identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0399CF10C439AA58FF7D6584FF2BFCBC.text	0399CF10C439AA58FF7D6584FF2BFCBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diogenes matabiru Rahayu & Pratiwi 2022	<div><p>Diogenes matabiru n. sp.</p><p>Material examined. Holotype, male, 2.7 mm (MZB Cru 5343), Jerowaru, Lombok Island, coll. R. Pratiwi, 29 September 2014 . Paratype, male, 2.5 mm (ZRC 2021.0900), same locality as holotype .</p><p>Comparative material. Diogenes pallescens Whitelegge, 1897 . 1 ovigerous female, 1.8 mm (MZB Cru 5344), Kuta, Lombok, June 2007; 1 ovigerous female, 2.5 mm (MZB Cru 5345), Tanjung Merah, Sulawesi, 31 March 2003 ; 1 male, 2.4 mm, 1 female, 1.3 mm (MZB Cru 5346), Medana, Lombok, 24 July 2006 ; 1 female, 2.0 mm, 1 ovigerous female, 2.7 mm (MZB Cru 5347), Teluk Kodek, Lombok, 20 August 2021 .</p><p>Diogenes biramus Morgan, 1987 . 1 female, 2.7 mm (MZB Cru 5348), Sekotong, Lombok, 16 May 2007 ; 1 female 4.9 mm (MZB Cru 5349), Medana, Lombok, 12 June 2007 ; 1 female, 2.0 mm (ZRC 2021.0901), Pulau Osi, Seram, Maluku, 16 June 1989 ; 1 female, 1.8 mm (ZRC 2021.0902), Medana, Lombok, 2 September 2021 .</p><p>Diogenes viridis Haig &amp; Ball, 1988 . 1 ovigerous female, 2.2 mm, Santo, Vanuatu, 4 October 2006 (MNHN).</p><p>Description. Shield (Figs. 1A, 2A) about 1.2 times as long as broad; anterior margin between rostral lobe and lateral projections somewhat concave; anterolateral angle rounded, margin sloping; posterior margin truncate; dorsal surface with scattered spines, prominent spines along lateral margin, and tufts of sparse setae. Rostrum broadly rounded. Lateral projections triangular, exceeding tip of rostral lobe, each with moderately large marginal spine. Dorsal margins of branchiostegites with row of 4 or 5 small spines.</p><p>Ocular peduncle (Figs. 1A, 2A) slender, cylindrical, approximately as long as shield, inflated proximally, with tufts of very sparse setae dorsoproximally; cornea not dilated, diameter about 0.2 times peduncular length. Ocular acicles broad, subrectangular, basally separated by approximately 0.3 width of 1 acicle, with row of spines on terminal margin, mesialmost spine largest. Intercalary rostral process slender, simple at tip, overreaching tip of mesial spine of ocular acicle, with few long setae; no ventral spine.</p><p>Antennular peduncle (Figs. 1A, 2A), when fully extended, not reaching cornea. Ultimate and penultimate segments glabrous. Basal segment with 2 or 3 small spines on ventrolateral distal margin</p><p>Antennal peduncle (Fig. 1A) reaching 0.5 length of ocular peduncle; fifth segment with 1 long and 1 short setae distally, fourth segment with few short setae distally, third segment short, stout, also with few short setae, second segment with dorsolateral distal angle produced into prominent spine, dorsomesial distal angle unarmed, small spine on dorsodistal margin medially, mesial and lateral margins with few setae, first segments unarmed. Antennal acicle overreaching distal margin of fourth segment, terminating in strong spine; lateral margin with 1 large spine, mesial margin with 3 moderately large spines, and few setae. Antennal flagellum slightly longer than shield, consisting of 16–18 articles, each with 1 or 2 simple setae.</p><p>Third maxilliped (Fig. 1B) with coxa and basis unarmed; crista dentata on ischium obsolete; merus longer than carpus, both segments with long setae on mesial margin, lateral margin with fewer setae; propodus slightly longer than carpus; dactyl shorter than propodus, both with setae ventrally.</p><p>Left cheliped (Figs. 1C, 2A) larger than right, narrow hiatus between dactyl and fixed finger. Dactyl about 0.8 times as long as upper margin of palm, slightly arched; cutting edge with row of blunt calcareous teeth, 1 large tooth proximally, terminating in small calcareous claw, slightly overlapped by fixed finger; outer surface convex with partial covering of spines, and with long stiff setae; upper margin with 2 rows of moderately large spines decreasing in size distally; inner surface slightly convex, with covering tubercles and tufts of short setae. Fixed finger outer surface slightly convex, armed with moderately large sharp and blunt spines and long stiff setae; lower margin with row of moderately small, subacute spines, forming weak curve with lower, similarly armed margin of palm; cutting edge of fixed finger with row of calcareous teeth, large tooth subproximally, terminating in large calcareous claw; inner surface with scattered small tubercles and few tufts of