taxonID	type	description	language	source
03A7BA3FC92FFFDFFF53C18FFC2D35F7.taxon	materials_examined	Type species: Topalia problematica Balogh & Csiszar, 1963, p. 477, Figs. 27, 28. Diagnosis: With the following combination of character states: broad lamellae covering most of prodorsum; setae ro, le and in minute; ten pairs of minute notogastral setae; pedotectum I and II prominent. Subcapitulum narrow; chelicerae slim, sub-pelopsiform, dentate; Epimeral plates I trapezoid or sub-rectangular, much broader than long; epimeral plates III and IV sub-rectangular, separate or fused; considerably shorter / narrower than I and II. Discidium well-developed, pointed; enantiophysis E 4 and perigenital carina present; with four pairs of genital setae. Tibia I with solenidion φ 1 lacking apophysis, or at most with a slight bulge; set back from the anterior margin of tibia. Tarsi monodactylous. Description: Non-poronotic brachypyline oribatid mites. Species with both males and females, though no indication of sexual dimorphism. With cerotegument of granular, reticulate and / or tuberculate microsculpture covering dorsal and ventral surfaces. Rostrum acute, entire or incised. Lamellae wide, covering most of prodorsum and bothridia (Figs. 1 a, 2 a); lamellar cusps extending anterior of rostrum, rounded apically, fused medially at base, with a short translamella; rostral, lamellar and interlamellar setae minute; lamellar setae positioned anterodorsally on cusp, with long insertion canal (xx). Bothridial seta club-shaped; stalk generally short; about as long as head (Fig. 2 e). Bothridia with internal ring-like thickenings. Tutorium positioned ventral of lamellae, with or without cusp. Notogaster only slightly longer than wide, with rounded humeral projections and ten pairs of minute setae; c 1, c 3, da, dp and lm absent; dm in centrodorsal position, others marginal. Pedotectum I and II massive; pedotectum I covering acetabulum I, extending dorsally at least one-third of length of lamellae, pedotectum II triangular in lateral aspect, free end pointed and strongly curved dorsally. Discidium strongly developed between acetabula III and IV; acutely pointed in ventral aspect. Subcapitulum narrow, about twice as long as broad; rutellum attenuated, with curved, sharp teeth; mentum without tectum (Fig 6 b). Chelicera narrow, sub-pelopsiform, dentate, with opposing digits (Fig. 2 c). Palp tarsus with seven setae; solenidion not on apophysis, separate from seta acm (Fig. 2 d). Epimeral plates I trapezoid or sub-rectangular, much broader than long; epimeral plates II longer than broad; epimera III and IV sub-rectangular, much shorter than I and II, their lateral margins extending distance of about half length of epimeral plates II (Fig. 3 a); epimeral setation typically 2 - 1 - 2 - 3 or 2 - 1 - 2 - 2. Enantiophysis E 4 present; E 2 and / or V occasionally present (Fig. 4 b). Anteriorly-projecting condyle of E 4 on apex of lath-like carina lateral of genital plate (Fig. 1 b, pc). With four pairs of genital setae, posterior two pairs separated from anterior ones; one pair of aggenital setae between genital plate and lateral ridge; two pairs of anal setae on main body of anal plate, not on margin; two pairs of adanal setae (Fig. 1 b). Anal and genital plates adjacent or close together, each surrounded by ring of sclerotised cuticle; anal and genital plates similar in shape: oval-to-sub-rectangular; anal plates never lozenge-shaped; lyrifissure iad present in para-anal position; ventral plate rounded caudally. With tectum on ventral plate at level of circumgastric scissure (vpt), visible in lateral view; gouttière de coaptation au notogaster (tcn) of Grandjean (1958, p. 122). Legs lacking porose areas or saccules (Figs. 3, 8 a); seta d absent from genua I, II and tibiae I – III. Trochanters and femorae broadly ovoid, flattened and keeled ventrally; solenidia φ 1 