taxonID	type	description	language	source
039BC700D760076DFF66FAF8FCC0DF08.taxon	distribution	Distribution and habitat: — D. virgata subsp. tenuirostris is endemic to Morocco. It is mostly distributed across the western and central plateau, extending approximately from Fès and Taza (Region Fès-Mèknes) in the north to Béni Mellal and Benguerir (Region Marrakech-Safi) in the south. Additional specimens examined: — MOROCCO. Region Fès-Mèknes: Jbel Jalaph, above Fez, 300 – 400 m, 12 April 1971, Davis 51194 (BM, E); Camping de Fes, 350 m, 3 June 1971, Boquet & al. 10337 (BM); Au N de Taza, 19 April 1924, Humbert s. n. (MPU). Region Rabat-Salé-Kénitra: Ez-Zhiliga, 27 April 1987, Martínez-Laborde 344 (MA 01 - 00916854); Hadra, Maïat, 19 February 1951, Nègre 966 (RAB 18006); Souk Tleta du Rharb, in wheat field, 1 March 1982, Smith 28 (RNG). Region Béni Mellal-Khénifra: Beni Amir, 1 April 1949, Nègre s. n. (MPU); ibidem, 1 April 1949, Sauvage 7141 (RAB 17993); ibidem, Domaine de la Sacima, 9 March 1952, Nègre 1947 (RAB 18009); Bled el Harcha, 2 km NNE de l’Azib, 23 March 1953, Nègre 3245 (MPU); Kasba Tadla, 17 March 1927, leg.? (RAB 17995); 9 km al N de Kasba Tadla, 3 May 1986, Martínez-Laborde 324 (MA 01 - 00916853); ± 10 km de Oued Zem hacia Fquih Ben Salah, 27 April 1987, Martínez-Laborde 343 (MA 01 - 00916883). Region Marrakech-Safi: Koudiat es Sloughi, â 5 km au S du signal 547 et 15 km à l’est de Benguerir, 27 November 1951, Nègre 767 (G, K, RAB 18005); ibidem, 27 February 1951, Nègre 1091 (RAB 18014); E de Benguerir hacia El Kelaa des Sraghna, 25 April 1987, Martínez-Laborde 341 (MA 01 - 00916884); 54 km N de Marrakech, 25 April 1987, Martínez-Laborde 342 (MA 01 - 00916880); Piste de Sidi Bou-Heddaj à Sidi Abd er Rahmane, 23 March 1953, Nègre 3231 (RAB 18004). Specimens obtained from cultivated material. SPAIN. Madrid: Ciudad Universitaria, spring 1985, Martínez-Laborde 356, culta ex semina BGV-UPM 3644 (MA 01 - 00916881); ibidem, spring 1985, Martínez-Laborde 357, culta ex semina BGV-UPM 3644 (MA 01 - 00916882). Cultivated material: — BGV-UPM 3644, BGV-UPM 7479, BGV-UPM 7480, BGV-UPM 7509, BGV-UPM 7525, BGV-UPM 7526, BGV-UPM 7530, BGV-UPM 7531, BGV-UPM 7532, BGV-UPM 7533, BGV-UPM 7535. Observations: — This taxon includes populations growing principally in central Morocco and resembling Iberian D. virgata mainly in being ± hirsute, with hairs mostly patent on stems, that tend to be somewhat retrorse in Moroccan plants, while those on leaves are also patent in Iberian plants, but to some extent reclining towards the leaf apex in Moroccan material. Besides, they have generally longer siliques and V-shaped (instead of capitate) stigmata. Note 1: — This plant was at first erroneously named D. catholica var. rivulorum or D. virgata subsp. rivulorum (Gómez-Campo 1978, 1990, Martínez-Laborde 1988), but examination of the type material of D. rivulorum Braun-Blanquet & Maire (1925: 23) [MOROCCO. Region Béni Mellal-Khénifra: Plaine de Mrirt, 15 February 1915, Dr. Nain s. n. (MPU!); Region Fès-Mèknes: Azrou, Plaine du Tigrira, 27 March 1923, Maire