short setae. Palm slightly shorter than carpus, with row of moderately large spines on upper margin, second row shorter and with smaller spines; outer surface convex, with sparse short setae, longitudinal row of sharp, large spines on midline, extending beyond midlength of palm but not reaching articulation with dactyl; additional longitudinal row of small spines between midline and upper margin merging into scattered spines adjacent to articulation with dactyl; lower half of outer surface with sparse spines forming irregular longitudinal rows; inner surface with few small tubercles and short setae. Carpus as long as merus; upper margin with row of irregularly spaced prominent spines and sparse long setae; outer surface convex with longitudinal row of large spine on midline; lower margin with large spine distally and few small spines medially; inner surface with few tubercles and sparse setae. Merus subtriangular in dorsal view; dorsodistal margin with strong spine; dorsal surface with rows of moderately small spines and tufts of long plumose setae; lateral face tuberculate, ventrolateral margin with row of large spines, distal spines strongest and accompanied by long setae; mesial face with sparse tubercles, ventromesial margin with row of moderately large spines, distal spines strongest and accompanied by plumose setae; ventral surface tuberculate. Ischium unarmed, but with sparse short setae on ventromesial margin.</p><p>Right cheliped (Figs. 1D, 2A) reaching proximal third of palm of left cheliped; narrow hiatus between dactyl and fixed finger. Dactyl longer than palm; upper margin with row small spines and numerous long setae; outer surface with scattered small spines and long setae; cutting edge with row of low, blunt, calcareous teeth, terminating in moderately large calcareous claw overlapped by fixed finger. Fixed finger with row of tubercles in midline partially obscured by long setae on outer surface; lower margin smooth but with sparse long setae; cutting edge with row of small calcareous teeth, terminating in moderately large calcareous claw. Palm with row of moderately large spines and tufts of long setae on upper margin; outer surface with longitudinal row of small spines in midline and with long setae. Carpus with row of large spines on upper margin; outer face with row of small spines in midline, lower half of surface with sparse tubercles, inner surface with setae distally. Merus triangular, dorsal margin with 1 large spine distally, and with numerous long, simple or plumose setae; ventrolateral margins with irregular row of small spines, ventromesial margin with few small spines. Ischium unarmed, but with few long setae.</p><p>P2 and P3 (Fig. 2A) slender. P2 (Fig. 1E) with dactyls 1.3 times as long as propodi, somewhat curved ventrally, but not twisted in dorsal view, terminating in moderately small corneous claws; dorsal and ventral margins each with mixture of short and long, plumose and simple setae; mesial surfaces each with row of setae in midline. Propodi 1.4 times as long as carpi, unarmed, each with sparse long and short setae on dorsal and ventral margins; mesial faces each with few setae. Carpi 0.7 times as long as meri; moderately large spine on dorsodistal and dorsoproximal and sparse, long setae; ventral margins unarmed. Meri each with sparse setae on dorsal and ventral margins, otherwise unarmed. Ischia unarmed, each with sparse setae. P3 (Fig. 1F) with dactyls and propodi same as in P2; carpi each with prominent dorsodistal spine, smaller spine on subdistal and subproximal dorsally, and sparse setae, lateral faces and ventral margins unarmed; meri and ischia also unarmed but with few setae. P4 semichelate (Fig. 1G); propodi with tiny dorsodistal spine, carpi with moderately large dorsodistal spine.</p><p>Telson (Fig. 1H) with distinct median cleft, long setae on terminal margin; left posterior lobe longer than right, left terminal margin with row of small spines, stronger spines extending onto lateral margin; right terminal margin also with row of small spines, but not extended onto lateral margin.</p><p>Colour in life. Shield cream with large dark brown spot medially, smaller dark brown patches anteriorly and laterally. Ocular peduncle generally white, greenish brown patch proximally, orangish red band on third distal, blue areas proximal to cornea, cornea light brown. Ocular acicle white with brown spot mesially. Antennular peduncles cream with dark brow ring on ultimate and penultimate segments distally. Antennal peduncles light brown, white band on fifth segment distally. Chelipeds cream, greenish brown on dactyl and fixed finger, palm with dark brown patches, spines white. P2 and P3 cream mottled with light brown, dark brown band on each propodi and carpi medially, meri with dark brown medially and subproximally (Fig. 2A).