taxonID	type	description	language	source
03981525FFA5FFA8FF11FDA5FBE90E3D.taxon	materials_examined	BANGLADESH: Sundarbans, 22 13 N, 89 50 E, May 1984, Sandom, J. H. 28 (fr.) (K); Chittagong, 8 km N of Teknaf, 16 Mar. 1979, van der Maesen, L. G. J. 3771 (fl.) (K) CHINA: Guizhou, Li Po County, Jan. 1988, Xianghou, S. 664 (fl.), (K); S. China, Dec. 1850, Seemann 2484 (fl.) (K); S. China, Prov. Quangtung, 1863, Tate, G. R. s. n. (fl.) (K); Fortuni A 123, (fl.) (K); Nan-t’o and mountains to north, Henry, A. 4629 (fr.) (K); Hong Kong, 28 Mar. 1968, Hu, S. J. 5097 (fl.) (K); Hong Kong, Sa Lo Tong to Hok-Tau, 9 Apr. 1992, Hu, S. Y. 20319 (fl.) (K); Hong Kong, Fo Tan South Valley, 20 Jun. 1970, Hu, S. Y. 10489 (fr.) (K); Hong Kong, 28 Mar. 1968, Hu, S. Y. 5097 (fl.) (K); Hong Kong, Kouy-tcheou, 1914, Cavaliere, M. 4492 (fl.) (K); Hong Kong, Lungchow, Morse, H. B. 545 (fl.) (K); Hong Kong, 300 yards Rifle Range Ravine, Urquhart s. n. (st.) (K); Hong Kong, Champion, M. s. n. (fl.) (K); Hong Kong, Wright, C. s. n. (fl) (K): Hong Kong, Wu Kau Tin, 2 Apr. 1928, Chun, W. Y. 6210 (fl.) (K); Hong Kong, Tank Ravine, 1861, Urquhart s. n. (fl.) (K); Hong Kong, Shek Pik Reservoir, 21 Feb. 1969, Shiu Ying Hu 6625 (fl.) (K) INDIA: Mangalor, Wight 618, (fl.) (K); Penins. Ind. Orientalis, Heyne s. n. (fl., fr.) (K); S. coll., 842 (Herb. Wight), (fr.) (K); S. Andaman, Manpur, 8 Oct. 1892, (fr.) (K); Andamans, 1915, Parkinson, C. E. 231 (fl., fr.) (K); Chittagong, 300 m, J. D. H. & J. J. 255 (fr.) (K); Cuttack, 7 May 1917, Herb. H. Haines 4690 (fr.) (K); Sambalpur, Cuttack District, Mahanadi Delta, Jambu, 4 Jun. 1949, Mooney, H. F. 3350 (fl., fr.) (K); Tamil Nadu, Viluppuram, Auroville, Matrimandir Nursery, 10 m, 22 Dec. 2002, Matthew, K. M. & Gastmans, W. F. RHT 74358 (fl., fr.) (K); Tamil Nadu, Viluppuram, Auroville, Matrimandir Nursery, 10 m, 22 Dec. 2002, Matthew, K. M. & Gastmans, W. F. RHT 73073 (fr.) (K); Chittagong, H. I. 5831 (fl.) (K); Bengal, 11 May 1883, Clarke, C. B. 33516 (K); Karwar, Dec. 1920, Bell, J. R. 7750 (fr.) (K); Bombay, North Kanara, 1885, Talbot, W. A. 1256 (fl.) (K); Kuppam River, Taliparamba, 8 Feb. 1913, Barber, C. A. 8788 (fl., fr.) (K) INDO-CHINA: Tonkin, environs de Haiphong, Apr. 1886, Balansa, B. 2143 (fl.) (K) INDONESIA: Aru Archipelago, Kobroor Riverside upstream from Jirlay, 6 Nov. 1994, Nooteboom 6111 (fr.) (K) JAPAN: Prov. Ryukyu, Yaeyama Islands, Is. Iriomote, Sonai to Shirahama, 31 Mar. 1974, Furuse, M. 5493 (fl.) (K); Prov. Ryukyu, Is. Ishigaki, Fukitooshi river mouth, Idona, 4 Mar. 1973, Furuse, M. 2561, (fl.) (K); Formosa, Tamsuy, 1864, Oldham, R. 184 (fl.) (K); Lunchow, Liukiu Islands, Yakuji Island, 7 Mar. 1917, Wilson, E. H. 8114 (fl.) (K); Formosa, Koshun, 28 May 1912, Price, W. R. 582 (fr.) (K); Ryukyus, Okinwa Pref., Taketomi cho, Funaura, Inda Point, 24 24 18.4 N, 123 49 47.5 E, 2 May 2003, Yasuda, K. 1735 (fr.) (K); Ryukyu, Okinawa Pref., Is. Ishigaki, Idona, Fukitooshi river mouth, 24 Sep. 1973, Furuse, M. 4103 (fr.) (K) MALAY PENINSULA: Pulau Jimun, Pulau Langkawi, 24 Nov. 1934, Henderson, M. R. 29110 (fr.) (K); Dayang Bunting, Langkawi Islands, W. Coast Malay Peninsula, Nov. 1916, Robinson, H. C. 6185 (K); Perak, Sumpai Krian Estate, 0 m, 9 Sep. 1937, Share, G. H. 33276 (fl.) (K); Port Dickson, 23 Feb. 1978, Kochummen, K. M. FRI 26561 (fr.) (K); Johor, Tanjung Penyabong Pontai, Pasir Lanun, 18 Apr. 2004, Lee, O. E. EL 47 (fr.) (K); Rompin-Nenasi road, 10 Feb. 1983, s. coll. FRI 27396 (st.) (K); Langkawi, Jul. 1892, Curtis’s collector 2867 (fl.) (K); Perak, Dec. 1880, Dr King’s collector 1029 (fl.) (K) MYANMAR: Rangoon, Thamaing, 15 m, Khant, P. 1079, (fl.) (K); Myebon, 0 m, 20 Feb. 1958, McKee, H. S. 6069 (fl.) (K); Tenassarim Division, Tawer District, west of Kanbauk, Heinze River estuary, Yebyu Township, 22 Oct. 1998, Maxwell, J. F. 98 - 1037 (fr.) (CMU); Bay of Bengal, Arkan Coast, Akyab Island, Fakir Point, 100 ft., 15 Sep. 1951, Vogt, G. B. BU- 223 (fl.) (K) A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 55 NEW CALEDONIA: Yate, Touaourou, 31 May 1984, Jaffre, T. 2503 (fl.) (K) NEW GUINEA: Papua, Lower Mori River, 28 May 1926, Brass, L. J. 1559 (fr.) (K); Irian Jaya, District Merauke, Ds. Sawa, Kec. Sawaerma, Agats, 10 m, 23 Oct. 1992, Widjaja, Wally & Subari EAW. EW 6234 (fr.) (K); Papua, Mount Jaya, Mimika Regency, PT-Freeport Indonesia Concession Area, Aikwa Estuary, along tidal waters of Aikwa River, with Nypa and Diospyros, 4 50 S, 136 59 E, 6 Mar. 1999, Willis, F. R. et. al. 82 (fr.) (K); New Guinea, Miscool Sorong, Waigama, 12 Sept. 1948, Pleyte, D. R. 820 (fr.) (K); New Guinea, Netherlands, Radjah Ampat District, Waigeo Island, Warmaserim Creek, E of Kabare, 7 m, 2 Feb. 1955, van Royen, P. 5428 (fr.) (K); Papua, Milne Bay District, Menapi, Cape Vogel Peninsula, 1 Apr. 1953, s. coll., s. n. (fl.) (K); New Britain, West Nakanai, Rapuri Village near Cape Hoskins, 5 Aug. 1954, Floyd, A. 6467 (fl., fr.) (K); New Britain, Estuary of Pani River, Hixon Bay, District W., Subdistrict Hoskins, 4 56 S, 151 26 E, 0 m, 8 Jun. 1973, Croft, J. R. & Katik, P. 34506 (fr.) (K); Papua, Mount Jaya, Mimika Regency, PT-Freeport Indonesia Concession Area, Pulau Puriri, a small island at the mouth of the Alkwa Estuary, 4 54 S, 136 52 E, 1 m, 2 Mar. 1999, Utteridge, T. M. A. 50 (fr.) (K); Papua, Omati, Gulf Division, 30 m, 14 Jan. 1955, Womersley, J. S. & Simmonds, N. W. 5081 (fl.) (K); Oeta, Sg. Oemar, 1 m, 12 Jun. 1941, Aet 229 (exp. Lundquist), (fl.) (K); New Britain, Central Nakanai, Walo Beach, 13 Aug. 1954, Floyd, A. 6539 (fl.) (K) PHILIPPINES: Sabtang, Savidug, 25 Mar. 1991, Barbon, Romero & Fernando PPI 1620, (fl.) (K); Mindoro, Bo. Manaul, Mansalay, 23 Dec. 1952, Sulit, M. D. 4895 (fr.) (K) SOLOMON ISLANDS: N. W. Kolombangara, Rei Area, 4 Jul. 1968, Mauriasi, R. et al. BSIP 11668 (fl.) (K); Cula River Area, South Vella Lavella, 17 Aug. 1968, Kotali, C. et al. BSIP 11203 (fl.) (K); S. E. of Paehena Pt., N. W. Sta Ysabel, 3 Dec. 1965, Beer, W. BSIP 7783 (fl., fr.) (K); New Georgia, 1927, Waterhouse, J. H. L. s. n. (st.) (K); Bougainville, Buin, 7 Apr. 1936, Voyce, A. H. 20 (fl.) (K); S. W. Guadalcanal, Lambi Bay area, 2 Nov. 1968, Mauriasi, R. et al. BSIP 12422 (fr.) (K); Oima Atoll, Juppy, B. 268 (fl.) (K); Russell Islands, Mbanika Island, Lever Point, 5 Feb. 1991, Herbst, D. 9350 (fl., fr.) (K) SRI LANKA: Walker 5833; Walker 68 (comm. Hooker, W. J.), (fl.) (K) TAIWAN: Formosa, Ape Hill, May 1889, Peayfair, G. M. H. 325 (fl. (K); Tamsuy, Formosa, 1864, Oldham, R. 309 (fl.) (K); South of Formosa, 28 Feb. 1867, Hance, A. 391 (fl.) (K); Formosa, Nov. 1918, Wilson, E. H. 11008 (fl.) (K); 20 Feb. 1918, Wilson, E. H. 9856 (fl.) (K) THAILAND: Kanchanaburi Province, Sangklaburi District, Toong Yai Naresuan Wildlife Reserve, Lai Wo Subdistrict, Ban Saneh Pawng, along Ro Kee stream, 200 m, 12 Oct. 1993, Maxwell, J. F. 93 - 1231, (fl.) (CMU); Kanchanaburi Province, Sangklaburi District, Toong Yai Naresuaz Wildlife Reserve, Lai Wo Subdistrict, Ban Saneh Pawng area, along Ro Kee stream, 200 m, 15 Jan. 1994, Maxwell, J. F. 94 - 56, (fr.) (CMU); La-Ngu, Salut, 8 Jan. 1918, Kerr, A. F. G. 13945 (fl., fr.) (K) VANUATU: Aniwa Island, Isavai village, 15 Dec. 1993, Curry, P. 1447 (fr.) (K); New Hebrides, Erromanga, between Nouanko Camp and Ipota, about 10 km E of Ipota, 9 Aug. 1971, Green, P. S. RSNH 1318 (fl., fr.) (K); Banks Islands, Port Patteson, 24 May 1896, Morrison, A. s. n. (fr.) (K); New Hebrides, Erromanga Island, Dillon Bay, 15 May 1928, Kajewski, S. F. 262 (fr.) (K); New Hebrides, Erromanga Island, de Cook Bay, 120 m, 23 Jul. 1983, Cabalion, P. 2185 (fl.) (K); New Hebrides, Port Vila, 12 May 1896, Morrison, A. s. n. (fl.) (K); New Hebrides, Banks Group, Vanua Lava Island, 6 Jul. 1928, Kajewski, S. F. 419 (fl.) (K); Port Vila, 3 Mar. 1994, Curry, P. 1524 (st.) (K) VIETNAM: Huu Lung District, Huu Lien Municipality, Huu Lien Protected Area, 21 40 15 N, 106 23 24 E, 510 m, 4 Apr. 1998, Harder, D. K., Loc, P. K., Hiep, N. T., Binh, N. Q. 4216 (fl.) (K, MO)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFA4FFA7FF11F9B9FC5C0908.taxon	materials_examined	CHINA: Guangxi Province, Luzhai Co., Mt. Guting, 260 m, 15 Apr. 2005, Wang, Y. - M. 62 (fl.) (K); Hunan Province, Yizhang Co., Mangshan Mt., 600 m, 21 Jun. 2004, Xiao, B. - Z. 3631 (fr.) (K); Western Hupeh, Apr. 1907, Jul. 1907, Wilson, E. H. 106 (fl., fr.) (K); Szechuan, Nanchuan Hsien, 5000 – 6000 ft., 15 May 1928, Fang, W. P. 773 (fr.) (K); Anhwei, Wangshan, 1973, Chow, K. S. 102 (fr.) (K) ETHIOPIA: Harar Province, 23 Sep. 1961, Burger, W. 992 (fr.) (K) INDIA: Tamil Nadu, Nilgiris, along old road Thalakunda-Mysore, Kalhatty, 2000 m, 3 Oct. 1982, Klackenberg, J. & Lundin, R. 212 (fl., fr.) (K); Herb. Heyne 5834 (fl., fr.) (K) JAPAN: Hondo, 24 Sep. 1914, Wilson, E. H. 7151 (fr.) (K) 56 • Phytotaxa 274 (1) © 2016 Magnolia Press	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFA4FFA7FF11F9B9FC5C0908.taxon	description	CLARK	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFA4FFA7FF11F9B9FC5C0908.taxon	materials_examined	MOZAMBIQUE: Tete Province, on Malawi border, 8 km north of Mlangeni, on Nchen-Dedza road, 1550 m, 30 Apr. 1970, Brummitt, R. K. 10289 (fl.) (K) SRI LANKA: Kandy District, Central Province, between Madugoda and Hunnasgiriya mile marker 24 / 19, 6 Mar. 1970, V. E Rudd & Balakrishnan, N. 3246 (fl., fr.) (K) TANZANIA: Arusha Region, Arusha District, Mount Meru, 1 km past Olmotonyi Forest Research Training Institute, 19 Dec. 1997, Herendeen, P. & Mbago, F. 19 - XII- 97 - 1 (fl., fr.) (K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFABFFA7FF11FE55FAAE0A75.taxon	materials_examined	CHINA: Kwangtung, T’aan Faan, Fang Ch’eng District, Kung P’ing Shan and vicinity, 25 – 30 Aug. 1936, Tsang, W. T. 26752 (fr.) (K); Yunnan, Si-chour-hsien, Hsin-chieh, 1100 – 1200 m, 24 Oct. 1947, Feng, K. M. 12602 (fr.) (KUN); Yunnan, Si-chour-hsien, Ping-chai, 1200 – 1300 m, 26 Oct. 1947, Feng, K. M. 12522 (fr.) (KUN)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFABFFA7FF11FD81FB140AC5.taxon	materials_examined	CHINA: Kwangtung, Lofoushan, 18 Dec. 1928, Tsiang Ying 1639 (fr.) (K); Kwangtung, Aug. 1877, Ford, C. 46 (fl.) (K); Millett s. n. (fl.) (K); Hong Kong, 1000 ft., 17 May 1883, Lo Fan Shan 141 (fl.) (K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD0FFDBFF11F89EFB3C08B8.taxon	materials_examined	AUSTRALIA: Queensland, Wide Bay, 0.7 km SW of Mt. Biggenden, WSW of Biggenden, 25 32 37 S, 151 58 60 E, 17 Nov. 2001, Bean, A. R. 18033 (fl.) (K); Queensland, Burnett District, 4.5 km S of Binjour, Mundubbera 9146 – 443727, 320 m, 25 54 S, 151 27 E, 12 Oct. 1988, Forster, P. I. PIF 4801 (fl.) (K, MO); Queensland, Burnett 68 • Phytotaxa 274 (1) © 2016 Magnolia Press	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD0FFDBFF11F89EFB3C08B8.taxon	description	CLARK District, Coalstoun Lakes, E of Goryndah, 25 35 S, 151 590 E, 27 May 1969, Telford, I. R. 961 (st.) (CANB); Queensland, Mt. Mueller, near Edgecombe Bay, 12 Dec. 1863, J. Dallachy 20 (fl.) (MEL)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFABFFA7FF11FD11FC400C10.taxon	materials_examined	PHILIPPINES: Palawan, Puerto Princesa, Mt. Pulgar, April 1911, Elmer, A. D. E. 12969 (fr.) (K) THAILAND: Chanburi, Pong Nam Rawn, Tamun, 200 m, 21 Oct. 1956, Smitinand, T. 3587 (fl.) (K); Kanchanaburi, Sai Yoke, 150 m, 14 09 N, 99 10 E, 26 Nov. 1971 (fl.) (K); Muok Lek, 200 m, 10 Nov. 1924, Marcan, A. 1866 (fl.) (K); Pak Chong, 300 m, 30 Dec. 1923, Marcan, A. 1532 (fr.) (K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD7FFDAFF11F9B5FB700B55.taxon	materials_examined	NEW CALEDONIA: Haut Diahot, Tende – Exp. Frouin, 12 May 1970, McKee, H. S. 21910 (fr.) (K, NOU); Haut Diahot, Tende, 650 m, Mar. 1979, McKee, H. S. 36669 (fl.) (NOU); Haute Diahot, 600 m, 12 May 1983, McKee, H. S. 41454 (fr.) (K) (MO, NOU); Mandjelia, 20 24 S, 164 32 E, 715 m, 4 Jan. 1978, Bamps, P. 6149 (fl.) (NOU, P); Pouebo, Mandjelia, Salandané, 26 Feb. 1970, McKee, H. S. 21648 (fr.) (K, NOU); Pouebo, Mt. Mandjelia, 750 m, Feb. 1980, McKee, H. S. 37777 (fl.) (NOU); Pouebo, Mandjelia, 28 Mar. 1980, McKee, H. S. 37992 (st.) (NOU); Mandjelia, above Pouebo, 700 m, 18 Aug. 1981, McPherson, G. 4080 (fr.) (MO, NOU, P); Mandjelia, above Pouebo, c. 700 m, 13 Nov. 1981, McPherson, G. 4375 (fl.) (MO, NOU, NSW, P); Mandjelia, J., 12 Jan. 2004, Munzinger, J. 1920 (fl.) (NOU); Mandjelia, 650 m, 15 Mar. 1983, Veillon, J. M. 5419 (fr.) (NOU, P); Auf den Bergen bei Oubatche, 800 m, 26 Dec. 1902, Schlechter, F. R. R. 15572 (fl.) (BR, K, L, M); Pouebo, Ouangati, 700 m, Oct. 1978, McKee, H. S. 35937 (fl.) (NOU); Prov. Nord, Mont Panie, 20 53 25 S, 164 47 46 E, 173 m, 27 Jul. 2009, A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 69 Clark, R. P. 146 (st.) (K, NOU, P); Prov. Nord, Mont Panie, 20 33 27 S, 164 47 10 E, 497 m, 28 Jul. 2009, Clark, R. P. 147 (st.) (K, NOU); Province du Nord, Mt. Panie, above Haut Coulna, 20 36 82 S, 164 44 40 E, 500 – 550 m, 24 Oct. 1999, McPherson, G., van der Werff, H. 17724 (fl.) (CANB, K, MO, NOU); Tanghene – Ouaieme, Oct. 1978, McKee, H. S. 35882 (fl.) (NOU); Poindimie, 1861 – 67, Vieillard, E. 2520 (fl.) (NY, P); Plateau de Tango, 400 m, 22 Oct. 1981, Veillon, J. M. 4683 (fl.) (K, NOU, P); Crête entre haute Tchamba et haute Amoa, 600 m, 14 Nov. 1968, McKee, H. S. 19943 (fl.) (NOU); Haute Tchamba, 21 0 55 S, 165 15 6 E, 10 Nov. 2002, Munzinger, J. et. al. 1454 (fl.) (MO); Forêt Plate, 21 8 – 21 10 S, 165 3 – 165 7 E, 500 – 550 m, 12 Aug. 1965, Bernardi, L. 10107 (fl.) (K); Region de Houailou, Aug. 1966, Odricourt 842 (st.) (NOU); Bourail, 500 m, 27 Feb. 1978, McKee, H. S. 34758 (fl.) (NOU); Vallee de Thy, 25 Aug. 1956, McKee, H. S. 5109 (st.) (K); Prov. Sud, near to Farino, 573 m, 21 36 51 S, 165 46 27 E, 22 Jul. 2009, Clark, R. P. 132 (fl.) (CHIC, K, NOU, P); Farino, foret Mépéou, Expl. Germain, 15 Apr. 1969, McKee, H. S. 20567 (fl.) (NOU); Prov. Sud, near Sarramea, 21 36 49 S, 165 48 49 E, 435 m, 23 Jul. 2009, Clark, R. P. 133 (fl.) (K, NOU); Prov. Sud, near Sarramea, 21 36 49 S, 165 48 49 E, 435 m, 23 Jul. 2009, Clark, R. P. 134 (st.) (K, NOU); 1 km S. of Col d’Amieu, 500 m, 16 Jan. 1961, McKee, H. S. 8148 (fl.) (L, P); Yaté, Les Dalmates, 3 Nov. 1984, McKee, H. S. 42358 (fl.) (NOU); Prov. Sud, Mont Koghis, 22 10 46 S, 166 30 36 E, 431 m, 21 Jul. 2009, Clark, R. P. 131 (fl.) (BRI, CHIC, K, NOU, P); Thy, 130 m, 8 Jun. 1980, Brinon, H. 750 (fl.) (NOU); Thy River Valley, c. 12 km NE of Nouméa, 8 Jun. 1980, McPherson, G. 2764 (fl.) (AAU, MO, NOU, NSW); nord de la Conception, Dec. 1870, Balansa, B., M. 3338 (fl.) (P); Riviere Bleue (gros Kaoris), 1974, Schmid, M. 5017 (fl.) (NOU); Locality uncertain: Mont Pembai, 700 m, 31 May 1990, McKee, H. S. 44930 (fl.) (NOU); 130 m, 08 Jun. 1980, Brinon, H. 750 (fl.) (K, NOU); Franc, I. 3056 (fl.) (NY); 1861, Pancher s. n. (fl.) (P)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFABFFA7FF11FA99FC150DAD.taxon	materials_examined	CHINA: Western China, Wilson, E. H. 3404 (fl.) (K); Kwangtung, Lokchong, 9 Jun. 1929, Tso, C. L. 21028 (fl.) (K); Szechuan, Kiating Fu, 300 – 500 m, Wilson, E. H. 3255 (fl.) (BM, E, K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFABFFA7FF11FA29FDFE0E8D.taxon	materials_examined	CHINA: Hong Kong, 1858, Wilford 169 (fl.) (K); Hong Kong, 1853 – 56, Wright, C. 148 (fl., fr.) (K); Hong Kong, Wilford s. n. (fl.) (K); Hong Kong, Dragons Back, Victoria Island, 11 Apr. 1970, Shiu Ying Hu 9873 A (fl.) (K); Hong Kong, Champion in Herb. Bentham 502 (fl.) (K); Hong Kong, Kowloon Reservoir, 27 Sep. 1969, Shiu, Y. H. 7982 (fr.) (K); Kwangtung, Naam Kwan Shan, Tsengshing District, 24 Apr. 1932, Tsang, W. T. 20317 (st.) (K); Hong Kong, Tai Mo Shan, 28 Mar. 1970, Shiu Ying Hu 9759 (fl.) (K); Hong Kong, Sai Kong, N. T. Lan Nai Chung, 4 Nov. 1969, Shiu Ying Hu 8570 (fr.) (K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFABFFA6FF11F8A8FB690BED.taxon	materials_examined	COCHINCHINA: Province Bien-koa, km 73 de la route col. 20, 17 Oct. 1931, Poilane, M. 19765 (fl.) (L, P) INDIA: Bengal, Chittagong, Garjania Range, Cowan, J. M. 523 (fl.) (NY); S. Andaman, Whani Khari, 13 Oct. 1894, King’s collector s. n. (fl.) (K); S. Andaman, Kalatang, 2 Dec. 1893, Dr King’s collector s. n. (fr.) (K); A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 57 Tenasserim and Andamans, s. coll. (Herb. Helfer) 1901 (fr.) (P); S. Andamans, Rangachang, 16 Nov. 1889, Prain, D. s. n. (fr.) (K) LAOS: Khammouan, Ban Mak Pheuang, 17 49 59 N, 105 20 59 E, 560 m, 2 Nov. 2005, Newman, M. F. et. al. LAO 824 (fr.) (E) MALAYA: Perak, Ipoh, Kinta Hill F. R., 1500 ft., 12 Jun. 1966, Zainuddin, R. 99756 (fl.) (L); Singapore, March 1908, s. coll., s. n. (fr.) (MO); Malaya, Perak, Sunga Ryah, Oct. 1880, King’s collector (Kunstler) 895 (fl.) (CAL, K) MYANMAR: Tennaserim Division, Tawer District, east of Kanbauk, c. 5 km east of Dawei River, Yebyu Township, 125 m, 18 Oct. 1998, Maxwell, J. F. 98 - 1186 (fl.) (CMU) SUMATRA: Prov. Lampung, NW of Kota Agung, 5 23 S, 104 25 E, 350 – 450 m, 17 May 1968, Jacobs, M. 8470 (fl.) (L) THAILAND: Chieng Rai, Mae Sai, 5 Feb. 1970, Sutisorn, S. 1585 (fr.) (BKF); E Pangnga, Ban Ko Road, Raad Island, 75 m, 1 Dec. 1986, Maxwell, J. F. 86 - 1019, (fl.) (L); Kaw Zao Zai, Pang-nga, 5 m, 5 May 1929, Kerr, A. F. G. 17343 (fr.) (BKF, K, L, P); Kaw Chang Ranawng, 7 Jan. 1929, Kerr, A. F. G. 16553 (fr.) (BKF, K, L); Nan Province, Sakoen National Park, Yod, Song Kwae 19 23 158 N, 100 32 11 E, 744 m, 16 Dec. 2010, La-ongsri, W., Romkham, N. 1246 (fr.) (QBG); Muang District, Trang, Kao Chong Forest, 2 Feb. 1985, Maxwell, J. F. 85 - 148 (fr.) (P); Sichon, N. Sritamarat, 50 m, 12 May 1928, Kerr, A. F. G. 15689 (fr.) (BKF, K, L, P); Songkhla Province, Boripath waterfall, 07 00 N, 100 12 E, 100 – 200 m, 9 Nov. 1990, Larsen, K., Larsen, S. S., Barfod, A. S., Nanakorn, W., Ueachirakan, W., Sirirugsa, P. 41234 (fl.) (P); Songkla, Hat Yai District, Ton Nga Chang Park, 50 m, 14 Feb. 1985, Maxwell, J. F. 85 - 191 (fr.) (K, L, P); Songkla, Nah Mawm, Klong Rhang Hill, 25 m, 10 Dec. 1985, Maxwell, J. F. 85 - 1092 (fr.) (MO, L); Trang, Rassada, Wat Nohng Si Chan limestone hill, 07 59 53.8 N, 99 44 11.1 E, 165 m, 17 Nov. 2011, Clark, R. P. et al. 251 (fr.) (K); Yanyau, Surat, 50 m, 21 Feb. 1930, Kerr, A. F. G. 18191 (fr.) (BKF, K, L, P)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF98FF94FF11FEA9FA280024.taxon	materials_examined	Type: — INDIA: South Andaman, near Port Blair at Protheropur, Rangachang, Prain s. n. Lectotype (here designated): — CAL (CAL 0000011174, image!); isolectotypes CAL (CAL 0000011172, image!), K (K 000264172!, K 000264173!) Synonym. Caesalpinia andamanica (Prain) Hattink. Reinwardtia 9 (1): 15. 1974. Synonym. Mezoneuron kunstleri Prain. J. As. Soc. Beng. ii 66 (1): 233. 1897. (Prain 1897 a) Type: — MALAYA, Perak, Sunga Ryah, Oct. 1880, Kunstler (King’s collector) 895. Holotype CAL; isotype K (K 000789322!). Woody climber or scandent shrub, spreading to 20 m in length. Stems with scattered recurved prickles to 5 mm; glabrous. Stipules caducous, triangular, ca. 1.5 – 2 mm × up to 2 mm broad at base, apex acute. Leaves with (3 –) 4 – 11 pairs pinnae; 8 – 12 leaflets per pinna; petiole 3 – 11.5 cm; leaf rhachis 4.7 – 28 [– 50] cm, with recurved prickles in pairs at pinna insertion points, and scattered on the internodes; pinnae (4 –) 7 – 15 cm, pinna rhachis with recurved prickles paired at leaflet insertion points, and sometimes scattered on the internodes, with a single prickle at the apex, ca. 1 – 2 mm; leaf rhachis, petiole, and pinna rhachis glabrous. Leaflets alternate (sometimes subopposite towards apex of pinna), with a subopposite apical pair; the terminal leaflets elliptic, oblong, obovate, or ovate, base oblique, apex rounded to retuse, 2.5 – 5.0 × 1.5 – 2.9 cm; lateral leaflets elliptic, oblong, or obovate, base oblique, apex rounded, emarginate, 2.5 – 4.8 × 1.4 – 2.9 cm; all leaves glabrous on both surfaces; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, faintly visible on both surfaces, midvein flat or sunken above, raised below. Inflorescence a panicle, axillary or terminal, to 60 cm [– 75 cm]; a few recurved prickles sometimes present on lower parts of peduncle; mostly glabrous, or with some hairs on pedicels; pedicels 10 – 20 mm, articulated 1 – 2 mm below the hypanthium in flower, 3 – 4 mm in fruit, very sparsely white or orange tomentose, hairs denser where pedicels are inserted on inflorescence, and at pedicel articulation. Bracts sometimes present, persistent, triangular, ca. 1 – 2 × 1 – 2 mm, subglabrous; bracteoles caducous, not seen. Flowers with a hypanthium ca. 2 – 3 × 4 – 6 mm, glabrous or sparsely white or orange tomentose; lower calyx lobe 7 – 12 × ca. 8 mm, other lobes ca. 3 – 6 × ca. 5 mm, all lobes glabrous. Median petal obovate, inrolled, slightly reflexed backwards, a pronounced transverse flap of tissue on inner surface between claw and blade; blade 3 – 4 × 5 – 7 mm, claw 3 – 4 × 3 – 4 cm; margin of flap of tissue covered with with dense, villous, orange hairs, petal otherwise glabrous. Upper lateral petals ovate, blade 4 – 10 × 4 mm, claw 1 mm; glabrous apart from a few hairs on the inner surface of the claw. Lower lateral petals slightly obovate, 5 – 10 × 4 – 5 mm, including claw 1 mm long; glabrous apart from a few hairs on the inner surface of the claw. Stamen filaments slightly flattened, 6 – 15 mm long; with dense, villous orange hairs on the inner surface, on the basal ½; anther 2 – 3 mm long. Ovary 4 – 7 mm long, style 4 – 8 mm long, both glabrous, stigma funnel-shaped, to 2 mm wide, the rim papillate. Fruit coriaceous; elliptic, base cuneate, oblique, apex obtuse, the central nerve extending to form a beak; 9 – 15 × 2.8 – 4.7 cm, wing 7 – 13 mm wide; fruit surface glabrous; venation reticulate, raised and clearly visible on fruit body, anastomosing and slightly visible on the wing; the persistent hypanthium reflexed. Seeds 3 – 4 per fruit; oval to circular, visible in outline on fruit exterior when mature, perpendicular to the wing or angled slightly towards the apex of the fruit.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF98FF94FF11FEA9FA280024.taxon	distribution	Distribution: — Myanmar, India, Malaya, Thailand, Laos, Vietnam, Sumatra. (Fig. 5). Habitat and ecology: — Forest; disturbed scrub; coastal; sandy beaches; river banks; chalk rocks; limestone. Elevation usually low, but to 500 m. Phenology: — Myanmar: Flowering October; Thailand, Laos, Vietnam: Flowering October – December, fruiting December – May; Malesia: Flowering May, October, fruiting October – March (Larsen & Larsen 1980). Preliminary conservation assessment: — Extent of Occurrence 2,306,205.590 km 2 = LC; Area of Occupancy 88.000 km 2 = EN. Category assigned: — Least Concern (LC). 12 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFAAFFA5FF11FC76FEE00D88.taxon	materials_examined	ANGOLA: Cazengo, 15 Jun. 1935, Gossweiler, J. 10.271 (fl., fr.) (BM, MO); Welwitsch, F. M. J. 608 (fr.) (BM, P); Loanda, Malange District, May – Aug. 1903, Gossweiler, J. 1364 (fr.) (K, P); Pungo Andongo District, Apr. 1857, Welwitsch, F. M. J. 606 B (fl.) (BM, K); Golungo Alto, between Sange and Camilungo, Welwitsch, F. M. J. 607 (fl., fr.) (BM, K, LISU, P); Dajalane, Gossweiler, J. 5930 (fr.) (BM); Cuanza-Sul, Gabela, Jul. 1908, Gossweiler, J. 4473 (fl.) (BM, K) BURUNDI: Bugarama, terr. Muramyva, 1900 m, Jun. 1969, Lewalle 3614 (fr.) (K) CAMEROON: Mont. Ngoyang, from 8 km to 1 km W of Ngoyang, village situe 12 km to the N of Lolodorf, 24 Jan. 1974, Letouzey, R. 12771 (st.) (P) CONGO: Belgian Congo, between Kasindi and Tubengo, Jan. 1932, Lebrun, J. 729 (fr.) (P); Belgian Congo, 9 Dec. 1937, 470 m, Louis, J. 6898 (fr.) (BM, P); Belgian Congo, Yomgambi, 70 m, 24 Aug. 1938, Louis, J. 10.984 (fl.) (BM, P); Belgian Congo, Kisozi, 30 May 1933, Lejeune, J. B. 41 (fl.) (NY); Belgian Congo, Kisozi, route de Kitego, Jul. 1933, Lejeune, J. B. 73 (fl., fr.) (K, P); Belgian Congo, 21 Jul. 1914, Bequart, J. 5087 (fl.) (P); Belgian Congo, mountains to the West of Lac Kivu, Tshibinda, 2000 m, 1929, Humbert, H. 7381 (fl.) (P); Belgian Congo, Nioka, 13 Aug. 1952, Liben, L. 366 (fl.) (K); Kivu Province, Kitshanga, 1400 m, Feb. 1959, Leonard, A. 2842 (fl.) (K); Manderi, 20 km SE of Sibiti, 3 45 S, 13 30 E, 20 Aug. 1965, Farron, C. 4518 (st.) (P); Piste Berunda-Vienne Deugese, 1500 m, 29 May 1959, Deville, A. 429 (st.) (K); Yanyambi, Ile Esali, 7 Nov. 1938, Louis, J. 12471 (fl.) (K); Yomgambi, Ile Baoke Mbole, 25 Aug. 1938, Louis, J. 10984 (fl.) (K); Zaire, Haut-Zaire, env. de Yangambi, Ile Esali sur le Zaire. 