taxonID	type	description	language	source
0399FA61285BFFF4FF2086E1E29F6866.taxon	materials_examined	Holotype. Male, [Indonesia] Sumatra, Prapat, HW 3 [= “ Holzweg 3 ”], 12. VIII. 1983, leg. Dr. Diehl, slide No.: LGN 2158 (coll. HNHM). Paratypes. Indonesia, Sumatra: 1 male, from the same locality as the holotype, 31. I. 1984; 1 male, from the same site, 4. X. 1983; 2 males, from the same site, 28. X. 1983; 1 male, from the same site, 28. IV. 1985; 1 male, Sitahoan, 24. XI. 1981, leg. Dr. Diehl (coll. HNHM); 1 male, Dolok Merangir, 180 m, 2 – 13. VII. 1970, leg. Dr. E. Diehl, slide No.: LGN 2062; 1 male, Doulou, 1250 m, 18. IX. 1969, leg. Dr. E. Diehl, slide No.: LGN 2061 (coll. SMNK).	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61285BFFF4FF2086E1E29F6866.taxon	diagnosis	Diagnosis. The new species resembles externally mostly Nanola basalactifera and N. subbasalactifera (Plate 1, Figs 5 – 8) having very similar forewing pattern. Their typical marking is the light basal part bordered by broad, sharply defined and evenly arcuate reddish-brown stripe in the medial area of forewing included the blackish, also evenly arcuate medial line. Despite this superficial similarity, M. pseudobasalactifera shows no closer relationship with the two Nanola species, the large genital differences (gen. figs 1 – 7) support placing these three species into two distinct genera. Meganola pseudobasalactifera displays closer relationship, according to the configuration of the male genitalia, with M. seima. The new species can be distinguished from M. seima (Plate 1, Figs 1, 2) by its conspicuously larger whitish basal area extending from the costa to the ventral margin and more distally positioned dark medial area (in M. seima the pale basal part is only an ovoid patch in the middle of the wing and the dark area is present only between the antemedial and medial lines). The median fascia of M. pseudobasalactifera is evenly arcuate, sharply defined, and blackish, while that of M. seima is diffuse, poorly visible. In the male genitalia, the new species (gen. figs 1, 2) has, in comparison with M. seima (gen. fig. 3), slightly longer uncus, somewhat narrower valva with shorter and narrower basal dilation, and considerably longer vinculum. The aedeagi of the two species have the same basic configuration, but that is somewhat longer in the new species than in M. seima. Female unknown. Meganola suisharyonensis (Strand, 1917) Roeselia fumosa ab. suisharyonensis Strand, 1917, Archiv für Naturgeschichte 82 A (1): 130. Type-locality: Taiwan, Suisharyo. Two syntypes, in coll. DEI Müncheberg.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612859FFF3FF2083B4E0986AEA.taxon	materials_examined	Type species: Selca indistincta Hampson, 1894, Fauna of British India, Moths 2: 147. Type-locality: [India] [Nagaland] Naga Hills. Holotype: male, in coll. BMNH.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612859FFF3FF2083B4E0986AEA.taxon	diagnosis	Diagnosis. The externally very diverse species of this new genus has been treated so far as Meganola, in spite of their conspicuously different, apomorphic genitalia structures of both sexes. In the hindwing venation, the veins M 1 + R 1 – 5 stalked, and the M 3 + CuA 1 are entirely fused. The diagnostic features of the male genitalia are the characteristically bilobate valva, the much slenderer (or in some cases reduced) uncus, the more uniform, arched and apically acute harpe and the simple, short or mediumlong aedeagus with the vesica lacking cornuti; those of the female genitalia are the poorly sclerotized, usually simple, V-shaped ostium bursae and the much narrower, membraneous ductus bursae (the female genitalia of Meganola have usually much broader, more strongly sclerotized ostium bursae and conspicuously thicker and more sclerotized ductus bursae). The genus Hampsonola displays closer relationship with the South African genus, Vandamia van Son, 1933, due to the similar configuration of their (otherwise dissimilarly bilobate) valvae, but Hampsonola has much slenderer and generally longer uncus (with the exception of those taxa where the uncus is reduced) while the uncus of Vandamia is more robust, thicker and shorter. An additional difference between the two genera that the dorsal valval lobe of Hampsonola lacks hair tuft, while Vandamia has a rather dense hair tuft in the narrowest part of the dorsal lobe of valva.