</p><p>Etymology: matabiru means blue eye in Bahasa Indonesia, alluding to the blue colour of ocular peduncles; use as a noun in apposition.</p><p>Remarks. Within the eight species of the D. pallescens group, four species have the antennular peduncle shorter than the ocular peduncle, namely, D. pallescens, D. biramus, D. viridis, and D. patae . Among these four species, D. matabiru most closely resembles D. pallescens in having strong spines on the shield and the left cheliped, the ocular peduncle as long as the shield, and P4 with distal spine on the dorsal margin of the carpi. However, the shield of the new species is clearly longer than broad (Fig. 1A) (as long as broad in D. pallescens), the intercalary rostral process is very slender, reaching well beyond the tip of the longest spine of the ocular acicle (Fig. 1A) (larger at base to tapering at tip, as long as or only slightly reaching beyond the tip of the ocular acicle in D. pallescens), the left cheliped with outer surface of fixed finger slightly convex, armed with strong spines, outer surface of the palm with row of strong spines on the midline and row of smaller spines between the midline and the upper margin (Fig. 1C), and the ventral margin of the merus of P2 unarmed (Fig. 1E) (outer surface of the fixed finger somewhat flattened, the palm with scattered tubercles or spines and with longitudinal row of strong spines beginning near the midpoint of proximal margin not reaching midlength, and the ventral margin of the merus of P2 with 1 to 5 spines in D. pallescens) (McLaughlin 2002: 91, figs.1–4).</p><p>Diogenes biramus and D. viridis differ in having the ocular peduncles much longer than the shield, the intercalary rostral process is short, triangular, reaching only midlength of the ocular acicle, and the terminal margin of the telson lacks the medial cleft (Morgan 1987; Haig &amp; Ball 1988) (the ocular peduncles as long as the shield, the intercalary rostral process is long, overreaching well beyond the ocular acicle, and the medial cleft of the telson is present in D. matabiru n. sp.). Diogenes patae is different in having dense setae on the chelipeds, P2 and P3 (Asakura &amp; Godwin 2006: fig. 2).</p><p>The life coloration will easily separate D. matabiru n. sp. from D. pallescens, D. biramus and D. viridis . The ocular peduncle of Diogenes matabiru is white, with greenish brown large patch proximally, orangish red broad band on third distal, and a blue area near cornea, the cheliped palm is cream with dark brown patches on dorsolateral and dorsomesial surface (Fig. 2A). In D. pallescens the ocular peduncle is white, light and dark brown patches on proximal half, brown band on distal third, the cheliped palm is cream with dark brown patches proximally on dorsal surface (Fig. 2B). In D. biramus the ocular peduncle is uniformly cream, the cheliped palm is light brown with pinkish brown patches on dorsolateral surface (Fig. 2C), while in D. viridis the ocular peduncle is uniformly light pink, the cheliped chela, palm, and carpus white, the merus green (Fig. 2D). The life colouration of D. patae is unknown.</p><p>Rahayu &amp; Forest (1995) discussed the similarity between D. biramus and D. viridis, and suggested the possibility that the latter is a junior synonym of the former. The respective type localities of each species are not distant: D. biramus was described from Coral Bay, the Northern Territory, Australia, while D. viridis was from Maluku, Indonesia. However, examination of the specimens from Indonesia and Vanuatu revealed several characters that separate these two species. The most distinctive character is the colouration of P2 and P3. In D. biramus P2 and P3 cream, dactyl with light brown band medially, propodus with dark brown band medially and proximally, carpus and merus mottled with light brown, dark brown band medially (Fig. 2C). In D. viridis the dactyls of P2 and P3 are white, propodi, carpi and meri green with small white spots, white band distally (Fig. 2D). Morphological differences of these two species are also remarkable. Diogenes biramus has the following features: shield slightly longer than broad, left cheliped dactyl much longer than palm, fixed finger slightly flattened and lower margin with row of large spines that continued to lower margin of palm, propodus of P4 without any spine, telson subrectangular, broader than long. Diogenes viridis has the following features: shield as long as broad, left cheliped with dactyl as long as palm, fixed finger flattened, deflexed, lower margin with very small spines continued to lower margin of palm, P4 with distal spine on propodus, often also on carpus, telson longer than broad. The differences mentioned above confirmed the validity of these two species.