5 Mar. 1978, Lisowski, S. 47943 (fr.) (K); Zaire, Ile Esali, en face de Jangambi, 9 Dec. 1937, Louis, J. 6898 (fr.) (K); Zaire, Orientale Province, Mahaqi Territory, Nioka, 1700 m, 10 Sep. 1957, Froment, D. 239 (fl.) (K); environs de Dimonika, piste Manenge-La Loubomo, 17 Dec. 1982, Cusset, G. 1253 (st.) (P) GABON: Ogooue-Ivindo, Lope Reserve, 200 m, 0 15 S, 11 40 E, 14 Dec. 1992, White, L. 649 (fl., fr.) (MO); Moyen-Ogooue, road Lambarene to Bifoun, after Weliga 2, 90 m, 0 32.83 S, 10 12.73 E, 25 Jan. 2005, Wieringa, J. J., Mackinder, B. A., Niangadouma, R. 5627 (fr.) (K); Doudou Mts., c. 20 km NNW of Doussala, 260 m, 28 Mar. 1988, de Wilde, J. J. & Jongkind, C. 9596 (fr.) (MO, P) GHANA: Berekum, Sampa Road, 12 Dec. 1954, Morton, J. K. A 1057 (fr.) (K); Pra Suhien Forest Reserve, 10 Nov. 1971, Deaw, J. 357 (fl.) (MO) GUINEA-CONAKRY: Diomandou village, 900 m, 17 Feb. 1949, Jacques-Georges 3769 (st.) (MO); Bouloukountou, 2 Dec, 1905, Caille 14780 (fl.) (P) KENYA: Mpiji Ravine, 24 Jul. 1971, Flock, O. 552 (fr.) (MO) 58 • Phytotaxa 274 (1) © 2016 Magnolia Press	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFAAFFA5FF11FC76FEE00D88.taxon	description	CLARK	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFAAFFA5FF11FC76FEE00D88.taxon	materials_examined	LIBERIA: Lamco Hq. Camp, 600 m, 1 Dec. 1964, Adames, P. 800 (fl.) (K); Mount Nimba mining road, 15 Dec. 1966, Bos, J. J. 2414 (fl.) (K); 3 miles northeast of Suacoco, Gbargana, Central Province, Jan. 1951, Daniel, P. M. 112 (fr.) (BM, MO, NY); Yekepa, Granfield, 1965, Jacques-Georges, A. 27057 (fl.) (MO) MOZAMBIQUE: between Imala and Muite, 24 Oct. 1948, Barbosa, L. A. G. 2549 (fr.) (BM); Cabo Delgado, Distrito de Ancuabe, 6 Aug. 1983, Groenendijk, E. M. C., Dungo, J. 520 (fr.) (MO); Cabo Delgado, Macondes, between Mueda and Chomba, 22 Sep. 1948, Barbosa 2230 (fr.) (BM, K); Niassa Province, near Malema R. bridge, W of Malema, 620 m, 16 May 1941, Hanby, A. J. W. 4454 (fr.) (K); Niassa Province, Okoewangoe, 16 km, on road from Marrupa to Nungo, 720 – 750 m, 6 Aug. 1981, Jansen, P. C. M., de Koning, J., de Wilde, J. J. F. E. 86 (fr.) (K); Zambezia, between Namacurra and Nicuadala, 29 km from Namacurra, 29 Aug. 1949, Barbosa, L. A. G. 3886 (fl., fr.) (K); Sofala Province, Cheringoma District, Cheringoma Plateau, c. 20 km SW of Inhaminga, Campira Village, Mazamba River, 185 m, 18 32 32 S, 34 55 25 E, 22 Jul. 2006, Goyder, D. J. et al. 4083 (fl.) (K); Cabo Delgado, between Makondo and Mueda on road from Diaca, 659 m, 11 36 55 S, 39 38 22 E, 9 Sep. 2009, Burrows, J. E. et al. 11419 (fl) (K) RWANDA: Vallee Bukingwe-Bweza, Jan. 1933, Bequet, A. 361 (fl.) (K); Nyanza-Ruanda, 1926, Robyns 2436 (fl.) (K); Riumba, N. E. Koekitumba, 1400 m, 11 Feb. 1954, Christiaensen, A. R. 354 (fl.) (K) TANZANIA: Lindi Rural District, Mchinga Division, Kilangala Village, 150 m, 09 45 13 S, 39 33 41 E, 22 Nov. 2003, Kayombo, C. J., Sangu, G., Suleiman, H. 4792 (fr.) (MO); Mlalakwa River, 2 Jul. 1939, Vaughan, J. H. 2830 (fl.) (K); Minziro Forest Reserve, Jinja to Kakindo area, Kagera, Bukoba Rural District, 1130 m, 01 05 S, 31 31 E, Festo, L., Bayona, W. 563 (fr.) (MO); Minziro Forest Reserve, Kakindo Village, 1148 m, 01 08 54 S, 31 31 17 E, 7 Jun. 2001, Festo, L., Bayona, W., Wilbard, W. 1531 (fl.) (MO); Ubungo, neaer D’Salaam, 200 ft., 20 Oct. 1968, Harris, B. J. 2463 (fr.) (K); Morogoro District, Kimbosa Forest Reserve, Jul. 1952, Semsei, G. R. 791 (fl.) (K); Mpanda District, Kabesi in Mahali Mts., 6 S, 30 E, 4500 ft., 4 Sep. 1958, Jefford, T. G., Newbold, J. 2330 B (fr.) (K); Pugu Hills Forest Reserve, Kisarawe, 23 Aug. 1972, Harris, Mabberley & Mwasumbi DSM 2621 (fr.) (MO); Usambara, 900 m, 19 Aug. 1916, Peter, A. K 298 (fl.) (K); Usambara, 900 m, 21 Jan. 1916, Peter, A. K 129 (st.) (K); Usambara, Amani, 650 m, 15 Jan. 1914, Peter, A. K 3 (st.) (K); Usambara, Amani, 800 m, 16 May 1914, Peter, A. K 55 (st.) (K); Usambara, Amani, 900 m, 24 May 1916, Peter, A. 48793 (st.) (K) UGANDA: Ankole District, Kalinzu Forest, Jul. 1938, Eggeling, W. J. 3779 (fl.) (K); banks of Nyagak River, near Neapea, West Nile, May 1936, Eggeling, W. J. 3012 (fl.) (K); Buhweju, Bushenyi District, Nyakibungo, 1820 m, 0 17 N, 30 24 E, 10 Mar. 2000, Rwaburindore, P. K. 4647 (fr.) (K); Buhweju, W. Ankola, Karungu, 1600 m, 11 Apr. 1982, Rwaburindore, P. K. 896 (fr.) (MO); Buhweju, W. Ankole, Kamukaaki, 1550 m, 0 22 S, 30 28 E, 24 Sep. 1982, Rwaburindore, P. K. 1018 (fr.) (MO); Entebbe, Victoria Nyanga, 09 Aug. 1905, Bagshawe, A. G., Camb., M. B. 720 (fl.) (BM); Entebbe, Victoria Nyanga, 12 Jun. 1905, Bagshawe, A. G., Camb., M. B. 707 (fl.) (BM); Omukwatangwe, Mpebo, Kigezi, 5500 ft. Jul. 1939, Purseglove, J. W. 911 (fl., fr.) (K); Rukungiri District, Kinkizi County, Bwindi Impenetrable Forest, Kayonza Sector, 1365 m, 0 53 – 1 08 S, 29 53 – 29 50 E, 17 Feb. 1998, Eilu, G. 256 (fl.) (K); Omukwatangwe (Lukiga), Kayonza, Kigezi, 5000 ft., Apr. 1948, Purseglove, J. W. 2644 (fl., fr.) (K) ZAMBIA: Mwinilunga District, by R. Matonchi, 12 Feb. 1938, Milne-Redhead, E. 4551 (st.) (K) ZANZIBAR: Chukwani, 9 Sep. 1960, Faulkner, H. G. 2706 (fr.) (K); Chukwani, 28 Aug. 1961, Faulkner, H. G. 2892 (fl., fr.) (K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF95FF97FF11FE1AFC010C92.taxon	description	Fl. Trop. Afr. 2: 261. 1871. Type: — ANGOLA, Golungo Alto and Pungo Andongo, Dr Welwitsch s. n. Lectotype: — ANGOLA, Pungo Andongo, Welwitsch 606 LISU (LISU 208797, image!), (Brenan 1967); isolectotypes LISU (LISU 208798, image!), BM (BM 000842328!). Remaining syntype: — Golungo Alto, between Sange and Camilungo, Welwitsch 607 LISU; isosyntypes BM!, K (K 000232355!), P (P 000374798!). Synonym. Caesalpinia angolensis (Welwitsch ex Oliver) Herendeen & Zarucchi. Ann. Missouri Bot. Gard. 77 (4): 854. 1990. Scandent shrub or liana, to 20 m in length. Stems with scattered recurved prickles ca. 8 mm; glabrous or sparsely tomentose. Stipules semi-persistent, triangular, 1 – 2 × 1 – 2 mm; glabrous to moderately tomentose, or with hairs only on the margins. Leaves with (2 –) 3 – 11 pairs pinnae; petiole 1.8 – 6.5 cm; rhachis 5.7 – 30 cm, with recurved prickles in pairs at the pinna insertion points, and scattered on the internodes, sometimes densely; pinnae ca. 2.5 – 11 cm long, sometimes with recurved prickles in pairs at the leaflet insertion points, and scattered on the internodes; 4 – 8 [– 9] pairs of leaflets per pinna; leaf rhachis, petiole, and pinna rhachis sparsely to moderately whitish to orange pilose to tomentose. Leaflets opposite or subopposite; the terminal leaflets obovate, base oblique, apex obtuse to retuse, 1.0 – 2.6 [– 3.5] × 0.6 – 1.5 cm; lateral leaflets broadly oblong, base oblique, apex obtuse to retuse, 0.7 – 2.2 × 0.4 – 1.2 cm; upper surface glabrous, or sparsely tomentose, lower surface with long hairs on basal portion of the midvein, or along whole midvein, sometimes on basal part of blade, occasionally sparsely across whole blade; 2 ˚ veins anastomosing, 3 ˚ veins reticulate, scarcely visible above, slightly raised or faintly visible below. Inflorescence a raceme or panicle (often few-branched), axillary or terminal, to ca. 50 cm, axes sparsely to moderately orange tomentose, sometimes glabrous on the basal portion; pedicels 5 – 11 mm, not articulated, sparsely to moderately orange tomentose. Bracts sometimes present, persistent, oblong with acuminate apex, or triangular, 1 – 2 × 1 – 2 mm, subglabrous, with few hairs at margins, or sparsely tomentose; bracteoles caducous, lanceolate, sometimes broad in basal portion, tapering to a long-acuminate apex, 3 – 6 × 1 mm, sparsely to moderately orange tomentose on both surfaces. Flowers with hypanthium 2 – 4 × 5 – 10 mm; lower calyx lobe 7 – 9 × 3 – 6 mm; other lobes 5 – 7 × 3 – 4 mm; inner surface of lobes distinctly striate, outer surface of hypanthium and lobes glabrous to moderately orange tomentose, with margins ciliate. Median petal obovate, sometimes reflexed backwards, laterally inrolled, with a transverse flap of tissue on inner surface between claw and blade, 5 – 10 × 5 – 10 mm, claw 1 – 4 mm, the flap of tissue sparsely hairy, the margins of the blade with a fringe of hairs, and some hairs on the inner surface behind flap of tissue, and at the edges; outer surface glabrous. Upper lateral petals suborbicular to obovate, 6 – 12 × 5 – 8 mm, including claw ca. 1 mm long. Lower lateral petals oblong to elliptic, 8 – 13 × 4 – 10 mm, including claw 1 – 2 mm long; all laterals villous on inner surface of claw, and sometimes the basal ⅓ of blade, outer surface glabrous. Stamen filaments 11 – 17 mm long, densely villous on the basal ½ on inner surface; anthers 1.5 – 2 mm long. Ovary ca. 5 – 7 mm long, glabrous, or sparsely, or densely tomentose, sometimes partially or only along upper edge (see discussion); style ca. 10 – 12 mm long; stigma funnel-shaped, to 2 mm wide, the rim papillate. Fruit oblong, base cuneate, apex with wing extending to form a beak, 6.5 – 15.5 × 2.3 – 3.9 [– 4.3] cm, wing 5 – 9 mm wide; fruit surface glabrous to sparsely tomentose, sometimes with hairs only at base; venation reticulate, raised and clearly visible on fruit body, anastomosing and somewhat raised and visible on the wing; hypanthium persistent. Seeds 2 – 5 per fruit; elliptic, visible in outline on fruit exterior when mature, ca. 8 – 11 × 6 – 9 mm, brown. A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 7 8 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF95FF97FF11FE1AFC010C92.taxon	distribution	Distribution: — Liberia, Ghana, Guinea-Conakry, Cameroon, Gabon, Congo, Rwanda, Burundi, Uganda, Kenya, Tanzania, Zanzibar, Mozambique, Zambia, Angola (Fig. 2). Habitat and Ecology: — Moist evergreen forest; secondary forest; grassland; scrub; roadsides. Elevation 90 – 1900 m. Phenology: — Flowering and fruiting throughout the year. Preliminary Conservation Assessment: — Extent of Occurrence 8,238,225.345 km 2 = LC; Area of Occupancy 216.000 km 2 = EN. Category assigned: — Least Concern (LC). Taxonomic notes: — The indumentum of the ovary of M. angolense varies from being completely glabrous, to partially to densely tomentose. The distribution of populations with a hairy ovary corresponds broadly with the Guineo-Congolian phytogeographical region (White 1979), of which the dominant vegetation types are rainforest on well-drained sites and swamp forest on hydromorphic soils (White 1983). Those with a glabrous ovary correspond with the Swahelian regional centre of endemism and the Swahelian-Maputaland regional transition zone (Clarke 1998), of which the dominant natural climax vegetation is wet and dry forest, but has largely been replaced by secondary wooded grassland and cultivation (White 1983).	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFA9FFA4FF11F9D2FEF5092D.taxon	materials_examined	NEW CALEDONIA: Ile Art, Belep, Plateau Nord, Te Waala, 19 42 2 S, 163 36 50 E, 255 m, 24 Aug. 2009, Barrabe, L., Swenson, U., Munzinger, J. 952 (st.) (K, NOU); Ile Art, Belep, Plateau Nord, Te Waala, 19 42 9 S, 163 39 52 E, 248 m, 26 Aug. 2009, Barrabe, L., Swenson, U., Munzinger, J. 965 (fr.) (K, NOU, P); Ile Art, Plateau Nord, 200 m, 10 Dec. 1975, McKee, H. S. 30507 (fl.) (K, NOU); Prov. Nord, base of Tiebaghi, 20 29 25 S, 164 11 53 E, 15 m, 25 Jul. 2009, Clark, R. P. 141 (fr.) (BRI, CHIC, K, NOU, P); Prov. Nord, base of Tiebaghi, 20 29 24 S, 164 11 52 E, 15 m, 25 Jul. 2009, Clark, R. P. 142 (fr.) (K, NOU, P); Prov. Nord, base of Tiebaghi, 20 29 24 S, 164 11 54 E, 15 m, 25 Jul. 2009, Clark, R. P. 143 (fr.) (K, NOU, P); Dome de Tiebaghi, 400 m, 25 Apr. 1976, McKee, H. S. 31140 (st.) (K); Dome de Tiebaghi, 400 m, 25 Apr. 1976, McKee, H. S. 31191 (fl.) (K, P); Tiebaghi, 13 Jun. 1974, Sevenet, T. 677 (fr.) (MOU); Dome de la Tiebaghi, 400 m, 11 Jun. 1974, Veillon, J. M. 3068 (fr.) (K, NOU, P); Nétéa, Poirin Néa, 21 15 3 S, 165 11 56 E, 10 Jul. 2007, Barrabé, L., Mouly, A. 513 (fl.) (NOU); Prov. Nord, Poya, Tribu de Nétéa, 21 14 59 S, 165 11 46 E, 130 m, 1 Aug. 2009, Clark, R. P. 162 (fr.) (K, NOU, P); Prov. Nord, Poya, Tribu de Nétéa, 21 15 06 S, 165 11 21 E, 130 m, 1 Aug. 2009, Clark, R. P. 161 (fr.) (K, NOU); Prov. Nord, Poya, Tribu de A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 59 Nétéa, 21 15 26 S, 165 12 05 E, 90 – 100 m, 1 Aug. 2009, Clark, R. P. 164 (fr.) (K, NOU); Thy, 200 m, 27 Aug. 1978, Brinon, H. 392 (fr.) (NOU); Prov. Sud, Mont Mou, 22 04 29 S, 166 19 54 E, 130 – 170 m, 2 Aug. 2009, Clark, R. P. 165 (st.) (K, NOU); Route de Yaté, Dalmates, 150 m, 08 Dec. 1966, McKee, H. S. 16043 (fl.) (K, NOU); Locality uncertain: Paita, 21 Oct. 1923, White, C. T. 2173 (st.) (BRI); Nouméa, Baudouin 249 (fl.) (P); Nouméa, Pancher 176 (fl., fr.) (P)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB6FFB8FF11F983FD9408B8.taxon	description	Contribution à la Flore de la Nouvelle-Calédonie. Bulletin du Muséum National d’Histoire Naturelle 31: 482. 1925. Lectotype: — NEW CALEDONIA: Nouméa, Pancher 176, P (P 00379673!), Clark & Gagnon 2015; isolectotypes P (P 00379675!, P 00379674!). Remaining syntype: — NEW CALEDONIA: Nouméa, Baudouin 249, P (P 00751918!). Scrambling shrub, becoming lianescent, to ca. 5 m in length. Stems with scattered small recurved prickles, older stems with corky spine-tipped tubercles; sparsely to moderately whitish to ferruginous tomentose. Stipules semi-persistent, triangular, 0.5 – 2 × 0.5 – 2 mm, glabrous to densely tomentose. Leaves with (4 –) 5 – 11 pairs pinnae; (3 –) 5 – 15 pairs leaflets per pinna; petioles 2 – 7.5 cm; leaf rhachis (6 –) 9 – 25 cm long, unarmed (especially older leaves), or with small recurved prickles in pairs at the pinna insertion points, and sometimes scattered on the internodes; 38 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK pinnae (2 –) 3 – 7.5 (– 9) cm long; leaf rhachis, petiole and pinna rhachis glabrous or sparsely to moderately tomentose with whitish to ferruginous hairs. Leaflets opposite, sometimes subopposite on lower parts of pinna; oblong, strongly oblique at base, apex rounded to retuse; the terminal leaflets 0.7 – 2.2 × 0.4 – 1.2 (– 1.7) cm; lateral leaflets 1 – 2.3 × 0.4 – 1.5 cm; all leaflets glabrous on both surfaces, or sometimes with few ferruginous hairs on lower portion of the midvein, on upper or lower surface; midvein usually excentric, slightly diagonal, sunken above, slightly raised below, 2 ˚ veins anastomosing, 3 ˚ veins reticulate, veins faintly to clearly visible. Inflorescence a panicle, usually few-branched, axillary or terminal, 9 – 20 cm long, glabrous, or very sparsely, occasionally densely, white to ferruginous tomentose on the axes, hairs sometimes only at pedicel insertion points; pedicels 2 – 8 mm, articulated 1 – 2 mm below calyx. Bracts sometimes present, persistent, triangular, 1 – 2 × 0.5 mm, moderately to densely orange tomentose; bracteoles caducous, lanceolate to elliptic, apex acute, inrolled, 1 – 1.5 × 0.5 – 1 mm, sparsely to moderately ferruginous tomentose. Flowers with a hypanthium ca. 1 mm deep, usually glabrous, occasionally moderately ferruginous tomentose; lower calyx lobe ca. 5 × 3 mm, other lobes ca. 4 × 2 mm, all glabrous. Median petal (3 –) 4 – 6 × 2 – 2.5 mm, with a circular patch of dense, dark ferruginous hairs on inner surface between claw and blade, and hairs on margins of claw, outer surface glabrous. Upper lateral petals elliptic to slightly obovate, 4 – 5 × 2 – 2.5 mm, blade glabrous, claw very sparsely ferruginous tomentose on inner surface. Lower lateral petals elliptic to slightly obovate, 4 – 5 × 2 – 3 mm; blade glabrous, claw very sparsely ferruginous tomentose on inner surface. Stamen filaments 3 – 6 mm long, densely ferruginous tomentose on the basal ½; anthers ca. 0.5 – 0.75 mm long. Ovary 1 – 1.5 mm long, densely ferruginous pubescent, style 2 – 3 mm long, glabrous, or ferruginous pubescent on the basal ½ of length or less; stigma funnel-shaped, the rim papillate. Fruit elliptic, the base cuneate, apex acute or the wing extending to form a short beak; (3.4 –) 4.2 – 6.4 × 2 – 3.5 cm, wing 3 – 5 mm wide, extending along upper ⅔ to entire length of fruit; fruit surface glabrescent, sometimes with a few hairs at the base and on the margins; venation reticulate, raised and visible on the fruit body, no veins visible on the wing; hypanthium persistent. Seeds 1 per fruit; suborbicular, generally visible in outline on fruit exterior when mature, flat 1 – 1.1 cm diameter, brown, smooth.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB6FFB8FF11F983FD9408B8.taxon	distribution	Distribution: — Endemic to New Caledonia. Ile Art; Grand Terre, North and South Provinces, West side of island. (Fig. 25). Habitat and ecology: — Occurs in wet forest, or maquis, on ultramaphic soil. Elevation 0 – 400 m. Phenology: — Flowering December – July, fruiting June – August. Preliminary conservation assessment: — Extent of Occurrence 11,469.543 km ² = VU; Area of Occupancy 36.000 km ² = EN. Category assigned: — Data Deficient (DD). This species has been only occasionally observed. From its known distribution, this species might be more common than the actual herbarium data reveal. Therefore, M. baudouinii is A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 39 assigned as Data Deficient and should be re-assessed when more observations have been made that will clarify its distribution (RLA Flore NC 2016).	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFA8FFA3FF11FEB6FD6308DD.taxon	materials_examined	GAMBIA: 1890 – 1891, Brown-Lester, J. 32 (fl., fr.) (E, K); Abuko Game Reserve, Starin, E. D. 42 (fr.) (K) GHANA: Cape Coast Road, 22 Jan. 1970, Enti, A. A. 1637 (fl.) (MO); Vigue, C. 3487 (fl.) (NY); 500 ft., 27 Nov. 1953, Adams, C. D. 1915 (fl.) (K); Cape Coast, Dec. 1951, Taylor, C. J. 5447 (fr.) (K, P); Mankrong R. H., bank of River Afran, Morton, J. K. A 550 (fr.) (K); W side of Volta, 2 miles from Senchi Ferry, 29 Nov. 1953, Morton, J. K. 9437 (fl.) (K); Aburi Scarp, 2 Nov. 1953, Morton, J. K. s. n. (fl.) (K); Aburi Scarp, 500 ft., 27 Nov. 1953, Adams, C. D. 1915 (fl.) (K); Bomkrom, Afram Plain, Kwalui, E. P. Colony, 2 Jan. 1914, Chipp, T. T. 594 (fr.) (K); Dodowa, Akobzwerewtwi, Dec. 1928, Irvine, F. R. 1024 (fr.) (K) GUINEA: Diomandou, 20 Jan. 1949, Jacques-Georges, A. 3382 (fl., fr.) (MO); Portuguese Guinea, Bissau, Bisslanca, 13 Jan. 1945, s. coll. 1646 (fl.) (K, MO, P); Bissora, Naga, 3 Jan. 1954, d’Ory, J. 98 (fl.) (K); Jemberem, 15 m, 22 Jan. 1995, Malaisse, F. 14605 (fl.) (P); Kindia Coidier, 1 Jan. 1939, Chillou, J. 1121 (fl.) (P); environs de Kindia, 1929 – 1932, Jacques, F. s. n. (fl., fr.) (P); Dionandon, 1949, Adam, J. G. 3382 (fl., fr.) (P); Niayes, Cap. Vert, Dakar, vers la Lac Ratha, 1948, Adam, J. G. 963 (fl.) (P); Village Konin, 1906, Pobeguin, H. 1456 (fl., fr.) (P); Toukoumba, 6 Jan., MacLaud 455 (fl.) (P); Guinee Moyenne, Mamou Prefecture, Oure Kaba, between Marella and Mamou, 423 m, 10 9 33.5 N, 11 41 40.5 W, 26 Nov. 2011, Haba, P. K. 978 (fl.) (K, NHGC); Guinee Maritime, Kindia Prefecture, Kindia, Kolakhoure to Sotueta, top of fall, 180 m, 10 8 49.2 N, 12 27 11.6 W, 22 Nov. 2011, Cheek, M. 16240 (fl.) (K); Mamou, 10 Mar. 1927, Dalziel, J. M. 8406 (fr.) (K); Trepadeira, Catio, 25 Jan. 1954, d’Ory, J. 225 (fr.) (K) IVORY COAST: Ivory Coast, Bouna, 3 Dec. 1967, Geerling, C. & Bokdam, J. 1620 (fl., fr.) (MO); near ” Yaoblé boka ”, 3 Dec. 1986, Gautier-Beguin, D. & L. 449 (fl.) (MO); Ake Assi, L. 12.172 (fl.) (MO); Bouroukrou, Gemin de Fu, km 92, Chevalier, A. 16861 (st.) (P); Piste de Zoukoum, 5 Jul. 1985, le Thomas, A. & Dewin, T. 42 (st.) (P) LIBERIA: Loffa county, on road between St Paul River and Zorzor, 20 Dec. 1966, Bos, J. J. 2510 (fl.) (K) NIGERIA: Western State, 17 Dec. 1975, Brown, R. C., Opayemi, J. 923 (fr.) (MO); University of Ife campus, 08 Jan. 1971, Opayemi, J., Hall, J. B. 1871 (fl.) (MO); Ado Odo, 24 Dec. 1971, Opayemi, J. 1302 (fl.) (E); Biological Garden, Nigerian College, Ibadan, 28 Dec. 1960, Hambler, D. J. 1090 (fl.) (BM); Lagos, Olokemeji, Foster, E. W. 145 (fl.) (K); Ibadan, 26 Nov. 1958, Olorunfemi, J., Daramala, B. O. FHI 40315 (fl.) (K); Mbo-ugo (Ibo), 3 to 4 miles W of Ibadan, 600 ft., Jan. 1937, Newberry, R. J., Etim, A. E. 185 (fl.) (K) SENEGAL: Sine Saloum National Park, 0 – 50 m, 13 39 N, 16 29 W, 26 Jan. 1994, Madsen, J. E., Goudiaby, A., Traoré, S. 3055 (fl., fr.) (MO); Saligon Region, Casamance, 19 Jan. 1900, Chevalier, A. 3334 (fr.) (P); Saligon Region, Casamance, 20 Feb. 1900, Chevalier, A. 3335 (st.) (P); Casamance, bord de route Ziguinchon, 26 Feb. 1966, Fotius, K 182 (fl., fr.) (P); between Tobor and Bignona, 6 Feb. 1931, Trochain, J. 1568 (fl., fr.) (P); Bonngual, Casamance, Jan. 1900, Chevalier, A. 3332 (fl.) (P); Bignona, Chevalier, A. 3327 (fr.) (P); Casamance, Jan. 1900, Chevalier, A. 3333 (fl.) (P); Heudelot 163 (fr.) (P); 1837, Heudelot s. n (fr.) (P) SENEGAMBIA: Heudelot, 962 (fl., fr.) (K, P) SIERRA LEONE: Nov. 1944, Jaeger, P. 772 (st.) (P); 6 Dec., Elliot, G. F. S. 4167 (fl.) (BM); Kain Kordu, Kono District N. P., 12 Dec. 1965, Morton & Jarr. SL 3295 (fl.) (K); Kambui Hills, Kpindi-yali, Mende, 10 Jan. 1953, Small, D. 880 (fr.) (K); 1887, Oldfield s. n. (fr.) (K); Sasseni, Tambakka, 14 Jan. 1899, Scott Elliott, G. F. 4519 (fl., fr.) (K); Pangumu, Smythe, C. W. 65 (fl.) (K); Jama-Kangama, 16 Apr. 1914, Aylmer, G. 2 (fl.) (K); Deighton, F. C. 2580 (fl.) (K); Tonkolili District, Southern Sula Mountains, north of Farangbaya village near Bumbuna, along road to Kegbema, 350 m, 9 0 15 N, 11 41 50 W, 3 Dec. 2009, Clark, R. 188 (fl.) (BR, FBC, G, K, SL) TOGO: near Amoussoukope, 21 Dec. 1971, Jansen, J. W. A. 2588 (fr.) (MO); near Lome, 1900 – 1902, Warnecke 411 (BM, P); Dahomey, Poisson, E. s. n. (fl., fr.) (P); Dahomey, Cercle de Zagnanado: Pays de Hollis, Village de Alba, 4 Feb. 1910, Chevalier, A. 23028 (fr.) (P); Adetikope, 20 km north of Lome, 2 Nov. 1977, Ern. H. 2337 (fl.) (K); Koveto, 20 km from Tsevie, 4 Dec. 1977, Ern, H. 2602 (fr.) (K); Chito, 18 Dec. 1925, Howe, F. N. 1061 (fr.) (K) 60 • Phytotaxa 274 (1) © 2016 Magnolia Press	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFA8FFA3FF11FEB6FD6308DD.taxon	description	CLARK W. AFR.: Jacques-Georges, A. 2328 (fl.) (MO)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF9BFF96FF11FAE8FD180D9D.taxon	materials_examined	Type: — SENEGAMBIA: 1837, Heudelot s. n. Holotype P (P 00374799!); isotype P!. Synonym. Caesalpinia benthamiana (Baillon) Herendeen & Zarucchi. Ann. Missouri Bot. Gard. 77 (4): 854. 1990. Scandent shrub or liana to 10 m in length. Stems with scattered large recurved prickles, to ca. 1.5 cm, corky spine-tipped tubercles on older stems; glabrous or very sparsely orange tomentose. Stipules persistent, triangular, 1 × 1 – 3 mm, the margins tomentose. Leaves with 4 – 7 pairs pinnae; 4 – 6 pairs leaflets per pinna; petiole 2.5 – 7 cm; rhachis 7 – 26 cm, with recurved prickles in pairs at the pinna insertion points, and scattered on the internodes; pinnae 5 – 12 cm, sometimes with scattered recurved prickles at the leaflet insertion points, and scattered on the internodes; leaf rhachis, petiole, and pinna rhachis glabrous or sparsely whitish to orange tomentose. Leaflets alternate; the terminal leaflets obovate, base oblique, apex obtuse to retuse, 1.8 – 4.6 × 1 – 2.9 cm; lateral leaflets broadly oblong, base oblique, apex obtuse to retuse, 1.5 – 6 × 0.9 – 3.3 cm; all leaflets with both upper and lower surface glabrous, or sparsely tomentose, the hairs sometimes semi-appressed; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, visible and slightly raised on both surfaces. Inflorescence a raceme or panicle, axillary or terminal, to 50 cm, the axis up to 1 cm wide at base; sometimes with recurved prickles scattered on the lower axes; densely greyish or fawn to orange tomentose on flower-bearing parts of racemes, basal portions lacking flowers are glabrous, or sparsely hairy; pedicels 6 – 13 mm, articulated ca. 2 mm below the hypanthium, sparsely to moderately whitish to orange tomentose, hairs denser around articulation. Bracts sometimes present, persistent, triangular, c 1 – 2 × 1 – 2 mm, A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 9 sparsely tomentose, or on the margins only; bracteoles caducous, elliptic, acuminate; 2 × 1 mm, sparsely tomentose, the acuminate apex glabrous. Flowers with a hypanthium ca. 2 × 6 – 9 mm; lower calyx lobe ca. 7 – 9 × 4 – 6 mm, other calyx lobes 5 – 8 × 2 – 4 mm; hypanthium and calyx lobes moderately to densely fawn to orange tomentose on outer surface, inner surface sparsely tomentose. Median petal blade obovate, claw oblong, laterally inrolled, with a transverse flap of tissue on inner surface between claw and blade, 9 – 10 × 6 – 9 mm, the flap of tissue sparsely hairy, hairs on the margins of the blade and claw. Upper lateral petals broadly elliptic to suborbicular, 7 – 9 × 7 – 8 mm, including claw 1 mm long; inner surface of claw pilose, petal otherwise glabrous. Lower lateral petals elliptic to suborbicular, 7 – 10 × 6 – 7 mm, including claw 2 mm long; inner surface of claw sparsely pilose, petal otherwise glabrous. Stamen filaments 10 – 18 mm long, tomentose on inner surface, along the basal ¼ - ½ of length. Ovary 5 – 7 mm long, style 11 – 16 mm long, ovary densely fawn, greyish or pinkish tomentose, style glabrous or with few hairs; stigma funnel-shaped, to 2 mm wide, the rim papillate. Fruit oblong, base cuneate, apex rounded, sometimes with the central vein extending to form a beak; 8 – 10.7 × 2.3 – 2.9 cm, wing arising basally 1 – 5 mm along central vein, 4 – 7 mm wide; fruit surface sparsely to moderately tomentose, densely so when young; veins clearly visible on body, slightly or clearly in wing; pink when mature; hypanthium persistent. Seeds 2 – 4 per fruit; oval in outline and clearly visible when mature under the raised surface of the fruit body.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF9BFF96FF11FAE8FD180D9D.taxon	distribution	Distribution: — Senegal, Gambia, Guinea-Bissau, Guinea-Conakry, Liberia, Ivory Coast, Ghana, Togo, Nigeria (Fig. 3). Habitat and ecology: — Forest, scrub, savannah. Elevation to 900 m. Phenology: — Flowering November – April, fruiting December – April Preliminary conservation assessment: — Extent of Occurrence 1,399,533.101 km 2 = LC; Area of Occupancy 136.000 km 2 = EN. Category assigned: — Least Concern (LC).	