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612859FFF3FF2083B4E0986AEA.taxon	description	Description. Due to the high diversity of the external morphological features observable within the genus (including the body size, wing shape, colouration and wing pattern etc.), a general characterization of the external features of the genus is hardly presented, thus the description is focused on the genitalia characters. Male genitalia. Uncus variable in length, it may be fully reduced (basinigra species-group) or long to very long and slim (indistincta species-group); tegumen with variably developed medial process; valva bilobate, medial incision between dorsal and ventral lobes generally deep, dorsal lobe generally longer than ventral one and apically may be dilated; ventral lobe rounded or triangular, may bear androconial hair scales at its distal margin (sijthoffi and rufa species-groups); dorsal valval lobe with sclerotized costal margin projecting partly to apex; ventral valval lobe membraneous. Harpe well developed, strongly sclerotized, elongate, most often slightly curved, claw- or beak-like; sacculus very short, without processi; vinculum relatively short, apically rounded. Aedeagus generally straight, sometimes S-shaped curved (e. g. in H. rufa, H. longshengensis, H. angustifasciata), relatively short and thin, without apical carinal process, coecum penis rather variable, apically bilobate; vesica without cornuti. Female genitalia. Ovipositor short, relatively broad, conical, apophyses relatively long, thin; ostium bursae relatively broad, simple, V- or U-shaped; ductus bursae narrow in its full length, membraneous, only its short distal section may be somewhat stronger sclerotized; corpus bursae generally large, ovoid, with a pair of remote, generally triangular signa.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61285EFFF1FF2083CCE3C56CD1.taxon	materials_examined	Holotype. Male, [Indonesia] Sumatra, Kebon Balok, NNW of Medan, 24. XI. 1968, leg. Dr. E. Diehl, slide No.: LGN 2057 (coll. SMNK). Paratypes. Indonesia, Sumatra: 1 male, Dolok Merangir, Najaradja, 280 m, 8. XII. 1969, leg. Dr. E. Diehl (coll. SMNK). Malaysia: 1 male, Sabah, Gunung Kinabalu, Sayap, 950 m, 116 ° 34 ’ E, 06 ° 10 ’ N, primary forest edge, in front of ranger house near Sungai Kemantis, 8. III. 2001, at light, J. P. Duffels & M. A. Schouten, slide No. MEV 001; 1 male, from the same area, 1000 m, Gua Malayu trail, clearing in primary forest, 13. III. 2001, J. P. & M. J. Duffels (coll. RMNH).	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61285EFFF1FF2083CCE3C56CD1.taxon	diagnosis	Diagnosis. The new species resembles superficially solely the rather remote species Xenonola limbata (Plate 2, Figs 5, 6; gen. figs 11, 12) due to the unusual and in Hampsonola unique forewing pattern, the extensive blackish inner part of the forewing, but the two species differ in several details of the forewing shape and markings. The most conspicuous differences, besides a number of fine differences in the wing pattern, are the much broader forewing and more expanded blackish part of the wing reaching the tornus in H. diehli, while the blackish forewing area is shorter in X. limbata, being terminated at the proximal two-thirds of ventral margin There are no other similar species known in Meganola or in any Eurasian noline genus. The male genitalia of the new species display, surprisingly, a closer relationship with H. indistincta (Hampson, 1894) (gen. figs 8, 9), according to the following distinctive features: H. diehli has conspicuously shorter, apically rounded uncus (that of H. indistincta much longer, apically pointed), somewhat thicker tegumen without process (H. indistincta has rounded, finger-like process on tegumen), narrower, apically less dilated dorsal and considerably shorter ventral lobe of valva; similarly shaped, but medially less bent harpe; much shorter, broadly rounded vinculum (it has triangular apical process in H. indistincta); the configuration of the aedeagus is practically identical in both species. Female unknown. Hampsonola basirufa (de Joannis, 1928) comb. n. (gen. figs 15, 16) Celama basirufa de Joannis, 1928, Bulletin de la Société Entomologique de France 97: 249. Type-locality: [N Vietnam] Tonkin. Syntypes: two females, in coll. MNHN Paris. PLATE II. Adults of Hampsonola and Xenonola species.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61285DFFF0FF208425E0466C26.taxon	materials_examined	Holotype. Male, China, Prov. N. Yunnan, Li-kiang, 15. VIII. 1934, leg. H. Höne, slide No.: LGN 1948 (coll. ZFMK). Paratypes. China. 1 female, with the same data as the holotype, slide No.: LGN 1949; 1 male, from the same site, 14. VIII. 1934 (coll. ZFMK); 1 female, from the same site, 29. III. 1935, leg. H. Höne, slide No.: LGN 2160 (coll. SMNK).	