taxonID	type	description	language	source
03996A093B3EFFFCFF04FBC3FF35FF18.taxon	description	(Figs 1 A – C, 2 A – C)	en	Andersen, Trond, Malla, Annui, Giłka, Wojciech (2024): Two new species of Riethia Kieffer, 1917 from Chile and Ecuador (Chironomidae: Chironominae, Pseudochironomini). Zootaxa 5448 (4): 499-507, DOI: 10.11646/zootaxa.5448.4.3, URL: http://dx.doi.org/10.11646/zootaxa.5448.4.3
03996A093B3EFFFCFF04FBC3FF35FF18.taxon	materials_examined	Type material. Holotype adult male (slide mounted under five coverslips), CHILE, Los Rios Region, Valdivia Province, Panguipulli, El Desague, Lago Riñihue, Limnological field station, 39.774388 ° S 72.453394 ° W, 115 m a. s. l., 5. xi. 1969, leg. F. Reiss (ZSM). Paratypes, 58 adult males as holotype (ZSM, ZMBN).	en	Andersen, Trond, Malla, Annui, Giłka, Wojciech (2024): Two new species of Riethia Kieffer, 1917 from Chile and Ecuador (Chironomidae: Chironominae, Pseudochironomini). Zootaxa 5448 (4): 499-507, DOI: 10.11646/zootaxa.5448.4.3, URL: http://dx.doi.org/10.11646/zootaxa.5448.4.3
03996A093B3EFFFCFF04FBC3FF35FF18.taxon	etymology	Etymology. The epithet, chilensis, is used as an adjective and meaning “ from Chile ” in reference to the country of origin of the holotype. Diagnostic characters. Riethia chilensis differs from its close relative, R. truncatocaudata, by uniformly coloured legs, without bands, and curved, stout superior volsellae without microtrichia dorsally. A light brown abdomen, with darker anterior bands on the tergites, and a superior volsella without microtrichia are the characters that separate R. chilensis from the second new species described here (see below). Both new species + R. truncatocaudata can be separated from all other Riethia by having the inferior volsella strongly bent dorsally, with apical part subtriangular and dorsally covered with broad, flattened, apically fringed setae. See also the diagnosis for the second species.	en	Andersen, Trond, Malla, Annui, Giłka, Wojciech (2024): Two new species of Riethia Kieffer, 1917 from Chile and Ecuador (Chironomidae: Chironominae, Pseudochironomini). Zootaxa 5448 (4): 499-507, DOI: 10.11646/zootaxa.5448.4.3, URL: http://dx.doi.org/10.11646/zootaxa.5448.4.3
03996A093B3EFFFCFF04FBC3FF35FF18.taxon	description	Description. Adult male (n = 5) Body size and proportions. Total length 5.21 – 5.71, 5.44 mm. Wing length 3.03 – 3.23, 3.09 mm. Total length / wing length 1.70 – 1.88, 1.76. Wing length / length of profemur 2.30 – 2.47, 2.35. Colouration. Head pale brown with brown pedicel; thorax brown with pale brown vitae, scutellum, and posterior part of preepisternum; legs brown without darker bands (Fig. 1 B); abdominal tergites brown in anterior ¼, light brown in posterior ¾ (Fig. 1 C). Wing hyaline (Fig. 1 A). Antenna. AR 1.78 – 2.04, 1.91. Terminal flagellomere 760 – 920, 846 µm long. Head. Temporal setae 31 – 36, 34, bi- to tri-serial. Clypeus with 18 – 22, 20 setae. Tentorium 193 – 221, 205 µm long, 57 – 66, 61 µm wide. Stipes 199 – 221, 209 µm long, 12 – 16, 14 µm wide. Palpomere lengths (in µm): 52 – 64, 58; 92 – 104, 97; 164 – 200, 181; 228 – 252, 237; 256 – 332, 289. Third palpomere with 4 – 7, 5 sensilla clavata subapically, longest about 29 µm long. Thorax. Antepronotum with 8 – 12, 9 ventrolateral setae. Acrostichals strong, 9 – 16, 12, in double row; dorsocentrals 11 – 17, 14 in single row; prealars 3 – 5, 4; supraalar 1. Scutellum with 14 – 16, 15 setae in partly double row. Wing (Fig. 1 A). VR 1.06 – 1.12, 1.08. Brachiolum with 3 setae; R with 23 – 31, 28; R 1 with 19 – 24, 21; R 4 + 5 with 32 – 39, 36 setae; other veins and membrane bare. Squama with 10 – 15, 13 setae. Legs. Spur of fore tibia 76 – 86, 81 µm long; spurs of mid tibia 58 – 69, 62 µm and 75 – 94, 82 µm