taxonID	type	description	language	source
03A41512FFFA482CFEBF2BC242AF02E4.taxon	diagnosis	Emended diagnosis. — The genus Carpomys comprises two extant, relatively small cloud rats and an extinct giant cloud rat, Ca. dakal n. sp. Carpomys is defined phylogenetically as including the most recent common ancestor of Ca. melanurus, Ca. phaeurus, Ca. dakal n. sp., and all of its descendants. Living members of Carpomys possess cranial traits and features of external appearance that were summarized by Musser and Heaney (1992) and Heaney et al. (2014). The dentition of Carpomys is distinguished from all others by the combination of the following morphological features: upper and lower first molars have four rows of lophs; second and third upper molars have three rows of lophs; a welldeveloped anterior labial cusp is present in m 2 and m 3; and posterior cingula in all upper molars are laminar and elongate (Figs. 2 A, 2 B, 4 A, and 4 B). Carpomys has moderately hypsodont molars and chevronate molar lophs. In the M 1, a well-developed and elongate posterior cingulum forms the fourth lamina, while in the m 1 a bicuspid anterocentral forms the first loph (Thomas 1898; Musser and Heaney 1992). The posterior loph of the m 3 of Carpomys usually has an irregular “ W ” outline formed by an additional cusplet between the hypoconid and entoconid (Figs. 5 A and 5 B).	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFA482CFEBF2BC242AF02E4.taxon	description	Description and comparisons. — In Carpomys, the height of crowns is medium andtooth morphology is intermediate between the hypsodont (high-crowned) and more chevronate Crateromys and the lophodont (elongated and transverse) Phloeomys; Carpomys cuspsaretypicallyseparatedby deepclefts (Figs. 2 – 5). As described by Heaney et al. (2009, 2014), the largely fused and lamellate cusp patterns of Carpomys and Musseromys aresimilar, but certain dental traits differentiate the two. Musseromys teeth and body sizes are substantially smaller than those of Carpomys (Heaney et al. 2014). In M 3 (Figs. 2 A, 4 B, and 4 C), Carpomys has a well-developed and elongate posterior cingulum that either is reduced or absent altogether in Musseromys. Carpomys has more chevronate lophs (Figs. 5 A and 5 B) compared to the more transversely oriented laminae of m 1 (second and third lophs) and m 3 (anterior loph) in Musseromys (Fig. 5 C). The m 3 cusplet found in Carpomys is absent in Musseromys and the m 3 posterior lamina is transverse in Musseromys (Fig. 5 C). The Carpomys M 1 anterocone is slightly arcuate and intermediate in curvature between the transverse anterocone of Phloeomys and the arcuate to chevronate anterocones of Batomys and Crateromys. In the M 2 and M 3, the third lamina is represented by a well-developed posterior cingulum, which is absent in Batomys and Crateromys, and absent in M 3 of all Musseromys. The Carpomys m 1 anteroconidiscomposedof two laminae, in contrast to the more typical triangular anteroconid in other phloeomyines. In Carpomys, the anteroconid of the m 1 has a midline cusp that forms a ridge connecting the anterocentral and second lamina (anterolabial – anterolingual loph) when it is worn (Figs. 2 B and 5 A; Supplementary Data SD 1 G and I); in Crateromys, Batomys, and Musseromys, this ridge is not observed, even when the anteroconid is worn down. Carpomys has more arcuate laminae in m 1 and m 2 compared to Musseromys, but less strongly chevronate compared to Crateromys and Batomys (Fig. 5). The posterior cingulum is absent in the m 3 of Carpomys, but present in about 25 % of Cr. schadenbergi included in this study (listed in Table 3). Carpomys upper and lower second and third molars typically have greater occlusal widths than occlusal lengths, but the opposite is seen in Crateromys and Batomys (Tables 2 and 3).	