taxonID	type	description	language	source
039987AF8039FFDCFF4B6DA4FCD934D1.taxon	etymology	Etymology. The word titan has its origin in the Greek mythology. The titans were six elder, giant gods, sons of Uranus and Gaia. Titan is also a noun related to one that is gigantic in some attribute. Here, we use the prefix titanto refer to the large size of the species of this new gryllid genus.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8039FFDCFF4B6DA4FCD934D1.taxon	materials_examined	Type species. Titanogryllus salgado Jaiswara, Souza-Dias, Desutter-Grandcolas & de Mello n. sp. Species included. Titanogryllus salgado Jaiswara, Souza-Dias, Desutter-Grandcolas & de Mello n. sp., Titanogryllus oxossi Souza-Dias & de Mello n. sp., Titanogryllus oxente Souza-Dias & de Mello n. sp.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8039FFDCFF4B6DA4FCD934D1.taxon	distribution	Distribution. Brazilian Atlantic Forest, in the states of Espírito Santo and Bahia.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8039FFDCFF4B6DA4FCD934D1.taxon	diagnosis	Diagnosis. Species large, robust, more than 3 cm of body length. Head wide, glabrous. Ocelli almost in line. Pronotum DD saddle-shaped, glabrous, wider than long. TIII subapical spurs 3 / 3; serrulation only anterior to subapical spurs. Male. FWs reaching half of abdomen; anal vein area bulged dorsally; harp crossed with 4 – 6 almost transverse veins; mirror undivided, distinct from apical venation; apical field short, reduced; CuPb short; lateral field with 3 – 5 longitudinal veins. Male genitalia. Pseudepiphallic sclerite elongated; apex of lateral lophi bifurcated; median lophi short; rami elongated, bent towards ventral side, almost connected; ectophallic arc dorsal, thin, almost indistinguishable, below pseudepiphallic medial lophi; ectophallic fold elongated, strongly sclerotized; dorsal cavity short. Female. Larger than males. FWs small, very reduced. Ovipositor short; apex of ovipositor modified, with conspicuous ventral concavity; apex pointed and curved downwards in lateral view, with ventral protuberance.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8039FFDCFF4B6DA4FCD934D1.taxon	description	Description. Species large, robust, more than 3 cm of body length (Figs. 1 A – C, 2 A, 3 A, 4 A, 7 A, B, 8 A, 11 A). Head large, glabrous, wider than long, as wide as pronotum (Figs. 1 A – C, 2 A, D, 3 A, 4 A, B, 7 A, 8 A, 11 A, B). Occiput, vertex and fastigium glabrous (Figs. 2 B, D, F, 4 C, 7 B, C, I, 11 B – D). Epistomal suture angular. Vertex flat. Eyes small, not protruding. Ocelli arranged almost in straight line; median ocellus very slightly ahead of lateral ocelli, distance between lateral ocelli twice the distance between one lateral and median ocellus. Fastigium twice wider than scape (Figs. 2 D, 3 A, 7 B, 11 B). Lower margin of eyes and antennal pit almost at same level. Maxillary palpi short, 3 rd and 4 th joints subequal, 5 th joint longest, regularly widened on its length, apex rounded (Figs. 2 B, 7 I, 11 C). Pronotum DD wider than long, glabrous (Figs. 1 A – C, 2 A, D, 3 A, 4 A, 7 A, B, 8 A, 11 A, B). Tympanum on both sides of TI. TI with 2 ventral spurs and 1 dorsal (inner). TII with 4 apical spurs. Legs III not specially elongated; TIII with spines before subapical spurs; subapical spurs 3 / 3 (Figs. 2 I – K). Double row of spines on basitarsus-III. Male. Metanotum without metanotal structures. FWs covering half of abdomen; stridulatory file present; harp crossed by 4 - 6 almost transverse veins; mirror undivided, wider than long, clearly distinct from surrounding veins; chords 1 and 2 elongated, close to each other, well-separated from chord 3; apical field reduced; lateral field with 3 – 5 longitudinal, parallel veins (Figs. 1 A – C, 2 A, C, E, 4 A, D, 7 A, D – F, 11 A, E). Supra anal plate constricted medially; subgenital plate