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB2FFBDFF11FDC3FD180BEB.taxon	description	Flora Australiensis (2): 278. 1864 (as Mezoneurum brachycarpum). Lectotype: — AUSTRALIA, New South Wales, Richmond River, C. Moore s. n., K (K 000264179!), Hattink 1974 Remaining syntype: — AUSTRALIA, Queensland, Araucaria Range, Moreton Bay, F. Mueller s. n., K! Original syntype: — AUSTRALIA, Queensland, Wide Bay, Bidwell s. n., K! (this specimen is re-identified as M. scortechinii F. v. Mueller) Synonym. Caesalpinia brachycarpa (Benth.) Hattink nom illeg. non C. brachycarpa (A. Gray) Fisher. Reinwardtia 9: 53. 1974. Synonym. Caesalpinia subtropica Pedley. Austrobaileya 1 (1): 33. 1977. syn. nov. Scrambling shrub or liana to 5 m in length. Stems with minute scattered recurved prickles to 2 mm; sparsely to moderately whitish to pale orange pilose or tomentose. Stipules persistent, triangular, 1 – 1.5 × 2 mm, sparsely to densely appressed-pilose. Leaves opposite, with (3 –) 4 – 8 pairs pinnae; (3 –) 5 – 7 pairs leaflets per pinna; petioles 1.4 – 5.4 cm; leaf rhachis 4.5 – 25 cm, with small recurved prickles in pairs at the pinna insertion points; pinnae subopposite, 2.2 – 7 cm; leaf rhachis, petiole, and pinna rhachis sparsely to moderately densely whitish to pale orange pilose or tomentose. Leaflets alternate; the terminal leaflets obovate, the base oblique, apex obtuse, rounded, or retuse, 0.8 – 2.1 × 0.5 – 1.1 cm; lateral leaflets elliptic to oblong, base oblique, apex obtuse, rounded, or retuse, 0.7 – 1.9 × 0.5 – 1.2 cm; all leaflets with upper surface glabrous or with sparse whitish appressed to patent or tomentose hairs, lower surface with sparse whitish appressed to patent or tomentose hairs; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, veins faintly visible above, slightly raised below. Inflorescence a panicle to ca. 21 cm, the racemes opposite; terminal or axillary, with small, sometimes dense, recurved prickles scattered on the axes; moderately to densely whitish to pale orange tomentose; pedicels 3 – 5 mm (– 6 mm in fruit) long; articulated 1 – 2 mm below hypanthium; sparsely to moderately orange tomentose. Bracts persistent, triangular, 1 – 2 × 1 – 2 mm, sparsely to densely tomentose; bracteoles semi-persistent, broadly lanceolate, the apex acuminate, 2 – 4 × 1 mm, moderately to densely orange or golden tomentose on both surfaces. Flowers with a hypanthium 1 × 2 – 3 mm; lower calyx lobe 4 – 5 × 2 mm, other lobes 3 – 4 × 1 – 2 mm; hypanthium and all calyx lobes sparsely orange to golden tomentose (or hairs appressed), inner surface glabrous. Median petal oblong, laterally inrolled, ca. 4 × 2 mm, with circular patch of ferrugineous hairs on inner surface between claw and blade, claw with hairs on margins, outer surface glabrous. Upper lateral petals elliptic to obovate, 4 × 2 – 3 mm, including claw ca. 1 mm long, a few hairs on inner surface of claw, petal otherwise glabrous. Lower lateral petals elliptic to obovate, ca. 5 × 2 – 3 mm, including claw ca. 1 mm long, a few hairs on inner surface of claw, petal otherwise glabrous. Stamen filaments 4 – 6 mm long, tomentose on the basal ½; anthers ca. 0.75 mm long. Ovary ca. 2 mm long, style ca. 4 mm long, both glabrous; stigma cupular, the rim papillate. Fruit chartaceous, elliptic, base cuneate, apex acute, sometimes slightly beaked, 2.5 – 4.2 × 1.3 – 3 cm, wing 2 – 4 mm wide; fruit surface glabrous; venation reticulate, raised and clearly visible on fruit body, no veins visible on the wing; hypanthium persistent. Seeds 1 per fruit; large, flat, reniform, slightly visible in outline on fruit exterior when mature.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB2FFBDFF11FDC3FD180BEB.taxon	distribution	Distribution: — Endemic to Australia (New South Wales, Queensland). (Fig 21) Habitat and ecology: — Rainforest; secondary forest margin; savannah; roadside; sandy soil. Elevation 0 – 900 m. Phenology: — Flowering July – November, fruiting November – January Preliminary conservation assessment: — Extent of Occurrence 304,497.872 km 2 = LC; Area of Occupancy 228.000 km 2 = EN. 34 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK Category assigned: — Least Concern (LC).	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFAFFFA2FF11FCE6FDEB0DF0.taxon	materials_examined	BHUTAN: Sarbhang District, Singi Khola, 10 km SW of Sarbhang, 390 m, 26 50 N, 90 12 E, 9 Mar. 1982, Grierson, A. J. C., Long, D. G. 3263 (fr.) (E, K); Phuntsholing, 200 m, 26 52 N, 89 23 E, 3 May 1979, Grierson, A. J. C., Long, D. G. 798 (st.) (E); Riang, 2000 ft., 26 May 1923, Cave, G. H. s. n. (fr.) (E) BORNEO: Bangarmassing, 1857 1858, Motley, J. 965 (fl.) (K) CHINA: Yunnan, Szemao, Henry, A. 12835 (fl.) (K, NY); Yunnan, Szemao, 4500 ft., Henry, A. 12215 (fl.) (NY); Yunnan, Szemao, Henry, A. 13528 (K, NY); Yunnan, Henry, A. 12876 (fr.) (K, NY); Yunnan, Szemao, Henry, A. 13528 (fl.) (K); Yunnan, Tengyueh, 1500 m, Jul. 1912, Forrest, G. 8519 (fl.) (K); Yunnan, 1917 – 1919, Forrest, G. 307 (18406) (fl.) (K); Yunnan, Henry, A. 11404 (fr.) (K) INDIA: Assam, 4 May 1952, Rupehand 5332 (fr.) (L); Assam, Lushai Hills, Aijal, 900 m, Oct. 1927, Parry, N. E. 400 (fl.) (K); Assam, Masters s. n. (fr.) (L); Assam, Colonel Jenkins s. n. (fr.) (L); Assam, Palasbari, 20 Jul. 1949, Koelz, W. N. 24279 (fl.) (L); Assam, Phulbair, Garo Hills, Chand, T. R. 3073 (fr.) (L); Assam, Phulbari, Garo Hills, near sea level, 13 Apr. 1950, Koelz, W. N. 25073 fr. (L); Assam, Palasbari, 10 Jan. 1952, Koelz, W. N. 29192 (fl.) (L); Assam, Sapacaty near Dalawade, Nov. 1898, Dr Prain’s collector 472 (fl.) (L); Bengal, Jan 1881, Gamble, J. S. 9205 (fr.) (K); Bengal, 4 Dec. 1875, Clarke, C. B. 26758 (fl.) (K); Assam, Tengali Bam near Naga Hills, Dr Prain’s collector 499 (st.) (L, P); Behar, 1000 ft., Hooker, J. D. s. n. (fl.) (K); Bengal, Katikund, 28 Jan. 1903, Lace, J. H. s. n. (fr.) (E); Bengal, Patiya, 24 Dec. 1920, Cowan, J. M. 2186 (st.) (E); Bengal, The Terai, Falakata, Julpai, 100 m, Nov. 1898, s. coll., s. n. (fl.) (K); Bombay, Khaudala, St. Xav. Ravine, 28 Jan. 1951, Santapau, S. J. 12197 (fr.) (MO); Sikkim, lower hills, 1852, Hooker, J. D. s. n. (fl.) (NY); Chittagong Hill Tracts, 1886, Dr King’s collector 203 (fl.) (L); Concan, Hooker, J. D., Thomson, s. n. (fl.) (K); Guddshalli — Karwar, 1000 ft., Ambo 6818 (st.) (K); Dec. 1873, Gamble, J. S. 4045 A (fl.) (K); Bengal, 2000 ft., 1852, Hooker, J. D. s. n. (fl.) (K); Gamble J. S., 399 A (st.) (K); Gamble J. S., 399 C (fr.) (K); Bengal, 12 Feb. 1876, Clarke, C. B. 26950 (fl.) (K); Assam, Tezbal or Tezmui, 23 Nov. 1946, Mukheyie, D. C. 11 (fl.) (K); Jalhaiquri, Kumerqram, 300 ft., 14 Jan. 1900, Wood, H. s. n. (fr.) (K); Kerala, Ranni Valley, Kakkada River, Pamba Dam, 13 Mar. 1980, Ridsdale, C. E. 611 A (st.) (L); Lebong, 1800 m, Maeule, 4046 A (fl.) (K); India, Tavancore, 1817, Wallich cat. no. 5830 B (st.) (K); Mysore, Hassan District, 10 Dec. 1969, Saldanha, C. J. 15811 (fl.) (K); Mysore, Hassan District, 17 Dec. 1969, Saldanha, C. J. 15832 (fl.) (MO); Mysore, Hassan District, 30 Jan. 1969, Saldanha, C. J. 12590 (fr.) (MO); Salem, Namakkal, Kolli Hills, 1000 m, 24 Feb. 1980, Matthew, K. M. 26783 (fl.) (K); Sambalpur, 1500 ft., 7 Nov. 1940, Mooney, H. 1633 (fl.) (K); Sikkim, 1 – 4000 ft., Hooker, J. D. s. n. (fl.) (L); Sikkim, 3000 ft., Hooker, J. D. s. n. (fl.) (K); Sikkim, Darjeeling, Mongpo, 24 Sep. 1875, Clarke, C. B., 26763 (st.) (K); Sikkim, W. side of Nanga Chhu, Tista Valley, 1525 m, 27 32 16 N, 88 37 50 E, 11 Jul. 1996, Long, D. G., Noltie, H. J. 114 (fl.) (E); Tamil Nadu, Palni Hills, Madurai, Preiakulam District, Kumbakkarai-Vengayapparai stream, 600 m, 17 May 1987, Matthew, K. M., Rajendren, N. 49538 (fr.) (K); Tamil Nadu, Topslip, 14 Apr. 1992, McKey, D. 92 - 1 (st.) (K); Upper Assam, 1841, Jenkins (for Hooker) s. n. (fl.) (K, NY); Wallich cat. no. 5828 E (fl., fr.) (K) A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 61 JAVA: Mt. Gedeh, 1400 m, 19 Jul. 1939, van Steenis, C. G. G. J. 11496 (fl.) (K, L); purchased 1959, Herb. T. Horsfield, s. coll. s. n. (fl.) (K); 1802 1818, Horsfield, T. 140 (fl.) (K); ex Herbario Lugduno-Batavo, s. coll., s. n. (st.) (K); Madjenang, 30 m, 1915, Backer 18880 (fl.) (L); Semarang, 1300 m, 15 May 1899, Koorders, S. H. 35245 (st.) (L); s. coll., s. n., [Herb. Lugd. Bat. 908.120 – 163] (st.) (L); Singaparna, 1913, Backer 8481 (fl.) (L); E. Java, Besuki, Nusa Barung Island, E. Point, 2 km WE of Teluk Djeruk, 0 – 50 m, 8 40 S, 113 20 E, Jacobs, M. 4791 (st.) (L); Batav 2100 (fl.) (L) MYANMAR: North Myanmar, Ngawchang Valley, near Black Rock, 4000 – 5000 ft., 28 Feb. 1939, Kingdon Ward, F. 356 (fr.) (NY); Mergui, Thamihla, 2 Feb. 1927, Parker, R. N. 2536 (fl.) (K); Myitkyina, Namma, 1000 ft., 28 Feb. 1910, Lace, J. H. 5142 (fr.) (K); Prome District, Nganswi, 1 Feb. 1905, Lace, J. H. 2742 (fl.) (K) NEPAL: East Nepal, Helok-Iladanda, 8 Nov. 1963, Kanai, S., Murata, G., Togashi, M. 6301611 (fr.) (NY); Okhaldhunga a Manabhangjyang, 1700 m, 4 Nov. 1954, Zimmerman, A. 2021 (fl.) (K); East Nepal, Sangkuri Lekh, 1200 m, 26 52 N, 87 20 E, 3 Sep. 1967, Williams & Stainton 8327 (fl.) (K); Narayani, Chitwan National Park, River Rapti, Sauraha, 150 m, 27 34 21 N, 84 29 44 E, 21 Nov. 2004, DNEP 2 B 33 (fl.) (E); Trijuba Khola, 750 ft., 16 Feb. 1982, Stainton, J. D. A. 805 (fr.) (E); Kathgara-Rangali, 14 Dec. 1963, Kanai, H. et. al. 6301708 (fl.) (E); Phulchoki, south of Kathmandu, 1700 m, 06 Sep. 1972, Kanai, H. 722155 (fl.) (E); Wallich cat. no. 5830 C (fr.) (K); Hooker, J. D. 299 (fr.) (K) THAILAND: Chiang Mai Province, Huay Mae Mae, 700 800 m, 24 Jan. 1996, BGO staff 13 (fl.) (QBG); Chiang Mai Province, Muang District, Doi Sutep, St Louis Church area, 1075 m, 29 Dec. 1988, Maxwell, J. F. 88 - 1411 (st.) (L); Chiang Mai, Mae Dang District, Ban Pa Deng, Bah Miang Subdistrict, 900 m, 20 Jan. 1990, Maxwell, J. F. 90 - 102 (fl.) (L, MO); Chieng Rai, Chiengmai Province, upper slopes of Doi Chang Mountain, 4600 ft., 11 Jan. 1922, Rock, J. F. 1750 (fr.) (NY); Chonburi Province, Siricha District, Kow Kieo, c. 800 m, 13 16 N, 101 05 E, 4 Sep. 1976, Maxwell, J. F. 76 - 633 (st.) (L); Doi Khun Guam, 800 m, 14 Jan. 1814, s. coll., s. n. (fl.) (K); Chantabun, Makham, 120 m, 10 Dec. 1924, Kerr, A. F. G. 9592 (fl.) (K); Kanchanaburi Province, Sangklaburi District, Toong Yai Naresuan Wildlife Reserve, Lai Wo Subdistrict, Ban Saneh Pawng, along Ro Kee stream, 200 m, 18 Jan. 1994, Maxwell, J. F. 94 - 99 (fl.) (CMU, L); Mae Hong Son Province, Baan Pa Pae, Mae Sa Riang, 800 m, 10 Jan. 2001, Pongamornkul, W. 602 (fl.) (QBG); Mae Hong Son Province, Baan Pa Pae, Mae Sa Riang, 800 m, 10 Jan. 2001, Pongamornkul, W. 602 (fl.) (QBG); Mae Sanam, Hod, Chiang Mai, Mae Hong Son, 27 Dec. 1978, Niyomdham, C., Suanggho, M., Sangkhachard, B. 162 (fl.) (K); Nan Province, Dio Phu Waae, 14 Nov. 2000, Srisanga, P. 1890 (st.) (QBG); Nan Province, Doi Phu Kha National Park, 1550 m, 19 10 N, 101 07 E, 07 Nov. 1998, Srisanga, P. 357 (fr.) (QBG); Nan Province, Doi Phu Kha National Park, 1580 m, 19 10 N, 101 07 E, 12 Feb. 1999, Srisanga, P. 502 (fr.) (QBG); Pak Chong, 300 m, 1 Jan. 1924, Marcan, A. 1581 (fl.) (K); Wangka, Kanburi, 9 Feb. 1926, Kerr, A. F. G. 10479 (fr.) (K) VIETNAM: Indo-China, Tonkin, Cho Ganh, Province of Ninh Binh, February 1923, Petelot, A. 1116 (fl.) (NY); Annam Merid, col de Braian, pres de Djining, prov. Ht. Donnai, 4 Mar. 1935, Poilane, M. 24701 (fr.) (K, L); Annam, col d’Ailao, Prov. Duang-tri, near to Lant Hon Rao, 15 Mar. 1927, Poilane, M. 13612 (fr.) (K); Tonkin, Cu Cu, Jan. 1888, Balansa, B. 2150 (fl.) (K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF9FFF91FF11FBBFFCCF09DF.taxon	description	Prod. Fl. Penins. Ind. Or. 1: 283 (1834). Type: — INDIA: “ A native of the Delta of the Ganges where it was found by Dr William Carey, and by him introduced into the Botanic Garden at Calcutta where it blossoms in February and March ”. (Voigt, J. O. 1845). No specimen matching this description was found at CAL. “ Au BM, un spécimen Roxburgh s. n. pourrait être considéré comme type ” (Vidal & Thol 1976). A specimen of Caesalpinia cucullata Roxb., Roxburgh s. n. at BM (BM 000958800) has been annotated as the type by Vidal, Jul. 1974. Lectotype (here designated): — India Orientalis, Roxburgh s. n., BM (BM 000958800!) Epitype (here designated): — Wallich cat. n. 5828 E (2 sheets), K (K 000264194!, K 000264195!) Basionym. Caesalpinia cucullata Roxb. ex Roxb. Flora Indica 2: 538. 1832; Hort. Bengal: 32. 1814. nom. nud., as C. cuculata Roxb. Synonym. Mezoneuron macrophyllum Blume ex Miq. Fl. Bat. 1 (1): 104. 1855. Type: — JAVA: Batav 2100. Holotype L (L 0018781!); isotypes L (L 0018780!, L 0018782!). Synonym. Mezoneuron cucullatum (Roxb.) Wight & Arn. var. grandis Heyne ex Baker in Hook f., Fl. Br. Ind. 2: 258. 1878 (Caesalpinia grandis Heyne ex Wall., Cat. n. 5830 B). Type: — INDIA: Madras, Wallich cat. n. 5830 B, K (K 000264186!). Synonym. Mezoneuron cucullatum (Roxb.) Wight & Arn. var. robustum Craib. Fl. Siam. Enum. 1: 499. 1928. Type: — THAILAND: Wangkia, Kanburi [Rachaburi], 9.2.1926, Kerr 10479, K (K 000264158!). Robust woody climber or scandent shrub to 20 m in length. Stems with scattered recurved prickles, older stems with spine-tipped tubercles, glabrous. Stipules caducous, triangular or oblong, 1 × 1 – 2 mm, glabrous. Leaves with (2 –) 3 – 6 [– 7] pairs pinnae; 2 – 5 pairs leaflets per pinna; petiole 6.5 – 13 cm; leaf rhachis (10 –) 16 – 45 (– 55) cm, with paired recurved prickles at the pinna insertion points, and scattered on the internodes, prickles also on petiole; pinnae 6 – 21 cm; sometimes with recurved prickles at the leaflet insertion points and scattered on the internodes; leaf rhachis, petiole, and pinna rhachis glabrous. Leaflets opposite or subopposite; the terminal leaflets elliptic to slightly ovate or obovate, base symmetric or oblique, apex acuminate, slightly rounded, mucronulate, 4 – 14.5 × A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 13 2.2 – 6.5 cm; lateral leaflets elliptic to ovate, base symmetric to oblique, apex acuminate, slightly rounded, mucronulate, (3 –) 3.8 – 15.7 × 2.3 – 6.2 cm; all leaflets glabrous on both surfaces, upper surface glossy; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, veins raised and clearly visible on both surfaces. Inflorescence a raceme or panicle, axillary or terminal, to 65 cm, axes and pedicels sometimes with scattered recurved prickles; usually glabrous, but sometimes (very) sparsely short orange or golden tomentose; pedicels (4 –) 7 – 17 mm, articulated 2 – 5 mm below the hypanthium, glabrous or very sparsely orange tomentose. Bracts sometimes present, caducous, triangular, semicircular or oblong, 1 × 2 – 3 mm, glabrous; bracteoles caducous, 2 – 4 mm, narrowly lanceolate, glabrous or with few hairs. Flowers with a hypanthium 2 – 5 × 6 – 12 mm; lower calyx lobe 6 – 16 × 4 – 8 mm, other calyx lobes 4 – 10 × (3 –) 4 – 6 mm; hypanthium and calyx usually glabrous, sometimes (very) sparsely orange tomentose on outer surface. Median petal broad, bilobed to half of length, 7 – 10 × 12 – 17 mm, entirely glabrous, or ciliate on margins of basal portion. Upper lateral petals suborbicular, 6 – 12 × 5 – 8 mm, including claw ca. 1 mm long, moderately orange tomentose on inner surface of basal portion of blade, sometimes only on margins. Lower lateral petals elliptic to slightly obovate, apex rounded to truncate, 8 – 15 × (4 –) 5 – 7 mm, including claw ca. 1 mm long, moderately orange tomentose on inner surface of basal portion of blade, sometimes only on margins. Stamen filaments strongly exserted, flattened, 12 – 33 mm long, sparsely orange tomentose on the basal ½ on inner surface. Ovary 5 – 12 mm long, style 15 – 32 mm long, both glabrous; stigma truncate, oblique, ca. 0.5 mm diameter, glabrous. Fruit chartaceous; elliptic, the base oblique, apex rounded, or wing extending to form a curved beak; 7 – 14.5 (– 17) × 1.7 – 4.4 (– 5.2) cm, including stipe ca. 4 – 5 mm, and wing 4 – 10 mm wide; fruit surface glabrous; venation reticulate, raised and visible on fruit body, anastomosing and slightly visible on the wing; hypanthium persistent. Seeds 1 (rarely 2) per fruit; positioned centrally in fruit, generally visible in outline on fruit exterior when mature; flattened, orbicular, ca. 1 cm diameter, glossy, brown.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF9FFF91FF11FBBFFCCF09DF.taxon	distribution	Distribution: — China, Nepal, Myanmar, India, Thailand, Laos, Vietnam, Java, Borneo. (Fig 6). Habitat and ecology: — In evergreen or mixed evergreen-deciduous forest, forest fringes and scrub, stream margins, on limestone or granite. Elevation to 1800 m. Seems to prefer a strong seasonal drought, although is also known from regions without a dry season (Borneo). Phenology: — China: flowering July – February, fruiting March – October. India, Myanmar, Bhutan: flowering October – July, fruiting January – May. Nepal: flowering September – December, fruiting November – February. Thailand, Laos, Vietnam: flowering January – February, fruiting November – March. Malesia: flowering April – October, fruiting time unknown. 14 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK Preliminary conservation assessment: — Extent of Occurrence 10,973,076.606 km 2 = LC; Area of Occupancy 212.000 km 2 = EN. Category assigned: — Least Concern (LC). Taxonomic Notes: — The lectotype was selected on the basis of being a Roxburgh specimen that has been previously indicated by Vidal and Thol (1976) as a type, and annotated as such by Vidal, and that no more suitable collections have been found amongst the specimens studied. The leaf of this specimen is incomplete, however, and an epitype with a more complete and representative leaf is thus also here designated. The epitype also bears fruit, which is described in the protologue, but lacking on the lectotype.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFAEFFA1FF11FA0AFCC60998.taxon	materials_examined	BANGLADESH: Chittagong, towards Chandranath Hill, 400 ft, 22 30 N, 91 45 E, 13 Apr. 1994, Huq, A. M., Mia, M. K. 10262 (fr.) (F) CHINA: Morse, H. B. 216 (fr.) (K) INDIA: Cachar, Assam, D. H. & T. T. s. n. fr. (L); Chittagong, 0 – 1000 ft. J. D. H. & J. J., 120 (fr) (K); Cachar, 1 Dec. 1872, Clarke, C. B. 18523 (fr.) (K); Chittagong, 26 Feb. 1873, Clarke, C. B. 19984 A (fr.) (K) MYANMAR: District Rhyoon, Tankkyanson, 21 m from Rgn, 30 Jun. 1932, Parkinson, C. E. 14611 (fr.) (K, NY); C. V. 3601 (fl.) (K); Upper MYANMAR, Wuntho, [Herbarium of Henry Haselfoot Haines], 24 Dec. 1914, s. coll., 5065 (fr.) (K); Rangoon, McLelland, M. s. n. (fr.) (K); Rangoon, 4 May 1854, McLelland, M. s. n. (fr.) (K); Tavoy, H. 5834 G. H. (fl.) (K); Rangoon, 4 May 1854, McLelland s. n. (fr.) (K) SRI LANKA: Sinharaja forest, Weddegalle entrance, 200 m, 19 Oct. 1979, Kostermans, A. 27888 (fr.) (L) THAILAND: Kanburi, 50 m, 8 Dec. 1925, Kerr, A. F. G. 10100 (fr.) (BKF, K); Ban Nap Conarit, 150 m, 14 Jan. 1927, Kerr, A. F. G. 11424 (fl., fr.) (BKF, K); Ban Kao, 60 m, 16 Nov. 1961, Larsen, K. 8276 (fr.) (E); Chiang Mai Province, Siriphoom Waterfall, Doi Inthanon, Chomthong District, 1320 – 1460 m, 6 Jan. 1983, Koyama, H. et al. 62 • Phytotaxa 274 (1) © 2016 Magnolia Press	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFAEFFA1FF11FA0AFCC60998.taxon	description	CLARK 31986 (fr.) (L); Kopah, 12 Nov. 1897, Haniff s. n. (fl.) (K); Sukotie Prov., Muang Gow, Soke Pra Ruang Falls, near Sahwang Ah Rome Temple, 400 m, 22 Jan. 1974, Maxwell, J. F. 74 - 72 (fr.) (BKF); Phetchabun, Wang Chon Pu, 14 Jan. 1969, Ratchanapong 301 (fr.) (BKF); Sara Buri, Botanic Garden, at rear of accommodation, 14 40 04.1 N, 100 53 05, 17 Nov. 2013, Clark, R. 379 (fl.) (BKF, K, US)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFAEFFA1FF11FA0AFCC60998.taxon	materials_examined	VIETNAM: Lang Son Province, Huu Lung District, Huu Lien Municipality, 202 m, 21 40 437 N, 106 22 743 E, 8 Dec. 2005, Atha, D. et al. NY-HN 33 (fr.) (MO, NY); Dak Gley District, Dak Man Municipality, 850 m, 15 10 10 N, 107 45 12 E, 07 Jan. 2009, Averyanov, L. V. et al. HAL 11996 (fr.) (K, MO); Tonkin, Cho ba, Jul. 1908, d’Alleizette s. n. (fl.) (L) JAVA: Pajaman, 1917, Beumee, J. 871 (fl.) (L); Jatte Kalangan, s. coll. s. n. (Herb. Lugd. Batav.) (fr.) (L); Junghuhn, F. 722 (fl.) (K); Langku, 875 ft., Junghuhn 42 (fr.) (K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF81FF8CFF11FC0DFD180D8F.taxon	description	Pl. Jungh.: 258. 1852; Roxb. Hort. Bengal: 32. 1814. nom. nud. Basionym. Caesalpinia enneaphylla Roxburgh, Fl. Ind., ed. 2: 363. 1832. Holotype: — Roxburgh’s drawing n. 1425, K!. Synonym. Mezoneuron glabrum Desf., Mém. Mus. Par. iv: 245. 1818. Holotype: — TIMOR, Leschenault s. n., P (P 00293980!). Scandent shrub, 3 – 5 [– 15] m in length. Stems with scattered recurved prickles 2 – 3 mm; sparsely whitish tomentose when young, glabrous when older. Stipules triangular, flat, 1 – 3 × 1 – 3 mm, glabrous. Leaves with 5 – 14 pairs of pinnae; 7 – 10 [– 12] pairs leaflets per pinna; petiole 3 – 6.5 cm; leaf rhachis 6 – 32.5 [– 40] cm, with recurved prickles in pairs at pinna insertion points, and sometimes scattered on the internodes; pinnae 4.1 – 7.5 cm, occasionally with recurved prickles paired at the leaflet insertion points, and scattered on the internodes; leaf rhachis and petiole glabrous to sparsely to moderately whitish to pale orange tomentose; pinna glabrous to sparsely to moderately whitish to pale orange tomentose. Leaflets opposite to subopposite; the terminal leaflets obovate, base oblique, apex rounded to emarginate, 10 – 23 × 5 – 9 mm; lateral leaflets oblong, sometimes asymmetric, base oblique, apex rounded to retuse, 9 – 21 × 4 – 7 mm; all leaflets with upper surface usually glabrous, occasionally very sparsely appressed pale hirsute, or with few hairs at base, lower surface usually glabrous, occasionally with sparse appressed pale hairs; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, veins slightly raised, flat or sunken above, raised below. Inflorescence a raceme or panicle, terminal or axillary; occasionally with scattered recurved prickles on the peduncle; glabrous, or sparsely to moderately whitish to orange tomentose; pedicels 8 – 15 [– 20] mm, articulated 1 – 3 mm below hypanthium, glabrous or very sparsely whitish to orange tomentose. Bracts sometimes present, persistent, triangular or semicircular with acuminate apex, 1 – 2 × 0.5 – 2 mm, subglabrous to sparsely tomentose; bracteoles lanceolate, acuminate, 1 – 2 mm, sparsely orange tomentose on both surfaces. Flower with a hypanthium ca. 1 – 2 × [4 –] 6 – 8 mm; lower calyx lobe [5 –] 7 – 10 × 4 – 5 mm, other lobes [4 –] 5 – 6 × 3 – 5 mm, hypanthium and calyx sparsely tomentose on outer surface, glabrescent, inner surface glabrous. Median petal reflexed backwards, inrolled, blade obovate, claw distinct, oblong; a pronounced transverse flap of tissue on the inner surface between the claw and the blade; blade 4 – 6 × 5 – 7 mm, claw 3 – 4 × 2 – 3 mm; a few hairs along margins of claw, or the whole glabrous. Upper lateral petals blade ovate to elliptic, 7 – 9 × 4 – 7 mm, including claw 1 mm long, some hairs on inner surface of claw and base of petal, otherwise glabrous. Lower laterals blade ovate to A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 19 elliptic, 5 – 9 × 6 – 8 mm, including claw 1 mm long, a few hairs on inner surface of claw and base of blade, otherwise glabrous. Stamen filaments flattened, 8 – 10 (– 13) mm long, the vexillary stamen glabrous or nearly so, remaining 9 stamens densely villous on the basal ⅓ or ½, on the inner surface; anther 1.5 mm long. Ovary 4 – 7 mm long, style 6 – 9 mm long, both glabrous, stigma funnel-shaped, ca. 1 mm wide, the rim papillate. Fruit coriaceous, elliptic, base cuneate, apex rounded or acute, the central nerve extending to form a beak, 8 – 12 × 2.4 – 4.2 cm, wing 4 – 10 mm wide; fruit surface glabrous; venation reticulate, raised and partially visible on fruit body, anastomosing and slightly visible on the wing; hypanthium persistent. Seeds 3 – 5 [– 6] per fruit; positioned close to central vein, perpendicular to the vein, or angled towards apex of fruit, visible externally as linear ridges when mature; flat, ca. 7 × 5 × 2 mm, brown, smooth.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF81FF8CFF11FC0DFD180D8F.taxon	distribution	Distribution: — China, Bangladesh, Myanmar, Vietnam, Thailand, Sri Lanka, Java, Lesser Sunda Islands. (Fig. 10). Habitat and ecology: — Wet forest, scrub. Elevation to 1450 m. Occurs in regions that experience a dry season. Phenology: — Flowering July – January, fruiting October – June. Preliminary conservation assessment: — Extent of Occurrence 6,034,965.788 km 2 = LC; Area of Occupancy 68.000 km 2 = EN. Category assigned: — Least Concern (LC).	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFADFFA1FF11FDE2FAEC0B78.taxon	materials_examined	AUSTRALIA: Queensland, Old Rocky River, 75 m, 13 50 S, 143 25 E, 06 Sep. 1973, Hyland, B. 6812 (fl, fr.) (K, L); Queensland, Kennedy Road, 4 miles from P. R. – J. R. junction, Aug. 1965, Gittins, C. H. 1068 (fl.) (NSW); Queensland, Cape York Peninsula, Archer River, Wenlock-Coen Road, 30 Jul. 1948, Brass, L. J. 19746 (fl., fr.) (CANB, L); Queensland, Cook District, Port Stewart Road, beside Stewart River, 14 03 53 S, 143 19 06 E, 23 Aug. 2008, Cooper, W. 2027 (fl.) (CNS); Queensland, Cook District, Wenlok River, Chuula Outstation, Kaanju Nation, Central Cape York, 13 07 00 S, 142 58 35 E, 19 Nov. 2006, Smith, N. M. 5058 a & Claudie, D. (fl., fr.) (CNS)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB1FFBCFF11FC73FCBF0A2C.taxon	materials_examined	Type: — AUSTRALIA, Queensland, Cook District, Rocky River, ca. 13 50 S, 143 25 E, 6 Sep. 1973, Hyland, B. 6812. Holotype BRI (BRI-AQ 0016442, image!); isotypes K, (K 000789364!, K 000789365!), QRS. Synonym. Caesalpinia erythrocarpa Pedley. Austrobaileya 5 (1): 97 – 102. 