long; spurs of hind tibia 72 – 88, 79 µm and 89 – 115, 99 µm long. Width at apex of fore tibia 75 – 94, 82 µm; of mid tibia 63 – 75, 69 µm; of hind tibia 72 – 79, 76 µm. Lengths and proportions of legs as in Table 1. Hypopygium (Figs 2 A – C). Tergite IX with straight posterior margin, with 17 – 27, 23 setae medially and 34 – 51, 42 setae in two to three rows along posterior margin. Laterosternite IX with 6 – 9, 8 setae. Phallapodeme 154 – 186, 166 µm long; with 41 – 58, 47 µm long, narrow, curved oral projection. Transverse sternapodeme straight, 69 – 97, 79 µm long, with comparatively long oral projections. Gonocoxite 242 – 275, 257 µm long. Superior volsella stout, curved, with long, narrowly triangular apex projecting anteromedially, 80 – 89, 86 µm long, 23 – 32, 26 µm wide medially, with 5 weak dorsal setae, without microtrichia. Inferior volsella strongly bent dorsally with bluntly subtriangular apical part, 54 – 69, 60 µm long, 37 – 44, 40 µm wide, with microtrichia, marginal setae, and 29 – 36, 32 broad flattened, apically fringed setae (Fig. 2 A 1) dorsally. Pseudovolsella consisting of low, bluntly triangular tubercle, with 2 – 3, 3 curved setae, longest seta 48 – 59, 55 µm long. Gonostylus 176 – 209, 194 µm long. HR 1.16 – 1.56, 1.33. HV 2.74 – 2.96, 2.80. Female and immature stages unknown.	en	Andersen, Trond, Malla, Annui, Giłka, Wojciech (2024): Two new species of Riethia Kieffer, 1917 from Chile and Ecuador (Chironomidae: Chironominae, Pseudochironomini). Zootaxa 5448 (4): 499-507, DOI: 10.11646/zootaxa.5448.4.3, URL: http://dx.doi.org/10.11646/zootaxa.5448.4.3
03996A093B3EFFFCFF04FBC3FF35FF18.taxon	distribution	Distribution. The species was collected resting on bushes and other vegetation near a lakeshore in southern Chile.	en	Andersen, Trond, Malla, Annui, Giłka, Wojciech (2024): Two new species of Riethia Kieffer, 1917 from Chile and Ecuador (Chironomidae: Chironominae, Pseudochironomini). Zootaxa 5448 (4): 499-507, DOI: 10.11646/zootaxa.5448.4.3, URL: http://dx.doi.org/10.11646/zootaxa.5448.4.3
03996A093B3BFFFDFF04FEB6FB9AF863.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 58 F 17 D 34 - 5 F 7 A- 47 E 1 - 8954 - 6 BBF 2 FBDA 93 B (Figs 3 A – C, 4 A – C)	en	Andersen, Trond, Malla, Annui, Giłka, Wojciech (2024): Two new species of Riethia Kieffer, 1917 from Chile and Ecuador (Chironomidae: Chironominae, Pseudochironomini). Zootaxa 5448 (4): 499-507, DOI: 10.11646/zootaxa.5448.4.3, URL: http://dx.doi.org/10.11646/zootaxa.5448.4.3
03996A093B3BFFFDFF04FEB6FB9AF863.taxon	materials_examined	Type material. Holotype adult male (slide mounted under five coverslips), ECUADOR, Carchi Province, Reserva ecológica El Ángel, LagUnas El Voladero, 0.691132 ° N 77.882543 ° W, 3.800 m a. s. l., 21. x. 1990, leg. K. Böttger (ZSM). Paratypes, 21 adult males as holotype (ZSM, ZMBN).	en	Andersen, Trond, Malla, Annui, Giłka, Wojciech (2024): Two new species of Riethia Kieffer, 1917 from Chile and Ecuador (Chironomidae: Chironominae, Pseudochironomini). Zootaxa 5448 (4): 499-507, DOI: 10.11646/zootaxa.5448.4.3, URL: http://dx.doi.org/10.11646/zootaxa.5448.4.3
03996A093B3BFFFDFF04FEB6FB9AF863.taxon	etymology	Etymology. The epithet, ecuadoriensis, is used as an adjective and meaning “ from Ecuador ” in reference to the country of origin of the holotype. Diagnostic characters. Riethia ecuadoriensis is a dark brown species, with uniformly coloured abdomen, without bands, except for subrectangular pale brown area on tergite VII, and with scattered microtrichia dorsally on the superior volsella. For other key characters see the diagnosis for R. chilensis given above.	en	Andersen, Trond, Malla, Annui, Giłka, Wojciech (2024): Two new species of Riethia Kieffer, 1917 from Chile and Ecuador (Chironomidae: Chironominae, Pseudochironomini). Zootaxa 5448 (4): 499-507, DOI: 10.11646/zootaxa.5448.4.3, URL: http://dx.doi.org/10.11646/zootaxa.5448.4.3
