taxonID	type	description	language	source
0398D71B5248735A5FFAFA573DB2EFC4.taxon	materials_examined	Type genus — Heatherella walter, 1997.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B5248735A5FFAFA573DB2EFC4.taxon	diagnosis	Diagnosis: Gamasine Mesostigmata. Adults covered in a thick cerotegument; oval idiosoma ringed with bifurcate marginal (r - R) setae (up to 28 pairs) and gland-bearing protuberances (up to ten pairs), each with or without emergent, spout-like lip. Protuberances and s - S series setae on upper margin of extensive opisthogastricperitrematal-marginal shield; this shield with numerous spicules and / or microtubercles; dorsum with most of the usual setae for Mesostigmata except setae s 3 absent, setae z 2 present or absent, r - R series hypertrichous; setae j 1 and z 1 ventral; dorsal shields sometimes sexually dimorphic but always with series of pre-pygidial platelets and pygidial shield bearing setae J 5 flanked by small lateral shields bearing setae Z 4. Anal region raised, with five circumanal setae. Gnathosoma ventral, between first pair of legs, and protected dorsally by idiosoma. Chelicerae slender, elongate, with short digits, setiform pilus dentilis, without excrescences; cheliceral seta elongate, inserted laterally on fixed digit. Palpal setation, trochanter, femur, genu, tibia, respectively: 2 – 5 – 6 – 14. Leg I slender, antenniform, without apotele; leg setation: coxae I – IV, respectively: 2 – 2 – 2 – 1; trochanters 6 – 5 – 5 – 5; femora 13 – 11 – 7 – 6; genua: 13 – 11 – 10 – 10, formulas: 2 3 / 2, 3 / 1 2; 2 3 / 1, 2 / 1 2; 2 2 / 1, 2 / 1 2; 2 2 / 1, 3 / 1 1; tibiae 12 / 13 (± av 2) - 10 9 10; formulas: 2 3 / 2, 3 / 1 2; 2 2 / 1, 2 / 1 2; 2 1 / 1, 2 / 1 2; 2 2 / 1, 2 / 1 2. Female with dorsal shield undivided, fused with or free from submarginal and marginal shields anteriorly, bearing nine pairs of setae (j 3 – j 6, J 1 – J 4, z 5); lateral submarginal shields with four pairs of setae (z 6, Z 1 – 3); setae s 2 – s 6, S 1 – S 5 and Z 5 on marginal shield; sternal shield subdivided, st 1 on jugularia or in soft cuticle; st 2 on single plate; st 3 – 4 on 1 – 2 pairs of platelets; a single subtriangular genital shield with one pair of setae. Male genital opening intercoxal, covered by pair of sclerites; st 1 on jugularia or in soft cuticle; chelicera without spermatodactyl; femur II and sometimes genu II with seta av 1 a blunt spur.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B5248735A5FFAFA573DB2EFC4.taxon	discussion	Comments. The setation of the legs and palps is mostly consistent throughout the family including the new taxa described below. The only variable seta is av 2 on femur I, which is absent in the new genus. However, Walter (1997) missed a small dorsal seta on trochanter III: the correct setation for trochanters I-IV, respectively, is 6 – 5 – 5 – 5, not 6 – 5 – 4 – 5. The spout-like gland openings are characteristic of the Heatherellidae, but the number of well-developed protuberances varies interspecifically (7 – 10 spout-like protuberances; Table 1). The pores associated with setae s 6 are sometimes indistinguishable from other large idiosomal pores (Fig. 15); those associated with setae s 4 are sometimes enlarged pores and raised only slightly (Fig. 25); while those associated with setae s 5 often appear as a raised pustule (Figs 34 – 35). This latter form appears similar to that found in the Epicriidae. Alberti (2013) explored the fine structure of the single pair of idiosomatic protuberances in Epicrius mollis (Kramer, 1876) and pointed out the glandular nature of the pair he refers to as the pustule. Although they appear very similar in Heatherellidae and Epicriidae, the pustules of E. mollis may not be homologous with those in the Heatherellidae because they are unlikely to be derived from the same pore. The pustule of Epicriidae lies behind seta s 6 (Moraza 2005; Alberti 2013) but in the Heatherellidae, the pustule-like structure is anterior to seta s 6, and