triangular, apex rounded (Figs. 2 G, H, 7 G, H, 11 F, G). Male genitalia. Male genitalia elongated. Pseudepiphallus: lateral lophi wide, apex bifurcated; medial lophi short; rami elongated, bent towards ventral side, ramus almost connected anteriorly (Figs. 5 A – C, 6 A – C, 9 A – F, 12 A – C). Pseudepiphallic apodemes absent. Pseudepiphallic parameres well sclerotized. Ectophallic invagination: ectophallic apodemes elongated; ectophallic arc dorsal, thin, almost indistinguishable, below pseudepiphallic medial lophi; ectophallic fold elongated, strongly sclerotized (Figs. 5 A – C, 6 A – C, 9 A – C, 12 A – C). Endophallus: endophallic sclerite small, related to ectophallic fold; endophallic cavity reduced (Figs. 5 A – C, 6 A – C, 9 A – C, 12 A – C). Female. Larger than males (Figs. 3 A, 8 A). FW small, very reduced (Figs. 3 B, 8 B). Ovipositor short; apex of ovipositor modified, with conspicuous concavity; apex pointed and curved in lateral view, with ventral protuberance (Figs. 3 A, E – G, 8 A, E – G). Female genitalia. Copulatory papilla small, rounded; base medially extended ventrally; apex membranous (Figs. 5 D – F, 10 A – C).	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF803FFFD7FF4B6DE2FBC73295.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 501927	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF803FFFD7FF4B6DE2FBC73295.taxon	materials_examined	Type locality. Brazil, Espírito Santo State, municipality of Linhares, Reserva Natural Vale.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF803FFFD7FF4B6DE2FBC73295.taxon	etymology	Etymology. Species name is a noun in apposition after the Brazilian humanist photographer Sebastião Ribeiro Salgado Júnior (born 1944).	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF803FFFD7FF4B6DE2FBC73295.taxon	distribution	Distribution. Atlantic Forest, in Espirito Santo State, municipality of Linhares, Reserva Natural Vale (Vale Natural Reserve).	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF803FFFD7FF4B6DE2FBC73295.taxon	materials_examined	Type material. Holotype, allotype, 2 male paratypes, 6 female paratypes. Holotype: male, with genitalia removed and kept with the specimen, labeled “ Brasil — ES [Espírito Santo] — Linhares. 23 - 25. vii / 2012 — Reserva Vale. Projeto SISBIOTA 563360 / 2010 - 0 Conjunto 23 Armadilha C ” (MZSP). Allotype: female, labeled “ Brasil — ES — Linhares. 23 - 25 / vii / 12 — Reserva Vale. Projeto SISBIOTA 5 63360 / 2010 - 0. Conjunto 16. Armadilha E ” (MZSP). Paratypes: 2 males, 6 females. 1 male, labeled “ Linhares, Réserve forestière, Compagnie Vale do Rio Doce, Etat de Espirito Santo, Brésil, 14. xi. 1992, Nuit, fn LD 31, terre nue avec cavités [bare ground with cavities], L. Desutter-Grandcolas (MNHN-EO-ENSIF 7004) ” (MNHN); 1 male labeled “ Fragment ‘ MEME’ S 19 ° 08 ’ 15,8 ’ W 40 ° 05 ’ 04,4 ”, 12. xi. 2000, 40 km NNE Linhares, Espirito Santo, Brésil, forêt semi-décidue (extraction sélective), P. Grandcolas & R. Pellens (MNHN-EO-ENSIF 7005) ” (MNHN); 1 female labeled “ Reserva Natural Vale, Linhares ES [Espírito Santo]. Trilha da Gávea / Farinha Seca. 14. i. 2012. J. P. P. Pena-Barbosa et al. ” (MZSP); 1 female labeled “ Brasil — ES — Linhares. 23 - 25 / vii / 12 — Reserva Vale. Projeto SISBIOTA 563360 / 2010 - 0. Conjunto 30. Armadilha E ” (MZSP); 1 female labeled “ Brasil — ES — Linhares. 23 - 25 / vii / 12 — Reserva Vale. Projeto SISBIOTA 5 63360 / 2010 - 0. Conjunto 1. Armadilha B ” (MZSP); 1 female labeled “ Brasil — ES — Linhares. 23 - 25 / vii / 12 — Reserva Vale. Projeto SISBIOTA 563360 / 2010 - 0. Conjunto 24. Armadilha C ” (MZSP); 1 female labeled “ BR, ES, Linhares, Reserva Vale do Rio Doce. Mata. i- 96. 19 o 09 ’ 10 ” S / 40 o 03 ’ 93 ” W. F. A. G. Mello, S. S. Nihei, leg. ” (BOTU); 1 female labeled “ Linhares — ES [Espírito Santo], viii / 2003 ” (BOTU).	