taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03A787D8FFC90B2EA3EEFC903091CEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12821050/files/figure.png	https://doi.org/10.5281/zenodo.12821050	Fig. 1. Male Austruca variegata (Heller, 1862) (a–d) and Cranuca inversa (Hoffmann, 1874) (e). (a) habitus; (b, c) major cheliped; (d, e) merus of right major cheliped. (a, b) lectotype, CW 17.2 mm (PL 24.7 mm, NHMW 25656; Madras, India), originally identified as A. perplexa. (c) “Gelasimus perplexus” in Heller (1865: pl. 5(4)). (d) male (CW 18.6 mm, ZRC 2018.1375; Tamil Nadu, India). (e) CW 19.9 mm (NCHUZOOL 14904; Al Darb, Arabia). Scale bars = 5.0 mm.	Fig. 1. Male Austruca variegata (Heller, 1862) (a–d) and Cranuca inversa (Hoffmann, 1874) (e). (a) habitus; (b, c) major cheliped; (d, e) merus of right major cheliped. (a, b) lectotype, CW 17.2 mm (PL 24.7 mm, NHMW 25656; Madras, India), originally identified as A. perplexa. (c) “Gelasimus perplexus” in Heller (1865: pl. 5(4)). (d) male (CW 18.6 mm, ZRC 2018.1375; Tamil Nadu, India). (e) CW 19.9 mm (NCHUZOOL 14904; Al Darb, Arabia). Scale bars = 5.0 mm.	2019-07-02	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish		Zenodo	biologists	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish			
03A787D8FFC90B2EA3EEFC903091CEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12821052/files/figure.png	https://doi.org/10.5281/zenodo.12821052	Fig. 2. Austruca variegata (Heller, 1862). a–d, male (CW 18.6 mm, CL 10.7 mm, PL 26.6 mm, ZRC 2018.1375). (a) habitus; (b) major cheliped; (c) pleon; (d) minor cheliped. Scale bar = 5.0 mm.	Fig. 2. Austruca variegata (Heller, 1862). a–d, male (CW 18.6 mm, CL 10.7 mm, PL 26.6 mm, ZRC 2018.1375). (a) habitus; (b) major cheliped; (c) pleon; (d) minor cheliped. Scale bar = 5.0 mm.	2019-07-02	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish		Zenodo	biologists	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish			
03A787D8FFC90B2EA3EEFC903091CEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12821054/files/figure.png	https://doi.org/10.5281/zenodo.12821054	Fig. 3. Carapaces of male Austruca variegata (Heller, 1862) (a, b), A. bengali (Crane, 1975) (c) and A. triangularis (A. Milne-Edwards, 1873) (d). (a) CW 16.6 mm (ZRC 2001.0853; left-handed; Tamil Nadu, India); (b) CW 14.5 mm (ZRC 2017.0917; left-handed; West Bengal, India); (c) CW 14.5 mm (NCHUZOOL 14345; right-handed; Selangor, Malaysia); (d) CW 14.8 mm (NCHUZOOL 13574; lefthanded; Cebu, Philippines).	Fig. 3. Carapaces of male Austruca variegata (Heller, 1862) (a, b), A. bengali (Crane, 1975) (c) and A. triangularis (A. Milne-Edwards, 1873) (d). (a) CW 16.6 mm (ZRC 2001.0853; left-handed; Tamil Nadu, India); (b) CW 14.5 mm (ZRC 2017.0917; left-handed; West Bengal, India); (c) CW 14.5 mm (NCHUZOOL 14345; right-handed; Selangor, Malaysia); (d) CW 14.8 mm (NCHUZOOL 13574; lefthanded; Cebu, Philippines).	2019-07-02	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish		Zenodo	biologists	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish			