closely associated with seta s 5 (Figs 34 – 35). The classification of the Heatherellidae is problematic. Walter (1997) reasoned that the family belonged in the suborder Monogynaspida, and of its cohorts, regarded the family most closely related to the Epicriina, Microgyniina or Zerconina, but no synapomorphy could place the Heatherellidae in any of these cohorts. Lindquist et al. (2009 a) essentially followed Walter (1997) by classifying the Heatherellidae in its own monogynaspid cohort, the Heatherellina, but Lindquist et al. (2009 b) suggested a close relationship with the Epicrioidea. In regards to the Epicrioidea, and in contrast to the state of classification in 1997, Lindquist et al. (2009 a) treated the Epicriina and Zerconina as superfamilies of the Epicriiae, a subcohort of the cohort Gamasina. This classification is supported by molecular evidence, which placed the Epicriiae as the first branch within the Gamasina (Klompen et al. 2007). This molecular evidence also excluded the Microgyniina and Uropodina from the Gamasina. Thus, if the Heatherellidae share a sister-relationship with the Epicrioidea, then under the classification of Klompen et al. (2007) and Lindquist et al. (2009 b), the Heatherellidae are gamasine mites within the Epicriiae. Evidence for a Heatherellidae-Epicriidae sister relationship was provided by Lindquist et al. (2009 b), who noted the possible synapomorphies of a thick cerotegument (Figs 30 – 31), hypertrophied cheliceral seta, fragmented sternal shields and loss of the peritreme. Our new genus has a short but well-developed peritreme, thus eliminating this synapomorphy, although we note that the Epicriiae show considerable variation in peritreme development. Usually, nymphs have well-developed peritremes that are reduced, sometimes entirely, in adults (Solomon 1978; Lindquist & Moraza 1998; Moraza & Lindquist 1998). Heatherellidae also show this ontogenetic reduction. Deutonymphs of the new genus have long peritremes, which are greatly reduced in the adult, and some poorly preserved deutonymphs of H. acanthocharis and H. callimaulos have short but distinct peritremes; these are obsolete in the adult stage. Further putative synapomorphies with the Epicriidae are the laterally placed cheliceral seta, leg I without apotele and, usually in Epicriidae, a large, idiosomal pustule near seta s 5 (see Alberti 2013). The dorsal setation is also similar to some Zerconidae, with a trend towards the marginal placement of the z - Z series (except z 5) close to the s - S series, and hypertrichy in the r - R series. In three Heatherellidae species this hypertrichy is similar to some Zerconidae in that the r - R setae are duplicated (i. e. there are 22 pairs). Therefore, we propose that the Heatherellidae should be considered as a superfamily of the subcohort Epicriiae, and represent the only known southern hemisphere representatives of this subcohort.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524A735B5FFAFF6D3C27EA26.taxon	materials_examined	Type species — Heatherella acanthocharis walter, 1997, by original designation.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524A735B5FFAFF6D3C27EA26.taxon	diagnosis	Diagnosis: With the revised characteristics of the family (see above). Adult: sternal seta st 1 in soft cuticle; peritremes absent; ventral seta JV 3 small, simple, inserted on anal rim; opisthogaster lacking deep concavities flanking anus, and with nine pairs of setae (ZV 5 absent), setae JV 5 placed caudally, well posteromesad setae JV 4; marginal protuberances with their setae disassociated, usually arising anterior to each protuberance; tibia I with 13 setae (av 2 present). Dorsal shields sexually dimorphic: female with podonotal-mesonotal shield and latinotal shield, males with podonotal shield (anterior latinotal shield fused with podonotal shield) and three mesonotal shields (one medial, two lateral). Both sexes with podonotal shield fused anteriorly with marginal shield. Female: lateral shields entire, not divided into podonotal and mesonotal sections. Male: setae st 5 on medial extension of opisthogastric-peritrematal-marginal shield, separate from st 1 – 4.