taxonID	type	description	language	source
03A6879FA845FFA9E5BCFB740217658D.taxon	description	(Figures 2 D, E, I, 4 A – E)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA845FFA9E5BCFB740217658D.taxon	diagnosis	Diagnosis Males resemble those of T. rothorum and T. andrewdavisi in having a relatively large apical conductor (Figure 4 A) but can be distinguished by the greatly elevated retrolateral side of the tegulum (Figure 4 B); females are easily recognized by the asymmetrical insemination ducts opening underneath the slit-like atrium (Figure 4 E).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA845FFA9E5BCFB740217658D.taxon	materials_examined	Type material Holotype. ♀, Malaysia, Pahang State, Penang Island (MSNG, examined). Original label: Teutamus politus Thor., Pulo Penang, Doria et Fea. New material Thailand: Krabi Province: Muang District, Thab Khaek – Hang Nark Hill nature trail, 50 – 80 m, semi-evergreen rainforest, 13 June 2008, 1 ♂ (MHNG, THMA- 08 / 07); Ban Khlong Jillat, 6 km west of Krabi City, 60 m, remnant patch of evergreen rainforest at foot of limestone hill, 16 June 2008, 1 ♀ (MHNG, THMA- 08 / 10); Khao Panom Benja NP, 100 – 190 m, evergreen rainforest, 11 October 1999, 4 ♂, 2 ♀ (MHNG, THA- 99 / 1); Khlong Thom District, Khao Pra Bang Khram Wildlife Sanctuary, 80 m, semi-evergreen rainforest, 2 – 3 June 2009, 1 ♀ (MHNG, TH- 09 / 04); Aou Luk District, limestone hill c. 1 km east of Aou Luk Tai, 50 m, semi-evergreen rainforest, 21 June 2008, 3 ♀ (MHNG, THMA- 08 / 11); 6 km south of Aou Neua, 50 m, swamp forest, 11 June 2008, 3 ♂, 1 ♀ (MHNG, THMA- 08 / 05); mountain east of Aou Luk Neua, 60 – 90 m, semi-evergreen rainforest on limestone, 12 September 2007, 1 ♂, 4 ♀ (MHNG, TH- 07 / 08); Aou Luk District, Than Boke Thoranee NP, 50 m, semi-evergreen rainforest, 18 – 19 November 2006, 1 ♀ (MHNG, TH- 06 / 06); limestone hill c. 1 km east of Aou Luk Tai, 60 – 80 m, remnant of semi-evergreen rainforest, 9 – 10 June 2009, 4 ♂, 2 ♀ (MHNG, TH- 09 / 08); Aou Luk District, limestone hill c. 1 km east of Aou Luk Tai, 60 – 80 m, semi-evergreen rainforest, 9 – 10 June 2008, 4 ♂, 3 ♀ (MHNG, THMA- 0 8 / 04); 15 km north of Krabi City, near Saeng-Phet Cave, 80 m, evergreen forest at foot of limestone hill, 24 July 2005, 1 ♂ (MHNG, TH- 05 / 1 1). Phuket Province: Siray Island, 30 m, old secondary forest, 28 September 2006, 1 ♂, 2 ♀ (MHNG, TH- 06 / 11); 20 July 2007, 7 ♂, 1 6 ♀ (MHNG, TH- 07 / 12); Khao Pra Taew, Tone Sai Waterfall, 18 November 2003, 1 ♀ (MHNG, Teu- 34234). Phang Nga Province: Yao Noi Island, 150 m, semi-evergreen rainforest near stream, 21 – 22 September 2006, 2 ♀ (MHNG, Teu- 12). Malaysia: Kedah State: Penang Island, Penang Hill, 710 m, evergreen rainforest, 8 – 9 December 1997, 6 ♂, 4 ♀ (MHNG, Teu- 12). Langkawi Island, Raya Hill, northeastern side, 300 – 500 m, 29 – 30 December 2004, 3 ♂, 1 ♀ (MHNG, AS- 04 / 28); 1 – 7 January 2005, 3 ♂, 3 ♀ (MHNG, AS- 04 / 29); 700 – 800 m, 31 August 2004, 1 ♀ (MHNG, AS- 04 / 18). All specimens leg. P. J. Schwendinger.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA845FFA9E5BCFB740217658D.taxon	description	Description Male (MHNG, THA- 99 / 1). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 9 r 9; metatarsi p 5 r 5. Leg II: Tibiae p 8 r 8; metatarsi p 5 r 4. Carapace reddish brown; submarginal lobes blunt, slightly elevated (Figure 2 D). Sternum reddish brown. Legs hairless, bi-coloured: Femora to tarsi brown, other segments yellowish brown. Opisthosoma elongate-ovoid. Dorsal scutum reddish brown, covering three-quarters of length of opisthosoma. Palp (Figure 4 A – C): Palpal tibia relatively short. RTA triangular, its apex blunt. DTA short and stout, weakly sclerotized. Tegulum ovoid, strongly protruding meso-retrolaterally, with field of pale, transparent area. Conductor triangular, broad at base, its apex blunt, curving backward. Embolic base well-developed, clearly visible. Female (holotype, MSNG). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 10 r 10; metatarsi p 6 r 6. Leg II: Tibiae p 9 r 9; metatarsi p 5 r 5. General appearance as in male except submarginal lobes on carapace broader at base, their apices blunt; dorsum of opisthosoma pale, posteriorly with a series of faint chevrons (Figure 2 E). Venter pale. Genitalia (Figure 4 D, E): Epigyne with deep cleft on left side. Copulatory orifices opening on crescent-shaped atrium. Insemination ducts asymmetrical, connected to middle portion of spermathecae on their ventral surfaces. Secretory ampullae digitiform, originating sub-proximally on insemination ducts. Spermathecae elliptic. Fertilization ducts lanceolated.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA845FFA9E5BCFB740217658D.taxon	distribution	Distribution Southern Thailand and west coast of the Thai-Malay Peninsula.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA848FFAFE629FF0303C4672E.taxon	description	(Figures 1 C, 5 A – E)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA848FFAFE629FF0303C4672E.taxon	diagnosis	Diagnosis Males of T. rama sp. nov. bear close resemblance to those of T. fertilis (Figure 4 F) but can be distinguished by the triangular dorsal flange on the conductor (Figure 5 D, E), the excavation on pro-apical and baso-retrolateral sides of the tegulum (Figure 5 A), and the globular submarginal lobes on the carapace (Figure 1 C); females can be recognized by the parallel-sided spermathecae, the short insemination ducts and the spatulate secretory ampullae (Figure 5 G).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA848FFAFE629FF0303C4672E.taxon	materials_examined	Type material Holotype. ♂, Thailand: Nakhon Sri Thammarat Province: Khao Luang NP, near Karom Waterfall, 380 m, rainforest, 17 July 2005, leg. P. J. Schwendinger (MHNG, TH- 05 / 07). Paratypes. Khao Luang NP, Ai Khieo Waterfall, 190 – 250 m, semi-evergreen rainforest, 25 November 2001, 4 ♂, 1 ♀ (MHNG, THMA- 01 / 03). Krabi Province: Ao Luk District, limestone hill c. 1 km east of Ao Luk Tai, 60 – 80 m, semi-evergreen rainforest, 9 – 10 June 2008, 4 ♂, 3 ♀ (MHNG, THMA- 08 / 04). Pattani Province: Nam Tok Sai Khao NP, Sai Khao Waterfall, c. 20 km northwest of Yala City, 260 – 350 m, evergreen forest near a stream, 22 October 1999, 2 ♂, 1 ♀, (MHNG, THA- 99 / 4); 23 October 1999, 3 ♂, 4 ♀ (MHNG, THA- 99 / 5). Malaysia: Perak State: Pangkor Island, 30 – 150 m, evergreen rainforest, Ipoh, 15 – 16 December 1997, 1 ♂, 1 ♀ (MHNG, 16). All specimens leg. P. J. Schwendinger.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA848FFAFE629FF0303C4672E.taxon	etymology	Etymology The specific epithet is an eponymous patronym (nominative form of a noun) in honour of His Majesty King Bhumibol Adulyadej the Great of Thailand, King Rama IX on the auspicious celebration of His Majesty’s 84 th Birthday in December, 2011.