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61285DFFF0FF208425E0466C26.taxon	diagnosis	Diagnosis. The new species is closely related to H. longshengensis (Hu, Han, László, Ronkay & Wang, 2014), but is easily distinguishable by the following characters: the new species has very conspicuous, arcuate and tapering blackish medial stripe on the whitish-grey forewing, while H. longshengensis has only a pale quadrangular costal patch and the forewing ground colour is somewhat darker, more greyish. In spite of the largely different external appearance, the two species have very similar configuration of male genitalia with the following differences: H. angustifasciata has somewhat shorter apical process of tegumen, considerably narrower, medially less dilated dorsal lobe of valva and slightly longer and thinner harpe compared to those of H. longshengensis. The almost identical configuration of the curiously S-shaped aedeagus also supports the close relationship of the two species. As the female of H. longshengensis is still unknown, the female genitalia of the new species is compared with H. subbasirufa. H. angustifasciata has considerably thicker and shorter apophyses, much narrower V-shaped ostium bursae, somewhat longer ductus bursae and conspicuously larger, claw-like signa bursae compared to those of H. subbasirufa.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61285DFFF0FF2080D5E2C26F02.taxon	materials_examined	Holotype. Male, China, Prov. N. Yunnan, Li-kiang, 7. VIII. 1934, leg. H. Höne, slide No.: LGN 1946 (coll. ZFMK). Paratypes. China, Yunnan. A series of both sexes from the same locality as the holotype, with the following collecting data: 19. VII. 1934, 20. VII. 1934, 3. VIII. 1934, 8. VIII. 1934, 9. VIII. 1934, 10. VIII. 1934, 15. VIII. 1934, 27. VIII. 1934, 13. VIII. 1934, 14. VIII. 1935, 17. VIII. 1935, 18. VIII. 1935, 25. VIII. 1935, 28. VIII. 1935, 1. IX. 1935, 5. IX. 1935, 6. IX. 1935, 9. IX. 1935, slide Nos: LGN 1947, LGN 2021 (females) (coll. ZFMK); 2 males, from the same site, 29. VII. 1935 and 2. VIII. 1935 (coll. SMNK). China, Shaanxi: 1 male, Tapaishan in Tsinling, ca. 3000 m, 10. VIII. 1936, leg H. Höne (coll. ZFMK).	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61285DFFF0FF2080D5E2C26F02.taxon	diagnosis	Diagnosis. Hampsonola subbasirufa is similar externally to H. basirufa (de Joannis, 1928) (Plate 3, Figs 1, 2), due to their similar elongate reddish patch covering the basal area of forewing, extending broadly on the costal margin towards the middle of forewing. The main distinctive external characters are as follows: the new species has graphite-grey head, collar and tegulae, darker, unicolorously grey forewing ground colour with only traces of medial and postmedial lines and the darker, pale greyish hindwing while H. basirufa has bright white head and collar, reddish tegulae, whitish forewing ground colour, sharply defined transverse lines and fine, dense darker suffusion in the terminal area; in addition, the hindwing is paler, whitish-grey. In the male genitalia the new species has, in comparison with H. basirufa (gen. figs 13 – 16), considerably shorter uncus, shorter apical extension of tegumen, narrower and shorter medial dilatation of tegumen, somewhat narrower dorsal lobe of valva, somewhat longer, more elongate harpe, slightly longer apical process of vinculum and somewhat thicker, but similarly simple, short and tubular aedeagus. There are considerable differences in the female genitalia as the new species has broad V-shaped ostium bursae, considerably smaller, thorn-like signa and somewhat thicker apophyses compared to those of H. basirufa, which has narrower U-shaped ostium bursae.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61285DFFFFFF208709E5746E3B.taxon	materials_examined	Holotype. Male, China, Prov. N. Yunnan, Li-kiang, 3. VII. 1934, leg. H. Höne, slide No.: LGN 1971 (coll. ZFMK). Paratypes. China, Yunnan. 2 females from the same site as the holotype, but collected at 16. VIII. 1934 and 12. VII. 1934, slide Nos: LGN 1972, LGN 1988; 1 male, from the same site, 28. VII. 1935, slide No.: Dufay gen prep. 764; 1 female, from the same site, 19. VIII. 1934 (coll. ZFMK).	