identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039887C4A31CF4727F387E3C31FD553E.text	039887C4A31CF4727F387E3C31FD553E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mahea Distant 1909	<div><p>Mahea Distant, 1909</p><p>Mahea Distant, 1909: 32, pl. 4 (description, figures). Type species: Mahea sexualis Distant, 1909, by monotypy.</p><p>Muschalea Cachan, 1952: 312 (description, figures). Type species: Muschalea andriai Cachan, 1952, by original designation (syn. KUMAR 1974).</p><p>Mahea: KUMAR (1974): 43 -44 (diagnosis, taxonomy).</p><p>Redescription. Body deltoid in shape; head shorter than pronotum; pronotum arched, sloping posteriad, highest between humeral angles, then descending caudad; dorsum flat; ventral side convex. Body surface slightly shining, covered with coarse dark punctures.</p><p>Head (Fig. 1). Mandibular plates sinuated in front of eyes, apically curved inwards, never spinously produced; clypeus with apex narrowly exposed or completely enclosed (intraspecific variable character). Eyes large, protruding from head outline by most of its width; ocelli situated behind imaginary line through posterior margins of eyes; each ocellus nearer to eye than to each other. Antenniferous tubercles visible from above; antennae 4-segmented (Fig. 11); antennomere 1 (scape) short, not or only very slightly surpassing apex of head; antennomere 2 (pedicel) very long, not subdivided, with erect pubescence; antennomeres 3 and 4 subequal in length, together about as long as antennomere 2, without erect pubescence. Bucculae low, diverging posteriad; maxillary plate tubercle developed; rostrum long, reaching or surpassing metacoxae; first rostral segment not surpassing posterior margin of eyes, hidden between bucculae.</p><p>Pronotum trapezoidal (Fig. 28); anterior margin more or less slightly concave, anterolateral angles touching posterior margins of eyes; antero-lateral margins rounded, more or less diverging posteriad; humeral angles always very prominent and spinuously produced (Fig. 2); posterior margin skewed posteriad to antero-lateral scutellar angles, margin anterior to scutellum concavely sinuate. Scutellum triangular, longer than clavi. Hemelytra narrow; clavi narrowly triangular (anteriorly with two rows of punctures, posteriorly with one row); corium long, triangular, acutely attenuated posteriorly and reaching middle of membrane, its posterior inner margin concave; suture between corium and membrane brown; membrane large, long oval, slightly brownish infuscated, translucent, slightly surpassing (exceptionally nearly surpassing) apices of postero-lateral angles of abdominal sternum 7 (Fig. 14).</p><p>Prosternum with deep median groove, flanked by longitudinal, prominently elevated carinae. Mesosternal carina flattened laterally, extending anteriorly between procoxae and reaching posterior margin of prosternal groove, posteriorly reaching between metacoxae. Metathoracic scent gland complex inconspicuous (Fig. 31); ostiole small, situated laterally between meso- and metacetabula; peritreme horizontal, slightly raised above metapleura, oval, shining; evaporatorium small, narrowly surrounding ostiole and peritreme. Metapleura swollen (Fig. 14) or not. Legs pale, ochraceous, with short to very short pubescence; femora twice as long as tibiae; tarsi 2-segmented; claws slender, long, straight, apically abruptly rectangularly curved; parempodia and pulvilli developed.</p><p>Abdomen with median carina well developed on sterna 3-6 in males (Fig. 4) and on sterna 3-4 in females (Fig. 5), produced anteriad as abdominal spine reaching between metacoxae, resting against mesosternal carina. Connexiva (especially its posterolateral corners) with more or less prominent spines (variable between species and sexes) (Figs. 4-5, 14, 16); especially postero-lateral angles of sternum 7 in males conspicuously produced; sternum 8 slightly incised posteromedially.</p><p>Male genitalia. Pygophore subquadrate (Figs. 7, 17, 25), more or less dorso-ventrally flattened (Figs. 6, 16, 24); margins of external opening setose; dorso-lateral angles slightly produced, with tuft of conspicuous setae; proctiger large, convex, exposed; hypophyses of parameres exposed from pygophore (Figs. 7, 17, 25); parameres flat, head enlarged, not bifurcate, bearing setae (Figs. 18, 26); vesica of aedeagus long, deeply sinuated (Fig. 19) or looped (Fig. 27).</p><p>Sexual dimorphism strongly developed. Body of females behind humeral angles less narrowed; metapleura only slightly swollen; abdomen broader, median carina short, posterior part of abdomen flat; connexival spines with more than one spine per sternum, especially on sterna 5-7; Pendergrast’s organ not developed. Hind femora and tibiae are most probably sexually dimorphic in some/all species; I cannot confirm that given the lack of material.</p><p>Differential diagnosis. According to KUMAR (1974), the presence of both abdominal spine and mesosternal carina places Mahea as a member of the subfamily Acanthosomatinae . Within the Acanthosomatinae, Mahea and Catadipson are the only genera with 4-segmented antennae. Catadipson differs from Mahea in the following characters: elongate oval species; head very broad, slightly broader than two thirds of pronotal width; apex of clypeus free; mandibular plates projected as conspicuous apical spines; humeral angles of pronotum not produced, rounded; metathoracic scent gland complex as in Fig. 32; connexival margins without apparent spines; head and pronotum with very large black punctures.</p><p>Distribution. This genus is known only from Madagascar (four species) and the Seychelles (Island of Mahé) (one species).</p><p>Comments. KUMAR (1974) noticed the resemblance in the descriptions of Mahea and Muschalea and considered them to be synonymous. He, however, did not examine Cachan’s type specimens of Muschalea . This synonymy is herein confirmed based on the study of type material.