taxonID	type	description	language	source
1F52DF65FAC270F55F190926B99313CB.taxon	materials_examined	Material examined. Holotype: male (17.6 x 13.3 mm) (ZRC 1992.7918), Danum Valley Field Centre, station 507, in dry stump on ridge, Lahad Datu, Sabah, Borneo, leg. H. K. Voris, 23 October 1990. Paratype: female (25.7 x 18.6 mm) (ZRC 1992.7919), Danum Valley, Lahad Datu, Sabah, Borneo, leg. S. C. Choy, 21 July 1989. Others: 1 male (30.8 x 20.5 mm), 1 female (30.1 x 20.5 mm, with 26 juvenile crabs) (ZRC 2017.1205), from water-filled tree buttress, ca. 35 cm above ground Danum Valley, Lahad Datu, Sabah, Borneo, Malaysia, 20 July 2017.	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
1F52DF65FAC270F55F190926B99313CB.taxon	materials_examined	Comparative material. Arachnothelphusa kadamaiana (Borradaile, 1900): 1 female (23.2 x 17.1 mm) (ZRC 2009.0094), Poring, Basin 1 A, Sabah, Malaysia, Borneo, coll. R. F. Inger et al., 12 August 1992; 3 males (21.1 x 15.8 mm, 22.8 x 16.5 mm, 25.3 x 18.5 mm) (ZRC 2002.0097), Crocker Range, Sabah, 5 ° 27 ' N 116 ° 03 ' E, coll. I. Das, 24 April 2001. Arachnothelphusa aff. kadamaiana: 1 female (19.0 x 14.2 mm) (ZRC 2002.0098), Bako National Park, Sarawak, coll. I. Das and L. Grismer, 27 March 2001. Arachnothelphusa merarapensis Grinang, Pui & Ng, 2015: Holotype male (22.5 x 16.8 mm) (ZRC 2016.0297), water-filled tree-hole, ca. 100 cm above ground, steep dipterocarp forest, Merarap Hot Spring Resort, Lawas, northern Sarawak, Malaysia, Borneo, 4 ° 22 ' 25.4 " N, 115 ° 26 ' 10.1 " E, 485 m asl, coll. J. Grinang and Y. M. Pui, 31 October 2014.	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
50DE103300730BFE71D77ADE0F070406.taxon	materials_examined	Material examined. Holotype: male (14.8 x 14.6 mm) (ZRC 2017.1298), in pitfall trap, primary forest, Danum Valley Field Centre, Sabah, Malaysia, coll. C. Colon, 22 November 1996. Others: 1 ovigerous female (14.9 x 15. mm) (ZRC 2017.1273), in water filled rotting log, Nature Trail, Danum Valley Field Centre, Sabah, Borneo, coll. local rangers, 21 July 2017.	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
50DE103300730BFE71D77ADE0F070406.taxon	materials_examined	Comparative material. Geosesarma sabanum Ng, 1992: holotype male (13.1 x 13.6 mm) (ZRC 2018.0296), on leaf of herb in forest, ca. 50 m from nearest stream, Tawau Hills Park, eastern Sabah, Malaysia, Borneo, coll. R. F. Inger, 3 November 1991.	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
E419790782B30585D0B076F303A49422.taxon	materials_examined	Material examined. Holotype: male (44.3 x 33.3 mm) (ZRC 1997.0796), Sungei Tawau, Tawau Hills Park, Tawau, Sabah, coll. P. Yam, 14 December 1991. Paratypes: 1 female (41.4 x 31.0 mm) (ZRC 1997.0797), same data as holotype. Others: 1 male, 2 females, 1 juvenile (ZRC 2000.2217), Lower Segama River, Danum Valley Field Centre, Lahad Datu, Sabah, coll. K. Martin-Smith, June 1996; 1 male (57.4 x 44.8 mm) (ZRC 1997.0798), Sungei Palum Tambum, near Danum Valley Field Centre, Lahad Datu, Sabah, coll. K. Martin-Smith, August 1996; 5 males (largest 57.4 x 43.5 mm), 1 female (ZRC 1997.0799), Sungei Palum Tambun, near Danum Valley Field Centre, Lahad Datu, Sabah, coll. K. M. Martin-Smith, 9 October 1996; 1 female (ZRC 2000.2218), Sepat Kalisun, stream 200 m from 4 ° 58 ' 04.8 " N, 117 ° 48 ' 56.5 " E, Danum Valley Field Centre, Lahad Datu, Sabah, coll. H. H. Tan et al., 1 October 1996; 1 male, 2 females (ZRC 2000.2210), Sungei Bole Ketabil tributary, 4 ° 57 ' 33.5 " N, 117 ° 51 ' 34.1 " E, Danum Valley Field Centre, Lahad Datu, Sabah, coll. H. H. Tan et al., 2 October 1996; 1 male, 2 females (ZRC 2000.2220), Cabin Stream, 50 km on road to Danum Valley Conservation