1997. Scandent shrub or liana, to 20 m in length. Stems unarmed or with scattered recurved prickles; glabrous to sparsely whitish to orange tomentose. Stipules persistent, triangular, ca. 1 – 2 × 1 – 2 mm, subglabrous to sparsely tomentose. Leaves (sub) opposite, with (3 –) 4 – 7 pairs pinnae; 5 – 7 pairs leaflets per pinna; petiole 2 – 4.5 cm; leaf rhachis (4 –) 7 – 16.5 cm, with recurved prickles in pairs at pinna insertion points, and sometimes scattered on the internodes; pinnae 3.5 – 8 cm; leaf rhachis, petiole, and pinna rhachis sparsely to moderately whitish pilose to appressed-pilose. Leaflets alternate, occasionally opposite or subopposite; the terminal leaflets slightly obovate, base oblique, apex rounded to retuse, 1.2 – 2.1 × 0.7 – 1.4; lateral leaflets elliptic, base oblique, apex rounded to retuse, 1.1 – 1.8 × 0.6 – 1.3; all leaflets with upper surface glabrous or with scattered appressed hairs, lower surface glabrous or with sparse appressed hairs; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, 2 ˚ veins slightly visible above, 2 ˚ and 3 ˚ veins slightly visible below. Inflorescence a terminal panicle (occasionally a raceme), 13 – 40 cm (or more), the racemes opposite; axes subglabrous to moderately whitish to orange tomentose; pedicels 3 – 8 mm, articulated 0 – 1 mm below hypanthium; axes with few prickles on lower parts, sparsely to moderately orange tomentose, denser around articulation. Bracts persistent, triangular, ca. 1 – 2 × 1 – 2 mm, glabrous to sparsely tomentose; bracteoles caducous, lanceolate, the apex acute, 3 – 4 × 1 mm, sparsely tomentose. Flowers with a hypanthium 2 – 3 × 4 – 5 mm, outer surface moderately to densely orange tomentose; lower calyx lobe 6 – 7 × 3 – 4 mm, other lobes 5 – 6 × 2 – 3 mm, all (subglabrous to) sparsely to moderately orange tomentose, hair denser on lower lobe. Median petal obovate, 6 – 7 × 3 – 4 mm, with transverse flap of tissue on inner surface between claw and blade, the flap of tissue densely orange tomentose, petal otherwise glabrous. Upper lateral petals broadly elliptic, narrowed to claw at base, 5 – 6 × 4 – 5 mm, including claw 0.5 – 1 mm long, few hairs on inner surface of claw, petal otherwise glabrous. Lower laterals broadly elliptic, narrowed to claw at base, 5 – 6 × 3 – 4 mm, including claw 0.5 – 1 mm long, petal with few hairs on inner surface of claw, otherwise glabrous. Stamen filaments 6 – 7 mm long, moderately villous on the basal ½; anther ca. 1.5 mm long. Ovary ca. 3 mm long, densely appressed tomentose; style ca. 5 mm long, glabrous or with sparse hairs along entire length; stigma cupular or funnel-shaped, the rim papillate. Fruit chartaceous, elliptic, base cuneate, sometimes oblique, apex acute, sometimes with a slight beak, 3 – 5 × 2 – 2.7 cm, wing arising basally up to 5 mm along central vein, 4 – 5 mm wide; fruit surface with sparse, semi-appressed, whitish to pale orange hairs; A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 35 venation reticulate, raised and visible on the fruit body, no veins visible on the wing; hypanthium persistent. Seeds 1 per fruit, slightly visible in outline on fruit exterior when mature.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB1FFBCFF11FC73FCBF0A2C.taxon	distribution	Distribution: — Endemic to Australia (Queensland) (Fig. 22). Habitat and ecology: — Riverine rainforest. Elevation 75 – 850 m. Phenology: — Flowering and fruiting July – September. Preliminary conservation assessment: — Extent of Occurrence 16,461.975 km 2 = VU; Area of Occupancy 92.000 km 2 = EN. Category assigned: — Vulnerable (VU). Taxonomic notes: — Mezoneuron erythrocarpum does not occur in New Guinea, as it has been previously reported to. The specimen Brass 7498 (BM, BRI) from New Guinea, cited by Pedley (1997) as Caesalpinia erythrocarpa, is Mezoneuron scortechinii. The specimen Schodde 2246, listed in the Australian Virtual Herbarium (AVH 2015) as M. erythrocarpa, is also M. scortechinii.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFADFFA1FF11FC82FB4D0DAD.taxon	materials_examined	MYANMAR: Tenasserim, Ataran River, Pabang Hill, 1827, Wallich cat. no. 5835 (fr.) (K); Toungoo District, 10 Dec. 1909, Lace, J. H. 5024 (fl.) (K); Bawbui, 11 Feb. 1905, Lace, J. H. 2751 (fr.) (K); Dec. 1927, Parkinson, C. E. 5065 (fr.) (K) THAILAND: Ban Ta Dua, 28 Nov. 1920, Kerr, A. F. G. 4665 (fl.) (BKF, K); Chiang Mai Province, Chiang Dao District, the way to Waeng Haeng, 970 m, 20 Aug. 1999, Watthana, S., Suksathan, P., Argent, G. 557 (fl.) (QBG); Chiang Mai Province, Muang District, Doi Sutep, Pui National Park, east side of Doi Sutep, along the main road at c. km 11.5, 825 m, 27 Nov. 1995, Maxwell, J. F. 95 - 1185 (fl.) (CMU); Chiang Mai, Muang District, Doi Sutep, east side, along the main road, near km 11, 800 m, 13 Nov. 1988, Maxwell, J. F. 88 - 1305 (fl.) (L): Huay Gom Pao, Chiluquiar, 900 ft., 24 Mar. 1915, Lao Nome et. al. 168 (fr.) (K); Kanchanaburi Province, Sangklaburi District, Toong Yai Naresuan Wildlife Reserve, Lai Wo Subdistrict, Ban Saneh Pawng area, along Ro Kee stream, 200 m, 15 Jan. 1994, Maxwell, J. F. 94 - 63 (fr.) (CMU); Mae Saamlaep, Maesairang, Maehongson, 20 Jan. 1995, Pooma, R. 1015 (fr.) (CMU); near Neeckay (n. Wangka), 150 m, 25 28 Apr. 1946, Kostermans, 98 A (fr.) (L); Prov. Kanchanaburi, District Sangkhla Buri, Toong Yai Naresuan Wildlife Reserve, Lai We Subdistrict, Ban Saneh Pawng area (Karen village), along Ro Kee stream, 200 m, 15 Jan. 1994, Maxwell, J. F. 94 - 63 (fr.) (K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF80FF8AFF11F9D7FD180893.taxon	materials_examined	Type: — MYANMAR, Tennasserim, Martaban, Attran River, Wallich cat. n. 5835. Holotype K (K 000264175!); isotype BM (BM 000958805!) Paratypes: — MYANMAR, Pegu, Makhoye Hill, King’s collectors s. n.; SRI LANKA: “ Ceylon ”, Thwaites 3601 (location of these specimens unknown). Based on Caesalpinia furfuracea Wall., Wallich cat. n. 5835, 1831 – 32 (nomen). Synonym. Caesalpinia furfuracea (Prain) Hattink. Reinwardtia 9 (1): 1 – 69. 1974. Woody climber or scrambling shrub, height unknown. Stems with scattered recurved prickles up to 5 mm long; sparsely, sometimes irregularly, whitish to orange tomentose. Stipules persistent, ellipsoid, sometimes with acuminate apex, ca. 0.5 × 1 – 3 mm, glabrous to moderately tomentose, or on margins only. Leaves with 6 – 10 pairs pinnae; (5 –) 7 – 11 pairs leaflets per pinna, petioles 2 – 6 cm; leaf rhachis 9.5 – 29 cm, paired recurved prickles on 20 • Phytotaxa 274 (1) © 2016 Magnolia Press	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF80FF8AFF11F9D7FD180893.taxon	description	CLARK rhachis at pinna insertion points, and usually scattered on the internodes; pinnae 4.5 – 11.5 cm, pinna rhachis with single prickle at the apex, ca. 1 mm; leaf rhachis, petiole, and pinna rhachis sparsely or moderately whitish or orange tomentose. Leaflets opposite or subopposite; the terminal leaflets obovate, base oblique, apex rounded, obtuse or retuse, 1.8 – 2.9 × 0.8 – 1.3 cm; lateral leaflets oblong to slightly obovate, base oblique, apex rounded, obtuse to retuse, mucronulate, 1.5 – 2.8 × 0.6 – 1.1 cm; all leaflets with upper surface glabrous, lower surface glabrous apart from a few tomentose hairs around the base and along the midvein; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, veins faintly visible on both surfaces, or not visible. Inflorescence a raceme or panicle, axillary or terminal; sometimes with scattered recurved prickles on axes; axes glabrous or sparsely to moderately tomentose; pedicels 17 – 34 mm, articulated 2 – 4 mm below the hypanthium, glabrous. Bracts caducous, sometimes present, semicircular, 1 × 1 × 2 mm, glabrous or the margins tomentose; bracteoles caducous, narrowly lanceolate, 7 – 12 × 1 mm, sparsely tomentose with pale orange hairs. Flower s with an asymmetric hypanthium, 2 – 4 × 6 – 8 mm; lower calyx lobe highly cucullate, ca. 10 – 15 × 4 – 8 mm, other lobes ca. 6 – 8 × 3 – 6 mm; hypanthium and calyx glabrous. Median petal strongly reflexed backwards, laterally inrolled, with a circular patch of dense orange hairs on inner surface between claw and blade, outer surface glabrous or both surfaces glabrous if from the Lesser Sunda Islands, 4 – 10 mm long, blade 4 – 5 mm × 5 – 6 mm, claw 3 – 6 mm long. Upper lateral petals ovate to orbicular, blade 9 – 12 × 10 – 20 mm, claw ca. 1 mm long, glabrous. Lower lateral petals ovate to orbicular, blade 12 – 14 × 11 – 14 mm, claw 1 – 2 mm, glabrous. Stamens filaments flattened, 14 – 20 mm long, moderately orange villous on the basal ⅓ to ½ on inner surface; anthers 2 – 4 mm. Ovary 7 – 10 mm long, style 9 – 20 mm long, thickened towards apex; both glabrous, stigma funnel-shaped, ca. 1.5 mm wide, the rim slightly papillate. Fruit oblong-elliptic, base cuneate to rounded, sometimes oblique, apex cuneate to acute, sometimes with the central vein extending to form a beak; 12.5 – 20.5 × 4 – 6.6 cm, wing 10 – 18 mm wide; the persistent, asymmetric hypanthium wider below winged side of fruit; fruit surface glabrous, with a waxy whitish coating; venation reticulate, raised and visible on the fruit body, anastomosing and not or slightly visible on the wing; hypanthium persistent. Seeds 3 – 4 per fruit; elliptic, spaced ca. 1 cm apart, close to the central vein and more or less perpendicular to it, visible in outline on fruit exterior when mature, ca. 11 × 6 × 4 mm, brown, smooth, dull.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF80FF8AFF11F9D7FD180893.taxon	distribution	Distribution: — Myanmar, Thailand, Laos, Lesser Sunda Islands (Fig. 11). Habitat and ecology: — Open alluvial thicket along stream; mixed evergreen and deciduous forest; savannahs; scrub; rugged limestone terrain; sandy loam soil. Occurs in regions that experience a dry season. Elevation to 1000 m. (Fig. 11). Phenology: — Myanmar: flowering December, fruiting December – February. Thailand: August – November, fruiting January – April. Lesser Sunda Islands: flowering May – June, fruiting July. flowering A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 21 Preliminary conservation assessment: — Extent of Occurrence 1,669,797.015 km 2 = LC; Area of Occupancy 44.000 km 2 = EN. Category assigned: — Least Concern (LC).	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF9AFF94FF11F9D9FACA092D.taxon	materials_examined	Type: — MADAGASCAR: NW Madagascar, Nosi-Be, Hildebrandt 3399. Lectotype (here designated): — JE (JE 00000390, image!); isolectotypes JE (JE 00000391, image!, JE 00000392, image!), (K 000232354!), L (L 0651495!), M (M 0108401, image!), P (P 00131733!). Synonym. Caesalpinia hildebrandtii (Vatke) Baillon Bull. Mens. Soc. Linn. Paris i: 371. 1883. Synonym. Caesalpinia grevei Baillon. In Grandidier, Hist. Phys. Madagascar, Plantes, Atlas 1: plate 24 B & 24 CA. 1887. Type: — Illustration in loc. cit., Atlas 1: plate 24 B & 24 C K Scrambling or thicket forming shrub, or liana, to 10 m in length. Stems with recurved prickles, older stems with robust, woody, spine-tipped tubercles; glabrous. Stipules caducous, triangular, 1 – 2 × 1 – 2 mm, glabrous. Leaves with (2 –) 3 – 8 (– 9) pairs pinnae; 4 – 7 (– 8) pairs leaflets per pinna; petiole 3 – 6.5 cm; rhachis 8 – 25 (– 30) cm, with recurved prickles paired at the pinna insertion points, and scattered on the internodes; pinnae 6 – 10 cm; leaf rhachis, 10 • Phytotaxa 274 (1) © 2016 Magnolia Press	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF9AFF94FF11F9D9FACA092D.taxon	description	CLARK petiole, and pinna rhachis sparsely to moderately tomentose. Leaflets opposite to alternate; the terminal leaflets slightly obovate, base oblique, apex rounded to retuse, 1.4 – 3 × 0.8 – 2 cm; lateral leaflets elliptic to oblong, suborbicular, or slightly obovate, base oblique, apex rounded to retuse, 1.3 – 2.6 × 0.7 – 2 cm; all leaflets with upper surface glabrous, lower surface glabrous, or with few hairs on lower midvein; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, not or scarcely visible above, slightly visible below. Inflorescence a panicle, terminal and in the upper leaf axils, to 30 cm; sometimes with scattered recurved prickles on the lower axes; the axes glabrous; pedicels 3 – 7 mm, not articulated, glabrous. Bracts persistent, semicircular to triangular, ca. 1 × 1 mm, glabrous; bracteoles caducous, lanceolate, long-acuminate, 2 – 3 × 0.5 – 1 mm, glabrous. Flowers with a hypanthium ca. 1 × 3 – 4 mm; lower calyx lobe 5 – 6 × 2 – 3 mm, the margin usually with small, irregular teeth; other lobes 3 – 5 × 2 – 3 mm; hypanthium and calyx lobes glabrous. Median petal obovate, with pronounced transverse flap of tissue on inner surface between claw and blade; 3 – 5 × 2 – 3 mm, the flap of tissue and margins of the claw tomentose. Upper lateral petals obovate to suborbicular, ca. 3 – 6 × 2 – 3 mm including claw ca. 1 mm; outer surface glabrous, inner with few hairs on claw and lower blade. Lower lateral petals obovate, ca. 6 × 2.5 mm, including claw ca. 1 mm long; outer surface glabrous, inner with few hairs on claw and lower blade. Stamen filaments flattened, 7 – 10 × 0.5 mm long; densely villous on the basal half on inner surface; anther 1 – 1.5 mm. Ovary 5 – 10 mm long, style ca. 5 – 8 mm long; both glabrous; stigma funnel-shaped, ca. 1 mm wide, the rim papillate. Fruit chartaceous, elliptic, base attenuate, with stipe 2 – 5 mm, apex acute to acuminate, or retuse; 6.5 – 14 × 2 – 4 cm; wing arising basally ca. 6 – 10 mm along the central vein, 5 – 12 mm wide, fruit surface glabrous, glossy; venation reticulate, raised and clearly visible on fruit body, not visible on the wing; hypanthium persistent. Seeds 1 – 2 (– 3) per fruit; discoid to ovatediscoid, visible in outline on fruit exterior when mature, 7 – 9 × 6.5 – 8 × 2.5 – 3 mm, smooth, matte, olive green or tinged brown.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF9AFF94FF11F9D9FACA092D.taxon	distribution	Distribution: — Endemic to Madagascar: W & N Madagascar, from Antsiranana south to Morondava, including the Sambirano (but absent from south-west Madagascar) (Du Puy et al. 2002). (Fig. 4). Habitat and ecology: — Deciduous woodland and scrubland, often in exposed situations including bare limestone ‘ tsingy’ and along river margins, on limestone or sandy soils. Deciduous forest, scrub, red laterite soil. Elevation sea level to 600 m. Phenology: — Flowering April – June, fruiting May – June. Conservation assessment: — The species has a wide distribution range across Madagascar, where it appears to A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 11 be generally common and no major threats are known to affect the species at present, therefore Mezoneuron hildebrandtii is rated as Least Concern (IUCN 2015). Field notes: — Flowers white. Taxonomic notes: — The lectotype was selected on the basis of being housed at JE, the institution at which the specimens of author of the species name, Vatke, are primarily stored, and on being representative of the species.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFADFFA1FF11FA2CFDEF0F2B.taxon	materials_examined	MADAGASCAR: aux environs de Bounamary, 17 Jun. 1965, Peltier, M. 5252 (fr.) (MO); Diego-Suarez / Antsiranana Ambahatra, c. 330 m, 13 59 S, 48 26 E, 4 Jun. 2000, Wohlhauser, S., Andriamalaza, H. SW 60288 (fr.) (MO); Eritampiky, 25 May 1930, Decary, M. R. 7791 (fl.) (BM); l’Antsingy, vers Bevary (E. d’Antsalova), 400 – 600 m, 27 Jan. 5 Feb. 1960, Leandri, J., Saboureau, P. 2846 (st.) (MO); Majunga, s. coll., s. n. (fr.) (L); N. Madagascar, Antsiranana Province, c. 20 km S of Anivorano Nord, area east of the Ankarana Massif, c. 3 km S of Ambodromifehy, c. 270 m, 12 55 S, 49 12 E, 12 Jun. 1989, Du Puy, D. J. & B. P., Rafamantanantsoa, G. M 273 (fr.) (MO); N. Madagascar, Antsoranana Province, western margin of the Ankarana Massif, SE of Maysaborimanga, c. 150 m, 12 55 S, 49 04 E, 28 May 1989, Du Puy, D. J. & B. P., Rafamantanantsoa M 245 (fr.) (MO); Nosibe, Apr. 1880, Hildebrandt, J. M. 3399 (fl.) (BM)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFADFFDFFF11F8A9FCC60AA0.taxon	materials_examined	AUSTRALIA: Queensland, Cape York Peninsula, 15 m, 14 23 S, 144 13 E, 21 Nov. 1992 Fell, D. G. DGF 2746 (st.) (BRI); Cook District, Arthur Creek, 90 m, 12 50 S, 142 36 E, 27 Jul. 1985, Clarkson, J. R 6094 (fr.) (BRI); Queensland, Cook District, 110 m, 13 04 S, 142 40 E, 22 Apr. 1991, Clarkson, J. R. 8991 (fr.) (BRI); Queensland, A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 63 Cook District, 16 03 S, 144 07 E, 13 Aug. 1998, Godwin, M. MG 4281 (fr.) (BRI); Queensland, Cook District, 12 44 44 S, 142, 42, 39 E, 09 May 2003, Gray, B. 8594 (fr.) (BRI) CHINA: Yunnan, Bons d’Anty, M. s. n. (recu 19 Jul. 1898) (fr.) (K) INDIA: North Andamans, Lekmipur, 24 Nov. 1976, Nair, N. G. 4896 (fl.) (L); S. Andaman, Hobdaypur, 15 Oct. 1892, Dr King’s collector s. n. (fl.) (K); S. Andaman, Mount Harriett, 5 Nov. 1892, Dr King’s collector s. n. (fl.) (K); S. Andaman, Port Monatravine, 10 Nov. 1894, Dr King’s collector s. n. (fl.) (K); S. Andaman, Anikhet, 24 Dec. 1892, Dr King’s collector s. n. (fr.) (K); S. Andaman, W-gunj, 29 Dec. 1980, Heinig, R. L. s. n. (fr.) (BM) LESSER SUNDA ISLANDS: Kl. Sunda Inseln, Alor. Zentrales Hochland, Lantoka Pido, 850 m, 18 May 1938, Jaag, O. 1267 (fr.) (L); Kleine Sunda Inseln. Alor., Kabola Halbinsel, Adang-Sabandar-Alor ketjil. Strand. Flores, 300 m, Schmutz, P. A. 183 a (fr.) (K); Alor Kabola peninsula, Adang-Sabandar-Alor ketjil, 4 May 1938, Jaag, O. 422 (fr.) (K, L); Flores, Manggarai, Nisar, 250 m, 20 May 1982, Schmutz, E. 5362 (fr.) (L); Flores, Komodo, Tekaq, Kodeng, 3 Jun. 1982, Verheijen, J. A. J. 5264 (fr.) (L); W. Sumbawa, Semongkat Atas, 600 m, 3 May 1961, Kuswata 136 (fr.) (BM, BRI, K, L); Tanimbar Is., Jamdena, 4 Apr. 1956, van Borssum Waalkes, J. 3267 (fl.) (BRI, P); Tanimbar Is., Jamdena, 7 Apr. 1956, van Borssum Waalkes, J. 3320 (fl.) (BRI, L, P) MYANMAR: Wallich cat. no. 5832 (fr.) (K); Maymyo Plateau, 3500 ft., 20 Aug. 1912, Lace, J. H. 5896 (fl.) (K); Upper Myanmar, Meiktita, Aug. 1888, Collett, H. 839 (fl.) (K); Rubymines District, 21 Oct. 1912, Lace, J. H. s. n. (fr.) (K); 12 miles below Ava, Wallich cat. no. H. 5834 G (fl.) (K) PHILIPPINES: Luzon, Bulacan, Mt. Biak na Bato, San Miguel, 15 07 5 N, 121 04 7 E, 12 Nov. 1994, Garcia, H. G. PPI 15084 (fr.) (L) SRI LANKA: Gardner, CP 2784 (no label data) (BM, K); 8 Sept. 1931, Simpson, N. D. 8548 (fl.) (BM) SULAWESI: Kabaena, Batu Awu, 50 m, 30 Jun. 1978, Widjaja, E. A. 661 (fr.) (L) THAILAND: Chantabun, 20 m, 10 Dec. 1924, Kerr, A. F. G. 9589 (fr.) (BKF, K); Chiang Mai Province, Mae Rim, 850 m, 10 Feb. 1999, Nanakorn, W. et al. 11748 (fr.) (QBG); Chiang Mai Province, Maesa Falls, Maerim- Samoeng Rd., Mae Rim, 2 Nov. 1995, BGO staff 5075 (fr.) (QBG); Chiang Mai Province, Queen Sirikit Botanic Garden, Mae Rim, 27 Jul. 1995, BGO staff 3950 (fl.) (QBG); Chiang Mai Province, Queen Sirikit Botanic Garden, Mae Rim, 700 m, 3 Dec. 1993, BGO staff 243 (fr.) (QBG); Chiang Mai Province, Queen Sirikit Botanic Garden, Mae Rim, 700 m, 27 Aug. 1995, BGO staff 4648 (fl.) (QBG); Chiang Mai Province, San Kamphaeng District, Doi Lon, west side, above Huay Gayo Village, Huay Gayo Subdistrict, Mae Gahm Bawng Village, 650 m, 11 Sept. 1996, Maxwell, J. F. 96 - 1184 (fl.) (CMU, L); Chiang Mai, 20 Aug. 1910, Kerr, A. F. G., 1329 (fl., fr.) (K, L, P); Chiang Mai, 300 m, 10 Feb. 1911, Kerr, A. F. G. 1329 A (fr.) (K); Chiang Mai, Chiang Dao, Doi Chiang Dao Animal Sanctuary, Ban Yang Toeng Bong Forest Station, 450 m, 27 Oct. 1990, Maxwell, J. F. 90 - 1182 (fr.) (L, MO), (fl.) (L); Chiang Mai, Sangampang, Doi Muang Awn, 475 m, 26 Dec. 1989, Maxwell, J. F. 89 - 1598 (fr.) (E, L, MO); Chiang Rai, Amphoe Chiangkham, 27 Dec. 1967, Prayad 1154 (fr.) (BKF); Chiang Rai, Mae Suai Ban Saen Sa-at, 10 Aug. 1967, Bunchuai, K., Nimanong, B., 1489 (fl.) (E, K, L, P); Chieng Mai, Nam Tok Mai Klang, 22 Nov. 1965, Prayad, S. 83 (fr.) (BKF); Chompoo Mountain, Bahn Beung District, Chonburi Province, 13 16 N, 101 05 E, 25 Oct. 1976, Maxwell, J. F., 76 - 704 (fl.) (BKF, L); E., District Nakhon Ratchasima, Salika forest, 40 km SE of Pakthongchai, 400 m, 14 40 N, 102 2 E, Beusekom, C. F., Geesink, R. 3378 (fr.) (L, MO); Eastern, NW of Phu Khieo, 700 m, 16 25 N, 102 05 E, 6 7 Aug. 1972, Larsen, K., Larsen, S. S., Nielsen, I., Santisuk, T. 31497 (fl.) (P); Kanburi, 30 m, May 1917, Winit, K. 487 (fr.) (K); Kanchanaburi Province, Thong Pha Phum District, Chalae Subdistrict, Forest Research and Demonstration Centre, 700 m, 15 Dec. 1995, van Welzen, P. C. 84 (fr.) (K); Kanchanaburi, Sai-yok, Namtok (station), 15 Aug. 1967, Uaseun 577 (fl.) (BKF); Kanchanaburi, Tawng Pa Poom District, Huay Bahn Gow, 650 m, 14 Jul. 1973, Maxwell, J. F. 73 - 295 (fl.) (BKF); Kerr, A. F. G. 4700 (fr.) (BKF, E); Kow Kieo, Siricha District, Chonburi Province, 125 m, 18 Oct. 1975, Maxwell, J. F. 75 - 1024 (fl.) (BKF, L); Loei Province, Na Haew, 1000 m, 29 Jul. 1995, BGO staff 3990 (fl.) (QBG); N. Tak, Mae Sot, 5 Apr. 1973, Phengkhlai, C. 3142 (fr.) (K, L); Nakon Sitamarat, Ta Sala District, Khao Luang National Park, Groong Ching Falls, 300 m, 15 Dec. 1985, Maxwell, J. F. 85 - 1122 (fl.) (L); Nakorn Najok Province, Muang District, Kow Yai National Park, road to Ha Jo Sawat Falls, km 44, Hin Dahng Subdistrict, 700 m, 10 Jul. 2003, Maxwell, J. F. 03 - 175 (fl.) (CMU); Nakorn Sawan Province, Mae Wong District, east side of Mae Gratu mountain, near Tahn Ma Yom stream, Wang Sahn Subdistrict, 200 m, 26 Nov. 2005, van de Bult, M. 853 (fl.) (CMU); Nan Province, Doi Phu Kha National Park, 1050 m, 19 13 N, 101 05 E, 9 Oct. 1998, Srisanga, P. 298 (fr.) (QBG); Nan Province, Doi Phu Kha National Park, 1050 m, 19 13 N, 29 Jul. 1998, Srisanga, P. 250 (fl.) (QBG); Northern, Maehongson, Khun Yuam, 600 – 700 m, 18 15 N, 98 E, Larsen, K. 34232 (fl., fr.) (L); Payao Province, Muang District, Doi Luang National Park, east side, 64 • Phytotaxa 274 (1) © 2016 Magnolia Press	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFADFFDFFF11F8A9FCC60AA0.taxon	description	CLARK Dtawm Nai Village area, 525 m, 12 Aug. 1997, Maxwell, J. F. 97 - 866 (fl.) (CMU); Peninsular, Ranong, Kapos, 22 Nov. 1965, Sangkhachand, B. 1158 (fl.) (K); Phayao Province, Muang District, Doi Luang National Park, Mae Dtawm Nai Village, 525 m, 12 Aug. 1997, Maxwell, J. F. 97 - 866 (fl.) (L); Sahm Lahn forest, Muang District, Saraburi Province, 100 m, 10 Aug. 1974, Maxwell, J. F. 74 - 761 (fl.) (BKF, L); Salika forest, 40 km S. E. from Pakthongchai, 400 m, 14 40 N, 102 2 E, 26 Oct. 1971, van Beusekom, C. F., Geesink, R. 3378 (fr.) (MO); Saraburi Province, Muang District, Sahm Lahn forest, 100 m, 17 Oct. 1973, Maxwell, J. F. 73 - 543 (fr.) (BKF); Sriracha, 30 ft., 4 Nov. 1927, Collins, D. J. 2036 (fl.) (K); Sriracha, 4 Dec. 1927, Collins, D. J. 1852 (fr.) (US); Suracha village, 20 ft., 15 Jan. 1927, Collins, D. J. 1351 (fl.) (K); Tak Province, Namdip, Tak-Lampang road, 150 m, 25 Nov. 1995, Pooma, R. 1288 (fr.) (CMU); Tak Province, Thung Yai East, 928 m, 15 28 23.1234 N, 99 0 37.908 E, 14 Jan. 2011, Watthana, S. 3672 (fr.) (QBG)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFADFFDFFF11F8A9FCC60AA0.taxon	materials_examined	LAOS: Vientiane Province, Hin Heup, Kon Phouk Village, along main road, near Nam Lik River, 200 m, 23 Aug. 1999, Maxwell, J. F. 99 - 169 (fl.) (CMU); Champasak Province, Khong District, Sadam Island, near Hang Don Sadam Village, Mekong River, 60 m, 14 Oct. 1997, Maxwell, J. F. 97 - 1175 (fl.) (CMU) TIMOR: Leschenault s. n. (fr.) (K) VIETNAM: Amamite, Cay Dong Tien, 18 Oct. 1919, Poilane, 606 (fl.) (L); Cochinchina, Apr. 1893, Pierre, L. s. n. (fr.) (K); Cochinchina, 1862 – 1866, le Thorel, M. s. n. (fr.) (K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF86FF89FF11FEEBFD180CB8.taxon	description	J. As. Soc. Beng. ii, 66: 233. 1897 (description) Type: — MYANMAR, Taong Doung, Wallich 5832. Holotype K (K 000264174!); isotype BM (BM 000958801!) M. hymenocarpum Wallich ex Wight & Arn. nom. nud. in Wight, R. A. & Arnott, G. A. Prodromus Florae Peninsulae Indiae Orientalis: 283. 1834. Based on Caesalpinia hymenocarpa Wallich cat. n. 5832 (nomen) 1832. Synonym. Caesalpinia hymenocarpa (Wight & Arn. ex Prain) Hattink. Reinwardtia 9 (1): 35. 1974. Synonym. Mezoneuron laoticum Gagnepin. Not. Syst. 2: 208. 