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFB482EFC6F2EDD44DE01A3.taxon	description	Figs. 3 A, 3 B, 4 A, and 5 A; Tables 1 and 2; Supplementary Data SD 1	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFB482EFC6F2EDD44DE01A3.taxon	materials_examined	Holotype. — PNM II- 1977 - J 11 A- 16301, left dentary with incisor and m 1 – 3; only a small portion of the dentary posterior to m 3 is present (Figs. 3 A and 3 B). The m 1 – 3 are lightly worn (Fig. 5 A), indicating that the animal was a young adult at the time of death. Type locality. — Minori Cave, Barangay Quibal, Peñablanca Municipality, Cagayan Province, Luzon Island, Philippines (Fig. 1), 200 m elevation, 17 ° 43 ′ 17 ″ N, 121 ° 49 ′ 42 ″ E, Chamber Dof Minori Cave, Square 27; derives from Holocene aceramic level 40 – 50 cm below cave surface. Paratypes. — (n = 10; Supplementary Data SD 1). Callao Cave: partial right dentary with m 2 (PNM II- 1977 - J 3 - 10196); partial left dentary with m 1 (PNM II- 1977 - J 3 - 10544); partial right maxillary with M 2 (PNM II- 1977 - J 3 - 11075 / 3); Minori Cave Chamber A: partial leftdentary with m 1 – 2 (PNM II- 1977 - J 11 B- 2986); Minori Cave Chamber D: partial left maxilla with M 1 – 3 (PNM II- 1977 - J 11 A- 15997); partial right maxilla with M 1 – 2 (PNM II- 1977 - J 11 A- 16447); partial left maxilla with M 1 – 2 (PNM II- 1977 - J 11 A- 16481); partial left maxilla with M 1 – 2 (PNM II- 1977 - J 11 A- 16825); partial right dentary with m 2 – 3 (PNM II- 1977 - J 11 A- 16951); Musang Cave: partial left dentary with m 1 – 3 (PNM II- 1976 - M- 101). Age of specimens. — Callao Cave specimens 10544 and 10196 originated in a layer dated at least ca. 67,000 BP; specimen 11075 is from Layer 11 of Callao Cave, which is older than ca. 30,000 cal BP but younger than 52,000 BP; seven specimens from Minori Cave, including the holotype, are from aceramic Holocene levels (prior to the introduction of pottery on Luzon at 4,500 – 4,000 cal BP); Musang Cave specimen 101 derives from aceramic-bearing level (Level I 3; Table 1).	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFB482EFC6F2EDD44DE01A3.taxon	etymology	Etymology. — The new species name comes from the languages of the Dupaningan Agta, Itawes, and Ibanag of NE Luzon, an adjective meaning “ large ” in all three languages (see Tharp and Natividad 1976; Greenhill et al. 2008; Robinson 2011); we use it as an adjective in the nominative singular neuter.	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFB482EFC6F2EDD44DE01A3.taxon	distribution	Distribution. — Known from three cave sites in the Callao Cave complex, Peñablanca Municipality, Cagayan Province (Fig. 1).	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFB482EFC6F2EDD44DE01A3.taxon	diagnosis	Diagnosis. — A member of the genus Carpomys as diagnosed by Musser and Heaney (1992) and as emended above (see Emended Diagnosis and Description and Comparisons); a giant species much larger than the two living species, Ca. melanurus and Ca. phaeurus (Table 2). The dentary is robust both absolutely and proportionately, with strongly marked inferior and superior masseteric ridges. Lower incisor is unusually wide, 2.3 mm at its midpoint. Upper toothrow length is 13.58 mm; lower toothrow length ranges from 13.25 to 13.46 mm. M 1 length ranges from 4.49 to 5.49 mm; M 1 width ranges from 3.42 to 4.41 mm. m 1 length ranges from 5.93 to 6.57 mm; m 1 width ranges from 3.37 to 4.53 mm (Table 2).	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFB482EFC6F2EDD44DE01A3.taxon	description	Description and comparisons. — A giant cloud rat, largest known member of Carpomys (Table 2; Fig. 6), easily distinguished from the two extant species by its much greater size. The dentary is both larger and proportionately more robust than in extant species, with strongly marked inferior and superior masseteric ridges. The mental foramen is moderately high relative to the apex of the masseteric ridges, situated close to the dorsal surface of the mandible and anterior to m 1. The lower incisor, as represented on the holotype, is robust, orange-yellow on the anterior surface, 2.3 mm wide at its midpoint and 8.5 mm long from its tip to the point it meets the dentary bone along its medial