</p></div>	https://treatment.plazi.org/id/039887C4A31CF4727F387E3C31FD553E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A31AF4767CE87C9D30E5504D.text	039887C4A31AF4767CE87C9D30E5504D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mahea sexualis Distant 1909	<div><p>Mahea sexualis Distant, 1909</p><p>(Figs. 1-7, 31, 40-41)</p><p>Mahea sexualis Distant, 1909: 32-33, pl. 4 (description, figure of female).</p><p>Mahea sexualis: KUMAR (1974): 44 (taxonomy, invalid lectotype designation).</p><p>Type locality. Seychelles, Mahé.</p><p>Type material. LECTOTYPE: ♁, ‘ Seychelles / Mahe / H P.T. 1906 [p] // Mahea / sexualis / Dist. [hw] // University Museum of / Zoology, CAMBRIDGE [p] // LECTOTYPUS / MAHEA / SEXUALIS / Distant, 1909 / des. P. KMENT 2005’ [p, red label] (CUMZ), here designated. The lectotype is glued onto the tip of a pentagonal piece of card, with the pygophore removed and mounted on the same piece of card. The original piece of card is pinned under the new one. Antennomeres 3 and 4 of both antennae, right middle and hind legs, and all tarsi from left legs of the lectotype are missing. PARALECTOTYPES: 1 ♁ and 1 ♀, ‘ Seychelles / Mahe / H P.T. 1906 [p] // Distant Coll. / 1911-383 [p] // SYN- / TYPE [p, white circle with blue margin] // PARALECTOTYPUS / MAHEA / SEXUALIS / Distant, 1909 / des. P. KMENT 2005’ [p, red label] (BMNH). The male paralectotype is pinned, its antennomeres 2-4 (left) and 3-4 (right), all left legs (except hind femur), right hind leg, apical part of scutellum and pygophore missing, and it has a large pinpoint in the scutellum. The female paralectotype is mounted on the tip of a pentagonal piece of card, with one separate leg glued on the same piece of card. Rest of legs and antennae are missing, right pronotal spine broken, and abdomen ruptured ventrally on the left side.</p><p>Redescription. Body slightly shining; basic colour ochraceous, with coarse dark punctures.</p><p>Male (lectotype) (Fig. 40). Length 7.5 mm, width of pronotum between humeral angles 4 mm. Head (Fig. 1) pale ochraceous with coarse brown punctures forming more or less apparent rows (two rows on each mandibular plate, posteriorly coalescent, forming a figure resembling an 8 next to each eye; two rows on clypeus, partly coalescent medially, reaching to posterior margin of head, on frons surrounded by two rows forming an incomplete circle). Head shorter than wide (1: 1.35), its width about half that of pronotum between humeral angles (1: 2.08). Mandibular plates basally almost parallel, suddenly curved inwards apically, not meeting; apex of clypeus free (Fig. 1). Eyes large, ochraceous; ocelli ochraceous, situated behind eyes, near anterior pronotal margin. Each antenniferous tubercle with black longitudinal spot laterally; antennae castaneous; antennomere 1 pale, basally ochraceous; antennomere 2 with erect pubescence, hairs slightly shorter than diameter of antennomere; length of antennomeres: 1 – 0.4 mm, 2 – 2.3 mm (ratio 1: 5.75). Head ventrally pale ochraceous, without punctures; apex of rostrum blackened, reaching middle of sternum 4.</p><p>Pronotum ochraceous with thick brown punctures; punctures on posterior half more dense than on anterior half; pronotal spines and narrow median line ochraceous; anterior margin concave; antero-lateral margins widening posteriad; humeral angles prominent, each abruptly produced into long, narrow spine directed laterad (Figs. 2-3). Scutellum triangular, ochraceous, anterior part medially brownish, with brown punctures; anterior part arched, sloping posteriad; posterior part flat, apex blackened, triangular. Hemelytra ochraceous with brown punctures; suture between corium and membrane brown; membrane slightly brownish, translucent, slightly surpassing postero-lateral angles of sternum 7.</p><p>Thorax ventrally with coarse brown punctures, which are more sparse on metapleura; peritreme ochraceous. Metapleura swollen, its postero-lateral corners visible in dorsal view.</p><p>Legs ochraceous, apices of tibiae and tarsomeres 2 slightly blackened; hind femur and tibia with inner margin granulated, femur slightly swollen and curved inwards, tibia straight. Abdomen with only connexival spines visible from above; ventrally ochraceous, sterna 3-6 each with small black spot on each side near middle; median carina on sterna 3-6 well developed; postero-lateral angles of sterna 3-5 with small spines, sternum 6 with long posterolaterally directed spine on each side; sternum 7 parallel-sided, postero-lateral angles strongly produced posteriad; posterior margin of sternum 8 slightly incised medially (Fig. 4).</p><p>Pygophore (Figs. 6-7) brownish, dorso-ventrally flattened, postero-lateral angles each with tuft of long setae; parameres apically pointed, ventrally with setigerous punctures.</p><p>Female (paralectotype) (Fig. 41). Length 8.2 mm, width of pronotum between humeral angles approximately 4.1 mm (right spine missing); punctures on body reddish; metapleura only slightly swollen, less prominent in dorsal view; abdomen (Fig. 5) less narrowing posteriad, more flattened, unicolorous, without black round spots, median carina developed only on sterna 3-4; connexival spines as in Fig. 5.</p><p>Variability. Male paralectotype differs from the lectotype in having the clypeus completely enclosed by mandibular plates; body length 7.4 mm, length of antennomere 2 equal to 2.4 mm. Differential diagnosis. Mahea sexualis differs from M. andriai, M. distanti sp. nov., and M. parvula by having each humeral angle abruptly produced into a long spine, not conical. From M. durrelli sp. nov. it differs by having the lateral pronotal margins regularly sinuated, the humeral spines not raised (Figs. 2-3), the metapleura distinctly swollen, differently shaped connexival spines (Figs. 4-5), and apex of each paramere pointed (Figs. 6-7) (see also the key).