taxonID	type	description	language	source
03A48795FFB8FFC4FF02FBBA5C80FE9F.taxon	diagnosis	Diagnosis. Eyebridge of three facet rows, interocular suture interrupted medially or reduced. Antennae with 16 articles, scape with sensory scales (" androconia "), flagellomeres nodiform (except f 13 in N. saltenia (Omad, Mangudo & Gleiser )), multiple filiform ascoids present as ring surrounding flagellomere nodes. Palp of four segments, 1 st palp segment reduced both in length and sclerotization, 4 th palp segment with reduced sclerotization. Wing broadly ovate, Rs with R 2 + 3 very short (radial fork in basal third of wing), sometimes pectinate (R 2 + 3 arising from R 4), R 5 terminating in wing apex, radial fork basal of medial fork. Sensory organ present near apex of S c (except in N. saltenia (Omad, Mangudo & Gleiser )). Aedeagus, parameres and gonopods asymmetrical. Aedeagus with two apical blade-shaped phallomeres of different size and shape. Gonocoxites connected via hypandrium, gonostyles jointed ventrally or fused to gonocoxites. Hypopod generally with both lateral epandrial branch and mesal cercal branch, epandrial branch carrying a single tenaculum (reduced or absent in N. saltenia), mesal branch with or without accessory tenacula.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFB8FFC4FF02FBBA5C80FE9F.taxon	discussion	Remarks. Omad et al. (2015) described the genus Chuspilepia based on a single species from the Salta province in Argentina. In their discussion, they contrasted the species from Neurosystasis in lacking an interocular suture, having the « cercopod » unbranched, having R 2 + 3 connected to R 4 and by the gonocoxites dimorphic. However, apart from the unbranched hypopod, all of these characters are indeed found in Neurosystasis; meaning that only one character separates Chuspilepia from Neurosystasis. We interpret the pilose mesal area and the bare lateral patch in the hypopod of Chuspilepia saltenia Omad, Mangudo & Gleiser to be a rudiment or evolutionary predecessor of the bifurcation present in the Caribbean species, and place Chuspilepia tentatively in synonymy with Neurosystasis. The diagnosis given by Quate & Brown (2004) is erroneous in several aspects, including the statements that the eyebridges can extend to the midline and that R 2 + 3 is not attached to R 4. Based on original descriptions of the type and other species, the eyebridges of all Neurosystasis are well separated and for three species, including the type, Rs is pectinate.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBBFFC4FF02FBD25C1CFA38.taxon	distribution	Distribution. Cuba. A record from USA: Florida by Quate & Brown (2004) was based on female specimens only and is likely to refer to N. bromeliphila Wagner & Hribar, 2010 (Wagner & Hribar 2010).	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBBFFC4FF02FBD25C1CFA38.taxon	discussion	Remarks. A full synonymy of N. amplipenna is given by Duckhouse (1974), who also provides redescriptions and figures.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBBFFC4FF02F99A5ED0F899.taxon	distribution	Distribution. USA (Florida).	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBBFFC4FF02F99A5ED0F899.taxon	discussion	Remarks. For descriptions and figures, see Wagner & Hribar (2010).	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBBFFC5FF02F8FD5CC0F8FF.taxon	materials_examined	Type material. Holotype male, Cuba, Sierra de Escambray, Profiles, 26 April 1992 leg. Stark. Coll. Museum für Naturkunde, Berlin (ZMHB).	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBBFFC5FF02F8FD5CC0F8FF.taxon	diagnosis	Diagnosis. Neurosystasis mira sp. nov. can be distinguished from other Neurosystasis species on the following combination of characters: wings infuscate with large translucent spots along margin between wing vein apices; M 3 not connected to M 1 + 2; left gonocoxite with an elongate curved lateral lobe with lateral margin smooth, right gonocoxite with a straight and short mesal lobe and an elongate and curved lateral lobe; hypopod with a blunt and setose cercal branch about twice as long as the epandrial branch; with single tenaculum apically on epandrial branch and a tubercle carrying four setiform accessory tenacula on the cercal branch near the base of the epandrial branch; cercal branch apically without tubercles.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBBFFC5FF02F8FD5CC0F8FF.taxon	description	Description. Adult male (n = 1). Head (fig. 1 A) wider than long, with large eyes extending to front and lateral; eyebridge of 3 facet rows, distance between eyes two facet diameters; interocular suture incomplete with two broad crescent-shaped extensions of the eyes; occiput narrow with median v-shaped incision; Frons with pair of oval patches of setae; antennae (fig. 1 B) with inner distal side of scape with bases of approximately 15 setae (androconia); pedicel spherical, shorter than scape; flagellomeres nodiform spindle-shaped, ascoids not discernable, probably several thin digitate ascoids along upper margin of antennomere setation; relative length of antennomeres: 22 - 17 - 20 - 23 - 25 - 24 - 25 - 24 - 21 - 23 - 21 - 20 - 21 - terminal segments missing; absolute length: 0.15 - 0.11 - 0.13 - 0.15 - 0.17 - 0.16 - 0.17 - 0.16 - 0.14 - 0.15 - 0.14 - 0.13 - 0.14 - mm; palp of four palpomeres (fig. 1 C), sclerotization of basal and distal segments weaker than that of segments 2 and 3; relative length of palpomeres: 13 - 32 - 33 - 38; absolute length: 0.09 - 0.21 - 0.22 - 0.25 mm; thorax: legs with apical tarsomeres modified: in lateral view (fig. 1 F) with dorsal prolongation; in dorsal (fig. 1 G) view prolongation consisting of two lateral tips, each with distal seta, between tips a serrate ridge; claws sickle-shaped; pulvilli present as large tufts of setae (fig. 1 H); wing (fig. 1 D) infuscate with elongate oval translucent areas between most vein tips; Sc short and sinuous, ends very basal in R 1; near tip of Sc is a parallel ‘ cross-vein’ with a hardly visible spherical structure with circular apertures (fig. 1 E); radial fork very basal, close to the incomplete fork R 2 + 3 / R 4 and basal of medial fork; R 5 straight, distinctly wider than other veins, ends in wing tip; four medial veins, faint cross-vein M 1 + 2 - M 3 present at base of both; CuA basally swollen, CuP short slightly bent; wing length: 3.05 mm; greatest wing width 1.61 mm; Abdomen with 8 segments and inverted genitalia (fig. 2 C); hypandrium unsclerotized, probably elastic, hardly visible; gonopods and aedeagus asymmetric; gonocoxites dark brown; left gonocoxite (fig. 2 A) thin, elongate bilobed, strongly bent; posterior lobe longer than anterior, its straight distal part behind bend about 1.5 times longer than basal part; at basal part the short anterior lobe arises whose distal blunt end is curved cranial and ventral to approximate the corresponding but straight lobe of the right gonocoxite; left gonostyle arising near base of gonocoxite, almost translucent, straight, blade-shaped, tip at same level as tip of gonocoxite; right gonocoxite (fig. 2 B) shorter than left, bilobed with short straight arm pointing towards short dorsally bent lobe of the left gonocoxite; the longer lateral lobe evenly bent with tip pointing right lateral; right gonostyle as well translucent, slightly curved, tip at about same level as left gonocoxite and gonostyle, but much more distal than tip of right gonocoxite; gonocoxites basally connected by posterior lobe of left gonocoxite that turns towards inner lobe of right gonocoxite, and is probably connected as well to the basiphallus; aedeagus (fig. 2 D) with large calabash-shaped basiphallus; on left side a short strong lobe pointing to the connection of left and right gonostyles; Distally with joints and two long blade-shaped phallomeres, the left one broader, twice as long as the right; morphologically dorsal of the calabash-shaped basiphallus lies a trapezoid transverse sclerite (fig. 2 G), forming a broad bridge between gonocoxites with medial basal prolongation with a keel; the lateral ends connected to left and right gonocoxite, the thin elongate keel probably connects with the basiphallus; epandrium (figs 2 E, 2 F) subrectangular, slightly narrowing distad, with a pair of narrow transversely oblong apertures in the basal third; lateral margins rolled up; subepandrial sclerite basally not clearly discernable; distal connection with hypopods bilobed; hypoproct with a microsetose ‘ nose’, epiproct small, setose; hypopods shorter than epandrium, Y-shaped; stronger mesal cercal lobe blunt, epandrial lobe narrower with single apical tenaculum; cercal lobe with a globular tubercle with 4 elongate straight accessory tenacula near origin of epandrial lobe.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBBFFC5FF02F8FD5CC0F8FF.taxon	distribution	Distribution. Only recorded from the type locality on Cuba.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBBFFC5FF02F8FD5CC0F8FF.taxon	biology_ecology	Biology. Unknown.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBBFFC5FF02F8FD5CC0F8FF.taxon	etymology	Etymology. From latin mira, exceptional, curious. The epithet is to be treated as an adjective and thus conform to the gender of the genus.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBFFFC0FF02FF065E90FE26.taxon	distribution	Distribution. Argentina	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBFFFC0FF02FF065E90FE26.taxon	discussion	Remarks. For descriptions and figures, see Omad et al. (2015).	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBFFFCEFF02FE695CF9FE2C.taxon	materials_examined	Type material. Holotype male, Cuba, Sierra de Escambray, Pretiles, 26 April 1992 leg. A. Stark, coll. ZMBH	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBFFFCEFF02FE695CF9FE2C.taxon	diagnosis	Diagnosis. Neurosystasis starki sp. nov. can be distinguished from other Neurosystasis species on the following combination of characters: wing infuscate with large translucent spots along margin between wing apices; M 3 not connected to M 4; left gonocoxite with an elongate curved lateral lobe with lateral margin serrate and a shorter, sinous median lobe, right gonocoxite with bilobed lateral lobe with lateral margin of lateral branch serrate and a slightly curved median lobe; hypopod with an incurved cercal branch about twice as long as the epandrial branch; with single tenaculum apically on epandrial branch and a tubercle carrying three setiform accessory tenacula on the cercal branch near the base of the epandrial branch; cercal branch apically with two tooth-like tubercles.