taxonID	type	description	language	source
0398EF3DE80BF2695F4BF8A63862FACA.taxon	diagnosis	Diagnosis. “ Medium-sized ” Japanese Cybaeus. Females of C. daimonji sp. nov. resemble those of C. communis Yaginuma, 1972, C. kirigaminensis Komatsu, 1963, C. maculosus Yaginuma, 1972 and C. shinkaii (Komatsu, 1970), which also are “ medium-sized ” and have a similar posteromedially located atrium and inverted V-shaped spermathecae [figs 2 - 2 - 30 - 18 – 23 in Ihara (2009 a)]. However, females of C. daimonji sp. nov. can be discriminated from those of the other four species by the copulatory ducts running toward the medial part of respective spermathecae, while in the other four congeners the copulatory ducts run directly toward the respective spermathecal heads [for C. communis, based on an unpublished observation by Yoh Ihara; for C. kirigaminensis, pl. 4, fig. H in Komatsu (1963); for C. maculosus, fig. 38 in Yaginuma (1972); and for C. shinkaii, fig. 4 in Komatsu (1970)]. Males of C. daimonji sp. nov. possess a palpal tibia, which is shorter than the palpal patella, and thus can be clearly discriminated from those of C. communis [fig. 37 in Yaginuma (1972)], C. kirigaminensis [pl. 4, fig. D in Komatsu (1963)] and C. shinkaii [fig. 2 - 2 - 30 - 17 in Ihara (2009 a)] bearing a tibia that is longer than the patella [see also figs 2 - 2 - 30 - 14 – 16 in Ihara (2009 a)].	en	Matsuda, Kenji, Ihara, Yoh, Nakano, Takafumi (2020): Description of a New Species of Cybaeus (Araneae: Cybaeidae) from Central Honshu, Japan. Species Diversity 25: 145-152, DOI: 10.12782/specdiv.25.145
0398EF3DE80BF2695F4BF8A63862FACA.taxon	materials_examined	Material examined. Holotype: KUZ Z 2753 (Fig. 1 A), male, under rotten wood in Mt. Daimonjiyama, Sakyoku, Kyoto City, Kyoto Prefecture, Japan (35.027462 ° N, 135.801530 ° E), Kenji Matsuda (KM), 1 November 2019. Paratypes (in total 13 specimens collected from the type locality by KM): 6 males, KUZ Z 2754 (35.027378 ° N, 135.801555 ° E), KUZ Z 2755 (35.027462 ° N, 135.801530 ° E), KUZ Z 2756 – Z 2757 (35.027280 ° N, 135.801641 ° E), KUZ Z 2758 (35.026685 ° N, 135.801831 ° E), KUZ Z 2759 (35.024971 ° N, 135.802875 ° E), and 4 females, KUZ Z 2761 (35.027302 ° N, 135.801637 ° E), KUZ Z 2762 (35.027142 ° N, 135.801608 ° E), KUZ Z 2763 (35.025709 ° N, 135.802005 ° E), KUZ Z 2764 (35.025709 ° N, 135.802005 ° E), 1 November 2019; 3 females, KUZ Z 2760 (35.026154 ° N, 135.80301 ° E), KUZ Z 2765 (35.026370 ° N, 135.802903 ° E), KUZ Z 2766 (collected near from the location of KUZ Z 2765), 26 November 2019 (KUZ Z 2754 – Z 2759, Z 2761 – Z 2766 were dissected). Additional materials (in total 28 specimens were collect- ed from 3 locations in Kyoto, by KM): 2 males, KUZ Z 2767 (35.035561 ºN, 135.798191 ºE), KUZ Z 2768 (35.035885 ºN, 135.798609 ºE), and female, KUZ Z 2783 (35.035507 ºN, 135.798138 ºE), Mt. Uryuyama, 24 October 2019; 6 males, KUZ Z 2769 (35.036335 ºN, 135.797853 ºE), KUZ Z 2770 (35.036796 ºN, 135.797790 ºE), KUZ Z 2771 (35.036862 ºN, 135.797984 ºE), KUZ Z 2772 – Z 2773 (35.036900 ºN, 135.797945 ºE), KUZ Z 2774 (35.037242 ºN, 135.798057 ºE), and 5 females, KUZ Z 2784 – Z 2785 (35.036597 ºN, 135.797874 ºE), KUZ Z 2786 (35.036547 ºN, 135.797793 ºE), KUZ Z 2787 (35.037240 ºN, 135.798016 ºE), KUZ Z 2788 (35.037191 ºN, 135.798158 ºE), Mt. Uryuyama, 30 October 2019; 7 males, KUZ Z 2775 (35.042871 ºN, 135.797571 ºE), KUZ Z 2776 (35.042327 ºN, 135.798311 ºE), KUZ Z 2777 (35.041602 ºN, 135.799438 ºE), KUZ Z 2778 (35.041573 ºN, 135.799448 ºE), KUZ Z 2779 (35.040904 ºN, 135.799914 ºE), KUZ Z 2780 (35.040474 ºN, 135.800204 ºE), KUZ Z 2781 (35.040553 ºN, 135.800283 ºE), and 6 females, KUZ Z 2789 (35.042341 ºN, 135.798351 ºE), KUZ Z 2790 (35.041719 ºN, 135.799351 ºE), KUZ Z 2791 (35.041602 ºN, 135.799438 ºE), KUZ Z 2792 (35.041638 ºN, 135.799287 ºE), KUZ Z 2793 (35.040974 ºN, 135.800063 ºE), KUZ Z 2794 (35.040912 ºN, 135.799824 ºE), Mt. Uryuyama, 8 November 2019; male, KUZ Z 2782, Mt. Hieizan (35.055603 ºN, 135.814328 ºE), 25 November 2019 (KUZ Z 2767 – Z 2771, Z 2773 – Z 2789, Z 2791 – Z 2794 were dissected).	