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524A73515FFAFD313C4AE94D.taxon	diagnosis	Diagnosis: With the characteristics of the genus. Adult: sternal shield (bearing st 2) subhexagonal, without endopodal elements, poorly differentiated anteriorly; ventral setae all short, bulbous and barbed; dorsal and marginal setae with ciliate crests; dorsal shield heavily ornamented; idiosomal margin with ten pairs of idiosomatic protuberances (protuberances P - s 4 and P - s 6 present), distinct pustule associated with seta s 5 absent, P - s 5 similar to other protuberances; dorsum with seta z 2; seta z 6 anterior to s 6, seta Z 1 anterior to S 1, seta z 5 placed obliquely to seta j 5; marginal r - R series setae not duplicated, 25 – 28 setae on each side, sometimes asymmetrical. Female: female with st 3 in soft cuticle and st 4 on small shields. Male: seta av 1 on genu II a blunt spur. Idiosomal dimensions: Females (n = 3) 360 – 375 long x 255 – 265 wide (holotype 360 x 265, small lateral split on one side); males (n = 9) 345 – 370 long x 220 – 255 wide.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524A73515FFAFD313C4AE94D.taxon	description	Female. Venter (Fig. 1). Tritosternum bifurcate; sternal region with a pair of small presternal platelets; st 1 in plicate cuticle, st 2 on subhexagonal, lineate-microtuberculate shield with pair of large pores in posterior corners; st 3 in soft cuticle; st 4 on small, subtriangular shields; all sternal setae short (st 1 4 – 7, st 2 9 – 10, st 3 9, st 4 8 – 9). Genital shield broadly subtriangular, flaring posteriorly, rounded anteriorly, irregularly reticulate with short (8 – 9), simple st 5; shield length 84 – 86, maximum width 90 – 98. Opisthogaster (Fig. 1) with seven pairs of short (7 – 14) setae (JV 1 – 2, JV 4, ZV 1 – 4), bulbous basally, tapering to fine point and ciliate on one side, lengthening posteriorly; two pairs of gland openings with raised rims posterolaterad coxae IV; one pair laterad para-anal setae; one pair laterad ZV 1; one pair anteriad ZV 4. Setae JV 5 caudal, posterior to final R seta and beneath most posterior spout. Anal region raised, with pair short (~ 5) simple setae on anterior corners anal opening (JV 3) and para-anal and postanal setae similar to ventral setae. Peritrematal shield obsolete, lateral region spiculate-microtuberculate, microtubercles becoming subreticulate across anterior ventral region and evanescent posteriorly; peritremes absent, stigma in raised tuberculate process (Fig. 36). Dorsum (Fig. 2). Dorsal shields comprising single median podonotal-mesonotal shield, latinotal shields, eight pre-pygidial platelets (often concealed), three pygidial shields (one medial, two lateral), all surrounded by dorsomarginal portion of opisthogastric-peritrematal-marginal shield; anteriorly, podonotal shield extending ventrally and fused with marginal shield. Median dorsal shield irregularly areolate, bearing eight pairs of setae (j 4 – 6, J 1 – 4, z 5; not including setae on vertex, i. e., ventrally placed j 1, z 1 and dorsally placed j 2 – 3, z 2, s 1); latinotal shields fused with podonotal shield anteriorly, each with four pairs of setae (z 6, Z 1 – 4); pre-pygidial platelets nude; median pygidial shield with two setae (J 5), lateral pygidial shield each with one seta (Z 4). All dorsal and marginal setae thickened, barbed, with ciliate crests, setal length 17 – 23. Seta z 6 anterior to s 6, seta Z 1 anterior to S 1, seta z 5 placed obliquely to seta j 5. Dorsal margin of shield bearing ten pairs of gland-bearing protuberances mostly aligned with s - S series setae (s 1, s 2, s 4 – 6, S 1 – 5) but also j 2, z 2 – 4; protuberance P - s 5 not pustule-like, similar in form to other protuberances (Table 1). Marginal setae (r - R series) hypertrichous, with 25 – 28 setae on each side (sometimes asymmetrical). Gnathosoma (Figs 3 – 7). Subcapitulum with strong row of denticles between palpcoxal setae; and three shorter, less distinct rows between setae h 3 and posterior to palpcoxal setae; all but most posterior row extend across deutosternum as fine denticles; deutosternal gutter narrow, with two additional rows of fine denticles anterior to palpcoxal setae; all subcapitular setae simple, acuminate, length h 1 11 – 12, h 2 9 – 10, h 3 11 – 12, pc 8 – 10 (h 3 expressed asymmetrically in holotype); corniculi simple. Chelicerae (~ 75 long), second article (55 – 65 long) with 3 subdistal teeth around pilus dentilis, deep notch receives fixed digit; moveable digit (25 – 28) tridentate; cheliceral seta (~ 12 long) thick, somewhat flattened, acuminate, inserted on paraxial side of chelicera. Tectum coarsely denticulate, with median prong (sometimes bifid) and 2 – 3 pairs of lateral teeth. Legs (Figs 8 – 11). Leg setae generally unmodified; dorsal femoral setae on leg I barbed (Fig. 8); femoral and genual setae subequal in length. Male. Similar to female except as follows. Venter (Fig. 12). Shield bearing st 2 – 4 and subcircular genital opening covered by a pair of platelets; st 5 on ventral, subtriangular extension of opisthogastric-peritrematalmarginal shield (and so eight pairs of ventral setae appearing to be present). Dorsum (Fig. 13). Dorsally with podonotal and mesonotal shields; podonotal shield fused with lateral shields, bearing five pairs of setae (j 4 – 6, z 5 – 6) excluding setae at vertex (j 1 – 3, z 1 – 3, s 1); mesonotal shield mostly free from lateral mesonotal shields (fused anteriorly), mesonotal shield bearing four pairs of setae (J 1 – 4), lateral mesonotal shields bearing three pairs of setae (Z 1 – 3); eight pre-pygidial platelets and pygidial shields as in female; protuberances and s - S and r - R setae as in female. Gnathosoma (Figs 5, 7). Similar to female; teeth on gnathotectum less developed. Legs (Fig. 14). Leg II with setae av 1 on femur and genu a blunt spur (8 – 9 x 5 wide, 5 – 6 x 2 – 3 wide, respectively).	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524A73515FFAFD313C4AE94D.taxon	materials_examined	Material examined. Holotype female, 2 female, 9 male paratypes, Australia, New South Wales, Lansdowne State Forest, 31 ° 45 ′ S 152 ° 37 ′ E, Graham Osler, 22 February 1996, Tullgren extraction of top 5 cm of soil in mixed hardwood forest with Eucalyptus pilularis & E. propinqua. Holotype, 1 female and 7 male paratypes deposited in QM; 1 female (ANIC 51 - 006365), 1 male paratype deposited in ANIC (ANIC 51 - 006366); 1 male paratype deposited in each of CNC and NZAC (NZAC 02016473).	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524A73515FFAFD313C4AE94D.taxon	etymology	Etymology: The new species is named in honour of the young acarologist who collected these specimens many years ago, Graham Osler.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524A73515FFAFD313C4AE94D.taxon	discussion	Comments. This species is found in wet sclerophyll forests; more details on the mite’s ecology may be found in Osler & Beattie (2001). This species is obviously different from the two species from Queensland in having the dorsal setae with ciliate crests (simple setae in other Heatherella), seta z 2 present (absent in other Heatherellidae), and ten pairs of lateral protuberances (8 – 9 pairs in other Heatherella including P - s 5; see Table 1). Heatherella osleri seems more closely related to H. callimaulos in sharing the position of the dorsal setae (i. e. seta z 6 anterior to s 6, seta Z 1 anterior to S 1, seta z 5 placed obliquely to seta j 5), having marginal protuberance P - s 4 present and having males with seta av 1 on genu II modified into a spur-like process. The presence of ten pairs of protuberances, the maximum number known so far in Heatherellidae, suggests a trend towards an increase in protuberance number. The presence of protuberance P - s 4 may be a synapomorphy, but protuberance P - s 5 is an autapomorphy, and we interpret protuberance P - s 6 as a homoplasy as it is present only in H. osleri and the new genus described below (Table 1).	