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61285DFFFFFF208709E5746E3B.taxon	diagnosis	Diagnosis. The closest relative of H. stueningi is H. angustifasciata, the differences between them are the following: the new species has pale grey forewing ground colour with rather broad darker grey medial part bordered by the somewhat sharper defined antemedial and postmedial crosslines, while H. angustifasciata has much paler whitish-grey forewings with darker, almost blackish, and much narrower, tapering medial stripe. Comparing the male genitalia of the two sister-species, H. stueningi has somewhat longer uncus, conspicuously different, shorter and more dilated, apically truncate, more or less triangular dorsal valval lobe (that is evenly rounded in H. angustifasciata) and longer, narrower, apically pointed ventral valval lobe (being rounded in the related species), and considerably shorter harpe. The aedeagus of the new species is curved once only in its basal third, while that of H. angustifasciata is characteristically S-shaped. The rather similar configuration of the female genitalia of H. stueningi and H. angustifasciata supports the close relationship of the two species, with the following distinctive features: H stueningi has considerably thinner and somewhat longer apophyses, funnel-like ostium bursae (that is V-shaped in H. angustifasciata) and somewhat thinner but similarly claw-like signa bursae.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612852FFFCFF208573E2876D4E.taxon	materials_examined	Holotype. Male, Indonesia, Sumatra, Prapat, HW 2, 15 – 19. V. 1985, leg. Dr Diehl, slide No.: LGN 2567 (coll. HNHM). Paratypes. Indonesia, Sumatra. 1 female, from the same locality as the holotype, but collected at 26. V. 1985, slide No.: LGN 2214 (coll. HNHM); 1 female, HW II, 28 km S Pematang Siantar, near Tigadoluk, 1050 m, 02 ° 45 ’ 52 ” N, 099 ° 58 ’ 20 ” E, 10. II. 2002, leg. K. Larsen & M. Fibiger, slide No.: LGN 1654 (coll. ZMUC); 1 male, Dolok Merangir, 180 m, 6. X. - 5. XI. 1970, leg. Dr. E. Diehl, slide No.: LGN 2157 (coll. SMNK).	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612852FFFCFF208573E2876D4E.taxon	diagnosis	Diagnosis. The closest relative of the new species is H. tarkabarka (László, Ronkay & Witt, 2010) (Plate 4, Figs 1, 2), despite the large differences in size (wingspan of H. micra and H. tarkabarka are 13 – 14 mm and 18 – 19 mm, respectively) and wing pattern. Their sister species relationship is indicated by the genitalia characters in both sexes (gen. figs 21 – 24), with a number of distinctive features which are as follows: The new species has, in comparison with H. tarkabarka, somewhat thicker uncus, less dilated tegumen, considerably longer, apically more dilated dorsal, and basally much broader ventral lobe of valva, much thinner, more curved harpe and considerably longer vinculum, while the aedeagi and vesicae of the two related species are practically identical. In the female genitalia, the new species has considerably shorter and thinner apophyses, narrower, ovoid ostium bursae (that of M. tarkabarka is broad U-shaped) and shorter ductus bursae, and a conspicuous quadrangular dilatation at cervix bursae (which is absent in M. tarkabarka). In addition, the configuration of the conspicuous, thorn-like pair of signa is different in the two species: the signa of H. micra are considerably smaller, armed with thinner, more stick-like processes, and the distal one is larger than proximal one, while the signa are larger, broader, more triangular in M. tarkabarka, and the proximal one is bigger than the distal one.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612851FFFBFF2086E1E5206F99.taxon	diagnosis	Diagnosis. The external appearance of the type-species of the new genus is rather unique, resembling somewhat Ctenane labuana (Swinhoe, 1904) (Plate 4, Figs 7, 8; gen. figs 27, 28) due to the presence of the similar, oblique dark forewing medial stripe and the postmedial line which is replaced by a wavy row of blackish dots. This similarity is only superficial, the genitalia of both sexes show the closer relationship of Wittonola not with Ctenane but with the Meganola lineage, according to the robust, long, evenly tapering, apically pointed uncus, the simple, relatively narrow, slightly arcuate valva with parallel margins and the harpe arising rather medially on the basal part of valva, however the very short, button-like harpe is unknown in Meganola. The generic level separation of Wittonola from Ctenane is supported also by the hindwing venation as in Wittonola the vein M 3 + CuA 1 is entirely fused while in Ctenane they are variably long stalked. The configuration of the female genitalia displays further distinctive characters between Wittonola and Meganola. Wittonola has narrow, elongate papillae anales (this phenomenon is also known in certain speciesgroups of Nola) with extraordinarily long apophyses; these characters are unknown in Meganola. The single, thorn-like signum of Wittonola resembles rather to a signum of the Manoba species than to any other Meganola taxa. The genus Wittonola shares certain genitalia characters with Meganola and Manoba (Plate 5, figs 1 – 2 and gen. figs 29 – 30) but, due to the above combination of the external and genital characters, it cannot be unified with any of them. Thus, this lineage is considered here as a distinct genus.