identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A687D8FFCED7090AC8F44FFCF6FBF8.text	03A687D8FFCED7090AC8F44FFCF6FBF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marphysa Quatrefages 1865	<div><p>Marphysa Quatrefages, 1865</p><p>Common name: Bloodworm</p><p>Marphysa Quatrefages, 1865: 331 .</p><p>Nauphanta Kinberg, 1865: 564 . – Fauchald, 1987: 375.</p><p>Diagnosis (after Fauchald et al. 2003). Anterior body segments cylindrical, usually becoming dorsoventrally flattened posteriorly; epidermis iridescent anterodorsally. Prostomium with deep or faint anteroventral notch which may give it a bilobed appearance; bearing five similar-looking appendages—three antennae and two (lateral) palps arranged in more or less curved line near posterior edge. Eyespots present or absent. Peristomium divided into two rings, anterior one longer than posterior one; peristomial cirri absent. Branchiae present over most of body or restricted to anterior half, pectinate or palmate arrangement. Chaetae include in superior position, limbate capillaries and pectinate chaetae which may be symmetrical or asymmetrical, and in inferior position, compound falcigers (may be absent), compound spinigers (rarely absent) and subacicular hooks. Jaw apparatus includes ventral mandibles and dorsally, maxillae comprising short, winged, or slender, carriers and four paired (and one unpaired) maxillary plates.</p><p>Remarks. Fauchald (1987) resurrected the genus Nauphanta to accommodate two species lacking compound falcigers and compound spinigers, including N. novaehollandiae Kinberg, 1865, originally from Sydney Harbour, and Eunice mossambica Peters, 1854 originally from Mozambique. He considered Nauphanta to differ from Marphysa in having a unique type of chaeta present only in posterior chaetigers, ‘fan chaetae’, or ‘asymmetrical pectinate chaetae’ as they are referred to here. However, our observations suggest that fan and pectinate chaetae are both in the same position in the neuropodium—on the anteromedial edge of the supra-acicular bundle of chaetae—so it is highly likely that the two types of chaetae are homologous. Further, present observations on Marphysa fauchaldi n. sp. (Table 1) and those of Crossland (1903: 140) and Treadwell (1922: 152) indicate that asymmetrical pectinate chaetae are not restricted to posterior chaetigers but may occur also in more anterior chaetigers. In Marphysa fauchaldi n. sp., as in other species of the genus with asymmetrical pectinate chaetae (see Table 2), asymmetry becomes more pronounced in posterior pectinate chaetae. The presence of pectinate chaetae is best determined by use of SEM, but they are also readily visible under high powered light microscopy provided the parapodium is mounted anterior side up.</p><p>The only other differences between Nauphanta and most Marphysa are the absence of both compound falcigers and spinigers in the former. The absence of compound falcigers is probably an ontogenetic loss as species of Marphysa whose embryology and juvenile development has been investigated have this type of chaetae initially as juveniles, but they are replaced in adults (Borradaile 1901; Southern 1921; Aiyar 1931, Pillai 1958; pers. obs. CG). The absence of compound spinigers is likely to be paedomorphic because juveniles of several Marphysa species, for example M. borradailei (see Pillai 1958), lack compound spinigers but they develop later in adults. Therefore continued absence of this type of chaetae in adults can be viewed as a retained juvenile characteristic. The new species described here, M. fauchaldi n. sp., exhibits what might be called ‘partial paedomorphosis’ in which compound spinigers are absent in all parapodia except the anterior ones (see below). Therefore, the absences of both types of compound chaetae in Nauphanta are most likely attributable to development novelties restricted to this particular taxon (here considered to represent a single species), rather than synapomorphic characteristics of a group of species—thus we propose returning Nauphanta to junior synonymy with Marphysa .