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA848FFAFE629FF0303C4672E.taxon	description	Description Male (holotype). Total length 4.82; prosoma 2.30 long, 1.52 wide; opisthosoma 2.52 long, 1.16 wide. Leg formula: 1423; I 10.76 (2.42, 0.98, 3.42, 2.36, 1.58); II 8.52 (2.00, 0.98, 2.64, 1.82, 1.08); III 6.88 (1.56, 0.82, 2.04, 1.54, 0.92); IV 10.22 (2.52, 0.92, 2.92, 2.50, 1.36). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 9 r 9; metatarsi p 5 r 5. Leg II: Tibiae p 7 r 7; metatarsi p 4 r 6. Carapace dark reddish brown; submarginal lobes triangular, their apices globular (Figure 1 C). Sternum dark reddish brown. Legs hairless, bi-coloured: Femora to tarsi orange-brown, other segments brown. Opisthosoma elongate-ovoid. Dorsal scutum brown, covering almost entire length of opisthosoma. Palp (Figure 5 A – E): RTA broad in ventral view, its apex blunt, anterior margin with weakly sclerotized, transparent lamina. DTA digitiform. Tegulum slightly excavated on pro-apical and baso-retrolateral sides. Conductor triangular, its apex sharply pointed, provided with triangular dorsal flange. Embolic base clearly visible. Female (paratype, MHNG, THMA- 01 / 03). Total length 5.62; prosoma 2.50 long, 1.52 wide; opisthosoma 3.12 long, 1.54 wide. Leg formula: 1423; I 11.30 (2.52, 0.94, 3.84, 2.52, 1.48); II 8.56 (1.96, 0.94, 2.92, 1.62, 1.12); III 7.32 (1.62, 0.84, 2.10, 1.70, 1.06); IV 10.80 (2.52, 0.94, 3.12, 2.72, 1.50). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 10 r 10; metatarsi p 6 r 6. Leg II: Tibiae p 9 r 9; metatarsi p 5 r 5. General appearance as in male except submarginal lobes on carapace broader at base, their apices blunt; dorsum of opisthosoma pale. Genitalia (Figure 5 F, G): Epigyne with widely separated circular copulatory orifices situated anteriorly. Insemination ducts short and curved, connected to anterior lateral portion of spermathecae. Secretory ampullae spatulated, originating terminally on insemination ducts. Spermathecae elongated, lateral margins parallel-sided, situated close to one another, anteriorly with large apical glandular pore. Fertilization ducts lanceolated.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA848FFAFE629FF0303C4672E.taxon	distribution	Distribution Southern Thailand and Malaysia (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA84EFFADE5D3FD63022D6649.taxon	description	(Figures 3 A, B, G, 6 A – F)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA84EFFADE5D3FD63022D6649.taxon	diagnosis	Diagnosis Males of T. serrulatus sp. nov. can be easily distinguished by a row of serrated denticles on baso-prolateral side of the conductor (Figure 6 B, D); females can be recognized by the large, circular epigynal atrium (Figures 3 A, 6 E) and the simple internal genitalia without clear boundary between insemination ducts and spermathecae (Figure 6 F).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA84EFFADE5D3FD63022D6649.taxon	materials_examined	Type material Holotype. ♂, Malaysia: Johor State: Mt Arong, 20 m, rainforest 15 km north of Mersing, 29 May 2004, leg. P. J. Schwendinger (MHNG, MAL- 04 / 06). Paratypes. Data as holotype, 1 ♂, 1 ♀ (MHNG, MAL- 04 / 06). Pahang State: Cameron Highlands, 1600 m, trail 14, Mt Mentiga, sifting at the summit, 23 March 1993, leg. I. Löbl and F. Calame, 1 ♀ (MHNG, WM- 93 / 16 a).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA84EFFADE5D3FD63022D6649.taxon	etymology	Etymology The specific epithet is a nominative form of an adjective, derived from Latin (serrulatus, - a, - um = with small teeth), referring to the serrated prolateral margin of the conductor.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA84EFFADE5D3FD63022D6649.taxon	description	Description Male (holotype). Total length 4.84; prosoma 2.32 long, 1.52 wide; opisthosoma 2.52 long, 1.16 wide. Leg formula: 1423; I 10.80 (2.46, 0.96, 3.46, 2.34, 1.58); II 8.52 (1.96, 0.94, 2.66, 1.84, 1.12); III 7.00 (1.60, 0.86, 2.06, 1.54, 0.94); IV 10.38 (2.56, 0.96, 2.96, 2.54, 1.36). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 9 r 9; metatarsi p 5 r 5. Leg II: Tibiae p 8 r 8; metatarsi p 5 r 4. Carapace reddish brown; submarginal lobes triangular, their apices globular. Sternum dark reddish brown. Legs hairless, bi-coloured: Femora to tarsi yellowish brown, other segments orange-brown. Opisthosoma elongate-ovoid. Dorsal scutum reddish brown, covering two-thirds of length of opisthosoma. Palp (Figure 6 A – D): RTA digitiform in ventral view, strongly excavated medially, its apex blunt. DTA digitiform, weakly sclerotized, provided with small, elevated lump at base. Tegulum protruding apico-retrolaterally, strongly excavated on baso-retrolateral side, with colourless and transparent area. Conductor triangular in ventral view, columnar with sharply pointed apex in prolateral view, its prolateral margin strongly serrated. Embolic base clearly visible. Female (paratype, MHNG, MAL- 04 / 06). Total length 5.68; prosoma 2.52 long, 1.54 wide; opisthosoma 3.16 long, 1.56 wide. Leg formula: 1423; I 11.44 (2.56, 0.94, 3.88, 2.54, 1.52); II 8.72 (2.02, 0.94, 2.96, 1.64, 1.16); III 7.54 (1.66, 0.86, 2.14, 1.76, 1.12); IV 10.86 (2.54, 0.92, 3.14, 2.72, 1.54). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 10 r 10; metatarsi p 6 r 6. Leg II: Tibiae p 10 r 9; metatarsi p 5 r 5. General appearance as in male except legs orange-brown, uniform in colour; dorsum of opisthosoma greenish, provided with a series of pale chevrons. Genitalia (Figures 3 A, B, G, 6 E, F): Epigyne with large, circular atrium filled with dark, hardening secretion. Copulatory orifices elongate-ovoid, situated anteriorly on epigynal atrium. Boundary between insemination ducts and spermathecae indistinct. Secretory ampullae digitiform, originating on inner surface of insemination ducts. Spermathecae elongated, with glandular pore situated posteriorly on dorsal surface. Fertilization ducts lanceolated.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA84EFFADE5D3FD63022D6649.taxon	distribution	Distribution Known from localities on the Malay Peninsula (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA84CFFB3E5CDFCC4022C6670.taxon	description	(Figures 1 E, 2 F, 7 A – F)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA852FFB1E5D2FC8D023367B8.taxon	description	(Figures 3 D, F, 8 A – F)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA852FFB1E5D2FC8D023367B8.taxon	diagnosis	Diagnosis Males of T. apiculatus sp. nov. closely resemble those of T. fertilis but can be distinguished by the peculiar shape of conductor (Figure 8 A, D); females can be recognized by the zigzagged insemination ducts and the apical glandular pore on the spermathecae (Figures 3 D, 8 F).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA852FFB1E5D2FC8D023367B8.taxon	materials_examined	Type material Holotype. ♂, Malaysia: Pahang State: Cameron Highlands, Orang Asil, village near Mt Perdah, 1575 m, sifting leaf litter in shallow ravine, 2 – 14 May 2009, leg. P. Banar (MHNG, MAL-PB 5). Paratypes. Data as holotype, 1 ♂, 2 ♀ (MHNG, MAL-PB 5). Fraser Hill, south side, 1300 m, 26 August 2004, 2 ♀ (MHNG, MAL- 04 / 10).