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF803FFFD7FF4B6DE2FBC73295.taxon	diagnosis	Diagnosis. Species large (3 – 4 cm body length), general coloration medium to yellowish brown. TIII inner apical spurs: dorsal longest, median slightly shorter, ventral smaller (iad> iam> iav); TIII outer apical spurs: median longest, dorsal slightly shorter, ventral smaller (oam> oad> oav). Male genitalia: anterior margin of pseudepiphallic sclerite with medial furrow; lateral lophi wide, posterior margin dorsally concave, bifurcated, rounded laterally; medial lophi short, straight, with shallow depression; apex of rami almost connected; pseudepiphallic parameres elongated, apex rounded, ventrally surpassing apex of lateral lophi; ectophallic apodemes elongated, longer than in T. oxossi n. sp. and T. oxente n. sp. Female. Body medium to yellowish brown; copulatory papilla small, rounded; basis convex dorsally, medially extended ventrally; apex membranous, rounded.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF803FFFD7FF4B6DE2FBC73295.taxon	description	Description. In addition to the characters of the genus: Species large (~ 3 cm body length), general coloration medium to yellowish brown (Figs. 1 A, 2 A, 3 A, 4 A, B). Head. Head large, wider than long, medium brown (Figs. 1 A, 2 A, D, 3 A, 4 A). Posterior margin of occiput medium brown, with 2 lines going from occiput to vertex and 2 lines going from occiput to dorsal margin of eyes (Figs. 1 A, 2 A, B, D, 3 A, 4 A). Occiput and vertex glabrous (Figs. 2 B, D, F, 3 A, 4 C). Eyes small, with small unpigmented area on dorsal margin (Figs. 1 A, 2 A, D, 4 A). Fastigium glabrous, wide as vertex, at least 2 x wider than antennal scape (Figs. 2 D, 3 A, 4 A). Antennal scape and antennomeres medium brown. In lateral view, gena medium brown (Fig. 2 B). Three ocelli, almost in line; lateral ocellus larger than in T. oxossi n. sp. and T. oxente n. sp., circular; median ocellus small, flat (Figs. 2 B, D, F, 4 C). Maxillary palpi light brown; apex rounded, whitish (Fig. 2 B). Frons yellowish brown, very short, shorter than clypeus (Figs. 2 F, 4 C). Clypeus, labrum and mandibles yellowish brown (Figs. 2 F, 4 C). Thorax. DD saddle-shaped, glabrous, wider than long, yellowish brown, with medium brown maculae; cephalic margin concave, caudal margin almost straight (Figs. 1 A, 2 A, D, 3 A, 4 A). LL ventro-cephalic angle rounded, ventral margin straight, ventro-caudal angle ascendant (Figs. 2 B, 4 B). Legs. FI, FII light brown; TI, TII medium brown (Figs. 1 A, 2 A, 3 A, 4 A, B). Tympanum on both sides of TI. TI with 2 ventral spurs and 1 dorsal (inner). TII with 4 spurs. Legs III not specially elongated, longer than in T. oxossi n. sp. and T. oxente n. sp. FIII yellowish brown, covered with dark brown pubescence; FIII medium brown in the articulation with TIII (Figs. 1 A, 2 A, 3 A, 4 A, B). TIII and basitarsus – III yellowish brown, with medium brown maculae (Figs. 1 A, 2 A, 3 A, 4 A, B). TIII with spines before subapical spurs; subapical spurs 3 / 3 (Fig. 2 I). Inner apical spurs: dorsal longest, median slightly shorter, ventral smaller (iad> iam> iav) (Fig. 2 K); outer apical spurs: median longest, dorsal slightly shorter, ventral smaller (oam> oad> oam) (Fig. 2 J). Double row of 5 spines on basitarsus – III (Fig. 2 I). Abdomen. Tergites pubescent, medium brown, yellowish brown medially. Cerci medium brown (Figs. 1 A, 2 A, 3 A, 4 A). Male. Legs III longer than body length (Figs. 1 A, 2 A, 4 A, B). Male FWs medium brown, reaching half of abdomen; anal vein area bulged dorsally; harp crossed with 6, almost transverse veins; mirror undivided, distinct from apical venation; apical field very short, almost reduced; CuPb short; lateral field with 4 – 5 longitudinal veins (Figs. 1 A, 2 A, C, E, 4 A, B). Metanotum without metanotal structures. Male supra anal plate pubescent, light brown with medium brown