</p><p>Bionomy. Unknown.</p><p>Distribution. Island of Mahé (Seychelles). Known only from the type series.</p><p>Comments. DISTANT (1909) described M. sexualis based on both sexes, but he did not mention the number or the location of syntypes. The type locality was given unambigously as ‘Locality. Mahé’, but on the next line DISTANT (1909) wrote: ‘Dr. Schouteden has kindly sent me a male specimen of this species from Madagascar.’ The description was accompanied by a figure of a female from Mahé and a male from Madagascar.</p><p>KUMAR (1974) examined three specimens of M. sexualis deposited in the BMNH – one male and one female from Mahé, and one male from Madagascar. He wrote: ‘… the Madagascar male is a distinct species, quite different from Mahé specimens. … Distant’s description covers both species and his type label is affixed to the Madagascar specimen. In these circumstances, I have taken the Madagascar specimen to be the type and have placed a label on it indicating this is the holotype. The species represented by the specimens from Mahé is being described elswhere.’ The Madagascar specimen, however, did not originate from the type locality, and thus it cannot be a syntype of M. sexualis either by DISTANT (1909) or KUMAR (1974). The lectotype designation by KUMAR (1974) is therefore unjustified and in valid.</p><p>I had the opportunity to study three specimens of M. sexualis – one male and one female from BMNH, and one male from CUMZ – with the same locality labels and determination labels written most probably by the same hand (see Type material). Therefore, I regard all these specimens to be syntypes of M. sexualis . The male from CUMZ was choosen as the lectotype because it is better preserved and has the pygophore, which is lost in the male from BMNH. The Madagascar specimen designated as a lectotype by Kumar (1970) is currently described as M. distanti sp. nov.</p></div>	https://treatment.plazi.org/id/039887C4A31AF4767CE87C9D30E5504D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A319F4787C9C7FDD3504526D.text	039887C4A319F4787C9C7FDD3504526D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mahea andriai (Cachan 1952)	<div><p>Mahea andriai (Cachan, 1952)</p><p>(Figs. 8-10, 42)</p><p>Muschalea Andriai Cachan, 1952: 312-313, Figs. 193 -194 (p. 311), 13 (Pl. XIV, p. 439) (description, figures).</p><p>Mahea andriai: KUMAR (1974): 43 -44 (diagnosis, taxonomy).</p><p>Type locality. South-east Madagascar, valley of Iantara by Ivohibe.</p><p>Type material. HOLOTYPE: ♀, ‘ Valleé de l’Iantara [hw] / Inst. Scient. Madagascar [p] / 10-XI- 50, A. R. [hw] // TYPIS [p, pink label] // Muschalea / andriai / Cachan [hw] // face side : HOLOTYPUS / Muschalea / andriai / Cachan 1952 / reverse: P. Kment 2005’ [hw, red label] (MNHN). Holotype pinned; left antennomeres 3-4 and left middle and hind legs missing.</p><p>Redescription. Female (holotype) (Fig. 42). Body yellowish ochraceous. Length 9 mm, width of pronotum between humeral spines 6 mm.</p><p>Head yellowish ochraceous, margins and antenniferous tubercles laterally darkend; punctures on the head finer than in other species, slightly contrasting, their coalescence hardly intimated; slightly shorter than wide, less than 0.33 times as wide as pronotum. Mandibular plates sinuated before eyes, meeting in front of clypeus. Eyes black. Antennae dark brown, antennomeres 3-4 slightly paler; antennomere 2 with short pubescence (ca half of its diameter); length ratio of antennomeres – 1: 5: 2.25: 2. Head ventrally ochraceous, with coarse and thick dark punctures; apex of rostrum blackened, reaching base of abdomen.</p><p>Pronotum coarsely and thickly punctate, punctures brown, darker on margins; anterior and posterior margins slightly concave; antero-lateral margins concave; each humeral angle prominent, produced gradually into sharp, conical, black spine directed laterad and somewhat raised (Figs. 8-9). Scutellum triangular, with sparse concolorous punctures, basal angles slightly blackened; proximal part convex, apical part depressed laterally, with vague median carina; apex narrowly rounded, with black spot. Hemelytra with coarse concolorous punctation, membrane hyaline, translucent, apically distinctly surpassing abdominal apex.</p><p>Thorax yellowish brown similar to upper surface; pleura with dispersed small brown punctures. Legs ochraceous.</p><p>Abdomen ventrally without apparent punctures, with small black roundish spots near middle on each side of sterna 3-6; median carina on abdominal venter nearly obsolete; connexival spines as in Fig. 10.</p><p>Differential diagnosis. Mahea andriai differs from both M. sexualis and M. durrelli sp. nov. by having the lateral pronotal angles conically produced and gradually tapering to a spinose apex. From M. parvula, it differs by the larger body, more prominent lateral pronotal angles, and relatively narrower head. From M. distanti sp. nov. it differs by the following characters: Body almost uniformly yellowish ochraceous with small dark punctures (except darkened lateral corners of pronotum, apex of scutellum, antennae and paired black spots laterally on abdominal venter). Head with only indicated impressed rows of pale punctures. Apex of scutellum more broadly rounded. Sternum without black punctures. Hairs on antennomere 2 shorter than its diameter. Thorax not swollen. Lateral connexival spines as in Fig. 10.</p><p>Bionomy. Unknown.</p><p>Distribution. South-east Madagascar. Known only from the type locality (CACHAN 1952).</p></div>	https://treatment.plazi.org/id/039887C4A319F4787C9C7FDD3504526D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A317F47D7CEB7D3D33AC52CD.text	039887C4A317F47D7CEB7D3D33AC52CD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mahea distanti	<div><p>Mahea distanti sp. nov.</p><p>(Figs. 11-19, 43)</p><p>Mahea sexualis (misidentification): DISTANT (1909): 33, pl. 4 (record, figure of male).