodo.3990451	Figure 5. Updated distribution of the six species of Gromphas.	Figure 5. Updated distribution of the six species of Gromphas.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC5F2A4616A3FB363796FA3B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990455/files/figure.png	https://doi.org/10.5281/zenodo.3990455	Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).	Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC5F2A4116B5FA323419FC77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990451/files/figure.png	https://doi.org/10.5281/zenodo.3990451	Figure 5. Updated distribution of the six species of Gromphas.	Figure 5. Updated distribution of the six species of Gromphas.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC5F2A4116B5FA323419FC77.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990455/files/figure.png	https://doi.org/10.5281/zenodo.3990455	Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).	Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC582A5A16D9FBF735D0FBC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990451/files/figure.png	https://doi.org/10.5281/zenodo.3990451	Figure 5. Updated distribution of the six species of Gromphas.	Figure 5. Updated distribution of the six species of Gromphas.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC582A5A16D9FBF735D0FBC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990455/files/figure.png	https://doi.org/10.5281/zenodo.3990455	Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).	Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC582A5A16D9FBF735D0FBC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990461/files/figure.png	https://doi.org/10.5281/zenodo.3990461	Figure 8. Lateral view of male protibia, ornamentation of ventral carina. (A) Gromphas inermis: a row of tubercles (indicated by red arrow), a synapomorphy of clade B of Gromphas. (B) Oruscatus davus: a single large tooth. (C) Bolbites onitoides: a large central tooth with minor lateral teeth.	Figure 8. Lateral view of male protibia, ornamentation of ventral carina. (A) Gromphas inermis: a row of tubercles (indicated by red arrow), a synapomorphy of clade B of Gromphas. (B) Oruscatus davus: a single large tooth. (C) Bolbites onitoides: a large central tooth with minor lateral teeth.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC582A5A16D9FBF735D0FBC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990443/files/figure.png	https://doi.org/10.5281/zenodo.3990443	Figure 1. Gromphas jardim sp. nov. (A–D) Holotype. (A) Dorsal view. (B) Lateral view. (C) Aedeagus. From left to right, dorsal, lateral and ventral views. (D) Labels. (E) Female paratype from Bolivia. (F) Dorsal view of head (holotype). (G) Frontal view of cephalic projection (male paratype).	Figure 1. Gromphas jardim sp. nov. (A–D) Holotype. (A) Dorsal view. (B) Lateral view. (C) Aedeagus. From left to right, dorsal, lateral and ventral views. (D) Labels. (E) Female paratype from Bolivia. (F) Dorsal view of head (holotype). (G) Frontal view of cephalic projection (male paratype).	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC582A5A16D9FBF735D0FBC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990447/files/figure.png	https://doi.org/10.5281/zenodo.3990447	Figure 3. Lateral view of pronotum. (A) Gromphas jardim sp. nov. (B) Gromphas amazonica. Note that the granulation extends much more posteriorly in G. jardim, reaching the pronotal posterior margin, than in G. amazonica, wherein the granulation is absent or rudimentary in posterolateral region after lateral fossa.	Figure 3. Lateral view of pronotum. (A) Gromphas jardim sp. nov. (B) Gromphas amazonica. Note that the granulation extends much more posteriorly in G. jardim, reaching the pronotal posterior margin, than in G. amazonica, wherein the granulation is absent or rudimentary in posterolateral region after lateral fossa.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC582A5A16D9FBF735D0FBC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990449/files/figure.png	https://doi.org/10.5281/zenodo.3990449	Figure 4. Male paratype of Gromphas jardim sp. nov. and its labels (braces ‘{’ joining two sides of a same label). We believe that the French zoologist and explorer Alcide d’Orbigny was the collector of this specimen in 1832 (see the text).	Figure 4. Male paratype of Gromphas jardim sp. nov. and its labels (braces ‘{’ joining two sides of a same label). We believe that the French zoologist and explorer Alcide d’Orbigny was the collector of this specimen in 1832 (see the text).	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC582A5A16D9FBF735D0FBC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990459/files/figure.png	https://doi.org/10.5281/zenodo.3990459	Figure 7. Metepisterna. (A) Gromphas jardim sp. nov.: gromphas is unique among the Phanaeini by not showing any trace of a metepisternal tab (MTab) covering the epipleural margin, a synapomorphy of this genus. (B) Oruscatus davus: Both species of Oruscatus have a small but evident MTab (indicated by red arrow). (C) Bolbites onitoides: The MTab of Bolbites is strongly developed and the dorsal margin of metepisternum is highly curved. This is the same form seen in the rest of Phanaeina and is a synapomorphy of this taxon. (D) Dichotomius fissus (Harold, 1867): MTab in a non-Phanaeini. This structure is present also in a series of other Coprini, as well as in at least some Oniticellini and Onitini.	Figure 7. Metepisterna. (A) Gromphas jardim sp. nov.: gromphas is unique among the Phanaeini by not showing any trace of a metepisternal tab (MTab) covering the epipleural margin, a synapomorphy of this genus. (B) Oruscatus davus: Both species of Oruscatus have a small but evident MTab (indicated by red arrow). (C) Bolbites onitoides: The MTab of Bolbites is strongly developed and the dorsal margin of metepisternum is highly curved. This is the same form seen in the rest of Phanaeina and is a synapomorphy of this taxon. (D) Dichotomius fissus (Harold, 1867): MTab in a non-Phanaeini. This structure is present also in a series of other Coprini, as well as in at least some Oniticellini and Onitini.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
039B9101BC582A5A16D9FBF735D0FBC3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3990445/files/figure.png	https://doi.org/10.5281/zenodo.3990445	Figure 2. Protibia of Gromphas jardim sp. nov. (A, B) Male protibia: (A) Ventral view, (B) Dorsal view. Red arrow indicates apical tubercle modified in a tapered spur. (C) Female protibia. Note that the apical tuft of setae is much denser and longer in male than in female.	Figure 2. Protibia of Gromphas jardim sp. nov. (A, B) Male protibia: (A) Ventral view, (B) Dorsal view. Red arrow indicates apical tubercle modified in a tapered spur. (C) Female protibia. Note that the apical tuft of setae is much denser and longer in male than in female.	2015-09-30	Cupello, Mario;Vaz-de-Mello, Fernando Z.		Zenodo	biologists	Cupello, Mario;Vaz-de-Mello, Fernando Z.			