maculae, medially constricted; proximal margin almost straight, distal margin rounded (Fig. 2 G). Subgenital plate pubescent, medium brown, triangular, proximal margin concave, distal margin rounded (Fig. 2 H). Male genitalia. Pseudepiphallus: anterior margin with medial furrow (Figs. 5 A, 6 A); lateral lophi wide, posterior margin dorsally concave, bifurcated, rounded laterally (Figs. 5 A, C 6 A, C); medial lophi short, straight, with shallow depression (Figs. 5 A, 6 A). Rami elongated, apically bent towards ventral side; apex of each ramus almost connected (Figs. 5 A – C, 6 A – C). Pseudepiphallic parameres elongated, apex rounded, ventrally surpassing the apex of lateral lophi (Figs. 5 B, 6 B). Ectophallic invagination: ectophallic apodemes elongated, longer than in T. oxossi n. sp. and T. oxente n. sp., reaching two-third of rami length (Figs. 5 A, B, 6 A, B); ectophallic arc dorsal, thin, amost indistinguishable, below pseudepiphallic medial lophi; ectophallic fold furrowed (Figs. 5 B, 6 B). Endophallus: endophallic sclerite small, related to ectophallic fold (Figs. 5 B, 6 B); endophallic cavity reduced. Female. Species large (body length ~ 4 cm), larger than males (Fig. 3 A). General coloration: body medium to yellowish brown; head, pronotum and legs same colour as males (Fig. 3 A). FW small, very reduced (Figs. 3 A, B). Abdomen glabrous, medium brown (Fig. 3 A). Supra anal plate pubescent, medium brown with light brown maculae, medially constricted; proximal margin almost straight, distal margin rounded (Fig. 3 C). Subgenital plate small, proximal margin almost straight, distal margin medially convex, angles rounded (Fig. 3 D). Ovipositor short, shorter than FIII; apex of ovipositor modified, with conspicuous concavity; apex pointed and curved downwards in lateral view, with ventral protuberance (Figs. 3 E, F, G). Female genitalia. Copulatory papilla small, rounded; basis convex dorsally, medially extended ventrally; apex membranous, rounded, as in Figs. 5 D – F. Measurements (mm). Males (n = 3) – mean (range): BL – 33.63 (32.4 – 35); PL – 5.05 (4.8 – 5.4); PW – 7.45 (7 – 7.7); FWL – 13.33 (13 – 13.5); FWW – 8.41 (8 – 8.7); LFIII – 22.83 (21 – 25); WFIII – 6.5 (6 – 7.5); LTIII – 13.46 (11.4 – 17.5). Females (n = 7) – mean (range): BL – 37.45 (36 – 39.15); HW – 8.57 (7.95 - 9); IOD – 5.01 (4.65 – 5.4); PL – 5.2 (4.65 – 5.55); AWP – 8.22 (7.65 – 8.7); PWP – 6.66 (6.15 – 7.05); PW – 8.16 (7.5 – 8.7); LFIII – 20.31 (19.8 – 21.15); WFIII – 6.21 (6 – 6.75); LTIII – 10.54 (10.05 – 11.1); LBt-III – 4.92 (4.65 – 5.4); OL – 9 (8.4 – 10.05).	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8032FFD0FF4B6BB7FD4530C5.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 501945	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8032FFD0FF4B6BB7FD4530C5.taxon	materials_examined	Type locality. Brazil, Bahia State, municipality of Mucuri.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8032FFD0FF4B6BB7FD4530C5.taxon	etymology	Etymology. Ọ ṣọ ́ ̀ ọ gy ̀ (Oxóssi in portuguese) is an orisha of the Yoruba religion in West Africa, and the canbomblé in Brazil. Oxóssi is the spirit associated with the animals, forests, and hunt. Distribution. Atlantic Forest, in Bahia State, municipality of Mucuri.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8032FFD0FF4B6BB7FD4530C5.taxon	materials_examined	Type material. Holotype, allotype, 1 male paratype. Holotype: male, with genitalia removed and kept with the specimen, and the right leg II missing, labeled “ Brasil, BA [Bahia]. Mucuri, Faz. Farol. 14 - 16 / vii / 1989. 2 - M. 89 / 81. F. A. G. Mello col. ” (BOTU). Allotype: female, with the copulatory papilla removed and kept with the specimen, labeled “ Brasil, BA [Bahia], Mucuri, Fazenda Farol - mata. i- 96. 18 o 04 ’ 01 ’’ S / 39 o 40 ’ 23 ” W. F. A. G. Mello & S. S. Nihei, leg. 5 - M ” (BOTU). Paratype. 