taxonID	type	description	language	source
03A487A3FFBBFF922D99FD56B39CFC5E.taxon	type_taxon	Type species. Syllis spongicola Grube, 1855	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBBFF922D99FD56B39CFC5E.taxon	diagnosis	Diagnosis. Body cylindrical. Prostomium with four eyes. Three antennae. Palps robust, fused at bases. Two pairs of tentacular cirri. Antennae, tentacular cirri and dorsal cirri distinctly moniliform, the latter sometimes with a single small article on posterior parapodia (giving a smooth appearance). Ventral cirri digitiform. Pharynx with one anterior tooth surrounded by crown of 10 – 12 soft papillae and ring of cilia, occasionally a trepan of small teeth. Chaetae all simple, with bi- or unidentate tips, 1 – 12 per parapodia. Capillary chaetae absent. Reproduction by sexual stolons (schizogamic scissiparity).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBAFF952D99FFC4B25DFA8B.taxon	description	Figs 1 A – G, 2 A – D, 3 A – D	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBAFF952D99FFC4B25DFA8B.taxon	materials_examined	Examined material. AUSTRALIA: NEW SOUTH WALES. 18 specimens AM W 29374 (plus 3 specimens mounted for SEM), 100 m south of Split Solitary Island, 30 º 15 ' S 153 º 10 ' 30 '' E, 16 m, sponge attached to rocky bottom, coll. R. T. Springthorpe, 23 June 1992. 3 specimens, AM W 26340, 100 m north west of Split Solitary Island, 30 º 14 ' 0 '' S 153 º 10 ' 48 '' E, 17 m, mixed red algae, coll. S. J. Keable, 7 March 1992.> 100 specimens AM W 26384 (plus 2 specimens mounted for SEM), Gap Bluff, 33 ° 50 ' 43 '' S 151 ° 17 ' 09 '' E, 20 m, in unidentified Ianthellidae sponge with spikey surface, dark reddish black in colour, coll. K. B. Attwood and G. San Martín, 14 June 2000. Additional material: PAPUA NEW GUINEA: BISMARK SEA, LAING ISLAND. 04 ° 09 ' S 114 ° 52 ' E, inside Anomoiantella lamella, 15 m, coll. R. Danovaro & S. Fraschetti, 12 August 1986. Personal collection, D. Martin. Comparative material: ISLAND OF GUAM, MICRONESIA. 20 Paratypes of H. basticola MNCN 16.01 / 8439.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBAFF952D99FFC4B25DFA8B.taxon	description	Description. Body fragile, translucent, small, 2.5 – 4.5 mm long for 15 – 33 segments, 0.15 – 0.4 mm wide excluding parapodia, widest at proventricle level (Figs 1 A, 2 A). Preserved specimens pale yellow to brown purple. Small dorsal granules throughout body. Dorsal pigment pattern absent. Prostomium subpentagonal, wider than long, with two pairs of small red eyes in trapezoidal arrangement. Median antennae inserted on middle of prostomium, between eyes (8 – 28 articles); lateral antennae on anterior margin (6 – 14 articles). Palps long, broadly triangular, fused at their bases but separated all along their length; sensory organs as rows of cilia on ventral upper side. Nuchal organs not seen. Pharynx orange, extending through about four segments; with a large anterior tooth, crown of 10 – 12 soft papillae and inner ring of cilia (Fig. 2 B); upper side of papillae with groups of cilia making up the sensory organs. Proventricle cylindrical, dark brown, extending through 2 – 4 segments, with 30 – 35 muscle cell rows (Fig. 1 A). Peristomium well defined, shorter than subsequent segments. Dorsal tentacular cirri longer than ventral ones (10 – 18 and 2 – 8 articles, respectively). Dorsal cirri slender, similar to antennae and tentacular cirri. Anterior cirri long, alternating in length as follows: first cirri long (10 – 28 articles); second short (3 – 14 articles), third and fourth progressively longer (4 – 14 and 5 – 19 articles, respectively) and fifth small (2 – 10 articles). After proventricle, longest cirri does not exceed half of body width, gradually decreasing in length toward posterior end (2 – 6 articles) (Fig. 1 B). Ventral cirri digitiform, anterior ones slender, longer than parapodial lobes; median and posterior ones gradually decreasing in length (Figs 1 C – D). Chaetae all bidentate, 1 – 3 per parapodia, stout, broad, all similar in shape but anterior ones slightly smaller; LMF similar in length to SW; MJP short, curved; US of MF without denticles; apical teeth similar in length (Figs 1 E – F, 2 C – D). All parapodia with one stout acicula, with slightly upwards-directed curved tip (Fig. 1 G). Reproduction. The reproductive bodies of Haplosyllis basticola are cephalic stolons. Both male and female stolons have 10 – 12 chaetigers, a well defined head with two small antennae and two pairs of reddish eyes, and long swimming chaetae (Fig. 3 A – D).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBAFF952D99FFC4B25DFA8B.taxon	biology_ecology	Ecology. Haplosyllis basticola has been found only on sponges of family Ianthellidae, such as Ianthella basta (Pallas, 1776) from Micronesia (Sardá et al. 2002), Anomoianthella lamella Pulitzer-Finali & Pronzato, 1999 from Papua-New Guinea (Magnino et al. 1999), and an unidentified species from Australia. Nevertheless, it is not possible to confirm if H. basticola is a specific endobiont of iantellid sponges due to the absence of host sponge identifications for most of the Australian material cited here.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBAFF952D99FFC4B25DFA8B.taxon	discussion	Remarks. The Australian specimens differ from the type population (Micronesia) in size, reaching up to 33 segments and up to 25 segments, respectively. Similar differences occur for the cirri length, especially in the anterior ones, which are smaller in Micronesian than in Australian specimens. Additionally, the specimens from Micronesia, Papua-New Guinea, and some of the Australian populations were violet or brown purple (even preserved), mimicking the colour of the respective host sponges. The other Australian populations had creamy-yellow bodies. Despite the differences in size, the stolons, aciculae and chaetae were identical in all populations, the latter being the most diagnostic feature for the identification of H. basticola.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBAFF952D99FFC4B25DFA8B.taxon	distribution	Distribution. AUSTRALIA (New South Wales); MICRONESIA (Island of Guam); PAPUA-NEW GUINEA (Laing Island, Bismark Sea).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBCFF992D99FADBB221FDF2.taxon	description	Figs 4 A – G, 5 A – F	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBCFF992D99FADBB221FDF2.taxon	materials_examined	Examined material. AUSTRALIA: NEW SOUTH WALES. 250 specimens AM W 36621 (plus 4 specimens mounted for SEM), Jervis Bay, half way along west side of Bowen Island, 35 º 06 ' 55 '' S 150 º 45 ' 55 '' E, grey sponge with orange flesh, large oscular chamber, 8 m, coll. G. Wilson and P. Serov, 7 Dec 1993. Comparative material: Japan, Boso Peninsula. Holotype (MNCN 16.01 / 10135); 2 Paratypes (MNCN 16.01 / 10136).