1911. Lectotype: — LAOS, Me Kong, Steung Streng (= Stung Treng), 1866 – 68, Thorel s. n., P (P 02142674!), Hattink 1974. Woody climber or scandent shrub to 30 m in length. Stems with scattered recurved prickles, older stems with spine-tipped corky tubercles; glabrous to sparsely whitish tomentose. Stipules persistent, oblong to ellipsoid, sometimes with acuminate apex, 1 – 2 × 1 – 5 mm, subglabrous. Leaves with 6 – 9 (– 12) pairs pinnae; (4 –) 6 – 9 (– 11) pairs leaflets per pinna; petiole 2.4 – 9 cm; leaf rhachis (12 –) 15.5 – 30 (– 33) cm, with recurved prickles paired at pinnae insertion points and sometimes scattered on the internodes; pinnae opposite or subopposite, 4 – 10 (– 12.5) cm, pinna rhachis sometimes with recurved prickles paired at leaflet insertion points and a sometimes scattered on the internodes, with terminal prickle 1 – 3 mm; leaf rhachis, petiole, and pinna rhachis sparsely to moderately whitish or pale orange tomentose. Leaflets opposite to subopposite, terminal leaflets obovate or elliptic, base oblique, apex rounded to retuse, 1 – 3.8 × 0.5 – 2.2 cm; lateral leaflets oblong to elliptic to slightly obovate, sometimes oblique, especially at base, apex rounded to retuse, 1.0 – 3.3 (– 4.0) × 0.6 – 2.0 cm; all leaflets with upper surface glabrous or with sparse, semi-appressed, or tomentose, whitish hairs, on base of lamina and midrib, or on entire surface, lower surface usually with sparse, semi-appressed, or tomentose, whitish hairs, these sometimes only on basal midrib, occasionally glabrous. Inflorescence a panicle or raceme, axillary, supra-axillary, or terminal, to 60 cm; sometimes with few recurved prickles on basal parts of axes; axes and pedicels sparsely to densely white or golden to pale orange tomentose; pedicels 6 – 28 mm, articulated 2 – 3 mm below the hypanthium, indumentum denser at articulation. Bracts sometimes present, caducous, triangular, ca. 1 × 1 mm, subglabrous to sparsely tomentose; bracteoles caducous, elliptic-lanceolate, tapering to an acute apex, narrowing at base, 1 – 5 × 1 – 2 mm, moderately to densely golden or orange tomentose on both surfaces. Flowers with an asymmetric hypanthium 2 – 4 × 6 – 12 mm, sparsely to moderately golden tomentose; lower lobe highly cucullate, 9 – 15 × 4 – 6 mm, other lobes 4 – 7 × 3 – 6 mm; outer surface of lobes moderately to densely golden to orange tomentose, inner surface of lower lobe moderately to densely tomentose, inner surface of other lobes glabrous or with few hairs, sometimes denser at margins. Median petal coriaceous, the blade broadly elliptic to oblong, and distinct narrow claw, strongly reflexed backwards, with transverse flap of tissue between claw and blade, this bilobed or dentate; blade 4 – 5 × 4 – 14 mm long, claw [3 –] 4 – 5 mm × 1.5 – 2 mm, the whole 7 – 10 mm long; few hairs along margins of claw, and along margin of the flap of tissue. Upper lateral petals orbicular to obovate, 7 – 15 × 6 – 13 mm, including claw 1 – 2 mm; glabrous apart from few hairs on inner surface of claw. Lower lateral petals orbicular to obovate, 7 – 15 × 9 – 13 mm, including claw 1 – 2 mm long; glabrous apart from few hairs on inner surface of claw. Stamen filaments [7 –] 9 – 18 mm long, vexillary stamen shorter than the others, glabrous or nearly so, other 9 stamens moderately to densely orange villous on the basal ½ on inner surface; anthers ca. 3 mm long. Ovary ca. 5 – 8 mm long, style ca. 7 – 14 mm long, both glabrous, stigma broadly funnel-shaped, ca. 1 mm wide, the rim papillate. Fruit oblong-elliptic, base cuneate, sometimes oblique, apex acute, sometimes oblique, (5.5 –) 6.5 – 17 × 1.9 – 4.6, wing 4 – 16 mm wide; fruit surface glabrous; venation reticulate, raised and visible on the fruit body, anastomosing and slightly visible on the wing; hypanthium semi-persistent. Seeds 2 – 6 per fruit, elliptic to orbicular, often clearly visible in outline when mature as concave or convex shape in the fruit, adjacent to and perpendicular to the wing, pale brown.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF86FF89FF11FEEBFD180CB8.taxon	distribution	Distribution: — India, Sri Lanka, Myanmar, Thailand, Vietnam; Malesia: Java, Lesser Sunda Islands, Philippines; Australia. (Fig. 12). Habitat and ecology: — Deciduous forest, evergreen forest, primary or secondary; open disturbed thicket, roadside; granite, sandstone, shale or limestone bedrock. Elevation to 850 m. Occurs in regions that experience a dry season. 22 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK Phenology: — India, Sri Lanka: flowering September – November, fruiting December. Myanmar: flowering August, fruiting October. Thailand, Laos, Vietnam: flowering July – January, fruiting August – May. Malesia: flowering April, fruiting May – November. Australia: flowering time unknown, fruiting April – August. Preliminary conservation assessment: — Extent of Occurrence 10,239,581.796 km 2 = LC; Area of Occupancy 220.000 km 2 = EN. Category assigned: — Least Concern (LC).	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF8AFF84FF11FBA3FE550EFD.taxon	description	Flora of the Hawaiian Islands. W. F. Hillebrand, London, Williams & Norgate: 110 – 111. 1888. Type: — HAWAII, Kauai, Waimea, Kaawaloa, Mann, H. & Brigham, W. T. s. n. Holotype CU; isotype BISH (BISH 1009127, image!). Basionym. Caesalpinia kavaiensis H. Mann. Proc. Amer. Acad. Arts vii: 164. 1867. Tree to ca. 12 m high. Stems unarmed; sparsely to densely whitish to orange tomentose. Stipules caducous; triangular; ca. 1 – 2 × 1 mm, partially tomentose. Leaves with 2 – 4 pairs pinnae; 4 – 5 [– 8] pairs leaflets per pinna; petioles 2 – 9.5 cm; leaf rhachis 5 – 22 cm, unarmed; pinnae opposite or subopposite, 3.5 – 11 cm, the basal pair shorter than the others; leaf rhachis, petiole, and pinna rhachis sparsely to densely orange tomentose or villous, petiolules sometimes more densely hairy than rhachis. Leaflets opposite to alternate; the terminal leaflets elliptic to oblong, base strongly oblique, apex retuse to emarginate; 2 – 5 × 1 – 2 cm; lateral leaflets elliptic to oblong, base strongly oblique, apex retuse to emarginate; 1.5 – 4.5 × 0.9 – 1.9 cm; all leaflets with upper surface glabrous, lower surface with a dense tuft of villous hairs at the base, the hairs extending part way or all the way along the midvein, otherwise glabrous; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, veins slightly visible above, clearly visible below. Inflorescence a raceme to ca. 50 cm, usually terminal, occasionally axillary or arising from the stem independently from the leaves; the axes glabrous, or subglabrous, with hairs only at base of pedicels, or sparsely to moderately orange tomentose; pedicels 23 – 50 mm, articulated 5 – 10 (– 15 in fruit) mm below hypanthium, glabrous, or occasionally with few hairs at articulation. Bracts not seen, apparently absent; bracteoles caducous, triangular, the apex acuminate, ca. 4 – 5 × 2 mm; densely pale orange tomentose at base and sides, glabrescent towards apex. Flowers with a hypanthium 8 – 11 × 3 – 4 mm; lower calyx lobe 13 – 14 × 6 – 7 mm; other lobes 9 – 11 × 5 – 7 mm; hypanthium and calyx lobes glabrous; veins clearly visible in calyx lobes. Median petal broadly obovate, tapering to a broad claw, strongly laterally inrolled, 8 – 10 × 7 – 12 mm, blade ca. 4 – 5 × 8 – 12 mm, claw ca. 1 – 5 × 1 – 4 mm, glabrous or with few hairs on claw. Upper lateral petals elliptic to suborbicular, ca. 10 – 12 × 8 – 12 mm, including 26 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 27 claw ca. 1 mm long, glabrous. Lower laterals petals elliptic, tapering to a claw, ca. 12 – 14 × 6 – 9 mm, including claw ca. 2 – 3 mm long, glabrous; veins prominent on all petals. Stamen filaments flattened, 10 – 22 mm long, with translucent orange hairs on the basal ½, on both surfaces, or sometimes on margins only; anthers ca. 2 mm long. Ovary glabrous, ca. 6 – 9 mm long, style ca. 12 – 20 mm long; style tapering to a narrowly cupular stigma, this very slightly papillate. Fruit ligneous, usually glaucous with white waxy coating; broadly elliptic to oblong, base obliquely attenuate to cuneate, apex retuse, or the wing extending to form a beak; 6.5 – 12 × 4 – 6 cm, wing 5 – 12 mm wide; glabrous; venation reticulate, faintly visible or slightly raised on fruit body, not visible on the wing; hypanthium persistent. Seeds 2 – 4; pale brown, transversely arranged, ovoid, flattened, 1.8 – 2 cm long, 1.4 – 1.6 cm wide; not visible individually in outline on fruit exterior. (Fig. 15).	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF8AFF84FF11FBA3FE550EFD.taxon	distribution	Distribution: — Endemic to Hawaii, islands of Hawaii, O’ahu and Kaua’i. (Fig. 16). Habitat and ecology: — Dry hills; dry and moist forest; degraded native vegetation, mostly Aleurites lowland mesic forest. Elevation 80 – 920 m. Phenology: — Flowering and fruiting throughout the year. Conservation Assessment: — Critically Endangered (CR) ver. 3.1 (IUCN 2015). Mezoneuron kauaiense is endemic to the islands of Hawaii and Oahu. Eight subpopulations are known, totalling 60 individuals, however, many of these are senescent and probably non-reproductive. Threats include pigs, cattle, deer, goats, introduced plants, rats, fire, volcanic eruptions, sheep, black twig borer and collection by humans (IUCN 2015). Field notes: — Petals red.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD3FFDDFF11FA36FE9D0F20.taxon	description	A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 65 Kalimantan, Coal mining company KP, Sangatta, Tanjung Bara area, 0 5 N, 117 4 E, 14 Apr. 1995, Kessler et al. 1175 (fr.) (K, L); East Kalimantan, road from Desa Batu Kajang to Penajam, 200 m, 1 49 S, 115 56 E, 16 Nov. 1996, Kessler et al. PK 1992 (fl.) (K, L); East Kalimantan, Sabulu (5 km point), 50 – 100 m, 117 E, 0 10 S, 1 Dec. 1980, Kato, M., Wiriadinata, H. 7025 (fr.) (L); East Kalimantan, Tanah Grogot, Kec. Batu Kajang, Desa Kasungai, 400 m, 29 Nov. 1979, Ma’roef, A. 279 (fr.) (K); Elopura, Sandakan Mile 15, Labuk Rd, 15 Mar. 1949, Cuadra, A. A 2210 (fr.) (K); Kalimantan Selatan, Djaro Dam, c. 10 km NE of Muara Uja, 80 m, 1 50 S, 115 40 E, 14 Nov. 1971, Vogel, E. F. 794 (fr.) (K, L); Kalimantan Timur, 30 km north of Sebulu, 50 – 100 m, 28 Dec. 1978, Murata, G., Iwatsuki, K., Kato, M., Mogea, Y. P. 554 (fr.) (L); Kalimantan, Berau, Inhutani I concession area, Labanan, along logging road to kampong Merasa, 150 m, 0 51 N, 117 10 E, 4 Mar. 2000, Kessler, P. J. A. et al. B 1584 (fl.) (K, L); Kalimantan, East Kutai Reserve, vicinity of Sengata and Mentoko Rivers, below 300 m, 6 Jul. 1979, Leighton, M. 758 (fr.) (L); Kalimantan, Timur, Sepaku, P. T. ITCI, near plot 76 - 3 B, 50 m, 1 S, 117 E, 03 Nov. 1990, Ambri & Arifin AA 35 (fr.) (K, L); Kampung Melangkap Tomis, 4 km dari Melangkap Tomis, Pisoh, 4 May 1996, Lugas, L. 2041 (fr.) (K); Kayoep, 27 May 1908, Winkler, H. 2183 (fr.) (L); Kinabatangan District, Malui river valley, E. S. E. of kpg. Enteleben and S. W. of bkt. Tawai summit, 75 m, 5 31 N, 117 07 E, 7 Jul. 1984, Sands, M. J. S. 3846 (fr.) (K); Labuan, Jan. 1886, Hullett, R. W. s. n. (fl., fr.) (K); Lahad Datu, Segama Upak, 60 m, 8 Aug. 1961, Chai, M. SAN 26010 (fr.) (L); Loa Haur, west of Samarinda, 60 m, 9 May 1952, Kostermans 6808 (fl.) (L); Mului, 1 26 S, 115 58 E, 15 Nov. 2005, Raes, N. et. al. 562 (fr.) (K); N. Borneo, Sandakan, Labuk Road, Mile 3, 25 Jun. 1957, Sinclair, J. 9286 (fr.) (K, L); Papar West Coast Interior District, Kuada Papar, 29 Apr. 1948, Cuadra, A. 1423 (fr.) (K); Pengkalan Madai, Lahad Datu, 15 m, 23 Jan. 1953, Rivera A 1908 (fl.) (K, L); Ranau District, Kampung Perancangan, 15 Sep. 1929, Tadong, D. 147 (fl.) (K); Ranau, Ulu Tungud Forest Reserve, Gunung Monkobo Expedition, 471 m, 5 49 50 N, 117 00 01 E, 25 Jul. 2005, Saw, L. G. et. al. SAN 146689 (fr.) (K); Sabah, Beaufort District, Beaufort Hill, 250 m, 5 22 N, 115 45 W, 6 May 1984, Beaman, J. H. 9642 (fr.) (K); Sabah, Danum Valley, 150 m, 21 Mar. 1987, Argent, G. et. al. 1987162 (fl., fr.) (E, K); Sabah, Kampung Melangkap Tomis, 1 km dari Melangkap Tomis, 16 Dec. 1995, Lugas, L. 1514 (fl.) (K); Sabah, Kampung Melangkap Tomis, 5 km dari Melangkap Tomis, 16 Nov. 1995, Lugas, L. 1284 (fl.) (K); Sabah, Keningau District, 05 Mar. 1988, Mantor, A. SAN 121998 (fr.) (K); Sabah, Kota Marudu District, Kampung Serinsim 3 km ke arah barat dari serinsim, 6 Mar. 1995, Bakia, K. 295 (fl.) (K); Sabah, Nabawan District, KM 55 Jalan Nabawan / Sepulut, 22 Feb. 1990, s. coll., SAN 128389 (fr.) (L); Sabah, Pandewan District, 17 Feb. 1987, Mantor, A. SAN 119992 (fl.) (K); Sabah, Pensiangan, 24 Jan. 1994, Mantor, A. SAN 136905 (fr.) (K); Sabah, Ranau District, Paka, 8 Apr. 1988, Amin & Suin SAN 121584 (fr.) (K, L); Sabah, Sandakan, Sungai Wonod, Telupid, along main road between mile 8 & 7, 16 Mar. 1974, Aban & Saikeh s. n., SAN 79391 (fr.) (L); Sabah, Sipitang District, Mandulong Forest SFI area, 960 m, 3 Oct. 1992, Mandani, L. & Patrick, L. s. n., SAN 134021 (fl.) (K, L); Sabah, Sungai Meliau, Beluran, Sigin et al. s. n., SAN 99705 (fr.) (K); Sabah, Sungai Meliau, Beluran, Sigin et al. s. n., SAN 99706 (fl.) (L); Sabah, Tawai, 10 Apr. 1994, Cheksum Tawan, Good, Ramlan & Zainuddin CST 260 (fl.) (K, L); Sandakan, Compartment 6, Sepilok Forest Reserve, 15 miles W of Sandakan, 23 Feb. 1956, Wood, G. H. S. s. n., SAN 15474 (fr.) (L); Sandakan, Feb. – Mar. 1920, Wood, D. D., 7 62 (fr.) (K); Sandakan, Mile 17, Lebuk Road, 85 ft., 26 Mar. 1963, Elleh, S. s. n. S. 35590 (fl.) (K); Sandakan, Sibuga Forest Reserve, 2 Jun. 1959, Meijer, W. s. n. S AN 19209 (fr.) (K, L); Sapagaya Forest Reserve, 5 m, 13 Jun. 1939, Kadir & Enggoh 10333 (fr.) (K); Sarawak, 1929, Clemens J. & M. S. 21295 (fr.) (K); Sarawak, Baram District, 1894, Hose, C. 70 (fr.) (BM, K, L, PNH); E. Borneo, 13 Jul. 1954, Kostermans, A. 9722 (fl., fr.) (K, L); Sarawak, Baram District, 4 th Division, Sg. Chipidi, Btg. Tinjar, 7 Aug. 1974, Chai, P. S. 34663 (fr.) (L); Sarawak, Bkt. Sepako kiri, Ulu Balaga, Kapit, 7 th Division, 6 Nov. 1981, Ismawi, O. et al. S 43868 (fr.) (L); Sarawak, Bkt. Sepako kiri, Ulu Belaga, Kapit, 7 th Division, 6 Nov. 1981, Ismawi, O. et. al. S 43869 (fr.) (K, L); Sarawak, Bukit Raya, Kapit, 3 rd Division, 7 Feb. 1971, Ismawi, O. S 26774 (fl.) (K, L); Sarawak, foot of Gunung Api, Sungai Melinau, Mulu National Park, 5 May 1985, Mohtar, A., Othman, I. S 49575 (fr.) (L, MO); Sarawak, foot of Hose Mountains, Ulu sg. Merirai, Batang Balleh, 7 th Division, 600 m, 25 Mar. 1987, Ching, Y. P. S 53684 (fl.) (K, L); Sarawak, Gunung Mulu National Park, Camp 5, 4 5 N, 114 54 E, 1977 – 78, Hansen, C. 208 (fr.) (K, L); Sarawak, Kapit, Upper Rejang River, Clemens, J. & M. S. 21293 (fr.) (K); Ulu Segama, Danum Valley, 150 m, 21 Mar. 1987, Argent, G., Aik, J., Kasil, K. 1987162 (fl., fr.) (K); Brunei, Belait, Labi, Sungai Rampayoh, 30 m, 4 21 N, 114 28 S, 12 Jan. 1994, Coode, M. J. E. 7817 (fl.) (K, L); Bt. Teraja M. 19 Hillside, 300 ft, Ashton, P. S. 989 (fr.) (st.) (K, L); Sabah, Papar District, Kampung Wolit, 1 km from Crocker Range Park, 1997, Jinaun, J. 79 (fr.) (K); Kalimantan Timur, near Malinau, 20 m, 3 36 N, 116 40 E, 2 Jul. 1981, Geesink, R. 8919 (fr.) (L); Sabah, Danum Valley and environs, 22 Jun. 1989, Ridsdale, C. E. 2134 (fl., fr.) (L) 66 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD3FFDDFF11FA36FE9D0F20.taxon	materials_examined	MOLUCCAS: Halmahera, Gunung Jailolo, 250 m, 1 4 N, 127 27 E, 14 Oct. 1974, Vogel, E. F. 3450 (fr.) (L) NEW GUINEA: Morobe Province: Herzog Mountains, Gabensis logging track, 685 m, 06 45 S, 146 45 E, 13 Jul. 1991, Takeuchi, W. 7168 (fl.) (F, L, MO, NY); Morobe District, Busu River, 100 m, 6 30 S, 147 10 E, 21 Jun. 1965, Gillison, A. N. G. 22252 (fl.) (K, L, NY, US); Palawan, Taytay municipality, Lake Manguao (Danao), valley stream leading nito NNW bay of lake, 60 – 80 m, 7 Apr. 1984, Ridsdale, C. E. 354 (fr.) (L) PHILIPPINES: 1903 / 4, Hallier, H. 488 (fr.) (NY); Antipolo, 28 Mar. 1891, Loher, A. 2202 (fr.) (K); Luzon, Bataan Province, Morong Municipality, Bgy. Mabajo, road to Natib Mountain, Subic Bay, 150 m, 11 Feb. 1991, Soejarto, D. D., Fernando, O., Madulid, D. A. 7519 (fl.) (MO, NY); Basilan, Jan. 1904, Hallier, H. 4488 (fl.) (L); Panay Island, Miagas, Mar. 1886, Villar, F. 2667 (fr.) (K); Cabadbaran (Mt. Urdaneta), Prov. of Agusan, Mindanao, Jul. 1912, Elmer A. D. E. 13386 (fl.) (BM, F, K, L, NY); Cebu, Tagbao, 450 m, 22 Apr. 1971, s. coll. (University of San Carlos) 149 (fr.) (L); Central Luzon, Bataan Province, Mt. Mariveles, Lamao River, 9 Mar. 1904, Williams, R. 701 (fl., fr.) (K, NY); Island of Paragua, Point Separation, 20 Feb. 1903, Merrill, E. D. 805 (fr.) (K); Los Banos (Mt. Maquiling), Prov. of Laguna, Luzon, Jun. – Jul. 1917, Elmer, A. D. E. 17983 (fl.) (BM, F, K, L, NY); Luzon, Albay Province, 1841, Cuming, H. 1193 (fl.) (K, L); Luzon, Angat, Bulacan Province, Sept. 1913, Merrill 294 (fl.) (BM, F, K, MO, NY); Luzon, Bataan Province, Morong Municip., Natib Mt., 300 m, 10 Feb. 1991, Soejarto, D. D., Fernando, O., Madulid, D. A. 7501 (fl.) (F); Luzon, Benguet Province, Baguio, Mar. 1907, Elmer, A. D. E. 8466 (fl.) (K, L, NY); Luzon, Camarines Sur, Mt. Isarog, Naga City, 11 Jun. 1992, Barbon, Romero & Fuentes s. n., PPI 8398 (fr.) (L); Luzon, Isabela Province, San Mariano, Feb. to Mar. 1926, Ramos, M.; Edano, G. Bureau of Science no. 47106 (fl.) (NY); Luzon, Laguna Prov., Apr. 1908, Robinson, C. B. 6095 (fr.) (MO); Luzon, Laguna Province, Los Banos, Apr. 1906, Elmer, A. D. E. 8138 (fr.) (K); Luzon, Mar. 1909, Ramos, M. Bureau of Science number 7737 (fl.) (L); Luzon, Merrill 875, (fl.) (BM, F, K, L, NY, US); Luzon, Mt. Bulusan, Sorsogon Prov. 2789, Jul. – Aug. 1947, Sulit, M. D. 2789, (fr.) (MO); Luzon, Pampanga Province, Arayat, Mar. 1903, Merrill, E. D. 1438 (fl., fr.) (K); Luzon, Province of Laguna, Apr. 1910, Tamensis, F. 11955 (fr.) (F); Luzon, Province of Laguna, Los Banos, Elmer, A. D. E. 17983 (fl.) (NY); Luzon, Province of Rizal, Ramos, Jan. 1914, Merrill 2089 (fr.) (MO); Luzon, Province of Rizal, Sept. 1905, Ahern’s collector 3370 (fl.) (MO); Luzon, Rizal Province, Antipolo, Mar. 1903, Merrill, E. D. 1711 (fr.) (K); Luzon, Sorsogon Prov., Mt. Bulusan, Jul. – Aug. 1947, Sulit, M. D. 2789 (fr.) (MO); Magallanes (Mt. Giting-Giting), Province of Capiz, Island of Sibuyan, Apr. 1910, s. coll. 12187 (fr.) (BM, F, K, L); Mindanao, District of Davao, Mar. 1904, Copeland, E. B. 470 (fl.) (K, NY); Mindoro Province, Puerto Galera, Aguada Mountain, 7 May 952, Santos, J. V. 5310 (fr.) (L); Mindoro, Paluan, Apr. 1921, Ramos, M. s. n. (Bureau of Science no. 39656) (fr.) (L); Palauan, Mindoro, 1903, Merrill 956 (fr.) (NY); Palawan, Municipality Bataraza, Sandoval, 100 m, 9 51 N, 118 37 E, 5 – 11 Apr. 1989, Soejarto, D. D., Reynoso, E., Sagcal, E., Edrada, R. 6532 (fl., fr.) (F, K); Palawan, Quezon Municipality, Bgy Tuangan, Ladayon village, c. 30 km S of Quezon on road Quezon — Ransang, 60 m, 09 07 N, 117 54 E, 22 Jun. 1994, Soejarto, D. D., Gaerlan, F., Sagcal, E., Fernando, O. 8525 (fr.) (F, L); Palawan, Quezon Municipality, Bgy. Tuangan, Ladayon village, 30 km S of Quezon on road Quezon – Ransang, 60 m, 9 7 N, 117 54 E, 24 Jan. 1995, Soejarto, F., Gaerlan, F., Sagcal, E., Fernando, O. 8525 (fr.) (L); Penablanca, Cagayan Province, Mar. – Apr. 1929, Ramos, M. 146 (fr.) (NY); Visayas, Antique Province, Mt. Baloy, Brgy., San Agustin, Valderama, 670 m, 11 0 N, 122 8 E, Garcia, H., Romero E. 15768 (fr.) (L) SULAWESI: Soroako-Wasuponda Road, km 19, 2 15 S, 121 45 E, 7 Jul., van Balgooy 3918 (fr.) (K, L, MO); Bolaang, Mongondoro, Jul. 1917, Kaudern, W. 254 (fr.) (L, NY); Selatan, Soroako, S. shore of Lake Matano, 400 m, 2 32 S, 121 19 E, 13 Jun. 1979, Vogel, E. F. 5750 (fr.) (K, L, US)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD3FFDDFF11FA36FE9D0F20.taxon	description	SUMATRA: Teysmann, J. E. s. n. (L)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD3FFDDFF11FA36FE9D0F20.taxon	materials_examined	VIETNAM: Tonkin, Aug. 1885, Balansa, B. 1298 (fr.); Annam, Tourane, 1 Jun. 1920, Poilane, M. 1452 (fl.) (K); Tonkin, 19 Jun. 1884, Bon, R. P. 2695 (fr.) (L); Annam, Tourane, 100 km south of Hue, May Jul 1927, Clemens, J. & M. S. 3167 (fl.) (K, NY, US); Hao Binh Province, Yen Thuy District, Com Village, 40 m, 11 Dec. 2000, Soejarto, D. D. & Cuong, N. M. 11714 (fr.) (MO); Tonkin, Kau Nga Shan, Tien-Yen, 1 – 9 Jan. 1937, Tsang, W. T. 27463 (st.) (E, K); Tonkin, Ho Yung Shan, Tien-Yen, 13 Oct. – 22 Nov. 1940, Tsang, W. T. 30749 (fl.) (K) (fr.) (E); Tonkin, Tu-Phap, 1887, Balansa, B. 2149 (fl., fr.) (P); Tonkin, Kien Khe, Hong Hain, 14 May 1883, Bon, R. P. 2132 (fl.) (P)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF9CFF8EFF11FB82FDE60A73.taxon	description	Phillip. J. Sci. 4. Bot.: 268. 1907. Type: — PHILIPPINES, Luzon, Cavanille’s plate excl. leaves, which are bauhinioid, Epitype (here designated): — PHILIPPINES, Luzon, Albay Province, Cuming 1193, K (K 000980294!); isoepitypes K (K 000980295!), L (L 0651597!). Basionym: Bauhinia? latisiliqua Cav. Icones et Descriptiones Plantarum vol. 5. 1799. Synonym. Caesalpinia latisiliqua (Cav.) Hattink. Reinwardtia 9 (1): 37. 1974. Synonym. Caesalpinia torquata Blanco. Fl. Filip.: 336. 1837. Type: lost fide Hattink 1974. Synonym. Mezoneurum procumbens Blanco. Fl. Filip. II: 235. 1845. Type: lost fide Hattink 1974. Synonym. Mezoneuron rubrum Merr. Publ. Bur. Sci. Gov. Lab. 6: 7. 1904. Type: — PHILIPPINES, Paragua (= Palawan), Point Separation, 20 Feb. 1903, Merrill 805, K (K 000264162!), PNH †, US. Synonym. Mezoneuron platycarpum Merr. Phillip. J. Sc., Bot. 11: 85. 1916. Type: — BORNEO, Sarawak, Baram District, Miri River, Jan. 1895, Hose 70, BM!, K (K 000264166!), L (L 0018788!), PNH. Synonym. Mezoneuron cabadbarense Elmer. Leafl. Phillip. Bot. 10: 3757. 1939. Type: — PHILIPPINES, Mindanao, Cabadbaran, Jul. 1912, Elmer 13386, BM!, BO, F!, GH, K (K 000264161!), L!, P, PNH, NY (0004516!), U. Synonym. Mezoneuron balansae Prain ex Gagnep. Fl. Gén. Indoch. 2: 198. 1913. Lectotype: — INDO-CHINA: Tonkin, pres de Luang Yen, Aug. 1885, Balansa 1298, P (barcode unknown); isolectotypes K (K 000264169!), L (L 0018786!, L 0018787!), Hattink 1974. Synonym. Mezoneuron keo Gagnepin. Bull. Mus. Hist. Nat. Par. 2: 318. 1952. Type: — INDO-CHINA, Annam, Tourane, Jun. 1920, Poilane 1452. Holotype P; isotypes K (K 000264168!), L (L 0018785!). Synonym. Mezoneuron oxyphyllum Gagnepin. Bull. Mus. Hist. Nat. Par. 2: 319. 1952. Type: — VIETNAM, Annam, N’ghe-an, reserve for. de Co-ba, Fleury, F. for Chevalier, 32512. Holotype P (P 02142678!); isotype P (P 02142677!). Woody climber or scandent shrub to 20 m in length. Stems with scattered recurved prickles, sometimes with corky spine-tipped tubercles on older stems; glabrous, or sparsely whitish to orange tomentose. Stipules oblong or linear, 16 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK sometimes with acuminate apex, or triangular, 1 – 2 × 1 – 5 mm, glabrous or partially tomentose. Leaves with 3 – 9 pairs pinnae; 3 – 8 pairs leaflets per pinna; petiole 3.5 – 12 cm; leaf rhachis 12 – 50 cm, recurved prickles usually present, paired at pinna insertion points, and sometimes also scattered on the internodes; pinnae (4 –) 6 – 12 (– 13) cm; pinna rhachis sometimes with recurved prickles paired at leaflet insertion points and sometimes also scattered on the internodes; leaf rhachis, petiole, and pinna rhachis glabrous, or sparsely, occasionally moderately, whitish to orange tomentose. Leaflets alternate to subopposite; the terminal leaflets 2 – 7.8 × (1 –) 1.9 – 5.2 cm, obovate, base oblique, apex rounded to retuse with small mucro; lateral leaflets 1.5 – 7.5 × 0.7 – 3.8 (– 4.6) cm, elliptic to oblong or slightly obovate, base oblique, apex rounded to retuse, mucronulate; all leaflets with upper surface glabrous or very sparsely tomentose, sometimes hairs only on margins, lower surface glabrous, or sparsely to moderately tomentose, or with semi-appressed hairs, sometimes hairs only at base or margins; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, veins visible on both surfaces, distinctly raised below, midvein sunken above, raised below. Inflorescence a raceme or panicle, terminal or axillary, up to 100 cm long, the axis to 0.6 cm diameter at base, axes sparsely to moderately whitish or golden or orange tomentose; recurved prickles occasionally present on basal portion of peduncle; pedicels (8 –) 10 – 25 mm, not articulated; sparsely to densely pale orange to golden tomentose, or glabrous. Bracts caducous, semicircular, 1 × 1 – 3 mm, sparsely tomentose; bracteoles caducous, broadly elliptic, acuminate, 1.5 – 3 × 0.5 – 1 mm, margins or entire outer surface orange tomentose, apex glabrous. Flowers with an asymmetric hypanthium, 4 – 6 × (6 –) 8 – 12 (– 15) mm; lower calyx lobe highly cucullate, [7 –] 11 – 19 × 6 – 8 mm; other lobes elliptic to ovate, [4 –] 5 – 10 × 3 – 5 mm; hypanthium and calyx both glabrous to moderately to densely golden or orange tomentose. Median petal strongly bilobed, to ⅓ or ½ of length, inrolled laterally, (8 –) 11 – 17 × (8 –) 11 – 13; glabrous. Upper lateral petals obdeltate, apex rounded or acuminate, 2 – 5 × 2 – 3 mm, glabrous. Lower lateral petals obdeltate, apex rounded or acuminate, 2 – 5 × 2 – 3 mm, glabrous. Stamen filaments flattened, (8 –) 11 – 31 mm, sparsely to densely orange villous, along the basal ¼ – ⅔, on inner surface or on margins only; anthers 1 – 3 mm. Ovary 4 – 10 mm long, style 8 – 20 mm long, ovary and style glabrous, or partially, or sparsely to densely golden or orange sericeous, hairs sparser towards apex of style; stigma laterally positioned, narrowly cupular 0.25 – 1 mm wide, the rim papillate. Fruit: pedicel in fruit to 30 mm, sometimes with a swelling 8 – 10 mm below hypanthium; elliptic, base oblique, apex with wing extending to form a beak, or occasionally emarginate at apex of midvein; wider part of the persistent, asymmetric hypanthium below winged side of fruit; 8.5 – 16.5 × 3 – 5.5, wing 7 – 15 mm wide; fruit surface glabrous, or sparsely to moderately orange tomentose, sometimes partially; venation reticulate, raised and visible on fruit body, anastomosing and slightly visible on the wing. Seeds 6 – 9 (– 13) per fruit; elliptic, visible in outline on fruit exterior when mature, glossy, dark brown, ca. 9 – 12 × 6 – 7 × 1 mm.