Area, drains from Bukit Rafflesia, Lahad Datu, Sabah, coll. H. H. Tan et al., 2 October 1996; 1 male (ZRC 2000.2221), West Eight, forest stream, 800 m into conservation area, tributary of Sungei Segama, Danum Valley Field Centre, Lahad Datu, Sabah, coll. H. H. Tan et al., 4 October 1996; 2 females (ZRC 2008.0431), Danum Valley Rainforest Lodge, 5 ° 03 ' 2.9 " N, 117 ° 34 ' 34.8 " E, Lahad Datu, Sabah, 3 October 1996; 1 female (56.0 x 42.0 mm) (ZRC 1989.3419), Sungei Madai, Madai Caves, Sabah, coll. R. Goh, 27 January 1985; 1 male (ZRC 1997.0802), Tawau, Sungei Matarid, Gua Madai, Jalan Madai, 4 ° 43 ' 8.7 " N, 118 ° 9 ' 14.7 " E, Tawau, Sabah, H. H. Tan et al., 6 October 1996.	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
65F0178A1FDE79E0B1A0C3F1CD362224.taxon	materials_examined	Material examined. Holotype: male (40.0 x 30.0 mm) (ZRC 1989.2024), stream outside Simud Hitam Cave, Gomantong, Sabah, Borneo, ca 5 ° 33 ' N 118 ° 06 ' E, coll. P. Chapman, 27 March 1986. Paratypes: 1 male (ZRC 1989.2192), Simud Puteh Cave, Gomantong, Sabah, Borneo coll. P. Chapman, 27 March 1986; 2 males (ZRC 1990.0445 - 0446), 1 male (ZRC 1989.3402), Sungei Madai, in stream just outside Madai Caves, Kunak, Lahad Datu district, Sabah, Borneo, ca. 04 ° 44 ' N 118 ° 12 ' E, coll. January 1985; 1 male (73.0 x 73.0 mm) (ZRC 1990.0444), Sungei Binuang, stream adjacent to Baturung Caves, Kunak, Lahad Datu district, Sabah, Borneo, 4 ° 43 ' N 117 ° 59 ' E, coll. R. Goh, 20 March 1985. Others: 1 female (ZRC 1989.3403), Sungei Madai, in stream just outside Madai Caves, Kunak, Lahad Datu district, Sabah, Borneo, ca. 04 ° 44 ' N 118 ° 12 ' E, coll. R. Goh, January 1985; 1 male (ZRC 1996.1897), Gomantong Caves Sabah, coll. C. L. Chan, January 1995; 1 male, 1 female (ZRC 2009.0091), Gomantong Caves, coll. D. Chia, 21 December 1999; 1 female (ZRC 1996.1999), Sungei Binuang, Banturung Caves, Lahad Datu, coll. R. Goh, 20 March 1989; 1 male (ZRC 1990.0571), Danum Valley, Sabah, coll. R. Stuebing, 1980 s; 1 male (ZRC 1990.0568), Danum Valley, Lahad Datu, Sabah, coll. R. Stuebing, 23 July 1989; 1 male, 2 females (ZRC 1008.1346), Sungei Palun Tambun, tributary of Sungei Segama, upstream of Danum Valley Field Centre, Lahad Datu, Sabah, coll. H. H. Tan et al., 1 October 1996; 2 males, 5 females (ZRC 1996.1998), Kallang Sebaru stream, 4 ° 58 ' 4.8 " N, 107 ° 48 ' 56.5 " E, Danum Valley, Lahad Datu, Sabah, coll. H. H. Tan et al., 1 October 1996; 1 male, 1 female (ZRC 1996.1997), Sepat Kalisun, Danum Valley, Lahad Datu, Sabah, coll. H. H. Tan et al., 1 October 1996; 2 males (ZRC 1996.1995) Ca Gin Stream Right, 4 ° 59 ' 8.5 " N, 107 ° 54 ' 5.1 " E, Danum Valley, Lahad Datu, Sabah, coll. H. H. Tan et al., 2 October 1996; 2 females (ZRC 1996.2000), Sungei Bole Ketabil tributary, 4 ° 57 ' 33.5 " N, 117 ° 51 ' 34.1 " E, Danum Valley Field Centre, Lahad Datu, Sabah, coll. H. H. Tan et al., 2 October 1996; 1 male, 2 females (ZRC 1996.1994), Sungei Bole Ketabil tributary, 4 ° 57 ' 33.5 " N, 117 ° 51 ' 34.1 " E, Danum Valley Field Centre, Lahad Datu, Sabah, coll. H. H. Tan et al., 2 October 1996; 2 males, 2 females (ZRC 2010.0045), West Six stream, tributary of Sungei Segama, 600 m inside conservation area, coll. H. H. Tan et al., 4 October 1996; 1 male, 1 female (ZRC 1996.2004), West Eight, forest stream, 800 m into conservation area, tributary of Sungei Segama, Danum Valley Field Centre, Lahad Datu, Sabah, coll. H. H. Tan et al., 4 October 1996; 2 males (ZRC 2009.0309), Sungei Palum Tambum, near Danum Valley Field Centre, Lahad Datu, Sabah, coll. K. Martin-Smith, 9 October 1996; 1 female (34.7 x 28.4 mm) (ZRC 2017.1268), Danum Valley Field Centre, in forest streams, Lahad