s. n. (MPU!); Plaine Azrou, April 1923, G. et J. Braun-Blanquet s. n. (MPU!)] has shown that it is a different plant, much closer to D. catholica (if not a synonym). Note 2: — Nègre (1960) appropriately transferred Maire’s var. tenuirostris to D. virgata as f. tenuirostris, but the material he cited also included specimens belonging to other taxa (see below) and even to the genus Erucastrum [MOROCCO. Region Fès-Mèknes: Aïne Cheggag, February 1913, Mouret 1803 and Mouret 1974 (P!); Sidi-Abdallah, entre Taza et Fès, 24 March 1921, Maire s. n. (MPU!)]. On the other hand, some specimens cited by him under D. virgata f. platystylos belong, however, to D. virgata subsp. tenuirostris.	en	Martínez-Laborde, Juan B. (2018): Taxa related to Diplotaxis virgata (Brassicaceae) in northwest Africa. Phytotaxa 371 (3): 205-216, DOI: 10.11646/phytotaxa.371.3.4, URL: http://dx.doi.org/10.11646/phytotaxa.371.3.4
039BC700D767076CFF66F984FC1ADF9C.taxon	distribution	Distribution and habitat: — Diplotaxis sahariensis is endemic to southern and southeastern Morocco south of the High Atlas (Regions Drâa-Tafilalet and Oriental) and northwestern Algeria (Provinces Béchar and Naâma). It is known to grow at elevations between 800 and 900 m above sea level. Additional specimens examined: — ALGERIA. Province Béchar: Béni Ounif, 1906, Herb. Battandier s. n. (MPU). Province Naâma: “ Duveyrier ” [currently Zoubia], 10 April 1938, Bertram s. n. (G, MPU); de Mograr [Moghrar] à Djenien bou Resg [Djeniene Bourezg], 16 April 1888, Bonnet & Maury s. n. (MPU); plaine argilo-sablonneuse entre Tyout [Tiout] et Asla, 9 May 1856, Cosson s. n. (MPU, P); ibidem, 11 May 1856, Cosson s. n. (MPU); ibidem, Kralik 11590 (BM); ibidem, 4 April 1875, Warion s. n. (P). MOROCCO. Region Drâa-Tafilalet: Erfoud-Alnif, stony desert, 800 – 900 m, 20 February 1968, Holliday 1 a (BM); Erfoud, 27 April 1939, P. & J. Davis 580 (E); Erfoud, at Bordj-Est, 5 April 1969, Davis 49131 (BM, E); Erfoud, 1 May 1927, Maire s. n. (MPU); Ouarzazate, 6 June 1936, Balls B 2640 (K); Zagora, 4 May 1986, Martínez-Laborde 327 (MA 01 - 00916875); Agdz, 850 m, 7 March 1992, Flügel M 38 & D- 34593 (B). Region Oriental: Djahifat, April 1913, Pitard 3065 (P); Djebel Grouz, Pitard 3064 (P); Djebel Melias, Pitard 3069 (P); Djebel Ouazzani, April 1913, Pitard 3063 (BM, K, P); Figuig, April 1924, Humbert s. n. (P). Region unknown: Aïne Yalau, Pitard 3066 (P); Djebel el Hainer, April 1913, Pitard 3068 (K, P); Sidi Youcef, March 1913, Pitard 3067 (K, P). Specimens obtained from cultivated material. SPAIN. Madrid: Ciudad Universitaria, spring 1985, Martínez-Laborde 202, culta ex semina BGV-UPM 5550 (MA 488554); ibidem, spring 1985, Martínez-Laborde 203, culta ex semina BGV-UPM 5545 (MA 488555); ibidem, spring 1985, Martínez-Laborde 204, culta ex semina BGV-UPM 5552 (MA 488556). Cultivated material: — BGV-UPM 5550, BGV-UPM 5552, BGV-UPM 5545, BGV-UPM 7515. Observations: — Two groups of populations close to D. virgata growing farther south and east had been preliminarily distinguished by Martínez-Laborde & Gómez-Campo (1993). In