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF9CFF8EFF11FB82FDE60A73.taxon	distribution	Distribution: — Vietnam, Borneo, Philippines, Sulawesi, Papua New Guinea. (Fig. 8). Habitat and ecology: — Foothill forest; lowland rainforest; alluvial forest; riparian forest; swamp forest; primary and secondary forest; roadsides in open places; coastal vegetation; substrate sandstone, limestone, or shale. Elevation 0 – 1500 m. A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 17 Phenology: — Vietnam: flowering May – June, fruiting June – December. Malesia: flowering and fruiting throughout the year. Preliminary conservation assessment: — Extent of Occurrence 6,601,692.749 km 2 = LC; Area of Occupancy 388.000 km 2 = EN. Category Assigned: — Least Concern (LC). Taxonomic notes: — The type of M. latisiliquum is an illustration of a specimen that bears the leaves of a species of Bauhinia. An epitype is here selected with a leaf representative of M. latisiliquum. The epitype also bears flowers, that are absent from the type illustration. The ovary indumentum of this species is very variable, ranging from glabrous to partially to densely hairy. The leaflets also vary considerably in size, from 1.5 to nearly 8 cm in length, and 1 to ca. 5 cm in width. The patterns of variation correlate with locality (Fig 8).	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD1FFDCFF11F8BAFCD508B8.taxon	materials_examined	PHILIPPINES: Biliran, Jun. 1914, McGregor, R. C. 18672 (fl.) (NY); Mindanao, Mati, Davao Prov., Mar. – Apr. 1927, Ramos, M., Edano, G. 49280 (fl.) (NY); Pinamalayan, Mindoro, Merrill s. n., For. Bur. no. 479 (fr.) (L); A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 67 Mindoro, Mt Yagaw (SE slope), 16 Aug. 1953, Conklin, H. C. 398 (fl.) (L); Mindoro, Bulalacao, Aug. – Sep. 1906, Bermejos, J. s. n. (Bureau of Science no. 1514) (fr.) (K, NY, US)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF8BFF86FF11FE7BFD3E0C5B.taxon	description	Phillip. J. Sc., Bot. 3: 232. 1908. Type: — PHILIPPINES, Mindoro, Pinamalayan, Oct. 1906, For. Bur. Merritt 5383 (PNH †) Neotype (here designated): — PHILIPPINES, Mindoro, Pinamalayan, 20 Oct. 1906, Merritt s. n., For. Bur. no. 479, L (L 0651589!) Synonym. Mezoneuron mindorense Merr. var. inermis Merr. Phillip. J. Sci., Bot. 232. 1908. Type: — PHILIPPINES: Mindoro, Bulalacao, 8 Sep. 1906, Bermejos 1514. Syntypes BO, GH, K, (K 000264160!), PNH, NY (00004517!), US. Climber, height unknown. Stems with scattered small recurved prickles to 3 mm; glabrous. Stipules linear to oblong, the apex acuminate, or triangular, ca. 1 × 1 – 2 mm, glabrous. Leaves with (6 –) 10 – 13 pairs pinnae; 8 – 12 pairs leaflets per pinna; petiole 2.8 – 5.5 cm; leaf rhachis 11 – 30 cm, with recurved prickles paired at pinna insertion points, sometimes in threes, and sometimes scattered on the internodes, or leaf unarmed; pinnae 4.5 – 8 cm; leaf and pinna rhachis both channelled or ridged on the upper side, with sparse whitish to pale orange tomentose hairs, glabrescent, the hairs mostly on the upper (adaxial) side, petiole with sparse whitish tomentose hairs on the upper (adaxial) side. Leaflets opposite or subopposite; terminal leaflets oblong to obovate, base sometimes oblique, apex rounded, obtuse, or retuse, mucronulate, 0.8 – 2.0 × 0.5 – 1.0 cm; lateral leaflets oblong, base sometimes oblique, apex rounded, obtuse or retuse, mucronulate, 1.0 – 2.2 × 0.4 – 0.9 cm; all leaflets with upper surface glabrous, lower surface glabrous or sometimes with few hairs on petiole and extending on to a small basal area of leaf blade; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, veins indistinct above, more clearly visible below. Inflorescence an axillary or terminal panicle; pedicels 9 – 14 mm, articulated 0 – 1 mm below hypanthium; all parts (axes, pedicels, calyx) sparsely orange tomentose, hairs short, on pedicels slightly denser around articulation. Bracts persistent, triangular, ca. 1 – 2 × 1 – 1.5 mm, glabrous, or tomentose on margins; bracteoles caducous, 3 – 4 × 0.5 – 1 mm, lanceolate, tapering to an acute apex, sparsely pubescent on inner surface, glabrous or very sparsely tomentose on outer surface. Flowers with a strongly asymmetric hypanthium, this wider on side of median petal, 1 – 2 mm × 5 – 8 mm; lower calyx lobe 6 × 4 mm, other lobes 4 – 6 × 2 – 3 mm. Median petal strongly reflexed backwards, 7 – 8 × 4 mm, blade obovate, 4 × 4 mm, claw oblong, 5 × 3 mm, with hairs along outer edges of claw, otherwise glabrous. Upper lateral petals obovate to suborbicular, ca. 8 × 7 mm, including claw 1 mm long. Lower lateral petals obovate to suborbicular, ca. 8 × 7 mm, including claw 1 mm long. Stamen filaments 12 – 13 mm long, orange tomentose on the basal ⅓ to ½, or along most of length; anther 1 – 1.5 mm long. Ovary 3 – 5 mm long, style ca. 10 mm long, both glabrous, stigma funnel-shaped, ca. 1 mm wide, the rim papillate. Fruit elliptic, base oblique, apex acute, wing usually arising basally ca. 2 – 10 mm along central vein, the central nerve extending to form a small mucro, 6.4 – 8 × 2 – 2.5 cm, wing 5 – 6 mm wide; fruit surface glabrous; venation reticulate, raised and visible on fruit body, anastomosing and not or slightly visible on the wing; hypanthium semi-persistent. Seeds 1 per fruit; elliptic, positioned centrally, adjacent to wing, visible in outline on fruit exterior when mature, ca. 7 mm diameter.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF8BFF86FF11FE7BFD3E0C5B.taxon	distribution	Distribution: — Endemic to the Philippines. (Fig. 14). Habitat and ecology: — Secondary forests, edge of swamps, thickets, at low altitude. Phenology: — Flowering April – August, fruiting August – September. Preliminary conservation assessment: — Extent of Occurrence 95,122.598 km 2 = LC; Area of Occupancy 20.000 km 2 = EN. Category assigned: — Vulnerable (VU). Although the EOO determines this species to be of Least Concern, consideration of other factors suggests that the level of threat may be higher than this. Only a small number (6) of herbarium specimens have been seen by the present author, suggesting that the species is not common, at least in herbaria. Additionally, the most recent of these specimens was collected in 1953. However, it is also noted that the A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 25 species appears to be able to grow in disturbed or secondary vegetation, and is thus able to survive anthropogenic influences to some extent. Taxonomic notes: — Note from Hattink 1974: “ Specimen FB 479 Merritt, 20 Oct. 1906, Pinamalyan, Mindoro, agrees very well with Merrill’s description (e. g. prickles sometimes in threes), while also the locality is the same. It might have exactly resembled the lost type. ”	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD0FFDCFF11FEC2FD390E18.taxon	materials_examined	NEW CALEDONIA: Ile Art, 25 Aug. 1978, Morat, P. 6186 (fl.) (NOU); Ile Art, 25 Aug. 1978, Veillon, J. M. 3713 (fl.) (NOU); Massif de Poum, versant Nord, captage Tich, 150 – 200 m, 8 Mar. 2008, Dagostini, G., Rigault, F. 1691 (st.) (NOU); Nehoue, Mine Fantouche, 100 m, 18 Jun. 1967, McKee, H. S. 16913 (fr.) (K, NOU); Nehoue, Mine Fantouche, 50 – 100 m, 20 Apr. 1967, McKee, H. S. 16615 (fl.) (NOU); Nehoue, Cap Tonerre, 0 – 30 m, 4 Feb. 1968, McKee, H. S. 18349 (fl.) (K); Nehoue, Cap Tonnerre, 9 Sep. 1969, McKee, H. S. 20734 (st.) (K, NOU); Prov. Nord, Cap Deverd, 38 – 50 m, 20 45 36 S, 164 22 44 E, 30 Jul. 2009, Clark, R. P. 150 (fl., fr.) (K, NOU); Prov. Nord, Cap Deverd, 38 – 50 m, 20 45 36 S, 164 22 44 E, 30 Jul. 2009, Clark, R. P. 151 (fl., fr.) (K, NOU); Cap Deverd, 1861 – 67, Deplanche 341 (fl.) (K, NY, P); Cap Deverd, Gomen, 10 Apr. 1975, McKee, H. S. 30023 (st.) (K, NOU); Cap Deverd, south of Kaala-Gomen, 30 m, 22 Dec. 1983, McPherson, G. 6211 (fl.) (K, MO); Cap Deverd, 1861 – 67, Vieillard 2523 (or 2533) (fl.) (P); Tinip, 20 45 10 S, 164 27 10 E, 84 m, 30 Jul. 2009, Clark, R. P. 152 (fl., fr.) (K, NOU, P); Tinip, 20 45 10 S, 164 27 10 E, 84 m, 30 Jul. 2009, Clark, R. P. 153 (fl., fr.) (BRI, K, NOU, P); Tinip, 20 45 10 S, 164 27 10 E, 84 m, 30 Jul. 2009, Clark, R. P. 154 (fl., fr.) (K, NOU); Tinip, 20 45 10 S, 164 27 6 E, 70 m, Clark, R. P. 156 (fl., fr.) (K, NOU); Prov. Nord, Kaala-Gomen, Tinip, 26 m, 20 46 45 S, 164 27 26 E, 27 Nov. 2009, Labat, J. - N. et. al. 4116 (fl., fr.) (K, P); Tinip, entre Gomen et Ouaco, vers 20 m, 17 Jun. 1967, McKee, H. S. 16895 (st.) (K, NOU); Tinip, 12 May 1978, McKee, H. S. 35126 (fr.) (NOU); Tinip, 150 m, 29 Jul. 1981, McKee, H. S. 39353 (fl.) (K); Taom, Mt. Homedeboua, 800 – 900 m, 16 Oct. 1969, McKee, H. S. 20992 (fl.) (NOU); Prov. Nord, base of Katepahie, near Voh, 20 56 31 S, 164 41 8 E, 13 – 100 m, 24 Jul. 2009, Clark, R. P. 137 (fl.) (BRI, CHIC, K, NOU, P); Prov. Nord, base of Katepahie, near Voh, 20 56 25 S, 164 41 11 E, 25 – 30 m, 30 Jul. 2009, Clark, R. P. 157 (fr.) (K, NOU); Prov. Nord, base of Katepahie, near Voh, 25 – 35 m, 20 56 25 S, 164 41 10 E, 31 Jul. 2009, Clark, R. P. 158 (fl.) (K, NOU); Prov. Nord, base of Katepahie, near Voh, 20 56 31 S, 164 41 9 E, 25 – 30 m, 31 Jul. 2009, Clark, R. P. 159 (fl., fr.) (K, NOU, P); Prov. Nord, base of Katepahie, near Voh, 20 56 23 S, 164 41 11 E, 40 m, 31 Jul. 2009, Clark, R. P. 160 (fl.) (K, NOU); Base du Katepahie, 15 Dec. 1973, Jaffré, T. 1255 (fl.) (K, NOU); Voh, Tribu de Gatope, Presqu’ile de Gatope, 20 59 3 S, 164 40 4 E, 141 m, 16 Nov. 2004, Labat, J. - N. et. al. 3500 (fl.) (NOU); Voh, Katepahie, 1 Jan. 1976, McKee, H. S. 30567 (st.) (K, NOU, P); Mt Katepahie, 2 km NW of Voh, 20 – 100 m, 20 55 S, 164 40 E, 13 Dec. 1973, Webster, G. L., Jaffré, T. 19282 (fl.) (P); Oundjo, 0 – 50 m, 17 Feb. 1972, McKee, H. S. 25032 (fl.) (K, NOU); Kone, Pinjen, 16 Aug. 1970, McKee, H. S. 22424 (fl.) (NOU); Riviere Coco, base du Koniambo, Dec. 2006, Munzinger, J. 3987 (fl.) (NOU); Base du Pic Tiaoué, 21 10 40 S, 164 57 30 E, 130 m, 12 Dec. 2006, Hequet, V., Butin, J. - P., Henocque, S. 3526 (fl.) (NOU); Plateau de Tiea, 5 km S of Pouembout, 300 m, 4 May 1983, McPherson, G. 5682 (fl.) (MO); Presqu’ile de Nepoui, 100 ft., 10 Jul. 1952, McMillan, C. 5047 (fl.) (P); presqu’ile de Nepoui, 18 Nov. 1987, Veillon, J. M. 6588 (fl.) (NOU); Poya, Station de Mueo, 06 May 1998, Veillon, J. M. 8090 (fl.) (NOU); Mépouiri, 15 Mar. 2007, Hequet, V., Papineau, C., Paulaud, D. 3636 (fl.) (NOU); Cote Est, Vallee de la Poro, 21 18 58 S, 165 42 40 E, 100 m, 8 Oct. 2007, Barrière, R. 146 (fr.) (NOU); Canala, Route a Horaire, 21 34 5 S, 166 5 4 E, 294 m, 19 Aug. 2009, Barrabe, L., Nigote, W. 944 (fl.) (K, NOU); Route de Canala, Nov. 1869, Balansa, B. 2447 (fl.) (L, P); prés l’embouchure du Dothio, Balansa, B. 3668 (fr.) (P); Ouroue (embouchure de la Dothio), 10 – 100 m, 8 Jan. 1966, McKee, M. 14203 (fr.) (K); N. E. de St. Louis, Balansa, B. 1384 (fl.) (NY, P); Taulé, 1861 – 67, Deplanche 338 (fl. / fr.) (NY, P); Taulé, 1861 – 67, Deplanche 340 (fl. / fr.) (K, NY); Locality uncertain: Taulé, Vieillard 2521 (fl.) (P); Taulé, Vieillard 2522 (fl.) (P); Deplanche 558 (fl.) (P); 1861 – 1867, Vieillard 20525 (fr.) (P)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFBBFFB5FF11FD49FD200AD0.taxon	description	Lectotype: — NEW CALEDONIA: Taulé, Deplanche 338, P (P 00379688!); isolectotypes: — K!, L (L 0651493!), NY!, P (P 00625915!), Clark & Gagnon 2015. Remaining syntypes: — NEW CALEDONIA: Taulé, Deplanche 340 K (K 000264184!), L (L 0651493!), NY!, P (P 00379688!, P 00625915!, P 02142824!); Taulé, Vieillard 2521, P (P 00379687!, P 02142820!, P 02142822! P 02142823!); Ile Art (49 in herb. Lugd.) Mueller: 3, 12; Deplanche 558 P! (barcode unknown). Synonym. M. deverdiana Guillaumin, Bulletin de la Société Botanique de France 83: 310. 1936. Syntypes: — NEW CALEDONIA: Cap Deverd, Vieillard 2533 (specimen not seen, the number as written in the protologue is possibly a typographic error, as the specimen Vieillard 2523! is determined as M. deverdiana); Cap Deverd, Deplanche 341 K (K 000556379!), NY!, P (P 00379664!, P 00379665!, P 00379689!, P 00379690!); Cap Deverd, Pancher 41 (location of specimen unknown). Synonym. M. deverdiana Guillaumin var. glabrifoliolula Guillaumin. Bulletin de la Société Botanique de France 83: 310. Syntypes: — NEW CALEDONIA: Route de Canala, Balansa, B. 2447 L (L 0651586!), P (P 00379662!, P 00379663!, P 00624907!); prés de l’embouchure du Dothio, Balansa, B. 3668 P (P 00379661!, P 00379686!); Taulé, Vieillard 2522 P!. Scrambling shrub to ca. 6 m in length. Stems unarmed, or with scattered recurved prickles, glabrous to very sparsely to moderately whitish to ferruginous tomentose. Stipules semi-persistent, triangular, ca. 1 – 2 × 0.5 mm, moderately tomentose, or hairs on margins only. Leaves with 1 – 6 opposite pairs pinnae; 1 – 6 pairs leaflets per pinna; petioles 1.8 – 8.5 cm; leaf rhachis (1.5 –) 5.4 – 25 (– 28.8) cm, unarmed or with recurved prickles in pairs at pinna insertion points, and scattered on the internodes (particularly on younger leaves); pinnae (0.8 –) 2.4 – 7 cm; leaf rhachis, petiole, and pinna rhachis sparsely to moderately whitish to orange pilose, tomentose, or villous. Leaflets opposite to subopposite, shape variable, with irregular margin, elliptic to orbicular, base oblique, apex obtuse, rounded or retuse, margin inrolled; the terminal leaflets (1.2 –) 1.5 – 3.6 (– 4.3) × 0.8 – 2.5 (– 3.3) cm; lateral leaflets (1.1 –) 1.6 – 3.4 × 0.8 – 2.6 cm; both surfaces glabrous to sparsely, occasionally moderately, tomentose, with somewhat appressed, whitish to ferruginous hairs; 2 ˚ veins anastomosing, 3 ˚ veins reticulate, veins slightly sunken above and raised below. Inflorescence a panicle, often few-branched, axillary or terminal, flowers spaced up to 6 mm apart, 8 – 30 cm long, glabrous or sparsely to densely ferruginous tomentose; pedicels 2 – 7 mm, articulated ca. 1 – 2 mm below calyx, sparsely to moderately (densely when very young) ferruginous tomentose. Bracts sometimes present, triangular, c. 0.5 – 1 × 0.5 mm, partially tomentose; bracteoles linear or lanceolate, apex acute, 1.5 – 4 × 0.5 – 1 mm, sparsely to densely ferruginous tomentose. Flowers with a hypanthium ca. 1 mm deep, glabrous or sparsely to moderately ferruginous tomentose, or hairs on margins only; lower calyx lobe 4 – 6 × 2 – 3 mm; other lobes 3 – 6 × 2 – 3 mm, glabrous or sparsely to moderately ferruginous tomentose. Median petal 3 – 6 × 1.5 – 3 mm, with a patch of translucent or ferruginous hair on inner surface between claw and blade, and hairs on margins of claw, outer surface glabrous. Upper lateral petals elliptic to slightly obovate, 3 – 6 × 1.5 – 2 mm, blade glabrous, claw very sparsely ferruginous tomentose on inner surface. Lower lateral petals elliptic to slightly obovate, 3 – 6 × 1.5 – 2 mm, blade glabrous, claw very sparsely ferruginous tomentose on inner surface. Stamen filaments flattened, (2 –) 3 – 6 mm long, sparsely to densely tomentose, with long, translucent or whitish to ferruginous hairs on the basal ½ of the inner face or both faces; anthers ca. 0.75 – 1 mm long. Ovary 2 – 3 mm long, glabrous, or densely pubescent with translucent, whitish, or A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 41 ferruginous hairs, style 3 – 5 mm long, very sparsely pubescent on the basal ½; stigma funnel-shaped, the rim papillate. Fruit elliptic, the base cuneate, apex acute or the the wing extending to form a short beak; 2.4 – 5.4 × 1.4 – 2.4 cm, wing 2 – 4 mm; fruit surface glabrous or subglabrous, or sparsely to moderately ferruginous tomentose, glossy; venation reticulate, raised and visible on the fruit body, no veins visible on the wing; hypanthium persistent. Seeds 1 per fruit, suborbicular, not or faintly visible in outline on fruit exterior when mature.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFBBFFB5FF11FD49FD200AD0.taxon	distribution	Distribution: — Endemic to New Caledonia. Distributed mostly on the west coast in the northern half of the island, with two localities on the East coast, in the southern part of the island. Ile Art; Grand Terre, North and South Provinces (primarily North), East to West (primarily West). (Fig. 27). Habitat and ecology: — Maquis and sclerophyllous forest, on ultramaphic or non ultramaphic soil. Elevation 1 – 900 m. Phenology: — Flowering December – August, fruiting November – July. Preliminary conservation assessment: — Extent of Occurrence 15,220.484 km ² = VU; Area of Occupancy 60.00 km ² = EN. Category Assigned: — Least Concern (LC). This species does not seem directly threatened. (RLA Flore NC 2016). New Combinations (fossil taxa) Mezoneuron claibornensis (Herendeen & Dilcher) R. Clark & E. Gagnon comb. nov. Holotype: — Warman (Indiana University (I. U.) Locality 15826: Como Quad., Weakley Co., Tennessee, 36 17 15 – 30 N, 88 31 35 – 45 W) and Puryear (I. U. Locality 15820: Puryear Quad., Henry Co., Tennessee, 36 26 17 N, 88 20 15 W) clay pits, Claiborne Formation, western Tennessee, IU 15826 - 5853, 5853 '. Basionym. Caesalpinia claibornensis Herendeen & Dilcher. Am. J. Bot. 78 (1): 3. 1991. Mezoneuron flumen-viridensis (Herendeen & Dilcher) R. Clark & E. Gagnon comb. nov. Holotype: — Little Mountain, Green River Formation, southwestern Wyoming (I. U. Locality 15882: NW ¼ SE ¼ Sec. 23, T 13 N, R 106 W, Sweetwater Co.), IU 15882 - 7388, 7388 '. Basionym. Caesalpinia flumen-viridensis Herendeen & Dilcher. Am. J. Bot. 78 (1): 8. 1991. Mezoneuron spokanensis (Knowlton) R. Clark & E. Gagnon comb. nov. Lectotype: — Deep Creek, north of Spokane, Washington (United States Geological Survey locality 7557), USNM 37011. Basionym. Caesalpinia spokanensis (Knowlton) Herendeen & Dilcher. Am. J. Bot. 78 (1): 8. 1991. 42 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK Synonym. Cercis? spokanensis Knowlton. Knowlton 1926, U. S. Geol. Surv. Prof. Pap. 140: 43, P 1. 29, Fig. 9. Synonym. Cercis idahoensis Berry. Berry 1931, Bull. Torrey Bot. Club. 57: 240 – 243, P 1. 1, Figs. 1 – 3. Berry 1934, U. S. Geol. Surv. Prof. Pap. 185: 114, P 1. 22, Figs. 1 – 3; P 1. 23, Figs. 5, 6. Synonym. Cercis spokanensis Knowlton. Brown 1937, J. Wash. Acad. Sci. 27: 177, P 1. 54, Figs. 10 – 12. Excluded Names	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD0FFDCFF11F962FECE0F45.taxon	materials_examined	VIETNAM: Annam, Nui Hon-heo, Nhatrang, 3 Oct. 1932, Poilane, M. 4855 (fl.) (K, L); Annam, prov. Nhatrang, Suoi-cat, 28 Sep. 1918, Chevalier, M. A. 39.005 (fl.) (K, L); Annam, Nhatrang, a 7 km au N., 27 Sep. 1922, Poilane, M. 4.749 (fl.) (K, L)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF9DFF91FF11FE27FA140022.taxon	description	Bull. Mus. Natl. Hist. Nat. sér. 2, 24: 318. 1952. Type: — VIETNAM: Prov. Khanh Koa, Nui Han Heo, pres de Nha Trang, Poilane 4855. Lectotype: — P (P 03232908!), Larsen et al. 1980; isolectotypes K (K 000980293!), L (L 0651585!). Remaining syntypes: — VIETNAM: à 7 km de Nhatrang, route de Ninh-hoa, Poilane 4749; Nhatrang, réserve de Suoi-cat, Chevalier 39005, K!. Scandent shrub to 8 m in length. Stems with scattered recurved prickles, glabrous, lenticellate. Stipules persistent, oblong, apex acuminate, 1 × 2 mm. Leaves with 4 – 5 pairs pinnae; 4 – 5 [– 6] pairs leaflets per pinna; petioles 2 – 2.8 cm; leaf rhachis 8.5 – 14.5 cm, with recurved prickles in pairs and pinna insertion points, and usually scattered on the internodes; pinnae 4 – 7 cm; leaf rhachis, petiole, and pinna rhachis glabrous. Leaflets opposite to subopposite; the terminal leaflets elliptic, base oblique, apex obtuse to acute, 3.4 – 6.1 × 1.6 – 3 cm; lateral leaflets elliptic, base strongly oblique, apex obtuse to acute, 3.5 – 5.3 × 1.5 – 2.7 cm; all leaflets glabrous on both surfaces, glossy above; 2 ˚ veins anastomosing, 3 ˚ veins reticulate, veins visible on both surfaces, more prominent below, midvein sunken above, raised below. Inflorescence a panicle, terminal or axillary, to 30 [– 40] cm; sometimes with scattered recurved prickles on axes and pedicels, these ca. 1 mm long; axes with (very) sparse pale orange pubescent hairs, glabresecent on basal peduncle; pedicels slender, 5 – 14 [– 20] mm, articulated 0 – 1 mm below hypanthium, with few minute recurved prickles ca. 0.5 mm, very sparsely orange tomentose, hair denser at articulation. Bracts persistent, semicircular or triangular, ca. 1 – 1.5 × 1 – 2 mm, subglabrous or sparsely tomentose; bracteoles caducous, lanceolate, acuminate, 0.5 – 1 × 1 – 2 mm, sparsely tomentose. Flowers with a hypanthium 1 – 2 × 3 mm, glabrous or sparsely orange tomentose, sometimes at base only; lower calyx lobe 7 – 8 × 2 – 3 mm, other lobes 4 – 6 × ca. 2 mm; all lobes lobes sparsely orange tomentose on both surfaces, glabrescent. Median petal strongly reflexed backwards, 6 – 7 mm long, blade 3 – 4 × 3 mm, claw 2 – 3 × 1 – 2 mm; inner claw sparsely to moderately orange pubescent, petal otherwise glabrous. Upper lateral petals 7 – 10 mm long, blade ovate, 6 – 7 [– 9] × 4 [– 6] mm, glabrous, claw ca. 1 mm, with a few hairs on the inner surface, petal otherwise glabrous. Lower lateral petals 7 – 10 mm long, blade obovate, 6 – 7 × 3 mm glabrous, claw 1 mm long, with a few hairs on the inner surface, petal otherwise glabrous. Stamen filaments flattened, ca. 7 mm long, orange pubescent on inner face to the basal ½ of, or nearly entire length; anther 0.5 – 1 mm long. Ovary laterally somewhat inflated; 2 – 3 mm long, style ca. 5 – 12 mm long; ovary moderately orange pilose, style glabrous or subglabrous; stigma funnel-shaped, the rim papillate. Fruit unknown. Seeds unknown.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF9DFF91FF11FE27FA140022.taxon	distribution	Distribution: — Endemic to Vietnam, vicinity of Nhatrang. (Fig. 7) Habitat and ecology: — Unknown. Phenology: — Flowering September – October. Preliminary conservation assessment: — Extent of Occurrence 131.115 km 2 = EN; Area of Occupancy 12.000 km 2 = EN. Category assigned: — Endangered (EN). In the absence of further data, this is the appropriate assessment for this species. However, considering that there are only three herbarium specimens of this species known, and that the most recent of these was collected in 1932, it is likely that the species is more endangered than EOO and AOO values conclude, and may even be extinct. It is not possible to verify either of these possibilities without a detailed survey of the Nhatrang area. A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 15	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB3FFBEFF11FF51FA8609BB.taxon	materials_examined	Holotype: — AUSTRALIA: Queensland, Mt. Mueller, near Edgecombe Bay, 12 Deca. 1863, J. Dallachy s. n., MEL (MEL 48871!) Basionym. Pterolobium nitens F. Muell. ex Benth. Fl. Austral. 2: 279. 1864. Synonym. Caesalpinia nitens (F. Muell. ex Benth.) Pedley. Austrobaileya 5: 99. 1997. Scrambling shrub or liana, height unknown. Stems with scattered recurved prickles, older stems with spine-tipped tubercles and longitudinal corky ridges; sparsely whitish tomentose, glabrescent when older. Stipules persistent, triangular, 1 × 0.5 – 1 mm, sparsely to moderately pale orange tomentose. Leaves opposite to subopposite; 2 – 4 [– 5] pairs pinnae; 2 – 4 (– 5) pairs leaflets per pinna; petioles 1 – 2.5 cm; leaf rhachis 1.5 – 6 (– 8) cm, with recurved prickles in pairs at the pinna insertion points, and sometimes scattered on the internodes; pinnae 1.5 – 5 cm, pinna rhachis with a single recurved prickle at the base of each leaflet, sometimes with scattered prickles on the internodes; leaf rhachis, petiole, and pinna rhachis sparsely to moderately pale orange to pale golden tomentose. Leaflets alternate to subopposite; the terminal leaflets elliptic to obovate, base oblique, apex rounded to retuse, (1.2 –) 1.7 – 2.5 (– 3) × (0.8 –) 1.2 – 2.2 cm; lateral leaflets elliptic, base oblique, apex rounded to retuse, (0.9 –) 1.7 – 2 × (0.7 –) 1 – 1.5; all leaflets glabrous or sparsely appressed hairy above and below, hairs denser on midvein; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, veins slightly raised on both surfaces. Inflorescence a terminal few-branched panicle, the racemes opposite, relatively short, to ca. 12 cm, with scattered recurved prickles up to 2 mm in length along entire length of peduncle; moderately golden to orange tomentose; pedicels 2 – 3 mm, articulated ca. 1 mm below the hypanthium, moderately golden or orange tomentose. Bracts persistent, triangular, 1 × 1 mm, sparsely pale orange tomentose; braceoles caducous, lanceolate, 2 – 4 × 1 mm, moderately pale orange tomentose on both surfaces. Flowers with a hypanthium 1 – 2 × 2 – 3 mm, sparsely golden or orange tomentose; lower lobe 5 – 6 [– 7.5] × 2 – 3 mm; sparsely to moderately golden or orange tomentose; other lobes 4 – 5 × 2 mm; sparsely to moderately golden or orange tomentose, sometimes hairs only on central area and the margins, inner calyx glabrous. Median petal obovate, ca. 6 × 3 mm, blade ca. 4 mm long, claw ca. 1.5 – 2 mm long; circular patch of orange hair between claw and blade, the margins of the claw tomentose, outer surface glabrous. Upper lateral petals ca. 6 × 3 – 4 mm; blade obovate, ca. 4 mm long, claw ca. 2 mm long; claw tomentose on inner surface, petal otherwise glabrous. Lower lateral petals obovate, ca. 6 × 3 – 4 mm; blade ca. 5 mm long, claw ca. 1 mm long; claw tomentose on inner surface, petal otherwise glabrous. Stamen filaments flattened, [6 –] 7 – 8 mm long, moderately pale orange tomentose on basal ½ on inner surface; anther 0.5 – 1 mm long. Ovary ca. 2 mm long, style ca. 7 mm long, both glabrous; stigma narrowly funnel-shaped, the rim papillate; 0.5 – 0.75 mm wide. Fruit unknown. Seeds unknown.