03A787D8FFC90B2EA3EEFC903091CEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12821056/files/figure.png	https://doi.org/10.5281/zenodo.12821056	Fig. 4. Floors of orbit of male Austruca variegata (Heller, 1862) (a–d), A. bengali (Crane, 1975) (e, f) and A. triangularis (A. Milne-Edwards, 1873) (g, h). (a, b) CW 18.6 mm (ZRC 2018.1375; right-handed; Tamil Nadu, India); (c, d) CW 14.5 mm (ZRC 2017.0917; left-handed; West Bengal, India); (e, f) CW 14.5 mm (NCHUZOOL 14345; right-handed; Selangor, Malaysia); (g, h) CW 14.8 mm (NCHUZOOL 13574; left-handed; Cebu, Philippines). Scale bars = 5.0 mm.	Fig. 4. Floors of orbit of male Austruca variegata (Heller, 1862) (a–d), A. bengali (Crane, 1975) (e, f) and A. triangularis (A. Milne-Edwards, 1873) (g, h). (a, b) CW 18.6 mm (ZRC 2018.1375; right-handed; Tamil Nadu, India); (c, d) CW 14.5 mm (ZRC 2017.0917; left-handed; West Bengal, India); (e, f) CW 14.5 mm (NCHUZOOL 14345; right-handed; Selangor, Malaysia); (g, h) CW 14.8 mm (NCHUZOOL 13574; left-handed; Cebu, Philippines). Scale bars = 5.0 mm.	2019-07-02	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish		Zenodo	biologists	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish			
03A787D8FFC90B2EA3EEFC903091CEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12821058/files/figure.png	https://doi.org/10.5281/zenodo.12821058	Fig. 5. Outer and inner sides of major palm of Austruca variegata (Heller, 1862) (a–d), A. bengali (Crane, 1975) (e, f) and A. triangularis (A. Milne- Edwards, 1873) (g, h). (a, b) CW 16.6 mm (ZRC 2001.0853; left-handed; Tamil Nadu, India); (c, d) CW 14.5 mm (ZRC 2017.0917; left-handed; West Bengal, India); (e, f) CW 14.5 mm (NCHUZOOL 14345; right-handed; Selangor, Malaysia); (g, h) CW 14.8 mm (NCHUZOOL 13574; lefthanded; Cebu, Philippines).	Fig. 5. Outer and inner sides of major palm of Austruca variegata (Heller, 1862) (a–d), A. bengali (Crane, 1975) (e, f) and A. triangularis (A. Milne- Edwards, 1873) (g, h). (a, b) CW 16.6 mm (ZRC 2001.0853; left-handed; Tamil Nadu, India); (c, d) CW 14.5 mm (ZRC 2017.0917; left-handed; West Bengal, India); (e, f) CW 14.5 mm (NCHUZOOL 14345; right-handed; Selangor, Malaysia); (g, h) CW 14.8 mm (NCHUZOOL 13574; lefthanded; Cebu, Philippines).	2019-07-02	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish		Zenodo	biologists	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish			
03A787D8FFC90B2EA3EEFC903091CEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12821060/files/figure.png	https://doi.org/10.5281/zenodo.12821060	Fig. 6. Right G1s of Austruca variegata (Heller, 1862) (a, b, c, f), A. bengali (Crane, 1975) (d, g) and A. triangularis (A. Milne-Edwards, 1873) (e, h). (a, f–h) lateral view of distal part; (b) mesial view of distal part; (c–e) lateral view. (a, b) CW 16.6 mm (ZRC 2001.0853; Tamil Nadu, India); (c, f) CW CW 14.5 mm (ZRC 2017.0917; West Bengal, India); (d, g) 14.5 mm (NCHUZOOL 14345; Selangor, Malaysia); (e, h) 14.8 mm (NCHUZOOL 13574; Cebu, Philippines). Scale bars: a, b, f–h = 0.5 mm, c–e = 1.0 mm.	Fig. 6. Right G1s of Austruca variegata (Heller, 1862) (a, b, c, f), A. bengali (Crane, 1975) (d, g) and A. triangularis (A. Milne-Edwards, 1873) (e, h). (a, f–h) lateral view of distal part; (b) mesial view of distal part; (c–e) lateral view. (a, b) CW 16.6 mm (ZRC 2001.0853; Tamil Nadu, India); (c, f) CW CW 14.5 mm (ZRC 2017.0917; West Bengal, India); (d, g) 14.5 mm (NCHUZOOL 14345; Selangor, Malaysia); (e, h) 14.8 mm (NCHUZOOL 13574; Cebu, Philippines). Scale bars: a, b, f–h = 0.5 mm, c–e = 1.0 mm.	2019-07-02	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish		Zenodo	biologists	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish			