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA852FFB1E5D2FC8D023367B8.taxon	etymology	Etymology The specific epithet is a nominative form of an adjective (Latin: apiculatus, - a, um), referring to the large glandular pore locating on the apex of spermathecae.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA852FFB1E5D2FC8D023367B8.taxon	description	Description Male (holotype). Total length 5.06; prosoma 2.42 long, 1.62 wide; opisthosoma 2.64 long, 1.30 wide. Leg formula: 1423; I 11.06 (2.52, 0.98, 3.48, 2.42, 1.66); II 8.76 (2.02, 0.98, 2.72, 1.88, 1.16); III 7.42 (1.66, 0.92, 2.16, 1.64, 1.04); IV 10.78 (2.62, 1.04, 3.06, 2.62, 1.44). Carapace reddish brown; submarginal lobes triangular, their apices bluntly pointed. Sternum reddish brown. Legs hairless, orange-brown. Opisthosoma elongate-ovoid. Dorsal scutum covering two-thirds of length of opisthosoma. Palp (Figure 8 A – C): RTA triangular, strongly excavated medially, its apex blunt, anterior margin with colourless and transparent lamina. DTA short and stout. Tegulum ovoid, slightly excavated on baso-retrolateral side. Conductor parallel-sided, abruptly narrowing towards apex. Embolic base clearly visible. Female (paratype, MHNG, MAL-PB 5). Total length 5.02; prosoma 2.38 long, 1.62 wide; opisthosoma 2.64 long, 1.30 wide. Leg formula: 1423; I 11.08 (2.48, 1.04, 3.52, 2.38, 1.66); II 8.78 (2.02, 1.02, 2.68, 1.88, 1.18); III 7.40 (1.68, 0.92, 2.14, 1.62, 1.04); IV 10.66 (2.58, 1.04, 3.06, 2.60, 1.38). General appearance as in male except submarginal lobes on carapace broader at base, their apices blunt; dorsum of opisthosoma pale, posteriorly with a series of faint chevrons. Genitalia (Figures 3 D, F, 8 E, F): Epigyne with shallow median depression situated between crescent-shaped copulatory orifices. Insemination ducts zigzagged, connected to anterior lateral portion of spermathecae on ventral surface. Spermathecae elongate-ovoid, widest subapically, glandular pore situated apically. Fertilization ducts lanceolated.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA852FFB1E5D2FC8D023367B8.taxon	distribution	Distribution Known only from the type localities on the Malay Peninsula.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA850FFB7E624FCD5024D66B1.taxon	description	(Figures 1 F, 2 G, 9 A – E)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA850FFB7E624FCD5024D66B1.taxon	diagnosis	Diagnosis Males of T. lioneli sp. nov. can be recognized by the presence of a circular pit on retrolateral side of the tegulum (Figure 9 A – C), the relatively small RTA and the triangular, sharply pointed submarginal lobes on the carapace (Figures 1 D, 2 G); females resemble those of T. jambiensis but can be distinguished by the curved insemination ducts connecting to anterior portion of the reniform spermathecae instead of middle portion (Figure 9 E).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA850FFB7E624FCD5024D66B1.taxon	materials_examined	Type material Holotype. ♂, Malaysia: Pahang State: Tioman Island, Mt Kajang, 900 – 1000 m, 27 April 2005, leg. A. Schulz (MHNG, AS- 05 / 08). Paratypes. Data as holotype, 1 ♀ (MHNG, AS- 05 / 08); Tioman Island, path from Mt Paya to Mt Kajang, 200 – 600 m, 6 June 2007, leg. A. Schulz, 1 ♀ (MHNG, ASWM- 0 7 / 7); Tioman Island, path from Mt Paya to Mt Kajang, 600 – 800 m, 9 June 2007, leg. A. Schulz, 1 ♂ (MHNG, ASWM- 07 / 9); Tioman Island, near Mt Genting, 100 – 250 m, 20 August 2004, leg. P. J. Schwendinger, 1 ♂ (MHNG, AS- 04 / 04); Tioman Island, Mt Kajang, 600 – 1000 m, 22 August 2004, leg. P. J. Schwendinger, 1 ♂ (MHNG, AS- 04 / 06).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA850FFB7E624FCD5024D66B1.taxon	etymology	Etymology The specific epithet, an eponymous patronym (genitive case), is established in honour of Dr Lionel Monod (MHNG) who collected many spider specimens from Tioman Island.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA850FFB7E624FCD5024D66B1.taxon	description	Description Male (holotype). Total length 4.62; prosoma 2.20 long, 1.42 wide; opisthosoma 2.42 long, 1.10 wide. Leg formula: 1423; I 9.86 (2.30, 0.90, 3.32, 2.18, 1.16); II 7.96 (1.80, 0.88, 2.54, 1.72, 1.02); III 6.70 (1.44, 0.90, 1.96, 1.56, 0.84); IV 9.90 (2.40, 0.92, 2.84, 2.46, 1.28). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 8 r 8; metatarsi p 4 r 4. Leg II: Tibiae p 7 r 7; metatarsi p 4 r 3. Carapace dark reddish brown; submarginal lobes triangular, apices sharply pointed; integument not punctate. Sternum dark reddish brown. Legs hirsute, bi-coloured: Coxae to femora reddish brown, other segments yellowish, with dark greenish annulation on distal portion of femora, patellae and tibiae. Opisthosoma elongate-ovoid. Dorsal scutum covering two-thirds of length of opisthosoma. Palp (Figure 9 A – C): RTA triangular, minute, its apex bluntly pointed, entirely sclerotized. DTA digitiform. Tegulum ovoid, protruding posteriorly, slightly excavated on pro-apical side, provided with deep circular pit on meso-retrolateral side. Conductor triangular, its apex sharply pointed. Embolic base clearly visible. Female (paratype, MHNG, AS- 05 / 08). Total length 5.54; prosoma 2.48 long, 1.50 wide; opisthosoma 3.06 long, 1.44 wide. Leg formula: 1423; I 10.88 (2.42, 0.90, 3.72, 2.40, 1.44); II 8.34 (2.00, 0.90, 2.88, 1.52, 1.04); III 710 (1.52, 0.88, 2.02, 1.66, 1.02); IV 10.32 (2.40, 0.92, 3.00, 2.62, 1.38). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 9 r 9; metatarsi p 5 r 5. Leg II: Tibiae p 9 r 8; metatarsi p 5 r 4. General appearance as in male except submarginal lobes on carapace broader at base, their apices bluntly pointed; legs reddish brown, uniform in colour; dorsum of opisthosoma dark greenish. Genitalia (Figure 9 E, F): Epigyne with circular copulatory orifices situated anteriorly. Insemination ducts curved, connected to apical portion of spermathecae. Secretory ampullae spherical, originating subproximally on insemination ducts. Spermathecae reniform, widely separated, provided with glandular pore situated laterally on dorsal surface. Fertilization ducts lanceolated.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA850FFB7E624FCD5024D66B1.taxon	distribution	Distribution Known only from Tioman Island, Malaysia (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA856FFB5E5C9FBCC0285618B.taxon	description	(Figure 10 A – E)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA856FFB5E5C9FBCC0285618B.taxon	diagnosis	Diagnosis Teutamus calceolatus sp. nov. can be easily distinguished from its congeners by the rostrated tegulum of the male palp greatly extending retrolaterally (Figure 10 A, C) and the sharply pointed submarginal lobes on the carapace; females can be easily recognized by the calceolus spermathecae with simple, curved ducts (Figure 10 E).