1 male with left leg I missing, and the genitalia and FWs removed and kept with the specimen, same data as the holotype (BOTU).	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8032FFD0FF4B6BB7FD4530C5.taxon	diagnosis	Diagnosis. Species large, but smaller than T. salgado n. sp. (~ 3 cm body length), general coloration light to yellowish brown. Male genitalia: anterior margin of pseudepiphallic sclerite without medial furrow; lateral lophi bifurcated, inner projection pointed, spine-like; medial lophi below inner margins of lateral lophi; pseudepiphallic parameres thin, shorter than in T. salgado n. sp., apex rounded. Female. General coloration darker than males: body light brown, head and pronotum medium brown. Copulatory papilla small, rounded; base convex dorsally, with medial invagination, medially extended ventrally; apex membranous, rounded, smaller than in T. salgado n. sp., as in Figs. 10 A – C.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8032FFD0FF4B6BB7FD4530C5.taxon	description	Description. In addition to the characters of the genus: General coloration light to yellowish brown (Figs. 1 B, 7 A, 8 A). Head. Head large, wider than long, yellowish brown (Figs. 7 A, B). Posterior margin of occiput yellowish brown, with 4 lines going from occiput to vertex (Figs. 1 B, 7 A, B). Occiput and vertex glabrous (Figs. 1 B, 7 A, B, C, I, 8 A). Eyes small, with very reduced unpigmented area on dorsal margin (Figs. 7 B, C, I). Fastigium glabrous, wide as vertex, at least 2 x wider than antennal scape (Figs. 1 B, 7 A, B, C, I). Antennal scape and antennomeres light brown (Figs. 1 B, 7 A – C, I). In lateral view, gena yellowish brown (Fig. 7 C). Three ocelli, almost in line; lateral ocellus small, circular; median ocellus small, flat (Figs. 7 C, I). Maxillary palpi light brown; apex rounded, whitish. Frons light brown, very short, shorter than clypeus (Fig. 7 I). Clypeus, labrum and mandibles light brown (Fig. 7 I). Thorax. DD saddle-shaped, glabrous, wider than long, light brown, with medium brown maculae; cephalic margin concave, caudal margin almost straight (Figs. 1 B, 7 A – C, 8 A). LL ventro-cephalic angle and ventral margin rounded, ventro-caudal angle ascendant (Fig. 7 C). Legs. Legs I, II light brown (Figs. 1 B, 7 A). TI with two auditory tympana, two ventral spurs and one dorsal (inner). TII with 4 spurs. Legs III not specially elongated, shorter than body length. FIII light brown, with longitudinal medium brown line; FIII medium brown in the articulation with TIII. TIII and basitarsus – III light brown. Double row of spines on basitarsus – III. TIII with spines before the subapical spurs; subapical spurs 3 / 3. Apical spurs: median longest on both sides, dorsal slightly shorter, ventral smaller. Abdomen. Abdomen light brown, glabrous; cerci light brown (Figs. 1 B, 7 A). Male. Male FWs reaching tergite 4, medium brown; anal vein area bulged dorsally; harp crossed with 4 almost transverse veins; stridulatory vein with 102 teeth (n = 1); mirror undivided, distinct from apical venation; apical field very short, almost reduced; lateral field with 4 - 5 longitudinal veins (Figs. 1 B, 7 A, D – F, 13 A). Metanotum without metanotal structures. Male supra anal plate light brown (Fig. 7 G). Subgenital plate light brown, shieldshaped, proximal margin almost straight, distal margin rounded, extended medially (Fig. 7 H). Male genitalia. Pseudepiphallus: anterior margin without medial furrow (Figs. 9 A, D, F); lateral lophi bifurcated, inner projection pointed, spine-like (Figs. 9 A, B D); medial lophi below inner margins of lateral lophi (Figs. 9 A, D, F). Rami elongated, apically bent towards ventral side; apex of each ramus separated (Figs. 9 – C). Pseudepiphallic parameres thin, shorter than in T. salgado n. sp., apex rounded (Figs. 9 B, E). Ectophallic invagination: ectophallic apodemes shorter than in