identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A487FEFF88FF9B05B1FEC57F6FFDC8.text	03A487FEFF88FF9B05B1FEC57F6FFDC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phasmister hkamticus Caterino and Yamamoto 2023	<div><p>Phasmister hkamticus Caterino and Yamamoto, new species zoobank.org/ urn:lsid:zoobank.org:act: C32C48F8-68A5-4313-A6E1-DE609FF0ADE1 (Figs. 1, 3B)</p> <p>Type Material. Holotype specimen, of unknown sex (SEHU-0000121202); deposited in HUM. Type locality: Northern Myanmar: Sagaing Region, Hkamti District. The holotype is embedded in a small, elongate-oval, yellowish transparent amber piece (9.8 × 4.9 × 1.7 mm).</p> <p>434 Diagnostic Description. Full body length, in situ: with posterior and lateral margins raised, antennal 1.21 mm (apex of head to posterior end of abdomen); insertions open anteriorly; antennal club with disbody (pronotum + elytra) length along midline: tinct transverse annuli; prosternum with lateral slots 1.02 mm; maximum (humeral) width: 0.71 mm; for passage of the antennal funicle; hypomeral head width: 0.26 mm; pronotal length: 0.34 mm; antennal depression open beneath. Phasmister pronotal width: 0.56 mm. This species shares many hkamticus differs from P. cristatus in the following of the generic level characters described for specific characters: prothorax less strongly trans- Phasmister (Caterino 2021): body subquadrate, verse, length/width ratio slightly greater (see Fig. 3); weakly compressed; elytra with every second inter- anterior corners of pronotum not as produced, less stria strongly raised to form a carina; humeral elytral deeply explanate; anterior margin of pronotum angles expanded; pronotum with raised lateral and rather narrow; median pronotal carina less dorsally anterior margins, depressed in anterior corners, with projecting (as seen in lateral view), little higher than various carinae on disc; pronotal disc strongly car- elytral carinae, more obtusely divergent near apex, inate; posterior pronotal margin sinuate, produced then angulate to anterior margin, enclosing a disposteriad in front of small scutellar shield; frons tinctly (irregularly) pentagonal space; sublateral elongate; propygidium deeply concave in basal half, punctate apically; pygidium with apex directed nearly anteriad.</p> <p>Derivation of Specific Name. The name hkamticus is an adjective referring to the novel source of amber (the Hkamti amber deposits) in which the specimen was found.</p> <p>Remarks. The differences between this species and P. cristatus are well-marked, despite their clearly close relationship. The distinct conformation of pronotal carinae separates them most clearly, with the broader, pentagonal area between the median carinae (Fig. 3B) of the new species quite distinct. The apparently lower overall elevation of the median pronotal carina also appears distinctive. The differences of both of these species from Phasmister parallelus, new species are more noteworthy. Most distinct are the differences in tibial shape, particularly in the broad, outwardly angulate protibia of P. parallelus, with a medial rather than marginal groove. Phasmister hkamticus and P. cristatus are both distinctly flattened dorsally, where the species described below is more convex.</p> <p>Syninclusions. No arthropod inclusions are included, but there is some small organic debris and a partial remnant of a boring of Pholadidae (Bivalvia: Myoida) within the amber piece.</p> </div>	https://treatment.plazi.org/id/03A487FEFF88FF9B05B1FEC57F6FFDC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Caterino, Michael S.;Yamamoto, Shûhei	Caterino, Michael S., Yamamoto, Shûhei (2023): New Onthophiline Fossil Species (Coleoptera: Histeridae: Onthophilinae) from Mid-Cretaceous Burmese Amber. The Coleopterists Bulletin 77 (3): 432-438, DOI: 10.1649/0010-065X-77.3.432, URL: http://dx.doi.org/10.1649/0010-065x-77.3.432
03A487FEFF8EFF9905B3FC417918F886.text	03A487FEFF8EFF9905B3FC417918F886.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phasmister parallelus Caterino and Yamamoto 2023	<div><p>Phasmister parallelus Caterino and Yamamoto, new species zoobank.org/ urn:lsid:zoobank.org:act: FD8814F7-CC2C-454F-BD7E-9C7726DDEC83 (Figs. 2, 3C)</p> <p>carinae extending further anteriad, meeting posterior corners of this pentagon, rather than meeting the median carina before the middle. Details visible in this specimen, not fully described in P. cristatus, include: labrum short, rounded, separated from epistoma by distinct suture; mandible (left) weakly excavate on outer surface, with outer margin strongly angulate to acute apical tooth; maxillary palp 4-segmented, basal palpomere barely visible, third palpomere shorter than second, apical palpomere about twice as long as penultimate, rather bulbous at base, narrowed to weakly obliquely truncate apex; antennal scape curved, pedicel nearly as wide as scape, about twice as long as succeeding segment; antennomeres 3–8 (funicle) gradually decreasing in width; antennal club oval, only slightly</p> <p>Type Material. Holotype specimen, of unknown sex (SEHU-0000121203); deposited in HUM. Type locality: Northern Myanmar: Kachin State, Hukawng Valley, Noije Bum Hill. The holotype is embedded in a small, rather flattened, more-or-less rectangular, yellowish transparent amber piece (6.1 × 3.8 × 1.9 mm). The right mesotarsus is partially lost due to over-polishing, but all mesotarsomeres are preserved.