and φ 2 on tibia I emerging from flat cuticular surface, not on prominent cylindrical apophyses. Tarsi long, slender, monodactylous. Remarks: Balogh & Csiszar (1963) defined Topalia as follows: “ lamellae meeting in median line, wide, covering prodorsum and rostrum, lamellar and interlamellar setae very small. Tectopedium I very large, shell-shaped. Four pairs of genital setae. Legs monodactylous. Body with granulated cerotegument. ” This definition, while adequate at the time, would now include the morphologically most similar genus Nosybea and, apart from it having five (rather than four) genital setae, Lamellocepheus. Further, the following species of Topalia have broad lamellae that do not meet in the midline: T. caliginosa sp. nov., T. katyae sp. nov., T. royi sp. nov. and T. velata. Mahunka (1993) distinguished Nosybea from Topalia based on the following character states: (1) notogaster with one unpaired median crest rather than 1 – 2 pairs of longitudinal crests; (2) notogaster with lateral excavations (cavernae) rather than unexcavated; (3) coxisternal region without opposite apophyses (i. e. enantiophysis E 4 and associated perigenital carinae) and with 12 ring-shaped thickenings rather than with opposite apophyses but without ring-shaped thickenings; (4) epimeral setal formula of Nosybea 2 - 0 - 2 - 3 rather than 3 - 1 - 3 - 3 (which is not the formula for Topalia; cf. above). This diagnosis requires modification in light of the current conception of Topalia because the presence of ring-shaped excavations on the coxisternite (epimeral foveolae) is shared by Topalia katyae sp. nov. (cf. below, Fig. 6 b). However, in this species, there are only two pairs, located medially between epimera I, II and III. In Nosybea genavensis some of the epimeral foveolae are not paired, there are twelve of them, and more heavily sclerotised than in T. katyae sp. nov., and distributed over the entire epimeral region (Fig. 12 b). Nevertheless, the narrow median notogastral carina of Nosybea is clearly differentiated from the broad, flat centrodorsal ridge of Topalia, as is the reduced epimeral setation of 2 - 0 - 2 - 3, the lateral cavernae and the absence of enantiophysis E 4 and perigenital carinae. Additional characters that distinguish Nosybea from Topalia are: (1) bothridial seta asymmetrically dilated and with spicules, rather than club-shaped with blunt spines (Figs. 10 a, b); (2) with pedotectum II divided by median carina (visible in lateral view), rather than entire; (3) 11 pairs of notogastral setae rather than ten: Nosybea retains seta lm, which is absent in Topalia (Figs. 1 a, 11 a, b).	en	Colloff, Matthew J. (2019): The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera. Zootaxa 4647 (1): 290-321, DOI: 10.11646/zootaxa.4647.1.18
03A7BA3FC92EFFD9FF53C373FEEE3008.taxon	description	(Figs. 1 – 3, 11 a, 12 a, 14 a)	en	Colloff, Matthew J. (2019): The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera. Zootaxa 4647 (1): 290-321, DOI: 10.11646/zootaxa.4647.1.18
03A7BA3FC92EFFD9FF53C373FEEE3008.taxon	materials_examined	Dimensions. Holotype length 378 μm, breadth 233 μm. Paratypes: mean length (n = 11) 370 μm (range 340 – 394 μm); mean breadth 226 μm (range 204 – 236 μm). Ratio of prodorsum to total length: 0.34 (holotype). Adult. Integument: prodorsum with sparsely-distributed tubercles. Notogaster with coarse irregular granules and tubercles overlaying fine pattern of angular foveolae. Cerotegument sparing or absent in centrodorsal region. Underlying cuticle smooth. Prodorsum: rostrum incised, with two lateral teeth and shorter median tooth. Lamellae separated along midline, incurved apically, with narrow anterior translamella positioned ventrally. Interlamellar region with sub-semicircular ridge bearing setae in on squat tubercles. Bothridial seta short, with attenuated club-shaped apices with blunt spines. Notogaster: ratio of