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524073515FFAFE953D14EB90.taxon	description	Heatherella acanthocharis walter, 1997: 168.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524073515FFAFE953D14EB90.taxon	diagnosis	Diagnosis: With the characteristics of the genus. Adult: sternal shield (bearing st 2) rounded anteriorly, without endopodal elements; most ventral setae short, smooth, including setae ZV 2 – 3; setae JV 4, ZV 4 thickened, barbed; dorsal setae thickened, barbed, without ciliate crests; marginal setae bifurcate but crest small, not highly ciliate; dorsal shield heavily ornamented; idiosomal margin with seven pairs of idiosomatic protuberances (protuberances P - s 4 and P - s 6 absent), distinct pustule P - s 5 present; dorsum lacking seta z 2; seta z 6 posterior to s 6, seta Z 1 posterior to S 1; seta z 5 placed obliquely to seta j 5; marginal r - R series setae duplicated, 22 pairs. Female: setae st 3 and st 4 on single shield. Male: seta av 1 on genu II unmodified.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524073515FFAFE953D14EB90.taxon	materials_examined	New material examined: 1 female, 1 deutonymph, Queensland, Conondale Ranges, Lobster Creek, 26 ° 40 ′ S 152 ° 39 ′ E, 14 July 1998, D. E. Walter & H. Klompen, ex riparian rainforest leaf litter.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524073515FFAFE953D14EB90.taxon	discussion	Comments. Our diagnosis above is based on Walter (1997) and the re-examination of specimens held in the Queensland Museum, including the holotype. The new specimens examined are in poor condition but include the only known nymph of H. acanthocharis.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524073535FFAFBEE3CD7E9D9.taxon	description	Heatherella callimaulos walter, 1997: 173.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524073535FFAFBEE3CD7E9D9.taxon	diagnosis	Diagnosis: With the characteristics of the genus. Adult: sternal shield (bearing st 2) with angular anterior corners in both sexes, endopodal elements fused to shield; ventral setae JV 1 – 2, ZV 1 short, smooth, setae ZV 2 – 4, JV 4 thickened, barbed; dorsal setae thickened, barbed, without ciliate crests; marginal setae bifurcate but crest small, not highly ciliate; dorsal shield moderately ornamented, less so than other species; idiosomal margin with nine pairs of idiosomatic protuberances (protuberances P - s 4 present, P - s 6 absent), distinct pustule P - s 5 present; dorsum lacking seta z 2; seta z 6 anterior to s 6, seta Z 1 anterior to S 1; seta z 5 placed almost transversely with seta j 5; marginal r - R series setae duplicated, 22 pairs. Female: setae st 3 and st 4 on separate shields. Male: seta av 1 on genu II a small blunt spur.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524073535FFAFBEE3CD7E9D9.taxon	materials_examined	New material examined: Lamington National Park, Queensland, Australia, total of 9 females, 8 males, 3 deutonymphs, 1 protonymph, as follows. 1 male, Sarabah, Mount Cainbable, 28 ° 08 ′ S 153 ° 07 ′ E, 28 July 2002, dry rainforest transit S of cabins, Brush-box ecotone, soil & litter # 2, F. Beaulieu; 1 female, 2 males, same data except sclerophyll-rainforest transit; 1 female, same data except rainforest soil & litter # 3. 