</p><p>Species included. Two of the species of Marphysa described below ( M. mullawa and M. fauchaldi n. sp.) are closest in morphology to Group B2 of Fauchald (1970), that is, having compound spinigers only and branchiae present to the end of the body. The only other species listed as Group B2 Marphysa reported from Australia is M. macintoshi, originally described from Zanzibar. However, Group B2 is well represented in the tropical/subtropical Indo-Pacific waters by many other species including M. gravelyi Southern, 1921 (type locality, Chilka Lake, India), M. orientalis Treadwell, 1936 (China), M. tamurai Okuda, 1934 (Japan), M. teretiuscula (Schmarda, 1861) (Sri Lanka) and M. borradailei Pillai 1958 (Sri Lanka). Marphysa borradailei was classified as Group C2 (only compound falcigers present) by Fauchald, presumably because falcigerous chaetae are mentioned in Pillai’s description (p. 104), but the chaetae illustrated by Pillai are clearly compound spinigers, albeit with short blades. Marphysa sanguinea (Montagu, 1813) is also a Group B2 species, but Australian records of this species are all thought to be misidentifications, having been described as a new species, M. mullawa Hutchings &amp; Karageorgopoulos, 2003 .</p><p>Day (1962) regarded Marphysa simplex Crossland, 1903 and M. furcellata Crossland, 1903 to be junior synonyms of M. macintoshi Crossland, 1903 —all three species having Zanzibar as the type locality. The main difference between the three species—whether or not an anterior notch exists between the two globular buccal lips—was attributed by Day to intraspecific variation. Examination of the types of all three species together with the hundreds of specimens of M. fauchaldi n. sp., suggest that the form of the prostomium does not vary significantly between individuals, and this taken together with the other small, but significant, differences between the three species suggest that M. simplex and M. furcellata are each diagnosable, as indicated in the Key. However Marphysa simplex is considered here to be a junior synonym of M. teretiuscula (the possibility was considered likely by Crossland (1903: 136)) as both species present the unusual condition of having a more or less circular (in cross section) body along the entire length of the worm, and antennae are all approximately the same length, being twice the length of the prostomium (see Key). Notwithstanding the relegation of M. simplex to junior synonymy, the name still remains a secondary homonym of Marphysa simplex Langerhans, 1884 (as Amphiro simplex) from Madeira.</p></div>	https://treatment.plazi.org/id/03A687D8FFCED7090AC8F44FFCF6FBF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Glasby, Christopher J.;Hutchings, Pat A.	Glasby, Christopher J., Hutchings, Pat A. (2010): A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352: 29-45, DOI: 10.5281/zenodo.193484
03A687D8FFCCD70C0AC8F68FFBC1F81E.text	03A687D8FFCCD70C0AC8F68FFBC1F81E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marphysa fauchaldi	<div><p>Marphysa fauchaldi n. sp.