length to breadth: 1.1. Anterior margin of notogaster straight, transverse, then indented posterior of bothridia. Humeral projections (h) rounded, bearing seta c 2, incurved posteriorly at level of lyrifissurae ia. Margins of centrodorsal ridge well-developed, converging anterior of setae dm, then diverging; posterior part of ridge rounded. Setae of p series visible only in ventral aspect. notogastral setae on squat tubercles. Ventral aspect: subcapitulum narrow; with very short spiniform setae h; rutellum pointed with three minute, curved, sharp teeth. Chelicerae slim, narrow; digits with reduced number of teeth. Palp setal formula 0 - 2 - 1 - 2 - 7. Epimeral plates I one-third broader (anterior to posterior) than long (lateral to medial); sub-rectangular to ovoid; epimera III and IV partly fused transversely and along mid-line; epimeral setation 3 - 1 - 2 - 2. Perigenital carina curved, ca. 35 µm long, extending posteriorly to point level with seta g 3 (Fig. 1 b). Genital plates 40 µm long, same length as anal plates and twice length of distance between them. Lateral aspect: apices of lamellar cusps curved ventrally and medially. Interlamellar region slightly domed. Anterior margin of rostrum between tutorium (tu) and rostral seta strongly concave, becoming vertical dorsally. Tutorium with short free cusp. Anterior margin of pedotectum I rounded, convex; apex of pedotectum II extending almost as far as level of dorsal margin of acetabulum of leg I. Centrodorsal ridge protruding to level of interlamellar region, initially concave, becoming flattened, extending posteriorly as far as level of insertion points of setae h 2; region immediately posterior slightly concave. Legs: Tarsus I with 16 setae including short setiform famulus (Fig. 3 a); solenidia ω 1 and ω 2 subequal in length (33 µm); tibia I with solenidion φ 1 very long (122 µm), flagelliform. Genu I bearing short solenidion σ. Femur IV with one dorsal and one ventral seta. Tarsus IV with 12 setae (Fig. 3 b). Material Examined and Locality Data. Holotype and 24 paratypes: ANIC 4016, moss on Sassafras (Atherosperma moschatum), cool temperate rainforest, 1077 m., Errinundra Saddle, Errinundra National Park, Victoria, 37 ° 19 ’ S 148 ° 51 ’ E, coll. M. J. Colloff, 29. ix. 2009. Etymology. The specific name is derived from the Latin caligo (mist), referring to the weather conditions common at its type locality. Diagnosis. Topalia caliginosa sp. nov. can be distinguished from all other species in the genus by the following combination of characters: (1) the rostrum incised medially; (2) the anteromedial edges of lamellae separated almost as far posteriorly as insertion points of setae in; (3) the domed interlamellar region; (4) the anterior margin of notogaster transverse medially, then stepped posterior of bothridia; (5) epimera III and IV partly fused; (6) epimeral setation 3 - 1 - 2 - 2. Remarks. Topalia caliginosa sp. nov. is morphologically most similar to T. royi sp. nov., with which it shares the semi-fused epimera III and IV, setae in on squat tubercles, the medially-separated lamellae and the pattern of angular foveolae of the notogastral cerotegument. Topalia caliginosa sp. nov. differs from T. royi sp. nov. in the transverse and stepped anterior margin of the notogaster, rather than wavy and indented medially; the smooth lamellae with a sparingly granular cerotegument, rather than with fine striae and with a divided rostrum rather than an entire one.	en	Colloff, Matthew J. (2019): The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera. Zootaxa 4647 (1): 290-321, DOI: 10.11646/zootaxa.4647.1.18
03A7BA3FC92BFFDBFF53C513FAD4300D.taxon	description	(Fig. 4)	en	Colloff, Matthew J. (2019): The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera. Zootaxa 4647 (1): 290-321, DOI: 10.11646/zootaxa.4647.1.18