T. salgado n. sp. (Fig. 9 A); ectophallic arc dorsal, thin, almost indistinguishable, below pseudepiphallic medial lophi; ectophallic fold furrowed, longer than in T. salgado n. sp. and T. oxente n. sp. Endophallus: endophallic sclerite small, related to ectophallic fold (Fig. 9 B); endophallic cavity reduced. Female. Larger than male, shorter than females of T. salgado n. sp. (Fig. 8 A). General coloration darker than males: body light to medium brown; head and pronotum medium brown (Fig. 8 A). FW small, very reduced (Figs. 8 A, B). Abdomen glabrous, yellowish to medium brown (Fig. 8 A). Supra anal plate pubescent, medially constricted, light brown with medium brown small maculae; proximal margin almost straight, distal margin rounded (Fig. 8 C). Subgenital plate small, proximal margin almost straight, distal margin medially convex, angles rounded (Fig. 8 D). Ovipositor short, shorter than FIII; apex of ovipositor modified, with conspicuous concavity; apex pointed and curved in lateral view, with ventral protuberance, as in T. salgado n. sp. (Figs. 8 E – G). Female genitalia. Copulatory papilla small, rounded; base convex dorsally, with medial invagination, medially extended ventrally; apex membranous, rounded, smaller than in T. salgado n. sp., as in Figs. 10 A – C. Measurements (mm). Males (n = 2) – mean (range): BL – 29.77 (28.8 – 30.75); PL – 3.9 (3.75 – 4.05); PW – 6.22 (6 – 6.45); FWL – 9.9 (9.75 – 10.05); FWW – 6.52 (6.3 – 6.75); LFIII – 17.4 (16.5 – 18.3); WFIII – 5.1 (4.95 – 5.25); LTIII – 9.22 (8.7 – 9.75). Female (n = 1): BL – 30.3; HW – 7.35; IOD – 4.2; PL – 4.05; AWP – 7.05; PWP – 5.25; PW – 7.05; LFIII – 17.25; WFIII – 5.1; LTIII – 9.15; LBt-III – 4.2; OL – 8.25.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8035FFCFFF4B6EB3FE813390.taxon	description	http: // lsid. speciesfile. org / urn: lsid: Orthoptera. speciesfile. org: TaxonName: 501949	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8035FFCFFF4B6EB3FE813390.taxon	materials_examined	Type locality. Brazil, Bahia State, municipality of Porto Seguro.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8035FFCFFF4B6EB3FE813390.taxon	etymology	Etymology. Oxente is a common expression used in the daily speech of the inhabitants of Northeastern Brazil. Oxente is an interjection of astonishment, doubt, frustration, impatience, surprise. The word is a noun in apposition.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8035FFCFFF4B6EB3FE813390.taxon	distribution	Distribution. Atlantic Forest, in Bahia State, municipalities of Porto Seguro and Itamaraju.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8035FFCFFF4B6EB3FE813390.taxon	materials_examined	Type material. Holotype, 4 male paratypes. Holotype: male, with genitalia removed and kept with the specimen, labeled “ Campo 0 0 0 3 Mison [Misonete Costa, leg.]. Sul da Bahia (Porto Seguro). 23 / 10 / 2003. (BOTU). Paratypes: 4 males. 1 male, subgenital plate and genitalia removed, labeled “ Brasil (Ba) [Bahia], Trancoso. Sperber, leg. 22. ix. 2006 ” (MZSP). 1 male labeled “ BR, BA [Bahia], Itamaraju. Parque Nacional do Monte Pascoal. 08 - 11. iii. 2012. Dias, P. G. B. S. et al., col. ” (MZSP); 1 male, same data as the previous paratype (BOTU); 1 male labeled “ Brasil (Ba) [Bahia], Trancoso. Sperber, leg. 22. ix. 2006 ” (BOTU).	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8035FFCFFF4B6EB3FE813390.taxon	diagnosis	Diagnosis. Species large (~ 3 cm body length), general coloration medium to dark brown. Male genitalia: anterior margin of pseudepiphallic sclerite without medial furrow; lateral lophi bifurcated, inner margins of distal part touching each other; medial lophi wide, rounded, posterior margin concave; pseudepiphallic parameres not surpassing the apex of lateral lophi, apex pointed.	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