en	Matsuda, Kenji, Ihara, Yoh, Nakano, Takafumi (2020): Description of a New Species of Cybaeus (Araneae: Cybaeidae) from Central Honshu, Japan. Species Diversity 25: 145-152, DOI: 10.12782/specdiv.25.145
0398EF3DE80BF2695F4BF8A63862FACA.taxon	description	Description. Males. Measurements [KUZ Z 2753 (holotype)]. Body length 6.27; carapace 3.04 long, 2.21 wide, head 1.26 wide; abdomen 2.98 long, 2.46 wide; sternum 1.52 long, 1.38 wide; labium 0.45 long, 0.45 wide. Leg formula, IV> I> II> III; length of legs (femur + patella + tibia + metatarsus + tarsus): leg I 10.24 (2.56 + 0.88 + 2.54 + 2.45 + 1.81); leg II 9.29 (2.25 + 0.88 + 2.29 + 2.29 + 1.58); leg III 8.45 (2.24 + 0.80 + 1.92 + 2.16 + 1.33); leg IV 11.11 (2.78 + 0.90 + 2.66 + 3.14 + 1.63). Carapace (Fig. 2 A): head narrow, 0.57 times as wide as thoracic region; thoracic region as high as head. Anterior median eyes smallest, approximately one-half diameter of other eyes; anterior eye row straight in frontal view; posterior eye row almost straight in dorsal view; ocular area twice as wide as long. Clypeus length 0.62 times length of median ocular area in holotype. Chelicerae geniculate, promargin of fang furrow with 3 teeth, retromargin with 4 teeth and 5 denticles (= small teeth), and basal with lateral condyle. Abdomen (Fig. 2 B) globular. Leg spination: Leg I: tibia p 3, r 3 (left) or 2 (right), v 2 - 2 - 2 - 2; metatarsus p 4 (left) or 3 (right), r 3 (left) or 2 (right), v 2 - 2 - 3. Leg II: tibia p 3, r 3 (left) or 2 (right), v 2 - 2 - 1 - 2; metatarsus p 4, r 4 (left) or 3 (right), v 2 - 2 - 3. Palp [KUZ Z 2755 (paratype)] relatively slender (Fig. 3 A). Patellar apophysis prominent on retrolateral anterior margin of patella, extended distally, slightly arched dorsally, triangular in dorsal view, distodorsal surface with 6 – 7 (left) or 6 – 8 (right) peg setae (Fig. 3 B). Tibia slightly shorter than patella, retrolateral tibial apophysis (RTA), plate-like, occupying most of length of tibia, distal margin slightly extended (Fig. 3 B). Cymbium slender, prolaterally unexpanded (Fig. 3 C), 2.7 times longer than wide; 0.9 times as long as femur, 1.3 times as long as patella + tibia. Bulb elliptic (Fig. 4 A); embolus simple, curved, originated and terminated, respectively, at ca. 11 o’clock and ca. 4 o’clock positions in ventral view (Fig. 4 A, B); conductor simple, triangular, small (Fig. 4 B, C). Females. Measurements [KUZ Z 2760 (paratype)]. Body length 6.02; carapace 3.14 long, 2.19 wide, head 1.47 wide; abdomen 3.28 long, 2.51 wide; sternum 1.47 long, 1.34 wide; labium 0.42 long, 0.48 wide. Legs shorter than those of male; leg formula, IV> I> II> III; length of legs (femur + patella + tibia + metatarsus + tarsus): leg I 7.96 (2.29 + 0.72 + 2.14 + 1.74 + 1.07); leg II 7.57 (2.29 + 0.69 + 1.82 + 1.65 + 1.12); leg III 6.69 (1.95 + 0.69 + 1.50 + 1.62 + 0.93); leg IV 8.51 (2.13 + 0.80 + 2.11 + 2.22 + 1.25). Carapace (Fig. 2 C) longer than that of male: head 0.67 times as wide as thoracic region, slightly wider than that of male. Abdomen (Fig. 2 D) oval, slightly larger than that of male. Leg spination. Leg I: tibia p 2, v 2 - 2 - 2 - 2; metatarsus p 1, r 2 (left) or 1 (right), v 2 - 2 - 2. Leg II: tibia p 3 (left) or 4 (right), v 2 - 2 - 1 - 2; metatarsus p 3, r 2 (left) or 1 (right), v 2 - 2 - 3. Genitalia [KUZ Z 2764 (paratype)]. Atrium slightly concave, located posteromedially on epigyne, anterior margin slightly curved, with two distinct copulatory openings (Fig. 5 A). Copulatory ducts, running anterolaterally toward medial part of respective spermathecae (Fig. 5 A, C). Paired spermathecae inverted V-shaped in ventral view; each spermatheca consisting of 3 well-defined parts, head, stalk and base (Fig. 5 B, C): head globular, located anteromedially above epigynum, with 2 simple pores ventromedially, heads well separated from each other; stalk cylindrical, loosely coiled, with developed Bennett’s gland at junction with base; base larger than head or stalk, globular, located posterolaterally above epigynum. Pair of fertilization ducts narrow, running posteromedially (Fig. 5 B, C). Variation. Males. Measurements (mean ± 1 SD, followed by ranges in parentheses; n = 23, including holotype;): body length 5.26 ± 0.42 (4.40 – 6.27); carapace length 2.91 ± 0.15 (2.48 – 3.14), width 2.14 ± 0.13 (1.81 – 2.35), head width 1.26 ± 0.08 (0.99 – 1.36); abdomen length 2.45 ± 0.27 (2.00 – 2.98), width 1.81 ± 0.24 (1.33 – 2.46); leg I 9.40 ± 0.54 (7.97 – 10.24); leg II 8.89 ± 0.52 (7.36 – 9.78); leg III 7.92 ± 0.43 (6.61 – 8.72); leg IV 10.07 ± 0.57 (8.30 – 11.10). Leg I metatarsus with v 2 - 2 - 3 spines (v 2 - 2 - 1 only on right leg of KUZ Z 2758). Distodorsal surface of patellar apophysis with 5 – 10 (left) or 5 – 11 (right) (usually 6, or 7) peg setae. Females. Measurements [mean ± 1 SD, followed by ranges in parentheses; n = 19, including KUZ Z 2760: for body length, abdomen length and width, n = 7 (KUZ Z 2760, Z 2761, Z 2763, Z 2765, Z 2783, Z 2790, Z 2794)]: body length 5.90 ± 0.54 (5.36 – 6.43); carapace length 3.13 ± 0.19 (2.78 – 3.52), width 2.18 ± 0.13 (2.02 – 2.29), head width 1.49 ± 0.09 (1.34 – 1.58); abdomen length 3.37 ± 0.39 (2.85 – 3.71), width 2.38 ± 0.37 (1.82 – 2.70); leg I 8.15 ± 0.39 (7.60 – 8.59); leg II 7.59 ± 0.43 (6.94 – 8.26); leg III 6.90 ± 0.41 (6.03 – 7.44); leg IV 8.89 ± 0.49 (8.19 – 9.44). Leg I metatarsus with v 2 - 2 - 3 spines, but sometimes with v 2 - 2 - 2 spines. Spermathecal head location slightly variable, located from anterior margin to almost middle part of epigynum, but heads always separated. Coloration. Carapace yellowish brown with reticulate olive-black markings on lateral sides of head, and with radical olive-black bands on thorax. Chelicerae, maxillae, labium and sternum yellowish brown; chelicerae deeper than others. Legs also yellowish brown, but paler than carapace, with olive black annulations. Dorsum of abdomen olive black with dull-yellowish broken chevrons (Fig. 2). Retreat. This species constructs a silken tube-like retreat, often V-shapes, with an opening at each end (Fig. 1 B) and silk signal threads extending from each opening.	en	Matsuda, Kenji, Ihara, Yoh, Nakano, Takafumi (2020): Description of a New Species of Cybaeus (Araneae: Cybaeidae) from Central Honshu, Japan. Species Diversity 25: 145-152, DOI: 10.12782/specdiv.25.145