03A7BA3FC92BFFDBFF53C513FAD4300D.taxon	materials_examined	Dimensions. Holotype length 416 μm, breadth 252 μm. Paratype length 397 μm, breadth 237 μm. Ratio of prodor- sum to total length: 0.35 (holotype). Adult. Integument: prodorsum with sparsely-distributed tubercles in interlamellar region. Notogaster with small granules; angular foveolae lacking. Cerotegument sparing or absent in centrodorsal region. Underlying cuticle smooth. Prodorsum: rostrum incised, with two lateral teeth but no median tooth. Lamellae meeting along anterior midline and with pattern of fine, transverse striae. Interlamellar region slightly convex, smooth; setae in on squat tubercles on posteromedian margins of lamellae. Bothridial seta with elongated, ovoid, club-shaped apices bearing blunt spines. Notogaster: ratio of length to breadth: 1.13. Anterior margin of notogaster wavy, indented medially. Humeral projections (h) rounded, well-developed, bearing seta c 2, incurved posteriorly at level of lyrifissurae ia. Margins of centrodorsal ridge well-developed, converging anterior of setae dm, then diverging; posterior part of ridge rounded, extending posteriorly to point level with setae h 3. Ventral aspect: epimeral plates I trapezoid, twice as long as broad; epimera III and IV separate, rectangular. Enantiophysis V present; epimeral setation 2 - 1 - 2 - 2. Pedotectum II not extending laterally beyond circumgastric scissure. Perigenital carina more-or-less straight, ca. 38 µm long, extending posteriorly to point level with seta ag. Genital plates 44 µm long, same length as anal plates and almost three times distance between them. Cuticle ante- rior to genital plate granulate. Pedotectum I long, thin, sparingly granulate, extending anteriorly to point almost half way along length of lamellae in dorsal view. Lateral margins of pedotecta II extending to apices of bothridia setae. Gnathosoma the same as in Topalia caliginosa. Material Examined and Locality Data. Holotype and paratype: ANIC 3757, sifted litter and mossy logs, rainforest with Eucalyptus sp., State Forest, 2.2 km NE Corinna, Tasmania, 41 ° 38 ’ 43 ” S 145 ° 05 ’ 50 ” E., 45 m., coll. T. Weir & C. Lemann, 14. iii. 2008. Etymology. This species is named for its type locality; Corinna, Tasmania. Diagnosis. Topalia corinnensis sp. nov. can be distinguished from other species in the genus by the following combination of characters: (1) rostrum incised medially; (2) anteromedial edges of lamellae meeting; (3) setae in on squat tubercles; (4) anterior margin of notogaster wavy, indented medially; (5) epimera III and IV completely separated; (5) with enantiophysis V between epimera II and III. Remarks. Topalia corinnensis sp. nov. is morphologically most similar to T. royi sp. nov., with which it shares the wavy anterior notogastral margin with a median indentation and the lamellae with fine striae. Topalia corinnensis sp. nov. differs from T. royi sp. nov. in the separated epimera II and IV, the granular, non-polygonal cerotegument and the presence of enantiophysis V, a character state it shares with T. africana and T. problematica.	en	Colloff, Matthew J. (2019): The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera. Zootaxa 4647 (1): 290-321, DOI: 10.11646/zootaxa.4647.1.18
03A7BA3FC925FFD5FF53C0AFFD0435DF.taxon	description	(Fig. 5)	en	Colloff, Matthew J. (2019): The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera. Zootaxa 4647 (1): 290-321, DOI: 10.11646/zootaxa.4647.1.18