1 female, Caves Circuit, 28 ° 12 ′ S 153 ° 11 ′ E, 24 October 2002, rainforest soil # 4, F. Beaulieu; 1 female, 1 male, Border Track (1 km from Albert Track junction), 28 ° 15 ′ S, 153 ° 09 ′ E, 8 January 2002, rainforest soil & litter # 4, F. Beaulieu; 1 male, Border Track (1.5 km from Binna Burra), 28 ° 12 ′ S 153 ° 11 ′ E, 11 June 2002, litter # 7, F. Beaulieu; 2 females, same data except 14 April 2002, litter # 6; 1 female, Border Track (1.4 km from O’Reilly), 28 ° 14 ′ S 153 ° 09 ′ E, 17 May 2002, Asplenium nidus litter # 10, F. Beaulieu; 1 female, Mike O’Reilly’s farm, 28 ° 14 ′ S 153 ° 07 ′ E, 27 July 2002, wet sclerophyll-rainforest transition soil & litter # 5, F. Beaulieu; 1 male, Duck Creek Road, 28 ° 13 ′ S, 153 ° 09 ′ E, 1 June 2002, rainforest soil & litter # 3, F. Beaulieu; 1 female, same data except soil & litter # 1; 1 male, same data except ecotone soil & litter # 5; 1 male, Dave’s Creek Circuit, 28 ° 13 ′ S 153 ° 12 ′ E, 15 July 2002, rainforest soil & litter # 6, F. Beaulieu; 1 deutonymph, 1 protonymph, Park Monument, 28 ° 11 ′ S 153 ° 07 ′ E, 15 November 1996, ex rainforest litter above creek, D. E. Walter (slide in poor condition); 2 deutonymphs, Green Mts, 28 ° 13 ′ S 153 ° 07 ′ E, 8 June 1998, ex litter from open forest, D. E. Walter (slide in poor condition). Gambubal Forest Station, Queensland, total of 6 females and 8 males, as follows. 2 females, 3 males, 27 ° 08 ′ S 152 ° 09 ′ E, 20 January 2003, rainforest soil & litter # 1, F. Beaulieu; 2 females, 1 male, same data except sclerophyll soil & litter # 4; 1 female, 3 males, same data except sclerophyll soil & litter # 5; 1 female, same data except sclerophyll soil & litter # 2; 1 male, same data except rainforest soil & litter # 6.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524073535FFAFBEE3CD7E9D9.taxon	discussion	Comments. The above collection records were vouchers from Beaulieu et al. (2010); we also examined the specimens deposited in the Queensland Museum (including those from the University of Queensland Insect Collection), which includes the holotype, and the female reported by Seeman (2007). The original description did not illustrate the dorsum, which is provided here (Fig. 15) to provide diagnostic information on the position of setae, form of the protuberance P - s 5, presence / absence of protuberances P - s 4 and P - s 6, and amount of ornamentation. Two forms of pores are also apparent in H. callimaulos, a larger, more obvious type, of which there are 12 pairs, and a smaller type of which there are ten pairs (Fig. 15).	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524273535FFAFE213E03EBCF.taxon	materials_examined	Type species — Aheatherella mira n. sp.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524273535FFAFE213E03EBCF.taxon	diagnosis	Diagnosis: With the revised characteristics of the family (see above). Adult: sternal seta st 1 on sclerotised jugularia; peritremes present, short but well-developed; ventral seta JV 3 well-developed, not inserted on anal rim; opisthogaster with deep concavities flanking anus and with ten pairs of setae (ZV 5 present), setae JV 5 placed ventrally, directly behind JV 4; marginal protuberances with their setae closely associated, arising on dorsal surface of each protuberance; tibia I with 12 setae (av 2 absent). Dorsal shields monomorphic: both sexes with entire podonotal-mesonotal shield; lateral shields divided into podonotal and mesonotal sections; dorsal and lateral shields free anteriorly from marginal shield. Male: setae st 5 on sternogenital shield.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B524273535FFAFE213E03EBCF.taxon	etymology	Etymology: The generic name is derived from the type genus of the family and the prefix ‘ a’, Greek for ‘ not’ or ‘ away from’.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B5242734C5FFAFC0A3ACEEA31.taxon	diagnosis	Diagnosis: With the characteristics of the genus. Adult: sternal shield (bearing st 2) subrectangular, poorly differentiated anteriorly; most ventral setae short, weakly barbed, setae ZV 4 – 5 thicker, more barbed, setae JV 5 bifurcate, similar to marginal setae; dorsal setae simple, smooth; marginal setae bifurcate, each branch thickened, barbed; dorsal shield areolate; idiosomal margin with ten pairs of idiosomatic protuberances (protuberances P - s 4 and P - s 6 present), distinct pustule P - s 5 present; dorsum lacking seta z 2; seta z 6 anterior to s 6, seta Z 1 anterior to S 1; seta z 5 placed transversely with seta j 5; marginal r - R series setae duplicated, 22 pairs. Female: setae st 3 and st 4 on separate shields. Male: seta av 1 on genu II unmodified. Idiosomal dimensions: Females (n = 2) ca. 540 long x 430 wide (holotype split laterally; other female fragmented); males (n = 5) 460 – 500 long x 340 – 370 wide; deutonymph (n = 3) 410 – 480 long x 320 – 330 wide.