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612851FFFBFF2086E1E5206F99.taxon	description	Description. The only known species of the genus is rather small in size with the wingspan 11 – 15 mm, length of forewing 5 – 7 mm. Head and eyes relatively small; antennae bipectinate in male, filiform in female; frons, vertex, tegulae and thorax covered with pale brown scales, collar dark brown. Forewing elongate, apically rounded; ground colour pale brown with reddish sheen; costal margin dark brown from the base to the middle; median area with conspicuous, thick, oblique dark brown stripe; postmedial and terminal areas dark brown; all but one crosslines deleted, postmedial line present but replaced by a row of small dark brown dots. Hindwing pale brown, without discal spot and transverse lines. Male genitalia. Uncus rather long, robust, evenly tapering, apically pointed, tegumen short, relatively broad, valva simple, relatively narrow, slightly arcuate, margins parallel, costal sclerotization rather narrow and weak. Harpe very short, button-like, arising in the middle of basal part of valva, vinculum well developed, pointed, Vshaped. Aedeagus simple, tubular, relatively short and thin, without apical carinal process; vesica with small, finger-shaped, finely scobinated section. Female genitalia. Ovipositor relatively long, very narrow, apically expanded, papillae anales very long, thin, eighth tergite rather short, ostium bursae relatively broad, cup-shaped, ductus bursae medium-long, distal half strongly sclerotized, proximal half membraneous, corpus bursae elongate, distal half gradually broadening, proximal one ovoidal; signum bursae single, relatively large, acute, thorn-like.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612856FFF9FF2085C7E330682D.taxon	materials_examined	Holotype. Female, Cambodia, Mondolkiri Prov., Seima Biodiversity Conservation Area, between Seima and O’Rang, 12 º 12 ’ 12 ” N, 107 º 01 ’ 09 ” E, 300 m, 30. I. 2006, leg. G. Csorba & G. Ronkay, slide No.: LGN 1734 (W 22330) (coll. MWM). Paratypes. Cambodia: 46 specimens of both sexes, with same data as the holotype, slide Nos: LGN 1731 (W 22327), 1733 (W 22329) (males), LGN 1732 (W 22328) (female); 23 specimens of both sexes, Seima Biodiversity Conservation area, between Seima and O'Rang, 360 m, 12 º 15 ’ 44 ” ' N, 107 º 03 ’ 49 ” ' E, 27 – 29. I. 2006, leg. G. Ronkay (coll. MWM); 1 male, with the same data (coll. HNHM). Indonesia, Sumatra: 2 females, Rimbo Panti, 250 m, 00 º 20 ’ 50 ” N, 100 º 04 ’ 07 ” E, 24 – 25. II. 2002, leg. K. Larsen & M. Fibiger, slide Nos: LGN 1638, LGN 1639 (coll. ZMUC).	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612856FFF9FF2085C7E330682D.taxon	diagnosis	Diagnosis. The diagnosis of W. latifasciata is given in detail in the diagnosis of the genus Wittonola. Nanola László, Ronkay & Witt, 2010 (Plate 1, Figs 5 – 8, Plate 5, Figs 3 – 7; gen. figs 4 – 7, 31 – 34) Nanola László, Ronkay & Witt, 2010, Esperiana 15: 38. Type-species: Nanola hluchyi László, Ronkay & Witt, 2010, by original designation. PLATE IV. Adults of Hampsonola, Wittonola and Ctenane species. Taxonomic notes. When the genus was described in the monographic work on the Nolinae fauna of Thailand (László et al. 2010), only the type species was considered as member of the genus, due to the restricted area of that study. The detailed analysis of the genital morphology of Meganola s. l. revealed the existence of a distinct phyletic lineage being associated with Meganola by the former authors. According to the recent investigations, the taxonomic content of Nanola has been extended remarkably, comprising more than a dozen of species. The hindwing venation of the Nanola species is “ quadrifine ”, with the veins M 3 + CuA 1 are stalked. Diagnosis. The main generic characters of this lineage are the relatively short, apically pointed uncus, the generally broadly opened, simple, rather narrow and apically rounded valva, the origin of harpe from the ventral margin of the valval plate as a direct continuation of sacculus, unlike in Meganola, where the harpe is arising more medially from the valval plate (this feature displays closer connections between Nanola and Manoba Walker, 1864), and the usually short, relatively thick, simple aedeagus with or without cornuti in the vesica. The configuration of the female genitalia, at least in those taxa where the female is known, is also rather curious. The main diagnostic features are the extraordinarily thick, strongly sclerotized ductus bursae and the unusal (and very variable) shape of the signum (it is a large quadrangular plate in N. liaoningensis, a long, narrow, evenly arcuate transverse sclerotization in N. franzdanieli, horseshoe-shaped in N. patkosigna and a pair of scobinated rounded plates in N. lesarbena); or even fully reduced like in N. hluchyi). In spite of the rather large external diversity, all species are characterised by the above mentioned combination of genital characters, indicating their close relationship.