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA856FFB5E5C9FBCC0285618B.taxon	materials_examined	Type material Holotype. ♂, Malaysia: Terengganu State: Lake Kenyir, 5 km southwest of dam, 50 km southwest of Kuala Terengganu, 300 – 400 m, 10 July 2001, leg. A. Schulz and K. Vock (MHNG, M 01 - 167). Paratypes. Data as holotype, 1 ♀ (MHNG, M 01 - 167); from type locality, 10 October 2001, leg. A. Schulz and K. Vock, 1 ♀ (MHNG, M 01 - 168); 7 July 2001, leg. A. Schulz and K. Vock, 1 ♂ (MHNG, M 01 - 159); 7 July 2001, leg. A. Schulz and K. Vock, 1 ♂ (MHNG, M 01 - 160). Kelantan State: Jeram Pasu Waterfall, 8 km west of Kampung Pasang Pak Amat, 30 km south of Kota Baharu, 100 m, evergreen rain forest, 100 m, 10 January 1999, leg. P. Schwendinger, 1 ♀ (MHNG).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA856FFB5E5C9FBCC0285618B.taxon	etymology	Etymology The specific epithet is a nominative form of an adjective (Latin: calceolatus, - a, - um = slipper-shaped), and refers to the shape of the spermathecae.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA856FFB5E5C9FBCC0285618B.taxon	description	Description Male (holotype). Total length 4.78; prosoma 2.28 long, 1.48 wide; opisthosoma 2.50 long, 1.20 wide. Leg formula: 1423; I 10.54 (2.38, 0.92, 3.38, 2.32, 1.54); II 8.32 (1.92, 0.92, 2.62, 1.82, 1.04); III 6.96 (1.54, 0.90, 2.06, 1.52, 0.94); IV 10.26 (2.52, 0.92, 2.94, 2.60, 1.28). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 9 r 9; metatarsi p 5 r 5. Leg II: Tibiae p 8 r 8; metatarsi p 5 r 4. Carapace orange-brown; submarginal lobes triangular, their apices sharply pointed. Sternum orange-brown. Legs hirsute, uniformly brown. Opisthosoma elongate-ovoid. Dorsal scutum covering two-thirds of length of opisthosoma. Palp (Figure 10 A – C): RTA triangular, rather short, strongly excavated anteriorly. DTA short and minute, broad at base, its apex blunt, directed ventrad. Tegulum rostrated, strongly protruding meso-retrolaterally, slightly excavated pro-apically. Conductor triangular, its apex sharply pointed. Embolic base well-developed. Female (paratype, MHNG, M 01 - 167). Total length 5.56; prosoma 2.48 long, 1.50 wide; opisthosoma 3.08 long, 1.52 wide. Leg formula: 1423; I 11.26 (2.48, 0.90, 3.84, 2.50, 1.54); II 8.48 (1.92, 0.92, 2.88, 1.68, 1.08); III 7.32 (1.62, 0.92, 2.08, 1.66, 1.04); IV 10.78 (2.56, 0.92, 3.06, 2.66, 1.58). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 9 r 9; metatarsi p 5 r 5. Leg II: Tibiae p 9 r 8; metatarsi p 5 r 4. General appearance as in male; submarginal lobes triangular, their apices bluntly pointed; dorsum dark greenish; venter pale. Genitalia (Figure 10 D, E): Epigyne with widely separated circular copulatory orifices situated anteriorly. Insemination ducts slender, elongated, connected to middle portion of spermathecae. Spermathecae calceolus, provided with apical glandular pore. Fertilization ducts lanceolated.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA856FFB5E5C9FBCC0285618B.taxon	distribution	Distribution Known only from the type localities on the east coast of Malay Peninsula.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA854FFBBE5C9FB06056061CF.taxon	description	(Figures 1 A, 2 A, 3 E, 11 A – E)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA854FFBBE5C9FB06056061CF.taxon	diagnosis	Diagnosis Somatically, T. orthogonus sp. nov. bears little resemblance to its congeners; it has a distinctly shorter epigastric scutum (Figure 1 A) and a reduced number of spines on anterior legs. Males resemble those of T. rhino in having an elongate tegulum but the conductors are conical rather than spiniform (Figure 11 A); females share close similarities on the epigyne but can be distinguished by the rhomboidal spermathecae and the straight proximal portion of the insemination ducts (Figure 11 E).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA854FFBBE5C9FB06056061CF.taxon	materials_examined	Type material Holotype. ♂, Indonesia: Sumatra, Jambi Province: Mt Kirinci, foot path to the summit, west of Kersik Tua, 2160 m, evergreen hill forest, 17 February 2000, leg. P. J. Schwendinger (MHNG, Sum- 00 / 13). Paratypes. Data as holotype, 3 ♂, 2 ♀ (MHNG, Sum- 00 / 13).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA854FFBBE5C9FB06056061CF.taxon	etymology	Etymology The specific epithet is latinization of a Greek word (oρ ϑ o ζ (orthos) = straight), referring to the straight proximal portion of insemination ducts.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA854FFBBE5C9FB06056061CF.taxon	description	Description Male (holotype). Total length 4.86; prosoma 2.36 long, 1.52 wide; opisthosoma 2.50 long, 1.26 wide. Leg formula: 1423; I 10.76 (2.44, 0.92, 3.42, 2.36, 1.62); II 8.56 (2.02, 0.90, 2.64, 1.88, 1.12); III 7.16 (1.62, 0.92, 2.12, 1.54, 0.96); IV 10.56 (2.58, 0.92, 2.88, 2.62, 1.56). Spination: Leg I: Femora p 1 - 1; tibiae p 7 r 7; metatarsi p 4 r 4. Leg II: Tibiae p 7 r 6; metatarsi p 4 r 3. Carapace reddish brown; submarginal lobes blunt, slightly elevated (Figure 1 A). Sternum reddish brown. Legs hairless, bi-coloured: Femora to tarsi yellowish, other segments reddish brown. Opisthosoma elongateovoid. Dorsal scutum covering almost two-thirds length of opisthosoma, indented posteriorly. Epigastric scutum short (Figure 2 A). Post-epigastric scuta extending half length of opisthosoma. Palp (Figure 11 A – C): RTA triangular, strongly excavated medially, its apex blunt, anterior margin with colourless and transparent lamina. DTA digitiform, weakly sclerotized. Tegulum elongate-ovoid, slightly protruding posteriorly. Conductor triangular, its apex sharply pointed, gradually narrowing anteriorly. Embolic base clearly visible. Female (paratype, MHNG, MAL- 04 / 01): Total length 5.78; prosoma 2.62 long, 1.58 wide; opisthosoma 3.16 long, 1.60 wide. Leg formula: 1423; I 11.30 (2.48, 0.92, 3.88, 2.56, 1.46); II 8.76 (2.00, 0.92, 2.94, 1.72, 1.18); III 7.66 (1.66, 0.90, 2.14, 1.76, 1.20); IV 10.96 (2.54, 0.94, 3.18, 2.68, 1.62). Spination: Leg I: Femora p 1 - 1; tibiae p 7 r 7; metatarsi p 4 r 4. Leg II: Tibiae p 7 r 7; metatarsi p 4 r 3. General appearance as in male; legs reddish brown, uniform in colour; dorsum of opisthosoma dark greenish, posteriorly with a series of pale chevrons. Genitalia (Figure 11 D, E): Epigyne with circular copulatory orifices situated anterior medially. Insemination ducts descending obliquely then ascending anteriorly, connected to apical portion of spermathecae. Spermathecae trapezoidal, strongly narrowed and elongated anteriorly. Fertilization ducts lanceolated.