03A7BA3FC925FFD5FF53C0AFFD0435DF.taxon	materials_examined	Dimensions. Holotype length 347 μm, breadth 211 μm. Paratypes mean length (n = 8) 324 μm (range 299 – 340 μm); mean breadth 200 μm (range 183 – 211 μm). Ratio of prodorsum to total length: 0.32 (holotype). Adult. Integument: prodorsum and notogaster with network of large foveolae separated by angular ridges interspersed with minute granular tubercles. Cerotegument present in centrodorsal region. Underlying cuticle with fine striae (Fig. 5 c). Prodorsum: rostrum incised, with two lateral teeth and shorter median tooth. Lamellae meeting along anterior midline, not striate. Interlamellar region flat; setae in on squat tubercles posterior of posteromedian margins of lamellae. Bothridial seta with short, spherical club-shaped apices bearing blunt spines. Notogaster: ratio of length to breadth: 1.12. Anterior margin of notogaster smooth, convex, entire. Humeral projections (h) rounded, narrow, bearing seta c 2 at level of posterior incurve. Margins of centrodorsal ridge welldeveloped, slightly converging anterior of setae dm, then diverging; posterior part of ridge rounded, extending posteriorly to point posterior of level of setae h 3. Posterior notogastral setae on squat tubercles. Ventral aspect: epimeral plates I trapezoid, twice as long as broad; epimera III and IV separate, rectangular. Epimeral setation 2 - 1 - 2 - 2. Pedotectum I long, thin, extending anteriorly to point almost half way along length of lamellae. Lateral margins of pedotecta II extending only just further than margins of pedotecta I. Perigenital carina thin, curved, ca. 34 µm long, extending posteriorly to point anterior of seta ag. Genital plates 43 µm long, same length as anal plates and over twice length of distance between them. Sclerotised ring around genital plates with parallel striae. Ventral plate sparingly granulate. Material Examined and Locality Data. Holotype: ANIC 1025, leaf litter under Araucaria heterophylla, Red Road Track, Norfolk Island, coll. T. A. Weir, 14. xi. 1984. Eight paratypes: ANIC 3506, trough of flight intercept trap, Maurge Jowett’s [garden] (Red Road), Norfolk Island, 29 ° 0 ’ 37.73 ” S, 167 ° 56 ’ 43.49 ” E, ca. 250 m., coll. 1. ii. 1985. Etymology. This species is named in honour of my colleague Dr Michael Dunlop (CSIRO Land and Water) in recognition of his contribution to ecology and conservation in Australia. Diagnosis. Topalia dunlopi sp. nov. can be distinguished from other species in the genus by the following combination of characters: (1) the medially incised rostrum; (2) the parallel striae on the sclerotised ring around the genital plates; (3) the cerotegument consisting of a network of large foveolae separated by angular ridges; (4) the anteromedial edges of lamellae adjoining; (5) the anterior margin of the notogaster smooth, convex; (6) epimera III and IV separate; (7) the epimeral setation of 2 - 1 - 2 - 2. Remarks. Topalia dunlopi sp. nov. is morphologically most similar to T. corinnensis sp. nov., with which it shares the medially incised rostrum, the separate epimera III and IV and the medially-adjoining lamellae. Topalia dunlopi sp. nov. differs from T. corinnensis sp. nov. in the shape of the anterior margin of the notogaster, the pattern of cerotegument and the absence of enantiophysis V.	en	Colloff, Matthew J. (2019): The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera. Zootaxa 4647 (1): 290-321, DOI: 10.11646/zootaxa.4647.1.18
03A7BA3FC924FFD5FF53C2E3FBA93087.taxon	description	(Fig. 6)	en	Colloff, Matthew J. (2019): The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera. Zootaxa 4647 (1): 290-321, DOI: 10.11646/zootaxa.4647.1.18
03A7BA3FC924FFD5FF53C2E3FBA93087.taxon	materials_examined	Dimensions. Holotype length 344 μm, breadth 221 μm. Paratype length 350 μm, breadth 205 μm. Ratio of prodor- sum to total length: 0.36 (holotype). Adult. Integument: prodorsum and lamellae with minute granular tubercles in interlamellar region (Fig. 6 c); notogaster with sparse tubercles. Cuticle reticulate (Fig. 6 a). Prodorsum: rostrum incised, with two lateral teeth and shorter median tooth. Lamellae separated along midline, with narrow anterior translamella positioned ventrally, not striate. Interlamellar region flat; setae in on squat tubercles on posteromedian margins of lamellae. With pair of blunt condyles (c) medial to bothridia. Bothridial