</p> <p>Diagnostic Description. Full body length, in situ: 1.24 mm (apex of pronotum to posterior end of abdomen); body (pronotum + elytra) length: 1.18 mm; maximum (humeral) width: 0.89 mm; pronotal length: 0.45 mm; pronotal width: 0.75 mm. This species shares many of the generic level characters described for Phasmister (Caterino 2021): body subquadrate, weakly compressed; elytra with every second interstria strongly raised to form a carina; humeral elytral angles expanded; pronotum with raised lateral and anterior margins, depressed in anterior corners, with various carinae on disc; pronotal disc strongly punctate; posterior pronotal margin sinuate, produced posteriad in front of small scutellar shield; frons with dorsal and lateral 436 margins raised; antennal insertions open anteriorly; distinct, with paired and widely separate median antennal club with distinct, transverse, annulate carinae, joined only near the base, while both sutures; prosternum with lateral slots for passage of preceding species have a single median carina the antennal funicle; hypomeral antennal depression divergent only near the anterior margin. In open beneath. It differs from its congeners in the tibial characters in particular, P. parallelus ap- following specific characters: body thicker, more pears more plesiomorphic, with the slender tibiae convex, elytral surface more gradually descending of P. cristatus and P. hkamticus constituting (in lateral aspect) to apices; prothorax less strongly apomorphic states. transverse, length/width ratio slightly greater; paired Synclusions. There is a partial fragment of an median pronotal carinae divergent at basal fourth, arthropod leg located close to the holotype. subparallel to apex; curved paramedial pronotal carinae weaker; lateral longitudinal pronotal carinae Carinumerus intricatus Caterino and well developed, curving mediad near middle; Yamamoto, new species anterior corners of pronotum curved more inward, zoobank.org/urn:lsid:zoobank.org:act:</p> <p>less strongly explanate; sublateral elytral carina</p> <p>0E01A568-570D-465B-A7FA-3C666F043847 (between stria 1 and inner subhumeral stria) more (Figs. 4, 5C) evenly curved from base to apex, only weakly sinuate; tibiae, especially protibia, broader, with angu- Type Material. Holotype specimen, of unknown late lateral margin, less elongate, apical process sex (SEHU-0000121204); deposited in HUM. Type weak; tarsi similarly shorter; protarsal groove along locality: Northern Myanmar: Kachin State, Hukawng mid-axis of protibia, rather than along outer margin. Valley, Noije Bum Hill. The holotype is embedded Derivation of Specific Name. The name parallelus in a small, narrowly elongate-ovoid, yellowish transis an adjective referring to the paired pronotal carinae, parent amber piece (11.5 × 4.2 × 2.3 mm). which are parallel along much of their lengths. Diagnosis. Full body length, in situ: 1.27 mm Remarks. The body shape and tibial shape dif- (apex of pronotum to posterior end of abdomen); ferences of P. parallelus from the previously de- body (pronotum + elytra) length: 1.17 mm; maxiscribed species are detailed above. The pronotal mum (humeral) width: 1.19 mm; pronotal length: carinae (Fig. 3C) of P. parallelus are also quite 0.42 mm; pronotal width: 0.89 mm. Similar in most</p> <p>Remarks. While all Carinumerus species are very similar, the pronotal carinae of the three species are distinct. In C. yingae there are several more-or-less parallel carinae on each side, including two subparallel median ridges that extend two-thirds the length of the pronotum; in C. maddisoni there are no median ridges at all, with only one short and one longer lateral carina; and in C. intricatus there appears to be a single median ridge, dividing and diverging anteriorly. The elytral strial punctures of C. intricatus appear to be somewhat smaller than those of C. yingae, while those of C. maddisoni are larger and somewhat elongated along the striae. Other views of the species are not consistently wellpreserved enough to assess clear differences. Among the available fossils of the genus, the metathoracic leg of the C. intricatus holotype is particularly clear, revealing short spines along the outer margin, a series of fine setae along the inner margin, two small but distinct apical spurs, and an inwardly pointing apical process. The basal four metatarsomeres (as with all the others) have a single pair of distinct setae at their apices.</p> <p>Syninclusions. There are no other Arthropoda inclusions.</p> <p>respects to its two known congeners (C. yingae and C. maddisoni), with all elytral interstriae evenly carinate, expanded humeral angles, and raised frontal margins. Species-level differences of C. intricatus are subtle, and somewhat difficult to observe in the fossil, with the pronotal disc obscured by an airfilled fracture. The pronotum bears a single median ridge (or possibly very approximate, parallel median ridges) that diverge near the anterior margin; distinct submedian carinae converge toward the anterior arms of the median pronotal ridge near the anterior pronotal margin; the elytral strial punctures are small.</p> <p>Derivation of Specific Name. The species name intricatus is a participle referring to the intricate pattern of pronotal carinae that distinguish it.</p> </div>	https://treatment.plazi.org/id/03A487FEFF8EFF9905B3FC417918F886	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Caterino, Michael S.;Yamamoto, Shûhei	Caterino, Michael S., Yamamoto, Shûhei (2023): New Onthophiline Fossil Species (Coleoptera: Histeridae: Onthophilinae) from Mid-Cretaceous Burmese Amber. The Coleopterists Bulletin 77 (3): 432-438, DOI: 10.1649/0010-065X-77.3.432, URL: http://dx.doi.org/10.1649/0010-065x-77.3.432