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBCFF992D99FADBB221FDF2.taxon	description	Description. Body fragile, translucent, small, 2 – 3 mm long, for 20 – 26 segments (14 – 18 in juveniles or smaller specimens), 0.4 – 0.8 mm wide excluding parapodia, (Figs 4 A, E; 5 A), widest at proventricle level; pale yellow when preserved, with orange marks in dorsal cirri and anterior or posterior ends (probably from host sponge, not seen in all specimens); granulose dorsally. Prostomium subpentagonal to oval similar in width and length (Fig. 4 A, E), with two pairs of small red eyes in trapezoidal arrangement. Median antennae inserted on middle of prostomium, between eyes (9 – 21 articles); lateral antennae on anterior margin (6 – 13 articles). Palps long, broadly triangular, fused at their bases but separated all along their length, with sensory organs as rows of cilia on ventral upper side (Fig. 5 C). Pharynx orange, 0.18 – 0.35 mm long, extending through 3 – 4 segments; with large anterior tooth surrounded by crown of 10 soft papillae and inner ring of cilia (Fig. 5 B); sometimes with trepan of about 10 small triangular teeth; upper ventral side of papillae with groups of cilia making up the sensory organs. Proventricle cylindrical, dark-brown, 0.25 – 0.32 mm long, 0.2 – 0.6 mm wide, extending through 2 – 4 segments, with 22 – 26 muscle cell rows (Fig. 4 A, E). Peristomium well defined, shorter than subsequent segments. Dorsal tentacular cirri (10 – 13 articles), twice as long as ventral ones. First three dorsal cirri always slender, similar to antennae and tentacular cirri; subsequent ones broad (Fig. 4 A) or slender (Fig. 4 E). Anterior dorsal cirri long, with a well-defined length pattern: first long (exceed body width, with 12 – 20 articles); second short (4 – 9 articles), third and fourth progressively longer (5 – 12 and 7 – 13 articles respectively); fifth short (3 – 12 articles), six long (7 – 12). Median cirri long, all equal in length (6 – 14 articles), longest cirri exceed body width, gradually decreasing in length toward posterior end (1 – 5 articles) (Fig. 4 B). Ventral cirri digitiform, anterior ones similar or shorter than parapodial lobes (Fig. 4 C, 5 D), median cirri and posterior ones small (Fig. 4 D). Chaetae all bidentate, 2 – 4 per parapodia, equal throughout, anterior ones slightly smaller; LMF similar in length to SW; MJP short, curved; denticles of US of MF absent or very short; apical teeth equal in length, long, well separated (Figs 4 F, 5 E – F). All parapodia with one acicula, with curved, upwards-directed tip (Fig. 4 G). Reproduction. The reproductive bodies of Haplosyllis crassicirrata are cephalic stolons. About 15 females with 26 – 31 segments have the last 12 – 14 segments modified into reproductive bodies, with cirri smooth or with 2 – 3 articles. Cirri from non-reproductive segments were slender, some with broad articles at their tips.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBCFF992D99FADBB221FDF2.taxon	distribution	Distribution. AUSTRALIA (New South Wales); JAPAN (Boso Peninsula).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFBCFF992D99FADBB221FDF2.taxon	discussion	Remarks. The studied specimens showed different cirri shapes, from all broad to almost all slender (but still a few broad). As the types were juveniles having most cirri broad and the studied specimens have almost all cirri slender, we suggest that the differences in the presence of broad cirri are linked to the ontogeny. In turn, the chaetae and aciculae were identical in all studied specimens, this being a more robust character for the identification of H. crassicirrata. The presence of the ciliated sensory organs on palps and pharyngeal papillae is here reported for the first time in Haplosyllis crassicirrata.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFB0FF9D2D99FDF1B263FCCE.taxon	description	Figs 6 A – I, 7 A – G	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFB0FF9D2D99FDF1B263FCCE.taxon	materials_examined	Examined material. AUSTRALIA. WESTERN AUSTRALIA. 33 specimens AM W 36549, Kimberleys, south west corner of Lucas Island, 15 º 13 ' S 124 º 31 ' E, 2 – 30 m, coll. P. A. Hutchings, 24 July 1988. 90 specimens AM W 36556, Reef south of Lucas Island, Brunswick Bay, Kimberley region, 15 ° 16 ' S 124 ° 29 ' E, 2 m, coll. P. A. Hutchings, 24 July 1988. 60 specimens, AM W 36552, inshore reef off Ned's Camp, Cape Range National Park, 21 º 59 ' S 113 º 59 ' E, 1 m, in frilly Caulerpa, coll. J. K. Lowry, 2 January 1984. 8 specimens AM W 36553, Bush Bay, 30 km south of Carnarvon, 25 º 10 ' S 113 º 39 ' E, 0.5 m, in extensive shallow sand flats, lumps of algae, coll. H. E. Stoddart, 6 January 1984. 53 specimens AM W 36542, Goss Passage, Beacon Island, 28 º 25 ' 30 '' 0 S 113 º 47 ' E, 8 m, dead plates of Acropora, covered in coralline algae, 22 May 1994, coll. P. A. Hutchings. 20 specimens AM W 36559, Goss Passage, north end of Long Island, 28 º 28 ' 18 '' S 113 º 46 ' 18 '' E, 8 m, dead coral substrate, coralline algae, boring bivalves, coll. Bryce, 22 May 1994. 1 specimen AM W 36547, southeast end of Long Island, 28 º 28 ' 48 '' S 113 º 46 ' 30 '' E, 8 m, dead coral covered in coralline algae, coll. P. A. Hutchings, 22 May 1994. 5 specimens AM W 36551, Goss Passage, Beacon Island, 28 º 25 ' 30 '' S 113 º 47 ' E, 10 m, branching dead coral covered in coralline algae, coll. P. A. Hutchings, 18 May 1994. 24 specimens AM W 36555, north end of Long Island, Goss Passage, 28 º 27 ' 54 S 113 º 46 ' 18 '' E, 6 m, dead coral covered in coralline and brown algae, coll. Bryce, 22 May 1994. 90 specimens AM W 36554, Beacon Island, 28 ° 28 ' 48 '' S 113 ° 46 ' 18 '' E, 4 – 5 m, dead coral substrate, covered in coralline algae, coll. P. A. Hutchings, 25 May 1994. 8 specimens AM W 36513, Beacon Island, Goss Passage, 28 ° 25 ' 30 '' S 113 ° 47 ' E, 8 m, dead plates of Acropora, coll. P. A. Hutchings, 19 May 1994. 70 specimens AM W 36548, Goss Passage, Beacon Island, 28 º 25 ' 30 '' S 113 º 47 ' E, 20 m, dead plates of Acropora, covered in coralline algae, coll P. A. Hutchings, 20 May 1994. 28 specimens AM W 36557, Beacon Island, of jetty adjacent to Fisheries Hut, 28 º 25 ' 30 '' S 113 º 47 ' E, 12 m, dead coral, plate-like Acropora montipora, 23 May 1994. 2 specimens AM W 36558, Warnbro Sound, west of Penguin Island, 32 º 20 ' S 115 º 43 ' E, 5 m, coll. P. A. Hutchings, 21 March 1993. 