</p><p>Mahea sexualis (misidentification): KUMAR (1974): 44 (taxonomy).</p><p>Type locality. Madagascar, Tamatave [= Toamasina].</p><p>Type material. HOLOTYPUS: ♁, ‘Type / H. T. [p, white circle with red margin] // Mahea / sexualis / type Dist. [hw] // Madagascar / Tamatave / XII. [hw] // Distant Coll. / 1911-383 [p] // SYN- / TYPE [p, white circle with blue margin] // HOLOTYPUS / MAHEA / DISTANTI / sp. nov. / det. P. KMENT 2005 [p, red label]’ (BMNH). Holotype on micro pin; left antenna, right antennomere 4, left fore leg, right fore and middle tibiae and tarsi and both hind tarsi missing; pronotum ventrally detached from mesonotum, dorsally cracked on the right side, right humeral spine missing. The pygophore and male genitalia of the specimen had been dissected and stored in glycerol in a small glass microvial attached to the same pin.</p><p>Description. Male (Fig. 43). Length 8.2 mm. Body ochraceous, slightly shining, with thick coarse dark punctures.</p><p>Head pale ochraceous with coarse black or brown punctures, forming more or less distinct rows (two regular rows on each mandibular plate, rest of head with more irregular pattern than in M. sexualis, ‘circular’ figure on vertex not developed). Head shorter than wide across eyes (1: 1.14), its width about half of pronotal width between humeral angles (1: 2.75). Mandibular plates almost parallel basally, smoothly rounded apically, enclosing apex of clypeus. Eyes brownish; ocelli reddish, situated behind imaginary line through posterior margins of eyes, but more distant from anterior pronotal margin than in M. sexualis . Each antenniferous tubercle laterally with black longitudinal spot; antennomere 1 blackish brown; antennomere 2 pale castaneous, with erect pubescence, hairs slightly longer than diameter of antennomere; antennomere 3 castaneous; lengths of antennomeres: 1 – 0.4 mm, 2 – 2.9 mm, 3 – 1.3 mm (ratio 1: 7.25: 3.25) (Fig. 11). Head ventrally ochraceous with small brown punctures; rostrum ochraceous, apex blackened, reaching base of abdomen.</p><p>Pronotum ochraceous with uniform brown punctures, median carina not developed, humeral angles and spines reddish; anterior margin strongly concave, head more projected into pronotum than in M. sexualis; antero-lateral margins rounded, divergent laterad; each humeral angle conically produced into spine, directed laterad and slightly upwards (Figs. 12, 13). Scutellum triangular, its punctures finer than those on pronotum; anterior part only slightly arched, castaneous, only antero-lateral angles ochraceous; lateral margins slightly depressed medially; posterior part flat, ochraceous, with inconspicuous dark punctures; apex black, triangular. Hemelytra ochraceous with brown punctures; suture between corium and membrane brown; membrane with brownish tinge, translucent, slightly surpassing postero-lateral angles of sternum 7.</p><p>Thorax ventrally ochraceous with coarse brown punctures; brown colour of punctures locally coalescent, forming small to large irregular spots on median parts of pleura; acetabula, peritreme and lateral margins of pleura ochraceous. Metapleura strongly swollen, produced laterad, its posterior margin introverted, both well visible in dorsal view (Fig. 14). Legs ochraceous, hind femora darkened apically, tarsi brown. Hind femora and tibiae slightly curved inwards, with granulation on inner margins (Fig. 15).</p><p>Abdomen ochraceous with thick brown punctures; sterna 3-4 broadly, sternum 5 narrowly reddish brown medially, sternum 6 with sparse brown punctures, sternum 7 with sparse brown punctures and large blackish brown spots on each side near anterior margin and before postero-lateral angles; median carina well developed on sterna 3-6, gradually disappearing on sternum 7, ochraceous, acutely produced anteriad as abdominal spine between metacoxae (Fig. 16); connexiva ochraceous, spines on postero-lateral angles of sterna 3-4 very small, on sternum 5 small, on sternum 6 long, narrow, directed posteriad, and on sternum 7 produced, widely triangular with parallel outer margins (Figs. 14, 16).</p><p>Pygophore brownish; dorso-ventrally flattened (Fig. 17); hypophysis of paramere roughly semicircular, apically rounded, not pointed, inner margin deeply incised (Fig. 18); vesica of aedeagus deeply sinuated, without loop (Fig. 19).</p><p>Female. Unknown.</p><p>Differential diagnosis. Mahea distanti sp. nov. differs from both M. sexualis and M. durrelli sp. nov. by having the lateral pronotal angles conically produced and gradually tapering to a spinose apex. From M. parvula, it differs by the larger body, more prominent lateral pronotal angles, and relatively narrower head. From M. andriai, it can be separated by the following characters: Body more vividly coloured, covered with dark punctures; basic colour ochraceous, posterior part of pronotum with reddish tones; anterior part of scutellum reddish brown, antero-lateral angles and posterior part contrasting pale; apex of scutellum black, narrowly rounded. Dark punctures on head, sternum and abdominal venter coalescent, forming conspicuous rows (usually impressed and darkened). Hairs of antennomere 2 as long as or slightly longer than its diameter. Thorax conspicuously swollen. Lateral connexival spines as in Figs. 14 and 16.</p><p>Etymology. This species is named in honour of William Lucas Distant (1845-1922), wellknown English entomologist, who described many heteropteran taxa, including Mahea .</p><p>Bionomy. Unknown.</p><p>Distribution. East Madagascar (Toamasina). Only the holotype is known.</p><p>Comments. See comments under M. sexualis .</p></div>	https://treatment.plazi.org/id/039887C4A317F47D7CEB7D3D33AC52CD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A312F47E7CCC7E5D32AD500D.text	039887C4A312F47E7CCC7E5D32AD500D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mahea durrelli	<div><p>Mahea durrelli sp. nov.