Datu, Sabah, Borneo, at night, coll. locals, 20 July 2017; 1 male (27.8 x 22.7 mm), 1 young female (17.4 x 5.3 mm) (ZRC 2017.1269), Danum Valley Field Centre, just outside dorms in streams, at night, Lahad Datu, Sabah, Borneo, coll. 22 July 2017; 1 male, 1 female (ZRC 2008.483), Maliau Basin, stream draining into Sungei Maliau, coll. S. H. Tan and T. H. T. Tan, 15 - 17 May 1996; 2 males, 1 female (ZRC 2008.0607), Maliau Basin, tributary of Sungei Maliau, adjacent Camp 88, coll. S. H. Tan and T. H. T. Tan 15 - 17 May 1996; 1 male, 1 female (ZRC 1997.0104), Maliau Basin, coll. S. H. Tan and T. H. T. Tan, 13 - 17 May 1996; 1 female (ZRC 1989.2194), Maliau Basin, Sabah, coll. Sabah Foundation Expedition 1988; 1 male, 2 females (ZRC 1996.2005), Tawau Plateau, Telupid Sandakan stream, coll. R. Goh, 1990; 5 males (ZRC 2008.1345), stream by Air Panas, near base of Tawau Hills Park, coll. H. H. Tan et al., 5 October 1996; 1 male 4 females (ZRC 1996.2008), Tawau, Sabah, 4 ° 18 ' 03 " N, 117 ° 54 ' 20.7 " E, coll. H. H. Tan et al., 5 October 1996; 2 males, 4 females (ZRC 1996.2001), Tawau, Sungei Matarid, Gua Madai, Jalan Madai, 4 ° 43 ' 8.7 " N, 118 ° 9 ' 14.7 " E, Tawau, Sabah, H. H. Tan et al., 6 October 1996; 1 male (ZRC 1994.4201), Danau Biandum, Kinabatangan River, Sabah, coll. S. H. Tan et al., 8 April 1994.	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
13CD62343A5A106CAA2D6CF5235A056D.taxon	materials_examined	Material examined. Holotype male (29.2 x 20.4 mm) (ZRC 2018.0297), found dead in pool adjacent to Borneo Rainforest Lodge, next to Danum Valley Conservation Area, Lahad Datu, Sabah, 4 ° 58.2 ' N 117 ° 41.4 ' E, ca. 600 m asl, East Malaysia, Borneo, coll. local ranger, 28 May 2015.	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
376D79F29B3F10389E3AC429E40E69D6.taxon	materials_examined	Material examined. Holotype: male (23.9 x 18.4 mm) (ZRC 2017.1294), coll. Danum Valley, Lahad Datu, Sabah, Borneo, Malaysia, 22 July 2017. Paratypes: 1 male (18.8 x 15.5 mm) (ZRC 2017.1295), same data as holotype; 1 male (19.9 x 16.4 mm) (ZRC 2009.0080), in pitfall trap, Danum Valley Research Centre, Sabah, coll. C. Colon, October 1996. Others: 3 juveniles (3.7 x 3.0 mm, 6.5 x 5.4 mm, 6.7 x 5.5 mm), 1 young female (10.7 x 9.1 mm) (ZRC 1990.0548 - 0551), Danum Valley Research Centre, Lahad Datu, Sabah, coll. R. Stuebing, 23 July 1989.	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
376D79F29B3F10389E3AC429E40E69D6.taxon	diagnosis	Diagnosis. Carapace broader than long, not raised; dorsal surface with regions clearly demarcated; frontal median triangle absent (Figs 2 A-C, 3 E); epibranchial tooth low, distinct, separated from external orbital tooth by shallow cleft; epigastric regions raised, rugose, not cristate; postorbital cristae low, distinct, rugose, not confluent with epigastric cristae, not reaching anterolateral margin; cervical grooves and H-shaped gastric depression deep; gastric regions with prominent transverse striae; antero- and posterolateral regions with strong oblique striae (Fig. 2 A, B); median lobe on posterior margin of epistome triangular, tip rounded (Figs 2 C, 3 E). Third maxilliped with subrectangular ischium, distinctly longer than broad (Fig. 2 D). Chelipeds with outer surface of palm almost smooth, dorsal and lateral surfaces of adult male dactylus covered with dense short setae (Fig. 3 A-D). Ambulatory legs not prominently elongate, dorsal margin of merus gently serrated (Figs 2 A, 3 F-I). Thoracic sternum with surface evenly pitted to smooth, sternopleonal cavity reaching imaginary line joining anterior edges cheliped coxae (Fig. 2 F, G); pleon distinctly T-shaped, somite 6 rectangular, slightly more than twice as long as broad, telson triangular, longer than broad (Fig. 2 E, F). G 1 relatively slender, almost straight; terminal segment approx. a quarter length of subterminal segment (Fig. 4 A-C). G 2 approx. two-thirds length of G 1, distal article short (Fig. 4 D).	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