4 specimens AM W 36541, Exmouth Gulf, beach at north end of Bundegi Reef, 21 º 49 ' S 114 º 11 ' E, 1 – 2 m, rocky rubble, brown alga with epiphytic growth, sticky sediment, coll. H. E. Stoddart, 4 January 1984. NEW SOUTH WALES. 100 specimens AM W 36491 (plus 3 specimens mounted for SEM), Look-At-Me-Now Headland, 30 ° 18 ' 50 " S 153 ° 08 ' 55 " E, 4 – 6 m, Ecklonia radiata holdfast, coll. S. Smith, 27 April 1994. 4 specimens AM W 36545, 150 m east of Burrill Rocks, 35 ° 23 ' 25 '' S 150 ° 28 ' 11 '' E, dead bryozoan encrusted with algae, 17 m, coll. K. Attwood, 1 May 1997. 1 specimen AM W 36543, Murrays Beach, Jervis Bay, 35 º 7 ' 30 '' S 150 º 45 ' 30 '' E, 9 m, coll. P. A. Hutchings, 23 January 1973. 22 specimens AM W 36628, Coff’s Harbour Jetty, 30 º 18 ' 24 '' S 153 º 08 ' 30 '' E, 4 m, finger sponge on jetty pilings, coll. R. T. Sprinthorpe, 9 March 1992. 120 specimens AM W 26333, North Ledge, Cook Island, 28 º 11 ' 26 '' S 153 º 34 ' 40 '' E, 10 m, orange sponge, coll. A. R. Parker, 8 June 1993. QUEENSLAND. 4 specimens AM W 36546, Outer Yonge Reef, Great Barrier Reef, 14 ° 36 ' S 145 ° 38 ' E, 9 m, coral rubble, coll. P. A. Hutchings, 21 January 1977. 6 specimens AM W 36550, Outer Yonge Reef, Great Barrier Reef, 14 ° 36 ' S 145 ° 38 ' E, 30 m, coral rubble, coll. P. A. Hutchings, 24 January 1977. NORTHERN TERRITORY. 1 juvenile AM W 36516, Darwin Harbour, East Point, 12 ° 24 ' 36 '' S 130 ° 39 ' 57 " E, 7 – 10 m, dead coral rubble and algal washings, coll. P. A. Hutchings, 17 July 1993. 4 specimens AM W 36544, Lee Point, Darwin, 12 º 20 ' 0 '' S 130 º 53 ' 48 '' E, 3 m, dead coral rubble washings, coll. P. A. Hutchings, 11 July 1993. Haplosyllis cf. djiboutiensis. WESTERN AUSTRALIA: 16 specimens AM W 36512, Beacon Island, Goss passage, 28 ° 25 ' 30 '' S 113 ° 47 ' E, 23 m, dead plates of Acropora, covered in coralline algae, 19 May 1994, coll. P. A. Hutchings. 5 specimens AM – W 36514, Goss Passage, Beacon Island, 28 º 25 ' 30 '' S, 113 º 47 ' E, 33 m, dead coral, embedded in fine sediment, coll. P. A. Hutchings, 23 May 1994. TASMAN SEA: 75 specimens AM W 36515, Elizabeth Reef, reef flat near " Yoshin Maru Iwaki " wreck, 29 ° 55 ' 48 " S 159 ° 01 ' 18 " E, 0 m, from small heads of Acropora valida and Pocillopora damicornis, 14 Dec. 1987, coll. J. K. Lowry & R. Springthorpe.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFB0FF9D2D99FDF1B263FCCE.taxon	description	Description. Body robust, from small to long sized 3 – 17 mm, for 30 – 92 segments; slightly widest at proventricle level, gradually tapering towards posterior end (Fig. 7 A); pale white to yellowish when preserved. Round, small dorsal granules, abundant laterally at posterior end, almost absent in anterior end. Dorsal pigment pattern absent. Prostomium subpentagonal, wider than long, with two pairs of small red eyes in trapezoidal arrangement (Fig. 6 A). Median antennae inserted on middle of prostomium, between eyes (11 – 30 articles); lateral antennae on anterior margin (7 – 16 articles). Palps longer than prostomium, broadly triangular, fused at their bases but separated all along their length; ventral side of palps with sensory organs as rows of cilia (Fig. 7 C – D); nuchal organs as lateral groups of cilia between prostomium and peristomium. Pharynx orange, extending through 4 – 6 segments; with long, triangular anterior tooth, crown of 10 – 12 soft papillae and inner ring of cilia (Fig. 7 B); sometimes with trepan of very small triangular teeth, difficult to see as often covered by cilia. Papillae with groups of cilia on upper-ventral side making up the sensory organs. Proventricle long, cylindrical, dark-brown, as broad as pharynx, extending through 3 – 9 segments, with 30 – 50 muscle cell rows. Peristomium well defined, shorter than subsequent segments. Dorsal tentacular cirri longer than ventral ones, with 13 – 24 articles. Antennae, tentacular cirri and anterior dorsal cirri similar. Anterior dorsal cirri relatively long, with a well-defined length pattern: first cirri exceed body width (11 – 30 articles), second short (5 – 16 articles), third and fourth gradually longer, almost as body width (7 – 20 and 10 – 29 articles respectively), fifth short (4 – 16 articles), sixth intermediate (6 – 24 articles). Subsequent cirri alternating long (7 – 16 articles) and short (3 – 11 articles), gradually decreasing in length toward posterior end (1 – 9 articles) (Fig. 6 B); median cirri, similar or shorter than body width. Ventral cirri digitiform, anterior ones similar in length than parapodial lobes (Fig. 6 C); median and posterior ones shorter than parapodial lobes (Fig. 6 D). Chaetae all bidentate, anterior ones with small MF and short denticles on US (Figs 6 E, 7 E). Median and posterior parapodia with 2 – 3 chaetae of two different sizes; long ones (Figs 6 G, 7 F) with LMF similar in length to SW; MJP straight and relatively long; US of MF with few, short denticles; apical teeth equal in length, often distal one slightly smaller than proximal. Small chaetae (Figs 6 H, 7 G) with MJP short and curved, wide angle between apical teeth. Two aciculae in anterior and median parapodia, one straight, other with curved tip (Fig. 6 F); one in posterior parapodia with upwards-directed curved tip (Fig. 6 I). Pygidium with two long cirri (8 – 10 articles) (Fig. 6 B). Reproduction. Haplosyllis djiboutiensis reproduces by acephalous stolons. Two specimens were in a reproductive stage, with the last 10 segments having pairs of black parapodial ocular spots and fascicles of capillary (i. e. swimming) chaetae and 4 – 5 chaetae similar to anterior ones (AM W 36559).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFB0FF9D2D99FDF1B263FCCE.taxon	discussion	Remarks. Haplosyllis djiboutiensis can be easily identified by the alternation of cirri on the median body, the abundant posterior dorsal granules, and the differences between long and short chaetae of median and posterior parapodia. Haplosyllis djiboutiensis is very abundant and common in Australian waters allowing us to study its interpopulation variability. The specimens varied from small and medium-sized worms (3 – 10 mm long), to long specimens (up to 17 mm long), with variations in some chaetal measures. For example, in long-sized specimens, the median and posterior big chaetae have the distal tooth shorter than the proximal one, and the length of MJP is usually long. These features are very similar to those of the type specimen from Djubal (Red Sea) (see Lattig & Martin, 2009, fig. 12 H), while in small to medium-sized worms the MJP is shorter and the apical teeth are equal in length (Fig. 6 G). The descriptions of Haplosyllis spongicola from Japan (Imajima, 1966) and Korea (Lee & Rho, 1994), agree with the morphology of H. djiboutiensis, which