southeastern Morocco and adjacent areas of northwestern Algeria south of Oran one finds plants with rather long, patent but soft hairs and siliques with relatively short stylar segments, which could be identified as D. virgata f. sahariensis Cosson (1887: 165) and are here proposed to constitute a separate species. Note 1: — As the name of a new taxon, D. virgata var. cavanillesiana f. platystylos Nègre should be considered invalid, since the short descriptive phrase in the key is written in French. However, the phrase “ D. virgata f. sahariensis Coss., comp., II, p. 165 ” written just below Nègre’s name constitutes a full reference to a previously published diagnosis with its own valid name, of which f. platystylos should be considered a superfluous synonym, despite the fact that the abundant material cited by Nègre includes specimens belonging not only to D. sahariensis, but also to D. virgata subsp. tenuirostris, and to the new species described further below. Note 2: — At least eight out of 13 specimens cited under D. virgata var. cavanillesiana f. hispida Nègre (1960: 64) belong to D. sahariensis. However, in the diagnostic French phrase the hairs are described as “ raides ” (rigid) whereas they are in fact soft. Other specimens belonging to D. sahariensis are cited by Nègre (1960) under his f. platystylos. Note 3: — Diplotaxis sahariensis belongs to sect. Rhynchocarpum.	en	Martínez-Laborde, Juan B. (2018): Taxa related to Diplotaxis virgata (Brassicaceae) in northwest Africa. Phytotaxa 371 (3): 205-216, DOI: 10.11646/phytotaxa.371.3.4, URL: http://dx.doi.org/10.11646/phytotaxa.371.3.4
039BC700D7660761FF66F928FA29DB25.taxon	materials_examined	Type: — MOROCCO. Region Drâa-Tafilalet: Ouarzazate: prope oppidulum Amerzgane, ad 1300 m, in clivosis siccis, 31 May 1985, Blanché, Fernández Casas, Molero, Montserrat & Romo 9644 (holotype BC 813229, rightmost specimen; isotype SEV!)	en	Martínez-Laborde, Juan B. (2018): Taxa related to Diplotaxis virgata (Brassicaceae) in northwest Africa. Phytotaxa 371 (3): 205-216, DOI: 10.11646/phytotaxa.371.3.4, URL: http://dx.doi.org/10.11646/phytotaxa.371.3.4
039BC700D7660761FF66F928FA29DB25.taxon	diagnosis	Diagnosis: — It differs from Diplotaxis virgata and Diplotaxis sahariensis mainly by its hairs, predominantly retrorse on stems, generally antrorse on leaves, and more or less flexuose, pointing in several directions and apparently disorderly arranged on sepals; as well as by its anthers, with a purple spot at tip (to be observed in vivo before anthesis). Annuals, 10 – 50 cm tall. Stems erect, foliose, ± hirsute at least at base, with predominantly retrorse hairs. Leaves ± hairy, hairs 0.3 – 0.5 mm long, generally antrorse, especially towards the apex, rarely retrorse towards the base. Lower leaves with petiole 5 – 50 mm long; blade 20 – 130 × 10 – 70 mm, elliptic to scarcely obovate in outline, margin usually remotely dentate, generally lobate to pinnatisect, rarely subentire; lateral segments 1 – 3 pairs, ± elliptical, apical segment generally much larger, sometimes lobate towards the