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBFFFCEFF02FE695CF9FE2C.taxon	description	Description. Adult male (n = 1). Head wider than long, with large eyes extending to front and lateral; eyebridge of 3 facet rows, distance between eyes one facet diameter; Interocular suture incomplete at middle with two broad crescent-shaped extensions of the eyes; occiput narrow with median v-shaped incision; frons with pair of oval patches of setae; antennae with scape on inner distal side with bases of more than 10 strong setae (androconia); pedicel spherical, shorter than scape; flagellomeres (fig. 3 A) nodiform spindle-shaped, with many digitate ascoids in one row along upper rims of ‘ bulb’ and a few irregularly positioned additional ascoids; relative length of antennomeres: 17 - 12 - 15 - 18 - 18 - 17 - 16 - 16 - 15 - 16 - terminal segments missing; absolute length: 0.11 - 0.08 - 0.10 - 0.12 - 0.12 - 0.11 - 0.11 - 0.11 - 0.10 - 0.11 mm; palp of four palpomeres, sclerotization of all segments equal; relative length of palpomeres: 9 - 25 - 22 - 25; absolute length: 0.06 - 0.16 - 0.14 - 0.16 mm; thorax: Mid coxa with short frontal protuberance with apical area of microsetae (fig. 3 C); two or three rows of seta bases along coxae, regularly arranged on front and hind coxa, irregular on middle coxa; tarsi not modified; wing (fig. 3 B) infuscate with elongate oval translucent areas between most vein tips along wing margin; areas between costa and subcosta, and subcosta and radius darker brown; Sc short and sinuous, ending just short of R 1; near tip of Sc is a hardly visible spherical structure with circular apertures; radial fork very basal, close to the incomplete fork R 2 + 3 / R 4 and basal of medial fork; R 5 basally with small brown spot, straight, ends in wing tip; four medial veins; CuA basally swollen, darker than other veins; CuP short, straight; wing length: 2.20 mm; greatest wing width 1.10 mm; abdomen with 8 pregenital segments and inverted genitalia; hypandrium (figs. 3 D — E) unsclerotized, hardly recognizable; epandrium and hypopods symmetric (fig. 3 G), gonopods and aedeagus asymmetric (fig. 3 D); gonocoxites brown; left gonocoxite bilobed, slightly bent, ventral lobe longer than dorsal, its straight distal part behind bend about 2 times longer than basal part; at basal part a short dorsal lobe arises whose distal blunt end is curved medially and ventral to approximate the corresponding but straight lobe of the right gonocoxite; left gonostyle arising near base of inner gonocoxite lobe, almost translucent, longer than gonocoxite, with 4 or 5 seta bases along distal edge; right gonocoxite shorter than left, bilobed with straight inner ventral lobe longer than lateral dorsal; this lobe serrate along lateral edge; right gonostyle similar to left, tip sharper as well with four to five seta bases; gonocoxites basally connected by dorsal lobe of left gonocoxite that turns towards inner lobe of right gonocoxite and probably connects to the basiphallus as well; aedeagus with large calabash-shaped basiphallus, distally with two thin bladeshaped phallomeres of unequal length (fig. 3 H, 3 F); morphologically dorsal of the basiphallus lies a rounded trapezoid transverse sclerite, forming a broad bridge between gonocoxites with medial basal prolongation with a keel (fig. 3 E); the lateral ends connect to gonocoxites, thin keel probably connected with basiphallus; paramere (fig. 3 F) bilobed, distal ends meet tips of phallomeres; basally with an additional short digitate lobe; epandrium (fig. 3 G) subrectangular, with a pair of narrow transversely oblong apertures in the basal third; subepandrial sclerite basally not clearly discernable; distal connection with hypopods bilobed; hypoproct with a microsetose triangular projection, epiproct small, setose; hypopods shorter than epandrium, bilobed; mesal lobe blunt, with a globular tubercle with 3 elongate straight accessory tenacula near origin of lateral lobe, lateral lobe thin with single apical tenaculum.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBFFFCEFF02FE695CF9FE2C.taxon	distribution	Distribution. Recorded only from the type locality in Cuba.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBFFFCEFF02FE695CF9FE2C.taxon	biology_ecology	Biology. Unknown.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFBFFFCEFF02FE695CF9FE2C.taxon	etymology	Etymology. Dedicated to Dr. Andreas Stark, who collected the specimen and donated his fine collection of Psychodidae from Cuba to RW.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFB1FFCEFF02FE6E5E67FCAD.taxon	distribution	Distribution. Jamaica.	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
03A48795FFB1FFCEFF02FE6E5E67FCAD.taxon	discussion	Remarks. For descriptions and figures, see Satchell (1955).	en	Kvifte, Gunnar Mikalsen, Wagner, Rüdiger (2017): Review of Neurosystasis Satchell, with two new species from Cuba and a discussion of cerci and surstyli in Psychodinae (Diptera: Psychodidae). Zootaxa 4306 (1): 81-90, DOI: 10.11646/zootaxa.4306.1.4