</p><p>Common name: Barra bloodworm (Figs 1–3)</p><p>Material examined. HOLOTYPE: Australia, Northern Territory, Ardatek Barrumundi farm, Cox Peninsula Road, off Elizabeth River, Darwin region 12.70313˚ S 130.96233 ˚E (NTM W23040), coll. C. Glasby, 13 Dec. 2008. PARATYPES: same locality as for holotype, 1(BMNH ANEA 2010.1), 3(NTM W23041), 2(USNM 1133660), 4(NTM W23042), 1 specimen on several SEM stubs (AM W35419). NON-TYPES: Australia, Northern Territory, Ardatek Barramundi farm, Cox Peninsula Road, off Elizabeth River, Darwin region 12.70313˚ S 130.96233 ˚E several specimens (NTM W17319), coll. C. Glasby, 27 Feb. 2002, Darwin, Stn LC1, 12˚24.8’S 130˚50.0’E Ludmilla Ck mouth, 1(NTM W68), mangrove mud, coll. R.J. Hanley, 26 Nov. 1981; Charles Darwin National Park, 1(NTM W18710), Stn E1-2-W-C2, 12˚27.389’S 130˚52.135’E, mud in Rhizophora zone, coll. K. Metcalfe, 15 Jan. 2003; East Arm Port, 1(NTM W18854), Stn DP-21-Q1, 1(NTM W18857), 12˚29.3’S 130˚ 54.18’E, coll. K. Metcalfe, 12 Oct. 2001; Mindil Beach, creek near casino, 1(NTM W20189), 12˚27’S 130˚50’E, coll. C. Glasby, 10 Sept. 2003; Cobourg Peninsula, Port Essington, Record Point, Stn NTERP, 1(NTM W20375), 11˚ 20.01’S 132˚ 10.70’S, coll. CSIRO CRIMP survey team, 23 Aug. 1998.</p><p>Comparative material. M. mullawa Hutchings &amp; Karageorgopoulos, 2003, Moreton Bay, paratypes 2(AM W27394−5); M. gravelyi Southern, 1921, Rhomba Bay, Chilka Lake, India, paratype 1(BMNH 1938.5.7.55); M. borradailei Pillai 1958, Sri Lanka, lectotype BMNH 1960.3.13.6; Marphysa elityeni Lewis &amp; Karageorgopoulos, 2008, Western Cape of South Africa. Marphysa furcellata Crossland, 1903, town of Zanzibar and Zanzibar channel, Zanzibar, syntypes 2(BMNH 1924.3.1.139). Marphysa macintoshi Crossland, 1904, east and west coasts of Zanzibar, syntypes 3(BMNH 1924:3:1:22−23). Marphysa simplex Crossland, 1903, town of Zanzibar,, syntypes 2(BMNH 1924.3.1.1−2).</p><p>Measurements. Holotype, mature male, complete but fragmented into four parts, 424 chaetigers in total, about 190 mm long, 6.0 mm maximum width. Paratypes range in size from 78−155 mm long, 4.0− 5.5 mm maximum width, 225−415 chaetigers.</p><p>Description (of holotype, values for paratypes in parentheses). Body long and slender, initially cylindrical in cross-section but becoming slightly dorsoventrally flattened after first 6 or 7 chaetigers (6–10), more so in posterior body, which is coiled and fragile when preserved. Live worm, pink coloured with olive pigment, spotted white, dorsally on head and first 6 or 7 segments; prostomial appendages white, except at base which is also olive coloured (Fig. 1A −C).</p><p>Prostomium equal in length to first ring of peristomium mid-dorsally, appearing bilobed as buccal lips separated by deep ventral notch, continuing dorsally as narrow groove to base of median antenna (Fig. 1A, B). Eyespots not visible in holotype; in some paratypes small pair visible on prostomium at base of lateral antennae near border with peristomium. Prostomial appendages smooth (wrinkled in some paratypes), tapered to blunt point, arranged in shallow arc; median antenna equal in length, or slightly longer than (paratypes) lateral antennae; antennae longer than palps (1.2-1.3x) and about 2x (1.5−2.0x) length prostomium (Fig. 1C). Antennae and palps attached to prostomium by narrow peduncle (Fig. 1A, C). Mandibles dark, with lighter coloured cutting plates. Maxillary black, bases of Mx II–V paler; formula: I = 1 + 1, II = 5 + 6, III= 7 + 0, IV = 4 + 9, V = 1 + 1 (NTM W23041).</p><p>Parapodia similar throughout, comprising low, faintly bifid, prechaetal ridge and prominent rounded postchaetal lobe, conical dorsal cirrus and broader ventral cirrus: all three components extend laterally to about same level, except for first few chaetigers where parapodial lobe is shorter (Figs 2A −H). Branchiae arising from superior base of dorsal cirrus, beginning on chaetiger 31 (22−32) and extending posteriorly to last chaetiger (last to 10th last), up to 9 (6−9) pectinate filaments per branchia, best developed on mid-posterior body where they arch over dorsal surface for distance of 1/4−1/5 width of body (Figs 2F −H).