</p><p>Habitat. The species was collected from muddy sandy substrate in mangrove environment, together with D. tumidus .</p></div>	https://treatment.plazi.org/id/0399CF10C439AA58FF7D6584FF2BFCBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Rahayu, Dwi Listyo;Pratiwi, Rianta	Rahayu, Dwi Listyo, Pratiwi, Rianta (2022): Diogenes matabiru, a new species of hermit crab from Lombok Island, Indonesia (Crustacea, Decapoda, Anomura, Diogenidae). Zootaxa 5093 (4): 493-500, DOI: 10.11646/zootaxa.5093.4.7
0399CF10C43DAA5BFF7D6698FA40FCE8.text	0399CF10C43DAA5BFF7D6698FA40FCE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diogenes Dana 1851	<div><p>Key to Indonesian species of Diogenes</p><p>1. Intercalary rostral process well developed and marginally armed with spinules; antennal acicle weakly or strongly bifurcate. 2</p><p>- Intercalary rostral process well developed, reduced, vestigial or absent, marginally unarmed; antennal acicle straight or slightly concave mesially..................................................................................... 5</p><p>2. Outer surface of left cheliped palm with tubercles............................................................ 3</p><p>- Outer surface of left cheliped palm with strong spines........................................................ 4</p><p>3. Antennal acicle with inner branch not reaching midlength of fourth peduncular segment; outer surface of cheliped palm with covering of low, closely-set flattened tubercles..................................... Diogenes custos Fabricius, 1798</p><p>- Antennal acicle with inner branch reaching midlength of fourth peduncular segment; outer surface of cheliped palm with covering of small conical tubercles............................................... Diogenes dubius (Herbst, 1804)</p><p>4. Outer surface of left cheliped with irregular unarmed or weakly spinulous area near upper margin..................................................................................... Diogenes alias McLaughlin &amp; Holthuis, 2001</p><p>- Outer surface of left cheliped with 2 irregular longitudinal rows of strong spines in midline and 1 shorter row near upper margin.................................................................... Diogenes mixtus Lanchester, 1902</p><p>5. Intercalary rostral process absent/vestigial.................................................................. 6</p><p>- Intercalary rostral process well developed.................................................................. 7</p><p>6. Ocular peduncles long and slender, longer than shield length, anterior lobe of sternite of P3 subrectangular.................................................................................... Diogenes jousseaumei (Bouvier, 1897)</p><p>- Ocular peduncles slender, obviously shorter than shield, anterior lobe of sternite of P3 distinctly subquadrate....................................................................................... Diogenes jubatus (Nobili, 1903)</p><p>7. Antennal acicle short, subquadrate, not reaching beyond midlength of fourth peduncular segment...................... 8</p><p>- Antennal acicle elongate, triangular or subtriangular, overreaching distal margin of fourth peduncular segment........... 9</p><p>8. Ocular peduncles slightly inflated proximally and on cornea; subtriangular lobe of telson with distinct median cleft; terminal margin of left lobe armed with large spines, smaller spines on right lobe........ Diogenes dorotheae Morgan &amp; Forest, 1991</p><p>- Ocular peduncles inflated proximally and tapering to narrow cornea; subquadrangular lobe of telson with shallow median cleft; terminal margin armed with tiny spines, with 3 moderately large spines on left lobes........ Diogenes takedai Rahayu, 2012</p><p>9. Antennal peduncle as long as or longer than ocular peduncle; antennal flagella long and with pair long setae on each article 10</p><p>- Antennal peduncle shorter than ocular peduncle; antennal flagella short, lacking pair long setae on each article.......... 