</p><p>(Figs. 20-27, 44)</p><p>Type locality. North-east Madagascar, Vohémar.</p><p>Type material. HOLOTYPE: ♁, ‘Vohémar / Madagascar [p] // Collectio / J.L.Stehlík / Moravské museum, Brno [p] // ♁ [p] // HOLOTYPUS / MAHEA / DURRELLI / sp. nov. / det. P. KMENT 2005’ [p, red label] (MMBC). The holotype is mounted with the pygophore glued on the same piece of card ; I have dissected the aedeagus and left paramere and placed them in a plastic microvial with glycerol attached to the same pin. PARATYPE: ♁, ‘Vohémar / Madagascar [p] // COLLECTIO / NATIONAL MUSEUM / Praha, Czech Republic [p] // ♁ [p] // PARATYPUS / MAHEA / DURRELLI / sp. nov. / det. P. KMENT 2005’ [p, red label] (NMPC). The paratype is mounted on the tip of a pentagonal piece of card ; left antennomere 2 broken, left antennomeres 3-4 missing, and right spine on sternum 6 broken; I have removed the pygophore and glued it on the same piece of card.</p><p>Note: The locality label ‘Vohémar’ is identical with those found in material collected in Madagascar by C. Lamberton in 1937 and housed in NMPC (see HOBERLANDT 1942).</p><p>Description. Body narrow, ochraceous with reddish-brown punctures, slightly shining.</p><p>Male (holotype) (Fig. 44). Length 6 mm, width of pronotum between humeral spines 3.1 mm.</p><p>Head ochraceous with irregular rows and groups of coarse black or brown punctures (two rows on mandibular plates less distinct than in other species, vertex with U-shaped figure). Head shorter than wide across eyes (1: 1.42), ratio of width of head: width of the pronotum between humeral angles equal to 1: 1.97. Mandibular plates only slightly narrowing basally, apically arcuately curved inwards, not meeting in front of clypeus; apex of clypeus free, clypeus slightly shorter than mandibular plates. Eyes ochraceous with brownish spots; ocelli reddish, situated near anterior pronotal margin. Antenniferous tubercles each with black longitudinal spot laterally; antennae castaneous, antennomeres 1-2 basally paler; antennomere 2 with erect pubescence, some hairs longer than its diameter; lengths of antennomeres: 1 – 0.3 mm, 2 – 1.7 mm, 3 – 1.0 mm, 4 – 0.9 mm (ratio 1: 5.67: 3.33: 3). Head ventrally ochraceous with sparse reddish punctures and one pair of larger, blackish L-shaped spots postero-medially next to bucculae; rostrum ochraceous, apex blackened, reaching middle of metacoxae.</p><p>Pronotum ochraceous with thick, coarse, uniformly distributed reddish-brown punctures; two rows of black punctures in area of cicatrices; humeral angles reddish. Anterior margin of pronotum only slightly concave, almost straight medially; antero-lateral margins rounded, proximally almost parallel, medially abruptly curved laterad; humeral angles prominent, conically produced laterad and upwards, apically constricted, with long narrow spine (Figs. 20- 21); posterior margin of pronotum medially more concave than in other species. Scutellum narrowly triangular, slightly sinuated medially, almost flat, only anterior part slightly sloping caudad; ochraceous, with reddish-brown punctures, colouration of neighbouring punctures locally coalescent in anterior part; apex narrowly triangular, with small black spot. Hemelytra narrow, suture brownish, membrane slightly brownish infuscate, not surpassing postero-lateral angles of sternum 7.</p><p>Thorax ventrally ochraceous with reddish-brown punctures; colouration of punctures especially on meso- and metapleuron coalescent, forming brown spots alternating with ochraceous unpunctured spots; acetabula, peritreme, and elevated unpunctured spots on pleura ochraceous. Metapleura not swollen, its margin only slightly visible from above. Legs ochraceous, tarsomeres 2 brownish. Hind femora and tibiae straight, lacking granulation on inner margins (Fig. 22).</p><p>Abdomen ventrally ochraceous with small and shallow red punctures, sterna 3-6 with three pairs of small black spots: one oval spot on anterior margin next to median carina, one round spot medially on each side, one small, round, impressed spot laterally on suture between sterna; sternum 7 with pair of oval blackish spots on anterior margin, postero-lateral angles of sternum 7 black. Median carina well developed on sterna 3-6, only basally on sternum 7, disappearing caudad (Fig. 23). Connexiva ochraceous, postero-lateral angles without spines except sternum 6 with long spine curved postero-laterad; postero-lateral angles of sternum 7 lanceolate, produced postero-laterad (Fig. 23).</p><p>Male genitalia. Pygophore brownish; less dorso-ventrally flattened (Fig. 24); exterior opening with small triangular tubercle dorso-medially (Figs. 24-25); hypophysis of paramere oval, apically broadly rounded, not pointed, inner margin only slightly incised (Fig. 26); vesica of aedeagus long, looped medially (Fig. 27).</p><p>Female. Unknown.</p><p>Variability. Paratype male (length 5.9 mm, width of pronotum between humeral spines 3 mm, length of antennomeres: 1 – 0.3 mm, 2 – 1.5 mm, 3 – 0.9 mm, 4 – 0.9 mm) differs from the holotype by having the mandibular plates meeting in front of the clypeus, punctures on the head darker, and the pronotum with a vague ochraceous median line.</p><p>Differential diagnosis. Mahea durrelli differes from M. andriai, M. distanti, and M. parvula in having each humeral angle abruptly produced into a long spine. From M. sexualis, it differs by having the lateral pronotal margins deeply sinuated (Fig. 20); the humeral spines raised (Figs. 20-21); the metapleura not swollen; the shape of connexival spines; and the apices of parameres rounded, not pointed (Figs. 24-26) (see also the key).</p><p>Etymology. The species is named in honour of Gerald Durrell (1925-1995), the famous English ‘amateur naturalist’, nature conservator and writer who paid a lot of attention to the nature of Madagascar and adjacent islands.