03A787D8FFC90B2EA3EEFC903091CEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12821062/files/figure.png	https://doi.org/10.5281/zenodo.12821062	Fig. 7. Urocardiac ossicles of Austruca variegata (Heller, 1862) (a, b, e), A. bengali (Crane, 1975) (c, f) and A. triangularis (A. Milne-Edwards, 1873) (d, g). (a) male (CW 16.1 mm, ZRC 2003.0463; Tamil Nadu, India); (b, e) female (CW 14.5 mm, ZRC 2003.0463; Tamil Nadu, India); (c) male (CW 13.1 mm, QM W27320; Phuket, Thailand); (d) male (CW 14.8 mm, NCHUZOOL 13574; Cebu, Philippines); (f) male (CW 12.5 mm, QM W27320; Phuket, Thailand); (g) female (CW 14.6 mm, NCHUZOOL 14350; Cebu, Philippines). Scale bars = 0.5 mm.	Fig. 7. Urocardiac ossicles of Austruca variegata (Heller, 1862) (a, b, e), A. bengali (Crane, 1975) (c, f) and A. triangularis (A. Milne-Edwards, 1873) (d, g). (a) male (CW 16.1 mm, ZRC 2003.0463; Tamil Nadu, India); (b, e) female (CW 14.5 mm, ZRC 2003.0463; Tamil Nadu, India); (c) male (CW 13.1 mm, QM W27320; Phuket, Thailand); (d) male (CW 14.8 mm, NCHUZOOL 13574; Cebu, Philippines); (f) male (CW 12.5 mm, QM W27320; Phuket, Thailand); (g) female (CW 14.6 mm, NCHUZOOL 14350; Cebu, Philippines). Scale bars = 0.5 mm.	2019-07-02	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish		Zenodo	biologists	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish			
03A787D8FFC90B2EA3EEFC903091CEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12821064/files/figure.png	https://doi.org/10.5281/zenodo.12821064	Fig. 8. Female Austruca variegata (Heller, 1862) (a–d), A. bengali (Crane, 1975) (e) and A. triangularis (A. Milne-Edwards, 1873) (f). (a) habitus; (b) floor of right orbit; (c) right minor cheliped; (d–f) right vulva (gonopore). (a–d) CW 16.6 mm (NCHUZOOL 14365; Tamil Nadu, India); (e) CW 12.4 mm (QM W27320; Phuket, Thailand); (f) CW 14.6 mm (NCHUZOOL 14350; Cebu, Philippines). Scale bars: b, c = 5.0 mm, d–f = 0.5 mm	Fig. 8. Female Austruca variegata (Heller, 1862) (a–d), A. bengali (Crane, 1975) (e) and A. triangularis (A. Milne-Edwards, 1873) (f). (a) habitus; (b) floor of right orbit; (c) right minor cheliped; (d–f) right vulva (gonopore). (a–d) CW 16.6 mm (NCHUZOOL 14365; Tamil Nadu, India); (e) CW 12.4 mm (QM W27320; Phuket, Thailand); (f) CW 14.6 mm (NCHUZOOL 14350; Cebu, Philippines). Scale bars: b, c = 5.0 mm, d–f = 0.5 mm	2019-07-02	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish		Zenodo	biologists	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish			
03A787D8FFC90B2EA3EEFC903091CEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12821066/files/figure.png	https://doi.org/10.5281/zenodo.12821066	Fig. 9. The live coloration of Austruca variegata (Heller, 1862) (a–h). (a, b) dorsal view of male; (c, d) frontal view of male; (e) dorsal view of female; (f) two males fighting; (g) two males moving to low intertidal zone for feeding; (h) a male A. variegata (left) and a male A. annulipes (H. Milne Edwards, 1837) (right) sympatric in one locality. Specimens not captured. Photos taken from Vellar River estuary, Porto Novo, Tamil Nadu, India.	