base, attenuate. Cauline leaves smaller, uppermost leaves narrowly ovate, with 1 – 2 lobes at base. Sepals 3.5 – 4.5 mm long, densely to sparsely hairy, hairs mostly more or less flexuose, pointing in several directions and apparently disorderly arranged. Petals 6.5 – 9.5 × 3.0 – 4.5 mm, claw distinct, limb yellow, obovate to slightly oblong. Median stamens 4.5 – 5.0 mm long, lateral stamens ± 3.5 mm long, anthers 1.4 – 1.6 mm long, with a purple spot at tip (to be observed in vivo, before anthesis). Ovary with ± 30 – 50 ovules per locule. Fruiting pedicels 4 – 16 mm, erect-patent to patent, hairless or provided with generally ± retrorse hairs. Siliques (13 –) 20 – 30 (– 35) × 1.0 – 1.7 mm, erect-patent to patent, rarely suberect, elliptical to oblong in outline; carpophore 0.3 – 0.6 (– 1.1) mm; valves flattened; stylar segment (1 –) 1.5 – 5.0 (– 6.5) × 0.8 – 1.2 mm (equivalent to approximately 1 / 10 – 1 / 5 of the total length of the silique), linear to attenuate, flattened, 1 - to 3 - veined, containing 1 – 3 seeds (or aborted ovules), stigma ± capitate. Seeds in two rows per locule, 0.6 – 0.9 × 0.4 – 0.6 mm, ovoid to ellipsoid.	en	Martínez-Laborde, Juan B. (2018): Taxa related to Diplotaxis virgata (Brassicaceae) in northwest Africa. Phytotaxa 371 (3): 205-216, DOI: 10.11646/phytotaxa.371.3.4, URL: http://dx.doi.org/10.11646/phytotaxa.371.3.4
039BC700D7660761FF66F928FA29DB25.taxon	distribution	Distribution: — Diplotaxis australis is endemic to southwestern Morocco and is distributed south of the High Atlas in Regions Souss-Massa and Guelmim-Oued Noun, from the Atlantic coast eastwards up to at least Agdz (Zagora province), at elevations between 0 and 1400 m above sea level. Additional specimens examined: — MOROCCO. Region Souss-Massa: Agadir, 15 January 1970, Kaae (C); ibidem, 25 January 1970, Kaae (C); ibidem, 18 March 1972, P. & J. Davis 53486 (BM, E, RNG); ibidem, March 1935, Thretewy 76 (BM, K); ibidem, steppe, 200 m, 25 December 1983, Lewalle 10876 (BM); Agadir-m-Ighir, in ruderatis, 12 April 1931, Maire s. n. (MPU); Agadir, prope oppidulum Tafraute, 970 m, in dumosis secus viam, 26 May 1985, Blanché, Fernández Casas, Molero, Montserrat & Romo 9348 (BC 807550, BC 813651); 10 km N of Agadir, 1 April 1974, Miller & al. 416 (RNG); bords de l’Oued Tamerahrt, à 10 km au N d’Agadir, 2 July 1885, de Campou s. n. (P); plaine de Ksima, 2 April 1922, Maire s. n. (MPU); ad ripas fluminis Massa, in rupestris arenaceis, 3 April 1934, Maire & Wilczek s. n. (MPU, P); Tafraoute, to Souk Tleta de Tasserirt, 1300 – 1400 m, 27 March 1969, Davis 48801 (BM); Ammeln Valley, ca. Tafraoute, 29 March 1972, Davis 53858 (E, RNG); Taroudant, champs irrigués, 17 April 1922, Maire s. n. (MPU); Taroudant, vers le Ras-el-Oued, sous les arganiers, 14 April 1931, Jahandiez 146 (BM, K); in arganietis clivi septentrionalis Anti-Atlantis supra Tiout, 700 m, solo lapidoso calcareo, 5 April 1934, Maire & Wilczek s. n. (MPU); in Anti-Atlante: in arganietis lapidosis calcareis prope Tiout, 600 – 900 m, 19 April 1931, Maire s. n. (MPU); 6 km N of Tiznit, ca. El-Mader-El-Kebir, 