edge. The dorsal wear surface extends fully to the beginning of the dentary bone slightly below the anteriormost projection of bone along the medial edge, and the tip lies slightly below the occlusal plane of the mandibular molars. The lower incisor of Ca. phaeurus also is orange-yellow on the anterior surface, but is both shorter and more slender, 1.0 mm wide at its midpoint and 5.0 mm (n = 2) from its tip to the anteriormost projection of bone along the medial edge. In this species, the dorsal wear surface extends from the tip about 90 % of the way to the highest point of the dentary bone; the tip lies slightly below the occlusal plane of the mandibular molars, equivalent to Ca. dakal. The lower incisor of Ca. melanurus (n = 1) is 1.3 mm wide at the midpoint, and about 6.0 mm from its tip to the anteriormost projection of bone along the medial edge. Thus, the lower incisor of Ca. dakal is nearly twice the width of those of the two extant species. Overall molar morphology follows that shown in Figs. 2 A and 2 B. Measurements of individual teeth are provided in Table 3. Molar dimensions of the two living Carpomys species comprise only 40 – 80 % of corresponding measurements in Ca. dakal. Molar dimensions of Musseromys species are only 20 – 30 % of the corresponding measurements in Ca. dakal. The variation in molar characters observed in Ca. dakal also is seen in Ca. melanurus and Ca. phaeurus; size is the main distinguishing character that differentiates the dental morphology of fossil species from the living species. Additional cusplets are often present in M 1 and M 2 of Carpomys (Fig. 4 A; Supplementary Data SD 1 B and C): a labial cusplet between t 3 and t 6 of M 1 and a smaller cusplet between t 5 and t 8 of M 2, which are absent in other phloeomyine genera. The M 1 cusplet is found in three out of five Ca. dakal specimens, compared to two out of five Ca. phaeurus specimens and none in three Ca. melanurus specimens. The M 2 cusplet is present in one of five specimens of Ca. dakal and is absent in specimens of the two extant species. Molar tooth dimensions of Ca. dakal are within the size range of the giant cloud rats Cr. schadenbergi and Cr. heaneyi, but compared to them, Ca. dakal has a more slender dentary (Fig. 3). The maxillary region is robust (Fig. 7); our fragmentary material implies that the palate is rather smooth with a raised ridge supporting the molar teeth, and a large posterior palatal foramen (ppf) adjacent to M 2 (Fig. 7 A), as seen in the two extant Carpomys species. No trace of the deep palatal grooves of Crateromys spp. is evident. It appears that the bony palate terminates about one-third of the way posterior to the anterior edge of M 3, as in Ca. melanurus, rather than at the anterior edge of M 3, as in Ca. phaeurus. The anterior root of the zygomatic arch appears to originate labial to M 1 (Fig. 7 B), similar to the extant species of Carpomys, and perhaps a bit further posterior than that of Cr. schadenbergi and Cr. ballik n. sp.	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFE482BFEB42CFD44FA0244.taxon	description	Figs. 4 D, 5 D, 8, 9, and 10; Tables 1 and 3; Supplementary Data SD 2	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFE482BFEB42CFD44FA0244.taxon	materials_examined	Holotype. — PNM II- 1977 - I 8 - 402 right dentary with m 1 – 3; incisor and all posterior processes missing (Figs. 8 A and 8 B). The m 1 – 3 are moderately worn (Fig. 5 D), indicating that the animal was an adult at the time of death. Type locality. — Dalan Serkot Cave, Barangay San Roque, Peñablanca Municipality, Cagayan Province, Luzon Island, Philippines (Fig. 1), 165 m elevation, 17 ° 39 ′ 52.2 ″ N, 121 ° 49 ′ 12 ″ E, Square 2, Layer 2 (aceramic layer, spit 24, approximately 1.2 m below surface). Paratypes. — (n = 6; Supplementary Data SD 2) Callao Cave: partial maxilla with left and right M 1 – 3 (PNM II- 1977 - J 3 - 10389; Fig. 10), partial right dentary with incisor fragment and m 1 – 2 (PNM II- 1977 - J 3 - 11075 / 2); Minori Cave: partial