is different from the Mediterranean H. spongicola. The latter has long whip-shaped cirri and dorsal granules are absent (see Lattig et al. 2007; Lattig & Martin 2009). Haplosyllis djiboutiensis resembles the Mediterranean H. granulosa (Lattig, San Martín & Martin, 2007) in body shape, cirri length and presence of dorsal granules, but the latter has more chaetae (3 – 6 per parapodia), all similar, with long denticles on US of MF. Haplosyllis uncinigera (from the Philippines and Australia, see below) also resembles H. djiboutiensis, both in body shape and chaetal characters. The species may be distinguished by aciculae (4 – 5 straight or slightly upwards-directed posterior aciculae in H. unicinigera, vs. 1 – 2 in H. djiboutiensis), and proventricle and pharynx (extending for more than 10 segments in H. uncinigera and for 4 – 6 segments in H. djiboutiensis). Haplosyllis djiboutiensis resembles H. cf. spongicola (see below) in the presence of abundant dorsal granules and in the cirri length alternation; but the chaetae strongly differ, being similar in each parapodium, with short and curved MJP in H. cf. spongicola. Haplosyllis djiboutiensis is the most common and abundant species of genus Haplosylli s from Indo- Pacific waters. Small differences such as the presence of chaetal denticles, apical teeth size and length of MJP have been observed (with the use of SEM), between populations of different geographic areas (i. e. the Philippines, Coasts of Vietnam and Camboya, the Sea of Japan and Hawaii). According to our observations, we suggest that all these populations, including the Australian ones, constitute a species complex, the species of which are impossible to distinguish using morphological techniques. In order to clarify the geographic distribution of H. djiboutiensis and the taxonomic position of all these populations future research using molecular techniques will be necessary.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFB0FF9D2D99FDF1B263FCCE.taxon	distribution	Distribution. AUSTRALIA (Western Australia, South Australia, New South Wales, Northern Territory); RED SEA (Djubal), KOREA; JAPAN.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFB4FF802D99FC9EB56DFC95.taxon	description	Figs 8 A – I, 9 A – F	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFB4FF802D99FC9EB56DFC95.taxon	materials_examined	Examined material. AUSTRALIA. SOUTH AUSTRALIA: HOLOTYPE, AM W 14020 and 2 PARATYPES AM W 36593: American Bay, Kangaroo Island, 35 º 47 ' 08 '' S 137 º 46 ' 31 '' E, coll. by H. Duyverman, 1976. Additional material examined: VICTORIA: 2 specimens AM W 29550, Corio Bay, Port Phillip Bay, 38 ° 06 ' S 144 ° 25 ' 55 " E, 5 m, silty clay, coll. Marine Science Lab, Corio Bay Benthic survey, 1 July 1987. NORTHERN TERRITORY: 1 specimen AM W 36594, Darwin Harbour, East Point, 12 ° 24 ' 36 '' S 130 ° 39 ' 57 '' E, 7 – 10 m, dead coral rubble and algal washings, coll. P. A. Hutchings, 17 July 1993. WESTERN AUSTRALIA: 1 specimen AM W 36595, Goss passage, Beacon Island, 28 º 25 ' 30 '' S 113 º 47 ' E, 20 m, dead plates of Acropora, covered in coralline algae, coll P. A Hutchings, 20 May 1994. 2 specimens AM W 36597, Lafontaine Island, Kimberleys, 14 º 10 ' S 125 º 47 ' E, 15 m, coll. P. A. Hutchings, 19 July 1988. 4 specimens AM W 36596 (plus 1 mounted for SEM), Beacon Island, off south end of Long Island, 28 ° 28 ' 48 '' S 113 ° 46 ' 18 '' E, 5 m, dead coral substrate, covered in coralline algae, coll. P. A. Hutchings, 25 May 1994. 1 incomplete specimen AM W 28398, west of Penguin Island, Warnbro Sound, 32 º 18 ' S 115 º 41 ' E, 5 m, reef substrate, algae and sponge, coll. P. A. Hutchings, 21 March 1993. 1 specimen AM W 28378, Wallabi group of Islands, 28 º 24 ' 07 '' S 113 º 47 ' 23 '' E, 40 m, medium sand with scallops, rubble, coll. P. A. Hutchings, 28 May 1994.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFB4FF802D99FC9EB56DFC95.taxon	description	Description. (Measurements in parenthesis from additional material examined) Body robust, broadest in proventricular segments, gradually tapering towards posterior end (Fig. 9 A). Holotype and two paratypes measuring 2.6 – 2.8 (1.4 – 2.7) cm long, with 107 – 117 (46 – 126) segments. Body pale yellow when preserved, with round, small dorsal granules; dorsal pigment pattern absent. Prostomium subpentagonal, wider than longer, with two pairs of small red eyes in trapezoidal arrangement (Fig. 8 A). Median antennae broken in most observed specimens, inserted on middle of prostomium between eyes (about 30 articles); lateral antennae inserted on anterior prostomial margin, with 20 – 25 articles. Palps longer than prostomium, broadly triangular, fused at their bases but separated all along their length. Sensory organs of palps and nuchal organs not seen. Pharynx orange, broad, difficult to see, extending through 8 (7) segments, with a long triangular anterior tooth, surrounded by crown of about 10 soft papillae and inner ring of cilia; ventral-upper side of papillae with cilia (Fig. 9 B). Proventricle very long, cylindrical, dark-brown, as wide as pharynx, extending through 16 (9 – 11) segments; number of muscle cell rows impossible to assess. Peristomium well defined, shorter than subsequent segments, difficult to see when pharynx everted (Fig. 8 A). Dorsal tentacular cirri longer than ventral ones, with 32 (25 – 35) articles. Antennae, tentacular cirri and dorsal cirri similar. Dorsal cirri slender, first cirri longer than remaining ones, with 33 (30 – 31) articles; thereafter, all similar in length (19 – 30 articles), longest ones slightly exceeding body width. Ventral cirri digitiform, anterior ones broad, as long as parapodial lobes (Figs 8 B, 9 C); median and posterior ventral cirri small, slender, much shorter than parapodial lobes (Fig. 8 C). Chaetae all bidentate, 3 – 5 per parapodia, with few short denticles on US of MF; anterior chaetae with short main fang (Figs 8 D, 9 D); median and posterior chaetae with MJP straight, long; LMF similar in length or longer than SW, apical teeth short, similar in size, with narrow angle (Figs 8 G, 9 E – F). Most aciculae straight, some with curved tip bent at 90 º; 4 – 5 in anterior and median parapodia (Fig. 8 E – F); 2 – 3 in posterior parapodia (Fig. 8 H – I). Pygidium not seen. Reproduction. Not known.