3 April 1974, Miller & al. 541 (BM, RNG); Montagnes de Siggrat et Ghiliz, Sud-Ouest du Maroc, 1876, Mardoché, Herb. Cosson (P). Region Drâa-Tafilalet: 2 km E de Agdz, 4 May 1986, Martínez-Laborde 328 (MA 01 - 00916874); Ait Saoun, ca. 3 km westl. Oase am Trockental, 11 March 1992, Flügel M 97 (B). Region Guelmim-Oued Noun: Asserarid [Asserir, SE of Guelmim], 1875, Mardoché s. n. (P); Iter Ifniense, inter El Mers et El Had de Bifurna, 28 April 1935, Font Quer s. n. (BC 111698); in depressis cultis inter Labier [Labyar] et Notfia, 1 April 1933, Maire s. n. (MPU); in lapidosis arenaceis Anti-Atlantis prope Taghjicht, 700 m, 6 April 1935, Maire & Wilczek s. n. (MPU); Tifernit et Ida Oubakil, district de Tazeroualt, 1876, Mardoché (P); Uad sidi Ifni, 9 April 1935, Font Quer s. n. (BC 111703); Ifni-Uad Areksis, 7 May 1935, Font Quer s. n. (BC 111699); in cultis prope Sidi Ifni, 23 June 1934, Caballero s. n. (MA 46367); Oued Noun, March – June 1936, Ollivier s. n. (MPU); 20 – 30 km S of Goulimine, 50 – 100 m, 21 March 1972, P. & J. Davis 53624 (BM, E, RNG). Specimens obtained from cultivated material. SPAIN. Madrid: Ciudad Universitaria, spring 1985, Martínez-Laborde 194, culta ex semina BGV-UPM 3646 (MA 488559); ibidem, spring 1985, Martínez-Laborde 195, culta ex semina BGV-UPM 2973 (MA 488590); ibidem, spring 1985, Martínez-Laborde 196, culta ex semina BGV-UPM 4064 (MA 488592); ibidem, spring 1985, Martínez-Laborde 199, culta ex semina BGV-UPM 2976 (MA 488594); ibidem, spring 1985, Martínez-Laborde 197, culta ex semina BGV-UPM 2977 (MA 488591); ibidem, spring 1985, Martínez-Laborde 198, culta ex semina BGV-UPM 2991 (MA 488593); ibidem, spring 1985, Martínez-Laborde 200, culta ex semina BGV-UPM 1099 (MA 488595); ibidem, spring 1985, Martínez-Laborde 201, culta ex semina BGV-UPM 3003 - UPM (MA 488557); ibidem, spring 1985, Martínez-Laborde 206, culta ex semina BGV-UPM 1480 (MA 488558); ibidem, spring 1985, Martínez-Laborde 207, culta ex semina BGV-UPM 2980 (MA 488589); ibidem, spring 1985, Martínez-Laborde 208, culta ex semina BGV-UPM 5540 (MA 488635). Cultivated material. BGV-UPM 1099, BGV-UPM 1480, BGV-UPM 1498, BGV-UPM 1943, BGV-UPM 2973 BGV-UPM 2976, BGV-UPM 2977, BGV-UPM 2980, BGV-UPM 3003 BGV-UPM 3646, BGV-UPM 5540, BGV-UPM 7514. Observations: — Of the two groups of populations close to D. virgata previously distinguished by Martínez-Laborde & Gómez-Campo (1993), those growing in southwestern Morocco south of the High Atlas have a distinct pubescence, mostly retrorse-appressed and apparently disorderly arranged on sepals. They have not been described or named so far and therefore constitute a new species. Note 1: — Several specimens belonging to D. australis were cited by Nègre (1960) under D. virgata var. cavanillesiana f. tenuirostris and f. platystylos. Note 2: — Diplotaxis australis belongs to sect. Rhynchocarpum.	en	Martínez-Laborde, Juan B. (2018): Taxa related to Diplotaxis virgata (Brassicaceae) in northwest Africa. Phytotaxa 371 (3): 205-216, DOI: 10.11646/phytotaxa.371.3.4, URL: http://dx.doi.org/10.11646/phytotaxa.371.3.4