03996A093B3BFFFDFF04FEB6FB9AF863.taxon	description	Description. Adult male (n = 5) Body size and proportions. Total length 5.42 – 5.87, 5.65 mm. Wing length 3.30 – 3.53, 3.39 mm. Total length / wing length 1.59 – 1.75, 1.67. Wing length / length of profemur 2.72 – 2.90, 2.79. Colouration. Head light brown with dark brown pedicel; thorax and legs brown (Fig. 3 B); abdomen brown, tergite VII with subrectangular pale brown area in posterior 1 / 3 flanked by brown lateral areas (Fig. 3 C). Wing hyaline (Fig. 3 A). Antenna. AR 1.49 – 1.59, 1.54. Terminal flagellomere 817 – 850, 835 µm long. Head. Temporal setae 16 – 19, 18, bi- to tri-serial. Clypeus with 15 – 20, 17 setae. Tentorium 195 – 230, 210 µm long, 59 – 68, 64 µm wide. Stipes 174 – 209, 197 µm long, 12 – 16, 15 µm wide. Palpomere lengths (in µm): 52 – 68, 61; 88 – 100, 92; 152 – 168, 158; 216 – 236, 226; 260 – 300, 285. Third palpomere with 5 – 7, 6 sensilla clavata subapically, longest about 28 µm long. Thorax. Antepronotum with 1 – 4, 2 ventrolateral setae. Acrostichals weak, apparently 1 – 2, 2; dorsocentrals 8 – 12, 10 in single row; prealars 3; supraalar 1. Scutellum with 7 – 11, 9 setae in single row. Wing (Fig. 3 A). VR 0.99 – 1.04, 1.02. Brachiolum with 2 – 3, 3 setae; R with 19 – 25, 23; R 1 with 11 – 15, 13; R 4 + 5 with 21 – 26, 24 setae; other veins and membrane bare. Squama with 14 – 17, 16 setae. Legs. Spur of fore tibia 69 – 79, 74 µm long; spurs of mid tibia 64 – 79, 73 µm and 76 – 86, 81 µm long; spurs of hind tibia 75 – 86, 80 µm and 83 – 94, 89 µm long. Width at apex of fore tibia 58 – 68, 64 µm; of mid tibia 66 – 72, 70 µm; of hind tibia 70 – 79, 75 µm. Lengths and proportions of legs as in Table 2. Hypopygium (Figs 4 A – C). Tergite IX with slightly convex posterior margin, with 48 – 63, 54 setae mainly in two to three rows along posterior margin. Laterosternite IX with 7 – 11, 9 setae. Phallapodeme 131 – 149, 140 µm long; with 35 – 51, 41 µm long, narrow, curved oral projection. Transverse sternapodeme straight, 51 – 68, 59 µm long, with weak oral projections. Gonocoxite 252 – 277, 259 µm long. Superior volsella stout, weakly curved, with narrowly triangular apex projecting medially to anteromedially, 76 – 83, 80 µm long, 32 – 37, 34 µm wide medially, with 6 – 7, 6 weak dorsal setae and scattered microtrichia. Inferior volsella strongly bent dorsally with bluntly subtriangular apical part, 58 – 66, 62 µm long, 41 – 46, 43 µm wide, with microtrichia, marginal setae, and 20 – 25, 22 broad flattened, apically fringed setae (Fig. 4 A 1) dorsally. Pseudovolsella consisting of bluntly triangular tubercle, with single, 52 – 58, 56 µm long, curved seta, with second curved setae at base. Gonostylus 200 – 212, 209 µm long. HR 1.19 – 1.35, 1.24. HV 2.41 – 2.81, 2.67. Female and immature stages unknown.	en	Andersen, Trond, Malla, Annui, Giłka, Wojciech (2024): Two new species of Riethia Kieffer, 1917 from Chile and Ecuador (Chironomidae: Chironominae, Pseudochironomini). Zootaxa 5448 (4): 499-507, DOI: 10.11646/zootaxa.5448.4.3, URL: http://dx.doi.org/10.11646/zootaxa.5448.4.3
03996A093B3BFFFDFF04FEB6FB9AF863.taxon	distribution	Geographical distribution and bionomics. The species is known only from páramo grassland in the high Andes in northern Ecuador, where the males were resting on Espeltia sp. close to a lake.	en	Andersen, Trond, Malla, Annui, Giłka, Wojciech (2024): Two new species of Riethia Kieffer, 1917 from Chile and Ecuador (Chironomidae: Chironominae, Pseudochironomini). Zootaxa 5448 (4): 499-507, DOI: 10.11646/zootaxa.5448.4.3, URL: http://dx.doi.org/10.11646/zootaxa.5448.4.3