Fig. 9. The live coloration of Austruca variegata (Heller, 1862) (a–h). (a, b) dorsal view of male; (c, d) frontal view of male; (e) dorsal view of female; (f) two males fighting; (g) two males moving to low intertidal zone for feeding; (h) a male A. variegata (left) and a male A. annulipes (H. Milne Edwards, 1837) (right) sympatric in one locality. Specimens not captured. Photos taken from Vellar River estuary, Porto Novo, Tamil Nadu, India.	2019-07-02	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish		Zenodo	biologists	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish			
03A787D8FFC90B2EA3EEFC903091CEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12821068/files/figure.png	https://doi.org/10.5281/zenodo.12821068	Fig. 10. The live coloration of Austruca bengali (Crane, 1975) (a–g). (a, b) dorsal view of adult male; (c, d) dorsal view of juvenile male; (e) frontal view of adult male; (f) frontal view of juvenile female; (g) a female in the field (not captured; Phuket, Thailand); (h) habitat in Phuket, Thailand. (a) male (not captured; Selangor, Malaysia); (b) male (not captured; Phuket, Thailand); (c, d) male (CW 7.3, 7.8 mm, NCHUZOOL 14361; Phuket, Thailand); (e) male (CW 14.5 mm, NCHUZOOL 14345; Selangor, Malaysia); (f) female (CW 8.1 cm, NCHUZOOL 14361; Phuket, Thailand).	Fig. 10. The live coloration of Austruca bengali (Crane, 1975) (a–g). (a, b) dorsal view of adult male; (c, d) dorsal view of juvenile male; (e) frontal view of adult male; (f) frontal view of juvenile female; (g) a female in the field (not captured; Phuket, Thailand); (h) habitat in Phuket, Thailand. (a) male (not captured; Selangor, Malaysia); (b) male (not captured; Phuket, Thailand); (c, d) male (CW 7.3, 7.8 mm, NCHUZOOL 14361; Phuket, Thailand); (e) male (CW 14.5 mm, NCHUZOOL 14345; Selangor, Malaysia); (f) female (CW 8.1 cm, NCHUZOOL 14361; Phuket, Thailand).	2019-07-02	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish		Zenodo	biologists	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish			
03A787D8FFC90B2EA3EEFC903091CEB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/12821070/files/figure.png	https://doi.org/10.5281/zenodo.12821070	Fig. 11. The live coloration of Austruca triangularis (A. Milne-Edwards, 1873). (a–d) dorsal view of adult male; (e, f) dorsal view of juvenile male; (g, h) frontal view of adult male. (a, g) NCHUZOOL 14347 (140716, Bali, Indonesia); (b, h) male (CW 11.5 mm, NCHUZOOL 14349; Cebu, Philippines); (c, d) specimens lost (Baoli River estuary, Pingtung, Taiwan); (f) NCHUZOOL 14346 (Labuan, Malaysia).	Fig. 11. The live coloration of Austruca triangularis (A. Milne-Edwards, 1873). (a–d) dorsal view of adult male; (e, f) dorsal view of juvenile male; (g, h) frontal view of adult male. (a, g) NCHUZOOL 14347 (140716, Bali, Indonesia); (b, h) male (CW 11.5 mm, NCHUZOOL 14349; Cebu, Philippines); (c, d) specimens lost (Baoli River estuary, Pingtung, Taiwan); (f) NCHUZOOL 14346 (Labuan, Malaysia).	2019-07-02	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish		Zenodo	biologists	Lin, Yu-Jia;Qurban, Mohammad A.;Shen, Kang Ning;Chao, Ning Labbish			