</p><p>Bionomy. Unknown.</p><p>Distribution. North-east Madagascar. Known only from the type locality.</p></div>	https://treatment.plazi.org/id/039887C4A312F47E7CCC7E5D32AD500D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A311F4617CE57F1D329F5507.text	039887C4A311F4617CE57F1D329F5507.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mahea parvula	<div><p>Mahea parvula sp. nov.</p><p>(Figs. 28-30, 45)</p><p>Type locality. Madagascar.</p><p>Type material. HOLOTYPE: ♀, ‘4175 / 34 [hw; round label, upper side green without text, lower side with numbers 4175 (corresponding to the Accession book of MNHN) and 34 (= 1834)] // Plodyrensus Amyot – Ms. Paris. –. MADAGASCAR. [hw, mounted with sellotape] // ♀ [p] // HOLOTYPUS / Mahea / parvula / sp. nov. / des. P. Kment 2005’ [hw, red label] (MNHN). Holotype pinned; antennae, all legs except right middle one, left hemelytra, and two apical segments of rostrum missing, right eye partly damaged, basal part of scutellum and sternum with puncture from original pinning.</p><p>Description. Female (holotype) (Fig. 45). Body ochraceous with dark brown punctures. Length 7.3 mm, width of pronotum between humeral angles 3.9 mm.</p><p>Head ochraceous, 2-3 dark brown spots laterally in front of each eye; dark brown punctures coalescent (two impressed rows on each mandibular plate; two rows on vertex coalescent basally on clypeus and following anteriad as single median row; vertex with two rows forming U-shaped figure, additional punctures mesad from eyes and on hind margin of vertex). Head slightly shorter than wide (1: 1.15), width slightly less than one half of pronotal width. Mandibular plates only slightly narrowing basally, apically curved inwards, not meeting; apex of clypeus narrowly free. Eyes dark brown. Head ventrally ochraceous, dark brown spot on base of each buccula; two basal segments of rostrum ochraceous.</p><p>Pronotum ochraceous with dark brown punctures forming small irregular groups on anterior third of pronotum and almost regularly distributed on posterior two thirds of pronotum; humeral angles chocolate brown. Anterior and posterior margins slightly concave; anterolateral margins rounded, regularly concave, strongly widening posteriad; each humeral angle conically produced into obtuse spine directed laterad and slightly upwards (Figs. 28-29). Scutellum triangular, brownish with dark brown punctures, extreme apex black. Hemelytra of the same colour and puncturing as disc of scutellum; membrane hyaline, translucent, with about half its length surpassing apex of abdomen.</p><p>Thorax ventrally brownish with dark brown punctures; punctures laterally coalescent, forming irregular darker spots; pleuron along mesosternal carina dark brown, anterior half of carina ochraceous. Legs ochraceous.</p><p>Abdomen flat, median carina present only on sternum 3 (Fig. 30); sterna 2-3 brown, abdominal spine ochraceous; sterna 4-5 brownish with dark brown, rounded spots laterally, sternum 6 brownish with large dark brown lateral spots connected with those on following sternum; sternum 7 entirely dark brown with two small pale spots medially on its anterior margin and one larger pale spot medially on its posterior margin. Connexival spines as in Fig. 30.</p><p>Male. Unknown.</p><p>Differential diagnosis. Mahea parvula differs from both M. sexualis and M. durrelli by having the lateral pronotal angles conically produced, each gradually tapering to a spinose apex. From M. andriai and M. distanti, it differs in having a smaller body, less prominent humeral angles, relatively broader head, and the connexival spines as in Fig. 30 (see also the key).</p><p>Etymology. Adjective parvulus (Latin) = ‘little small’, minute. The female of M. parvula is distinctly smaller than females of M. andriai and M. sexualis .</p><p>Bionomy. Unknown.</p><p>Distribution. Madagascar (no exact locality). Only the holotype is known.</p></div>	https://treatment.plazi.org/id/039887C4A311F4617CE57F1D329F5507	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A30FF4637CFA796A306D524D.text	039887C4A30FF4637CFA796A306D524D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catadipson Breddin 1903	<div><p>Catadipson Breddin, 1903</p><p>Catadipson Breddin, 1903: 90 (description). Type species: Catadipson aper Breddin, 1903, by monotypy.</p><p>Catadipson: BREDDIN (1906): 198 -199 (taxonomy).</p><p>Catadipson: BERGROTH (1908): 192 (catalogue).</p><p>Catadipson: KIRKALDY (1909): 170 (catalogue).</p><p>Irsmia Cachan, 1952: 310 (description, figures). Type species: Irsmia imernensis Cachan, 1952, by original designation (syn. LESTON 1953).</p><p>Catadipson: LESTON (1953): 124 -125, Figs. 1-3, 5 (p. 125) (taxonomy, morphology).</p><p>Catadipson: KUMAR (1974): 43, Figs. 75-76 (p. 38), 77 (p. 50) (diagnosis, figures).</p><p>Distribution. Three described species are distributed in tropical Africa: Catadipson aper Breddin, 1903 from Equatorial Guinea (Fernando Póo Island) (BREDDIN 1903), Angola (LESTON 1953) and Ivory Coast (SCHOUTEDEN 1964a); C. sus Breddin, 1906 from Togo (BREDDIN 1906) and Ivory Coast (SCHOUTEDEN 1964a); and C. imernensis (Cachan, 1952) from Madagascar (CACHAN 1952).</p></div>	https://treatment.plazi.org/id/039887C4A30FF4637CFA796A306D524D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A30CF4637C9A7DDD32A35022.text	039887C4A30CF4637C9A7DDD32A35022.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catadipson imernensis (Cachan 1952)	<div><p>Catadipson imernensis (Cachan, 1952)</p><p>(Figs. 32, 46)</p><p>Irsmia imernensis Cachan, 1952: 310-312, Figs. 188 -192 (p. 311), 5 (Pl. XIV, p. 439) (description, figures, bionomy). Catadipson imernensis: LESTON (1953): 123 (new combination).</p><p>Type locality. Central Madagascar, Tananarive [= Antananarivo].