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612854FFC2FF208426E5106ABD.taxon	materials_examined	Holotype. Female, [Philippines] Luzon, Palali, 2000 m, Benguet, 27. XII. 1912, leg. A. E. Wileman, slide No.: LGN 2013 (coll. BMNH). Paratypes. Philippines: 1 male, Tawi Tawi, Lapid Lapid at Manalik Channel, 19. XI. 1961, Noona Dan Exp. 61 – 62, Caught by Mercury-light 19.00 – 02.00, slide No.: LGN 2133 (coll. ZMUC). Indonesia, Sumatra: 1 male, Dolok Merangir, 12. III. 1981, leg. Dr. Diehl, slide No.: LGN 2175 (coll. HNHM).	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612854FFC2FF208426E5106ABD.taxon	diagnosis	Diagnosis. The closest relative of N. rothschildi is N. hluchyi. The two species display easily recognisable external and genital differences which are as follows: The new species has pale grey head, collar and tegulae, pale brownish forewing ground colour with elongate quadrangular whitish basal patch and narrow, elongate whitish apical patch, continuing in a relatively broad, oblique, whitish line bordered proximally with the dark brownish, diffuse, rather wavy postmedial line towards the middle of the ventral margin of forewing. In comparison, N. hluchyi has whitish head and thorax, much darker brown forewing ground colour with larger, more rounded whitish basal and apical patches, lacking the oblique white line in the postmedial area. Comparing the male genitalia of the two species (gen. figs 31 – 34), N. rothschildi has considerably longer uncus with slightly shorter acute apical process, somewhat shorter medial dilatation of tegumen, somewhat narrower apical part of valva, thicker, more robust, more dentate harpe, and more broad-based, short, triangular cornutus of vesica. In the female genitalia, the new species has narrower, less sclerotized basin-shaped ostium bursae (that of N. hluchyi is much broader, strongly sclerotized and wrinkled, funnel-like), broader, only partially sclerotized, proximally more swollen ductus bursae, in addition, N. rothschildi has, well-developed, finger-like signum bursae, while the related species lacks the signum. PLATE V. Adults of Manoba, Nanola and Fragilonola species. Nanola laevis (László, Ronkay & Ronkay, 2014) comb. n. Meganola laevis László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 34, pl. 7, fig. 5; gen. fig. 10. Typelocality: Vietnam, Fan-si-pan Mts, Sa Pa. Holotype: male, in coll. MWM. Nanola franzdanieli (László, Ronkay & Ronkay, 2014) comb. n. Meganola franzdanieli László, Ronkay & Ronkay, 2014, Fibigeriana Supplement 2: 35, pl. 8, figs 1 – 2; gen. figs 13 – 14. Type-locality: [Vietnam], Tonking. Holotype: female, in coll. MWM.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61286FFFC2FF2081F5E44C6D1E.taxon	materials_examined	Type species: ‘ Meganola’ igorkostjuki László, Ronkay & Witt, 2010, Esperiana 15: 51, pl. 7, fig. 7; gen. fig. 45. Type-locality: North Thailand, Prov. Chiang Mai, 1100 m, 4 km SE of Pang Faen. Holotype: male in coll. MWM.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61286FFFC2FF2081F5E44C6D1E.taxon	diagnosis	Diagnosis. The external appearance of the members of the new genus is rather conspicuous; all known taxa are colorful, tiny species with the wingspan 12 – 16 mm. The forewings of Fragilonola have broad orange-brown or pale brownish basal field, red-brown or greyish-brown terminal area and dark costal margin, the postmedial line is conspicuous, almost straight, oblique, broad, and whitish. Fragilonola species are easily recognizeable due to this combination of characters, while the configuration of their genitalia shows close relationship with Nanola, sharing the characteristic origination of the harpe at the ventral margin of valva in both genera. The main distinctive character between Fragilonola and Nanola is the considerably longer and narrower uncus and the fine but conspicuous, needle-like apical carinal process of the aedeagus of the former genus, while Nanola has generally shorter, more robust uncus and the carina of aedeagus lacks the apical process. In the female genitalia, the differences are even larger as Fragilonola has the longer, thinner, less sclerotized ductus bursae and double signum, while the species of Nanola have rather thick, swollen, strongly sclerotized ductus bursae and a single signum of very variable shape. Comparing the hindwing venation of the two genera, the hindwing of Fragilonola is “ trifine ” with the vein M 3 + CuA 1 being entirely fused, while in Nanola it is “ quadrifine ” as the vein M 3 + CuA 1 is variably long stalked.