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA854FFBBE5C9FB06056061CF.taxon	distribution	Distribution Known only from the type locality on the west coast of Sumatra (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85AFFB9E5D3FB42028B601F.taxon	description	(Figures 3 C, 12 A – F)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85AFFB9E5D3FB42028B601F.taxon	diagnosis	Diagnosis Males of T. globularis sp. nov. closely resemble those of T. rothorum in having a relatively large conductor but can be easily distinguished by the globular shape of the tegulum (Figure 12 A – C); females also possess the large, oval epigynal atria but can be by the different shapes of insemination ducts and spermathecae (Figures 3 C, 12 D – F).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85AFFB9E5D3FB42028B601F.taxon	materials_examined	Type material Holotype. ♂, Malaysia: Pahang State: Charas Hill, 3 km north of Pancing 60 m, evergreen rainforest on limestone, 9 July 2001, leg. P. J. Schwendinger (MHNG, SIM- 01 / 12). Paratypes. Data as holotype, 2 ♂, 3 ♀ (MHNG, SIM- 01 / 12); from type locality, 60 m, rain forest remnants at foot of limestone hill, 3 June 2004, leg. P. J. Schwendinger, 3 ♂ (MHNG, MAL- 04 / 08).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85AFFB9E5D3FB42028B601F.taxon	etymology	Etymology The specific epithet is derived from Latin (globularis, globulare = globular), and refers to the shape of the tegulum of male palp.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85AFFB9E5D3FB42028B601F.taxon	description	Description Male (holotype). Total length 4.98; prosoma 2.36 long, 1.58 wide; opisthosoma 2.62 long, 1.30 wide. Leg formula: 1423; I 10.64 (2.48, 0.92, 3.26, 2.32, 1.66); II 8.54 (2.06, 0.90, 2.58, 1.92, 1.08); III 7.06 (1.64, 0.92, 2.08, 1.42, 1.00); IV 10.38 (2.56, 0.92, 3.06, 2.56, 1.28). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 9 r 9; metatarsi p 5 r 5. Leg II: Tibiae p 9 r 8; metatarsi p 4 r 4. Carapace reddish brown; submarginal lobes triangular, their apices globular. Sternum orange-brown. Legs hirsute, uniform in colour, orangebrown. Opisthosoma elongate-ovoid. Dorsal scutum covering almost entire length of opisthosoma. Palp (Figure 12 A – C): RTA digitiform, strongly excavated anteriorly, its apex blunt, anterior margin with colourless and transparent lamina. DTA short and stout, weakly sclerotized. Tegulum more or less spherical, slightly protruding apico-retrolaterally. Conductor conical, elongated, gradually narrowing apically, its apex sharply pointed. Embolic base almost indistinct. Female (paratype, MHNG, SIM- 01 / 12). Total length 5.58; prosoma 2.46 long, 1.50 wide; opisthosoma 3.12 long, 1.48 wide. Leg formula: 1423; I 11.62 (2.60, 1.00, 3.82, 2.54, 1.66); II 9.26 (2.08, 1.00, 3.02, 1.84, 1.32); III 7.92 (1.68, 0.90, 2.26, 1.88, 1.20); IV 11.32 (2.60, 1.00, 3.24, 2.86, 1.62). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 8 r 8; metatarsi p 5 r 5. Leg II: Tibiae p 8 r 8; metatarsi p 4 r 4. General appearance as in male; submarginal lobes on carapace blunt, slightly elevated; dorsum pale; venter pale. Genitalia (Figure 12 D – F). Epigyne with large, elongate-oval epigynal atria filled with dark hardening secretion. Copulatory orifices situated laterally on epigynal atria. Insemination ducts descending obliquely then moving transversely, connected to anterior portion of spermathecae. Spermathecae reniform, anterior portion distinctly narrowed. Fertilization ducts lanceolated.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85AFFB9E5D3FB42028B601F.taxon	distribution	Distribution Known only from the type locality on the west coast of Malay Peninsula.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA858FFBFE5D4FA72005B6338.taxon	description	(Figures 1 D, 13 A – E)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA858FFBFE5D4FA72005B6338.taxon	diagnosis	Diagnosis Males of T. seculatus sp. nov. resemble those of T. brachiatus sp. nov. in having an elongate conductor but can be distinguished by the presence of a minute accessory apophysis (Figure 13 A, B); females can be recognized by the elliptic spermathecae provided with short and tortuous anterior insemination ducts (Figure 13 E).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA858FFBFE5D4FA72005B6338.taxon	materials_examined	Type material Holotype. ♂, Indonesia: Ligga Island: waterfall c. 7 km north of Dalik, 60 m, rainforest along stream, 13 – 15 July 2001, leg. P. J. Schwendinger (MHNG, SIM- 01 / 03). Paratypes. Malaysia: Johor State: Mt Ophir, Puteri Waterfall, 80 – 130 m, rainforest near stream, 21 May 2004, leg. P. J. Schwendinger, 5 ♂, 3 ♀ (MHNG, MAL- 04 / 03).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA858FFBFE5D4FA72005B6338.taxon	etymology	Etymology The specific epithet is a nominative form of an adjective (Latin: secula = sickle), and refers to the sickle-shaped conductor on the male palp.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA858FFBFE5D4FA72005B6338.taxon	description	Description Male (holotype). Total length 4.82; prosoma 2.30 long, 1.52 wide; opisthosoma 2.52 long, 1.22 wide. Leg formula: 1423; I 10.58 (2.40, 0.90, 3.40, 2.32, 1.56); II 8.26 (1.92, 0.90, 2.60, 1.80, 1.04); III 6.88 (1.54, 0.82, 2.02, 1.50, 1.00); IV 10.24 (2.54, 1.00, 2.88, 2.46, 1.36). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 9 r 9; metatarsi p 5 r 5. Leg II: Tibiae p 9 r 8; metatarsi p 5 r 4. Carapace dark reddish brown; submarginal lobes triangular, their apices bluntly pointed. Sternum dark reddish brown. Legs hairless, bi-coloured: Coxae, metatarsi and tarsi yellowish, other segments orangebrown. Opisthosoma elongate-ovoid. Dorsal scutum covering two-thirds of length of opisthosoma. Palp (Figure 13 A – C). RTA digitiform, strongly excavated anteriorly, its apex blunt, anterior margin with colourless and transparent. DTA short and stout. Tegulum elongate-ovoid, with field of lightly sclerotized area on baso-retrolateral side. Conductor elongated, gradually narrowing towards sharply pointed apex. Embolic base almost indistinct. Accessory tegular apophysis minute, triangular, situated close to conductor. Female (paratype, MHNG, MAL- 04 / 03). Total length 5.58; prosoma 2.48 long, 1.50 wide; opisthosoma 3.10 long, 1.52 wide. Leg formula: 1423; I 11.52 (2.46, 0.90, 3.84, 2.50, 1.52); II 8.52 (1.96, 0.90, 2.96, 1.60, 1.10); III 7.26 (1.60, 0.80, 2.12, 1.70, 1.04); IV 10.72 (2.50, 0.90, 3.12, 2.70, 1.50). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 10 r 10; metatarsi p 6 r 6. Leg II: Tibiae p 9 r 9; metatarsi p 5 r 5. General appearance as in male except submarginal lobes on carapace distinctly broader, their apices blunt; dorsum of opisthosoma dark greenish. Genitalia (Figure 13 D, E): Epigyne with two large, circular copulatory orifices situated anteriorly. Insemination ducts descending mid-longitudinally then abruptly ascending anteriorly, connected to apical portion of spermathecae. Secretory ampullae digitiform, originating terminally on insemination ducts. Spermathecae elliptic, gradually narrowing anteriorly, provided with sub-apical glandular pore on inner surface and median pore on dorsal surface. Fertilization ducts lanceolated.