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFB4FF802D99FC9EB56DFC95.taxon	discussion	Remarks. Haplosyllis imajimai sp. nov., is characterized by its robust, broad and long body, 3 – 5 aciculae per parapodia, and the long MJP. The new species differs from H. djiboutiensis in its robust body, the median cirri (which are all similar in length) and chaetal form (which are all similar in each parapodia). H. imajimai sp. nov., resembles H. cf. spongicola in body robustness, but the latter has a clear alternation in cirri length, the MJP is curved and short, and it usually have one acicula in posterior parapodia. The Mediterranean Haplosyllis spongicola resembles H. imajimai sp. nov., in body size and acicular shape. However, in the former, the anterior and median chaetae have different morphologies (all similar in H. imajimai sp. nov.). In turn, H. uncinigera differs from H. imajimai sp. nov., in the shape of median and posterior chaetae (see below).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFB4FF802D99FC9EB56DFC95.taxon	distribution	Distribution. AUSTRALIA (Victoria, Western Australia and Northern Territory).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFB4FF802D99FC9EB56DFC95.taxon	etymology	Etymology. The species name refers to Dr. Minoru Imajima from the National Science Museum, Tokyo, who studied the syllids from Japan and significantly contributed to the knowledge on the whole family.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA9FF822D99FCD1B494F9EC.taxon	description	Figs 10 A – H, 11 A – D	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA9FF822D99FCD1B494F9EC.taxon	materials_examined	Examined material. AUSTRALIA: WESTERN AUSTRALIA. 4 specimens AM W 36618, Goss Passage, Beacon Island, 28 º 25 ' 30 '' S 113 º 47 ' E, 8 m, dead plates of Acropora, covered in coralline algae, coll. P. A. Hutchings, 22 May 1994. 60 specimens AM W 36619 (2 specimens mounted for SEM), Lafoutaine Island, Kimberleys, 14 º 10 ' S 125 º 47 ' E, 15 m, coll. P. A. Hutchings, 19 July 1988. 105 specimens AM W 36620, Lafoutaine Island, Kimberleys, 14 º 10 ' S 125 º 47 ' E, 15 m, coll. P. A. Hutchings, 19 July 1988.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA9FF822D99FCD1B494F9EC.taxon	description	Description. Body fragile, translucent, medium to long sized, 7 – 13 mm long for 37 – 53 segments (25 – 35 in juveniles), widest at proventricle segments, gradually tapering towards posterior end (Fig. 11 A); with dark brown pigment pattern on palps, prostomium and anterior segments (even preserved), disappearing at proventricle level, becoming pale cream. Prostomium subpentagonal, wider than long, with two pairs of small red eyes in trapezoidal arrangement (Fig. 10 A). Median antenna inserted on middle of prostomium, twice as long (29 – 58 articles) as lateral ones (13 – 23 articles); lateral antennae inserted on anterior margin of prostomium. Palps slightly longer than prostomium, broadly triangular, fused at their bases but separated all along their length (Fig. 10 A). Sensory organs of ventral side of palps, pharyngeal papillae and nuchal organs not seen. Pharynx pale orange, extending through 6 – 8 segments; with large anterior tooth, surrounded by crown of about 10 soft papillae and inner ring of cilia. Proventricle cylindrical, pale brown, extending through 3 – 6 segments, with 33 – 43 muscle cell rows (Fig. 10 A). Peristomium well defined, shorter than subsequent segments. Antennae, tentacular cirri and dorsal cirri long, slender, whip shaped. Dorsal tentacular cirri twice as long (21 – 33 articles) as ventral ones. Anterior dorsal cirri very long (some twice as long as body width), with a well-defined length pattern: first longer than remaining ones (32 – 63 articles), second shorter (as long as body width, for 12 – 20 articles), third and fourth intermediate, longer than second (19 – 24 and 32 – 42 articles, respectively), fifth shorter (14 – 24 articles), sixth intermediate (24 – 42) (Fig. 10 A). Median cirri alternating long (17 – 41) and short (11 – 18), slightly decreasing in length towards posterior end (Fig. 10 B). Ventral cirri digitiform, anterior ones longer and wider than median and posterior ones, slightly longer than parapodial lobes. Chaetae all bidentate, 2 – 3 per parapodia, apical teeth long, distal one slightly shorter than proximal tooth; anterior chaetae (Figs 10 C; 11 B) with 2 – 3 rows of short spines on US of MF; LMF similar in length than SW; MJP short and curved. Median and posterior chaetae (Figs 10 D, E; 11 C – D) with short, eroded spines on US of MF; LMF longer than SW; MJP straight. Anterior parapodia with 1 – 2 aciculae, straight (Fig. 10 G), and with curved tip bent at 90 º (Fig. 10 F); one acicula in median and posterior parapodia, broader than chaetae, with curved tip bent at 90 º (Fig. 10 H). Pygidium with two long anal cirri (13 – 15 articles). Reproduction. Not known.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA9FF822D99FCD1B494F9EC.taxon	discussion	Remarks. Haplosyllis ingensicola is easily identified by the whip-shaped cirri, longer than body width, the brown pigment pattern on anterior region and the median and posterior broad acicula with curved tip. The apparently cannibalistic behaviour reported for the Indonesian population (Lattig et al. 2010), has not been observed within the Australian one.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA9FF822D99FCD1B494F9EC.taxon	distribution	Distribution. AUSTRALIA (Western Australia); INDONESIA (Java and Sulawesi).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFACFF882D99FBD4B46FFD2E.taxon	description	Figs 14 A – G, 15 A – F	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFACFF882D99FBD4B46FFD2E.taxon	materials_examined	Examined material. AUSTRALIA. WESTERN AUSTRALIA: HOLOTYPE AM W 36580 and 70 PARATYPES AM W 36588 (2 specimens mounted for SEM), Kimberleys, south west corner of Lucas Island, 15 º 13 ' S 124 º 31 ' E, 2 – 30 m, coll. P. A. Hutchings, 24 July 1983. Additional material examined: WESTERN AUSTRALIA: 2 specimens AM W 36589, east side of Fenelon Island, Kimberleys, 14 º 02 ' S 125 º 43 ' E, 6 m, 18 July 1988. 7 specimens AM W 36590, south east end of Long Island, 28 º 28 ' 48 '' S 113 º 46 ' 30 '' E, 8 m, dead coral covered in coralline algae, coll. P. A. Hutchings, 22 May 1994. 2 specimens AM W 36579, north end of Long Island, Goss Passage, 28 º 27 ' 54 '' S 113 º 46 ' 18 '' E, 6 m, dead coral covered in coralline and brown algae, coll. C. Bryce, 22 May 1996. NORTHERN TERRITORY: 3 specimens AM W 36592, Darwin Harbour, East Point, 12 ° 24 ' 36 '' S 130 ° 39 ' 57 '' E, 7 – 10 m, dead coral rubble and algal washings, coll. P. A. Hutchings, 17 July 1993.