</p><p>Chaetae arranged in two bundles separated by oblique row of aciculae, where maximally developed in anterior-mid body (Figs 2C, 3B). Supra-acicular bundle comprising 10−25 limbate capillaries (numbers reducing posteriorly) and 2−4 asymmetrically pectinate chaetae; subacicular bundle comprising up to 20 compound spinigers in anterior chaetigers and posteriorly to chaetiger 55 (13−42) after which they disappear, 10−20 limbate capillaries and single subacicular hook from chaetiger 40 (31−50) (Figs 3 A-I). Supra-acicular limbate capillaries arranged in groups of two or three different lengths, but otherwise similar to each other and those of subacicular fascicle. Compound spinigers appear arranged in two groups each having slightly different blade length, but otherwise similar. Pectinate chaetae with long lateral teeth (about 2–3 times longer than inner teeth) show increasing teeth number, asymmetry and curvature posteriorly (Table 1; Figs 3 G-I); also inner tooth closest to lateral tooth occasionally prolonged producing additional elongated lateral tooth (Fig. 3 H). Posterior chaetigers with one or two types of pectinate chaetae: one with many small teeth (always present) and other with fewer larger teeth (sometimes present) (Table 1; Figs 3 I). Subacicular hooks dark brown, bluntly pointed, bidentate, close-fitting hood (Figs 3E, F). Neuroaciculae black, maximally 4(3) in anterior chaetigers (chaetigers 8−15), reducing to 1 in posterior chaetigers. Notoaciculae slender and curved, pale, 3−4 at base of dorsal cirri from about chaetiger 30 onwards (Figs 2F −H).</p><p>Pygidium bearing two pairs of anal cirri on ventral margin; dorsal most pair about ½ to several times (in paratypes) length of pygidium, ventral most pair papilliform to ½ length (paratypes) of pygidium.</p><p>Variation. Size-dependent variation in both branchial and chaetal patterns has been reported before for Marphysa (e.g., Lu &amp; Fauchald 1998). Although the number of specimens examined here was low (n = 9) and the size range small, we found the same strong size correlations as these authors, viz. larger specimens have branchiae that appear later, a higher number of branchial filaments, subacicular hooks that appear later; and compound spinigers that extend posteriorly over a greater number of chaetigers.</p><p>Remarks. In general appearance, Marphysa fauchaldi n. sp., may be distinguished from the other species of Marphysa occurring in Darwin Harbour ( M. mullawa and M. mossambica) by its smaller size, smooth dorsal epithelium, and the marked flattening of body after the first 6–10 chaetigers. The other two species grow to a larger size, have a rugose, glandular dorsal epithelium, and posterior flattening is much more gradual over the anterior-mid body region. Specifically, the new species differs uniquely from other Group B2 Marphysa species in having fewer compound spinigers, which are present over a smaller range of chaetigers (first 13−55 chaetigers only). Also, the compound spinigers are accompanied by limbate capillaries in the subacicular bundle of the anterior parapodia; initially the compound spinigers out-number the capillaries, but they are gradually replaced by the capillaries until from the anterior-mid body onward only capillaries are present. In this respect the species is approaching the condition of M. mossambica, which completely lacks compound chaetae. The other difference between M. fauchaldi n. sp. and other B2 species is the presence of pectinate chaetae from chaetiger 1, and that they do not exceed 2−4 per bundle (because of this and their small size they are difficult to observe under the compound microscope).</p><p>Etymology. The new species is named after Kristian Fauchald, friend and mentor. The common name, ‘Barra’, is the local name for Barramundi.