left dentary with m 1 – 3 (PNM II- 1977 - J 11 A- 17018), partial left dentary with m 1 – 2 (PNM II- 1977 - J 11 A- 16692), partial left dentary with m 2 – 3 (PNM II- 1977 - J 11 A- 896); Eme Cave: partial left dentary with m 1 – 3 (PNM II- 1977 - G- 292). Age of specimens. — The holotype comes from an aceramic layer (Layer 2) of Dalan Serkot Cave, dated to 7,254 – 6,995 cal BP (Table 1); specimen 10389 derives from the Late Pleistocene levels of Callao Cave at least ca. 66,700 BP; specimen 11075 / 2 comes from Layer 11 of Callao which is older than ca. 30,000 cal BP and less than 52,600 BP; specimen 292 derives from an aceramic layer in Eme Cave dated to 3,985 – 3,702 cal BP; specimens 17018 and 16692 come from aceramic Holocene levels of Minori Cave (prior to the introduction of pottery on Luzon at 4,500 – 4,000 cal BP); specimen 896 derives from aceramicbearing level from Minori.	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFE482BFEB42CFD44FA0244.taxon	etymology	Etymology. — The new species name comes from the Dupaningan Agta language of NE Luzon, an adjective meaning “ small ” (see Reid 1994; Robinson 2011); we use it as an adjective in the nominative singular neuter.	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFE482BFEB42CFD44FA0244.taxon	distribution	Distribution. — Known from four cave sites in the Callao Cave Complex, Peñablanca Municipality, Cagayan Province (Fig. 1), as noted above.	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFE482BFEB42CFD44FA0244.taxon	diagnosis	Diagnosis. — A relatively small cloud rat of the genus Crateromys as diagnosed and defined by Musser and Gordon (1981), Musser et al. (1985), and Musser and Heaney (1992), distinctly smaller and less robust in comparison to the only living congener on Luzon, Cr. schadenbergi (Table 3; Fig. 9). Incisive foramina (if) extend nearly (about 0.5 mm anterior) to the anterior margin of the M 1 alveolus (Fig. 10). Palate is narrow, and palatal grooves end about parallel to t 1 on M 1 (Fig. 10). Dentary is moderately robust relative to molar dimensions (Fig. 9 A). Lower incisors proportionately narrow, 1.8 mm at midpoint. Upper toothrow length is 11.45 mm; lower toothrow length ranges from 10.22 to 12.91 mm. M 1 length is 4.17 mm; M 1 width is 3.27 mm. m 1 length ranges from 4.2 to 5.41 mm; m 1 width ranges from 2.79 to 3.34 mm (Table 3). Labial cusps (t 3 and t 6) of M 1 and M 2 are less prominent relative to t 2 and t 5. M 1 and M 2 t 6 is directly fused with t 5, notches are absent between t 5 and t 6. M 1 anteroconid is not connected to the anterolabial – anterolingual loph.	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
03A41512FFFE482BFEB42CFD44FA0244.taxon	description	Description and comparisons. — Overall, Cr. ballik is a small member of the genus Crateromys, smaller than Cr. schadenbergi and Cr. heaneyi, similar in size but a bit smaller on average than Cr. paulus, and smaller than Ca. dakal (Tables 2 and 3; Fig. 9). The dentary of Cr. ballik is less robust relative to its molar dimensions compared to Cr. schadenbergi and Cr. heaneyi (Fig. 9 A), and less robust in absolute terms than Cr. paulus. The mental foramen of the dentary is situated at or slightly above the apex of the superior and inferior masseteric ridges on Cr. ballik (Fig. 8 A), whereas it is placed high above the apex in Cr. paulus, situated on the dorsal margin of the dentary and anterior to m 1, and is much lower on Cr. schadenbergi and Cr. heaneyi (see also Musser and Gordon 1981; Gonzales and Kennedy 1996; Reyes et al. 2017). The lower incisor, as represented by Callao Cave paratype specimen 11072 / 2, is robust, orange-yellow on the anterior surface. The tip is missing, but at a place estimated to have been the midpoint, it is 1.8 mm wide, and from the broken tip to the anteriormost projection of bone along the medial edge is 3.7 mm. The dorsal wear surface stops ca. 1 mm above the beginning of the dentary bone. It appears that the tip would have extended about to the level of the occlusal