seta with short, spherical club-shaped apices bearing blunt spines. Notogaster: ratio of length to breadth: 1.02. Anterior margin of notogaster smooth, convex, entire; faint ridge connecting bases of setae c 2. Humeral projections (h) bilobed, with prominent anterior lobe posterolateral of bothridia and separated from posterior lobe by pronounced indentation. Margins of centrodorsal ridge well-developed, wavy, connecting with faint transverse ridge at level with setae lp. Cuticle of centrodorsal ridge punctate. Ventral aspect: epimeral plates I trapezoid, only slightly longer than broad. With two pairs of pronounced, sclerotised epimeral foveolae (ef); anterior pair between epimera I and II; posterior pair between epimera II and III. Epimera III and IV separate, rectangular. Epimeral setation 2 - 1 - 3 - 3. Pedotectum I long, broad, with incurved apices, extending anteriorly to point almost half way along length of lamellae. Pedotectum II not extending laterally beyond circumgastric scissure. Perigenital carina broad, short, slightly curved, ca. 28 µm long, extending posteriorly to point anterior of seta g 3. Genital plates 47 µm long, slightly longer than anal plates and over four times distance between them. Ventral plate and sclerotised ring around genital plates sparingly granulate. Material Examined and Locality Data. Holotype and paratype: ANIC 3507, trough of flight intercept trap, Maurge Jowett’s [garden], Red Road, Norfolk Island, 29 ° 0 ’ 37.73 ” S, 167 ° 56 ’ 43.49 ” E, ca. 250 m., coll. M. Christian & M. Sexton, 8. iii. 1985. Etymology. This species is named in memory of Dr Ekaterina “ Katya ” Sidorchuk in recognition of her considerable contribution to the systematics and taxonomy of the Oribatida. Diagnosis. Topalia katyae sp. nov. can be distinguished from other species in the genus by the following combination of characters: (1) the presence of a pair of interlamellar condyles; (2) the transverse notogastral ridge; (3) the narrow, wavy centrodorsal ridge, not expanded posteriorly; (3) the lamellae divided medially; (4) the presence of two pairs of epimeral foveolae; (5) the epimeral setal formula 2 - 1 - 3 - 3; (6) the reticulate notogastral cuticle. Remarks. Topalia katyae sp. nov. has two unusual characters; the ⊥ - shaped pattern of notogastral ridges, not found in other Topalia spp., and the pair of condyles on the interlamellar region, and is thus not particularly similar in morphology to other Topalia spp. However, T. katyae sp. nov. shares interlamellar condyles with T. africana Mahunka, 1985 from South Africa and T. clavata Hammer, 1966 from New Zealand and divided lamellae with T. caliginosa sp. nov., T. royi sp. nov. and T. velata Hammer, 1966 from New Zealand.	en	Colloff, Matthew J. (2019): The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera. Zootaxa 4647 (1): 290-321, DOI: 10.11646/zootaxa.4647.1.18
03A7BA3FC927FFD1FF53C55AFC163300.taxon	description	(Figs. 7, 8, 9 b)	en	Colloff, Matthew J. (2019): The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera. Zootaxa 4647 (1): 290-321, DOI: 10.11646/zootaxa.4647.1.18
03A7BA3FC927FFD1FF53C55AFC163300.taxon	materials_examined	Dimensions. holotype: length 340, breadth 195; paratype 1: length 363, breadth 212; paratype 2: length 355, breadth 208. Ratio of prodorsum to total length: 0.36 (holotype). Adult. Integument: notogaster with network of large, deep, angular foveolae interspersed with globular tubercles of varying size. Underlying cuticle smooth. Prodorsum: rostrum incised. Lamellae separated along midline and with pattern of fine, curved striae, with narrow anterior translamella positioned ventrally. Interlamellar region flat, smooth, bearing squat tubercles of setae in. Bothridial seta short, with attenuated club-shaped apices with blunt spines. Notogaster: ratio of length to breadth: 1.12. Anterior margin of notogaster transverse, wavy, concave