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B5242734C5FFAFC0A3ACEEA31.taxon	description	Female. Venter (Fig. 16). Intercoxal region with bifurcate tritosternum; sternal shield fragmented into pair of jugularia bearing st 1, tuberculate area of soft cuticle bearing stp 1, subrectangular shield bearing st 2 + stp 2, subtriangular shield bearing st 3 + stp 3, and narrow metasternal shields bearing st 4 flanking genital shield; genital shield subrectangular, reticulate, bearing s t 5, flanked by stp 5 at posterior corners in soft cuticle; sternal setae simple, tapering except st 5 with basal barb; lengths: st 1 10, st 2 11, st 3 13, st 4 14, st 5 17. Opisthogaster with ten pairs of setae (JV 1 – 5, ZV 1 – 5) and 10 – 11 cuticular gland openings (pairing and placement erratic); pores laterad coxa IV in densely spiculate depression; anal region raised, flanked by large concave depressions, and with two pairs of para-anal setae, postanal seta, and pair of glands on anterior margin of anal opening; setae JV 3 not on anal rim; setae JV 5 ventral, posterior to setae JV 4, positioned close to small tubercle; all ventral setae slightly bulbous basally, tapering to fine point and smooth (anteromedially) to weakly to strongly barbed (laterally) on one side. Lateral peritrematal region and anterior margin of ventral region covered in dense vestiture of hair-like spicules, single, bifurcate or with several divisions; cuticle between exopodal and peritrematal margins smooth; exopodal and peritrematal shields fused but lines of fusion apparent (Figs 16, 37); peritremes short (~ 50 long), restricted to region of coxa III. Dorsum (Fig. 17). Dorsal shields irregularly areolate, setae tapering, acuminate, weakly barbed dorsally. Podonotal-mesonotal shield bearing nine pairs of setae (j 3 – j 6, J 1 – J 4, z 5; seta z 2 absent); anterior lateral shields each with two setae (z 4, z 6); posterior lateral shields each with three setae (Z 1 – Z 3); eight nude pre-pygidial platelets; median pygidial shield with two setae (J 5); lateral pygidial shields each with one seta (Z 5). Seta z 6 anterior to s 6, seta Z 1 anterior to S 1, seta z 5 placed transversely with seta j 5. Dorso-marginal part of opisthogastricperitrematal-marginal shield bearing nine pairs of large, spout-like gland-bearing protuberances; each protuberance with closely associated ciliate seta (j 2, s 1 – 2, s 5 – 6, S 1 – 2, S 4 – 5); protuberance P - s 5 different from other protuberances, being a raised pustule with two small openings; seta s 4 on small tubercle and associated gland on tiny protuberance. Marginal (r - R) setae of opisthogastric-peritrematal-marginal shield hypertrichous, formed by duplication (i. e. 22 pairs); all r - R setae strongly bifurcate, ciliate. Gnathosoma (Figs 18 – 19). Subcapitulum with four pairs smooth, scale-like lobes; basal row of denticles, two irregular rows more distally; deutosternal gutter narrow with single file of minute teeth; labial lobe rounded distally with lateral tooth; internal malae simple lobes; palpcoxal and hypostomal setae simple, acuminate; lengths: h 1 15, h 2> 10 (broken), h 3 20, pc 16; corniculi simple, blunt. Chelicerae (Fig. 19) slender (~ 220 long), second article (~ 125 long) ending in fixed digit with cluster of three small teeth, small setiform pilus dentilis, pocket for receiving tooth on movable digit; moveable digit (~ 34) with single distal tooth; cheliceral seta (~ 22 long) thick, somewhat flattened, acuminate, inserted on paraxial side of chelicera. Legs (Figs. 20 – 23). Leg setae unmodified; dorsal femoral setae on leg I smooth, some leg setae (e. g. ad setae on femora I-II, pd setae on femora III-IV) much shorter than other setae on same segment. Male. Similar to female except in intercoxal region and leg II. Venter (Fig. 24). Sternal shields comprised of jugularia bearing st 1, subtrapezoidal shield bearing st 2 and large pore, basal shield with subreticulate lines of denticles and bearing st 3 – 5 and nude genital platelets; all setae simple, acuminate. Dorsum (Fig. 25). Similar to female. Legs (Fig. 26). Leg II with setae av 1 on femur a blunt spur (13 – 14 x 4 – 5 wide); seta av 1 on genu II thickened, but not spur-like. Deutonymph. Venter (Figs 27 – 28). Intercoxal region with five pairs of setae; setae st 1 on smooth soft cuticle; setae st 2 on small subrectangular sternal shield; setae st 3 in striate cuticle; setae st 4 among irregular striate-rugose cuticle; setae st 5 in striate cuticle; medial round pore-like structure between coxae IV. Opisthogaster striate with numerous spicules medially; setae barbed and mostly shorter than in adult; ventrianal shield and anus projecting, with deep folds laterally; ventrianal shield capturing setae JV 3; ciliate cavities posterolaterad para-anal setae; marginal cuticle heavily rugose. Peritremes broad, longer than in adult, reaching base of coxae I. Dorsum (Fig. 29). Single shield covering idiosoma, densely ornamented with roseate fovaeoles but smooth laterally. Dorsal setation consistent with adult; median row of 1 one pairs of setae (j 1 – 6, J 1 – 5), setae z 2, z 4, z 5 – 6, s 1 – 2, s 4 – 6, S 1 – 5, and hypertrichous r - R series formed by duplication (i. e. 22 pairs); marginal tubercles similar to adult except pustule-like protuberance associated with seta s 5 not as raised and seta clearly disassociated from protuberance; vestibules of glands smaller than in adult. Gnathosoma and legs. Similar to adult female.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B5242734C5FFAFC0A3ACEEA31.taxon	materials_examined	Material examined: 2 females, 5 males, 3 deutonymphs, as follows. New Zealand, North Island, Manawatu- Whanganui: Holotype female (NZAC 02016468): Whanganui National Park, Whanganui River Road, Atene Pa (39 ° 43 ' 36 " S, 175 ° 08 ' 20 " E, NZTopo 50 BK 33 833 004), 90 m a. s. l., secondary broadleaf native forest (regenerating Beilschmiedia tawa, Knightia excelsa; understory Cyathea dealbata, Brachyglottis repanda, Olearia rani), ex leaf litter, Site 2, 29 January 2004, M. Minor. 1 paratype female (QMS 108782), 2 paratype males (NZAC 02016469; QMS 108783), 3 paratype deutonymphs (NZAC 02016470 – 71; QMS 108784), Whanganui National Park, Whanganui River Road, road marker 191 (39 ° 38 ' 07 " S, 175 ° 10 ' 23 " E, NZTopo 50 map reference BK 33 865 105), 104 m a. s. l. (above sea level), secondary broadleaf native forest (regenerating B. tawa; understory Melicytus ramiflorus, Schefflera digitata, Hedycarya arborea, Macropiper excelsum; abundant liana Ripogonum scandens), ex leaf litter, 29 January 2004, M. Minor, Site 1. 3 male paratypes (NZAC 02016472 (2 on same slide); QMS 108785): Whanganui District Council Block WN 31 B / 280, Kauarapaoa Road (39 ° 41 ' 29 " S, 175 ° 02 ' 20 " E, NZTopo 50 map reference BK 32 784 045), 159 m a. s. l., mixed black beech / broadleaf forest (canopy Nothofagus solandri with B. tawa, K. excelsa, Prumnopitys taxifolia, Dacydium cupressinum, Nestegis cunninghamii; understory C. dealbata, Myrsine australis, O. rani), ex leaf litter, Site 12, 31 January 2004, M. Minor. Type deposition: Female holotype, 3 males, 2 deutonymphs in NZAC; 1 female, 2 males, 1 deutonymph in QM.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
0398D71B5242734C5FFAFC0A3ACEEA31.taxon	etymology	Etymology: The specific name mira means “ wonderful ”, “ extremely impressive ” in Latin; gender feminine.	en	Seeman, Owen D., Minor, Maria, Baker, Michelle R., Walter, David Evans (2018): A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia. Zootaxa 4434 (3): 441-465, DOI: 10.11646/zootaxa.4434.3.3