</p><p>Distribution. Northern Australia, including Darwin Harbour and Cobourg Peninsula, Port Essington.</p><p>Biology. Densities and distributions of M. fauchaldi n. sp., in the aquaculture ponds varied considerably depending on the species farmed. When Tiger Prawns, Penaeus monodon, were present densities were low as a result of predation, and the worms occurred mainly in deep burrows in the clay banks; when Barramundi, Lates calcarifer were present densities were very high (of up to 900 individuals per square metre) and the worms appeared to be evenly distributed over the floor of the pond, although this needs further study. Marphysa fauchaldi n. sp., encapsulates its embryos and larvae in jelly cocoons (Table 2), which seems to be an adaptation for life in unstable estuarine environments. Details of its ecology and biology including larval development and growth are currently being investigated (Glasby et al. in prep.).</p></div>	https://treatment.plazi.org/id/03A687D8FFCCD70C0AC8F68FFBC1F81E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Glasby, Christopher J.;Hutchings, Pat A.	Glasby, Christopher J., Hutchings, Pat A. (2010): A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352: 29-45, DOI: 10.5281/zenodo.193484
03A687D8FFC7D7010AC8F7C4F809F9B3.text	03A687D8FFC7D7010AC8F7C4F809F9B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marphysa mossambica (Peters 1854) Peters 1854	<div><p>Marphysa mossambica (Peters, 1854)</p><p>Eunice mossambica Peters, 1854: 612 .</p><p>Nauphanta novae Hollandiae Kinberg, 1865: 564; 1910: 43, pl. 16, fig. 23, 23B, C, F, G.</p><p>Marphysa mossambica . – Gravier, 1900: 267, pl. 14, figs 89−90, text figs. 137−137; Crossland, 1903: 139 −140, pl. 15, figs 7−10; Day 1967, 395, fig. 17.5 i −m.</p><p>Marphysa simplex Treadwell, 1922: 151 −152, text-fig. 39, pl. 5, figs 8−12.</p><p>Nauphanta mossambica . – Fauchald, 1987: 376 −378, fig. 1.</p><p>Material examined. Australia, Northern Territory, Ludmilla Creek mouth, 12˚24.8' S 130˚50.0' E 1(NTM W60), 1(NTM W61), coll. R.J. Hanley, 18 Dec. 1981, inside rotting timber among mangroves; same location 1(NTM W67), coll. R.J. Hanley, 26 Nov 1981, burrowing in mangrove mud; same location, 2(NTM W153), coll. R.J. Hanley, 12 Mar. 1982, in mud among roots of Rhizophora stylosa, Gunn Point, 12˚9.5' S 131˚00.5' E, 2(NTM W154), coll. R.J. Hanley, 4 Oct. 1981, inside rotting timber, R. stylosa, Field Island, 12˚06.90’S 132˚25.20’E, muddy reef, coll. C.J. Glasby, 20 Aug. 2004, 1(NTM W23043). Western Australia, Kimberley region, coll. R.J. Hanley, 1994, RH 94/11, 1(NTM W23044), 5(NTM W23045).</p><p>Comparative material. Marphysa mossambica sensu Monro, 1931, Low Isles, Queensland, 16˚23’S 145˚34’E, Great Barrier Reef, Sts A19, 19a, coll. 18.4.1929, 2(AM W2956).</p><p>Description. Present material ranged from 2.2–9.0 mm maximum body width. Branchiae first present on chaetiger 14−46, maximum number of filaments (3−6) on mid-posterior body. Pectinate chaetae asymmetrical throughout, with 15–25 teeth in anterior chaetigers, 30−40 teeth in posterior chaetigers; bidentate subacicular hooks from chaetigers 23−68, continue as one per parapodium for several chaetigers then disappear for few chaetigers, but may reappear again later [Crossland (1903) also commented on the loss of chaetae in the mid- and posterior body parapodia, particularly subacicular hooks, in this species; but this feature may be more widespread across the family (Zanol et al. 2007)].