surface of the mandibular molars. The lower incisor of Cr. schadenbergi is a slightly paler version of orange-yellow on the anterior surface. At its midpoint, the incisor averages about 2.2 mm wide and about 8.7 – 10.7 mm long (n = 3) from the tip to the anteriormost projection of bone along the medial edge. The dorsal wear surface ends ca. 2.9 mm above the beginning of the dentary bone along the medial edge, and the tip extends to about the level slightly below the occlusal plane of the molars. The lower incisor of Cr. heaneyi is similar in color to Cr. schadenbergi. At its midpoint, the incisor averages about 2.5 mm wide, which is wider than the generally larger Cr. schadenbergi, but we suggest caution in interpreting this as the two specimens available to us were raised in captivity. Figure 4 in Musser and Gordon (1981) shows the lower incisor of Cr. paulus as having a dorsal wear surface that stops about 2 mm above the anteriormost projection of bone along the medial edge, with the tip ending somewhat below the occlusal plane of the molars. On our single maxillary of Cr. ballik (Callao Cave specimen 10389; Fig. 10), the incisive foramina (if) extend posteriorly to about 0.5 mm anterior to the anterior margin of the M 1 alveolus. In two Cr. heaneyi this distance is 1.8 mm and 3.3 mm, and in six Cr. schadenbergi, the range is 1.6 to 3.3 mm, averaging 2.7 mm. In Cr. ballik 10389, the palatal bridge is narrow and longitudinally scored by deep medial palatal grooves. This trait is also seen in Cr. schadenbergi and appears to be a shared character of Luzon Crateromys, but the grooves in Cr. ballik 10389 end about parallel to t 1 (Fig. 10), whereas in Cr. schadenbergi, the grooves flare laterally and become shallow anterior to M 1. On other Crateromys species, the palatal bridge is relatively wider and the medial palatal grooves (pg) are shallower and less conspicuous (Musser et al. 1985; Gonzales and Kennedy 1996). The anterior root of the zygomatic arch (za) appears to originate lateral to t 3 on M 1 (Fig. 10). The molar morphology of Cr. ballik differs from Batomys species on Luzon based on the traits listed above for the genus Crateromys. In comparison to the fossil Batomys described below, Cr. ballik is significantly larger, has higher crowns, and has a more robust dentary (Figs. 8 and 9 A). Overall molar morphology follows that shown in Figs. 2 C and 2 D, except as noted below. Measurements of individual teeth are provided in Table 3. Among Luzon cloud rats, Cr. ballik is intermediate in size between Cr. schadenbergi and B. cagayanensis n. sp. (Fig. 9). In comparison to Cr. schadenbergi, crown length and width of Cr. ballik are smaller (Table 3; Fig. 9). Apart from absolute size of the teeth and the dentary, Cr. ballik differs from Cr. schadenbergi in its m 1 configuration: the anteroconid is not connected to the second loph in Cr. ballik including in specimens with heavily worn teeth (Fig. 5 D; Supplementary Data SD 2 A – D), whereas this is connected in Cr. schadenbergi (Fig. 5 E). In comparison to Cr. heaneyi, crown length and width of Cr. ballik also are smaller (Fig. 9; Table 3). Instead, tooth measurements of Cr. ballik overlap with the range of measurements published for fossil Cr. paulus (Fig. 9; Reyes et al. 2017). M 1 and M 2 of Cr. ballik are differentiated from Cr. paulus (see figure 6 in Musser and Gordon 1981) and Cr. heaneyi (Fig. 4 F) by less prominent labial cusps (t 3 and t 6). The t 6 in M 1 and M 2 of Cr. paulus and Cr. heaneyi is separated by a relatively deep notch from t 5, and this notch is absent in Cr. ballik. On the m 3, the hypoconid and entoconid loph is chevronate on Cr. ballik and Cr. schadenbergi (i. e., both of the Luzon Crateromys), whereas it is fused in a nearly transverse loph in Cr. paulus and in Cr. heaneyi.	en	Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C., Heaney, Lawrence R. (2021): Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini). Journal of Mammalogy 102 (2): 1-22, DOI: 10.1093/jmammal/gyab023