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA858FFBFE5D4FA72005B6338.taxon	distribution	Distribution The southern tip of Malay Peninsula and Ligga Island of Indonesia (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA858FFBFE5D4FA72005B6338.taxon	discussion	The following species described below are based on single sex. Nevertheless, it has been possible to circumscribe all taxa with reasonable clarity on their species boundaries.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85FFFBDE595FF7B0561613A.taxon	description	(Figures 2 H, 14 A)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85FFFBDE595FF7B0561613A.taxon	diagnosis	Diagnosis The male holotype of T. hirtellus sp. nov. closely resembles that of T. rothorum in having a large conductor but can be easily distinguished by the more elongate conductor pointing anteriad instead of prolaterad (Figure 14 A). To our knowledge, only the male of this new species and those of T. andrewdavisi (Figure 14 B) and T. vittatus possess a group of bristles on the RTA. In other species the postepigastric scuta slightly extend posteriorly (Figure 2 I), whereas in T. hirtellus sp. nov. and T. orthogonus sp. nov. they extend further and fuse with the dorsal scutum to form a complete tubular ring enclosing anterior half of the opisthosoma entirely (Figure 2 H).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85FFFBDE595FF7B0561613A.taxon	materials_examined	Type material Holotype. ♂, Philippines: Palawan, Puerto Princesa Region: Sabang Underground NP, Daylight Hole and Lions Cave, 100 – 200 m, primary forest, 6 – 8 December 2009, leg. A. Schulz (MHNG, PAL- 09 / 07).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85FFFBDE595FF7B0561613A.taxon	etymology	Etymology The specific epithet is a nominative form of an adjective (Latin: hirtellus, - a, - um = slightly hairy), and refers to the tuft of bristles on RTA of the male palp.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85FFFBDE595FF7B0561613A.taxon	description	Description Male (holotype). Total length 4.74; prosoma 2.26 long, 1.44 wide; opisthosoma 2.48 long, 1.26 wide. Leg formula: 1423; I 10.48 (2.36, 0.92, 3.38, 2.32, 1.50); II 8.08 (1.86, 0.90, 2.54, 1.76, 1.02); III 6.84 (1.54, 0.88, 2.00, 1.44, 0.98); IV 10.16 (2.46, 0.92, 2.88, 2.56, 1.34). Spination: Leg I: Femora p 1 - 1; tibiae p 10 r 10; metatarsi p 6 r 6. Leg II: Tibiae p 10 r 8; metatarsi p 6 r 5. Carapace dark reddish brown; submarginal lobes blunt, slightly elevated. Sternum dark reddish brown. Legs hairless, bi-coloured: Coxae, trochanters and femora dark brown, other segments slightly paler, their distal portion white. Opisthosoma elongate-ovoid. Dorsal scutum covering two-thirds of length of opisthosoma. Venter covered by posteriorly extended post-epigastric scuta (Figure 2 H). Palp (Figure 14 A): RTA digitiform in ventral view, elongated-slender, its apex bluntly pointed, lateral margin covered with bristles. DTA short, almost indistinct. Tegulum elongate-ovoid, slightly excavated on apico-prolateral side. Conductor elongated, broad at base, its apex sharply pointed, almost reaching apex of cymbium. Embolic base clearly visible. Female. Unknown.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85FFFBDE595FF7B0561613A.taxon	distribution	Distribution Known only from the type locality on the island of Palawan, the Philippines.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85CFF83E5D7FB5D056066AA.taxon	description	(Figure 15 A, B)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85CFF83E5D7FB5D056066AA.taxon	diagnosis	Diagnosis Females of T. tortuosus sp. nov. can be easily recognized by the strongly convoluted terminal portion of the insemination ducts (Figure 15 B).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85CFF83E5D7FB5D056066AA.taxon	materials_examined	Type material Holotype. ♀, Indonesia: Sumatra, West Sumatra Province: Anai Valley, 6 km south of Padang Panjang, 500 m, primary forest, 1 June 2006, leg. P. J. Schwendinger (MHNG, Sum- 06 / 03). Paratypes. Data as holotype, 2 ♀, (MHNG, Sum- 06 / 03).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85CFF83E5D7FB5D056066AA.taxon	etymology	Etymology The specific epithet is a nominative form of an adjective (Latin: tortuosus, - a, - um = twist), and refers to the strongly tortuous duct of the internal female genitalia.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85CFF83E5D7FB5D056066AA.taxon	description	Description Female (holotype). Total length 5.46; prosoma 2.66 long, 1.82 wide; opisthosoma 2.80 long, 1.40 wide. Leg formula: 1423; I 11.98 (2.70, 1.12, 3.72, 2.58, 1.86); II 9.38 (2.18, 1.00, 2.88, 1.96, 1.36); III 8.18 (1.86, 0.98, 2.32, 1.78, 1.24); IV 11.14 (2.78, 1.12, 3.12, 2.46, 1.66). Spination: Leg I: Femora p 1 - 1; tibiae p 9 r 9; metatarsi p 5 r 5. Leg II: Tibiae p 8 r 8; metatarsi p 4 r 4. Carapace dark chestnut-brown; submarginal lobes blunt. Sternum brown. Legs hairless, bi-coloured: Tibiae, metatarsi and tarsi yellowish, other segments orange-brown. Opisthosoma elongate-ovoid. Dorsum dark greenish, posteriorly with series of pale chevrons. Venter dark greenish. Genitalia (Figure 15 A, B): Epigyne with elliptic copulatory orifices situated anteriorly. Insemination ducts elongated, descending posteriorly then abruptly bent, forming tortuous terminal complex, connected to anterior lateral portion of spermathecae. Secretory ampullae spherical, originating proximally on insemination ducts. Spermathecae ovoid, rounded anteriorly, gradually narrowing posteriorly, provided with apical glandular pore. Fertilization ducts lanceolated. Male. Unknown.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA85CFF83E5D7FB5D056066AA.taxon	distribution	Distribution Known only from the type locality on the west coast of Sumatra (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA862FF82E5D5FBEA02AC663C.taxon	description	(Figure 15 C, D)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA862FF82E5D5FBEA02AC663C.taxon	diagnosis	Diagnosis Females of T. rollardae sp. nov. can be recognized by the peculiar shape of spermathecae and the small insemination ducts (Figure 15 D).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA862FF82E5D5FBEA02AC663C.taxon	materials_examined	Type material Holotype. ♀, Indonesia: Sumatra, West Sumatra Province: Mt Merapi, c. 15 km southeast of Bukittiggi, 1650 – 1700 m, hill forest, 4 June 2006, leg. P. J. Schwendinger (MHNG, Sum- 06 / 07). Paratypes. Data as holotype, 1 ♀, (MHNG, Sum- 06 / 07).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA862FF82E5D5FBEA02AC663C.taxon	etymology	Etymology The specific epithet is an eponymous patronym (genitive form of a noun), dedicated to Dr Christine Rollard (MHNH) in gratitude for her generous hospitality.