</p><p>Remarks. The synonymy of Nauphanta novaehollandiae with M. mossambica, first proposed by Gravier (1900) was supported by Crossland (1903) and Augener (1922). However, Fauchald (1987) disagreed, considering that the types of both species differed sufficiently such that the two species could be recognised, viz. ‘branchiae are present from setiger 30 and subacicular hooks from setiger 44 in Nauphanta novaehollandiae; branchiae are present from setiger 37 or later and subacicular hooks not until setiger 58 in N. mossambica ’. Also, he found slight (but unspecified) differences in shape of the subacicular hooks. However as demonstrated below, the differences in where the branchiae and hooks start can be accounted for by size-related variation taking into consideration the much smaller size of the holotype of Nauphanta novaehollandiae compared to the lectotype of N. mossambica (117 chaetigers, 4 mm wide compared to 420 chaetigers, 10 mm wide) (Fig. 4A, B). The range of variation in these two characters in our material (branchiae from chaetigers 14−46, subacicular hooks from chaetigers 23−68) encompasses both species.</p><p>The present material shows a positive linear relationship between body size (x) and the chaetiger on which the branchiae and subacicular hooks appear, as follows:</p><p>Branchiae, y = 5.2999x – 1.8029 (r2 = 0.94; n = 15; P &lt;0.001)</p><p>Subacicular hooks, y = 7.4469x + 5.0369 (r2 = 0.89; n = 15; P &lt;0.001)</p><p>When the data for the lectotype of M. mossambica (which at 10 mm wide is about the same size as the larger specimens in this study) and the holotype of M. novaehollandiae were included in the analysis the regression values decreased slightly for both branchiae and subacicular hooks (viz. r2 = 0.86 and r2 = 0.87 respectively), but the change was not statistically significant (P &lt;0.001; Fig. 4A, B). Therefore, it is highly likely that the northern Australian forms, the lectotype of M. mossambica from Mozambique, and the holotype of M. novaehollandiae represent populations of a single species.</p><p>The synonymy of Marphysa simplex Treadwell, 1922, described from Suva Harbour, Fiji with Marphysa mossambica is newly proposed. Although Treadwell apparently misreported the chaetiger (=somite) on which the branchiae begin as somite 242 (the total number of somites was reported to be about 200!), all other key features agree with those of a small-sized specimen of Marphysa mossambica . Notwithstanding the relegation of Marphysa simplex Treadwell, 1922 to junior synonymy, this is the second case of secondary homonymy in the genus, the senior homonym being Marphysa simplex Langerhans, 1884 (as Amphiro simplex) from Madeira.</p><p>Other records of M. mossambica from Australian waters are confused. The record of the species from Saint Vincent Gulf South Australia by Fauvel (1917) is a misidentification as he illustrates the presence of a compound spiniger, which places it in the B2 Marphysa group. The second report of the species from Australia—from the Low Isles, Great Barrier Reef by Monro (1931) —could not be verified as the two specimens now lack anterior chaetigers including the head. The only other records of the species from Australia are Kinberg’s original description of N. novaehollandiae from Sydney Harbour (holotype later redescribed by Fauchald 1987), and Augener’s (1922) report of Marphysa novaehollandiae from Cape York, Queensland.</p><p>Distribution. Tropical and subtropical Indo-west Pacific including Zanzibar, Mozambique, Red Sea, northern Australia, Philippines and Fiji.</p><p>M. fauchaldi M. borradailei M. furcellata M. graveleyi M. macintoshi M. M. mullawa M. M. teretiuscula M. tamurai</p><p>n. sp. mossambica orientalis (= M. simplex)</p><p>Mandibles lighter black and lighter lighter black, no dark brown, brown, white dark brown, white edging dark brown,</p><p>coloured white coloured coloured white edging lighter cutting plate white present whitish</p><p>cutting plate cutting plate cutting plate coloured encrustation anterior plates</p><p>cutting plate on anterior</p><p>plates Maxillae black with black and? mostly light black, no brown, edges uniform light uniformly white edging?