0398EF3DE80BF2695F4BF8A63862FACA.taxon	distribution	Distribution. This species appears to be restricted to a very small montane area of approximately 1.5 km 2 located on the western side of the southernmost part of Lake Biwa, Japan (Fig. 6). Spiders identified as C. daimonji sp. nov. were collected from the western side of the Higashiyama- Sanjuroppo mountains. The confirmed northernmost habitat occupied by the species was located at Mt. Hieizan and the southernmost collection site was the type locality, Mt. Daimonjiyama. DNA sequences. In total 11 sequences were determined: paratype male (KUZ Z 2755), six sequences, ITS- 1 (LC 529208; 676 bp), 28 S (LC 529207; 790 bp), H 3 (LC 529206; 328 bp), COI (LC 529209; 658 bp), 12 S (LC 529211; 332 bp), and 16 S (LC 529210; 439 bp); and paratype female (KUZ Z 2764), five sequences, ITS- 1 (LC 529214; 676 bp), 28 S (LC 529213; 790 bp), H 3 (LC 529212; 328 bp), COI (LC 529215; 658 bp), and 12 S (LC 529216; 332 bp). According to the nuclear ITS- 1 and mitochondrial COI sequences obtained from the paratype male and paratype female, the males and females examined in this study clearly belong to the same species newly described in the present study. The ITS- 1 sequences obtained from the male and female are almost concordant with each other (675 / 676 bp), and their COI sequences are identical.	en	Matsuda, Kenji, Ihara, Yoh, Nakano, Takafumi (2020): Description of a New Species of Cybaeus (Araneae: Cybaeidae) from Central Honshu, Japan. Species Diversity 25: 145-152, DOI: 10.12782/specdiv.25.145
0398EF3DE80BF2695F4BF8A63862FACA.taxon	etymology	Etymology. The specific name is derived from the type locality, Mt. Daimonjiyama. The specific name is from a Japanese word, and thus treated as indeclinable.	en	Matsuda, Kenji, Ihara, Yoh, Nakano, Takafumi (2020): Description of a New Species of Cybaeus (Araneae: Cybaeidae) from Central Honshu, Japan. Species Diversity 25: 145-152, DOI: 10.12782/specdiv.25.145
0398EF3DE80BF2695F4BF8A63862FACA.taxon	discussion	Remarks. Females of C. daimonji sp. nov. may be most likely to be confused with females of C. shinkaii in their spermathecal characteristics. In addition to the feature of copulatory ducts, however, females of C. daimonji sp. nov. are distinguishable from those of C. shinkaii by spermathecal heads that are separated [vs. heads contiguous; fig. 2 - 2 - 30 - 23 in Ihara (2009 a)]. Although males of C. maculosus remains unknown, Ihara (2009 a) suggested a possibility that C. maculosus might belong to the same species as C. communis. Therefore, it is highly possible that males of C. daimonji sp. nov. are also distinguishable from those of C. maculosus by characteristics of the palpal tibia. In addition to the four “ medium-sized ” congeners, i. e., C. communis, C. kirigaminensis, C. maculosus, and C. shinkaii, males of C. daimonji sp. nov. and the other “ medium-sized ” C. tajimaensis Ihara and Nojima, 2004 share similar RTA characteristics, but the former can be distinguished from the latter by a triangular patella apophysis with peg setae concentrated distodorsally, an elliptic bulb, and a simple triangular conductor [vs. a trapezoidal patella apophysis with peg setae widely distributed dorsally, an almost circular bulb, and a complex hook-like conductor in C. tajimaensis; figs 8 E, 18 in Ihara and Nojima (2004)]. Although males of C. daimonji sp. nov. and the medium-sized C. tottoriensis Ihara, 1994 share the simple triangular conductor, C. daimonji sp. nov. differs from C. tottoriensis in its distally extended patellar apophysis with distodorsalpeg setae [vs. retrolaterally extended patellar apophysis with peg setae laterally in C. tottoriensis; figs 11 – 13 in Ihara (1994)]. Females of C. daimonji sp. nov. can be unquestionably differentiated from females of C. tajimaensis and C. tottoriensis by their epigynal features.	en	Matsuda, Kenji, Ihara, Yoh, Nakano, Takafumi (2020): Description of a New Species of Cybaeus (Araneae: Cybaeidae) from Central Honshu, Japan. Species Diversity 25: 145-152, DOI: 10.12782/specdiv.25.145