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61286FFFC2FF2081F5E44C6D1E.taxon	description	Description. Wingspan 12 – 16 mm, length of forewing: 6 – 8 mm. Head relatively small; eyes relatively large; male antenna bipectinate with ciliae gradually shortening towards tip, female antenna filiform. Forewing relatively narrow, apically rounded, ground colour brownish-grey, basal and terminal area lighter grey or orange-brown, costal margin dark brown. Transverse lines deleted, except postmedial line which is rather broad, oblique, more or less straight, and whitish. Hindwing pale grey, without discal spot and transverse lines. Male genitalia. Uncus relatively long, thin, slightly arcuate, apically pointed. Tegumen rather broad, medially slightly dilated. Harpe bifurcate, erected at the ventral margin of valva. Vinculum long and pointed in the type species, relatively short, rounded in F. fragilis. Aedeagus tubular, medium-long, relatively thin, apical carinal process, if present, straight, thorn-like apical; vesica without cornuti but with fine scobination. Female genitalia. Ovipositor short, conical, apophyses medium-long, relatively thin, eighth tergite very short, ostium bursae narrow, cup-shaped, ductus bursae long or relatively short, corpus bursae large, ovoid, signum bursae double, represented by a pair of rather remote, rounded, scobinated plates or short triangular processes.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61286CFFC1FF2081D1E3586F5D.taxon	materials_examined	Holotype. Male, [Indonesia] Sumatra, Sitahoan, 21 – 22. XI. 1981, leg. Dr. Diehl, slide No.: LGN 2154 (coll. HNHM). Paratypes. Indonesia, Sumatra: 1 female, Prapat, HW 3, 31. I. 1984, leg. Dr. Diehl, slide No.: LGN 2156 (coll. HNHM); 2 males, Dolok Merangir, 180 m, collected in 1971, and 5 – 25. XI. 1973, leg. Dr. E. Diehl, slide Nos LGN 2076, LGN 2153 (coll. SMNK).	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61286CFFC1FF2081D1E3586F5D.taxon	diagnosis	Diagnosis. The new species is similar to F. igorkostjuki but is easily distinguished by the following characters: F. fragilis has considerably longer ciliation of male antenna, somewhat shorter forewing, much darker colouration of forewing with less light areas, and conspicuously broader whitish postmedial line. In the male genitalia the new species has considerably thicker uncus, longer tegumen, broader valva, larger, more robust bifurcate harpe with more unequal processes and conspicuously shorter vinculum than in F. igorkostjuki. The aedeagus of F. fragilis lacks the apical carinal process, while the carina of F. igorkostjuki possesses relatively short but well developed thorn-like apical process. The vesica is with fine scobination in both species. The female genitalia of the new species has somewhat shorter apophyses, much broader, more strongly sclerotized ostium bursae, considerably shorter and thinner ductus bursae being sclerotized in its full length (that of F. igorkostjuki is sclerotized only in its distal section), somewhat larger corpus bursae, and the signa are a pair of remote, more or less equally small, short triangular processes, while the signa of F. igorkostjuki are inequal, relatively small, rounded scobinated plates.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61286CFFC1FF2084D4E3B66D15.taxon	materials_examined	Holotype. Male, Philippines, N. Palawan, S. Vicente, 20 km NEE Roxas, 10 º 21 ’ N, 119 º 10 ’ E, 600 m, 12. – 17. I. 1988, leg. Cerny & Schintlmeister, slide No.: LGN 2155 (coll. HNHM).	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61286CFFC1FF2084D4E3B66D15.taxon	diagnosis	Diagnosis. Fragilonola parentela appears as an allopatric sister species of F. igorkostjuki occurring in the Philippines. The two sister species are externally almost identical, the distinctive characters are expressed in the configuration of the male genitalia. F. parentela has, in comparison with F. igorkostjuki, somewhat shorter and thicker uncus, considerably shorter and broader valva, and slightly larger bifurcate harpe with apically rounded arms (these arms are apically pointed in F. igorkostjuki). Last but not least, the aedeagus of the new species is somewhat shorter than that of F. igorkostjuki.