</p><p>paler bases white coloured white edging and sutures brown dark present</p><p>darker brown</p><p>Mx II – no. teeth (one 5+6 6 5 5+6 4+5 5–7 4 3 3–4 4 side)</p><p>(present</p><p>observations) continued next page</p></div>	https://treatment.plazi.org/id/03A687D8FFC7D7010AC8F7C4F809F9B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Glasby, Christopher J.;Hutchings, Pat A.	Glasby, Christopher J., Hutchings, Pat A. (2010): A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352: 29-45, DOI: 10.5281/zenodo.193484
03A687D8FFC2D7070AC8F29FFE2DFAD0.text	03A687D8FFC2D7070AC8F29FFE2DFAD0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Marphysa mullawa Hutchings & Karageorgopolous 2003	<div><p>Marphysa mullawa Hutchings &amp; Karageorgopolous, 2003</p><p>Marphysa mullawa Hutchings &amp; Karageorgopolous, 2003: 90 −93, figs 2a −f; 3, b, d; 4b, d (note corrected figure references—figures and captions in original are jumbled).</p><p>Material examined. PARATYPES: Australia, Queensland, Moreton Bay, Fisherman's Island, mudflat, 27° 15' S, 153° 15' E, coll. Sep. 1996, 1 (AM W27394), 1(AM W27395). NON-TYPES: Australia, Queensland, Cairns marina, 16.9˚ S 145.8 ˚E, scraping off wharf pile, 3m, coll. CRC Reef Research Limited, 19 Nov. 2001, 1(AM W30786), 10 m sled sample, coll. CRC Reef Research Limited, 16 Nov. 2001, 1(AM W35251), Weipa, 12˚40.34’S 141˚52.12’E, scaping of pylon in port, 0.5 m, coll. CRC Reef Research Limited, Oct. 1999, 1 (AM W35250), Northern Territory, Darwin Harbour, Creek H, CH 9, mudflat, coll. R. Hanley, 13 Mar. 1986, 1(NTM W3897), 1(NTM W3898); Cullen Bay marina, Stn P3-3 NTDCBM, 12˚ 27.16’S 130˚ 49.30’E, coll. CSIRO CRIMP survey team, 14 Aug. 1998, 1(NTM W17768); Leaders Creek, 12˚10.766’S 131˚06.607”E, coll. C. Glasby &amp; T. Bakken, 21 Sept. 2000, 8(NTM W23047). Western Australia, Kimberley region, coll. R.J. Hanely, 1995, RH 95/5, 1(NTM W23046).</p><p>Remarks. The present material closely resembles the type material of M. mullawa, except subacicular hooks occur from chaetigers 26−36, which is just outside the range of 37−45 given by Hutchings &amp; Karageorgopolous (2003) for the type material. However, the early appearance of the hooks is accountable by the smaller size of the present specimens (2.8−3.5 mm wide). As none of the specimens are posteriorly complete we were unable to determine the types of pectinate chaetae in far posterior chaetigers; however, in the material at hand, pectinate chaetae in mid-posterior chaetigers had about 22 teeth—the coarse-tooth types were not observed. The most complete specimen with about 250 chaetigers lacks entirely any limbate capillaries in the subacicular bundle.</p><p>Marphysa mullawa belongs to the Sanguinea-group of species, which lack limbate capillaries in the subacicular bundle. Apart from M. sanguinea the group also includes M. orientalis Treadwell, 1936 from Amoy, China and M. elityeni Lewis &amp; Karageorgopolous, 2008 from South Africa. Another species that may belong to this group is M. tamurai Okuda, 1934, but information on the presence or absence of limbate capillaries in this species is lacking.</p><p>Habitat. Intertidal to 10 m; muddy sediments and amongst invertebrate fouling communities in protected coastal embayments including harbours and estuaries.</p><p>Distribution. Northern Australia from Kimberley, Western Australia in the west to Moreton Bay, Queensland in the east.</p></div>	https://treatment.plazi.org/id/03A687D8FFC2D7070AC8F29FFE2DFAD0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Glasby, Christopher J.;Hutchings, Pat A.	Glasby, Christopher J., Hutchings, Pat A. (2010): A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352: 29-45, DOI: 10.5281/zenodo.193484