</p><p>Type material. HOLOTYPE: ♀, ‘Tananarive / Tsimbazaza [p] // I. S. Madagascar [p] / Sur Ficus (figuàr) [hw] / Mai 1947 A. R. [hw] // Ficusia imernensis Cachan [hw] // TYPIS [p, pink label] // Irsmia / imernensis / n. sp. / Cachan det. [hw] // face side: HOLOTYPUS / Irsmia / imernensis / Cachan 1952, reverse: P. Kment 2005’ [hw, red label] (MNHN).</p><p>Bionomy. Collected from Ficus sp. ( Moraceae) (CACHAN 1952).</p><p>Distribution. Central Madagascar (Antananarivo env.) (CACHAN 1952).</p></div>	https://treatment.plazi.org/id/039887C4A30CF4637C9A7DDD32A35022	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A30CF4637CED7F02325F5761.text	039887C4A30CF4637CED7F02325F5761.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Noualhieridia Breddin 1898	<div><p>Noualhieridia Breddin, 1898</p><p>Noualhieridia Breddin, 1898: 266-267 (description). Type species: Noualhieridia ornatula Breddin, 1898, by monotypy.</p><p>Noualhieridia: BERGROTH (1908): 191 (catalogue).</p><p>Noualhieridia: KIRKALDY (1909): 178 (catalogue).</p><p>Noualhieridia: CACHAN (1952): 308 (redescription, figures).</p><p>Noualhieridia: KUMAR (1974): 23 -24 (diagnosis, figures).</p><p>Distribution. A genus endemic to Madagascar and including only three species – N. marginata Cachan, 1952, N. ornatula Breddin, 1898 and N. rufa Cachan, 1952 .</p></div>	https://treatment.plazi.org/id/039887C4A30CF4637CED7F02325F5761	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A30CF4637C9778C0329F56D8.text	039887C4A30CF4637C9778C0329F56D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Noualhieridia marginata Cachan 1952	<div><p>Noualhieridia marginata Cachan, 1952</p><p>(Fig. 47)</p><p>Noualhieridia marginata Cachan, 1952: 310, Fig. 187 (p. 311) (description, figure).</p><p>Type locality. Madagascar.</p><p>Type material. HOLOTYPE: ♀, ‘Muséum Paris / Madagascar / Coll. Sicard 1930 [p] // TYPE [p, red label] // Noualhieridia / marginata / n. sp. / Cachan det. [hw] // face side: HOLOTYPUS / Noualhieridia / marginata / Cachan 1952, reverse: P. Kment 2005’ [hw, red label] (MNHN).</p><p>Bionomy. Unknown.</p><p>Distribution. Madagascar (no exact locality). Only the holotype is known.</p></div>	https://treatment.plazi.org/id/039887C4A30CF4637C9778C0329F56D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A30DF4627CB47C9D32D55019.text	039887C4A30DF4627CB47C9D32D55019.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Noualhieridia ornatula Breddin 1898	<div><p>Noualhieridia ornatula Breddin, 1898</p><p>(Fig 48)</p><p>Noualhieridia ornatula Breddin, 1898: 267-268 (description).</p><p>Noualhieridia ornatula: BERGROTH (1908): 191 (catalogue).</p><p>Noualhieridia ornatula: KIRKALDY (1909): 178 (catalogue).</p><p>Noualhieridia ornatula: CACHAN (1952): 309, Figs. 5 (Pl. VII, p. 275), 183-186 (p. 311), (redescription, figures, distribution).</p><p>Noualhieridia ornatula: KUMAR (1974): 24, Figs. 33-34 (p. 22) (lectotype designation, figures).</p><p>Type locality. Madagascar.</p><p>Material examined. MADAGASCAR, coll. Noualhier 1898, 3 ♁♁ 5 ♀♀, Martin det. (part) or Cachan det. (part), P. Kment revid. (MNHN). Madagascar, 1 ♀, P. Kment det. (NMPC).</p><p>Bionomy. Unknown.</p><p>Distribution. Central Madagascar (Antananarivo) and West Madagascar (Maevatanana) (CACHAN 1952).</p><p>Note. Noualhieridia ornatula resembles (in body shape and colouration) another endemic Madagascar species, Cloequeria bourgini Cachan, 1952 ( Pentatomidae: Pentatominae: Coquerelini), which differs by having 3-segmented tarsi, a different shape of the peritreme, no conspicuous round black spot on the mesopleura, and by other details of colouration.</p></div>	https://treatment.plazi.org/id/039887C4A30DF4627CB47C9D32D55019	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A30DF4627C937F0932525670.text	039887C4A30DF4627C937F0932525670.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Noualhieridia rufa Cachan 1952	<div><p>Noualhieridia rufa Cachan, 1952</p><p>(Fig 33, 49)</p><p>Noualhieridia rufa Cachan, 1952: 309-310 (description).</p><p>Type locality. Madagascar.</p><p>Type material. LECTOTYPE: ♀, ‘Muséum Paris / Madagascar / Coll. Sicard 1930 [p] // TYPE [p, red label] // Noualhieridia / rufa / n. sp. / Cachan det. [hw] // LECTOTYPUS / Noualhieridia / rufa / Cachan 1952 / des. P. Kment 2005’ [hw, red label] (MNHN). PARALECTOTYPE: ♀, the same labels as lectotype (MNHN). Here designated. The lectotype is pinned; the paralectotype is pinned on a shortened pin which is put onto a piece of card and regularly pinned.</p><p>Other material examined. MADAGASCAR bor. or.: Maroansotra [= Maroantsetra] env., Ambodivoangy [= Ambodivoahangy], Institut Scientifique Madagascar, 1 ♀, P. Kment det. (MNHN). Sambava dct., R. N. XII – Marojejy, Ambatosoratra, 100 m, VIII-60, 1 ♀, P. Soga lgt., Institut Scientifique Madagascar, P. Kment det. (MNHN).</p><p>Bionomy. Unknown.</p><p>Distribution. North-east Madagascar (Maroantsetra env., Sambava env.).</p></div>	https://treatment.plazi.org/id/039887C4A30DF4627C937F0932525670	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
039887C4A30DF4657D89793A354A56E2.text	039887C4A30DF4657D89793A354A56E2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acanthosomatidae Signoret 1863	<div><p>Key to the Acanthosomatidae of Madagascar and the Seychelles</p><p>1. Mesosternal carina not developed. Antennae 5-segmented, antennomere 2 not exceptionally long. Metathoracic scent gland complex as in Fig. 33; mesopleura with lateral round black spot. Broadly oval bugs of pentatomid appearance. Noualhieridia Breddin, 1898 ........................................................................................................ 2</p><p>– Mesosternal carina well developed. Antennae 4-segmented, antennomere 2 very long. Metathoracic scent gland complex different (Figs. 31-32), mesopleura without lateral round black spot. Slender species, body more or less parallel. ......................................................................................................................................................... 