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFACFF882D99FBD4B46FFD2E.taxon	description	Description. Body translucent, fragile, small, wide at proventricle level, abruptly tapering towards posterior end (Fig. 15 A). Holotype 6 mm long, 0.5 mm wide (excluding parapodia), for 42 segments; paratypes 4 – 6.5 mm long for 25 – 41 segments; pale cream to yellowish when preserved. Scattered small, round dorsal granules on posterior segments. Pigment pattern absent. Prostomium subpentagonal, wider than long, with two pairs of small red eyes in trapezoidal arrangement (Fig. 14 A). Median antennae inserted on middle of prostomium, between eyes (20 – 36 articles); lateral antennae inserted on anterior prostomial margin (11 – 21 articles). Palps long, longer than prostomium, broadly triangular, fused at their bases. Sensory organs in palps and nuchal organs not seen. Pharynx pale orange, extending through 5 – 7 segments, with long triangular anterior tooth, crown of about 10 soft papillae (Fig. 14 B) and inner ring of cilia (Fig. 15 B); groups of cilia make up the sensory organs on upper-ventral side of papillae. Proventricle cylindrical, dark brown, long, extending through 3 – 6.5 segments, broader than pharynx, with 28 – 36 muscle cell rows. Peristomium well defined, shorter than subsequent segments. Dorsal tentacular cirri longer than ventral ones (14 – 24 articles). Antennae, tentacular cirri and dorsal cirri slender. Anterior dorsal cirri relatively long, with welldefined length pattern: first longer than remaining ones (20 – 36 articles), second shorter (7 – 15 articles), third and fourth gradually longer (10 – 14 and 14 – 30 articles respectively), fifth shorter (6 – 12 articles) and sixth intermediate (12 – 20 articles). After proventricle cirri gradually decrease in length towards posterior segments, slightly alternating long (10 – 20) and short (4 – 12); median longest cirri do not exceed ½ of body width; posterior cirri short, with 1 – 7 articles (Fig. 14 C). Ventral cirri digitiform, anterior ones similar or longer to parapodial lobes; median cirri gradually shorter. Chaetae all bidentate, similar throughout, 2 – 3 per parapodia; anterior ones slightly smaller (Figs 14 D; 15 C – D); LMF similar in length to SW; MJP short and curved; US of MF with deep crevice and some small denticles; apical teeth long, well separated, distal tooth slightly smaller than proximal one (Figs 14 F; 15 C – F). Aciculae with upward-directed curved tip, two in anterior parapodia (Fig. 14 E); only one, broader than chaetae, from median to posterior segments (Fig. 14 G). Pygidium not seen. Reproduction. Not known.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFACFF882D99FBD4B46FFD2E.taxon	discussion	Remarks. Haplosyllis sandii sp. nov., is characterized by the unusual deep crevice on the US of MF. It resembles H. tenhovei in body shape and cirri length, but differs in having broad, curved aciculae (which is straight in the latter), and in the absence of the marked serration on US of MF, present in H. tenhovei. The new species differs from H. ingensicola by the long cirri, the curved acicula bent at 90 º, and the long MF.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFACFF882D99FBD4B46FFD2E.taxon	distribution	Distribution. AUSTRALIA (Western Australia, Northern Territory).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFACFF882D99FBD4B46FFD2E.taxon	etymology	Etymology. The specific name refers to Jesús Sandi, the first author husband.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA1FF8B2D99FDBEB436FD05.taxon	description	Figs 16 A – H, 17 A – F	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA1FF8B2D99FDBEB436FD05.taxon	materials_examined	Examined material. AUSTRALIA. WESTERN AUSTRALIA: 3 specimens AM W 36565 (plus 2 specimens mounted for SEM), off Jetty adjacent to Fisheries Hut, Beacon Island, 28 º 25 ' 30 '' S 113 º 47 ' E, 12 m, dead coral, plate-like Acropora montipora, 23 May 1994. 1 specimen AM W 36511, Goss Passage, Beacon Island, 28 ° 25 ' 30 '' S 113 ° 47 ' E, 8 m, dead plates of Acropora, covered in coralline algae, coll. P. A. Hutchings, 19 May 1994. 9 specimens AM W 36566, of south end of Long Island, Beacon Island, 28 ° 28 ' 48 '' S 113 ° 46 ' 18 '' E, 5 m, dead coral substrate covered in coralline algae, coll. P. A. Hutchings, 25 May 1994. NEW SOUTH WALES: 1 specimen AM W 36567, 100 m south of Split Solitary Island, 30 º 15 ' S 153 º 10 ' 30 '' E, 16 m, sponge attached to rocky bottom, coll. R. Springthorpe, 23 June 1992. 1 specimen AM W 36569, south-west side of South Solitary Island, Jervis Bay, 30 º 12 ' S 153 º 16 ' E, 18 m, coral rubble, coll. R. Springthorpe, 24 June 1992. 2 specimens AM W 36568, Elizabeth Reef, south-west outer slope, 29 ° 57 ' 42 " S 159 ° 02 ' 48 " E, coll. P. A. Hutchings, 11 December 1987.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA1FF8B2D99FDBEB436FD05.taxon	description	Description. Body robust, widest at proventricular segments, gradually tapering towards posterior end (Fig. 17 A); small to long-sized, 5 – 12 mm long, for 46 – 80 segments; pale yellow when preserved. Small, round dorso-lateral granules present, more abundant in posterior than in anterior and median regions (Fig. 16 A, D). Prostomium subpentagonal, wider than longer, with two pairs of small red eyes in trapezoidal arrangement (Fig. 16 A). Median antennae inserted on middle of prostomium, between posterior eyes (13 – 24 articles); lateral antennae inserted on anterior prostomial margin (10 – 21 articles). Palps longer than prostomium, broadly triangular, separated all along their length but in close contact at the base. Groups of cilia make up the sensory organs on ventral-upper side of palps (Fig. 17 C); nuchal organs not seen. Pharynx orange, broad, extending through 7 – 10 segments; with long triangular anterior tooth, crown of about 10 soft papillae and inner ring of cilia (Fig. 17 B). Ventral, upper side of papillae with groups of cilia making up the sensory organs. Proventricle cylindrical, long, extending through 7 – 14 segments, dark-brown, as broad as pharynx, with 33 – 61 muscle cell rows. Peristomium well defined, shorter than subsequent segments, difficult to see with pharynx everted. Dorsal tentacular cirri slightly longer (17 – 26 articles) than ventral ones. Antennae, tentacular cirri and dorsal cirri similar. Anterior dorsal cirri with well-defined length pattern, longest ones not exceeding body width: first cirri longer than remaining ones (18 – 35 articles), second short (9 – 15 articles), third and fourth long (11 – 16 and 12 – 17 articles, respectively), fifth short (9 – 12 articles), sixth long (14 – 24 articles). Thereafter, alternating with long (10 – 20 articles) and short (4 – 14 articles), gradually decreasing in length towards posterior end (2 – 10 articles) (Fig. 16 D). Ventral cirri digitiform, anterior ones broad, longer than parapodial lobes (Fig. 16 B); median and posterior ones gradually shorter, smaller than