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61286CFFCCFF20861DE0B56867.taxon	materials_examined	Type species: Meganola apiensis Holloway, 2003, The Moths of Borneo 18: 30, pl. 1, fig. 20. Type-locality: Borneo, Sarawak, Gunong Mulu NP, Api, 900 m. Holotype: male, in coll. BMNH.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61286CFFCCFF20861DE0B56867.taxon	diagnosis	Diagnosis. The habitus of the type-species of the new genus is unique within the whole tribe. The pale greyishbrown forewing lacks all but one crosslines except the oblique, straight, pale, but well visible brownish postmedial line. The other markings of the wing are conspicuous blackish patches on the basal and medial part of the costa, an oblique row of three dots in the terminal area and a larger patch at the middle of the ventral margin. The hindwing venation is typically „ quadrifine ”, with the M 3 + CuA 1 long stalked. PLATE VI. Adults of Fragilonola and Maculonola species. The basic configuration of the male genitalia indicates the closer relationship of Maculonola rather with Nanola than Meganola according to the rather narrow-elongate valval shape and the configuration of harpe. The main autapomorphic character of the new genus is the presence of a rather curious, long, slim, strongly arcuate process of valva arising basally from the sclerotized costal margin, covered densely with short, fine, hair-like cornuti. This character is unknown in other species of the tribe, being eligible to delineate a distinct genus.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA61286CFFCCFF20861DE0B56867.taxon	description	Description. Wingspan 19 – 20 mm, length of forewing 9 – 10 mm. Head and eyes relatively large, frons, vertex, collar and tegulae covered with pale greyish-brown hair-scales. Forewing elongate, apically rounded, ground colour pale greyish-brown, all but one transverse lines deleted, postmedial line present, as an oblique, straight, pale but well visible, narrow brownish stripe. Forewing costal margin with conspicuous blackish patches at the basal and medial sections and oblique row of three relatively large blackish dots in the terminal area; the mid-section of the ventral margin is marked by a larger patch. Hindwing shining greyish brown, somewhat paler than forewing, discal spot and transverse lines absent. Male genitalia. Uncus short and broad, with relatively long, acute, fine medial process apically. Tegumen narrow, without medial dilatation; valvae long, rather narrow, apically barely dilated, rounded. Harpe arising at the ventral margin of valva, short ribbon-like with parallel margins, slightly arcuate, apically either with short, pointed process arising at the distal margin, or fully truncate. Valva is armed with long, slim, caudally curved process arising basally at the costal margin, covered densely with short, fine hair-like cornuti; vinculum relatively short, rounded. Aedeagus tubular, relatively short and narrow, without apical carinal process, coecum penis conspicuously short; vesica without cornuti. Female unknown.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612861FFCCFF208486E0B56D97.taxon	materials_examined	Holotype. Male, [Indonesia] Sumatra, Dolok Merangir, 3. IX. 1980, leg. Dr. Diehl, slide No.: LGN 2023 (coll. HNHM).	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
0399FA612861FFCCFF208486E0B56D97.taxon	diagnosis	Diagnosis. The new species is a twin species of M. apiensis with easily recognisable external and genital features which are as follows: The forewing ground colour of M. dolokmerangiri is somewhat paler with more brownish shine (M. apiensis is more greyish); the forewing patches of the new species are more blurred, except the large one at the ventral margin, which is larger and more sharply defined than in M. apiensis. In the male genitalia, the new species (gen. fig. 42) has, compared with M. apiensis (gen. fig. 41), slighly narrower valva, fully truncate harpe (that of M. apiensis has short, pointed apical process arising at distal margin) and much longer basal costal process. The aedeagus of the two species are rather similar, but the short coecum penis is quadrangular in the new species, and rounded in M. apiensis. Female unknown.	en	László, Gyula M., Ronkay, Gábor, Ronkay, László (2015): Contribution to the knowledge on the Palaearctic and Oriental taxa of the Meganola s. l. (Lepidoptera, Noctuoidea, Nolidae, Nolinae) generic complex with descriptions of 4 new genera and 11 new species. Zootaxa 4052 (2): 251-296, DOI: 10.11646/zootaxa.4052.3.1