medially. Rounded humeral projections (h) positioned anterolaterally on notogaster, bearing minute seta c 2. Margins of centrodorsal ridge well-developed, closely convergent anterior of setae dm, then diverging posteriorly. Posterior part of ridge broadly rounded, extending to point posterior of level of setae h 3. Notogastral setae on squat tubercles. Ventral aspect: epimeral plates I one-third longer than broad; sub-rectangular to trapezoid; epimera III and IV partly fused transversely and along mid-line; epimeral setation 2 - 1 - 2 - 3. Pedotectum I extending anteriorly about one-third of length of lamellae. Pedotectum II extending laterally as far as margin of humeral projections, with strong, dorsally-curved, pointed apex. Perigenital carina narrow, strongly curved, ca. 40 µm long, extending posteriorly to point level with seta ag. Genital plates 45 µm long, same length as anal plates and twice distance between them. Subcapitulum elongated (72 µm long) and narrow; chelicera thin, narrow, elongate, sub-pelopsiform; movable digit with reduced teeth (Fig. 8 b). Lateral aspect: apices of lamellar cusps slightly curved ventrally. Interlamellar region slightly domed, level with highest part of centrodorsal ridge. Anterior margin of rostrum between tutorium (tu) and rostral seta straight, diagonal. Tutorium with short free cusp. Anterior margin of pedotectum I rounded, convex. Centrodorsal ridge low and flat, extending posteriorly as far as level of lyrifissure ih; region just posterior slightly concave. Legs: Tarsus I with 15 setae, including short setiform famulus; solenidia ω 1 and ω 2 subequal in length (30 µm); tibia I with solenidion φ 1 very long (110 µm), flagelliform, arising free on tarsal cuticle and set back from articulation with tibia. Genu I without setae, bearing short solenidion σ only; femur I with one dorsal and one ventral seta (Fig. 8 a). Material Examined and Locality Data. Holotype and two paratypes: ANIC 460, leaf litter, rainforest, O’Reilly’s Rainforest Retreat, Lamington National Park, Queensland, 28 ° 13 ’ 58 ” S 153 ° 8 ’ 17 ” E, c. 920 m, coll. R. J. Kohout, 21. iii. 1973. Etymology. This species is named in honour and memory of Dr Alexander “ Sandy ” Bilton Roy in recognition of his nearly twenty years of volunteer work devoted to the curation and cataloguing of spiders, mites, ticks and termites in the Australian National Insect Collection. Diagnosis. Topalia royi sp. nov. can be distinguished from other species in the genus by the following combination of characters: (1) the rostrum incised medially; (2) the anteromedial edges of lamellae separated; (3) the partly fused epimera III and IV; (4) the medially indented anterior notogastral margin; (4) the laterally-positioned humeral projections; (5) the cerotegument consisting of a network of large, deep, angular foveolae; (6) the epimeral setation 2 - 1 - 2 - 3. Remarks. Topalia royi sp. nov. is morphologically most similar to Topalia caliginosa sp. nov., as detailed above. Topalia sp. 2 (Hunt et al., 1998) was collected from the same location as T. royi (leaf litter, O’Reilly’s Rainforest Retreat, Lamington National Park, Queensland (28 º 14 ’ S 153 º 08 ’ E) coll. R. Kitching) and I consider it to be T. royi sp. nov. They share the characteristic pattern of cerotegument, faint striae on the lamellar cusps, equal ratios of prodorsal length to total length, notogastral length to breadth, and the morphology of the bothridial setae (Fig. 9 b). When the figure of the dorsal aspect of T. royi sp. nov. is superimposed on the scanning electron micrograph of Topalia sp. 2, it fits almost exactly the outline of the lamellae and notogaster.	en	Colloff, Matthew J. (2019): The oribatid mite genus Topalia in Australia (Oribatida: Nosybeidae) and the taxonomic status of related families and genera. Zootaxa 4647 (1): 290-321, DOI: 10.11646/zootaxa.4647.1.18