parapodial lobes (Fig. 16 C). Chaetae all bidentate, 2 – 3 per parapodia; anterior ones with LMF similar in length to SW, with numerous short spines on US of MF (Figs 16 E; 17 D – E). Median and posterior chaetae with LMF longer than SW; MJP short and curved; US of MF with few, short denticles (Figs 16 H, 17 F). Apical teeth equal in length, with wide angle. Aciculae straight or with upward-directed curved tip; two in anterior and median parapodia (Fig. 16 F) one (sometimes two) in posterior parapodia (Fig. 16 G). Pygidium with two long cirri (about 10 articles). Reproduction. Haplosyllis cf. spongicola reproduces by acephalous stolons; two female specimens (AM W 36565) were observed in reproductive stage, with the last 9 (of a total of 46 body segments) with pairs of black parapodial ocular spots, capillary (i. e. swimming) chaetae, and 4 – 5 chaetae similarly to anterior ones. These reproductive segments, as well as the normal ones up to proventricular region contained numerous oocytes.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA1FF8B2D99FDBEB436FD05.taxon	discussion	Remarks. The Australian specimens resemble the Mediterranean H. spongicola in chaetal characteristics and body shape. However, their cirri are much shorter (not longer than body width), with a marked alternation length pattern type (long and short), while Mediterranean species have longer cirri, all similar in size. Moreover, type of H. spongicola lack dorsal granules and has aciculae with curved tip bent at 90 º. Therefore, the Australian specimens are here referred as Haplosyllis cf. spongicola, and future molecular studies seems to be the only way try to solve the identity of this morphotype. Haplosyllis cf. spongicola most resembles Haplosyllis imajimai sp. nov., but has short, curved MJP (long, straight in the latter). Additionally, H. imajimai sp. nov., has a smaller, strongly curved acicula, long cirri and a long, robust body.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA1FF8B2D99FDBEB436FD05.taxon	distribution	Distribution. AUSTRALIA (Western Australia and New South Wales)	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA2FF8F2D99FC41B451F8D6.taxon	description	Figs 18 A – F, 19 A – F, 20 A – G, 21 A – D	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA2FF8F2D99FC41B451F8D6.taxon	materials_examined	Examined material. AUSTRALIA. NEW SOUTH WALES: 10 specimens AM W 26629 (plus 2 specimens mounted for SEM), south west of Bowen Island, 35 º 7 ' 49 '' S 150 º 45 ' 77 '' E, ramified yellow sponge, 5 m, 8 December 1993, coll. P. Servo and G. D. Wilson. 2 specimens AM W 26336, north east of Bowen Island, Jervis Bay, 35 º 07 ' S, 150 º 46 ' E, 36 m, clumps of ascidians and bryozoans, 31 May 1987, coll. P. A. Hutchings and P. B. Berents. 3 specimens AM W 36598 (plus 1 specimen mounted for SEM), 150 meters east of Burrill Rocks, 35 ° 23 ' 25 '' S 150 ° 28 ' 11 '' E, 17 m, dead bryozoan encrusted with algae, 1 May 1997, coll. K. B. Attwood. TASMAN SEA: 5 specimens AM W 36599, Taupo Seamount, 33 º 14 ' 13 '' S 156 º 10 ' 41 '' E, 133 m, coll. J. K. Lowry, 2 May 1989. Haplosyllis cf. tenhovei: WESTERN AUSTRALIA. 1 specimen AM W 36600 (plus 1 specimen mounted for SEM), Goss Passage, Beacon Island, 28 º 25 ' 30 '' S 113 º 47 '' E, 33 m, dead coral, embedded in fine sediment, coll. P. A. Hutchings, 23 May 1994. 1 specimen AM W 36616, north end of Long Island, Goss Passage, 28 º 27 ' 54 '' S 113 º 46 ' 18 '' E, 6 m, dead coral covered in coralline and brown algae, coll. C. Bryce, 22 May 1994. NORTHERN TERRITORY: 1 specimen AM W 36617 Darwin Harbour, East Point, 12 ° 24 ' 36 '' S 130 ° 39 ' 57 " E, 10 m, dead coral rubble and algal washings, coll. P. A. Hutchings, 17 July 1993.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA2FF8F2D99FC41B451F8D6.taxon	description	Description. (Measurements in parenthesis for juveniles). Body strong, small, 3 – 6 (1.5 – 1.7) mm long for 25 – 42 (17 – 20) segments, less than 1 mm wide excluding parapodia, widest at proventricular level (Fig. 19 C); pale yellow, with some cirri having brownish round spots. Dorsal granules and pigment pattern absent. Prostomium subpentagonal, wider than long, with two pairs of small red eyes in trapezoidal arrangement (Fig. 18 A – B). Median antennae inserted on middle of prostomium, between eyes, twice as long as lateral ones, with 16 – 29 (10 – 16) articles; lateral antennae inserted on anterior prostomial margin, with 10 – 16 (6 – 13) articles. Palps long, slightly longer than prostomium, broadly triangular, fused at their bases. Sensory organs of palps and nuchal organs not seen. Pharynx orange, extending through 5 – 7 (3 – 4) segments; with large triangular anterior tooth, crown of about 10 soft papillae and inner ring of cilia (Fig. 19 B). Ventral side of papillae with rows of cilia making up sensory organs. Proventricle cylindrical, dark-brown, long, extending through 6 – 10 (3 – 6) segments, with 35 – 40 (22 – 26) muscle cell rows. Peristomium well defined, shorter than subsequent segments. Dorsal tentacular cirri longer than ventral ones, with 15 – 24 (10 – 17) and 7 – 18 (3 – 13) articles respectively. Antennae, tentacular cirri and dorsal cirri similar, long and slender. Anterior dorsal cirri long, with a well-defined length pattern: first cirri slightly longer than body width, longer than remaining ones, with 21 – 29 (12 – 18) articles, second shorter, 9 – 13 (3 – 7) articles, third and fourth intermediate, gradually longer than second, with 11 – 19 (3 – 10) and 15 – 26 (2 – 13) articles respectively, fifth shorter, 9 – 15 (2 – 7) articles, sixth intermediate, with 14 – 23 (3 – 8) articles. Thereafter, cirri slightly alternating with long (13 – 24 articles, almost as long as body width) and short (6 – 15 articles), gradually decreasing in length towards posterior end. Ventral cirri digitiform, anterior ones as long as parapodial lobes; median and posterior ones shorter. Chaetae all bidentate, 2 – 3 per parapodia, similar throughout, anterior ones slightly smaller (Figs 18 C; 19 D); LMF similar in length to SW; MJP short, curved and relatively long; US of MF with a long denticle and few smaller ones toward the tip; apical teeth similar in length, long, well separated (Figs 18 D; 19 E – F). Aciculae straight broad, with rounded tips (Fig. 18 E – F), 2 – 3 per anterior and median parapodia, one in the posterior end. Pygidium not seen. Reproduction. Not known.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA2FF8F2D99FC41B451F8D6.taxon	distribution	Distribution. INDONESIA (Komodo and Karang Kaledupa Islands). AUSTRALIA (New South Wales, Western Australia and Northern Territory).