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA862FF82E5D5FBEA02AC663C.taxon	description	Description Female (holotype). Total length 5.12; prosoma 2.46 long, 1.68 wide; opisthosoma 2.66 long, 1.34 wide. Leg formula: 1423; I 11.34 (2.56, 1.08, 3.54, 2.46, 1.70); II 8.94 (2.06, 1.00, 2.74, 1.94, 1.20); III 7.56 (1.72, 0.96, 2.14, 1.66, 1.08); IV 10.82 (2.68, 1.04, 3.00, 2.64, 1.46). Spination: Leg I: Femora p 1 - 1; tibiae p 10 r 9; metatarsi p 6 r 6. Leg II: Tibiae p 8 r 8; metatarsi p 5 r 4. Carapace orange-brown; submarginal lobes triangular, their apices blunt. Sternum brown. Legs hairless, bi-coloured: Tibiae, metatarsi and tarsi yellowish brown, other segments brown. Opisthosoma elongate-ovoid. Dorsum pale, posteriorly with a series of pale chevrons. Genitalia (Figure 15 C, D): Epigyne with circular copulatory orifices situated anteriorly. Insemination ducts elongated, slender, descending posteriorly then abruptly bent, moving transversely and curving posteriorly, connected to lateral portion of spermathecae without distinctive border. Secretory ampullae spherical, minute, originating proximally on insemination ducts. Spermathecae reniform, anterior portion strongly elongated, enlarged posteriorly, provided with glandular pore on inner surface. Fertilization ducts lanceolated. Male. Unknown.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA862FF82E5D5FBEA02AC663C.taxon	distribution	Distribution Known only from the type locality on the west coast of Sumatra (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA863FF81E5E0FC5105606635.taxon	description	(Figure 15 E, F)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA863FF81E5E0FC5105606635.taxon	diagnosis	Diagnosis Females of T. poggii sp. nov. share with those of T. rollardae sp. nov. a peculiar shape of the spermathecae but can be distinguished by the simple insemination ducts connecting to the very large, circular copulatory orifices (Figure 15 E, F).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA863FF81E5E0FC5105606635.taxon	materials_examined	Type material Holotype. ♀, Indonesia: Sumatra, West Sumatra Province: old secondary forest above Taman Hutan Raya Bung Hatta, near road Padang – Lubuk Sulasih, 1100 m, 29 – 30 May 2006, leg. P. J. Schwendinger (MHNG, Sum- 06 / 01). Paratypes. Data as holotype, 2 ♀, (MHNG, Sum- 06 / 01).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA863FF81E5E0FC5105606635.taxon	etymology	Etymology The specific epithet is a nominative possessive form of a noun in honour of Dr Roberto Poggi, director of MSNG who kindly made this study possible.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA863FF81E5E0FC5105606635.taxon	description	Description Female (holotype). Total length 5.22; prosoma 2.50 long, 1.72 wide; opisthosoma 2.72 long, 1.36 wide. Leg formula: 1423; I 11.54 (2.60, 1.08, 3.58, 2.52, 1.76); II 9.16 (2.10, 1.00, 2.82, 1.98, 1.26); III 7.76 (1.76, 1.00, 2.20, 1.72, 1.08); IV 11.04 (2.68, 1.08, 3.06, 2.70, 1.52). Spination: Leg I: Femora p 1 - 1; tibiae p 10 r 8; metatarsi p 5 r 6. Leg II: Tibiae p 9 r 8; metatarsi p 5 r 4. Carapace orange-brown; submarginal lobes blunt; integument not punctate. Sternum yellowish brown. Legs hairless, bicoloured: Tibiae, metatarsi and tarsi yellow, other segments brown. Opisthosoma elongate-ovoid. Dorsum pale, posteriorly with a series of faint chevrons. Genitalia (Figure 15 E, F): Epigyne with large circular copulatory orifices situated anteriorly. Insemination ducts broad proximally, gradually narrowing terminally, descending obliquely then abruptly twisted, connected to anterior lateral portion of spermathecae without distinctive border. Secretory ampullae spherical, originating subproximally on insemination ducts. Spermathecae elongated anteriorly, enlarged posteriorly, provided with glandular pore on inner surface. Fertilization ducts lanceolated. Male. Unknown.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA863FF81E5E0FC5105606635.taxon	distribution	Distribution Known only from the type locality on the west coast of Sumatra (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA860FF87E5C8FC48022E6740.taxon	description	(Figures 3 H, 16 A, B)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA860FF87E5C8FC48022E6740.taxon	diagnosis	Diagnosis Females of T. leptothecus sp. nov. can be recognized by the anteriorly narrowed, elliptic spermathecae and the simple curved ducts (Figure 16 A, B).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA860FF87E5C8FC48022E6740.taxon	materials_examined	Type material Holotype. ♀, Malaysia: Johor State: Kota Tinggi Waterfalls, at the foot of Mt Muntahak, 170 m, rainforest along stream, 24 – 26 June 2001, leg. P. J. Schwendinger (MHNG, SIM- 01 / 07). Paratype. Data as holotype, 1 ♀, (MHNG, SIM- 01 / 07).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA860FF87E5C8FC48022E6740.taxon	etymology	Etymology The specific epithet is a nominative form of an adjective (leptos = thin; thecus = sac), and refers to the shape of spermathecae.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA860FF87E5C8FC48022E6740.taxon	description	Description Female (holotype). Total length 5.86; prosoma 2.62 long, 1.62 wide; opisthosoma 3.24 long, 1.62 wide. Leg formula: 1423; I 11.78 (2.58, 1.00, 3.96, 2.62, 1.62); II 8.94 (2.06, 0.98, 3.04, 1.68, 1.18); III 7.84 (1.72, 0.92, 2.22, 1.82, 1.16); IV 11.30 (2.64, 1.00, 3.22, 2.80, 1.64). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 9 r 9; metatarsi p 6 r 6. Leg II: Tibiae p 8 r 8; metatarsi p 5 r 5. Carapace dark chestnut-brown; submarginal lobes blunt. Sternum brown. Legs hairless, bi-coloured: Tibiae, metatarsi and tarsi yellowish brown, other segments brown. Opisthosoma elongate-ovoid. Dorsum pale, posteriorly with a series of faint chevrons. Genitalia (Figure 16 A, B): Epigyne with circular copulatory orifices situated anteriorly. Insemination ducts curving posteriorly, connected to apical portion of spermathecae. Spermathecae elliptic, distinctly narrowed anteriorly, surface provided with numerous pore ductules. Fertilization ducts lanceolated. Male. Unknown.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA860FF87E5C8FC48022E6740.taxon	distribution	Distribution Known only from the type locality in Malaysia (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA866FF86E5CAFDBD026A6455.taxon	description	(Figure 16 C, D)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA866FF86E5CAFDBD026A6455.taxon	diagnosis	Diagnosis The female of T. deelemanae sp. nov. can be recognized by the elliptic spermathecae and the partially hidden glandular pore situated underneath the insemination ducts (Figure 16 D).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA866FF86E5CAFDBD026A6455.taxon	materials_examined	Type material Holotype. ♀, Malaysia: Johor State: Mt Ophir, Puteri Waterfall, 80 – 130 m, rainforest near stream, 21 May 2004, leg. P. J. Schwendinger (MHNG, MAL- 04 / 03).