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB3FFBEFF11FF51FA8609BB.taxon	distribution	Distribution: — Endemic to Australia (Queensland). (Fig. 20). Habitat and ecology: — Notophyll forest; monsoon rainforest; on basalt / volcanic rock. Phenology: — Flowering October – December, fruiting time unknown. A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 33 Preliminary conservation assessment: — Extent of Occurrence 69,060.480 km 2 = LC; Area of Occupancy 76.000 km 2 = EN. Category assigned: — Vulnerable (VU). Notophyll forests, which are characterised by a predominance of notophyll leaf sizes in the canopy layer (Webb 1958; www. wettropics. gov. au, accessed 2016), were subject to extensive logging prior to 1987, and are now heavily fragmented and restricted to inaccessible locations. These forests are subject to further threat from infection by the plant pathogen Phytophthora cinnamomi Rands, and degradation due to edge effects (www. wettropics. gov. au, accessed 2016). If M. nitens occurs only in notophyll forest, its habitat area is limited, and is subject to ongoing threats. Therefore, although the EOO of the species determines it as LC according to IUCN criteria, the higher threat category of Vulnerable is here assigned.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD7FFDBFF11FEC2FCEC092D.taxon	materials_examined	NEW CALEDONIA: Ile Ouen, 1868, Balansa, M. 312 (fl.) (P)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB0FFBBFF11FB15FA290C22.taxon	materials_examined	Holotype: — NEW CALEDONIA: 27 ‘ 7 bre’ 1868, Balansa, B. 312 (fl.) P (P 00379653!). Basionym. Caesalpinia ouenensis Guillaumin, Bulletin de la Société Botanique de France 83: 308 – 309. 1936. Shrub, 1 – 2 m. Stems without recurved prickles, glabrous. Stipules unknown. Leaves bipinnate, unarmed; pinnae 2 – 3 opposite pairs, paripinnate; leaflets opposite, 1 – 2 pairs per pinna; petioles 2.5 – 3.5 cm; rhachis 6.5 – 7.5 cm; pinnae 1.5 – 3.8 cm; leaf rhachis, petiole and pinna rhachis glabrous. Leaflets orbicular to obovate, base oblique, apex rounded; 3.3 – 5 × 2.7 – 4 cm; 2 ˚ veins anastomosing, visible on both surfaces. Inflorescence an axillary raceme, ca. 7 cm long, all parts glabrous. Pedicels 8 – 10 mm, not articulated. Bract at base of pedicel 2 – 3 mm long, lanceolate. Hypanthium ca. 3 mm deep, glabrous, lower calyx lobe ca. 9 × 3 mm, other lobes ca. 6 – 7 × 3 mm, glabrous. Median petal size unknown, lateral petals ca. 11 – 13 mm long. Stamen filaments ca. 12 – 15 mm long, glabrous; anthers c. 1 mm long; ovary not seen. Fruit unknown.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB0FFBBFF11FB15FA290C22.taxon	distribution	Distribution: — Endemic to New Caledonia. Known only from the type specimen, collected on Ile Ouen (South Province). (Fig. 23). Habitat and ecology: — Single known specimen found on the edge of a stream. Vegetation not stated on specimen label, inferred to be maquis on ultramaphic substrate, as this is the only native vegetation type on Ile Ouen. Phenology: — Unknown. Preliminary conservation assessment: — Known only from a single locality, so no AOO or EOO values can be generated. 36 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK Category assigned: — Critically Endangered CR B 1 ab (iii) + 2 ab (iii) (PE). Mezoneuron ouenensis is known from a single herbarium specimen on the Ile Ouen (ultramafic substrate) collected in 1868. Ile Ouen, the type locality, is a small island (c. 8.7 × 7.2 km), which has been subject to anthropogenic disturbance (mining activities & bush fires etc.) that may have destroyed the population from which the type specimen was derived. It is therefore assigned a status of Critically Endangered (CR) with criteria B 1 ab (iii) + 2 ab (iii )) with the tag Possibly Extinct with a continuing decline of the habitat quality. It is recommended to implement prospection of the island in order to confirm its probable extinction or implement urgent conservation actions if it was relocated. (RLA Flore NC 2016)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD7FFDBFF11FEB6FB8D0D38.taxon	description	JAVA: Horsfield, T. 141 (fr.) (K); Horsfield, T. 135 (fl.) (K); Leschenault s. n. (fr.) (P) LESSER SUNDA ISLANDS: Island of Wetar, between Camp I on the Meta Lahela and Ilwaki, 1 – 100 m, 8 – 9 Apr. 1939, Bloembergen, S. 3662 (fl.) (K, L, US); W. Sumbawa, Pernak, Olat Seli, + / - 12 km S. of Sumbawa Besar, 50 m, 17 May 1961, Kuswata, 217 (fr.) (K, L); S. Timor, S. Belu, Kalitin, 5 Jul. 1980, Verheien, J. A. J. 4851 (fr.) (L); Nusa Tenggara, Kp. Obe Naik, 300 m, 2 Mar. 1939, Bloembergen, S. 3342 (fl.) (K, L); Lombok, 2 Jul. 1909, Elbert, J. 2490 (fr.) (L); Lombok, 27 Apr. 1909, Elbert, J. 692 (fl.) (L); Lombok, 500 – 700 m, 30 Apr. 1909, Elbert, J. 799 (fl.) (L); W. Flores (S. part), Wt. Ndeki, 200 m, 12 Apr. 1965, Kostermans & Wirawan 176 (fl.) (L)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD7FFDBFF11FEB6FB8D0D38.taxon	materials_examined	PHILIPPINES: Luzon, Cagayan Prov., Alcala, Apr. 1927, Clemens, J. 17531 (fl.) (NY); Malinta bei Manila, 6 Dec. 1903, Hallier, E. D. s. n. (fr.) (NY); Luzon, Rizal Prov., Bosoboso, Sep. 1904, Ahern’s collector, 206 (fl.) NY; Luzon, Rizal Prov., Bosoboso, Sep. 1906, Ramos, M. s. n., Bureau of Science no. 1477 (fl.) (NY); Luzon, Prov. Rizal, Sep. 1905, Ahern’s collector, s. n. Forestry Bureau number 3370 (fl.) (K, NY); Luzon, Pampanga Prov., Mt. Arayat, 31 Oct. 1925, Clemens s. n., Bureau of Science no. 16150 (fl.) (BM, NY); Manila, San Francisco del Monte, 25. Aug. 1890, Loher, A. 2191 (fl.) (K); Loher, A. 5909 (fl.) (fr.) (K); Centre of Luzon, s. coll. (Com. Vidal) 1284 (fl.) (K); Luzon, Bulacan Province, Angat, Sept. 1913, Merrill, 294 A (fl.) (F, K, L, NY); Luzon, Rizal, Antipolo, Jan. 1914, Ramos, M. 2089 (fr.) (BM, L, MO); Manila, Shaw, W. R. 334 (fr.) (BM); Norzagaray, Bulacan Prov., Luzon, Jan. 1914, Foxworthy, F. W. s. n., Bureau of Science no. 12265 (fr.) (BM); Quezon City, 50 m, 14 Jan. 1953, Britton, B. B. 34 (fr.) (L); Luzon, Prov. Rizal, San Pedro Macati, 26 Dec. 1909, Robinson, C. B. s. n. (Bureau of Science number 9601) (fr.) (L); Luzon, Prov. Rizal, Antipolo, Jan 1914, Ramos, M. s. n. (Bureau of Science number 2089) (fr.) (L); Luzon, Bulacan Province, Angat, Sep. 1913, Merrill 294 A (fl.) (L) THAILAND: Kerr, A. F. G. 13111 (fl.) (E, K) TIMOR: [Herb. Lugd. Batav.], s. coll., s. n. (fl.) (K, L); T. T. U., 7 Jun. 1901, Koay, C. W. 1307 (fl.) (L); Timor, [coll. illegible], s. n. (fl.) (L); [Herb Spanoghe], Teones 34 (fl.) (L); s. coll., s. n., [Herb. Lugd. Bat.] (fl., fr.) (L); Kupang, s. coll., s. n. (fr.) (K) VIETNAM: Annam, Nha-trang and vicinity, 11 – 26. Mar. 1911, Robinson, C. B. 1355 (fl., fr.) (K, NY); Annam, Tourane and vicinity, May – Jul., 1927, Clemens, J. & M. S. 3092 (fr.) (K); Annam, Han-Neo, pres Nhatrang, 3 Oct. 1922, Poilane, M. 4844 (fl.) (K); Phan Thiet, 5 Nov. 1924, Evrard, F. 1676 (fr.) (L)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF85FF87FF11FAC2FDE809E3.taxon	description	Mem. Mus. Hist. Nat. Paris 4: 245. 1818. Holotype: — JAVA, Leschenault s. n., P (P 02142683!) Synonym. Caesalpinia pubescens (Desf.) Hattink. Reinwardtia 9 (1): 47. 1974. Mezoneuron pubescens Desf. var. longipes Craib. Fl. Siam. Enum. 1: 500. 1928. Holotype: — THAILAND, Surat, Kanchanandit, Aug. 1927, Kerr 13111, K (K 000264758!). Synonym. Mezoneuron glabrum Desf. Mem. Mus. Hist. Nat. Paris 4: 245. 1818. Holotype: — TIMOR, Leschenault s. n., P (P 00293980!). Synonym. Mezoneuron intermedium Zippelius ex Spanoghe. Linnaea 15: 200. 1841. Woody climber, or scandent shrub, to 4 m tall and spreading to 15 m. Stems sometimes with small, scattered recurved prickles; sparsely to moderately whitish to orange tomentose. Stipules caducous, triangular, 1.5 – 5 mm long, subglabrous to densely golden tomentose. Leaves with 7 – 15 (– 18) pairs pinnae; 5 – 11 pairs leaflets per pinna; petiole 2 – 7.8 cm; leaf rhachis (9 –) 15 – 34 [– 50] cm, with recurved prickles on leaf rhachis in pairs or threes at the pinna insertion points, and usually scattered on the internodes, a single prickle at apex of rhachis, ca. 2 mm long; pinnae 2 – 8.5 cm; leaf rhachis, petiole, and pinna rhachis sparsely to moderately whitish to orange tomentose. Leaflets alternate to opposite, often discolorous, darker above; the terminal leaflets obovate, base oblique, apex rounded to slightly acute, mucronulate, (5 –) 10 – 20 × 3 – 13 mm, lateral leaflets oblong, base oblique, midvein often excentric to almost diagonal, 5 – 20 × 2 – 13 mm; upper surface of all leaflets sparsely to moderately appressed tomentose, occasionally glabrous, lower surface sparsely appressed tomentose, often denser on midvein and margins, the hairs pale orange to golden to whitish; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, veins clearly A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 23 visible on lower surface, faintly visible above. Inflorescence a raceme or few-branched panicle, terminal or (supra-) axillary, to 40 cm, sparsely to moderately whitish or golden or orange tomentose on all parts, the hairs often denser and darker in colour on calyces and younger parts; occasionally small recurved prickles on the basal ⅓ of infloresence rhachis; pedicels 14 – 40 mm, articulated 1 – 3 mm below the hypanthium. Bracts sometimes present, lanceolate to triangular, 1 – 4 × 1 mm, moderately tomentose; bracteoles semi-persistent, lanceolate or elliptic, long-acuminate, 6 – 15 mm × 1 – 3 mm, orange tomentose on both surfaces, the hairs forming a distinct central longitudinal stripe on the outer surface. Flower with a hypanthium 2 – 4 mm × (6 –) 7 – 8 mm; lower calyx lobe 12 – 17 × 6 – 12 mm, other lobes broad, rounded, 8 – 11 × 5 – 7 mm, hypanthium and calyx lobes all moderately to densely orange pubescent on both surfaces. Median petal strongly reflexed backwards, with a pronounced transverse flap of tissue on the inner surface between claw and blade; 8 – 14 mm long, blade oblong to suborbicular, 4 – 8 × 5 – 11 mm, glabrous or with a few hairs on outer surface, claw 4 – 6 × 0.5 – 2.5 mm, glabrous or with a few hairs on outer surface. Upper lateral petals obovate to suborbicular, blade 9 – 14 [– 20] × 8 – 16 [– 20] mm, claw 2 – 3 mm long, few hairs at the base of claw on inner and outer surfaces, occasionally glabrous. Lower laterals obovate to suborbicular, blade 8 – 13 [– 20] × 8 – 14 [– 20] mm, claw 2 – 3 mm long, few hairs at the base of claw on inner and outer surfaces, occasionally glabrous. Stamen filaments flattened, (12 –) 14 – 18 mm long, the filament of vexillary stamen glabrous, the other filaments covered with dense villous orange hairs on the basal ½, along the inner surface; anther ca. 2.5 – 3.5 mm. Ovary 5 – 8 mm long, style [3 –] 8 – 13 mm long, the ovary and basal ⅓ of style moderately to densely orange sericeous, sometimes extending the length of the style; stigma funnel-shaped, ca. 0.5 – 1 mm wide, the rim papillate. Fruit narrowly elliptic, sometimes slightly falcate, base sometimes oblique, apex acute, the central nerve extending to form a slight beak, the persistent, asymmetric hypanthium wider below winged side of fruit; 8.5 – 24.5 × 3 – 5.5 cm, the wing arising basally up to 12 mm along central nerve, wing 8 – 20 mm wide; fruit surface densely orange tomentose when young, hairs usually becoming sparse when mature, glabrescent, venation reticulate, raised and visible on fruit body, anastomosing and slightly visible on the wing; hypanthium semi-persistent. Seeds 4 – 9 per fruit; arranged close together, adjacent to and perpendicular to the central vein, not visible in outline on fruit exterior, outline of entire seed chamber sometimes slightly visible; ellipsoid, 8 – 10 × 3 – 6 mm, brown, smooth.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF85FF87FF11FAC2FDE809E3.taxon	distribution	Distribution: — Thailand, Laos, Vietnam, Indonesia, Lesser Sunda Islands, Philippines, Java, Sumatra, Sulawesi. (Fig. 13). Habitat and ecology: — Scrub, dry thicket, savannah, monsoon forest. Elevation to 700 m. Occurs in regions that experience a dry season. Phenology: — Thailand, Vietnam: flowering March – October, fruiting March – November. Malesia: Flowering March – October, fruiting May – January. 24 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK Preliminary conservation assessment: — Extent of Occurrence 7,038,070.582 km 2 = LC; Area of Occupancy 96.000 km 2 = EN. Category assigned: — Least Concern (LC). However, the majority of the existing herbarium specimens were collected before 1950, and the most recently collected specimen known to the author is from 1980. This suggests that the species distribution may have been reduced since the majority of the specimens were collected, and is smaller than the specimen data appears to show. If further data becomes available, it is likely that a higher category of threat will be assigned to this species.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD7FFDBFF11FAB8FDD60E27.taxon	materials_examined	NEW CALEDONIA: Koumac, 250 m, 1 Dec. 1972, McKee, H. S. 26003 (fl.) (K, P); Lifou, 27 Oct. 1925, Daniker, A. U. 2390 (fl.) (P); Boguen River Valley, above Katrikoin, c. 20 km E of Bourail, c. 160 m, 20 Nov. 1979, McPherson, G. 2089 (fl.) (MO, NOU); Boguen River, c. 25 km NW of La Foa., 160 m, 20 Nov. 1979, Hartley, T. G. 14975 (fl.) (NOU); Boghen, 100 m, Oct. 1900, Cribs 982 (fl.) (P); Ouaia, 6 Oct. 1886, Gervais s. n. (fl.) (P); Mount Boakim, 800 m, Lecard, M. T. s. n. (fl.) (P)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB4FFB7FF11FEC2FD940A0D.taxon	description	Lectotype: — NEW CALEDONIA: Boghen, Cribs 982, P (P 02969638!); isolectotype: P (P 02969637!), Clark & Gagnon 2015. Remaining syntype: — NEW CALEDONIA: Mount Boakim, Lecard s. n., P (P 02969633!); isosyntype P (P 02969644!). Basionym. Caesalpinia rubiginosa Guillaumin, Bulletin de la Société Botanique de France 83: 309 – 310. 1936. Liana to 5 m in length (or more). Stems unarmed; sparsely to moderately whitish to ferruginous tomentose. Stipules semi-persistent, triangular, 1 – 1.5 × 0.5 mm, sparsely tomentose. Leaves with 4 – 6 pairs pinnae; 2 – 5 pairs leaflets per pinna; petiole 4.3 – 14 cm; rhachis 14.5 – 25.5 cm, unarmed or with recurved prickles in pairs at pinna insertion points, and sometimes scattered on the internodes; pinnae 4.8 – 10 cm; leaf rhachis, petiole, and pinna rhachis sparsely to densely whitish or ferruginous tomentose. Leaflets opposite, elliptic to obovate, base oblique, apex rounded to retuse; the terminal leaflets 3.2 – 5.1 × 2 – 3.3 cm; lateral leaflets 2.8 – 5.1 × 1.8 – 3 cm; 2 ˚ veins anastomosing, 3 ˚ veins reticulate, veins clearly visible on both surfaces; all leaflets with upper surface glabrous, or with few whitish to ferruginous hairs on basal midvein, lower surface glabrous or glabrate, with sparse hairs on petiole and midvein, or sparsely appressed hairy, denser on midvein. Inflorescence an axillary, supra-axillary, or terminal, raceme or few to many-branched panicle, congested, the flowers spaced ca. 2 – 3 mm apart; 12 – 18.5 cm long, sparsely to densely ferruginous tomentose on peduncle and pedicels; pedicels 2 – 3 mm, not articulated. Bracts sometimes present, persistent, triangular, ca. 1 × 0.5 mm, moderately to densely tomentose; bracteoles caducous, linear, inrolled, 3 × 0.5 – 1 mm, moderately to densely ferruginous tomentose. Flowers with a hypanthium ca. 2 mm deep, sparsely to moderately ferruginous tomentose; lower calyx lobe ca. 5 × 2 mm, sparsely ferruginous tomentose, other lobes ca. 4 × 2 mm, glabrous or with very few hairs on the margins. Median petal 6 × 2.5 – 3 mm, with a patch or band of ferruginous hairs on inner surface of claw, outer surface glabrous. Upper lateral petals elliptic to slightly obovate, 6 – 7 × 2.5 mm, blade glabrous, claw very sparsely ferruginous tomentose on inner surface or both surfaces very sparsely puberulous. Lower lateral petals elliptic to slightly obovate, 6 – 7 × 2.5 mm, blade glabrous, claw very sparsely ferruginous tomentose on inner surface or both surfaces very sparsely puberulous. Stamen filaments 6 – 8 mm long, ferruginous tomentose on the basal ⅓; anthers ca. 1 mm long. Ovary 3 – 4 mm long, densely ferruginous tomentose, style 4 – 10 mm long, moderately ferruginous pubescent along most or all of its length; stigma funnel-shaped, the rim papillate. Fruit unknown.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB4FFB7FF11FEC2FD940A0D.taxon	distribution	Distribution: — Endemic to New Caledonia. Lifou (Îles Loyauté); Grand Terre, North and South Province, Western side. (Fig. 26). 40 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK Habitat and ecology: — Wet forest, on non-ultramaphic soil (collections McPherson 2089, Daniker 2390, Hartley 14975). The species may also occur in sclerophyllous forest, according to the localities of some specimens collected in areas that appear to have been formerly vegetated with sclerophyllous forest (Jaffré et al. 1998). Due to the imprecise nature of the locality information available, however, it is not possible to confidently establish the vegetation type from which some of the specimens were collected (McKee 26003, Cribs 982, Gervais s. n.). Elevation 100 – 800 m. Phenology: — Flowering October – November, fruiting time unknown. Preliminary conservation assessment: — Extent of Occurrence 16,020.691 km ² = VU; Area of Occupancy 16.00 km ² = EN. Category assigned: — Data Deficient (DD). This species has been only occasionally observed due to the difficulty to see it on the top of trees. There is no recent herbarium material. Therefore, M. rubiginosum is assigned as Data Deficient and should be re-assessed when more observations have been made that will clarify its distribution (RLA Flore NC 2016).	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB7FFBAFF11FBB8FF540BED.taxon	materials_examined	Type: — NEW CALEDONIA: Auf den Bergen bei Oubatche, 800 m, 26 Deca. 1902, Schlechter, F. R. R. 15572 (fl.). Lectotype: — M (M 0217766, image!); isolectotypes: — BR, (000005110605, image!), K (K 000556378!), L (L 0018794!), Clark & Gagnon 2015. Basionym. Caesalpinia schlechteri Harms. Beiträge zur Kenntnis der Flora von Neu-Kaledonien. Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie: 1 – 274. 1906. Liana to 20 m or more in length. Stems unarmed, or with scattered recurved prickles, glabrous or sparsely to densely ferruginous tomentose. Stipules semi-persistent, triangular, 0.5 – 1 × 0.5 – 1 mm, subglabrous to sparsely tomentose. Leaves with (2 –) 3 – 7 (– 8) pairs pinnae; (3 –) 4 – 8 (– 14) pairs leaflets per pinna; petiole 2 – 5.5 (– 8) cm; leaf rhachis (3 –) 5 – 23 cm, unarmed, or with recurved prickles up to 5 mm in length, in pairs at pinna insertion points and scattered on the internodes; pinnae 2.8 – 12.7 cm; leaf rhachis, petiole sparsely to moderately (sometimes patchily) ferruginous pilose or tomentose, pinna rhachis glabrous to moderately (occasionally densely) ferruginous pilose or tomentose. Leaflets opposite, elliptic to rhomboid, strongly oblique at base, apex obtuse to retuse; the terminal leaflets 2 – 4.7 × 0.6 – 3.1 cm; lateral leaflets 1.8 – 4.4 × 0.6 – 2.3 cm; 2 ˚ veins anastomosing, 3 ˚ veins reticulate, veins clearly visible on both surfaces, or indistinct on upper surface; upper surface usually glossy, glabrous, or sometimes with a few hairs at the base or along the basal part of the margins, lower surface glabrous, with sometimes a few hairs at the base or along the basal part of the margins. Inflorescence a raceme or panicle, axillary, supra-axillary or terminal, 13 – 31 cm long, glabrous, or sparsely to moderately ferruginous tomentose on all parts, occasionally a few recurved prickles on the basal portion of the peduncle. Pedicels 3 – 10 mm, not articulated, glabrous to densely ferruginous tomentose. Bracts sometimes present, triangular, 0.5 – 2 × 0.5 – 1 mm, subglabrous to sparsely tomentose; bracteoles usually caducous, lanceolate, inrolled, 1 – 1.5 × 0.5 – 1 mm, sparsely to moderately orange pubescent, or with hairs only along margin, sometimes glabrescent. Flowers with a A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 37 hypanthium 1 – 2 mm deep, glabrous, lower calyx lobe ca. 8 × 3 – 4 mm, other lobes ca. 6 × 3 – 4 mm, all lobes glabrous or with hairs on margins. Median petal 4 – 6 × 1.5 – 3 mm, with a circular patch of dense, dark ferruginous hairs on inner surface between claw and blade, and hairs on margins of claw, outer surface glabrous. Upper lateral petals elliptic to slightly obovate, (4 –) 5 – 8 × 1.5 – 3 mm, blade glabrous, claw very sparsely ferruginous tomentose on inner surface. Lower lateral petals elliptic to slightly obovate, (4 –) 5 – 8 × 1.5 – 4 mm, blade glabrous, claw very sparsely ferruginous tomentose on inner surface. Stamen filaments 5 – 11 mm long, the basal ⅓ to ½ with long translucent or ferruginous tomentose hairs; anthers ca. 1 mm long. Ovary 3 – 4 mm long, glabrous, style 5 – 7 mm long, sparsely to moderately pale translucent or ferruginous tomentose on lower half, sometimes thickening towards apex; stigma funnel-shaped, the rim papillate. Fruit elliptic, the base cuneate, apex acute or the wing extending to form a short beak; 4 – 5.6 × 1.7 – 2.6 cm, wing 4 – 6 mm wide; fruit surface glabrous; venation reticulate, raised and visible on the fruit body, no veins visible on the wing; hypanthium persistent. Seeds 1 per fruit, suborbicular, not or faintly visible in outline on fruit exterior when mature.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFB7FFBAFF11FBB8FF540BED.taxon	distribution	Distribution: — Endemic to New Caledonia. Grande Terre, North and South Province, East to West (primarily East). (Fig. 24). Habitat and ecology: — Dense wet forest, on ultramaphic and non-ultramaphic (volcano-sedimentary) soil. Elevation 100 – 800 m. Phenology: — Flowering July – January, fruiting December – February Preliminary conservation assessment: — Extent of Occurrence 10,382.55 km ² = VU; Area of Occupancy 84.00 km ² = EN. Category assigned: — Least Concern (LC). This species does not seem directly threatened. (RLA Flore NC 2016).	