376D79F29B3F10389E3AC429E40E69D6.taxon	description	Description of male holotype. Carapace broader than long, not raised; dorsal surface gently convex, regions clearly demarcated, covered with very short setae which does not obscure surface; frontal margin almost straight, without distinct median concavity, not deflexed, approx. a third carapace width; frontal median triangle absent (Figs 2 A-C, 3 E); anterolateral margin not clearly separated from posterolateral margin; external orbital tooth low, broadly triangular; epibranchial tooth low but distinct, separated from external orbital tooth by shallow cleft; postfrontal surface slightly rugose; postorbital region surface; epigastric regions raised, rugose, not cristate, divided into 2 parts by narrow, deep median groove; postorbital cristae low but distinct, sharp, not confluent with epigastric cristae, not reaching anterolateral margin; cervical grooves and H-shaped gastric depression deep; gastric regions with prominent transverse striae; antero- and posterolateral regions with strong oblique striae; posterolateral margins concave, gently converging towards posterior carapace margin; posterior carapace margin straight (Fig. 2 A, B); pterygostomial, suborbital, sub-branchial, and subhepatic regions covered with striae (Fig. 2 C); orbits large, eyes occupying entire width, supra and suborbital margins entire, cristate; eyes pigmented, well developed, peduncle with low, sinuous median ridge (Fig. 2 B, C); median lobe on posterior margin of epistome triangular, tip rounded, lateral margins sinuous (Figs 2 C, 3 C). Mandibular palp 2 - segmented, terminal one distinctly bilobed. Third maxilliped covering majority of buccal cavity when closed; ischium subrectangular, distinctly longer than broad, with shallow submedian groove; merus quadrate, slightly broader than long; exopod long, slender, reaching median part of merus, flagellum long, exceeding width of merus (Fig. 2 D). Chelipeds asymmetrical, right larger; surface of merus slightly rugose, relatively long, trigonal in cross section, margins without teeth or spines; carpus surface distinctly rugose, subovate, inner distal angle with sharp spine with basal tubercle; palm relatively stout, longer than broad, outer surface slightly rugose to almost smooth; fingers subequal in length to palm, dactylus marginally longer than pollex, curving inwards, cutting margin of fingers lined with numerous denticles, fingers pitted (Figs 2 A, 3 A-D); dorsal and lateral surfaces of most of dactylus covered with dense short setae; lateral surface of pollex with mat of short, relatively less dense setae; tips of fingers strongly curved, corneous, glabrous (Fig. 3 B-D). Ambulatory legs not prominently elongate, third pair longest, fourth leg shortest; segments laterally flattened laterally, surfaces mildly rugose; dorsal margin of merus gently serrated, no visible subdistal tooth; carpus of first to third legs with low median ridge, absent on carpus of fourth leg; margins of propodus and dactylus lined with numerous short spines (Figs 2 A, 3 F-I). Thoracic sternum surface evenly pitted to smooth; sternites 1 and 2 completely fused forming triangular structure; suture separating sternites 2 and 3 relatively shallow, sinuous, medially convex with lateral parts concave (towards buccal cavity); sternites 3 and 4 completely fused; sternopleonal