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA866FF86E5CAFDBD026A6455.taxon	etymology	Etymology The specific epithet is an eponymous patronym (genitive form of a noun), dedicated to Dr Christa Deeleman-Reinhold, a prolific expert on Southeast Asian spider fauna.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA866FF86E5CAFDBD026A6455.taxon	description	Description Female (holotype). Total length 5.12; prosoma 2.46 long, 1.68 wide; opisthosoma 2.66 long, 1.32 wide. Leg formula: 1423; I 11.26 (2.54, 1.02, 3.54, 2.46, 1.70); II 8.98 (2.06, 1.00, 2.76, 1.94, 1.22); III 7.62 (1.72, 1.00, 2.16, 1.68, 1.06); IV 10.70 (2.58, 1.02, 3.00, 2.64, 1.46). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 8 r 8; metatarsi p 6 r 6. Leg II: Tibiae p 7 r 7; metatarsi p 4 r 5. Carapace reddish brown; submarginal lobes triangular, their apices bluntly pointed. Sternum brown. Legs hairless, uniform in colour, yellowish brown. Opisthosoma elongate-ovoid. Dorsum pale. Genitalia (Figure 16 C, D): Epigyne with circular copulatory orifices situated anteriorly. Insemination ducts elongated, curving posteriorly then forming two tortuous loops, connected to apical portion of spermathecae. Spermathecae more or less elliptic, provided with apical glandular pore on dorsal surface, partially hidden by insemination ducts. Fertilization ducts lanceolated. Male. Unknown.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA866FF86E5CAFDBD026A6455.taxon	distribution	Distribution Known only from the type locality in Malaysia (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA867FF85E59FFEA802DA618A.taxon	description	(Figure 17 A, B)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA867FF85E59FFEA802DA618A.taxon	diagnosis	Diagnosis Females of T. spiralis sp. nov. can be recognized by the spiral insemination ducts and the enlarged, spatulate secretory ampullae (Figure 17 B).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA867FF85E59FFEA802DA618A.taxon	materials_examined	Type material Holotype. ♀, Indonesia: Borneo, East Kalimantan Province: Berau Distinct, Hutan Wisata Sei Tangap, c. 8 km west of Tanjung Redeb, 30 m, primary forest, 2 October 2008, leg. P. J. Schwendinger (MHNG, IND- 08 / 13). Paratype. Berau District: 1 km off road Tanjung Redeb – Tanjung Selor, c. 45 km north of Tanjung Redeb, 190 m, primary forest, 29 September to 3 October 2008, P. J. Schwendinger, 1 ♀ (MHNG, IND- 08 / 07); near Kampung Suaran, c. 40 km south of Tanjung Redeb, 50 m, primary forest on limestone, 1 October 2008, leg. P. J. Schwendinger, 1 ♀ (MHNG, IND- 08 / 11).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA867FF85E59FFEA802DA618A.taxon	etymology	Etymology The specific epithet is derived from Latin (spiralis, - ale = spiral), and refers to the strongly convoluted insemination ducts of the female internal genitalia.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA867FF85E59FFEA802DA618A.taxon	description	Description Female (holotype). Total length 5.20; prosoma 2.50 long, 1.72 wide; opisthosoma 2.70 long, 1.36 wide. Leg formula: 1423; I 11.42 (2.58, 1.00, 3.58, 2.50, 1.76); II 9.16 (2.10, 1.00, 2.82, 1.98, 1.26); III 7.76 (1.76, 0.98, 2.22, 1.70, 1.10); IV 11.00 (2.70, 1.00, 3.08, 2.72, 1.50). Spination: Leg I: Femora p 1 - 1 - 1; tibiae p 10 r 10; metatarsi p 6 r 6. Leg II: Tibiae p 8 r 8; metatarsi p 4 r 4. Carapace yellowish brown; submarginal lobes blunt; integument not punctate. Sternum orange-brown. Legs hairless, uniformly brown. Opisthosoma elongate-ovoid. Dorsum dark greenish. Genitalia (Figure 17 A, B): Epigyne with elliptic copulatory orifices situated anteriorly. Internally represented by elongated proximal convoluted ducts and enlarged terminal spermathecae. Secretory ampullae spatulated, with long stalks and numerous pore ductules. Fertilization ducts lanceolated. Male. Unknown.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA867FF85E59FFEA802DA618A.taxon	distribution	Distribution On the east coast of Borneo (East Kalimantan Province) (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA864FF8BE5CBFB0705606707.taxon	description	(Figure 17 C, D)	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA864FF8BE5CBFB0705606707.taxon	diagnosis	Diagnosis Females of T. sumatranus sp. nov. can be recognized by the elongate insemination ducts connecting to the small, trapezoidal spermathecae with no distinctive boundary (Figure 17 D).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA864FF8BE5CBFB0705606707.taxon	materials_examined	Type material Holotype. ♀, Indonesia: Sumatra, West Sumatra Province: old secondary forest above Harau Canyon, north of Payakumbuh, 750 m, primary forest, 7 June 2006, leg. P. J. Schwendinger (MHNG, Sum- 06 / 11). Paratypes. Data as holotype, 1 ♀, (MHNG, Sum- 06 / 11).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA864FF8BE5CBFB0705606707.taxon	etymology	Etymology The specific is a nominative form of an adjective (Latin: sumatranus = of Sumatra), and refers to the collecting locality.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA864FF8BE5CBFB0705606707.taxon	description	Description Female (holotype). Total length 5.48; prosoma 2.70 long, 1.76 wide; opisthosoma 2.78 long, 1.36 wide. Leg formula: 1423; I 12.12 (2.74, 1.10, 3.76, 2.62, 1.90); II 9.44 (2.22, 1.00, 2.86, 2.06, 1.30); III 7.82 (1.90, 0.98, 2.34, 1.60, 1.00); IV 11.68 (2.86, 1.10, 3.26, 2.84, 1.62). Spination: Leg I: Femora p 1 - 1; tibiae p 9 r 10; metatarsi p 5 r 6. Leg II: Tibiae p 8 r 9; metatarsi p 4 r 5. Carapace dark chestnut-brown; submarginal lobes triangular, slightly elevated. Sternum dark brown. Legs hairless, uniformly brown, distal portion of tibiae and metatarsi yellowish. Opisthosoma elongate-ovoid. Dorsum greenish, posteriorly with a series of pale chevrons. Genitalia (Figure 17 C, D): Epigyne with elliptic copulatory orifices situated anteriorly. Insemination ducts elongated, tortuous, connected to apical portion of spermathecae without distinctive boundary. Secretory ampullae situated subproximally. Spermathecae trapezoidal, provided with glandular pore on its inner surface. Fertilization ducts lanceolated. Male. Unknown.	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
03A6879FA864FF8BE5CBFB0705606707.taxon	distribution	Distribution Known only from the type locality on the west coast of Sumatra (Figure 18).	en	Dankittipakul, Pakawin, Tavano, Maria, Singtripop, Tippawan (2012): Seventeen new species of the spider genus Teutamus Thorell, 1890 from Southeast Asia (Araneae: Liocranidae). Journal of Natural History 46: 1689-1730, DOI: 10.110.1080/00222933.2012.681314, URL: https://www.researchgate.net/publication/254272704_Seventeen_new_species_of_the_spider_genus_Teutamus_Thorell_1890_from_Southeast_Asia_Araneae_Liocranidae