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD6FFD9FF11FCAEFA8D0B30.taxon	materials_examined	AUSTRALIA: Budlein Mt, Woombe, Apr. – May 1911, s. coll., s. n. (fr.) (BRI); Coveny, R. 3880 (fr.) (BRI, K, L); Fraser Island, 17 – 18 Oct. 1930, Hubbard, C. E. 4581 (fl.) (L); Konorigan Range, 5 miles NE of Nimbin, 0.7 miles SW of Mt Nardi summit, 28 33 S, 153 16 E, 5 Sep. 1972, Coveny, R. 4495 (fr.) (BRI); Mt Slavian, May 1920, White, C. T. s. n. (fr.) (BRI); New South Wales, 1 – 2 miles S of Mt Nardi, 6 Jan. 1971, Salasoo, H. 4540 (st.) (BRI); New South Wales, 3.7 miles c. S of Murwillumba, 28 22 S, 153 24 E, 10 Dec. 1971, Thurtell, K. 3880 (fr.) (BRI, K, L); New South Wales, C. 10 miles SW of Murwillumbah, North Coast, 45 m, 28 26 S, 153 15 E, 10 Oct. 1969, Hoogland, R. D. 11665 (fl.) (BRI, E, K, L); Queensland, 1 km E of Mount Coorey, 130 m, 26 26 S, 152 58 E, 17 Nov. 1990, Forster, P. I. PIF 7584 (fl.) (BRI); Queensland, 26 S, 151 E, Aug. 1944, Clemens, M. S. s. n. (fr.) (BRI); Queensland, Benarkin, Pennings Rd, 13 Mar. 1985, 26 53 S, 152 08 E, Zink, D. s. n. (fr.) (BRI); Queensland, Blaudie River, 60 m, 12 45 S, 143 15 E, 11 Oct. 1972, Hyland, B. 6401 (fr.) (L); Queensland, Bromley Out Station, 60 m, 12 30 S, 143 12 E, 20 Sep. 1976, Hyland, B. 9022 (fl.) (L); Queensland, Brookfield, 27 30 S, 152 54 E, Hargreaves, J. 700 (fl.) (BRI); Queensland, Cape York Peninsula, Dec. 1873, Hann, W. 188 (fl.) (K); Queensland, Cook District, Mcilwraith Range, 13 35 S, 143 25 E, 25 Jul. 1978, Butler, G. 362 (fl., fr.) (L); Queensland, Daintree Area, Noah Creek, 5 m, 16 08.7 S, 145 26.2 E, 06 Mar. 2002, Wieringa, J. J. et. al. 4195 (st.) (K); Queensland, Dalrymple Heights, Crediton Parish, Aug. – Sept. 1947, Clemens, M. S. s. n. (fl., fr.) (BM, F, K, L, NY); Queensland, Double Island Point, Wide Bay District, 16 Oct. 1946, Clemens, M. S. s. n. (fr.) (K); Queensland, Fernleigh Rd., Fernleigh, 28 47 S, 153 29 E, 15 Oct. 1990, Wilson, K. L. 7648 (fl.) (BRI); Queensland, Fraser Island, 100 ft., May 1964, Smith, L. S., Tracey, J. G. 6372 (st.) (BRI); Queensland, Fraser Island, 17 – 18 Oct. 1930, Hubbard, C. E. 4581 (fl.) (K, L); Queensland, Kandanga, Mary Valley, 10 Nov. 1933, White, C. T. 9611 (fl.) (BRI); Queensland, Moreton District, 1 – 2 miles east of Nambour, off Paynter Creek Road, 26 38 S, 153 00 E, 4 May 1977, Elsol, J. A. 126 (fr.) (BRI); Queensland, Moreton District, 3 km SW of Mooloolah, Lower Mooloolah River, 26 47 S, 152 57 E, 7 Nov. 1982, Sharpe, P. R. 3237 (fl.) (BRI); Queensland, Moreton District, Mooloolah River, 3 Jan. 1962, Everist, S. L. 7021 (fr.) (K); Queensland, Moreton District, Mount Glorious, 21 Nov. 1946, Tyack-Bake, J. s. n. (fl.) (BRI, K); Queensland, Moreton District, near Yandina, 30 Oct. 1937, White, C. T. 12102 (fl.) (BRI, K); Queensland, Moreton, 24 May 1888, Simmonds, J. H. s. n. (fl., fr.) (BRI); Queensland, Moreton, 2 km north of Peachester township, McDonald Road Recreation Reserve, London Creek area, 26 49 S, 152 53 E, 1 Dec. 1993, Grimshaw, P. G 223 (fr.) (BRI); Queensland, Moreton, 3 km south of Cooroy, Ocean View Road, 26 26 S, 152 55 E, 2 Nov. 1997, Bean, A. R. 12507 (fl.) (BRI); Queensland, Moreton, Dianas bath area, 9 km E of Somerset Dam, 160 m, 27 06 S, 152 38 E, 19 Jun. 1993, Forster, P. I. 13369 (fr.) (BRI); Queensland, Moreton, NW of Brisbane near Bryces Road, Mount Glorious SF 1355 D’Aguilar Range, 27 20 08 S, 152 45 45 E, 30 Aug. 1997, Franks, A. J., Butler, D. W. AJF 9708018 (st.) (BRI); Queensland, Moreton, Wises Road, Immanuel Lutheran College, 26 40 16 S, 153 4 58 E, 70 • Phytotaxa 274 (1) © 2016 Magnolia Press	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD6FFD9FF11FCAEFA8D0B30.taxon	description	CLARK 9 Nov. 2003, Leiper, G., Bright, H. WGS 84 (fl.) (BRI); Queensland, Moreton, Yandina-Bli Bli road, 26 35 S, 152 59 E, 29 Oct. 1989, Bean, A. R. 1158 (fl.) (BRI); Queensland, Mount Brisbane, Timber Reserve 209, 27 06 S, 152 32 E, 21 Jun. 1990, Forster, P. I. 6877 (fr.) (BRI); Queensland, Mount Glorious, Moreton District, Jan. 1945, Clemens, M. S. s. n. (fr.) (F, K); Queensland, near Brisbane, Upper Brookfield, Jun. 1926, White, C. T. s. n. (fl.) (K); Queensland, North Brisbane, Cedar Creek, Jan. 1917, Longman, H. A. s. n. (fr.) (K); Queensland, nr. Brisbane, Upper Burkefield, White, C. T. s. n. (fr.) (BRI); Queensland, Pamona Kin-Kin Rd, Kin Kin, 3 km E of Cooran, 141 m, 26 19 12.5 S, 152 52 23 E, 22 Feb. 2007, Boyle, P. & Halford, J. BGQLD 451 (fr.) (K); Queensland, Petrie, 18 miles N. of Brisbane, 14 Dec. 1930, Blake, S. T. s. n. (fr.) (BRI); Queensland, South Kennedy, Kinguri road to Mount Ossa, 21 05 S, 148 44 E, Nov. 1990, Pearson, S. G. 403 (fl.) (BRI); Queensland, Wide Bay, 2 km north of Upper Widgee, Station Creek, 26 11 30 S, 152 26 30 E, 21 Oct. 2001, Phillips, S. P. 684 (fl.) (BRI); Queensland, Wide Bay, Conondale Ranges, Bundaroo Creek, 26 42 S, 152 36 E, 28 Nov. 1990, Bean, A. R. 2696 (fl.) (BRI); Queensland, Wide Bay, Jimna Range, Sunday Creek Road western section near Jimna Road intersection, 26 41 S, 152 28 E, 15 Dec. 1993, Grimshaw, P., Franks, A. J. G 281 (fl.) (BRI); Wide Bay, 1851, Bidwill (Hooker) s. n. (fr.); NSW; Queensland, Cook District, Kuranda, 16 49 S, 145 38 E, 7 Nov 1958, Smith, L. S. 10448, (fl., fr.) (BRI, K, L); Queensland, Daintree area, Noah Creek, 5 m, 16 08.7 S, 145 26.2 E, 29 Jun. 2001, Wieringa, J. J. 4195 (st.) (K); Queensland, Cook District, JCU Canopy Crane plot, 16 06 11 S, 145 26 49 E, 12 Dec. 2009, Costion, C. 2181 (fl.) (CNS); Noah Creek, Cape Tribulation Road, 5 m, 16 9 S, 145 26 E, 14 Oct. 2000, Gray, B. 7842 (fl.) (CNS); T. R. 1230, Boonjie L. A., 680 m, 17 24 S, 145 45 E, 7 Sep. 1976, Hyland, B. 8917 (st.) (CNS)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD6FFD9FF11FCAEFA8D0B30.taxon	materials_examined	PAPUA NEW GUINEA: Tage, Lake Kutubu, Southern Highlands, Papua, 2700 ft., 25 Sep. 1961, Schodde, R. 2246 (fl.) (CANB, K, L); Lake Daviumbu, Middle Fly River, Aug. 1936, Brass, L. J. 7498 (fl.) (BRI, BM)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF8EFF80FF11FC8CFA280024.taxon	description	South. Sc. Rec.: 73. 1882. Lectotype: — AUSTRALIA: Queensland, Wide Bay, Bidwill s. n., K (K 000264178!), Hattink 1974. Remaining syntypes: — AUSTRALIA: Queensland, Logan R., 1881, B. Scortechini 166, MEL (3 sheets), NSW; Burnett R., Qld, F. Mueller s. n. (location of specimen unknown). Synonym. Mezoneuron traceyi Pedley syn. nov. Type: — AUSTRALIA: Queensland, Cook District, Kuranda, 16 49 S, 145 38 E, 7 Nov. 1958, L. S. Smith 10448. Holotype BRI (BRI-AQ 0010072, image!); isotypes K (K 000789366!), L (L 0651494!) Basionym. Caesalpinia traceyi Pedley. Austrobaileya 5: 99. 1997. Woody climber or scandent shrub to 7 m in length. Stems sometimes with recurved prickles to 3 mm, corky spine-tipped tubercles or longitudinal ridges on older stems; sparsely orange tomentose, glabrescent. Stipules persistent, triangular, black, 0.5 – 2 × 0.5 – 2 mm, sparsely orange tomentose, glabrescent. Leaves alternate, with 4 – 9 pairs pinnae; (4 –) 5 – 9 pairs leaflets per pinna; petiole 1.5 – 7.5 cm; leaf rhachis (8.5 –) 11 – 25 (– 33) cm, with paired recurved prickles at pinna insertion points, and usually scattered on the internodes, including terminal prickle, to 4 mm; pinnae opposite to subopposite, 4.5 – 9.5 (– 11) cm, pinna rhachis sometimes with scattered recurved prickles, including a terminal prickle 1 – 2 mm; leaf rhachis, petiole, and pinna rhachis sparsely to moderately whitish to orange tomentose. Leaflets alternate to subopposite; the terminal leaflets oblong to elliptic to slightly obovate, base oblique, apex rounded to retuse, mucronulate 1.4 – 3 (– 3.5) × 0.7 – 1.7 (– 2) cm; lateral leaflets oblong to elliptic, base oblique, apex rounded to retuse, mucronulate, 1.1 – 2.7 (– 3.6) × 0.5 – 1.5 (– 2.2) cm; all leaflets with upper surface glabrous or sparsely whitish to orange subappressed hairy, denser at base of blade around midvein, lower surface sparsely, regularly, whitish to orange appressed pilose, the hairs denser at base of midvein, occasionally subglabrous; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, 2 ˚ veins slightly or not visible above, slightly or clearly visible below. Inflorescence a panicle, often large and complex, occasionally a raceme, axillary or terminal, to 40 cm, the racemes alternate; sometimes with recurved prickles to 2 mm on lower part of racemes, peduncle and racemes sparsely to moderately orange tomentose; pedicels 2 – 7 mm, elongating in fruit to 13 mm, articulated 1 – 2 mm below hypanthium; sparsely to moderately pale orange to golden tomentose, hair denser around articulation, glabrescent in fruit. Bracts persistent, minute, triangular, ca. 1 – 1.5 × 1 mm, sparsely to densely orange tomentose on both surfaces, or at margins; bracteoles semi-persistent, lanceolate, narrowing at base, tapering to acuminate apex, (2 –) 3 – 6 × 1 – 2 mm, sparsely to moderately orange or golden tomentose to pilose on both surfaces. Flowers: sometimes a few male flowers per inflorescence; hypanthium 1 – 3 × ca. 3 – 4 mm; sparsely to moderately pale orange to golden pilose, the hair often paler than on peduncle; lower calyx lobe 5 – 8 × 2 – 3 mm; other lobes 3 – 6 × 1 – 3 mm; all calyx lobes sparsely pale orange to golden pilose to tomentose, hairs covering the lower lobe and on 30 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 31 lower parts of other lobes, sometimes denser on margins of lobes. Median petal 5 – 8 × 2 – 5 mm, inrolled, with dense circular patch of orange or brown hairs between claw and blade, sometimes long orange tomentose hairs on outer surface of claw. Upper lateral petals 4 – 8 × 1.5 – 6, long orange tomentose on inner claw. Lower lateral petals 4 – 9 × 1 – 4, long orange tomentose on inner claw. Stamen filaments flattened, 7 – 12 mm long, densely long orange tomentose on the basal ⅓ to ½ of length, on inner surface; anthers ca. 1 mm. Ovary ca. 2 mm long, glabrous in Australia, densely orange hirsute in New Guinea; style 7 – 11 mm long, glabrous or sparsely hirsute along lower part of length; stigma funnel-shaped, 0.5 – 0.75 mm wide, the rim papillate. Fruit coriaceous, suborbicular, the base oblique, the wing very narrow at base, apex beaked; 3 – 5 × 2.4 – 3.4 (– 4) cm, wing 3 – 8 mm wide; fruit surface glabrous; venation reticulate, clearly visible on body of fruit, not visible on wing; dark brown; hypanthium semi-persistent; fruit when old breaking up, the wing detaching from fruit body, and the valves separating. Seeds 1 per fruit; elliptic, ca. 1.5 – 2 × 1.5 cm, not visible in outline on fruit exterior, pale green to brown, smooth. (Fig. 18)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF8EFF80FF11FC8CFA280024.taxon	distribution	Distribution: — Papua New Guinea, Australia. (Fig. 19). Habitat and ecology: — Rainforest; secondary forest; scrub. Phenology: — Papua New Guinea: flowering September, fruiting time unknown. Australia: flowering time May – December, fruiting May – March. Preliminary conservation assessment: — Extent of Occurrence 969,996.884 km 2 = LC; Area of Occupancy 252.000 km 2 = EN. Category assigned: — Least Concern (LC). Taxonomic notes: — The principal differences between C. traceyi as described in the protologue, and M. scortechinii, are the slightly more numerous leaflets (9 – 12 pairs vs. 5 – 9 pairs), slightly more numerous pinnae (7 – 11 pairs vs. 4 – 9 pairs), and smaller flowers (3.5 – 4.5 mm vs. 4 – 8 mm). However, the ranges of variation between the supposed species are contiguous or overlapping for each character. Additionally, the inrolled claw of the median petal of C. traceyi described in the protologue and indicated by Pedley (1997) as being a distinctive character for that proposed species is a character of M. scortechinii and does not distinguish C. traceyi. The morphological evidence overall does not support the existence of two distinct species. Moreover, the range of the specimens formerly named as C. traceyi falls within the range of M. scortechinii, and no ecological difference is known that would define two entities. Therefore, C. traceyi is here placed into synonymy with M. scortechinii. 32 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD5FFD9FF11FC4AFBA00DF0.taxon	materials_examined	CHINA: Central China, Prov. Hupeh, 1885 – 1888, Henry, A. 3113 A (fl.) (K, US); Guangxi, Liuzhou City, Longtan Park, 17 Sep. 1989, Qun, W. E. w 1 50 (fr.) (K); Henry, A. 1122 (fl.) (K, NY); Hupeh Province, Ichang, Henry, A. 3416 (fl.) (K, P); Ichang, Henry, A. 3113 B (fl.) (K); Ichang, Henry, A. 3416 A (fr.) (K); Kwangsi, me-Jon, Seh-Feng, Dar Shan, S. Nanning, 2600 ft., 5 Nov. 1928, Ching, R. C. 8435 (fr.) (US); Kwangsi, Shang-sze District, Kwangtung Border, Tang Lung Village, 26 Oct. 1934, Tsang, W. T. 24516 (fr.) (NY); Lungchow, Morse, H. B. 208 (fr.) (K); Nanto and mountains to northward, Henry, A. 3819 (fl.) (K); South China, Kwong Tung Prov., Teng Woo Mt., 18 Nov. 1916, Levine & Groff. 149 (fr.) (US); Southern Yunnan, between Szemao and Puerhfu, 13 – 15 Mar. 1922, Rock, J. F. 2849 (fl.) (NY, US); Western China, Apr. 1904, Wilson, E. H. 3450 (fl.) (BM, K); Western China, Nanto, Wilson, E. H. 248 (fl.) (K); Western Hupeh, 1907 – 1909, Wilson, E. H. 3256 (fl.) (BM, K, US); Hupeh Province, Ichang, Henry, A. 2238 (fl.) (K, P) LAOS: Phou Phung pres de Louang Prabang, 2 Mar. 1932, Poilane, M. 20257 (fl.) (K, L) MYANMAR: Bhamo District, Lapycke to Sinlum Kabo, 1500 m, 10 Apr. 1912, Lace, J. H. 5769 (fl.) (K) VIETNAM: Ninh Binh Province, Cuc Phuong National Park, 105 36 27 N, 20 21 17 E, 26 Aug. 1999, Cuong, N. M. 464 (fr.) (MO); Indo-China, Tonkin, Jun. 1933, Petelot, A. 4757 (fr.) (NY)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF88FF83FF11F979FA140022.taxon	description	J. Linn. Soc., Bot. 23: 204. 1887. Type: — CHINA: Hupeh, Ichang, Henry, A. s. n. Lectotype: — Henry, A. 3113, K (K 000264688!), Larsen et al. 1980. Synonym. Caesalpinia sinensis (Hemsley) J. E. Vidal. Bull. Mus. Natl. Hist. Nat., ser. 3, 395 (Bot. 27): 90. 1976. Synonym. Mezoneuron sinensis var. parvifolium Hemsley. J. Linn. Soc., Bot. 23: 205. 1887. Holotype: — Hupeh Province, Ichang, Henry, A. 2238, K (K 000264687!), P (P 00751902!). Synonym. Caesalpinia stenoptera Merrill. J. Arnold Arbor. 19: 35. 1938. 28 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK Type: — INDO-CHINA: Tonkin, Jun. 1933, Petelot, A. 4757, NY. Synonym. Caesalpinia tsoongii Merrill. Philipp. J. Sci. 27: 162. 1925. Type: — CHINA: Szechuen, Tsoong 4190. Holotype UC; isotype GH. Synonym. Caesalpinia chinensis Roxb. in Fl. Ind. ed. 2: 361. 1832., nom. rej. Li et al. Taxon 51: 816. 2002. Type: — “ Introduced from China, into the Botanic Garden at Calcutta, where it has now been for these five or six years and has not yet blossomed ” Scandent shrub or vine to 40 m in length. Stems with scattered recurved prickles to 4 mm; glabrous or sparsely whitish to pale orange tomentose. Stipules persistent, triangular, ca. 1 – 3 × 1 – 2.5 mm, glabrous, or the margins tomentose. Leaves with 2 – 4 pairs pinnae; 2 – 4 pairs leaflets per pinna; petiole (1.3 –) 3 – 7 cm; leaf rhachis 7 – 31 [– 40] cm, with paired recurved prickles at pinna insertion points, and scattered on the internodes, sometimes densely; pinnae 2.5 – 12.5 cm, sometimes with recurved prickles in pairs at the leaflet insertion points and sometimes scattered on the internodes; leaf rhachis and petiole glabrous to sparsely whitish to pale orange tomentose; pinna rhachis glabrous to sparsely pale orange tomentose. Leaflets opposite, elliptic, base cuneate to rounded, sometimes oblique, apex acute to rounded; terminal leaflets 3.5 – 10.5 × 1.3 – 5 cm; lateral leaflets 3.5 – 9.6 × 1.5 – 4.7; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, veins faintly visible above, raised and clearly visible below; all leaflets glabrous both surfaces, or lower surface sparsely orange tomentose at base and on midvein, sometimes at margins, glossy above. Inflorescence a panicle, axillary, supra-axillary or terminal, 8 – 42 cm, axes sparsely to densely ferruginous tomentose, sometimes with small recurved prickles; pedicels 4 – 12 (– 14 in fruit) mm, articulated 1 – 2 mm below hypanthium, constricted at articulation, sparsely to moderately ferruginous tomentose, hairs denser at articulation. Bracts caducous, triangular, 0.5 – 2 × 1 – 1.5 mm, glabrous to moderately tomentose; bracteoles caducous, 2 – 3 × 1 mm, elliptic, acute, outer surface moderately to densely orange tomentose, hairs denser at margins, inner surface glabrous; clusters of triangular scale-like bracts sometimes below base of raceme. Flowers with a hypanthium 1 – 2 × 3 – 5 mm, sparsely to moderately ferruginous tomentose; lower calyx lobe 6 – 8 × 3 – 5 mm; other lobes 5 – 6 × 2 – 3 mm; all lobes sparsely to densely pale orange to ferruginous tomentose on inner and outer surface. Median petal obovate, sometimes reflexed backwards, inrolled, 6 – 9 × 3 – 5 mm, including claw 1 – 2 mm long, circular patch of orange hairs between claw and blade, hairs on margins of claw. Upper lateral petals obovate, 5 – 10 × 3 – 6 mm, including claw ca. 1 mm long, petal glabrous or with few hairs on inner surface of claw. Lower lateral petals 5 – 10 × 2 – 6 mm, including claw ca. 1 mm long, petal glabrous or with few hairs on inner surface of claw. Stamen filaments flattened, 5 – 14 mm long, densely orange villous on the basal ½ on inner surface; anther 1 – 2 mm long. Ovary 2 – 5 mm long, sparsely to densely, sometimes partially, orange tomentose, occasionally glabrous, style 7 – 12 mm long, sparsely hairy on basal part of length; stigma funnel-shaped, ca. 1 mm wide, not or only very slightly papillate. Fruit ligneous; suborbicular, straight along upper suture, to lunate, base cuneate to rounded, not stipitate or stipe 1 mm or less, apex with the wing and central vein extending to form a beak 0.1 – 2.5 cm long; (2.5 –) 3 – 5.5 × 1.9 – 3.6 cm × ca. 3 – 5 mm deep, wing 0.5 – 3 mm wide; fruit surface sparsely ferruginous tomentose, hair densest at base and on margins, glabrescent; venation reticulate, raised and clearly visible on fruit body, no venation visible on the wing; hypanthium semi-persistent. Seeds 1 per fruit; ca. 2 cm diameter, not visible in outline on fruit exterior.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF88FF83FF11F979FA140022.taxon	distribution	Distribution: — China, Myanmar, Thailand, Laos, Vietnam. (Fig. 17). Habitat and ecology: — Forest; thicket; limestone. Elevation 100 – 1500 m. Phenology: — China: flowering March – May, fruiting March – October. Vietnam, Laos: flowering March, fruiting June – August. Preliminary conservation assessment: — Extent of Occurrence 837,078.951 km 2 = LC; Area of Occupancy 40.000 km 2 = EN Category assigned: — Least Concern (LC). However, the species occurs within a fairly restricted area, and if found only on limestone then its potential habitat is limited. A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 29	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD5FFD8FF11FA0AFA280024.taxon	materials_examined	INDIA: W. Krauji, 1891, Ridley, H. N. 2105 (fr.) (US) JAVA: Zollinger, 1002, (fl.) (BM, K); Insula Java, Preanger, 9 Apr. 1899, Koorders, S. H. 34442 B (fr.) (L); Tjidadap, Tjibeber, 1918, Winckel s. n. (fl.) (L); Banham, Tjibera, 40 m, 28 Jan. 1919, Backer, C. A. 26522 (st.) (L)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD5FFD8FF11FA0AFA280024.taxon	description	MALAYA: Pahang, 52 m. s. from K. L. towards Kuantan, 20 Feb. 1974, T. & P. 515 (fl.) (K, L); Lenggeng FR. near top of Jelebu Pass, 1100 ft., 27 Mar. 1968, Burgess, P. F. FRI 9126 (fr.) (L); Malacca, Griffith s. n. (fl., fr.) (K); Singapore Bot. Gard. (cult.), 1898, [collector illegible] 9197 (fl.) (K); Malay Peninsula, Prov. Wellesley, near Ara Kuda, Jan. 1900, Curtis, C. 3432 (fl., fr.) (K); Malacca, 1867 – 1868, Maingay, A. C. 534 (fr.) (K, L); Selangor, Upper Ulu Gombak, NW of Karak Highway, 300 m, 27 Mar. 1982, Stone, B. C. 15116 (fr.) (L, MO); Singapore Bot. A TAXONOMIC REVISION OF MEZONEURON Phytotaxa 274 (1) © 2016 Magnolia Press • 71 Gard. (cult.) Feb. 1905, s. coll, s. n. (fl.) (MO); Trengganu, Gunung Padang Expedition Ulu Brang, Camp 1 near K. Lallang, 91 m, 16 Oct. 1969, Whitmore, T. C. FRI 12619 (fl.) (L); Singapore, Bukit Timah Forest Reserve, Fern Valley, 25 Aug. 1951, Sinclair, J. 6887 (st.) (E) NEW BRITAIN: Nantamba, District West New Britain, Subdistrict Hoskins, 5 m, 4 54 S, 151 25 E, Stevens, P. F. & LeLean, Y. LAE 58622 (fr.) (K, L)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD5FFD8FF11FA0AFA280024.taxon	materials_examined	NEW GUINEA: Nunumai, c. 12 km N. of Amazon Bay, Central District, 30 m, 10 11 S, 149 23 E, 18 Jun. 1969, Pullen, R. 7619 (fl.) (K, L); Sudest Isl., Joe Landing, 150 m, 19 Aug. 1956, Brass, L. J. 2775 (fl.) (L, US); Mamberamo river, Jul. 1926, Doctors van Leeuwen, W. M. 9689 (L); New Guinea, Taun L. A. Buolo, Wau Subdistrict, Morobe District, 500 ft., 7 10 S, 146 40 E, 2, 28 Sep. 1969, Kairo, A. 30898 (fl., fr.) (K, L); New Guinea, distr. Hollandia, 70 m, 29 May 1961, van Royen, P. & Sleumer, H. 5637 (fl., fr.) (K, L); Papua, Central District, near Karema, Brown River, 50 ft., 19 Jul. 1962, Schodde, R. 2604 (fr.) (K, L); NW New Guinea, Hollandia, above Polimaeweg, 50 m, 19 Sep. 1956, Versteegh, C. BW 3934 (fl., fr.) (L); West Sepik Province, Ossima turnoff on Vanimo Bewani road, 23 km SSW of Vanimo Bewani Sub Province, 150 m, 2 51 S, 141 13 E, 23 Oct. 1982, Wiakabu, J., Kerenga, K. 50609 (fr.) (E, K, L); Solomon Islands, Kwalo Dolo, Umasami River, 5 miles inland, 400 ft., 03 Jul. 1965, Whitmore, T. C. 6003 (fl.) (L); Guadalcanal, 160 m, 9 28 S, 159 58 E, 17 Jun. 1991, Forster, P. I.; Liddle, D. J. PIF 8611 (fl.) (K); Tenaru River area, 11 km from main road, 9 29 S, 160 2 E, 20 Jun. 1991, Forster, P. I. & Liddle, D. J. PIF 8659 (fl.) (BRI, K) SUMATRA: Sibago, Teijsmann, J. E. s. n. (fl.) (K, L); Korinchi, 3000 ft., Robinson, H. C. & Kloss, C. B. s. n. (fl.) (BM, K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFD5FFD8FF11FA0AFA280024.taxon	description	72 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF82FF8DFF11FD8BFDA80BF1.taxon	description	Hort. Suburb. Calcutt. 245. 1845. Type: — SUMATRA, Campbell, C. s. n., 1797. Specimen untraced. Basionym: — Caesalpinia sumatrana Roxb. Hort. Beng.: 32. 1814. nom. nud.; Fl. Ind. 2: 366. 1832. Synonym. M. sulfureum Miq. Fl. Ind. Bat. 1 (1): 1081. 1855. Type: — JAVA, Zollinger 1002. Holotype U; isotypes BM!, K (K 000264164!), P!. Synonym. M. koordersii Backer ex Koorders-Schumacher. Syst. Verz. i. Fam. 128, 36. 1911. Type: JAVA: Koorders 34442. Holotype BO; isotype L. Synonym. M. peekelii Harms. Bot. Jahrb. 55: 58. 1917. Type: PAPUA NEW GUINEA, New Ireland, Peekel 453 (location of specimen unknown). Woody climber or scandent shrub to 20 m in length. Stems usually with scattered recurved prickles to 5 mm; glabrous, or sparsely whitish puberulous. Stipule flat or raised, oblong, sometimes with acuminate apex, 1 – 2 × 3 – 6 mm. Leaves with 4 – 7 [– 8] pairs pinnae; 2 – 6 pairs leaflets per pinna; petioles 4.5 – 14 cm; leaf rhachis 21 – 43 [– 50] cm, unarmed, or with recurved prickles paired at pinna insertion points and scattered on the internodes; pinnae opposite to subopposite, 7 – 16 cm, pinna rhachis sometimes with recurved prickles paired at leaflet insertion points and sometimes scattered on the internodes; leaf rhachis, petiole, and pinna rhachis all glabrous to sparsely pubescent, with short, erect, pale orange hairs. Leaflets alternate to subopposite; the terminal leaflets obovate, oblong, ovate, elliptic, or rhomboid, base oblique, apex rounded to retuse, 2.9 – 6.4 [– 7] × 1.6 – 3.9 [– 5.5] cm; lateral leaflets oblong to ovate to elliptic to rhomboid, base oblique, apex rounded to retuse, 2.8 – 5.2 × 1.2 – 3.3 cm; all leaflets with upper surface glabrous to very sparsely pubescent, lower surface glabrous to sparsely pubescent, with pale orange, erect hairs, often denser on midvein; 2 ˚ venation anastomosing, 3 ˚ venation reticulate, veins slightly raised above and below. Inflorescence a raceme to few-branched panicle, supra-axillary or terminal, up to 1.1 m long, the axis to 1 cm diameter at base; glabrous, or sparsely orange tomentose; pedicels not articulated, 10 – 15 mm (– 37 in fruit); glabrous. Bracts persistent, occasionally present at base of racemes, oblong, c. 0.75 × 1.5 mm; bracteoles caducous, broadly elliptic, tapering to acute apex, 1.5 – 2 × 0.75 – 1 mm, sparsely ferrugineous tomentose. Flowers with a hypanthium [3 –] 5 – 8 × 4 – 12 mm, glabrous; lower calyx lobe 4 – 14 × 3 – 8 mm, other lobes [2 –] 3 – 7 × 2 – 10 mm, all lobes with few hairs on margins, otherwise glabrous. Median petal blade obovate to orbicular, claw sometimes inrolled, petal 14 – 33 mm long, claw 7 – 21 mm, blade 7 – 12 × 7 – 17 mm; glabrous. Upper lateral petals narrowly obovate, claw attenuate to blade, 15 – 36 × 5 – 15 mm, glabrous. Lower lateral petals narrowly obovate, claw attenuate to blade, 12 – 33 × 3 – 11 mm, glabrous. Stamen filaments flattened, (9 –) 13 – 37 mm long, 0.5 – 2.5 mm wide, glabrous; anthers (0.5 –) 1 – 3.5 mm. Ovary 3 – 10 [– 15] mm long, style [6 –] 13 – 21 mm long, both glabrous; stigma cupular, ca. 0.5 mm wide, the rim papillate. Fruit elliptic, base obtuse, apex with the wing or central vein extending to form a beak; 9 – 19 × 3 – 6 cm, the wing arising basally ca. 2 – 4 mm along the central vein, wing 6 – 18 mm wide; fruit surface glabrous; venation reticulate, raised, and clearly visible on fruit body, anastomosing and faintly visible on the wing; widest part of the persistent, asymmetric hypanthium below the main body of the fruit. Seeds 3 – 8 per fruit; in outline oval and visible on fruit exterior when mature, perpendicular to central vein, ca. 10 × 7 mm, mid brown, smooth, with linear fissures.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FF82FF8DFF11FD8BFDA80BF1.taxon	distribution	Distribution: — Malay Peninsula, Borneo, Sumatra, Java, Papua New Guinea, Solomon Islands, New Britain. (Fig. 9). 18 • Phytotaxa 274 (1) © 2016 Magnolia Press CLARK Habitat and ecology: — Primary and secondary rainforest. Limestone or granite substrate. To 1000 m elevation. Phenology: — Flowering January – September, fruiting January – October. Preliminary conservation assessment: — Extent of Occurrence 7,674,670.082 km 2 = LC; Area of Occupancy 132.000 km 2 = EN. Category assigned: — Least Concern (LC). Field notes: — Petals pink-red.	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
03981525FFAFFFA3FF11FF26FE940A9D.taxon	materials_examined	AUSTRALIA: Mt. Warning, 4 Oct. 1939, Rodway, J. 2962 (fl.) (K); N. S. W., 4 miles NNW of Nimbin, on road between Murillumbah and Lismore, 90 m, 19 Feb. 1971, O’Hara, J., Coveny, R. 3486 (fr.) (K, L); N. S. W., Tweed River, 10 km SW of Murwillumbah, 29 Sep. 1973, Telford, I. R. 3323 (fl.) (CANB, K, L); New South Wales, Alstonville, W. B. 69 (fr.) (BM); New South Wales, Casino, 28 52 S, 153 03 E, Dec. 1910, McAuliffe, D. J. s. n. (fr.) (NSW); New South Wales, Goat Island, Richmond River, Sep. 1916, Cheel, E. s. n. (st.) (NSW); New South Wales, North Coast, Fernleigh Rd, Fernleigh, 28 47 S, 153 29 E, 15 Oct. 1990, Wilson, K. L. 7648 (fl.) (NSW); Queensland, Mar. 1910, Staer, J. s. n. (fr.) (E); Australia, Jan. 1882, Statter, J. W. (fr.) (BM); Queensland, Moreton District, Coolum, 27 Nov. 1960, Trapnell, W. G. s. n. (fl.) (BRI); Emunundi, Nov. 1895, s. coll., s. n. (fl.) (BRI); Queensland, Moreton District, Yarraman Rd, 26 Jul. 1947, Michael, N. 3032 (fl.) (BRI); Queensland, Mudgereeba, Moreton District, 27 Sep. 1930, Hubbard, C. E. 4181 (fl.) (K); S. E. Queensland, Tallebudgera Creek, 23 Nov. 1929, White, C. T. 6592 (fr.) (K, NY); N. S. W., Glen Ugie Peak (Mount Elaine), near Grafton, 28 Oct. 1966, Hayes, H. C. et. al. 2522 (fl.) (K); Queensland, Tomewin Range, Moreton District, Upper Currumbin Creek, 28 1 S, 153 2 E, 17 Oct. 1981, Williams, K. 81241 (fl.) (CANB, NSW); Rukinow (?) River, Moore, C. s. n. (fl.) (K); Wide Bay, Bidwill 107 (fr.) (K)	en	Clark, Ruth P. (2016): A Taxonomic Revision of Mezoneuron (Leguminosae: Caesalpinioideae: Caesalpinieae). Phytotaxa 274 (1): 448-450, DOI: 10.11646/phytotaxa.274.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.274.1.1