039987AF8035FFCFFF4B6EB3FE813390.taxon	description	Description. In addition to the characters of the genus: General coloration medium to dark brown (Figs. 1 C, 11 A). Head. Head large, wider than long, medium brown (Figs. 1 C, 11 A – D). Posterior margin of occiput medium brown (Figs. 11 A, B). Occiput and vertex glabrous (Figs. 11 A – D). Eyes small, without unpigmented area (Figs. 11 B, C). Fastigium medium brown, glabrous, wide as vertex, at least 2 x wider than antennal scape (Figs. 11 A, B). Antennal scape and antennomeres medium brown (Figs. 11 A – D). In lateral view, gena medium brown (Fig. 11 C). Three ocelli, almost in line; lateral ocellus smaller than in T. oxossi n. sp., circular; median ocellus small, flat (Figs. 11 B – D). Maxillary palpi medium brown; apex rounded, whitish (Fig. 11 C). Frons medium brown, very short, shorter than clypeus and labrum (Fig. 11 D). Clypeus, labrum and mandibles medium brown (Fig. 11 D). Thorax. DD saddle-shaped, glabrous, wider than long, medium brown; cephalic margin concave, caudal margin convex (Figs. 1 C, 11 A, B). LL ventro-cephalic angle rounded, ventral margin straight, ventro-caudal angle ascendant (Fig. 11 C). Legs. Legs I, II light brown. Tympanum on both sides of TI. TI with 2 ventral spurs and 1 dorsal (inner). TII with 4 spurs. Legs III not specially elongated, shorter than body length. FIII medium brown, articulation with TIII dark brown. TIII and basitarsus – III medium brown (Fig. 11 A). Double row of spines on basitarsus – III (Fig. 11 A). TIII with spines before the subapical spurs; subapical spurs 3 / 3. Apical spurs: median longest on both sides, dorsal slightly shorter, ventral smaller. Abdomen. Abdomen medium brown, glabrous (Fig. 11 A). Sternites medium brown. Cerci light brown. Male. Male FWs large, reaching tergite 6, medium brown; anal vein area bulged dorsally; harp crossed with 4 almost transverse veins; stridulatory vein with 56 – 86 teeth (n = 3); mirror undivided, distinct from apical venation; apical field very short, almost reduced; lateral field with 4 - 5 longitudinal veins (Figs. 11 A, F, 13 B). Metanotum without metanotal structures. Male supra anal plate light brown, with medium brown macula; constricted medially, proximal margin slightly concave, distal margin rounded (Fig. 11 F). Subgenital plate medium brown, triangular, proximal margin almost straight, distal margin rounded (Fig. 11 G). Male genitalia. Pseudepiphallus: anterior margin without medial furrow (Figs. 12 A); apex of lateral lophi bifurcated, inner margins of distal part touching each other (Fig. 12 A); medial lophi wide, rounded, posterior margin concave (Figs. 12 A, C). Rami elongated, apically bent towards ventral side; apex of each ramus separated (Figs. 12 A – C). Pseudepiphallic parameres not surpassing the apex of lateral lophi, apex pointed (Fig. 12 B). Ectophallic invagination: ectophallic apodemes shorter than in T. salgado n. sp. (Figs. 12 A, B); ectophallic arc dorsal, thin, almost indistinguishable, below pseudepiphallic medial lophi; ectophallic fold furrowed, longer than in T. salgado n. sp. (Fig. 12 B). Endophallus: endophallic sclerite small, related to ectophallic fold (Fig. 12 B); endophallic cavity reduced. Female. Unknown. Measurements (mm). Males (n = 6) – mean (range): BL – 29.17 (26.7 – 31.65); PL – 4.5 (3.9 – 4.95); PW – 6.65 (6 – 7.2); FWL – 12.17 (10.95 – 13.95); FWW – 7.2 (6.3 – 8.7); LFIII – 18.45 (16.65 – 20.1); WFIII – 5.45 (4.5 – 6.15); LTIII – 10.47 (9.75 – 11.4).	en	Jaiswara, Ranjana, Souza-Dias, Pedro G. B., Campos, Lucas Denadai De, Redü, Darlan R., De Mello, Francisco De A. G., Desutter-Grandcolas, Laure (2018): Titanogryllus n. gen., the largest Gryllinae cricket from the Neotropical Region with three new species from the Brazilian Atlantic Forest (Orthoptera, Grylloidea, Gryllidae). Zootaxa 4402 (3): 487-507, DOI: 10.11646/zootaxa.4402.3.4