16</p><p>10. Antennal peduncles without small spine on fourth peduncular segment.......................................... 11</p><p>- Antennal peduncles with small spine on fourth peduncular segment............................................ 12</p><p>11. Antennal acicle not reaching distal margin of fourth peduncular segment; row of spines on dorsal margin of propodus and carpus of P2.................................................................. Diogenes avarus Heller, 1865</p><p>- Antennal acicle overreaching distal margin of fourth peduncular segment; row of small, often relatively obscure, spines on dorsal margin of carpus of P2 and P3...................................... Diogenes moosai Rahayu &amp; Forest, 1995</p><p>12. Carpus of left cheliped with spinous protuberance medially......................... Diogenes goniochirus Forest, 1956</p><p>- Carpus of left cheliped without spinous protuberance........................................................ 13</p><p>13. Ventrolateral margin of left cheliped merus with row of strong spines............. Diogenes klaasi Rahayu &amp; Forest, 1995</p><p>- Ventrolateral margin of left cheliped merus without row of strong spines........................................ 14</p><p>14. Corneas dilated, dactyls of P2 and P3 slender, long, and curved, carpi with distal spine...... Diogenes foresti Rahayu, 2002</p><p>- Corneas not dilated; dactyls of P2 and P3 slender, carpi with row of spines....................................... 15</p><p>15. Left cheliped palm with continuation of proximal row of tubercles or spines as longitudinal granular ridge in 0.25 of upper outer surface............................................................. Diogenes fasciatus Rahayu &amp; Forest, 1995</p><p>- Left cheliped palm with continuation of proximal row of strong, outwardly directed spines on ventral margin as longitudinal spinose ridge in half of outer surface............................................ Diogenes rectimanus Miers, 1884</p><p>16. Antennular peduncles longer than ocular peduncle.......................................................... 17</p><p>- Antennular peduncle shorter than ocular peduncle.......................................................... 19</p><p>17. Dactyl of right cheliped longer than palm, large hiatus between dactyl and fixed finger, palm without any spines on upper margin.............................................................. Diogenes tumidus Rahayu &amp; Forest, 1995</p><p>- Dactyl of right cheliped same length or shorter than palm, no hiatus between fingers, palm with row of spines on upper margin............................................................................................. 18</p><p>18. Terminal margin of telson obliquely truncate, with row of spinules, without medial cleft... Diogenes leptocerus Forest, 1956</p><p>- Terminal margin of telson with large spines on left lobe, small spines on right lobe, medial cleft distinct................................................................................. Diogenes spinicarpus Rahayu &amp; Forest, 1995</p><p>19. Intercalary rostral process short, triangular, not reaching tip of mesial spine of ocular acicle......................... 20</p><p>- Intercalary rostral process long, reaching or overreaching tip of mesial spine of ocular acicle........................ 21</p><p>20. Shield longer than broad, P4 without any spine, telson broader than long................ Diogenes biramus Morgan, 1987</p><p>- Shield as long as broad, P4 with distal spine on propodus and/or carpus, telson longer than broad............................................................................................... Diogenes viridis Haig &amp; Ball, 1988</p><p>21. Outer surface of fixed finger of left cheliped slightly convex, armed with strong spine; merus of P2 without any spine on ventral margin........................................................................... Diogenes matabiru n. sp.</p><p>- Outer surface of fixed finger of left cheliped flattened, with sparse small spines or tubercles; merus of P2 with 1 to 5 spines on ventral margin.......................................................... Diogenes pallescens Whitelegge, 1897</p></div>	https://treatment.plazi.org/id/0399CF10C43DAA5BFF7D6698FA40FCE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Rahayu, Dwi Listyo;Pratiwi, Rianta	Rahayu, Dwi Listyo, Pratiwi, Rianta (2022): Diogenes matabiru, a new species of hermit crab from Lombok Island, Indonesia (Crustacea, Decapoda, Anomura, Diogenidae). Zootaxa 5093 (4): 493-500, DOI: 10.11646/zootaxa.5093.4.7