cavity almost reaching imaginary line joining anterior edges cheliped coxae, near suture between sternites 2 and 3; part of sternite 8 exposed when pleon closed; tubercle of male pleonal locking mechanism prominent, peg-like, on anterior third of sternite 5 (Fig. 2 F, G). Pleon distinctly T-shaped; somite 1 short, broad, reaching coxae of fourth ambulatory legs; somite 2 slightly longer than somite 1, as broad as somite 1; somite 3 short, broadest, with prominently convex lateral margins; somites 4 and 5 trapezoidal; somite 5 notably narrower than 4, trapezoidal with concave lateral margins; somite 6 rectangular, slightly more than twice as long as broad, lateral margins concave; telson triangular, longer than broad, tip rounded (Fig. 2 E, F). G 1 relatively slender, entire structure almost straight; terminal and subterminal segments clearly separated; terminal segment relatively short, approx. a quarter length of subterminal segment, cylindrical with tip tapering to subtruncate tip, margins with short stiff setae, surface just before tip with numerous squamiform setae; lower half of subterminal segment with numerous short setae (Fig. 4 A-C). G 2 approx. two-thirds length of G 1; basal segment long; distal segment short (Fig. 4 D).	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
376D79F29B3F10389E3AC429E40E69D6.taxon	etymology	Etymology. The name is derived from the Latin capillus for hair and digitus for finger. The name is used as a noun in apposition.	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
1AD6EEE48235CBB3A5F50F3FBE885B93.taxon	materials_examined	Material examined. Holotype: male (14.0 x 12.0 mm) (ZRC 1989.3588), in pitfall trap, adjacent to pool of rhinoceros mud wallow, Danum Valley, Lahad Datu, Sabah, ca. 4 ° 55 ' N 117 ° 46 ', coll. R. Stuebing, 4 March l 988. Paratypes: 1 female (18.3 x 15.0 mm), 2 juveniles (ZRC 1989.3592 - 3594), in pitfall trap, adjacent to pool of rhinoceros mud wallow, Danum Valley, Lahad Datu, Sabah, ca. 4 ° 55 ' N 117 ° 46 ', coll. R. Stuebing, 1 March 1988; 1 male (18.5 x 15.4 mm) (ZRC 1989.3591), in mist net in rhinoceros mud wallow pool, Danum Valley, Lahad Datu, Sabah, ca. 4 ° 55 ' N 117 ° 46 ', coll. R. Stuebing, 2 March l 988; 1 male, 1 female (ZRC 1989.3589 - 3590), in rhinoceros mud wallow pool, Danum Valley, Lahad Datu, Sabah, ca. 4 ° 55 ' N 117 ° 46 ', coll. R. Stuebing, 2 March l 988. Others: 1 female (11.0 x 9.5 mm) (ZRC 1997.0138), in pitfall trap, Danum Valley Field Centre, Lahad Datu, Sabah, coll. C. Colon, 14 October 1996; 1 male (16.4 x 13.5 mm) (ZRC 1997.0140), in pitfall trap, Danum Valley Field Centre, Lahad Datu, Sabah, coll. C. Colon, 10 October 1996; 1 male (20.0 x 15.8 mm) (ZRC 1997.0141), in pitfall trap, Danum Valley Field Centre, Lahad Datu, Sabah, coll. C. Colon, 17 October 1996; 1 male (22.0 x 17.6 mm) (ZRC 1997.0142), in pitfall trap, Danum Valley Field Centre, Lahad Datu, Sabah, coll. C. Colon, 19 October 1996; 1 male (10.5 x 9.0 mm), 1 female (17.8 x 14.9 mm) (ZRC 2017.1272), in mud and leaves under wooden walk-way, Orchid Trail, Danum Valley Field Centre, Lahad Datu, Sabah, at night, coll. local rangers, 20 July 2017; 1 female (26.1 x 21.3 mm) (ZRC 2017.1047), in pool along wooden walkway at night, Orchid Trail, Danum Valley Field Centre, Lahad Datu, Sabah, at night, coll. local rangers, 20 July 2017; 1 female (24.0 x 17.5 mm) (ZRC 1997.0139), Kunak, Baturong, Binuang River, Lahad Datu, Sabah, coll. R. Stuebing, 20 March 1989.	en	Ng, Peter K. L., Ng, Paul Y. C. (2018): The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). ZooKeys 760: 89-112, DOI: http://dx.doi.org/10.3897/zookeys.760.24787, URL: http://dx.doi.org/10.3897/zookeys.760.24787