4</p><p>2. Body dorsally (including connexiva), antennae and legs entirely terracotta orange to red. Antero-lateral margins of pronotum convex. ...................................................................................... Noualhieridia rufa Cachan, 1952</p><p>– Body dorsally, antennae and legs multicoloured. Antero-lateral margins of pronotum convex or concave with pale spot; base of corium with another pale spot laterally; postero-lateral angles of connexiva with black spots. ............................................................................................................................................................................. 3</p><p>3. Antero-lateral pronotal margins convex, with oval pale spot overlapping to pronotal disc; trochanters, apices of femora and entire tibiae, tarsi and antennae black. ............................... Noualhieridia ornatula Breddin, 1898</p><p>– Antero-lateral pronotal margins slightly concave, with long narrow pale spot not overlapping to pronotal disc; legs and antennae entirely pale, only apical halves of antennomeres 3-5 blackened. .......................................... ............................................................................................................ Noualhieridia marginata Cachan, 1952</p><p>4. Elongate oval species. Head very broad, slightly broader than two thirds of pronotal width.Apex of clypeus free, mandibular plates projected as conspicuous apical spines. Humeral angles of pronotum not produced, rounded. Metathoracic scent gland complex as in Fig. 32.Connexival margins without apparent spines.Head and pronotum with very large black punctures. ........... Catadipson Breddin, 1903: Catadipson imernensis (Cachan, 1952)</p><p>– Not conspicuously elongate, parallel species. Head only as broad as ca one half to one third of pronotal width). Mandibular plates without apical spines, meeting or not in front of clypeus. Humeral angles of pronotum each with conspicuous spine. Metathoracic scent gland complex as in Fig. 31. Connexival margins spinous. Head and pronotum with small dark punctures. Mahea Distant, 1908 ...................................................................... 5</p><p>5. Humeral angles gradually narrowed, each produced into conical spine, not constricted basally. ...................... 6</p><p>– Humeral angles abruptly narrowed, constricted, each with long needle-like spine. ........................................... 8</p><p>6. Smaller species (7.3 mm). Humeral angles of pronotum less prominent (Figs. 28-29). Head slightly narrower than half of pronotal width. Clypeus narrowly free apically. Lateral connexival spines as in Fig. 30. ................ ....................................................................................................................................... Mahea parvula sp. nov.</p><p>– Larger species (&gt; 8 mm). Humeral angles of pronotum more prominent (Figs. 8-9, 12-13). Head as wide as about one third of pronotal width. Clypeus completely enclosed apically. Lateral connexival spines different (Fig. 10). .............................................................................................................................................................. 7</p><p>7. Body almost uniformly yellowish ochraceous with small dark punctures (except darkened lateral corners of pronotum, apex of scutellum, antennae and paired black spots laterally on abdominal venter). Head with vague impressed rows of punctures. Apex of scutellum more broadly rounded. Sternum without black punctures. Hairs on antennomere 2 shorter than its diameter. Metapleura not swollen. Lateral connexival spines as in Fig. 10 .. ........................................................................................................................... Mahea andriai (Cachan, 1952)</p><p>– Body more colourful, covered with prominent black punctures. Basic colour ochraceous, posterior part of pronotum with reddish tones. Anterior part of scutellum reddish brown, antero-lateral angles and posterior part contrastly pale, apex of scutellum black. Dark punctures on head, pleura and abdominal venter coalescent, forming conspicuous rows (usually impressed and darkened). Apex of scutellum narrowly rounded. Sternum with black punctures. Hairs on antennomere 2 as long as or slightly longer than its diameter. Metapleura conspicuously swollen. Lateral connexival spines as in Fig. 16. ........................................................... Mahea distanti sp. nov.</p><p>8. Larger, more robust species (7.5-8 mm). Humeral angles not raised upwards (Figs. 2-3). Male: metapleura swollen; hind femora and tibiae curved inwards; abdomen ventrally with only one row of black spots mediolaterally on sterna 3-7; connexiva as in Fig. 4; postero-lateral angles of sternum 7 triangular, not curved laterad (Fig. 4). Female: connexiva as in Fig. 5. ............................................................. Mahea sexualis Distant, 1908</p><p>– Smaller, more slender species (5.9-6 mm). Humeral angles raised askew upwards (Figs. 20-21). Male: metapleura not swollen; hind femora and tibiae not curved inwards (Fig. 22); abdomen ventrally with three rows of black spots on each side of sterna 3-6; connexiva as in Fig. 23; postero-lateral angles of sternum 7 broadly lanceolate, distinctly curved laterad (Fig. 23). Female: unknown. ................................ Mahea durrelli sp. nov.</p></div>	https://treatment.plazi.org/id/039887C4A30DF4657D89793A354A56E2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Kment, Petr	Kment, Petr (2005): Revision of Mahea Distant, 1909, with a review of the Acanthosomatidae (Insecta: Heteroptera) of Madagascar and Seychelles. Acta Entomologica Musei Nationalis Pragae (suppl.) 45: 21-50, DOI: 10.5281/zenodo.4476396