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FFA2FF8F2D99FC41B451F8D6.taxon	discussion	Remarks. The original description of H. tenhovei is here complemented with the description of the long denticles on the US of MF, the sensory organs on the pharyngeal papillae and the nuchal organs. Some small specimens, likely juveniles, (AM W 36629) had shorter and broader cirri than those of adults, but all chaetal features remain identical (Figs 18 B; 19 A, D). Haplosyllis tenhovei resembles H. sandii sp. nov., in body shape and cirri length, but differs in the acicular shape (straight in the former, with curved tip in the latter). A few specimens are herein considered as Haplosyllis cf. tenhovei (see examined material) due their slender bodies (Figs 20 A – B; 21 A), but more importantly, by the presence of a small slender, simple chaetae on the posterior last 1 – 4 segments (Figs 20 E; 21 D), resembling that described by Imajima (1966, fig. 43 f), and never seen in any other Haplosyllis species. However, those specimens are not considered as a new species due to their chaetal and acicular shapes (Figs 20 C – D, F – G; 21 B – C) being very similar to those of H. tenhovei, the most characteristic features of the species. Furthermore, the length of their dorsal cirri, pharynx and proventricle, agree with the same measured values in specimens of H. tenhovei.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FF99FFB12D99FFC4B522F848.taxon	materials_examined	Examined material. AUSTRALIA, WESTERN AUSTRALIA: 1 specimen AM W 36624, reef south of Lucas Island, Brunswick Bay, Kimberley region, 15 ° 16 ' S 124 ° 29 ' E, 2 m, coll. P. A. Hutchings, 24 July 1988. 2 specimens AM W 36625 (plus 1 specimen mounted for SEM), south side of Long Reef, Kimberley region, 14 º 01 ' S 125 º 44 ' E, 20 m, 18 July 1988, coll. P. A. Hutchings. Haplosyllis cf. uncinigera: 1 specimen AM W 36626, northeast entrance to Goss Passage, Beacon Island, 28 º 27 ' 54 '' S 113 º 46 ' 42 '' E, 33 m, underneath isolated boulders embedded in coral sand, 25 May 1994, coll. P. A. Hutchings. Comparative material examined. South China Sea: Haplosyllis hainanensis 1 SYNTYPE SMF 10856; 1 specimen SMF 10855; 3 specimens SMF 10857; 2 specimens SMF 10858. Philippines: Haplosyllis uncinigera HOLOTYPE ZMB F. 1977.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FF99FFB12D99FFC4B522F848.taxon	description	Description. Body robust, long, 1.8 – 2 cm long, for 100 segments (Figs 22 A, 23 A), widest at proventricular region, gradually tapering towards posterior end. Two specimens with dark brown pigmented palps, prostomium and anterior segments; pale yellow from proventricular region to posterior end (Fig. 22 A). Prostomium subpentagonal, with two pairs of small red eyes in trapezoidal arrangement. Median antennae inserted on middle of prostomium (16 – 21 articles); lateral antennae on anterior prostomial margin (11 – 15 articles). Palps long, broadly triangular, fused at their bases. Sensory organs of palps not seen. Nuchal organs as two lateral ciliated grooves between prostomium and peristomium. Pharynx very long, extending through 12 segments, with large anterior tooth, crown of 10 – 13 soft papillae and inner ring of cilia (Fig. 23 B); upper ventral side of papillae with groups of cilia making up the sensory organs. Proventricle cylindrical, extending through about 11 segments, dark brown, with 58 – 84 muscle cell rows. Peristomium well defined, shorter than subsequent segments. Dorsal tentacular cirri (18 – 20 articles) longer than ventral ones. Dorsal cirri similar to antennae and tentacular cirri, long, slender, whip-shaped. Anterior dorsal cirri slightly longer than medium and posterior ones; first long (24 – 28 articles); second short (8 – 16 articles), third and fourth long (23 and 22 – 28 articles, respectively), fifth intermediate (14 – 21 articles), sixth long (20 – 25 articles); thereafter, slightly alternating with long (20 – 29 articles) and slightly shorter (10 – 18 articles), gradually decreasing in length towards posterior end. Posterior cirri relatively short (6 – 14 articles). Ventral cirri digitiform, anterior ones broad, as long as parapodial lobes (Fig. 22 B), median and posterior ones, slender (Fig. 22 C). Anterior chaetae bidentate, 2 – 3 per parapodia, with apical teeth long, similar in length; LMF shorter than SW, with short denticles on US of MF (Figs 22 D; 23 C – D). Median and posterior chaetae of two different sizes, long and short, each parapodia with 1 – 2 long chaetae and one short (Figs 22 G – H; 23 E – F); long chaetae with narrow angle between apical teeth, distal one very small, sometimes giving a unidentate appearance (Figs 22 G; 23 F), with LMF similar in length to SW; MJP straight and relatively long; US of MF with few, short denticles, sometimes eroded. Short chaetae markedly bidentate, with wide angle between apical teeth, proximal tooth shorter than distal one; MJP short (Figs 22 H; 23 E). Aciculae of three types: straight (Fig. 22 J); with upwarddirected curved tips (Fig. 22 I), and (more rarely) with curved tips bent at 90 º (Fig. 22 K); 4 – 5 in anterior parapodia, 3 – 4 in median and posterior regions. Pygidium with two (broken) anal cirri. Reproduction. Not known.	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FF99FFB12D99FFC4B522F848.taxon	discussion	Remarks. The Australian specimens are identified as H. uncinigera due to their long pharynx and proventricle, the presence of two different chaetal shapes on each parapodia (long, with unidentate appearance and short, markedly bidentate). Two specimens also had a brown pigment pattern as reported by Sun (1996) for the Chinese population. Accordingly, the presence of this pigmentation seems to be a variable character, being more frequent in long and robust specimens (Lattig & Martin 2009).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
03A487A3FF99FFB12D99FFC4B522F848.taxon	distribution	Distribution. AUSTRALIA (Western Australia); PHILIPPINES (Bohol); SOUTH CHINA SEA (Nansha and Hainan Islands). FIGUE 22. Haplosyllis uncinigera. A — anterior end, dorsal view; B — anterior parapodium; C — posterior parapodium; D — anterior chaeta; E and F — anterior aciculae; G — long chaeta with small distal tooth, midbody; H — small chaeta with bidentate tip; I, J, K — posterior aciculae. Scale: A = 500 µm; B, C = 200 µm; D – K = 20 µm. (AM W 36624).	en	Lattig, Patricia, Martin, Daniel, Martín, Guillermo San (2010): Syllinae (Syllidae: Polychaeta) from Australia. Part 4. The genus Haplosyllis Langerhans, 1879. Zootaxa 2552 (1): 1-36, DOI: 10.11646/zootaxa.2552.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.2552.1.1
